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<document id="DC7C00499C7B6D0AE6CCD8B385F31952" ID-DOI="10.11646/zootaxa.2104.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5315428" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629623837903" checkinUser="felipe" docAuthor="Barnich, Ruth &amp; Fiege, Dieter" docDate="2009" docId="03A61979C237FFCEFF18FB13FC031556" docLanguage="en" docName="zt02104p076.pdf" docOrigin="Zootaxa 2104 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2104.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Harmothoe aspera" docType="treatment" docVersion="5" lastPageNumber="24" masterDocId="FF9F6101C223FFD9FF8FFD4EFFA41036" masterDocTitle="Revision of the genus Harmothoe Kinberg, 1856 (Polychaeta: Polynoidae) in the Northeast Atlantic" masterLastPageNumber="76" masterPageNumber="1" pageNumber="21" updateTime="1699085409460" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="4CD9702A84A9882EE26F38E6434FD1A9">Revision of the genus Harmothoe Kinberg, 1856 (Polychaeta: Polynoidae) in the Northeast Atlantic</mods:title>
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<mods:namePart id="F318DE0160464EC3CB21C99B14245E89">Barnich, Ruth</mods:namePart>
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<paragraph id="8BB0A86FC237FFCDFF18FB13FD9E1641" blockId="20.[151,570,1629,1690]" box="[151,570,1629,1655]" pageId="20" pageNumber="21">
<heading id="D0F81F03C237FFCDFF18FB13FD9E1641" bold="true" box="[151,570,1629,1655]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<taxonomicName id="4C0FD3ECC237FFCDFF18FB13FD9E1641" ID-CoL="6LCF2" authority="(Hansen, 1879)" baseAuthorityName="Hansen" baseAuthorityYear="1879" box="[151,570,1629,1655]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="21" phylum="Annelida" rank="species" species="aspera">
<emphasis id="B97B747DC237FFCDFF18FB13FD9E1641" bold="true" box="[151,570,1629,1655]" pageId="20" pageNumber="21">
<emphasis id="B97B747DC237FFCDFF18FB13FEDC1641" bold="true" box="[151,376,1629,1655]" italics="true" pageId="20" pageNumber="21">Harmothoe aspera</emphasis>
(
<bibRefCitation id="EF9ED59EC237FFCDFE08FB13FD961641" author="Hansen, G. A." box="[391,562,1629,1655]" pageId="20" pageNumber="21" pagination="1 - 14" refId="ref37769" refString="Hansen, G. A. (1879) Annelider fra den norske Nordhavexpedition i 1876. Nyt Magazin for Naturvidenskaberne, Oslo, 24 (1), 1 - 14." type="journal article" year="1879">Hansen, 1879</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C315FBE4C237FFCDFF18FBCEFEFB16AC" box="[151,351,1664,1690]" pageId="20" pageNumber="21" type="description">
<paragraph id="8BB0A86FC237FFCDFF18FBCEFEFB16AC" blockId="20.[151,570,1629,1690]" box="[151,351,1664,1690]" pageId="20" pageNumber="21">
(
<figureCitation id="1334B4EAC237FFCDFF10FBCEFF5616AC" box="[159,242,1664,1690]" captionStart="FIGURE 2" captionStartId="10.[151,257,1783,1807]" captionTargetBox="[279,1395,201,1750]" captionTargetId="figure-136@10.[151,1436,179,1776]" captionTargetPageId="10" captionText="FIGURE 2. Identification characters of Leucia violacea, Harmothoe fraserthomsoni, Leucia nivea, Acanthicolepis asperrima, Harmothoe aspera, and H. serrata." figureDoi="http://doi.org/10.5281/zenodo.5315434" httpUri="https://zenodo.org/record/5315434/files/figure.png" pageId="20" pageNumber="21">Figs. 2</figureCitation>
,
<figureCitation id="1334B4EAC237FFCDFF70FBCEFEF316AC" box="[255,343,1664,1690]" captionStart="FIGURE 10" captionStartId="22.[151,257,1833,1857]" captionTargetBox="[209,1390,191,1803]" captionTargetId="figure-125@22.[151,1436,169,1822]" captionTargetPageId="22" captionText="FIGURE 10. Harmothoe aspera (small syntype, ZMBN 2000). (A) anterior end, palps and style of median antenna missing, left dorsal tentacular cirrus regenerating; (B) right anterior elytron from unknown segment; (C) detail of posterior margin of same; (D) right cirrigerous parapodium from unknown segment, posterior view, style of dorsal cirrus missing; (E) short notochaeta; (F) long notochaeta; (G) tip of same; (H) middle neurochaeta; (I) tip of same; (J) distal part of upper neurochaeta; (K) lower neurochaeta. (Scale: A, B = 2 mm; C, E, F, H, J, K = 250 µm; D = 1 mm; G = 50 µm; I = 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5315458" httpUri="https://zenodo.org/record/5315458/files/figure.png" pageId="20" pageNumber="21">10AK</figureCitation>
