301 lines
37 KiB
XML
301 lines
37 KiB
XML
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<mods:title id="6E0D08E4D3882659B2DB3EE063F92C70">A Revision Of The Didelphid Marsupial Genus Marmosops, Part 1. Species Of The Subgenus Sciophanes</mods:title>
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<mods:namePart id="AFBF470C6DA9E011D8234F093A5EE6EF">Díaz-Nieto, Juan F.</mods:namePart>
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<mods:namePart id="12DD610062BCA2AEC41D08CB662E21BF">Voss, Robert S.</mods:namePart>
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<treatment id="03A689729833FFF5055D7425D3E2F958" ID-DOI="http://doi.org/10.5281/zenodo.4630915" ID-GBIF-Taxon="180670567" ID-Zenodo-Dep="4630915" LSID="urn:lsid:plazi:treatment:03A689729833FFF5055D7425D3E2F958" httpUri="http://treatment.plazi.org/id/03A689729833FFF5055D7425D3E2F958" lastPageId="39" lastPageNumber="40" pageId="36" pageNumber="37">
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<subSubSection id="C3156BEF9833FFF6055D7425D6FDFAD7" box="[802,1166,1336,1360]" pageId="36" pageNumber="37" type="nomenclature">
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<paragraph id="8BB038649833FFF6055D7425D6FDFAD7" blockId="36.[802,1166,1336,1360]" box="[802,1166,1336,1360]" pageId="36" pageNumber="37">
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<heading id="D0F88F089833FFF6055D7425D6FDFAD7" box="[802,1166,1336,1360]" centered="true" fontSize="9" level="2" pageId="36" pageNumber="37" reason="2">
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<taxonomicName id="4C0F43E79833FFF6055D7425D6FDFAD7" ID-CoL="3Y83S" authority="(Pine, 1981)" baseAuthorityName="Pine" baseAuthorityYear="1981" box="[802,1166,1336,1360]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="handleyi">
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<emphasis id="B97BE4769833FFF6055D7425D672FAD7" box="[802,1025,1336,1360]" italics="true" pageId="36" pageNumber="37">Marmosops handleyi</emphasis>
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(
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<bibRefCitation id="EF9E45959833FFF6026F7424D6F5FAD7" author="Pine, R. H." box="[1040,1158,1336,1360]" pageId="36" pageNumber="37" pagination="55 - 70" refId="ref38708" refString="Pine, R. H. 1981. Reviews of the mouse opossums Marmosa parvidens Tate and Marmosa invicta Goldman (Mammalia: Marsupialia: Didelphidae) with description of a new species. Mammalia 45: 55 - 70." type="journal article" year="1981">Pine, 1981</bibRefCitation>
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)
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3156BEF9833FFF605017476D641FA05" box="[894,1074,1386,1410]" pageId="36" pageNumber="37" type="description">
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<paragraph id="8BB038649833FFF605017476D641FA05" blockId="36.[894,1074,1386,1410]" box="[894,1074,1386,1410]" pageId="36" pageNumber="37">
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<heading id="D0F88F089833FFF605017476D641FA05" box="[894,1074,1386,1410]" centered="true" fontSize="9" level="2" pageId="36" pageNumber="37" reason="2">
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<figureCitation id="133424E19833FFF605017476D196FA05" box="[894,997,1386,1410]" captionStart="FIG" captionStartId="6.[144,185,1651,1672]" captionTargetBox="[170,1223,248,1623]" captionTargetId="figure-9@6.[170,1223,248,1623]" captionTargetPageId="6" captionText="FIG. 2. Marmosops handleyi (above; photographed by J. Díaz-Nieto at Finca Costa Rica, Antioquia, Colombia) and M. parvidens (below; photographed by T. Semedo near Urucará, Amazonas, Brazil). In addition to obvious differences in body pelage color, these species differ in forefoot markings: whereas M. handleyi has dark metacarpal fur that contrasts with the pale fur of the manual digits, M. parvidens has all-pale forefeet." figureDoi="http://doi.org/10.5281/zenodo.4612616" httpUri="https://zenodo.org/record/4612616/files/figure.png" pageId="36" pageNumber="37">Figures 2</figureCitation>
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,
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<figureCitation id="133424E19833FFF6058F7476D67FFA05" box="[1008,1036,1386,1410]" captionStart="FIG" captionStartId="38.[144,186,1730,1751]" captionTargetBox="[205,1187,248,1703]" captionTargetId="figure-9@38.[205,1187,248,1703]" captionTargetPageId="38" captionText="FIG. 18. Dorsal and ventral views of skulls of Marmosops handleyi (left, CTUA 411) and M. invictus (right, USNM 337962). All views about ×3." figureDoi="http://doi.org/10.5281/zenodo.4612668" httpUri="https://zenodo.org/record/4612668/files/figure.png" pageId="36" pageNumber="37">18</figureCitation>
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,
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<figureCitation id="133424E19833FFF602677476D641FA05" box="[1048,1074,1386,1410]" captionStart="FIG" captionStartId="39.[144,186,1392,1413]" captionTargetBox="[308,1085,248,1364]" captionTargetId="figure-107@39.[308,1085,248,1364]" captionTargetPageId="39" captionText="FIG. 19. Lateral views of skulls and mandibles of Marmosops handleyi (above, CTUA 411) and M. invictus (below, USNM 337962). All views about ×3." figureDoi="http://doi.org/10.5281/zenodo.4612670" httpUri="https://zenodo.org/record/4612670/files/figure.png" pageId="36" pageNumber="37">19</figureCitation>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3156BEF9833FFF604AF7481D6EFF99C" pageId="36" pageNumber="37" type="reference_group">
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||
<paragraph id="8BB038649833FFF604AF7481D1FAFA51" blockId="36.[720,1248,1436,1761]" pageId="36" pageNumber="37">
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||
<treatmentCitationGroup id="AB1F1F4A9833FFF604AF7481D1FAFA51" pageId="36" pageNumber="37">
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<treatmentCitation id="0AAE1E759833FFF604AF7481D646FA33" author="Pine, R. H." box="[720,1077,1436,1460]" page="67" pageId="36" pageNumber="37" year="1981">
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<taxonomicName id="4C0F43E79833FFF604AF7481D646FA33" authority="Pine, 1981: 67" authorityName="Pine" authorityPageNumber="67" authorityYear="1981" box="[720,1077,1436,1460]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="handleyi">
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<emphasis id="B97BE4769833FFF604AF7481D1E5FA33" box="[720,918,1436,1460]" italics="true" pageId="36" pageNumber="37">Marmosa handleyi</emphasis>
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||
<bibRefCitation id="EF9E45959833FFF605E27480D646FA33" author="Pine, R. H." box="[925,1077,1436,1460]" pageId="36" pageNumber="37" pagination="55 - 70" refId="ref38708" refString="Pine, R. H. 1981. Reviews of the mouse opossums Marmosa parvidens Tate and Marmosa invicta Goldman (Mammalia: Marsupialia: Didelphidae) with description of a new species. Mammalia 45: 55 - 70." type="journal article" year="1981">Pine, 1981: 67</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
(original description).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BB038649833FFF604AF74FDD6EFF99C" blockId="36.[720,1248,1436,1761]" pageId="36" pageNumber="37">
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||
<treatmentCitationGroup id="AB1F1F4A9833FFF604AF74FDD6EFF99C" pageId="36" pageNumber="37">
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||
<treatmentCitation id="0AAE1E759833FFF604AF74FDD13DF99C" author="Gardner, A. L. & G. K. Creighton" page="4" pageId="36" pageNumber="37" year="1989">
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||
<taxonomicName id="4C0F43E79833FFF604AF74FDD13DF99C" ID-CoL="3Y83S" authority=": Gardner and Creighton, 1989: 4" authorityName=": Gardner and Creighton" authorityPageNumber="4" authorityYear="1989" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="handleyi">
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||
<emphasis id="B97BE4769833FFF604AF74FDD1D8FA7F" box="[720,939,1504,1528]" italics="true" pageId="36" pageNumber="37">Marmosops handleyi</emphasis>
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:
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<bibRefCitation id="EF9E45959833FFF605C774FCD13DF99C" author="Gardner, A. L. & G. K. Creighton" pageId="36" pageNumber="37" pagination="3 - 7" refId="ref36576" refString="Gardner, A. L., and G. K. Creighton. 1989. A new generic name for Tate's (1933) microtarsus group of South American mouse opossums (Marsupialia: Didelphidae). Proceedings of the Biological Society of Washington 102: 3 - 7." type="journal article" year="1989">Gardner and Creighton, 1989: 4</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
(first use of current binomial).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3156BEF9833FFF7048F7738D30AFEF2" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="materials_examined">
|
||
<paragraph id="8BB038649833FFF7048F7738D30AFEF2" blockId="36.[720,1248,1436,1761]" lastBlockId="37.[144,674,250,1629]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
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<materialsCitation id="3B6732399833FFF7048F7738D307FEF2" ID-GBIF-Occurrence="3059923304" collectingDate="1950-06-16" collectionCode="TYPE, MATERIAL" collectorName="Philip Hershkovitz" country="Philippines" elevation="1400" lastPageId="37" lastPageNumber="38" location="Valdivia" pageId="36" pageNumber="37" specimenCode="FMNH 69838, FMNH 70926" specimenCount="2" typeStatus="holotype">
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||
<collectionCode id="ED1EA0A19833FFF6048F7738D1D7F9BB" box="[752,932,1572,1596]" pageId="36" pageNumber="37">TYPE MATERIAL</collectionCode>
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||
: The
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<typeStatus id="54B486C69833FFF605947738D63FF9BB" box="[1003,1100,1572,1596]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
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(by original designation) consists of the skin, skull, and postcranial skeleton of an adult female (
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<specimenCode id="DBA9901F9833FFF602F2777AD167F918" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="36" pageNumber="37" type="Museum">FMNH 69838</specimenCode>
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; original number
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<collectingCountry id="F31878F49833FFF6059B779BD67BF919" box="[996,1032,1671,1694]" name="Philippines" pageId="36" pageNumber="37">PH</collectingCountry>
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4371) collected by
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<collectorName id="26FA5DB29833FFF604AF77B4D1D8F947" box="[720,939,1704,1728]" pageId="36" pageNumber="37">Philip Hershkovitz</collectorName>
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at a site
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<quantity id="4CF795819833FFF6025277B4D61EF947" box="[1069,1133,1704,1728]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="36" pageNumber="37" unit="km" value="9.0">9 km</quantity>
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south of
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<location id="8ED06EBF9833FFF604AF77D5D159F966" LSID="urn:lsid:plazi:treatment:03A689729833FFF5055D7425D3E2F958:8ED06EBF9833FFF604AF77D5D159F966" box="[720,810,1737,1761]" country="Philippines" name="Valdivia" pageId="36" pageNumber="37">Valdivia</location>
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||
(fig. 20: locality 24) in the Colombian department of Antioquia at an elevation of
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<quantity id="4CF795819832FFF7041371E6D2D5FEB5" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="37" pageNumber="38" unit="m" value="1400.0">
|
||
<elevation id="0022DF579832FFF7041371E6D2D5FEB5" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="37" pageNumber="38" unit="m" value="1400.0">1400 m</elevation>
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||
</quantity>
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||
on
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<date id="FFB11EA49832FFF706A77007D302FEB4" box="[216,369,283,307]" pageId="37" pageNumber="38" value="1950-06-16">
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||
<collectingDate id="EFF5E74C9832FFF706A77007D302FEB4" box="[216,369,283,307]" pageId="37" pageNumber="38" value="1950-06-16">16 June 1950</collectingDate>
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||
</date>
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||
. In addition,
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||
<bibRefCitation id="EF9E45959832FFF7046F7007D0D3FEB4" author="Pine, R. H." box="[528,672,283,307]" pageId="37" pageNumber="38" pagination="55 - 70" refId="ref38708" refString="Pine, R. H. 1981. Reviews of the mouse opossums Marmosa parvidens Tate and Marmosa invicta Goldman (Mammalia: Marsupialia: Didelphidae) with description of a new species. Mammalia 45: 55 - 70." type="journal article" year="1981">Pine’s (1981)</bibRefCitation>
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||
original material included an adult male
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||
<typeStatus id="54B486C69832FFF704197020D2CCFEF2" pageId="37" pageNumber="38" type="paratype">paratype</typeStatus>
|
||
(
|
||
<specimenCode id="DBA9901F9832FFF706B17041D31EFEF2" box="[206,365,349,373]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="37" pageNumber="38" type="Museum">FMNH 70926</specimenCode>
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)
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3156BEF9832FFF706CF7062D394FD66" pageId="37" pageNumber="38" type="distribution">
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||
<paragraph id="8BB038649832FFF706CF7062D394FD66" blockId="37.[144,674,250,1629]" pageId="37" pageNumber="38">
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||
DISTRIBUTION, HABITATS, AND SYMPATRY: Examined specimens of
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<taxonomicName id="4C0F43E79832FFF707E670BCD00BFE30" authorityName=": Gardner and Creighton" authorityYear="1989" box="[409,632,415,439]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="handleyi">
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||
<emphasis id="B97BE4769832FFF707E670BCD00BFE30" box="[409,632,415,439]" italics="true" pageId="37" pageNumber="38">Marmosops handleyi</emphasis>
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||
</taxonomicName>
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||
are all from the northern terminus of the Cordillera Central (central Andes) of
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||
<collectingCountry id="F31878F49832FFF707CD70FDD06CFE7E" box="[434,543,481,505]" name="Colombia" pageId="37" pageNumber="38">Colombia</collectingCountry>
|
||
, where they have been collected in humid premontane and montane rain forest at elevations ranging from
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<quantity id="4CF795819832FFF706EF7358D333FDDB" box="[144,320,580,604]" metricMagnitude="3" metricUnit="m" metricValue="1.675" metricValueMax="1.95" metricValueMin="1.4" pageId="37" pageNumber="38" unit="m" value="1675.0" valueMax="1950.0" valueMin="1400.0">1400 to 1950 m</quantity>
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||
(fig. 20). This species has been collected sympatrically with
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||
<taxonomicName id="4C0F43E79832FFF707AA737AD03AFDFA" authorityName="Díaz-Nieto & Voss" authorityYear="2016" box="[469,585,613,637]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="chucha">
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||
<emphasis id="B97BE4769832FFF707AA737AD03AFDFA" box="[469,585,613,637]" italics="true" pageId="37" pageNumber="38">M. chucha</emphasis>
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||
</taxonomicName>
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(a new species of the Bishopi Group; see below) and with
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<taxonomicName id="4C0F43E79832FFF706B773B4D346FD38" baseAuthorityName="Thomas" baseAuthorityYear="1900" box="[200,309,680,703]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="caucae">
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||
<emphasis id="B97BE4769832FFF706B773B4D346FD38" box="[200,309,680,703]" italics="true" pageId="37" pageNumber="38">M. caucae</emphasis>
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</taxonomicName>
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(a member of the nominotypical subgenus; Díaz-N. et al., 2011).
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="C3156BEF9832FFF706CF73F6D1E5F9F8" pageId="37" pageNumber="38" type="description">
|
||
<paragraph id="8BB038649832FFF706CF73F6D3FDFA94" blockId="37.[144,674,250,1629]" pageId="37" pageNumber="38">
|
||
DESCRIPTION: Body pelage dark brown to chestnut brown middorsally, indistinctly paler laterally, about
|
||
<quantity id="4CF795819832FFF707437230D3DBFCC4" box="[316,424,812,836]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="37" pageNumber="38" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
|
||
long at midback, and somewhat woolly in texture; ventral pelage superficially whitish, but hairs of throat, chest, and abdomen uniformly gray based (only the apex of the chin, the oral margins, the scrotum, and mammary region consistently have selfwhite fur).
|
||
<superScript id="7C7A952C9832FFF7077672ECD361FB86" attach="left" box="[265,274,1008,1025]" fontSize="7" pageId="37" pageNumber="38">9</superScript>
|
||
Metacarpals covered dorsally with dark hairs (contrasting in color with the pale manual digits); lateral carpal tubercles bladelike. Mammary formula 3–1–3 =
|
||
<quantity id="4CF795819832FFF707A7754AD076FBEA" box="[472,517,1109,1133]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="37" pageNumber="38" unit="in" value="7.0">7 in</quantity>
|
||
all examined females with conspicuous teats. Tail longer than combined length of head and body (mean LT/ HBL × 100 = 125%), dorsal caudal surface dark gray but somewhat paler distally; ventral caudal surface distinctly paler.
