treatments-xml/data/03/98/E9/0398E958487A1576FF4109031033FD21.xml
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<document id="5D865C33B80EBCEE426A4343E7490A78" ID-DOI="10.11646/zootaxa.1866.1.3" ID-ISSN="1175­5334" ID-Zenodo-Dep="5133305" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627156080233" checkinUser="felipe" docAuthor="Tabachnick, Konstantin R. &amp; Menschenina, Dorte Janussen And Larisa L." docDate="2008" docId="0398E958487A1576FF4109031033FD21" docLanguage="en" docName="zt01866p068.pdf" docOrigin="Zootaxa 1866 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1866.1.3" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Sympagella multihexastera Tabachnick &amp; Menschenina 2008, sp. nov." docType="treatment" docVersion="3" lastPageNumber="65" masterDocId="FFA191204842154DFFD60F491274FFC3" masterDocTitle="New Australian Hexactinellida (Porifera) with a revision of Euplectella aspergillum *" masterLastPageNumber="68" masterPageNumber="7" pageNumber="63" updateTime="1699021398582" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7989A78B4E74E73278C3828AADDD0D24">New Australian Hexactinellida (Porifera) with a revision of Euplectella aspergillum *</mods:title>
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<mods:namePart id="5D2469818DB9CD8C97341782FBE5FBFB">Tabachnick, Konstantin R.</mods:namePart>
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<mods:namePart id="DC04EFA24397B977CA1021273FE75CE9">Menschenina, Dorte Janussen And Larisa L.</mods:namePart>
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<mods:title id="4EEF1C20FEB5DAB804AA0ACAECC802C9">Zootaxa</mods:title>
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<mods:date id="1D2BC39FB3EBA403F71DF1606AE0BB73">2008</mods:date>
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<subSubSection id="C32B0BC5487A1575FF4109031149F9A7" box="[151,829,1610,1636]" pageId="56" pageNumber="63" type="nomenclature">
<paragraph id="8B8E584E487A1575FF4109031149F9A7" blockId="56.[151,829,1530,1636]" box="[151,829,1610,1636]" pageId="56" pageNumber="63">
<emphasis id="B945845C487A1575FF4109031037F9A7" bold="true" box="[151,579,1610,1636]" pageId="56" pageNumber="63">
<taxonomicName id="4C3123CD487A1575FF41090313AAF9A7" ID-CoL="7B6VM" authority="Tabachnick &amp; Menschenina, 2008" authorityName="Tabachnick &amp; Menschenina" authorityYear="2008" box="[151,478,1610,1636]" class="Hexactinellida" family="Rossellidae" genus="Sympagella" kingdom="Animalia" order="Lyssacinosida" pageId="56" pageNumber="63" phylum="Porifera" rank="species" species="multihexastera" status="sp. nov.">
<emphasis id="B945845C487A1575FF41090313AAF9A7" bold="true" box="[151,478,1610,1636]" italics="true" pageId="56" pageNumber="63">Sympagella multihexastera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2763927487A1575FE3009031037F9A7" box="[486,579,1610,1636]" pageId="56" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="130A44CB487A1575FD82090310DFF9A7" box="[596,683,1610,1636]" captionStart="FIGURE 31" captionStartId="57.[151,255,1295,1319]" captionTargetBox="[383,1211,200,1254]" captionTargetId="figure-237@57.[382,1215,190,1259]" captionTargetPageId="57" captionText="FIGURE 31. Sympagella multihexastera, sp. n., spicules, holotype. A, dermal pinular pentactin. B, atrial pinular hexactin. C, hypodermal pentactin. D­E, choanosomal hexactins. F­I, central parts and outer ends of choanosomal diactins. J, discohexaster. K, outer end of discohexaster. L, onychohexaster. M, oxyhexaster. N, oxyhemihexaster. O, oxyhexactin. P, strobiloplumicome." figureDoi="http://doi.org/10.5281/zenodo.5133365" httpUri="https://zenodo.org/record/5133365/files/figure.png" pageId="56" pageNumber="63">Fig. 31</figureCitation>
,
<figureCitation id="130A44CB487A1575FD60090310A2F9A7" box="[694,726,1610,1636]" captionStart="FIGURE 32" captionStartId="58.[151,255,1290,1314]" captionTargetBox="[151,1436,181,1264]" captionTargetId="figure-275@58.[151,1436,175,1264]" captionTargetPageId="58" captionText="FIGURE 32. Sympagella multihexastera, sp.n., holotype. Dermal pinules, scale 30 µm." figureDoi="http://doi.org/10.5281/zenodo.5133367" httpUri="https://zenodo.org/record/5133367/files/figure.png" pageId="56" pageNumber="63">32</figureCitation>
;
<tableCitation id="C6B36DF5487A1575FD3509031141F9A7" box="[739,821,1610,1636]" captionStart="TABLE 29" captionStartId="59.[151,239,802,826]" captionText="TABLE 29: Some measurements of spicules of Sympagella multihexastera, sp.n." pageId="56" pageNumber="63">Tab 29</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32B0BC5487A1576FF4109D21033FD21" lastPageId="59" lastPageNumber="66" pageId="56" pageNumber="63" type="etymology">
<paragraph id="8B8E584E487A1575FF4109D21287F91F" blockId="56.[151,1436,1690,1963]" pageId="56" pageNumber="63">
<emphasis id="B945845C487A1575FF4109D2136EF977" box="[151,282,1691,1716]" italics="true" pageId="56" pageNumber="63">Etymology.</emphasis>
The name refers to the multitude of hexaster
<typeStatus id="548AE6EC487A1575FCFB09D3111FF977" box="[813,875,1690,1716]" pageId="56" pageNumber="63">types</typeStatus>
characteristic of the spicule composition of this species.
</paragraph>
<paragraph id="8B8E584E487A1575FF1009A210B3F8F3" blockId="56.[151,1436,1690,1963]" pageId="56" pageNumber="63">
<emphasis id="B945845C487A1575FF1009A21359F8C7" box="[198,301,1771,1796]" italics="true" pageId="56" pageNumber="63">Material</emphasis>
.
<materialsCitation id="3B595213487A1575FEE909A210B3F8F3" collectionCode="NTM" elevation="405" latitude="-17.466667" longLatPrecision="1278" longitude="118.88333" pageId="56" pageNumber="63" specimenCode="Z 0002592" specimenCount="405" typeStatus="holotype">
<emphasis id="B945845C487A1575FEE909A213DEF8C7" box="[319,426,1771,1796]" italics="true" pageId="56" pageNumber="63">
<typeStatus id="548AE6EC487A1575FEE909A213DEF8C7" box="[319,426,1771,1796]" pageId="56" pageNumber="63" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectionCode id="ED20C08B487A1575FE6809A3138BF8C7" box="[446,511,1770,1796]" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="56" pageNumber="63">NTM</collectionCode>
<specimenCode id="DB97F035487A1575FDDC09A310E5F8C7" box="[522,657,1770,1796]" collectionCode="NTM" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="56" pageNumber="63">Z 0002592</specimenCode>
—Sta. NWS­0050, off Rowley Shoals, Northwest Shelf, Scampi trawl Grounds,
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17
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28S
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118
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53E
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, depth
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<quantity id="4CC9F5AB487A1575FDA3085F10B6F8F3" box="[629,706,1814,1840]" metricMagnitude="2" metricUnit="m" metricValue="4.05" pageId="56" pageNumber="63" unit="m" value="405.0">
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.
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</paragraph>
<paragraph id="8B8E584E487A1575FF10087710F7F840" blockId="56.[151,1436,1690,1963]" pageId="56" pageNumber="63">
<materialsCitation id="3B595213487A1575FF10087710F7F840" collectionCode="NTM" elevation="405" latitude="-17.466667" longLatPrecision="1278" longitude="118.88333" pageId="56" pageNumber="63" specimenCode="Z 0002594" specimenCount="405" typeStatus="paratype">
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<typeStatus id="548AE6EC487A1575FF1008771345F894" box="[198,305,1854,1879]" pageId="56" pageNumber="63" type="paratype">Paratype</typeStatus>
</emphasis>
:
<collectionCode id="ED20C08B487A1575FE9C087713FFF89B" box="[330,395,1854,1880]" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="56" pageNumber="63">NTM</collectionCode>
<specimenCode id="DB97F035487A1575FE4C08771051F89B" box="[410,549,1854,1880]" collectionCode="NTM" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="56" pageNumber="63">Z 0002594</specimenCode>
—Sta. NWS­0052, off Rowley Shoals, Northwest Shelf, Scampi trawl Grounds,
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17
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28S
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118
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53E
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, depth
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.
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</paragraph>
<paragraph id="8B8E584E487A1575FF1008D8173DF868" blockId="56.[151,1436,1690,1963]" box="[198,1353,1937,1963]" pageId="56" pageNumber="63">
<emphasis id="B945845C487A1575FF1008D8132CF869" box="[198,344,1937,1962]" italics="true" pageId="56" pageNumber="63">Description.</emphasis>
Body: Both specimens are presented by lamella­like fragments,
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in thickness.
</paragraph>
<caption id="DF4E08C6487B1574FF410A46131CFA4C" ID-DOI="http://doi.org/10.5281/zenodo.5133365" ID-Zenodo-Dep="5133365" httpUri="https://zenodo.org/record/5133365/files/figure.png" pageId="57" pageNumber="64" startId="57.[151,255,1295,1319]" targetBox="[383,1211,200,1254]" targetPageId="57">
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<emphasis id="B945845C487B1574FF410A461353FAE4" bold="true" box="[151,295,1295,1319]" pageId="57" pageNumber="64">FIGURE 31.</emphasis>
<taxonomicName id="4C3123CD487B1574FEFB0A59103BFAE4" authority="Tabachnick &amp; Menschenina, 2008" authorityName="Tabachnick &amp; Menschenina" authorityYear="2008" box="[301,591,1296,1319]" class="Hexactinellida" family="Rossellidae" genus="Sympagella" kingdom="Animalia" order="Lyssacinosida" pageId="57" pageNumber="64" phylum="Porifera" rank="species" species="multihexastera" status="sp. nov.">
<emphasis id="B945845C487B1574FEFB0A59103BFAE4" box="[301,591,1296,1319]" italics="true" pageId="57" pageNumber="64">Sympagella multihexastera</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2763927487B1574FD8B0A5910ECFAE4" box="[605,664,1296,1319]" pageId="57" pageNumber="64" rank="species">sp. n.</taxonomicNameLabel>
, spicules, holotype. A, dermal pinular pentactin. B, atrial pinular hexactin. C, hypodermal pentactin. D­E, choanosomal hexactins. F­I, central parts and outer ends of choanosomal diactins. J, discohexaster. K, outer end of discohexaster. L, onychohexaster. M, oxyhexaster. N, oxyhemihexaster. O, oxyhexactin. P, strobiloplumicome.
</paragraph>
</caption>
<paragraph id="8B8E584E487B1574FF100A8B1155F8AF" blockId="57.[151,1439,1474,1900]" pageId="57" pageNumber="64">
Spicules: Choanosomal spicules are diactins and hexactins. The diactins have stout shafts, rarely with a widening in the middle and conically pointed or rounded outer ends, smooth or rough, they measure 1.52.9/
<quantity id="4CC9F5AB487B1574FF41095B1332F9EF" box="[151,326,1554,1580]" metricMagnitude="-5" metricUnit="m" metricValue="7" metricValueMax="13.0" metricValueMin="1.0" pageId="57" pageNumber="64" unit="mm" value="0.07" valueMax="0.13" valueMin="0.01">0.010.13 mm</quantity>
. The hexactins have conically pointed outer ends, usually smooth, some hexactins have rough rays directed outside the body; the rays of these hexactins are 0.150.37/0.010.0.026 mm. Hypodermalia and probably hypoatrialia are pentactins with conically pointed, usually smooth outer ends, their tangential rays are
<quantity id="4CC9F5AB487B1574FF1209C31306F967" box="[196,370,1674,1700]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" metricValueMax="8.7" metricValueMin="2.5" pageId="57" pageNumber="64" unit="mm" value="0.56" valueMax="0.87" valueMin="0.25">0.250.87 mm</quantity>
long, the ray directed inside the body is
<quantity id="4CC9F5AB487B1574FCB309C31196F967" box="[869,994,1674,1700]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="57" pageNumber="64" unit="mm" value="0.65" valueMax="1.0" valueMin="0.3">0.31 mm</quantity>
, the diameter of these rays is
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<geoCoordinate id="EE053E89487B1574FA9809C317E4F967" box="[1358,1424,1674,1700]" degrees="0.014" direction="north" orientation="latitude" pageId="57" pageNumber="64" precision="55" value="0.014">0.014</geoCoordinate>
<geoCoordinate id="EE053E89487B1574FA4609C312ADF90F" degrees="0.026" direction="west" orientation="longitude" pageId="57" pageNumber="64" precision="55" value="-0.026"> 0.026</geoCoordinate>
mm
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. Dermalia and atrialia are pinular hexactins with a spindle­like pinular ray, the other rays are conically pointed or rounded, usually rough, sometimes smooth. The pinular ray of dermal hexactins is 0.066 0.143/
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, tangential rays are
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<geoCoordinate id="EE053E89487B1574FD9B084B10FAF8DF" box="[589,654,1794,1820]" degrees="0.041" direction="north" orientation="latitude" pageId="57" pageNumber="64" precision="55" value="0.041">0.041</geoCoordinate>
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mm
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, the proximal ray is
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<geoCoordinate id="EE053E89487B1574FBC7084B1626F8DF" box="[1041,1106,1794,1820]" degrees="0.041" direction="north" orientation="latitude" pageId="57" pageNumber="64" precision="55" value="0.041">0.041</geoCoordinate>
<geoCoordinate id="EE053E89487B1574FB84084B16D0F8DF" box="[1106,1188,1794,1820]" degrees="0.077" direction="west" orientation="longitude" pageId="57" pageNumber="64" precision="55" value="-0.077">0.077</geoCoordinate>
mm
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, the diameter of these rays is
<quantity id="4CC9F5AB487B1574FEF90863138FF887" box="[303,507,1834,1860]" metricMagnitude="-6" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="57" pageNumber="64" unit="mm" value="0.0025" valueMax="0.003" valueMin="0.002">
<geoCoordinate id="EE053E89487B1574FEF908631305F887" box="[303,369,1834,1860]" degrees="0.002" direction="north" orientation="latitude" pageId="57" pageNumber="64" precision="55" value="0.002">0.002</geoCoordinate>
<geoCoordinate id="EE053E89487B1574FEA4086313B6F887" box="[370,450,1834,1860]" degrees="0.003" direction="west" orientation="longitude" pageId="57" pageNumber="64" precision="55" value="-0.003">0.003</geoCoordinate>
mm
</quantity>
. The pinular ray of atrial hexactin is
<quantity id="4CC9F5AB487B1574FC670863160DF887" box="[945,1145,1834,1860]" metricMagnitude="-4" metricUnit="m" metricValue="1.48" metricValueMax="2.04" metricValueMin="0.9199999999999999" pageId="57" pageNumber="64" unit="mm" value="0.148" valueMax="0.204" valueMin="0.092">
<geoCoordinate id="EE053E89487B1574FC6708631187F887" box="[945,1011,1834,1860]" degrees="0.092" direction="north" orientation="latitude" pageId="57" pageNumber="64" precision="55" value="0.092">0.092</geoCoordinate>
<geoCoordinate id="EE053E89487B1574FC2508631630F887" box="[1011,1092,1834,1860]" degrees="0.204" direction="west" orientation="longitude" pageId="57" pageNumber="64" precision="55" value="-0.204">0.204</geoCoordinate>
mm
</quantity>
long, tangential rays are
<quantity id="4CC9F5AB487B1574FF41081B1316F8AF" box="[151,354,1874,1900]" metricMagnitude="-5" metricUnit="m" metricValue="5.6" metricValueMax="7.1" metricValueMin="4.1" pageId="57" pageNumber="64" unit="mm" value="0.055999999999999994" valueMax="0.071" valueMin="0.041">
<geoCoordinate id="EE053E89487B1574FF41081B12ADF8AF" box="[151,217,1874,1900]" degrees="0.041" direction="north" orientation="latitude" pageId="57" pageNumber="64" precision="55" value="0.041">0.041</geoCoordinate>
<geoCoordinate id="EE053E89487B1574FF0E081B135EF8AF" box="[216,298,1874,1900]" degrees="0.071" direction="west" orientation="longitude" pageId="57" pageNumber="64" precision="55" value="-0.071">0.071</geoCoordinate>
mm
</quantity>
, the proximal ray is
<quantity id="4CC9F5AB487B1574FD84081B1169F8AF" box="[594,797,1874,1900]" metricMagnitude="-5" metricUnit="m" metricValue="6.65" metricValueMax="8.2" metricValueMin="5.1" pageId="57" pageNumber="64" unit="mm" value="0.0665" valueMax="0.082" valueMin="0.051">
<geoCoordinate id="EE053E89487B1574FD84081B10E7F8AF" box="[594,659,1874,1900]" degrees="0.051" direction="north" orientation="latitude" pageId="57" pageNumber="64" precision="55" value="0.051">0.051</geoCoordinate>
<geoCoordinate id="EE053E89487B1574FD44081B1090F8AF" box="[658,740,1874,1900]" degrees="0.082" direction="west" orientation="longitude" pageId="57" pageNumber="64" precision="55" value="-0.082">0.082</geoCoordinate>
mm
</quantity>
.
</paragraph>
<caption id="DF4E08C648781577FF410A431634FAE2" ID-DOI="http://doi.org/10.5281/zenodo.5133367" ID-Zenodo-Dep="5133367" box="[151,1088,1290,1314]" httpUri="https://zenodo.org/record/5133367/files/figure.png" pageId="58" pageNumber="65" startId="58.[151,255,1290,1314]" targetBox="[151,1436,181,1264]" targetPageId="58">
<paragraph id="8B8E584E48781577FF410A431634FAE2" blockId="58.[151,1088,1290,1314]" box="[151,1088,1290,1314]" pageId="58" pageNumber="65">
<emphasis id="B945845C48781577FF410A431352FAE1" bold="true" box="[151,294,1290,1314]" pageId="58" pageNumber="65">FIGURE 32.</emphasis>
<taxonomicName id="4C3123CD48781577FEFB0A43103AFAE2" authority="Tabachnick &amp; Menschenina, 2008" authorityName="Tabachnick &amp; Menschenina" authorityYear="2008" box="[301,590,1290,1313]" class="Hexactinellida" family="Rossellidae" genus="Sympagella" kingdom="Animalia" order="Lyssacinosida" pageId="58" pageNumber="65" phylum="Porifera" rank="species" species="multihexastera" status="sp. nov.">
<emphasis id="B945845C48781577FEFB0A43103AFAE2" box="[301,590,1290,1313]" italics="true" pageId="58" pageNumber="65">Sympagella multihexastera</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A276392748781577FD8A0A4310E4FAE2" box="[604,656,1290,1313]" pageId="58" pageNumber="65" rank="species">sp.n.</taxonomicNameLabel>
, holotype. Dermal pinules, scale 30 µm.
</paragraph>
</caption>
<paragraph id="8B8E584E48781577FF100A1C17E8F9F4" blockId="58.[151,1436,1365,2031]" pageId="58" pageNumber="65">
Microscleres: Microscleres are discohexasters, rarely onychohexasters, oxyhexasters, rarely oxyhemihexasters, oxyhexactins and strobiloplumicomes. The discohexasters, with 2, rarely 3 rays, are uncommon; they are
<quantity id="4CC9F5AB48781577FF140AEC13FFFA7C" box="[194,395,1445,1471]" metricMagnitude="-4" metricUnit="m" metricValue="1.03" metricValueMax="1.05" metricValueMin="1.01" pageId="58" pageNumber="65" unit="mm" value="0.103" valueMax="0.105" valueMin="0.101">
<geoCoordinate id="EE053E8948781577FF140AEC1377FA7C" box="[194,259,1445,1471]" degrees="0.101" direction="north" orientation="latitude" pageId="58" pageNumber="65" precision="55" value="0.101">0.101</geoCoordinate>
<geoCoordinate id="EE053E8948781577FED50AEC1321FA7C" box="[259,341,1445,1471]" degrees="0.105" direction="west" orientation="longitude" pageId="58" pageNumber="65" precision="55" value="-0.105">0.105</geoCoordinate>
mm
</quantity>
in diameter with a primary rosette
<quantity id="4CC9F5AB48781577FCFD0AEC1187FA7C" box="[811,1011,1445,1471]" metricMagnitude="-5" metricUnit="m" metricValue="1.25" metricValueMax="1.7" metricValueMin="0.8" pageId="58" pageNumber="65" unit="mm" value="0.0125" valueMax="0.017" valueMin="0.008">
<geoCoordinate id="EE053E8948781577FCFD0AEC1119FA7C" box="[811,877,1445,1471]" degrees="0.008" direction="north" orientation="latitude" pageId="58" pageNumber="65" precision="55" value="0.008">0.008</geoCoordinate>
<geoCoordinate id="EE053E8948781577FCBB0AEC11CAFA7C" box="[877,958,1445,1471]" degrees="0.017" direction="west" orientation="longitude" pageId="58" pageNumber="65" precision="55" value="-0.017">0.017</geoCoordinate>
mm
</quantity>
in diameter. The single onychohexaster found is
<quantity id="4CC9F5AB48781577FEED0A8413C5FA24" box="[315,433,1485,1511]" metricMagnitude="-5" metricUnit="m" metricValue="6.3" pageId="58" pageNumber="65" unit="mm" value="0.063">0.063 mm</quantity>
in diameter with a primary rosette
<quantity id="4CC9F5AB48781577FC9A0A8411B7FA24" box="[844,963,1485,1511]" metricMagnitude="-6" metricUnit="m" metricValue="8.0" pageId="58" pageNumber="65" unit="mm" value="0.008">0.008 mm</quantity>
in diameter. The microscleres with oxyoidal secondary rays are
<quantity id="4CC9F5AB48781577FE150ABC10F9F9CC" box="[451,653,1525,1551]" metricMagnitude="-5" metricUnit="m" metricValue="9.45" metricValueMax="11.3" metricValueMin="7.6" pageId="58" pageNumber="65" unit="mm" value="0.0945" valueMax="0.113" valueMin="0.076">
<geoCoordinate id="EE053E8948781577FE150ABC1071F9CC" box="[451,517,1525,1551]" degrees="0.076" direction="north" orientation="latitude" pageId="58" pageNumber="65" precision="55" value="0.076">0.076</geoCoordinate>
<geoCoordinate id="EE053E8948781577FDD30ABC1020F9CC" box="[517,596,1525,1551]" degrees="0.113" direction="west" orientation="longitude" pageId="58" pageNumber="65" precision="55" value="-0.113">0.113</geoCoordinate>
mm
</quantity>
in diameter with a primary rosette
<quantity id="4CC9F5AB48781577FBEF0ABC1777F9CC" box="[1081,1283,1525,1551]" metricMagnitude="-5" metricUnit="m" metricValue="1.25" metricValueMax="1.7" metricValueMin="0.8" pageId="58" pageNumber="65" unit="mm" value="0.0125" valueMax="0.017" valueMin="0.008">
<geoCoordinate id="EE053E8948781577FBEF0ABC160FF9CC" box="[1081,1147,1525,1551]" degrees="0.008" direction="north" orientation="latitude" pageId="58" pageNumber="65" precision="55" value="0.008">0.008</geoCoordinate>
<geoCoordinate id="EE053E8948781577FBAD0ABC16B8F9CC" box="[1147,1228,1525,1551]" degrees="0.017" direction="west" orientation="longitude" pageId="58" pageNumber="65" precision="55" value="-0.017">0.017</geoCoordinate>
mm
</quantity>
in diameter. The strobiloplumicomes are
<quantity id="4CC9F5AB48781577FE33095410DFF9F4" box="[485,683,1565,1591]" metricMagnitude="-5" metricUnit="m" metricValue="4.6" metricValueMax="6.3" metricValueMin="2.9" pageId="58" pageNumber="65" unit="mm" value="0.046" valueMax="0.063" valueMin="0.029">
<geoCoordinate id="EE053E8948781577FE3309541053F9F4" box="[485,551,1565,1591]" degrees="0.029" direction="north" orientation="latitude" pageId="58" pageNumber="65" precision="55" value="0.029">0.029</geoCoordinate>
<geoCoordinate id="EE053E8948781577FDF109541003F9F4" box="[551,631,1565,1591]" degrees="0.063" direction="west" orientation="longitude" pageId="58" pageNumber="65" precision="55" value="-0.063">0.063</geoCoordinate>
mm
</quantity>
in diameter with a primary rosette
<quantity id="4CC9F5AB48781577FB920954177FF9F4" box="[1092,1291,1565,1591]" metricMagnitude="-5" metricUnit="m" metricValue="1.9" metricValueMax="2.5" metricValueMin="1.3" pageId="58" pageNumber="65" unit="mm" value="0.019" valueMax="0.025" valueMin="0.013">
<geoCoordinate id="EE053E8948781577FB92095416F2F9F4" box="[1092,1158,1565,1591]" degrees="0.013" direction="north" orientation="latitude" pageId="58" pageNumber="65" precision="55" value="0.013">0.013</geoCoordinate>
<geoCoordinate id="EE053E8948781577FB50095416A3F9F4" box="[1158,1239,1565,1591]" degrees="0.025" direction="west" orientation="longitude" pageId="58" pageNumber="65" precision="55" value="-0.025">0.025</geoCoordinate>
mm
</quantity>
in diameter.
</paragraph>
<paragraph id="8B8E584E48781577FF10090C1073F804" blockId="58.[151,1436,1365,2031]" pageId="58" pageNumber="65">
<emphasis id="B945845C48781577FF10090C1341F99D" box="[198,309,1605,1630]" italics="true" pageId="58" pageNumber="65">Remarks.</emphasis>
The new species
<emphasis id="B945845C48781577FDC2090C10A1F99D" box="[532,725,1605,1630]" italics="true" pageId="58" pageNumber="65">multihexastera</emphasis>
is erected according to its specific combination of microscleres: discohexasters, onychohexasters, oxyhexasters, oxyhexactins and strobiloplumicomes (the latter are characteristic for all species of the genus). Probably the closest species with most similar microscleres is
<taxonomicName id="4C3123CD48781577FA5009DC102BF914" authority="Tabachnick and Levi, 2004" authorityName="Tabachnick and Levi" authorityYear="2004" class="Hexactinellida" family="Rossellidae" genus="Sympagella" kingdom="Animalia" order="Lyssacinosida" pageId="58" pageNumber="65" phylum="Porifera" rank="species" species="clavipinula">
<emphasis id="B945845C48781577FA5009DC136EF915" italics="true" pageId="58" pageNumber="65">S. clavipinula</emphasis>
<bibRefCitation id="EFA025BF48781577FEF409F4102BF914" author="Tabachnick, K. R. &amp; Levi, C." box="[290,607,1725,1751]" pageId="58" pageNumber="65" pagination="11 - 71" refId="ref36646" refString="Tabachnick, K. R. &amp; Levi, C. (2004) Lyssacinosa du Pacifique sud-ouest (Porifera: Hexactinellida). Memoires du Museum National d'Histoire Naturelle, 191, 11 - 71." type="journal article" year="2004">Tabachnick and Levi, 2004</bibRefCitation>
</taxonomicName>
from off
<collectingCountry id="F32618DE48781577FD0609F411FDF914" box="[720,905,1725,1751]" name="New Caledonia" pageId="58" pageNumber="65">New Caledonia</collectingCountry>
, but this species has no oxyhexactins, dermal pinular rays are clavate in shape and atrial pinular hexactins differ strongly from the dermal ones in shape and length of their pinular rays. Two other species which possess similar dermal and atrial spicules are
<taxonomicName id="4C3123CD48781577FACC08441324F88C" authority="(Schulze, 1887)" baseAuthorityName="Schulze" baseAuthorityYear="1887" class="Hexactinellida" family="Rossellidae" genus="Sympagella" kingdom="Animalia" order="Lyssacinosida" pageId="58" pageNumber="65" phylum="Porifera" rank="species" species="johnsoni">
<emphasis id="B945845C48781577FACC084417E8F8E5" box="[1306,1436,1805,1830]" italics="true" pageId="58" pageNumber="65">S. johnsoni</emphasis>
(
<bibRefCitation id="EFA025BF48781577FF49087C133DF88C" author="Schulze, F. E." box="[159,329,1845,1871]" pageId="58" pageNumber="65" pagination="1 - 514" refId="ref36092" refString="Schulze, F. E. (1887) Report on the Hexactinellida collected by H. M. S. ' Challenger' during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. ' Challenger', 1873 - 1876. Zoology, 21, 1 - 514." type="journal article" year="1887">Schulze, 1887</bibRefCitation>
)
</taxonomicName>
(formerly
<taxonomicName id="4C3123CD48781577FE04087C1048F88D" authorityName="Schulze" authorityYear="1886" box="[466,572,1845,1870]" class="Hexactinellida" family="Rossellidae" genus="Aulascus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lyssacinosida" pageId="58" pageNumber="65" phylum="Porifera" rank="genus">
<emphasis id="B945845C48781577FE04087C1048F88D" box="[466,572,1845,1870]" italics="true" pageId="58" pageNumber="65">Aulascus</emphasis>
</taxonomicName>
) from off
<collectingCountry id="F32618DE48781577FD6E087C1124F88C" box="[696,848,1845,1871]" name="South Africa" pageId="58" pageNumber="65">South Africa</collectingCountry>
and
<taxonomicName id="4C3123CD48781577FC5D087C16CAF88C" authority="(Schulze, 1903)" baseAuthorityName="Schulze" baseAuthorityYear="1903" box="[907,1214,1845,1871]" class="Hexactinellida" family="Rossellidae" genus="Sympagella" kingdom="Animalia" order="Lyssacinosida" pageId="58" pageNumber="65" phylum="Porifera" rank="species" species="gracile">
<emphasis id="B945845C48781577FC5D087C1188F88D" box="[907,1020,1845,1870]" italics="true" pageId="58" pageNumber="65">S. gracile</emphasis>
(Schulze, 1903)
</taxonomicName>
(formerly
<taxonomicName id="4C3123CD48781577FAE9087C12A1F8B5" authorityName="Schulze" authorityYear="1899" class="Hexactinellida" family="Rossellidae" genus="Calycosoma" kingdom="Animalia" order="Lyssacinosida" pageId="58" pageNumber="65" phylum="Porifera" rank="genus">
<emphasis id="B945845C48781577FAE9087C12A1F8B5" italics="true" pageId="58" pageNumber="65">Calycosoma</emphasis>
</taxonomicName>
) from the Indonesian Archipelago. The difference here is in the microsclere composition:
<taxonomicName id="4C3123CD48781577FAC1081417E8F8B5" baseAuthorityName="Schulze" baseAuthorityYear="1887" box="[1303,1436,1885,1910]" class="Hexactinellida" family="Rossellidae" genus="Sympagella" kingdom="Animalia" order="Lyssacinosida" pageId="58" pageNumber="65" phylum="Porifera" rank="species" species="johnsoni">
<emphasis id="B945845C48781577FAC1081417E8F8B5" box="[1303,1436,1885,1910]" italics="true" pageId="58" pageNumber="65">S. johnsoni</emphasis>
</taxonomicName>
has (besides the characteristic strobiloplumicomes) discohexasters and onychohexasters,
<taxonomicName id="4C3123CD48781577FB6C08CC1759F85D" baseAuthorityName="Schulze" baseAuthorityYear="1903" box="[1210,1325,1925,1950]" class="Hexactinellida" family="Rossellidae" genus="Sympagella" kingdom="Animalia" order="Lyssacinosida" pageId="58" pageNumber="65" phylum="Porifera" rank="species" species="gracile">
<emphasis id="B945845C48781577FB6C08CC1759F85D" box="[1210,1325,1925,1950]" italics="true" pageId="58" pageNumber="65">S. gracile</emphasis>
</taxonomicName>
has oxyhexasters and onychohexasters.
</paragraph>
<paragraph id="8B8E584E48781576FF10089C1033FD21" blockId="58.[151,1436,1365,2031]" lastBlockId="59.[151,1437,152,738]" lastPageId="59" lastPageNumber="66" pageId="58" pageNumber="65">
<emphasis id="B945845C48781577FF10089C1333F82D" box="[198,327,2005,2030]" italics="true" pageId="58" pageNumber="65">Discussion</emphasis>
. Intraspecific variability and key to subspecies of
<emphasis id="B945845C48781577FC72089C16B2F82D" box="[932,1222,2005,2030]" italics="true" pageId="58" pageNumber="65">
<taxonomicName id="4C3123CD48781577FC72089C16B6F82D" authorityName="Owen" authorityYear="1841" box="[932,1218,2005,2030]" class="Hexactinellida" family="Euplectellidae" genus="Euplectella" kingdom="Animalia" order="Lyssacinosida" pageId="58" pageNumber="65" phylum="Porifera" rank="species" species="aspergillum">Euplectella aspergillum</taxonomicName>
.
</emphasis>
The result of this revision is the new definition of
<taxonomicName id="4C3123CD48791576FDC40FD110C8FF72" authorityName="Owen" authorityYear="1841" box="[530,700,152,177]" class="Hexactinellida" family="Euplectellidae" genus="Euplectella" kingdom="Animalia" order="Lyssacinosida" pageId="59" pageNumber="66" phylum="Porifera" rank="species" species="aspergillum">
<emphasis id="B945845C48791576FDC40FD110C8FF72" box="[530,700,152,177]" italics="true" pageId="59" pageNumber="66">E. aspergillum</emphasis>
</taxonomicName>
as
<taxonomicName id="4C3123CD48791576FD350FD1111DFF72" authorityName="Owen" authorityYear="1841" box="[739,873,152,177]" class="Hexactinellida" family="Euplectellidae" genus="Euplectella" kingdom="Animalia" order="Lyssacinosida" pageId="59" pageNumber="66" phylum="Porifera" rank="genus">
<emphasis id="B945845C48791576FD350FD1111DFF72" box="[739,873,152,177]" italics="true" pageId="59" pageNumber="66">Euplectella</emphasis>
</taxonomicName>
having principalia of stauractins, notable fusion of the choanosomal skeleton, formation of ridges, microscleres as floricomes, oxyhexasters and their derivatives, including abnormal oxyoidal forms. Furthermore, usually also graphiocomes and sometimes sigmatocomes (which are probably young stages of floricomes) occur. The four subspecies are justified by their morphological differences as well as the fact that they inhabit distal geographical locations:
<taxonomicName id="4C3123CD48791576FB380E71131BFEB9" authority="aspergillum Owen" authorityName="aspergillum Owen" authorityYear="1841" class="Hexactinellida" family="Euplectellidae" genus="Euplectella" kingdom="Animalia" order="Lyssacinosida" pageId="59" pageNumber="66" phylum="Porifera" rank="subSpecies" species="aspergillum" subSpecies="aspergillum">
<emphasis id="B945845C48791576FB380E711356FEBA" italics="true" pageId="59" pageNumber="66">E. aspergillum aspergillum</emphasis>
Owen
</taxonomicName>
off the
<collectingCountry id="F32618DE48791576FE360E29101CFEB9" box="[480,616,352,378]" name="Philippines" pageId="59" pageNumber="66">Philippines</collectingCountry>
;
<taxonomicName id="4C3123CD48791576FDA20E2911AFFEB9" authority="Schulze" authorityName="Schulze" authorityYear="1900" box="[628,987,352,378]" class="Hexactinellida" family="Euplectellidae" genus="Euplectella" kingdom="Animalia" order="Lyssacinosida" pageId="59" pageNumber="66" phylum="Porifera" rank="subSpecies" species="aspergillum" subSpecies="regalis">
<emphasis id="B945845C48791576FDA20E291102FEBA" box="[628,886,352,377]" italics="true" pageId="59" pageNumber="66">E. aspergillum regalis</emphasis>
Schulze
</taxonomicName>
Sea of Bengal;
<taxonomicName id="4C3123CD48791576FB7D0E291283FE62" authorityName="Tabachnick, Janussen &amp; Menschenina" authorityYear="2008" class="Hexactinellida" family="Euplectellidae" genus="Euplectella" kingdom="Animalia" order="Lyssacinosida" pageId="59" pageNumber="66" phylum="Porifera" rank="subSpecies" species="aspergillum" subSpecies="indonesicum">
<emphasis id="B945845C48791576FB7D0E291283FE62" italics="true" pageId="59" pageNumber="66">E. aspergillum indonesicum</emphasis>
</taxonomicName>
ssp. nov. Indonesian Archipelago;
<taxonomicName id="4C3123CD48791576FD730EC11190FE62" authority="Tabachnick &amp; Menschenina, 2008" authorityName="Tabachnick &amp; Menschenina" authorityYear="2008" box="[677,996,392,417]" class="Hexactinellida" family="Euplectellidae" genus="Euplectella" kingdom="Animalia" order="Lyssacinosida" pageId="59" pageNumber="66" phylum="Porifera" rank="subSpecies" species="aspergillum" status="subsp. nov." subSpecies="australicum">
<emphasis id="B945845C48791576FD730EC11190FE62" box="[677,996,392,417]" italics="true" pageId="59" pageNumber="66">E. aspergillum australicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A276392748791576FC3C0EC1163BFE61" box="[1002,1103,392,418]" pageId="59" pageNumber="66" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
off W
<collectingCountry id="F32618DE48791576FB600EC11751FE61" box="[1206,1317,392,418]" name="Australia" pageId="59" pageNumber="66">Australia</collectingCountry>
. Two subspecies are known from many specimens:
<emphasis id="B945845C48791576FD700EF91130FE0A" box="[678,836,432,457]" italics="true" pageId="59" pageNumber="66">aspergillum</emphasis>
and
<emphasis id="B945845C48791576FC510EF91653FE0A" box="[903,1063,432,457]" italics="true" pageId="59" pageNumber="66">australicum</emphasis>
, the other two only by single representatives. Their identification is based on the detailed investigation of oscularia (spicules located close to the lateral oscula) and of their microscleres: floricomes and oxyhexasters. No overlaps of important features for their identification was observed, therefore the identification key presented here seems justified. The other species of
<taxonomicName id="4C3123CD48791576FE8B0D191396FDAA" authorityName="Owen" authorityYear="1841" box="[349,482,592,617]" class="Hexactinellida" family="Euplectellidae" genus="Euplectella" kingdom="Animalia" order="Lyssacinosida" pageId="59" pageNumber="66" phylum="Porifera" rank="genus">
<emphasis id="B945845C48791576FE8B0D191396FDAA" box="[349,482,592,617]" italics="true" pageId="59" pageNumber="66">Euplectella</emphasis>
</taxonomicName>
, which show principalia of stauractins, require re­examination: It is very likely that
<taxonomicName id="4C3123CD48791576FF1C0D3110F4FD51" authority="Herklots and Marshal, 1868" authorityName="Herklots and Marshall" authorityYear="1868" box="[202,640,632,658]" class="Hexactinellida" family="Euplectellidae" genus="Euplectella" kingdom="Animalia" order="Lyssacinosida" pageId="59" pageNumber="66" phylum="Porifera" rank="species" species="oweni">
<emphasis id="B945845C48791576FF1C0D311344FD52" box="[202,304,632,657]" italics="true" pageId="59" pageNumber="66">E. oweni</emphasis>
Herklots and Marshal, 1868
</taxonomicName>
,
<taxonomicName id="4C3123CD48791576FD5A0D3111DEFD51" authority="Ijima, 1895" authorityName="Ijima" authorityYear="1895" box="[652,938,632,658]" class="Hexactinellida" family="Euplectellidae" genus="Euplectella" kingdom="Animalia" order="Lyssacinosida" pageId="59" pageNumber="66" phylum="Porifera" rank="species" species="marshalli">
<emphasis id="B945845C48791576FD5A0D311169FD52" box="[652,797,632,657]" italics="true" pageId="59" pageNumber="66">E. marshalli</emphasis>
Ijima, 1895
</taxonomicName>
and
<taxonomicName id="4C3123CD48791576FC350D311757FD51" authority="Ijima, 1901" authorityName="Ijima" authorityYear="1901" box="[995,1315,632,658]" class="Hexactinellida" family="Euplectellidae" genus="Euplectella" kingdom="Animalia" order="Lyssacinosida" pageId="59" pageNumber="66" phylum="Porifera" rank="species" species="curvistellata">
<emphasis id="B945845C48791576FC350D3116E2FD52" box="[995,1174,632,657]" italics="true" pageId="59" pageNumber="66">E. curvistellata</emphasis>
<bibRefCitation id="EFA025BF48791576FB4B0D311757FD51" author="Ijima, I." box="[1181,1315,632,658]" pageId="59" pageNumber="66" pagination="1 - 299" refId="ref35574" refString="Ijima, I. (1901) Studies on the Hexactinellida. Contribution I. (Euplectellidae). Journal of the College of Sciences, Imperial University of Tokyo, 15, 1 - 299." type="journal article" year="1901">Ijima, 1901</bibRefCitation>
</taxonomicName>
should be synonymized; as well as
<taxonomicName id="4C3123CD48791576FE1E0DE9108EFD79" authority="Ijima, 1894" authorityName="Ijima" authorityYear="1894" box="[456,762,672,698]" class="Hexactinellida" family="Euplectellidae" genus="Euplectella" kingdom="Animalia" order="Lyssacinosida" pageId="59" pageNumber="66" phylum="Porifera" rank="species" species="imperialis">
<emphasis id="B945845C48791576FE1E0DE91017FD7A" box="[456,611,672,697]" italics="true" pageId="59" pageNumber="66">E. imperialis</emphasis>
Ijima, 1894
</taxonomicName>
,
<taxonomicName id="4C3123CD48791576FCDF0DE91635FD79" authority=", Ijima, 1927" authorityName=", Ijima" authorityYear="1927" box="[777,1089,672,698]" class="Hexactinellida" family="Euplectellidae" genus="Euplectella" kingdom="Animalia" order="Lyssacinosida" pageId="59" pageNumber="66" phylum="Porifera" rank="species" species="timorensis">
<emphasis id="B945845C48791576FCDF0DE911D2FD7A" box="[777,934,672,697]" italics="true" pageId="59" pageNumber="66">E. timorensis</emphasis>
,
<bibRefCitation id="EFA025BF48791576FC610DE91635FD79" author="Ijima, I." box="[951,1089,672,698]" pageId="59" pageNumber="66" pagination="1 - 383" refId="ref35659" refString="Ijima, I. (1927) The Hexactinellida of the Siboga Expedition. In: Weber, M. (Ed.), Siboga-Expeditie. Uitkomsten op zoologisch, botanisch, oceanographisch en geologisch gebied verzameld in Nederlandsch Oost-lndie 1899 - 1900 aan boord H. M. ' Siboga' Monographie VI. (E. J. Brill: Leiden), i - viii, 1 - 383." type="journal article" year="1927">Ijima, 1927</bibRefCitation>
</taxonomicName>
and specimens of
<taxonomicName id="4C3123CD48791576FAFE0DE917E8FD7A" authorityName="Schulze" authorityYear="1900" box="[1320,1436,672,697]" class="Hexactinellida" family="Euplectellidae" genus="Euplectella" kingdom="Animalia" order="Lyssacinosida" pageId="59" pageNumber="66" phylum="Porifera" rank="species" species="regalis">
<emphasis id="B945845C48791576FAFE0DE917E8FD7A" box="[1320,1436,672,697]" italics="true" pageId="59" pageNumber="66">E. regalis</emphasis>
</taxonomicName>
from off S
<collectingCountry id="F32618DE48791576FECE0D8113F7FD21" box="[280,387,712,738]" name="Australia" pageId="59" pageNumber="66">Australia</collectingCountry>
(
<bibRefCitation id="EFA025BF48791576FE440D81104FFD21" author="Reiswig, H. M." box="[402,571,712,738]" pageId="59" pageNumber="66" pagination="25 - 36" refId="ref35985" refString="Reiswig, H. M. (1992) First Hexactinellida (Porifera) (glass sponges) from the Great Australian Bight. Records of the South Australian Museum, 26 (1), 25 - 36." type="journal article" year="1992">Reiswig 1992</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>