)
</paragraph>
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<subSubSection id="C315FBE4C237FFCDFF18FB83FCE31712" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="8BB0A86FC237FFCDFF18FB83FC1516D2" blockId="20.[151,1436,1741,1828]" box="[151,945,1741,1764]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="AB1F8F41C237FFCDFF18FB83FC1516D2" box="[151,945,1741,1764]" pageId="20" pageNumber="21">
<treatmentCitation id="0AAE8E7EC237FFCDFF18FB83FE5616D2" author="Hansen, G. A." box="[151,498,1741,1764]" page="1" pageId="20" pageNumber="21" year="1879">
<taxonomicName id="4C0FD3ECC237FFCDFF18FB83FE5616D2" ID-CoL="7862V" authority="Hansen, 1879: 1" authorityName="Hansen" authorityPageNumber="1" authorityYear="1879" box="[151,498,1741,1764]" class="Polychaeta" family="Polynoidae" genus="Polynoe" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="21" phylum="Annelida" rank="species" species="aspera">
<emphasis id="B97B747DC237FFCDFF18FB83FE9916D2" box="[151,317,1741,1764]" italics="true" pageId="20" pageNumber="21">Polynoe aspera</emphasis>
<bibRefCitation id="EF9ED59EC237FFCDFECBFB83FE5616D2" author="Hansen, G. A." box="[324,498,1741,1764]" pageId="20" pageNumber="21" pagination="1 - 14" refId="ref37769" refString="Hansen, G. A. (1879) Annelider fra den norske Nordhavexpedition i 1876. Nyt Magazin for Naturvidenskaberne, Oslo, 24 (1), 1 - 14." type="journal article" year="1879">Hansen, 1879: 1</bibRefCitation>
</taxonomicName>
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, pl.1;
<treatmentCitation id="0AAE8E7EC237FFCDFDB9FB83FD5516D2" author="Hansen, G. A." box="[566,753,1741,1764]" page="5" pageId="20" pageNumber="21" year="1882">
<bibRefCitation id="EF9ED59EC237FFCDFDB9FB83FD7316D2" author="Hansen, G. A." box="[566,727,1741,1764]" pageId="20" pageNumber="21" pagination="1 - 53" refId="ref37832" refString="Hansen, G. A. (1882) Annelida. The Norwegian North-Atlantic Expedition 1876 - 1878. Zoology, 3 (2), 1 - 53." type="journal article" year="1882">Hansen (1882)</bibRefCitation>
: 5
</treatmentCitation>
, pl. 2 figs. 1015.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB0A86FC237FFCDFF18FBA3FCE31712" blockId="20.[151,1436,1741,1828]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="AB1F8F41C237FFCDFF18FBA3FCE31712" pageId="20" pageNumber="21">
<treatmentCitation id="0AAE8E7EC237FFCDFF18FBA3FD641732" author="Hartmann-Schroder, G." box="[151,704,1773,1796]" page="44" pageId="20" pageNumber="21" year="1996">
<taxonomicName id="4C0FD3ECC237FFCDFF18FBA3FD3C1732" ID-CoL="6LCF2" authority=": Hartmann-Schroder (1996)" authorityName=": Hartmann-Schroder" authorityYear="1996" box="[151,664,1773,1796]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="21" phylum="Annelida" rank="species" species="aspera">
<emphasis id="B97B747DC237FFCDFF18FBA3FEC51732" box="[151,353,1773,1796]" italics="true" pageId="20" pageNumber="21">Harmothoe aspera</emphasis>
:
<bibRefCitation id="EF9ED59EC237FFCDFEFEFBA3FD3C1732" author="Hartmann-Schroder, G." box="[369,664,1773,1796]" pageId="20" pageNumber="21" pagination="1 - 648" refId="ref37937" refString="Hartmann-Schroder, G. (1996) Annelida, Borstenwurmer, Polychaeta. Die Tierwelt Deutschlands. 2. neubearbeitete Auflage, 58, 1 - 648." type="journal article" year="1996">Hartmann-Schröder (1996)</bibRefCitation>
</taxonomicName>
: 44
</treatmentCitation>
[part; not
<taxonomicName id="4C0FD3ECC237FFCDFCBAFBA3FBDF1732" ID-CoL="3JRGS" authority="(Moore, 1902)" baseAuthorityName="Moore" baseAuthorityYear="1902" box="[821,1147,1773,1796]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="21" phylum="Annelida" rank="species" species="multisetosa">
<emphasis id="B97B747DC237FFCDFCBAFBA3FC751732" box="[821,977,1773,1796]" italics="true" pageId="20" pageNumber="21">H. multisetosa</emphasis>
(
<bibRefCitation id="EF9ED59EC237FFCDFC6EFBA3FBD71732" author="Moore, J. P." box="[993,1139,1773,1796]" pageId="20" pageNumber="21" pagination="258 - 278" refId="ref39353" refString="Moore, J. P. (1902) Description of some new Polynoidae, with a list of other Polychaeta from North Greenland waters. Proceedings of the Academy of Natural Sciences Philadelphia, 258 - 278." type="book chapter" year="1902">Moore, 1902</bibRefCitation>
)
</taxonomicName>
];
<treatmentCitation id="0AAE8E7EC237FFCDFB00FBA3FF5B1712" author="Barnich, R. &amp; Fiege, D." httpUri="http://treatment.plazi.org/id/03DE3D65F440C6219038A4A54C0C3ABE" page="1915" pageId="20" pageNumber="21" year="2000">
<bibRefCitation id="EF9ED59EC237FFCDFB00FBA3FA3D1732" author="Barnich, R. &amp; Fiege, D." box="[1167,1433,1773,1796]" pageId="20" pageNumber="21" pagination="1889 - 1938" refId="ref36372" refString="Barnich, R. &amp; Fiege, D. (2000) Revision of the Mediterranean species of Harmothoe Kinberg, 1856 and Lagisca Malmgren, 1865 (Polychaeta, Polynoidae, Polynoinae) with descriptions of a new genus and a new species. Journal of Natural History, 34, 1889 - 1938." type="journal article" year="2000">Barnich &amp; Fiege (2000)</bibRefCitation>
: 1915
</treatmentCitation>
, fig. 13AD;
<treatmentCitation id="0AAE8E7EC237FFCDFE1EFA43FD1F1712" author="Barnich, R. &amp; Fiege, D." box="[401,699,1805,1828]" page="37" pageId="20" pageNumber="21" year="2003">
<bibRefCitation id="EF9ED59EC237FFCDFE1EFA43FD301712" author="Barnich, R. &amp; Fiege, D." box="[401,660,1805,1828]" pageId="20" pageNumber="21" pagination="1 - 167" refId="ref36428" refString="Barnich, R. &amp; Fiege, D. (2003) The Aphroditoidea (Annelida, Polychaeta) of the Mediterranean Sea. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 559, 1 - 167." type="journal article" year="2003">Barnich &amp; Fiege (2003)</bibRefCitation>
: 37
</treatmentCitation>
, fig. 14AD.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C315FBE4C237FFCEFF18FA1AFC031556" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8BB0A86FC237FFCDFF18FA1AFDB417A2" blockId="20.[151,1436,1876,1940]" pageId="20" pageNumber="21">
<emphasis id="B97B747DC237FFCDFF18FA1AFDBC175B" box="[151,536,1876,1901]" italics="true" pageId="20" pageNumber="21">
Type material.
<taxonomicName id="4C0FD3ECC237FFCDFEDAFA1AFDBC175B" authorityName="Hansen" authorityYear="1879" box="[341,536,1876,1901]" class="Polychaeta" family="Polynoidae" genus="Polynoe" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="21" phylum="Annelida" rank="species" species="aspera">Polynoe aspera</taxonomicName>
</emphasis>
:
<materialsCitation id="3B67A232C237FFCDFDA4FA1AFDA817A2" collectionCode="ZMBN" elevation="547" latitude="64.6" longLatPrecision="1007" longitude="-10.366667" pageId="20" pageNumber="21" specimenCode="ZMBN 2000" specimenCount="2" typeStatus="syntype">
<specimenCount id="9D0963E6C237FFCDFDA4FA1AFD171758" box="[555,691,1876,1902]" pageId="20" pageNumber="21" type="generic" typeStatus="syntypes">2 syntypes</specimenCount>
,
<specimenCode id="DBA90014C237FFCDFD4DFA1AFCC01758" box="[706,868,1876,1902]" collectionCode="ZMBN" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="20" pageNumber="21">ZMBN 2000</specimenCode>
, Norske Nordhavsexp. 1876, St. 48,
<geoCoordinate id="EE3BCEA8C237FFCDFAB9FA1AFA381758" box="[1334,1436,1876,1902]" degrees="64" direction="north" minutes="36" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="64.6">64°36'N</geoCoordinate>
<geoCoordinate id="EE3BCEA8C237FFCDFF18FA34FF5A17A2" box="[151,254,1914,1940]" degrees="10" direction="west" minutes="22" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="-10.366667">10°22'W</geoCoordinate>
,
<quantity id="4CF7058AC237FFCDFE85FA34FEF317A2" box="[266,343,1914,1940]" metricMagnitude="2" metricUnit="m" metricValue="5.47" pageId="20" pageNumber="21" unit="m" value="547.0">
<elevation id="00224F5CC237FFCDFE85FA34FEF317A2" box="[266,343,1914,1940]" metricMagnitude="2" metricUnit="m" metricValue="5.47" pageId="20" pageNumber="21" unit="m" value="547.0">547 m</elevation>
</quantity>
, sand and ooze
</materialsCitation>
.
</paragraph>
<caption id="DF70F8E7C236FFCCFF18FA74FE6817ED" ID-DOI="http://doi.org/10.5281/zenodo.5315456" ID-Zenodo-Dep="5315456" httpUri="https://zenodo.org/record/5315456/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,257,1850,1874]" targetBox="[222,1375,190,1801]" targetPageId="21">
<paragraph id="8BB0A86FC236FFCCFF18FA74FE6817ED" blockId="21.[151,1436,1850,2011]" pageId="21" pageNumber="22">
<emphasis id="B97B747DC236FFCCFF18FA74FEBA1764" bold="true" box="[151,286,1850,1874]" pageId="21" pageNumber="22">FIGURE 9.</emphasis>
<taxonomicName id="4C0FD3ECC236FFCCFEAAFA74FDA31767" authorityName=": Fauvel" authorityYear="1923" box="[293,519,1850,1873]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="areolata">
<emphasis id="B97B747DC236FFCCFEAAFA74FDA31767" box="[293,519,1850,1873]" italics="true" pageId="21" pageNumber="22">Harmothoe areolata</emphasis>
</taxonomicName>
(additional specimen, SMF 9199). (A) anterior end; (B) left anterior elytron from unknown segment; (C) detail of posterior margin of same; (D) inflated dorsal cirrus; (E) tapering dorsal cirrus; (F) right cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus missing; (G) long notochaeta; (H) tip of same; (I) middle neurochaeta; (J) tip of same; (K) tip of lower neurochaeta. (Scale: A, B = 2 mm; C, F, G, I = 250 µm; D, E = 1 mm; H, J, K = 50 µm).
</paragraph>
</caption>
<caption id="DF70F8E7C235FFCFFF18FA67FEF517C3" ID-DOI="http://doi.org/10.5281/zenodo.5315458" ID-Zenodo-Dep="5315458" httpUri="https://zenodo.org/record/5315458/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,257,1833,1857]" targetBox="[209,1390,191,1803]" targetPageId="22">
<paragraph id="8BB0A86FC235FFCFFF18FA67FEF517C3" blockId="22.[151,1436,1833,2037]" pageId="22" pageNumber="23">
<emphasis id="B97B747DC235FFCFFF18FA67FE8E1777" bold="true" box="[151,298,1833,1857]" pageId="22" pageNumber="23">FIGURE 10.</emphasis>
<taxonomicName id="4C0FD3ECC235FFCFFEBCFA64FDA51777" authorityName=": Hartmann-Schroder" authorityYear="1996" box="[307,513,1834,1857]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="23" phylum="Annelida" rank="species" species="aspera">
<emphasis id="B97B747DC235FFCFFEBCFA64FDA51777" box="[307,513,1834,1857]" italics="true" pageId="22" pageNumber="23">Harmothoe aspera</emphasis>
</taxonomicName>
(small syntype, ZMBN 2000). (A) anterior end, palps and style of median antenna missing, left dorsal tentacular cirrus regenerating; (B) right anterior elytron from unknown segment; (C) detail of posterior margin of same; (D) right cirrigerous parapodium from unknown segment, posterior view, style of dorsal cirrus missing; (E) short notochaeta; (F) long notochaeta; (G) tip of same; (H) middle neurochaeta; (I) tip of same; (J) distal part of upper neurochaeta; (K) lower neurochaeta. (Scale: A, B = 2 mm; C, E, F, H, J, K = 250 µm; D = 1 mm; G = 50 µm; I = 100 µm).
</paragraph>
</caption>
<paragraph id="8BB0A86FC234FFCEFF49FDD6FEA4117A" blockId="23.[151,1436,152,1376]" pageId="23" pageNumber="24">
<emphasis id="B97B747DC234FFCEFF49FDD6FE1D1087" box="[198,441,152,177]" italics="true" pageId="23" pageNumber="24">Additional material.</emphasis>
<materialsCitation id="3B67A232C234FFCEFE4BFDD6FF58117A" accessionNumber="SMF 17280, SMF 17279" collectingDate="1999-06-16" collectingDateMax="2005-06-06" collectingDateMin="1999-06-16" collectorName="NE Atlantic &amp; C. d'Udekem d'Acoz. &amp; Visund N &amp; Gullfaks B &amp; St. &amp; A. Sikorski" country="Norway" elevation="20" latitude="61.200886" location="Sammelsundet Ryggen" longLatPrecision="1" longitude="2.1917863" pageId="23" pageNumber="24" specimenCount="381">
<collectorName id="26FACDB9C234FFCEFE4BFDD6FDFE1084" box="[452,602,152,178]" pageId="23" pageNumber="24">NE Atlantic</collectorName>
: 1 spm. (af + mf), SMF 17280, &quot;Johan Ruud&quot; St. 294,
<geoCoordinate id="EE3BCEA8C234FFCEFA9DFDD6FA381084" box="[1298,1436,152,178]" degrees="70" direction="north" minutes="38.56" orientation="latitude" pageId="23" pageNumber="24" precision="9" value="70.64267">70°38.56N</geoCoordinate>
<geoCoordinate id="EE3BCEA8C234FFCEFF18FDF0FE8410EE" box="[151,288,190,216]" degrees="24" direction="east" minutes="10.46" orientation="longitude" pageId="23" pageNumber="24" precision="9" value="24.174334">24°10.46E</geoCoordinate>