|
||
</paragraph>
|
||
<paragraph id="8BB038649832FFF706CF7400D0E7F9DA" blockId="37.[144,674,250,1629]" pageId="37" pageNumber="38">Nasal bones long (extending well behind the lacrimals) and much wider posteriorly than anteriorly (laterally expanded at the maxillary-frontal suture). Upper lacrimal foramen not visible in lateral view, but lower foramen laterally exposed; zygomatic process of squamosal broadly overlapped dorsally by the jugal. Palatine fenestrae present (usually as a pair of large rounded perforations). Dorsolateral margin of ethmoid foramen with equal contribution of frontal and orbitosphenoid.</paragraph>
|
||
<paragraph id="8BB038649832FFF706EF778DD002F883" blockId="37.[144,673,1681,1797]" pageId="37" pageNumber="38">
|
||
<superScript id="7C7A952C9832FFF706EF778DD2E4F91A" attach="left" box="[144,151,1681,1693]" fontSize="5" pageId="37" pageNumber="38">9</superScript>
|
||
<bibRefCitation id="EF9E45959832FFF706DF778DD355F923" author="Pine, R. H." box="[160,294,1681,1701]" pageId="37" pageNumber="38" pagination="55 - 70" refId="ref38708" refString="Pine, R. H. 1981. Reviews of the mouse opossums Marmosa parvidens Tate and Marmosa invicta Goldman (Mammalia: Marsupialia: Didelphidae) with description of a new species. Mammalia 45: 55 - 70." type="journal article" year="1981">Pine (1981: 68)</bibRefCitation>
|
||
described the dorsal pelage as “fuscous,” a grayer shade than we observed in fresh skins. Unfortunately, none were compared by us with color swatches in
|
||
<bibRefCitation id="EF9E45959832FFF7042A77DDD2B9F96A" author="Ridgway, R." pageId="37" pageNumber="38" refId="ref38915" refString="Ridgway, R. 1912. Color standards and color nomenclature. Washington, DC: Published by the author." type="book" year="1912">Ridgway (1912)</bibRefCitation>
|
||
. One specimen (CTUA 415, a subadult male) has a 5 ×
|
||
<quantity id="4CF795819832FFF706EF77EDD2BEF882" box="[144,205,1777,1797]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="37" pageNumber="38" unit="mm" value="15.0">15 mm</quantity>
|
||
midventral streak of self-white abdominal hairs.
|
||
</paragraph>
|
||
<paragraph id="8BB038649832FFF7048F71E6D1C5FD19" blockId="37.[720,1248,250,1795]" pageId="37" pageNumber="38">Upper canine (C1) short in both sexes but sexually dimorphic in shape: male C1 with posterior accessory cusp only, female C1 with both anterior and posterior accessory cusps. Upper third molar (M3) anterolabial cingulum discontinuous with preprotocrista (anterior cingulum incomplete). Lower canine (c1) premolariform (procumbent, with posterior accessory cusp) and small, subequal in height to p1; c1 anterolingual accessory cusp present. Entoconid of m1 subequal in height to adjacent m2 paraconid; unworn m4 talonid with three distinct cusps.</paragraph>
|
||
<paragraph id="8BB038649832FFF7048F73B4D627FC85" blockId="37.[720,1248,250,1795]" pageId="37" pageNumber="38">
|
||
COMPARISONS: Comparisons of
|
||
<taxonomicName id="4C0F43E79832FFF7021D73B4D15DFD66" authorityName=": Gardner and Creighton" authorityYear="1989" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="handleyi">
|
||
<emphasis id="B97BE4769832FFF7021D73B4D15DFD66" italics="true" pageId="37" pageNumber="38">Marmosops handleyi</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C0F43E79832FFF7050573D5D1AAFD67" baseAuthorityName="Allen and Chapman" baseAuthorityYear="1897" box="[890,985,713,736]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="carri">
|
||
<emphasis id="B97BE4769832FFF7050573D5D1AAFD67" box="[890,985,713,736]" italics="true" pageId="37" pageNumber="38">M. carri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0F43E79832FFF7026373D5D6D2FD66" authorityName=": Gardner and Creighton" authorityYear="1989" box="[1052,1185,713,737]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="fuscatus">
|
||
<emphasis id="B97BE4769832FFF7026373D5D6D2FD66" box="[1052,1185,713,737]" italics="true" pageId="37" pageNumber="38">M. fuscatus</emphasis>
|
||
</taxonomicName>
|
||
have already been described (see above).
