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<mods:title id="5CBEB278A47462F8FAB84DADD865C4E4">A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species</mods:title>
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<heading id="D0C1C930FFB6FF862AB26971FDEBFBA9" bold="true" box="[151,541,1107,1134]" fontSize="11" level="1" pageId="3" pageNumber="548" reason="1">
<taxonomicName id="4C3605DFFFB6FF862AB26971FDEBFBA9" authority="Miyatake" authorityName="Miyatake" authorityYear="1976" box="[151,541,1107,1134]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="548" phylum="Arthropoda" rank="species" species="hisamatsui">
<emphasis id="B942A24EFFB6FF862AB26971FDEBFBA9" bold="true" box="[151,541,1107,1134]" pageId="3" pageNumber="548">
<emphasis id="B942A24EFFB6FF862AB26971FE68FBAA" bold="true" box="[151,414,1107,1134]" italics="true" pageId="3" pageNumber="548">Horniolus hisamatsui</emphasis>
Miyatake
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<emphasis id="B942A24EFFB6FF862AB269BAFE84FB74" box="[151,370,1176,1200]" italics="true" pageId="3" pageNumber="548">Horniolus hisamatsui</emphasis>
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<bibRefCitation id="EFA703ADFFB6FF862B5D69BAFDC8FB74" author="Miyatake, M." box="[376,574,1175,1200]" pageId="3" pageNumber="548" pagination="29 - 37" refId="ref5103" refString="Miyatake, M. (1976) Descriptions of five new species of the genus Horniolus Weise of Southeast Asia (Coleoptera: Coccinellidae). Transactions of the Shikoku Entomological Society, 13 (1 - 2), 29 - 37." type="journal article" year="1976">Miyatake, 1976: 29</bibRefCitation>
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(
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female; EUMJ).
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<taxonomicName id="4C3605DFFFB6FF862AB26994FDBEFB0A" authority=": Canepari 2003: 262" authorityName=": Canepari" authorityPageNumber="262" authorityYear="2003" box="[151,584,1206,1230]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="548" phylum="Arthropoda" rank="species" species="hisamatsui">
<emphasis id="B942A24EFFB6FF862AB26994FE84FB0A" box="[151,370,1206,1230]" italics="true" pageId="3" pageNumber="548">Horniolus hisamatsui</emphasis>
:
<bibRefCitation id="EFA703ADFFB6FF862B5A6994FDBEFB0A" author="Canepari, C." box="[383,584,1206,1230]" pageId="3" pageNumber="548" pagination="261 - 265" refId="ref4612" refString="Canepari, C. (2003) Coccinellidae (Insecta: Coleoptera) of Nepal from the collection of the Naturkundemuseum Erfurt. In: Hartmann, M. &amp; Baumbach, H. (Eds.), Biodiversitat und Naturausstattung im Himalaya, e. V. Verein der Freunde und Forderer des Naturkundemuseums, Erfurt, pp. 261 - 265." type="book chapter" year="2003">Canepari 2003: 262</bibRefCitation>
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.
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<bibRefCitation id="EFA703ADFFB6FF86287D6994FCC1FB0A" author="Chen, X. &amp; Xie, X. &amp; Ren, S. &amp; Wang, X." box="[600,823,1206,1230]" pageId="3" pageNumber="548" pagination="105 - 123" refId="ref4677" refString="Chen, X., Xie, X., Ren, S. &amp; Wang, X. (2016) A taxonomic review of the genus Horniolus Weise from China, with description of a new species (Coleoptera, Coccinellidae). ZooKeys, 623, 105 - 123. https: // doi. org / 10.3897 / zookeys. 623.10191" type="journal article" year="2016">
Chen
<emphasis id="B942A24EFFB6FF8628B36995FD35FB0A" box="[662,707,1206,1230]" italics="true" pageId="3" pageNumber="548">et al</emphasis>
. 2016: 114
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;
<treatmentCitation id="0A97584DFFB6FF8629676994FB64FB0A" ID-DOI="http://doi.org/10.5281/zenodo.4416935" ID-Zenodo-Dep="4416935" author="Szawaryn, K. &amp; Vetrovec, J." box="[834,1170,1206,1230]" httpUri="http://treatment.plazi.org/id/03F4878E4233400E22D9F882FBD7FF3A" page="576" pageId="3" pageNumber="548" year="2020">
<bibRefCitation id="EFA703ADFFB6FF8629676994FB64FB0A" author="Szawaryn, K. &amp; Vetrovec, J." box="[834,1170,1206,1230]" pageId="3" pageNumber="548" pagination="573 - 580" refId="ref5715" refString="Szawaryn, K. &amp; Vetrovec, J. (2020) A new species of Horniolus Weise with remarks on the systematic position of the genus (Coleoptera: Coccinellidae). Zootaxa, 4861 (4), 573 - 580. https: // doi. org / 10.11646 / zootaxa. 4861.4.5" type="journal article" year="2020">Szawaryn &amp; Větrovec 2020: 576</bibRefCitation>
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.
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<paragraph id="8B897E5CFFB6FF862AB269DEFD99FAFF" blockId="3.[151,1436,1276,1446]" pageId="3" pageNumber="548">
<emphasis id="B942A24EFFB6FF862AB269DEFEC7FAD2" bold="true" box="[151,305,1276,1302]" pageId="3" pageNumber="548">Distribution.</emphasis>
<collectingCountry id="F3213ECCFFB6FF862B1069DEFE99FAD2" box="[309,367,1276,1302]" name="India" pageId="3" pageNumber="548">India</collectingCountry>
(
<bibRefCitation id="EFA703ADFFB6FF862B5E69DFFD44FAD3" author="Szawaryn, K. &amp; Vetrovec, J." box="[379,690,1277,1303]" pageId="3" pageNumber="548" pagination="573 - 580" refId="ref5715" refString="Szawaryn, K. &amp; Vetrovec, J. (2020) A new species of Horniolus Weise with remarks on the systematic position of the genus (Coleoptera: Coccinellidae). Zootaxa, 4861 (4), 573 - 580. https: // doi. org / 10.11646 / zootaxa. 4861.4.5" type="journal article" year="2020">Szawaryn &amp; Větrovec 2020</bibRefCitation>
),
<collectingCountry id="F3213ECCFFB6FF8628E669DEFCF0FAD2" box="[707,774,1276,1302]" name="Nepal" pageId="3" pageNumber="548">Nepal</collectingCountry>
(
<bibRefCitation id="EFA703ADFFB6FF86293769DEFC4CFAD3" author="Canepari, C." box="[786,954,1276,1303]" pageId="3" pageNumber="548" pagination="261 - 265" refId="ref4612" refString="Canepari, C. (2003) Coccinellidae (Insecta: Coleoptera) of Nepal from the collection of the Naturkundemuseum Erfurt. In: Hartmann, M. &amp; Baumbach, H. (Eds.), Biodiversitat und Naturausstattung im Himalaya, e. V. Verein der Freunde und Forderer des Naturkundemuseums, Erfurt, pp. 261 - 265." type="book chapter" year="2003">Canepari 2003</bibRefCitation>
), and
<collectingCountry id="F3213ECCFFB6FF8629DD69DEFBCDFAD2" box="[1016,1083,1276,1302]" name="China" pageId="3" pageNumber="548">China</collectingCountry>
(
<collectingRegion id="49F2B0BEFFB6FF862E6D69DEFB39FAD2" box="[1096,1231,1276,1302]" country="China" name="Guangdong" pageId="3" pageNumber="548">Guangdong</collectingRegion>
,
<collectingRegion id="49F2B0BEFFB6FF862EFD69DEFADAFAD2" box="[1240,1324,1276,1302]" country="China" name="Hainan" pageId="3" pageNumber="548">Hainan</collectingRegion>
,
<collectingRegion id="49F2B0BEFFB6FF862F1069DEFA6EFAD2" box="[1333,1432,1276,1302]" country="China" name="Guangxi" pageId="3" pageNumber="548">Guangxi</collectingRegion>
,
<collectingRegion id="49F2B0BEFFB6FF862AB26802FF25FAFF" box="[151,211,1312,1339]" country="China" name="Xizang" pageId="3" pageNumber="548">Tibet</collectingRegion>
) (
<bibRefCitation id="EFA703ADFFB6FF862ACC6803FE6FFAFF" author="Ren, S. X. &amp; Wang, X. M. &amp; Pang, H. &amp; Peng, Z. Q. &amp; Zeng, T." box="[233,409,1312,1339]" pageId="3" pageNumber="548" refId="ref5448" refString="Ren, S. X., Wang, X. M., Pang, H., Peng, Z. Q. &amp; Zeng, T. (2009) Colored Pictorial Handbook of Ladybird Beetles in China. Science Press, Beijing, 336 pp." type="book" year="2009">
Ren
<emphasis id="B942A24EFFB6FF862B3A6803FEA1FAFE" box="[287,343,1312,1338]" italics="true" pageId="3" pageNumber="548">et al.</emphasis>
2009
</bibRefCitation>
;
<bibRefCitation id="EFA703ADFFB6FF862B806802FD95FAFF" author="Chen, X. &amp; Xie, X. &amp; Ren, S. &amp; Wang, X." box="[421,611,1312,1339]" pageId="3" pageNumber="548" pagination="105 - 123" refId="ref4677" refString="Chen, X., Xie, X., Ren, S. &amp; Wang, X. (2016) A taxonomic review of the genus Horniolus Weise from China, with description of a new species (Coleoptera, Coccinellidae). ZooKeys, 623, 105 - 123. https: // doi. org / 10.3897 / zookeys. 623.10191" type="journal article" year="2016">
Chen
<emphasis id="B942A24EFFB6FF862BC26803FDE9FAFE" box="[487,543,1312,1338]" italics="true" pageId="3" pageNumber="548">et al.</emphasis>
2016
</bibRefCitation>
).
</paragraph>
<paragraph id="8B897E5CFFB6FF862AE26867FD92FA62" blockId="3.[151,1436,1276,1446]" pageId="3" pageNumber="548">
<emphasis id="B942A24EFFB6FF862AE26867FEF0FA9B" bold="true" box="[199,262,1349,1375]" pageId="3" pageNumber="548">Note.</emphasis>
<bibRefCitation id="EFA703ADFFB6FF862B356866FE38FA9B" author="Miyatake, M." box="[272,462,1348,1375]" pageId="3" pageNumber="548" pagination="29 - 37" refId="ref5103" refString="Miyatake, M. (1976) Descriptions of five new species of the genus Horniolus Weise of Southeast Asia (Coleoptera: Coccinellidae). Transactions of the Shikoku Entomological Society, 13 (1 - 2), 29 - 37." type="journal article" year="1976">Miyatake (1976)</bibRefCitation>
described it from
<collectingCountry id="F3213ECCFFB6FF8628806867FCD9FA9B" box="[677,815,1349,1375]" name="Hong Kong" pageId="3" pageNumber="548">Hong Kong</collectingCountry>
.
<bibRefCitation id="EFA703ADFFB6FF86291E6866FBF9FA9B" author="Chen, X. &amp; Xie, X. &amp; Ren, S. &amp; Wang, X." box="[827,1039,1348,1375]" pageId="3" pageNumber="548" pagination="105 - 123" refId="ref4677" refString="Chen, X., Xie, X., Ren, S. &amp; Wang, X. (2016) A taxonomic review of the genus Horniolus Weise from China, with description of a new species (Coleoptera, Coccinellidae). ZooKeys, 623, 105 - 123. https: // doi. org / 10.3897 / zookeys. 623.10191" type="journal article" year="2016">
Chen
<emphasis id="B942A24EFFB6FF8629A56867FC42FA9A" box="[896,948,1348,1374]" italics="true" pageId="3" pageNumber="548">et al</emphasis>
. (2016)
</bibRefCitation>
redescribed it with illustrations of specimens from
<collectingCountry id="F3213ECCFFB6FF862B70684AFE6AFA46" box="[341,412,1384,1410]" name="China" pageId="3" pageNumber="548">China</collectingCountry>
. Recently
<bibRefCitation id="EFA703ADFFB6FF862833684BFC91FA47" author="Szawaryn, K. &amp; Vetrovec, J." box="[534,871,1385,1411]" pageId="3" pageNumber="548" pagination="573 - 580" refId="ref5715" refString="Szawaryn, K. &amp; Vetrovec, J. (2020) A new species of Horniolus Weise with remarks on the systematic position of the genus (Coleoptera: Coccinellidae). Zootaxa, 4861 (4), 573 - 580. https: // doi. org / 10.11646 / zootaxa. 4861.4.5" type="journal article" year="2020">Szawaryn &amp; Větrovec (2020)</bibRefCitation>
recorded it from northeastern
<collectingCountry id="F3213ECCFFB6FF862EEC684AFAF5FA46" box="[1225,1283,1384,1410]" name="India" pageId="3" pageNumber="548">India</collectingCountry>
(
<collectingRegion id="49F2B0BEFFB6FF862F36684BFA94FA47" box="[1299,1378,1385,1411]" country="India" name="Assam" pageId="3" pageNumber="548">Assam</collectingRegion>
) and illustrated the habitus and male genitalia.
</paragraph>
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</treatment>
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