, N
<collectingCountry id="F318E8FFC234FFCEFEC6FDF0FE0C10EE" box="[329,424,190,216]" name="Norway" pageId="23" pageNumber="24">Norway</collectingCountry>
,
<location id="8ED0FEB4C234FFCEFE3BFDF0FD6110EE" LSID="urn:lsid:plazi:treatment:03A61979C237FFCEFF18FB13FC031556:8ED0FEB4C234FFCEFE3BFDF0FD6110EE" box="[436,709,190,216]" country="Norway" latitude="61.200886" longLatPrecision="1" longitude="2.1917863" name="Sammelsundet Ryggen" pageId="23" pageNumber="24">Sammelsundet Ryggen</location>
,
<date id="FFB18EAFC234FFCEFD5FFDF0FCD310EE" box="[720,887,190,216]" pageId="23" pageNumber="24" value="2005-04-12">
<collectingDate id="EFF57747C234FFCEFD5FFDF0FCD310EE" box="[720,887,190,216]" pageId="23" pageNumber="24" value="2005-04-12">12 April 2005</collectingDate>
</date>
, dive;
<quantity id="4CF7058AC234FFCEFC4AFDF0FBA610EE" box="[965,1026,190,216]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="24" unit="m" value="20.0">
<elevation id="00224F5CC234FFCEFC4AFDF0FBA610EE" box="[965,1026,190,216]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="24" unit="m" value="20.0">20 m</elevation>
</quantity>
, leg. &amp; ded.
<collectorName id="26FACDB9C234FFCEFB17FDF0FA3810EE" box="[1176,1436,190,216]" pageId="23" pageNumber="24">C. dUdekem dAcoz.</collectorName>
1 spm.(af), Akvaplan-Niva Collection,
<collectorName id="26FACDB9C234FFCEFDEEFDABFD6B10C9" box="[609,719,229,255]" pageId="23" pageNumber="24">Visund N</collectorName>
2005, St. 1-3,
<date id="FFB18EAFC234FFCEFCF4FDABFBAD10C9" box="[891,1033,229,255]" pageId="23" pageNumber="24" value="2005-06-06">
<collectingDate id="EFF57747C234FFCEFCF4FDABFBAD10C9" box="[891,1033,229,255]" pageId="23" pageNumber="24" value="2005-06-06">6 June 2005</collectingDate>
</date>
,
<geoCoordinate id="EE3BCEA8C234FFCEFB9AFDABFB0A10C9" box="[1045,1198,229,255]" degrees="61.42849" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="61.42849">61.42849° N</geoCoordinate>
,
<geoCoordinate id="EE3BCEA8C234FFCEFB36FDABFA9B10C9" box="[1209,1343,229,255]" degrees="2.56782" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="2.56782">2.56782° E</geoCoordinate>
,
<specimenCount id="9D0963E6C234FFCEFAC4FDABFA3810C9" box="[1355,1436,229,255]" pageId="23" pageNumber="24" type="generic">
<quantity id="4CF7058AC234FFCEFAC4FDABFA3C10C9" box="[1355,1432,229,255]" metricMagnitude="2" metricUnit="m" metricValue="3.81" pageId="23" pageNumber="24" unit="m" value="381.0">
<elevation id="00224F5CC234FFCEFAC4FDABFA3C10C9" box="[1355,1432,229,255]" metricMagnitude="2" metricUnit="m" metricValue="3.81" pageId="23" pageNumber="24" unit="m" value="381.0">381 m</elevation>
</quantity>
.
</specimenCount>
1 spm., SMF 17279;
<collectorName id="26FACDB9C234FFCEFE1FFC42FDAB1110" box="[400,527,268,294]" pageId="23" pageNumber="24">Gullfaks B</collectorName>
1999,
<collectorName id="26FACDB9C234FFCEFDEEFC42FD241110" box="[609,640,268,294]" pageId="23" pageNumber="24">St.</collectorName>
9-2,
<date id="FFB18EAFC234FFCEFD31FC42FCF91110" box="[702,861,268,294]" pageId="23" pageNumber="24" value="1999-06-16">
<collectingDate id="EFF57747C234FFCEFD31FC42FCF91110" box="[702,861,268,294]" pageId="23" pageNumber="24" value="1999-06-16">16 June 1999</collectingDate>
</date>
,
<geoCoordinate id="EE3BCEA8C234FFCEFCE5FC42FB851110" box="[874,1057,268,294]" degrees="61" direction="north" minutes="12" orientation="latitude" pageId="23" pageNumber="24" precision="1" seconds="03.18" value="61.200886">61°12'03.18'' N</geoCoordinate>
,
<geoCoordinate id="EE3BCEA8C234FFCEFBA2FC42FB441110" box="[1069,1248,268,294]" degrees="02" direction="east" minutes="11" orientation="longitude" pageId="23" pageNumber="24" precision="1" seconds="30.43" value="2.1917863">02°11'30.43'' E</geoCoordinate>
,
<quantity id="4CF7058AC234FFCEFB62FC42FA9D1110" box="[1261,1337,268,294]" metricMagnitude="2" metricUnit="m" metricValue="1.41" pageId="23" pageNumber="24" unit="m" value="141.0">
<elevation id="00224F5CC234FFCEFB62FC42FA9D1110" box="[1261,1337,268,294]" metricMagnitude="2" metricUnit="m" metricValue="1.41" pageId="23" pageNumber="24" unit="m" value="141.0">141 m</elevation>
</quantity>
, ded.
<collectorName id="26FACDB9C234FFCEFAF0FC42FF58117A" pageId="23" pageNumber="24">A. Sikorski</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB0A86FC234FFCEFF4AFC17FCD71145" blockId="23.[151,1436,152,1376]" box="[197,883,345,371]" pageId="23" pageNumber="24">
(For further material see
<bibRefCitation id="EF9ED59EC234FFCEFE66FC17FD4B1145" author="Barnich, R. &amp; Fiege, D." box="[489,751,345,371]" pageId="23" pageNumber="24" pagination="1889 - 1938" refId="ref36372" refString="Barnich, R. &amp; Fiege, D. (2000) Revision of the Mediterranean species of Harmothoe Kinberg, 1856 and Lagisca Malmgren, 1865 (Polychaeta, Polynoidae, Polynoinae) with descriptions of a new genus and a new species. Journal of Natural History, 34, 1889 - 1938." type="journal article" year="2000">Barnich &amp; Fiege 2000</bibRefCitation>
and
<bibRefCitation id="EF9ED59EC234FFCEFCA7FC17FCC31145" author="Barnich, R. &amp; Fiege, D." box="[808,871,345,371]" pageId="23" pageNumber="24" pagination="1 - 167" refId="ref36428" refString="Barnich, R. &amp; Fiege, D. (2003) The Aphroditoidea (Annelida, Polychaeta) of the Mediterranean Sea. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 559, 1 - 167." type="journal article" year="2003">2003</bibRefCitation>
).
</paragraph>
<paragraph id="8BB0A86FC234FFCEFF49FCCEFAA611D1" blockId="23.[151,1436,152,1376]" pageId="23" pageNumber="24">
<emphasis id="B97B747DC234FFCEFF49FCCEFE9911AF" box="[198,317,384,409]" italics="true" pageId="23" pageNumber="24">Diagnosis</emphasis>
. Anterior pair of eyes dorsolateral at widest part of prostomium. Elytral margin with fringing papillae; surface covered by conical elongate to thorn-shaped pointed microtubercles and few scattered papillae, with a row of large triangular or pyramid-shaped macrotubercles near posterior margin.
</paragraph>
<paragraph id="8BB0A86FC234FFCEFF49FCBAFD3E1238" blockId="23.[151,1436,152,1376]" box="[198,666,500,526]" pageId="23" pageNumber="24">
<emphasis id="B97B747DC234FFCEFF49FCBAFEF4123B" box="[198,336,500,525]" italics="true" pageId="23" pageNumber="24">Description</emphasis>
(based on smaller
<typeStatus id="54B416CDC234FFCEFDA1FCBAFD2A1238" box="[558,654,500,526]" pageId="23" pageNumber="24" type="syntype">syntype</typeStatus>
).
</paragraph>
<paragraph id="8BB0A86FC234FFCEFF4AFF54FBE9129E" blockId="23.[151,1436,152,1376]" pageId="23" pageNumber="24">
Body with 38 segments. At anterior end (
<figureCitation id="1334B4EAC234FFCEFD35FF54FC8E1202" box="[698,810,538,564]" captionStart="FIGURE 10" captionStartId="22.[151,257,1833,1857]" captionTargetBox="[209,1390,191,1803]" captionTargetId="figure-125@22.[151,1436,169,1822]" captionTargetPageId="22" captionText="FIGURE 10. Harmothoe aspera (small syntype, ZMBN 2000). (A) anterior end, palps and style of median antenna missing, left dorsal tentacular cirrus regenerating; (B) right anterior elytron from unknown segment; (C) detail of posterior margin of same; (D) right cirrigerous parapodium from unknown segment, posterior view, style of dorsal cirrus missing; (E) short notochaeta; (F) long notochaeta; (G) tip of same; (H) middle neurochaeta; (I) tip of same; (J) distal part of upper neurochaeta; (K) lower neurochaeta. (Scale: A, B = 2 mm; C, E, F, H, J, K = 250 µm; D = 1 mm; G = 50 µm; I = 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5315458" httpUri="https://zenodo.org/record/5315458/files/figure.png" pageId="23" pageNumber="24">Fig. 10A</figureCitation>
), prostomium bilobed, with distinct cephalic peaks; ceratophore of median antenna in anterior notch, lateral antennae inserted ventrally, styles of antennae papillate, tapering to filiform tip; anterior pair of eyes situated dorsolaterally at widest part of prostomium, posterior pair dorsally near hind margin of prostomium; palps papillate, tapering.
</paragraph>
<paragraph id="8BB0A86FC234FFCEFF4AFFFBFB5B132A" blockId="23.[151,1436,152,1376]" pageId="23" pageNumber="24">Tentaculophores inserted laterally to prostomium, each with two notochaetae and a dorsal and ventral tentacular cirrus, styles of cirri papillate, tapering to filiform tip. Second segment with first pair of elytra, biramous parapodia, and long buccal cirri. Following segments with tapering, short ventral cirri.</paragraph>
<paragraph id="8BB0A86FC234FFCEFF4AFE67FB0413E8" blockId="23.[151,1436,152,1376]" pageId="23" pageNumber="24">
Fifteen pairs of elytra, covering dorsum, on segments 2, 4, 5, 7, then on every second segment to 23, 26, 29, 32; last six segments cirrigerous; elytral margin with fringing papillae; surface covered by conical elongate to thorn-shaped pointed microtubercles and few scattered papillae, with a row of large triangular or pyramid-shaped macrotubercles near posterior margin (
<figureCitation id="1334B4EAC234FFCEFCAFFED3FC021381" box="[800,934,925,951]" captionStart="FIGURE 10" captionStartId="22.[151,257,1833,1857]" captionTargetBox="[209,1390,191,1803]" captionTargetId="figure-125@22.[151,1436,169,1822]" captionTargetPageId="22" captionText="FIGURE 10. Harmothoe aspera (small syntype, ZMBN 2000). (A) anterior end, palps and style of median antenna missing, left dorsal tentacular cirrus regenerating; (B) right anterior elytron from unknown segment; (C) detail of posterior margin of same; (D) right cirrigerous parapodium from unknown segment, posterior view, style of dorsal cirrus missing; (E) short notochaeta; (F) long notochaeta; (G) tip of same; (H) middle neurochaeta; (I) tip of same; (J) distal part of upper neurochaeta; (K) lower neurochaeta. (Scale: A, B = 2 mm; C, E, F, H, J, K = 250 µm; D = 1 mm; G = 50 µm; I = 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5315458" httpUri="https://zenodo.org/record/5315458/files/figure.png" pageId="23" pageNumber="24">Fig. 10B,C</figureCitation>
). Cirrigerous segments with distinct dorsal tubercles; dorsal cirri with cylindrical cirrophore, style papillate, tapering to filiform tip.
</paragraph>
<paragraph id="8BB0A86FC234FFCEFF4AFEA4FF5A14F0" blockId="23.[151,1436,152,1376]" pageId="23" pageNumber="24">
Parapodia biramous; notopodia with elongate acicular lobe; neuropodia with elongate prechaetal acicular lobe with digitiform supra-acicular process; neuropodial postchaetal lobe shorter than prechaetal lobe, rounded; tips of noto- and neuroacicula penetrating epidermis (
<figureCitation id="1334B4EAC234FFCEFC44F976FBE71464" box="[971,1091,1080,1106]" captionStart="FIGURE 10" captionStartId="22.[151,257,1833,1857]" captionTargetBox="[209,1390,191,1803]" captionTargetId="figure-125@22.[151,1436,169,1822]" captionTargetPageId="22" captionText="FIGURE 10. Harmothoe aspera (small syntype, ZMBN 2000). (A) anterior end, palps and style of median antenna missing, left dorsal tentacular cirrus regenerating; (B) right anterior elytron from unknown segment; (C) detail of posterior margin of same; (D) right cirrigerous parapodium from unknown segment, posterior view, style of dorsal cirrus missing; (E) short notochaeta; (F) long notochaeta; (G) tip of same; (H) middle neurochaeta; (I) tip of same; (J) distal part of upper neurochaeta; (K) lower neurochaeta. (Scale: A, B = 2 mm; C, E, F, H, J, K = 250 µm; D = 1 mm; G = 50 µm; I = 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5315458" httpUri="https://zenodo.org/record/5315458/files/figure.png" pageId="23" pageNumber="24">Fig. 10D</figureCitation>
). Notochaetae stouter than neurochaetae, with distinct rows of spines and blunt tip (
<figureCitation id="1334B4EAC234FFCEFCC2F910FC44144E" box="[845,992,1118,1144]" captionStart="FIGURE 10" captionStartId="22.[151,257,1833,1857]" captionTargetBox="[209,1390,191,1803]" captionTargetId="figure-125@22.[151,1436,169,1822]" captionTargetPageId="22" captionText="FIGURE 10. Harmothoe aspera (small syntype, ZMBN 2000). (A) anterior end, palps and style of median antenna missing, left dorsal tentacular cirrus regenerating; (B) right anterior elytron from unknown segment; (C) detail of posterior margin of same; (D) right cirrigerous parapodium from unknown segment, posterior view, style of dorsal cirrus missing; (E) short notochaeta; (F) long notochaeta; (G) tip of same; (H) middle neurochaeta; (I) tip of same; (J) distal part of upper neurochaeta; (K) lower neurochaeta. (Scale: A, B = 2 mm; C, E, F, H, J, K = 250 µm; D = 1 mm; G = 50 µm; I = 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5315458" httpUri="https://zenodo.org/record/5315458/files/figure.png" pageId="23" pageNumber="24">Fig. 10EG</figureCitation>
); neurochaetae with distinct rows of spines, middle bidentate with small secondary tooth (often abraded), upper and lower unidentate (
<figureCitation id="1334B4EAC234FFCEFAE6F9CBFF5614F0" captionStart="FIGURE 10" captionStartId="22.[151,257,1833,1857]" captionTargetBox="[209,1390,191,1803]" captionTargetId="figure-125@22.[151,1436,169,1822]" captionTargetPageId="22" captionText="FIGURE 10. Harmothoe aspera (small syntype, ZMBN 2000). (A) anterior end, palps and style of median antenna missing, left dorsal tentacular cirrus regenerating; (B) right anterior elytron from unknown segment; (C) detail of posterior margin of same; (D) right cirrigerous parapodium from unknown segment, posterior view, style of dorsal cirrus missing; (E) short notochaeta; (F) long notochaeta; (G) tip of same; (H) middle neurochaeta; (I) tip of same; (J) distal part of upper neurochaeta; (K) lower neurochaeta. (Scale: A, B = 2 mm; C, E, F, H, J, K = 250 µm; D = 1 mm; G = 50 µm; I = 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5315458" httpUri="https://zenodo.org/record/5315458/files/figure.png" pageId="23" pageNumber="24">Fig. 10HK</figureCitation>
).
</paragraph>
<paragraph id="8BB0A86FC234FFCEFF4AF99DFD9A1525" blockId="23.[151,1436,152,1376]" pageId="23" pageNumber="24">
<emphasis id="B97B747DC234FFCEFF4AF99DFECB14DA" box="[197,367,1235,1260]" italics="true" pageId="23" pageNumber="24">Measurements</emphasis>
.
<taxonomicName id="4C0FD3ECC234FFCEFEF2F99DFE4E14DA" authorityName=": Hartmann-Schroder" authorityYear="1996" box="[381,490,1235,1260]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="aspera">
<emphasis id="B97B747DC234FFCEFEF2F99DFE4E14DA" box="[381,490,1235,1260]" italics="true" pageId="23" pageNumber="24">P. aspera</emphasis>
</taxonomicName>
,
<specimenCount id="9D0963E6C234FFCEFE77F99DFDD314DB" box="[504,631,1235,1261]" pageId="23" pageNumber="24" type="generic" typeStatus="syntypes">2 syntypes</specimenCount>
,
<collectionCode id="ED1E30AAC234FFCEFD0CF99DFD7C14DB" box="[643,728,1235,1261]" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="23" pageNumber="24">ZMBN</collectionCode>
2000: L
<quantity id="4CF7058AC234FFCEFCCDF99DFC3314DB" box="[834,919,1235,1261]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="23" pageNumber="24" unit="mm" value="18.0">18 mm</quantity>
, W
<quantity id="4CF7058AC234FFCEFC4AF99DFBAD14DB" box="[965,1033,1235,1261]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="24" unit="mm" value="6.0">6 mm</quantity>
for 39 segments; L 16.5 mm, W
<quantity id="4CF7058AC234FFCEFA02F99DFF611525" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="24" unit="mm" value="6.0">6 mm</quantity>
for 38 segments (spm. figured).
</paragraph>
<paragraph id="8BB0A86FC234FFCEFF4AF86EFAE6150C" blockId="23.[151,1436,152,1376]" box="[197,1346,1312,1338]" pageId="23" pageNumber="24">
<emphasis id="B97B747DC234FFCEFF4AF86EFEF0150F" box="[197,340,1312,1337]" italics="true" pageId="23" pageNumber="24">Distribution</emphasis>
. Arctic, North Pacific, North Atlantic, North Sea to Kattegat, and Mediterranean Sea.
</paragraph>
<paragraph id="8BB0A86FC234FFCEFF4AF809FC031556" blockId="23.[151,1436,152,1376]" box="[197,935,1350,1376]" pageId="23" pageNumber="24">
<emphasis id="B97B747DC234FFCEFF4AF809FEBA1556" box="[197,286,1351,1376]" italics="true" pageId="23" pageNumber="24">Habitat</emphasis>
. Occurs on a wide range of substrata in
<quantity id="4CF7058AC234FFCEFD79F808FC071556" box="[758,931,1350,1376]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="10.0" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="600.0" valueMax="1000.0" valueMin="200.0">200 to 1000 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>