|
||
</paragraph>
|
||
<paragraph id="8BB038649832FFF7048F7217D1E5F9F8" blockId="37.[720,1248,250,1795]" pageId="37" pageNumber="38">
|
||
Although
|
||
<taxonomicName id="4C0F43E79832FFF7050D7217D618FCA4" authorityName=": Gardner and Creighton" authorityYear="1989" box="[882,1131,779,803]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="handleyi">
|
||
<emphasis id="B97BE4769832FFF7050D7217D618FCA4" box="[882,1131,779,803]" italics="true" pageId="37" pageNumber="38">Marmosops handleyi</emphasis>
|
||
</taxonomicName>
|
||
averages slightly larger than
|
||
<taxonomicName id="4C0F43E79832FFF705DC7231D668FCC3" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[931,1051,813,836]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="invictus">
|
||
<emphasis id="B97BE4769832FFF705DC7231D668FCC3" box="[931,1051,813,836]" italics="true" pageId="37" pageNumber="38">M. invictus</emphasis>
|
||
</taxonomicName>
|
||
in most same-sex comparisons of external dimensions (tables 5, 6), sample sizes for both species are small, and it seems unlikely that any external measurement will prove to be diagnostic as more material becomes available. A more consistent external difference between these species is pelage color (distinctly brownish in
|
||
<taxonomicName id="4C0F43E79832FFF70593750FD638FBAC" authorityName=": Gardner and Creighton" authorityYear="1989" box="[1004,1099,1043,1067]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="handleyi">
|
||
<emphasis id="B97BE4769832FFF70593750FD638FBAC" box="[1004,1099,1043,1067]" italics="true" pageId="37" pageNumber="38">handleyi</emphasis>
|
||
</taxonomicName>
|
||
versus dark grayish brown to blackish gray in
|
||
<taxonomicName id="4C0F43E79832FFF7022E7529D6D7FBCB" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[1105,1188,1077,1100]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="invictus">
|
||
<emphasis id="B97BE4769832FFF7022E7529D6D7FBCB" box="[1105,1188,1077,1100]" italics="true" pageId="37" pageNumber="38">invictus</emphasis>
|
||
</taxonomicName>
|
||
) and fur length (
|
||
<quantity id="4CF795819832FFF70529754AD1B5FBEA" box="[854,966,1110,1134]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="37" pageNumber="38" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
|
||
middorsally in
|
||
<taxonomicName id="4C0F43E79832FFF702FD7549D693FBEA" authorityName=": Gardner and Creighton" authorityYear="1989" box="[1154,1248,1109,1133]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="handleyi">
|
||
<emphasis id="B97BE4769832FFF702FD7549D693FBEA" box="[1154,1248,1109,1133]" italics="true" pageId="37" pageNumber="38">handleyi</emphasis>
|
||
</taxonomicName>
|
||
versus
|
||
<quantity id="4CF795819832FFF7055D756AD1F6FB09" box="[802,901,1142,1166]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="37" pageNumber="38" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C0F43E79832FFF705CB756BD67AFB09" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[948,1033,1143,1166]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="invictus">
|
||
<emphasis id="B97BE4769832FFF705CB756BD67AFB09" box="[948,1033,1143,1166]" italics="true" pageId="37" pageNumber="38">invictus</emphasis>
|
||
</taxonomicName>
|
||
). The two species broadly overlap in craniodental measurements, none of which, therefore, is useful for identification. Instead,
|
||
<taxonomicName id="4C0F43E79832FFF7051975C6D194FB75" authorityName=": Gardner and Creighton" authorityYear="1989" box="[870,999,1242,1266]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="handleyi">
|
||
<emphasis id="B97BE4769832FFF7051975C6D194FB75" box="[870,999,1242,1266]" italics="true" pageId="37" pageNumber="38">M. handleyi</emphasis>
|
||
</taxonomicName>
|
||
is more readily distinguished from
|
||
<taxonomicName id="4C0F43E79832FFF7050175E7D671FA95" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[894,1026,1275,1298]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="invictus">
|
||
<emphasis id="B97BE4769832FFF7050175E7D671FA95" box="[894,1026,1275,1298]" italics="true" pageId="37" pageNumber="38">M. invictus</emphasis>
|
||
</taxonomicName>
|
||
by the consistent presence of palatine fenestrae (consistently absent in
|
||
<taxonomicName id="4C0F43E79832FFF7053F7422D1CFFAD2" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[832,956,1342,1365]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="invictus">
|
||
<emphasis id="B97BE4769832FFF7053F7422D1CFFAD2" box="[832,956,1342,1365]" italics="true" pageId="37" pageNumber="38">M. invictus</emphasis>
|
||
</taxonomicName>
|
||
), by the presence of both anterior and posterior accessory C1 cusps in females (females usually have only a posterior accessory C1 cusp in
|
||
<taxonomicName id="4C0F43E79832FFF705BA74BDD64CFA3F" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[965,1087,1441,1464]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="invictus">
|
||
<emphasis id="B97BE4769832FFF705BA74BDD64CFA3F" box="[965,1087,1441,1464]" italics="true" pageId="37" pageNumber="38">M. invictus</emphasis>
|
||
</taxonomicName>
|
||
), by lacking a complete M3 anterior cingulum (M3 has a narrowly complete anterior cingulum in
|
||
<taxonomicName id="4C0F43E79832FFF7020474FFD083F99C" baseAuthorityName="Goldman" baseAuthorityYear="1912" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="invictus">
|
||
<emphasis id="B97BE4769832FFF7020474FFD083F99C" italics="true" pageId="37" pageNumber="38">M. invictus</emphasis>
|
||
</taxonomicName>
|
||
), and by having an m1 entoconid that is subequal in height to the adjacent m2 paraconid (the m1 entoconid is shorter than the m2 paraconid in
|
||
<taxonomicName id="4C0F43E79832FFF7054C777BD1F5F9F9" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[819,902,1639,1662]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="invictus">
|
||
<emphasis id="B97BE4769832FFF7054C777BD1F5F9F9" box="[819,902,1639,1662]" italics="true" pageId="37" pageNumber="38">invictus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3156BEF9832FFF5048F7794D3E2F958" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" type="discussion">
|
||
<paragraph id="8BB038649832FFF5048F7794D3B1F9BE" blockId="37.[720,1248,250,1795]" lastBlockId="39.[144,672,1470,1759]" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">
|
||
REMARKS:
|
||
<taxonomicName id="4C0F43E79832FFF705187794D634F918" authorityName=": Gardner and Creighton" authorityYear="1989" box="[871,1095,1671,1695]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="handleyi">
|
||
<emphasis id="B97BE4769832FFF705187794D634F918" box="[871,1095,1671,1695]" italics="true" pageId="37" pageNumber="38">Marmosops handleyi</emphasis>
|
||
</taxonomicName>
|
||
had not been collected for 60 years before it was rediscovered and redescribed by Díaz-N. et al. (2011), whose report should be consulted for additional mor- phological, geographical, and ecological information. Using the dichotomous key in
|
||
<bibRefCitation id="EF9E45959830FFF5043B74C3D30FF99E" author="Gardner, A. L. & G. K. Creighton" pageId="39" pageNumber="40" pagination="61 - 74" refId="ref36627" refString="Gardner, A. L., and G. K. Creighton. 2008 (" 2007 "). Genus Marmosops Matschie, 1916. In A. L. Gardner (editor), Mammals of South America, vol. 1: marsupials, xenarthrans, shrews, and bats: 61 - 74. Chicago: University of Chicago Press." type="book chapter" year="2008">Gardner and Creighton (2008)</bibRefCitation>
|
||
, this species would be misdidentified as
|
||
<taxonomicName id="4C0F43E79830FFF50754773ED3CFF9BE" baseAuthorityName="Tate" baseAuthorityYear="1931" box="[299,444,1570,1593]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="juninensis">
|
||
<emphasis id="B97BE4769830FFF50754773ED3CFF9BE" box="[299,444,1570,1593]" italics="true" pageId="39" pageNumber="40">M. juninensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF7068EC9831FFF406EF77DED070F972" ID-DOI="http://doi.org/10.5281/zenodo.4612668" ID-Zenodo-Dep="4612668" httpUri="https://zenodo.org/record/4612668/files/figure.png" pageId="38" pageNumber="39" startId="38.[144,186,1730,1751]" targetBox="[205,1187,248,1703]" targetPageId="38">
|
||
<paragraph id="8BB038649831FFF406EF77DED070F972" blockId="38.[144,1248,1728,1781]" pageId="38" pageNumber="39">
|
||
FIG. 18. Dorsal and ventral views of skulls of
|
||
<taxonomicName id="4C0F43E79831FFF4041477DCD14FF95F" authorityName=": Gardner and Creighton" authorityYear="1989" box="[619,828,1728,1752]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="handleyi">
|
||
<emphasis id="B97BE4769831FFF4041477DCD14FF95F" box="[619,828,1728,1752]" italics="true" pageId="38" pageNumber="39">Marmosops handleyi</emphasis>
|
||
</taxonomicName>
|
||
(left, CTUA 411) and
|
||
<taxonomicName id="4C0F43E79831FFF4025877DCD6EBF950" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[1063,1176,1728,1751]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="invictus">
|
||
<emphasis id="B97BE4769831FFF4025877DCD6EBF950" box="[1063,1176,1728,1751]" italics="true" pageId="38" pageNumber="39">M. invictus</emphasis>
|
||
</taxonomicName>
|
||
(right, USNM 337962). All views about ×3.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7068EC9830FFF506EF746CD020FA24" ID-DOI="http://doi.org/10.5281/zenodo.4612670" ID-Zenodo-Dep="4612670" httpUri="https://zenodo.org/record/4612670/files/figure.png" pageId="39" pageNumber="40" startId="39.[144,186,1392,1413]" targetBox="[308,1085,248,1364]" targetPageId="39">
|
||
<paragraph id="8BB038649830FFF506EF746CD020FA24" blockId="39.[144,1248,1390,1443]" pageId="39" pageNumber="40">
|
||
FIG. 19. Lateral views of skulls and mandibles of
|
||
<taxonomicName id="4C0F43E79830FFF504EB7472D115FA01" authorityName=": Gardner and Creighton" authorityYear="1989" box="[660,870,1390,1414]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="handleyi">
|
||
<emphasis id="B97BE4769830FFF504EB7472D115FA01" box="[660,870,1390,1414]" italics="true" pageId="39" pageNumber="40">Marmosops handleyi</emphasis>
|
||
</taxonomicName>
|
||
(above, CTUA 411) and
|
||
<taxonomicName id="4C0F43E79830FFF502117472D693FA02" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[1134,1248,1390,1413]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="invictus">
|
||
<emphasis id="B97BE4769830FFF502117472D693FA02" box="[1134,1248,1390,1413]" italics="true" pageId="39" pageNumber="40">M. invictus</emphasis>
|
||
</taxonomicName>
|
||
(below, USNM 337962). All views about ×3.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB038649830FFF506CF775FD3E2F958" blockId="39.[144,672,1470,1759]" pageId="39" pageNumber="40">
|
||
SPECIMENS EXAMINED (
|
||
<emphasis id="B97BE4769830FFF507CB775FD3B4F9DD" box="[436,455,1603,1626]" italics="true" pageId="39" pageNumber="40">N</emphasis>
|
||
= 10):
|
||
<emphasis id="B97BE4769830FFF50469775ED0D3F9DD" bold="true" box="[534,672,1602,1626]" pageId="39" pageNumber="40">Colombia—</emphasis>
|
||
<emphasis id="B97BE4769830FFF506EF7778D370F9FC" box="[144,259,1636,1659]" italics="true" pageId="39" pageNumber="40">Antioquia,</emphasis>
|
||
Finca
|
||
<collectingCountry id="F31878F49830FFF507287778D3A0F9FB" box="[343,467,1636,1660]" name="Costa Rica" pageId="39" pageNumber="40">Costa Rica</collectingCountry>
|
||
(CTUA 410–412, 433); Finca El Bosque (CTUA 413, 414), Finca Villa Nueva (CTUA 415, 416),
|
||
<quantity id="4CF795819830FFF5079277BAD05BF939" box="[493,552,1702,1726]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="40" unit="km" value="9.0">9 km</quantity>
|
||
S Valdivia (FMNH 69823, 69838).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |