305 lines
39 KiB
XML
305 lines
39 KiB
XML
<document id="0B72421278EE63EDE776863B122CFE1E" ID-CLB-Dataset="279445" ID-DOI="10.11646/zootaxa.5365.1.1" ID-GBIF-Dataset="23a3b51c-63b7-4937-bdba-384f28db6be5" ID-ISSN="1175-5334" ID-Zenodo-Dep="10146969" ID-ZooBank="1DC72714-D0E8-49D8-821D-03C6B2A7AE80" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1700197580075" checkinUser="plazi" docAuthor="Colloff, Matthew J." docDate="2023" docId="03A2C77C4622FF88C79CB2E11495DC58" docLanguage="en" docName="zootaxa.5365.1.1.pdf" docOrigin="Zootaxa 5365 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pterozetidae Luxton 1988" docType="treatment" docVersion="4" lastPageNumber="69" masterDocId="FF9BBF044661FFCCC70BB31F1724DE65" masterDocTitle="The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families" masterLastPageNumber="93" masterPageNumber="1" pageNumber="68" updateTime="1701119607223" updateUser="ExternalLinkService">
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<mods:title id="430CE25989F69291703C78CF28F5FFD9">The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families</mods:title>
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<mods:namePart id="4A7B839C9A82C55F5FB2C0742A938BC6">Colloff, Matthew J.</mods:namePart>
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<mods:typeOfResource id="9D75AD38BC7D6EB8FD52020A33579EA9">text</mods:typeOfResource>
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<mods:title id="7C49BBE6B4E781BA2BDC19782F4197AB">Zootaxa</mods:title>
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<mods:part id="31FE21B43DC0B6D8BE87B954A501297D">
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<mods:date id="1F9784224DEA7FED75586CCCCEE69BC9">2023</mods:date>
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<mods:number id="8841EBE15C2D68955077F2F87339193E">2023-11-06</mods:number>
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<mods:number id="9ACC884754725DD30B1A0CAABBC8BAC4">5365</mods:number>
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<mods:identifier id="99B75E733A497F877C3963DB67FF91ED" type="DOI">10.11646/zootaxa.5365.1.1</mods:identifier>
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</mods:mods>
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<treatment id="03A2C77C4622FF88C79CB2E11495DC58" ID-GBIF-Taxon="219262726" LSID="urn:lsid:plazi:treatment:03A2C77C4622FF88C79CB2E11495DC58" httpUri="http://treatment.plazi.org/id/03A2C77C4622FF88C79CB2E11495DC58" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68">
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<subSubSection id="C31125E14622FF8FC79CB2E116FBDC7D" box="[151,479,510,537]" pageId="67" pageNumber="68" type="nomenclature">
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<paragraph id="8BB4766A4622FF8FC79CB2E116FBDC7D" blockId="67.[151,479,510,537]" box="[151,479,510,537]" pageId="67" pageNumber="68">
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<heading id="D0FCC1064622FF8FC79CB2E116FBDC7D" bold="true" box="[151,479,510,537]" fontSize="11" level="2" pageId="67" pageNumber="68" reason="3">
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<taxonomicName id="4C0B0DE94622FF8FC79CB2E116FBDC7D" authority="Luxton, 1988" authorityName="Luxton" authorityYear="1988" box="[151,479,510,537]" class="Arachnida" family="Pterozetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oribatida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="family">
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<emphasis id="B97FAA784622FF8FC79CB2E116FBDC7D" bold="true" box="[151,479,510,537]" pageId="67" pageNumber="68">Pterozetidae Luxton, 1988</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31125E14622FF8FC79CB15C16D1DC3E" box="[151,501,579,603]" pageId="67" pageNumber="68" type="reference_group">
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<paragraph id="8BB4766A4622FF8FC79CB15C16D1DC3E" blockId="67.[151,612,579,634]" box="[151,501,579,603]" pageId="67" pageNumber="68">
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<treatmentCitationGroup id="AB1B51444622FF8FC79CB15C16D1DC3E" box="[151,501,579,603]" pageId="67" pageNumber="68">
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<treatmentCitation id="0AAA507B4622FF8FC79CB15C16D5DC3E" author="Luxton, M." box="[151,497,579,603]" page="83" pageId="67" pageNumber="68" year="1988">
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<taxonomicName id="4C0B0DE94622FF8FC79CB15C16D5DC3E" authority="Luxton, 1988a, p. 83" authorityName="Luxton" authorityPageNumber="83" authorityYear="1988" box="[151,497,579,603]" class="Arachnida" family="Pterozetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oribatida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="family">
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Pterozetidae
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<bibRefCitation id="EF9A0B9B4622FF8FC610B15B1695DC3E" author="Luxton, M." box="[283,433,579,603]" pageId="67" pageNumber="68" pagination="71 - 91" refId="ref67435" refString="Luxton, M. (1988 a) A new mite superfamily (Acari: Cryptostigmata). Zoological Journal of the Linnean Society, 93, 71 - 91. https: // doi. org / 10.1111 / j. 1096 - 3642.1988. tb 01360. x" type="journal article" year="1988">Luxton, 1988a</bibRefCitation>
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, p. 83
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31125E14622FF8FC79CB17D1540DC1F" box="[151,612,610,634]" pageId="67" pageNumber="68" type="materials_examined">
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<paragraph id="8BB4766A4622FF8FC79CB17D1540DC1F" blockId="67.[151,612,579,634]" box="[151,612,610,634]" pageId="67" pageNumber="68">
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<typeStatus id="54B0C8C84622FF8FC79CB17D17EDDC1F" box="[151,201,610,634]" pageId="67" pageNumber="68">Type</typeStatus>
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genus:
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<taxonomicName id="4C0B0DE94622FF8FC613B17C1544DC1F" authority="Hammer, 1966, p. 78" authorityName=", Hammer" authorityPageNumber="78" authorityYear="1966" box="[280,608,610,634]" class="Arachnida" family="Eutegaeidae" genus="Pterozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
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<emphasis id="B97FAA784622FF8FC613B17C16A5DC1F" box="[280,385,611,634]" italics="true" pageId="67" pageNumber="68">Pterozetes</emphasis>
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<bibRefCitation id="EF9A0B9B4622FF8FC68CB17D1504DC1F" author="Hammer, M." box="[391,544,610,634]" pageId="67" pageNumber="68" pagination="1 - 108" refId="ref66631" refString="Hammer, M. (1966) Investigations on the Oribatid fauna of New Zealand. Part I. Biologiske Skrifter det Kongelige Dansk Videnskabernes Selskab, 15 (2), 1 - 108." type="journal article" year="1966">Hammer, 1966</bibRefCitation>
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, p. 78
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31125E14622FF8FC79CB1B713A2DAF3" pageId="67" pageNumber="68" type="diagnosis">
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<paragraph id="8BB4766A4622FF8FC79CB1B71431DC83" blockId="67.[151,1437,680,2039]" pageId="67" pageNumber="68">
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<emphasis id="B97FAA784622FF8FC79CB1B71636DCA7" bold="true" box="[151,274,680,706]" pageId="67" pageNumber="68">Diagnosis.</emphasis>
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The following diagnosis is based on that of
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<bibRefCitation id="EF9A0B9B4622FF8FC405B1B614E4DCA7" author="Luxton, M." box="[782,960,680,707]" pageId="67" pageNumber="68" pagination="71 - 91" refId="ref67435" refString="Luxton, M. (1988 a) A new mite superfamily (Acari: Cryptostigmata). Zoological Journal of the Linnean Society, 93, 71 - 91. https: // doi. org / 10.1111 / j. 1096 - 3642.1988. tb 01360. x" type="journal article" year="1988">Luxton (1988a)</bibRefCitation>
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, modified to accommodate the transfer of
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<taxonomicName id="4C0B0DE94622FF8FC79CB1D31619DC83" authorityName="J. Balogh & Mahunka" authorityYear="1966" box="[151,317,716,742]" class="Arachnida" family="Eutegaeidae" genus="Porrhotegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
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<emphasis id="B97FAA784622FF8FC79CB1D31619DC83" box="[151,317,716,742]" italics="true" pageId="67" pageNumber="68">Porrhotegaeus</emphasis>
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</taxonomicName>
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to
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<taxonomicName id="4C0B0DE94622FF8FC66AB1D31533DC83" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[353,535,716,742]" family="Porrhotegaeidae" pageId="67" pageNumber="68" rank="family" status="fam. nov.">Porrhotegaeidae</taxonomicName>
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<emphasis id="B97FAA784622FF8FC515B1D315A8DC83" bold="true" box="[542,652,716,742]" pageId="67" pageNumber="68">
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<taxonomicNameLabel id="A24C17034622FF8FC515B1D315A8DC83" box="[542,652,716,742]" pageId="67" pageNumber="68" rank="family">fam. nov.</taxonomicNameLabel>
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</emphasis>
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(cf. below).
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</paragraph>
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<paragraph id="8BB4766A4622FF8FC7CCB1EF13A2DAF3" blockId="67.[151,1437,680,2039]" pageId="67" pageNumber="68">
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Lamellae long, well developed, fused with each other apically, their anterior margin concave and curved or Vshaped, with lateral points, but anterior margin not folded ventrally and fused with rostrum. Bothridia fused with bases of lamellae and prodorsum. With long, blade-like humeral processes extending from lateral margin at widest part of notogaster to point at least level with mid-length of lamellae, reflexed ventrally (
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<taxonomicName id="4C0B0DE94622FF8FC383B0431220DD13" authorityName="J. Balogh" authorityYear="1970" box="[1160,1284,860,886]" class="Arachnida" family="Eutegaeidae" genus="Dudichella" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
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<emphasis id="B97FAA784622FF8FC383B0431220DD13" box="[1160,1284,860,886]" italics="true" pageId="67" pageNumber="68">Dudichella</emphasis>
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</taxonomicName>
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), but lacking marginal lobes posterior of main anterior lobe. Interlamellar setae present (
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<taxonomicName id="4C0B0DE94622FF8FC4FFB09E17F4DDDB" authority="Hammer, 1966" authorityName="Hammer" authorityYear="1966" class="Arachnida" family="Eutegaeidae" genus="Pterozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="novazealandicus">
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<emphasis id="B97FAA784622FF8FC4FFB09E1208DDFF" box="[1012,1324,896,922]" italics="true" pageId="67" pageNumber="68">Pterozetes novazealandicus</emphasis>
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<bibRefCitation id="EF9A0B9B4622FF8FC23FB09E17F4DDDB" author="Hammer, M." pageId="67" pageNumber="68" pagination="1 - 108" refId="ref66631" refString="Hammer, M. (1966) Investigations on the Oribatid fauna of New Zealand. Part I. Biologiske Skrifter det Kongelige Dansk Videnskabernes Selskab, 15 (2), 1 - 108." type="journal article" year="1966">Hammer, 1966</bibRefCitation>
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</taxonomicName>
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) or absent. Notogaster U-shaped, with eight or nine pairs of short, thin, smooth setae; those of
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<emphasis id="B97FAA784622FF8FC20CB0BB122BDDDB" box="[1287,1295,932,958]" italics="true" pageId="67" pageNumber="68">l</emphasis>
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and
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<emphasis id="B97FAA784622FF8FC24CB0BB1271DDDB" box="[1351,1365,932,958]" italics="true" pageId="67" pageNumber="68">h</emphasis>
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series positioned marginally. Pedotectum I large, broad, sub-rectangular in ventral outline; pd II rectangular or pointed; discidium sub-rectangular or ovoid. Anterior margin of pedotectum I and epimere I forming straight or curved transverse tectum covering posterior mentum. Perigenital carinae present (
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<taxonomicName id="4C0B0DE94622FF8FC30EB70F13A5DA4F" authorityName="J. Balogh" authorityYear="1970" box="[1029,1153,1040,1066]" class="Arachnida" family="Eutegaeidae" genus="Dudichella" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
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<emphasis id="B97FAA784622FF8FC30EB70F13A5DA4F" box="[1029,1153,1040,1066]" italics="true" pageId="67" pageNumber="68">Dudichella</emphasis>
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</taxonomicName>
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) or absent (
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<taxonomicName id="4C0B0DE94622FF8FC213B70E12A8DA4F" authorityName=", Hammer" authorityYear="1966" box="[1304,1420,1041,1066]" class="Arachnida" family="Eutegaeidae" genus="Pterozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
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<emphasis id="B97FAA784622FF8FC213B70E12A8DA4F" box="[1304,1420,1041,1066]" italics="true" pageId="67" pageNumber="68">Pterozetes</emphasis>
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</taxonomicName>
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). Enantiophysis E4 present as posterior tubercle only (
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<taxonomicName id="4C0B0DE94622FF8FC40FB72A14D4DA2B" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[772,1008,1076,1102]" class="Arachnida" family="Eutegaeidae" genus="Pterozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="lawrencei" status="sp. nov.">
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<emphasis id="B97FAA784622FF8FC40FB72A14D4DA2B" box="[772,1008,1076,1102]" italics="true" pageId="67" pageNumber="68">Pterozetes lawrencei</emphasis>
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</taxonomicName>
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<emphasis id="B97FAA784622FF8FC4F0B72B137DDA2B" bold="true" box="[1019,1113,1076,1102]" pageId="67" pageNumber="68">
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<taxonomicNameLabel id="A24C17034622FF8FC4F0B72B137DDA2B" box="[1019,1113,1076,1102]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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) or absent (
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<taxonomicName id="4C0B0DE94622FF8FC3E7B72B124CDA2B" authorityName="J. Balogh" authorityYear="1970" box="[1260,1384,1076,1102]" class="Arachnida" family="Eutegaeidae" genus="Dudichella" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B97FAA784622FF8FC3E7B72B124CDA2B" box="[1260,1384,1076,1102]" italics="true" pageId="67" pageNumber="68">Dudichella</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C0B0DE94622FF8FC79CB74616F6DA17" authorityName="Hammer" authorityYear="1966" box="[151,466,1112,1138]" class="Arachnida" family="Eutegaeidae" genus="Pterozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="novazealandicus">
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<emphasis id="B97FAA784622FF8FC79CB74616F6DA17" box="[151,466,1112,1138]" italics="true" pageId="67" pageNumber="68">Pterozetes novazealandicus</emphasis>
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</taxonomicName>
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). Six pairs of genital setae: aligned longitudinally in
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<taxonomicName id="4C0B0DE94622FF8FC358B74613E3DA17" authorityName=", Hammer" authorityYear="1966" box="[1107,1223,1113,1138]" class="Arachnida" family="Eutegaeidae" genus="Pterozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
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<emphasis id="B97FAA784622FF8FC358B74613E3DA17" box="[1107,1223,1113,1138]" italics="true" pageId="67" pageNumber="68">Pterozetes</emphasis>
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||
</taxonomicName>
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or with
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<emphasis id="B97FAA784622FF8FC23FB7461266DA17" box="[1332,1346,1113,1138]" italics="true" pageId="67" pageNumber="68">g</emphasis>
|
||
<subScript id="178F742F4622FF8FC249B775126EDA1D" attach="left" box="[1346,1354,1130,1144]" fontSize="6" pageId="67" pageNumber="68">2</subScript>
|
||
and
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<emphasis id="B97FAA784622FF8FC28DB74612B0DA17" box="[1414,1428,1113,1138]" italics="true" pageId="67" pageNumber="68">g</emphasis>
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||
<subScript id="178F742F4622FF8FC29FB77512B8DA1D" attach="left" box="[1428,1436,1130,1144]" fontSize="6" pageId="67" pageNumber="68">5</subScript>
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displaced laterally in
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<taxonomicName id="4C0B0DE94622FF8FC68CB7631527DAF3" authorityName="J. Balogh" authorityYear="1970" box="[391,515,1148,1174]" class="Arachnida" family="Eutegaeidae" genus="Dudichella" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
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<emphasis id="B97FAA784622FF8FC68CB7631527DAF3" box="[391,515,1148,1174]" italics="true" pageId="67" pageNumber="68">Dudichella</emphasis>
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||
</taxonomicName>
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||
. Three pairs of adanal setae. Anal plates lozenge-shaped.
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</paragraph>
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</subSubSection>
|
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<subSubSection id="C31125E14622FF88C7CCB7BF1495DC58" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68" type="discussion">
|
||
<paragraph id="8BB4766A4622FF8FC7CCB7BF1265DBBF" blockId="67.[151,1437,680,2039]" pageId="67" pageNumber="68">
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||
<emphasis id="B97FAA784622FF8FC7CCB7BF161FDADF" bold="true" box="[199,315,1184,1210]" pageId="67" pageNumber="68">Remarks.</emphasis>
|
||
<bibRefCitation id="EF9A0B9B4622FF8FC64FB7BE15F5DADF" author="Norton, R. A. & Behan-Pelletier, V. M." box="[324,721,1184,1211]" pageId="67" pageNumber="68" pagination="430 - 564" refId="ref68204" refString="Norton, R. A. & Behan-Pelletier, V. M. (2009) Suborder Oribatida. In: Krantz, G. W. & Walter, D. E. (Eds.), A Manual of Acarology. 3 rd Edition. Texas Tech University Press, Lubbock, Texas, pp. 430 - 564." type="book chapter" year="2009">Norton and Behan-Pelletier (2009)</bibRefCitation>
|
||
considered
|
||
<taxonomicName id="4C0B0DE94622FF8FC457B7BF14C2DADF" authorityName="Luxton" authorityYear="1988" box="[860,998,1184,1210]" class="Arachnida" family="Pterozetidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="67" pageNumber="68" phylum="Arthropoda" rank="family">Pterozetidae</taxonomicName>
|
||
a junior synonym of
|
||
<taxonomicName id="4C0B0DE94622FF8FC3E9B7BF124FDADF" authorityName="J. Balogh" authorityYear="1965" box="[1250,1387,1184,1210]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="family">Eutegaeidae</taxonomicName>
|
||
due to the apparent lack of diagnostic characters at the family-level. However, the humeral processes of
|
||
<taxonomicName id="4C0B0DE94622FF8FC219B7DB12B8DABB" authorityName="Luxton" authorityYear="1988" box="[1298,1436,1220,1246]" class="Arachnida" family="Pterozetidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="67" pageNumber="68" phylum="Arthropoda" rank="family">Pterozetidae</taxonomicName>
|
||
originate on the lateral margin of the notogaster at its widest part, well posterior of the dorsosejugal scissure, and are curved and convex along their external margins, forming triangular, blade-like structures extending almost to the rostrum, covering most or all of legs I and II, and quite different from those of the
|
||
<taxonomicName id="4C0B0DE94622FF8FC35AB62F13FADB2F" authorityName="J. Balogh" authorityYear="1965" box="[1105,1246,1328,1354]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="family">Eutegaeidae</taxonomicName>
|
||
. In her definition of
|
||
<taxonomicName id="4C0B0DE94622FF8FC7BDB64A1511DB0B" authority=", Hammer (1966, p. 78)" authorityName=", Hammer" authorityPageNumber="78" authorityYear="1966" box="[182,565,1364,1391]" class="Arachnida" family="Eutegaeidae" genus="Pterozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA784622FF8FC7BDB64A160EDB0B" box="[182,298,1365,1390]" italics="true" pageId="67" pageNumber="68">Pterozetes</emphasis>
|
||
,
|
||
<bibRefCitation id="EF9A0B9B4622FF8FC632B64A16C3DB0B" author="Hammer, M." box="[313,487,1364,1391]" pageId="67" pageNumber="68" pagination="1 - 108" refId="ref66631" refString="Hammer, M. (1966) Investigations on the Oribatid fauna of New Zealand. Part I. Biologiske Skrifter det Kongelige Dansk Videnskabernes Selskab, 15 (2), 1 - 108." type="journal article" year="1966">Hammer (1966</bibRefCitation>
|
||
, p. 78)
|
||
</taxonomicName>
|
||
pointed out that the humeral process ‘is different from that of
|
||
<taxonomicName id="4C0B0DE94622FF8FC3F1B64A124BDB0B" authorityName="Colloff" authorityYear="2023" box="[1274,1391,1365,1390]" class="Arachnida" family="Eutegaeidae" genus="Eutegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA784622FF8FC3F1B64A124BDB0B" box="[1274,1391,1365,1390]" italics="true" pageId="67" pageNumber="68">Eutegaeus</emphasis>
|
||
</taxonomicName>
|
||
’, in that it is more like a pteromorph than the cuticular projection from the anterior margin of the notogaster found in the latter genus. In
|
||
<taxonomicName id="4C0B0DE94622FF8FC643B68316E0DBD3" authorityName="J. Balogh" authorityYear="1970" box="[328,452,1436,1462]" class="Arachnida" family="Eutegaeidae" genus="Dudichella" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA784622FF8FC643B68316E0DBD3" box="[328,452,1436,1462]" italics="true" pageId="67" pageNumber="68">Dudichella</emphasis>
|
||
</taxonomicName>
|
||
, the ventrally curved median margins of the humeral processes overlap with the lateral margins of epimeres I and II, which are broadened and excavated, covering legs I and II (J. Balogh, 1970).
|
||
</paragraph>
|
||
<paragraph id="8BB4766A4622FF8FC7CCB6FB144ED89F" blockId="67.[151,1437,680,2039]" pageId="67" pageNumber="68">
|
||
The main character state that differentiates
|
||
<taxonomicName id="4C0B0DE94622FF8FC5A2B6FB1417DB9B" authorityName="Luxton" authorityYear="1988" box="[681,819,1508,1534]" class="Arachnida" family="Pterozetidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="67" pageNumber="68" phylum="Arthropoda" rank="family">Pterozetidae</taxonomicName>
|
||
from other families of
|
||
<taxonomicName id="4C0B0DE94622FF8FC33EB6FB13E8DB9B" authorityName="Balogh" authorityYear="1965" box="[1077,1228,1508,1534]" class="Pilidiophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heteronemertea" pageId="67" pageNumber="8" phylum="Nemertea" rank="superFamily" superFamily="Eutegaeoidea">Eutegaeoidea</taxonomicName>
|
||
is that the anterior parts of the lamellae are fused with each other and their anterior margin is concave and curved or V-shaped, but is not folded ventrally and fused with the rostrum and with lateral foramina as in
|
||
<taxonomicName id="4C0B0DE94622FF8FC306B5331393D823" authorityName="Hammer" authorityYear="1966" box="[1037,1207,1580,1606]" class="Arachnida" family="Cepheusidae" genus="Compactozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA784622FF8FC306B5331393D823" box="[1037,1207,1580,1606]" italics="true" pageId="67" pageNumber="68">Compactozetes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EF9A0B9B4622FF8FC3CEB5321249D823" author="Luxton, M." box="[1221,1389,1580,1607]" pageId="67" pageNumber="68" pagination="71 - 91" refId="ref67435" refString="Luxton, M. (1988 a) A new mite superfamily (Acari: Cryptostigmata). Zoological Journal of the Linnean Society, 93, 71 - 91. https: // doi. org / 10.1111 / j. 1096 - 3642.1988. tb 01360. x" type="journal article" year="1988">Luxton (1988a</bibRefCitation>
|
||
, pp. 84, 85) considered the lamellar cusps of
|
||
<taxonomicName id="4C0B0DE94622FF8FC568B54F15FBD80F" authorityName="J. Balogh" authorityYear="1970" box="[611,735,1616,1642]" class="Arachnida" family="Eutegaeidae" genus="Dudichella" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA784622FF8FC568B54F15FBD80F" box="[611,735,1616,1642]" italics="true" pageId="67" pageNumber="68">Dudichella</emphasis>
|
||
</taxonomicName>
|
||
are free and not fused, but J.
|
||
<bibRefCitation id="EF9A0B9B4622FF8FC33BB54F13FFD80F" author="Balogh, J." box="[1072,1243,1616,1642]" pageId="67" pageNumber="68" pagination="33 - 67" refId="ref64077" refString="Balogh, J. (1970 b) New oribatids from Ceylon. The scientific results of the Hungarian soil zoological expeditions. Opuscula Zoologica, 10, 33 - 67." type="journal article" year="1970">Balogh (1970b</bibRefCitation>
|
||
, p. 36) described them as ‘medially confluent and thus X-shaped’ and his
|
||
<figureCitation id="13306AEF4622FF8FC400B56B1468D8EB" box="[779,844,1652,1678]" captionStart="FIGURE 1" captionStartId="6.[152,255,1105,1129]" captionTargetBox="[151,1436,181,1081]" captionTargetId="figure-26@6.[151,1436,181,1081]" captionTargetPageId="6" captionText="FIGURE 1. Global distribution of genera and species of Eutegaeoidea and Cepheoidea, based on records of type localities of species described prior to the present work. Inset: detail of type localities of species in New Zealand." figureDoi="http://doi.org/10.5281/zenodo.10146971" httpUri="https://zenodo.org/record/10146971/files/figure.png" pageId="67" pageNumber="68">Fig. 1</figureCitation>
|
||
clearly shows the cusps as fused in the midline, with a deep V-shaped indentation between the lateral apices. This arrangement is completely different from the free lamellar cusps of the
|
||
<taxonomicName id="4C0B0DE94622FF8FC688B5A3152BD8B3" authorityName="J. Balogh" authorityYear="1965" box="[387,527,1724,1750]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="family">Eutegaeidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B0DE94622FF8FC512B5A315E8D8B3" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[537,716,1724,1750]" family="Neoeutegaeidae" pageId="67" pageNumber="81" rank="family" status="fam. nov.">Neoeutegaeidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0B0DE94622FF8FC40BB5A31492D8B3" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[768,950,1724,1750]" family="Porrhotegaeidae" pageId="67" pageNumber="68" rank="family" status="fam. nov.">Porrhotegaeidae</taxonomicName>
|
||
<taxonomicNameLabel id="A24C17034622FF8FC4B7B5A3130FD8B2" box="[956,1067,1724,1751]" pageId="67" pageNumber="68" rank="family">
|
||
<emphasis id="B97FAA784622FF8FC4B7B5A31300D8B3" bold="true" box="[956,1060,1724,1750]" pageId="67" pageNumber="68">fam. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
I consider the morphology of the anterior margin of the lamellae a synapomorphy of
|
||
<taxonomicName id="4C0B0DE94622FF8FC5D3B5FF1442D89F" authorityName="Luxton" authorityYear="1988" box="[728,870,1760,1786]" class="Arachnida" family="Pterozetidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="67" pageNumber="68" phylum="Arthropoda" rank="family">Pterozetidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB4766A4622FF8FC7CCB41B1661D9B7" blockId="67.[151,1437,680,2039]" pageId="67" pageNumber="68">
|
||
<taxonomicName id="4C0B0DE94622FF8FC7CCB41B1675D97B" authorityName="Luxton" authorityYear="1988" box="[199,337,1796,1822]" class="Arachnida" family="Pterozetidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="67" pageNumber="68" phylum="Arthropoda" rank="family">Pterozetidae</taxonomicName>
|
||
is morphologically most similar to
|
||
<taxonomicName id="4C0B0DE94622FF8FC5EAB41B1487D97B" authorityName="Luxton" authorityYear="1988" box="[737,931,1796,1822]" family="Compactozetidae" pageId="67" pageNumber="68" rank="family">Compactozetidae</taxonomicName>
|
||
in that the margins of the bothridia are fused with the prodorsum and bases of lamellae and the humeral processes originate on the margin of the notogaster at its widest part. However, they differ from those of the
|
||
<taxonomicName id="4C0B0DE94622FF8FC400B45314E9D903" authorityName="Luxton" authorityYear="1988" box="[779,973,1868,1894]" family="Compactozetidae" pageId="67" pageNumber="68" rank="family">Compactozetidae</taxonomicName>
|
||
in lacking the 2-4 lobes posterior of the apical humeral projection. As mentioned above, the lamellae of
|
||
<taxonomicName id="4C0B0DE94622FF8FC456B46F14C3D9EF" authorityName="Luxton" authorityYear="1988" box="[861,999,1904,1930]" class="Arachnida" family="Pterozetidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="67" pageNumber="68" phylum="Arthropoda" rank="family">Pterozetidae</taxonomicName>
|
||
are quite different in morphology from those of
|
||
<taxonomicName id="4C0B0DE94622FF8FC7F1B48B1680D9CB" authorityName="Hammer" authorityYear="1966" box="[250,420,1940,1966]" class="Arachnida" family="Cepheusidae" genus="Compactozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA784622FF8FC7F1B48B1680D9CB" box="[250,420,1940,1966]" italics="true" pageId="67" pageNumber="68">Compactozetes</emphasis>
|
||
</taxonomicName>
|
||
. Also, the anal plates of
|
||
<taxonomicName id="4C0B0DE94622FF8FC5B5B48B146CD9CB" authorityName="Luxton" authorityYear="1988" box="[702,840,1940,1966]" class="Arachnida" family="Pterozetidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="67" pageNumber="68" phylum="Arthropoda" rank="family">Pterozetidae</taxonomicName>
|
||
are lozenge-shaped while those of
|
||
<taxonomicName id="4C0B0DE94622FF8FC3D1B48B12B8D9CB" authorityName="Luxton" authorityYear="1988" box="[1242,1436,1940,1966]" family="Compactozetidae" pageId="67" pageNumber="68" rank="family">Compactozetidae</taxonomicName>
|
||
are rectangular.
|
||
</paragraph>
|
||
<paragraph id="8BB4766A4622FF88C7CCB4C3150BDFC8" blockId="67.[151,1437,680,2039]" lastBlockId="68.[151,1437,151,573]" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68">
|
||
Additional diagnostic characters for
|
||
<taxonomicName id="4C0B0DE94622FF8FC56DB4C315D4D993" authorityName="Luxton" authorityYear="1988" box="[614,752,2012,2038]" class="Arachnida" family="Pterozetidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="67" pageNumber="68" phylum="Arthropoda" rank="family">Pterozetidae</taxonomicName>
|
||
are that epimeres I and II are very broad and long, whereas epimeres III and IV are much narrower and shorter. Pedotectum I is very large with sub-rectangular lateral margins, with an anterolateral point in
|
||
<taxonomicName id="4C0B0DE94625FF88C6E3B3A41540DEB0" authorityName="J. Balogh" authorityYear="1970" box="[488,612,187,213]" class="Arachnida" family="Eutegaeidae" genus="Dudichella" kingdom="Animalia" order="Sarcoptiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA784625FF88C6E3B3A41540DEB0" box="[488,612,187,213]" italics="true" pageId="68" pageNumber="69">Dudichella</emphasis>
|
||
</taxonomicName>
|
||
, and extend to the medial margin of the humeral process (
|
||
<taxonomicName id="4C0B0DE94625FF88C20BB3A41250DEB1" authorityName=", Hammer" authorityYear="1966" box="[1280,1396,187,212]" class="Arachnida" family="Eutegaeidae" genus="Pterozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA784625FF88C20BB3A41250DEB1" box="[1280,1396,187,212]" italics="true" pageId="68" pageNumber="69">Pterozetes</emphasis>
|
||
</taxonomicName>
|
||
) or overlap it (
|
||
<taxonomicName id="4C0B0DE94625FF88C61DB3C016B6DE9C" authorityName="J. Balogh" authorityYear="1970" box="[278,402,223,249]" class="Arachnida" family="Eutegaeidae" genus="Dudichella" kingdom="Animalia" order="Sarcoptiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA784625FF88C61DB3C016B6DE9C" box="[278,402,223,249]" italics="true" pageId="68" pageNumber="69">Dudichella</emphasis>
|
||
</taxonomicName>
|
||
). The anterior margin of pedotectum I and epimere I forms a straight or curved transverse tectum covering the posterior part of the mentum and the ventro-lateral margins of the gnathosoma, as in most
|
||
<taxonomicName id="4C0B0DE94625FF88C79CB2381678DF24" authorityName="Luxton" authorityYear="1988" box="[151,348,295,321]" family="Compactozetidae" pageId="68" pageNumber="69" rank="family">Compactozetidae</taxonomicName>
|
||
, but this structure is not present in the
|
||
<taxonomicName id="4C0B0DE94625FF88C432B23814E6DF24" authorityName="J. Balogh" authorityYear="1965" box="[825,962,295,321]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="family">Eutegaeidae</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C0B0DE94625FF88C4FAB238138CDF24" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[1009,1192,295,321]" family="Neoeutegaeidae" pageId="68" pageNumber="81" rank="family" status="fam. nov.">Neoeutegaeidae</taxonomicName>
|
||
. I consider there are sufficient character states of diagnostic value to retain
|
||
<taxonomicName id="4C0B0DE94625FF88C400B25414B1DF00" authorityName="Luxton" authorityYear="1988" box="[779,917,331,357]" class="Arachnida" family="Pterozetidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="68" pageNumber="69" phylum="Arthropoda" rank="family">Pterozetidae</taxonomicName>
|
||
as a valid family. The interlamellar setae are absent in
|
||
<taxonomicName id="4C0B0DE94625FF88C60EB27016CBDFEC" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[261,495,367,393]" class="Arachnida" family="Eutegaeidae" genus="Pterozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="lawrencei" status="sp. nov.">
|
||
<emphasis id="B97FAA784625FF88C60EB27016CBDFEC" box="[261,495,367,393]" italics="true" pageId="68" pageNumber="69">Pterozetes lawrencei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97FAA784625FF88C6FCB2701577DFEC" bold="true" box="[503,595,367,393]" pageId="68" pageNumber="69">
|
||
<taxonomicNameLabel id="A24C17034625FF88C6FCB2701577DFEC" box="[503,595,367,393]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C0B0DE94625FF88C587B270142CDFEC" authorityName="J. Balogh" authorityYear="1970" box="[652,776,367,393]" class="Arachnida" family="Eutegaeidae" genus="Dudichella" kingdom="Animalia" order="Sarcoptiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA784625FF88C587B270142CDFEC" box="[652,776,367,393]" italics="true" pageId="68" pageNumber="69">Dudichella</emphasis>
|
||
</taxonomicName>
|
||
but present in
|
||
<taxonomicName id="4C0B0DE94625FF88C4BEB27012B8DFEC" authority="Hammer, 1966" authorityName="Hammer" authorityYear="1966" box="[949,1436,367,393]" class="Arachnida" family="Eutegaeidae" genus="Pterozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="novazealandicus">
|
||
<emphasis id="B97FAA784625FF88C4BEB27013C8DFEC" box="[949,1260,367,393]" italics="true" pageId="68" pageNumber="69">Pterozetes novazealandicus</emphasis>
|
||
<bibRefCitation id="EF9A0B9B4625FF88C3FFB27012B8DFEC" author="Hammer, M." box="[1268,1436,367,393]" pageId="68" pageNumber="69" pagination="1 - 108" refId="ref66631" refString="Hammer, M. (1966) Investigations on the Oribatid fauna of New Zealand. Part I. Biologiske Skrifter det Kongelige Dansk Videnskabernes Selskab, 15 (2), 1 - 108." type="journal article" year="1966">Hammer, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation id="EF9A0B9B4625FF88C7CFB28C164EDFC8" author="Luxton, M." box="[196,362,403,429]" pageId="68" pageNumber="69" pagination="431 - 433" refId="ref67516" refString="Luxton, M. (1988 c) The family Pterozetidae (Cryptostigmata: Eutegaeoidea) in New Zealand. New Zealand Journal of Zoology, 15, 431 - 433. https: // doi. org / 10.1080 / 03014223.1988.10422970" type="journal article" year="1988">Luxton, 1988c</bibRefCitation>
|
||
,
|
||
<figureCitation id="13306AEF4625FF88C67DB28C16EDDFC8" box="[374,457,403,429]" captionStart="FIGURE 1" captionStartId="6.[152,255,1105,1129]" captionTargetBox="[151,1436,181,1081]" captionTargetId="figure-26@6.[151,1436,181,1081]" captionTargetPageId="6" captionText="FIGURE 1. Global distribution of genera and species of Eutegaeoidea and Cepheoidea, based on records of type localities of species described prior to the present work. Inset: detail of type localities of species in New Zealand." figureDoi="http://doi.org/10.5281/zenodo.10146971" httpUri="https://zenodo.org/record/10146971/files/figure.png" pageId="68" pageNumber="69">Fig. 1A</figureCitation>
|
||
therein).
|
||
</paragraph>
|
||
<paragraph id="8BB4766A4625FF88C7CCB2A81495DC58" blockId="68.[151,1437,151,573]" pageId="68" pageNumber="69">
|
||
In
|
||
<taxonomicName id="4C0B0DE94625FF88C7E2B2A81641DFB4" authorityName="J. Balogh" authorityYear="1970" box="[233,357,439,465]" class="Arachnida" family="Eutegaeidae" genus="Dudichella" kingdom="Animalia" order="Sarcoptiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA784625FF88C7E2B2A81641DFB4" box="[233,357,439,465]" italics="true" pageId="68" pageNumber="69">Dudichella</emphasis>
|
||
</taxonomicName>
|
||
the lamellar setae emerge from the ventral surface of the lamellae, as in
|
||
<taxonomicName id="4C0B0DE94625FF88C3CDB2A81248DFB4" authorityName="J. Balogh & Mahunka" authorityYear="1966" box="[1222,1388,439,465]" class="Arachnida" family="Eutegaeidae" genus="Porrhotegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA784625FF88C3CDB2A81248DFB4" box="[1222,1388,439,465]" italics="true" pageId="68" pageNumber="69">Porrhotegaeus</emphasis>
|
||
</taxonomicName>
|
||
(cf. below).
|
||
<taxonomicName id="4C0B0DE94625FF88C7F9B2C4164ADF90" authorityName="J. Balogh" authorityYear="1970" box="[242,366,475,501]" class="Arachnida" family="Eutegaeidae" genus="Dudichella" kingdom="Animalia" order="Sarcoptiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA784625FF88C7F9B2C4164ADF90" box="[242,366,475,501]" italics="true" pageId="68" pageNumber="69">Dudichella</emphasis>
|
||
</taxonomicName>
|
||
also differs from
|
||
<taxonomicName id="4C0B0DE94625FF88C53EB2C4158DDF91" authorityName=", Hammer" authorityYear="1966" box="[565,681,475,500]" class="Arachnida" family="Eutegaeidae" genus="Pterozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA784625FF88C53EB2C4158DDF91" box="[565,681,475,500]" italics="true" pageId="68" pageNumber="69">Pterozetes</emphasis>
|
||
</taxonomicName>
|
||
in having what J.
|
||
<bibRefCitation id="EF9A0B9B4625FF88C47FB2C41300DF90" author="Balogh, J." box="[884,1060,475,501]" pageId="68" pageNumber="69" pagination="33 - 67" refId="ref64077" refString="Balogh, J. (1970 b) New oribatids from Ceylon. The scientific results of the Hungarian soil zoological expeditions. Opuscula Zoologica, 10, 33 - 67." type="journal article" year="1970">Balogh (1970b)</bibRefCitation>
|
||
described as ‘minute, peloptoide’ [
|
||
<emphasis id="B97FAA784625FF88C7AAB2E017E4DC7D" box="[161,192,511,536]" italics="true" pageId="68" pageNumber="69">sic</emphasis>
|
||
] chelicerae. It is unclear whether the chelicerae are truly pelopsiform (i.e. expanded basally) or are simply thin, attenuate and chelate-dentate as in
|
||
<taxonomicName id="4C0B0DE94625FF88C515B13C140DDC59" baseAuthorityName="Balogh & P. Balogh" baseAuthorityYear="1983" box="[542,809,547,573]" class="Arachnida" family="Eutegaeidae" genus="Atalotegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="mensarosi">
|
||
<emphasis id="B97FAA784625FF88C515B13C140DDC59" box="[542,809,547,573]" italics="true" pageId="68" pageNumber="69">Atalotegaeus mensarosi</emphasis>
|
||
</taxonomicName>
|
||
(cf. above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |