treatments-xml/data/03/98/36/03983615FFD1FA54FF34F897FCD8F79D.xml

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<mods:title id="F02C1AB4E2B30B22A42779983EB95A1F">A new species of Speleogobius (Teleostei: Gobiidae) from the Western Mediterraenean Sea</mods:title>
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<paragraph id="8B8E8703FFD1FA5DFF34F897FCDDF0EE" blockId="1.[151,892,1904,1964]" box="[151,892,1904,1931]" pageId="1" pageNumber="302">
<heading id="D0C6306FFFD1FA5DFF34F897FCDDF0EE" bold="true" box="[151,892,1904,1931]" fontSize="11" level="1" pageId="1" pageNumber="302" reason="1">
<emphasis id="B9455B11FFD1FA5DFF34F897FCDDF0EE" bold="true" box="[151,892,1904,1931]" pageId="1" pageNumber="302">
<taxonomicName id="4C31FC80FFD1FA5DFF34F897FCA9F0EE" authority="Kovacic, Ordines &amp; Schliewen" authorityName="Kovacic, Ordines &amp; Schliewen" box="[151,776,1904,1931]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="302" phylum="Chordata" rank="species" species="llorisi" status="sp. nov.">
<emphasis id="B9455B11FFD1FA5DFF34F897FE22F0EF" bold="true" box="[151,387,1904,1930]" italics="true" pageId="1" pageNumber="302">Speleogobius llorisi</emphasis>
Kovačić, Ordines &amp; Schliewen
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, 
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<figureCitation id="130A9B86FFD1FA5DFF34F874FEA1F0C9" box="[151,256,1939,1964]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="4.[151,250,1557,1579]" captionStartId-1="5.[151,250,910,932]" captionStartId-2="5.[151,250,1628,1650]" captionTargetBox-0="[151,1436,780,1535]" captionTargetBox-1="[151,1435,517,888]" captionTargetBox-2="[179,1414,1022,1594]" captionTargetId-0="figure@4.[151,1436,780,1536]" captionTargetId-1="figure@5.[151,1436,517,889]" captionTargetId-2="figure@5.[173,1414,1012,1605]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="5" captionText-0="FIGURE 2. Speleogobius llorisi sp. nov. Holotype: PMR VP 3510 (P 3), female, 25.6 + 5.6 mm, Spain, Balearic Islands, Mallorca. A: freshly collected specimen; B: preserved specimen. Photos by (A) F. Ordines, (B) M. Kovačić." captionText-1="FIGURE 3. Speleogobius llorisi sp. nov. Paratype: PMR VP 3512 (P 54), male, 24.4 + 5.5 mm, Spain, Balearic Islands, Mallorca. Preserved specimen. Photo by M. Kovačić." captionText-2="FIGURE 4. Speleogobius llorisi sp. nov., head LL sensory papillae and canal pores: paratype, PMR VP 3512 (P 54), male, 24.4 + 5.5 mm, Spain, Balearic Islands, Mallorca. Terminology in text. Drawing by M. Kovačić." httpUri-0="https://zenodo.org/record/262959/files/figure.png" httpUri-1="https://zenodo.org/record/262960/files/figure.png" httpUri-2="https://zenodo.org/record/262961/files/figure.png" pageId="1" pageNumber="302">Figs. 2–4</figureCitation>
</paragraph>
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<emphasis id="B9455B11FFD1FA5DFF34F83BFEA5F090" bold="true" box="[151,260,2012,2037]" pageId="1" pageNumber="302">Holotype</emphasis>
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. 
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<collectionCode id="ED201FC6FFD1FA5DFEF6F83BFEDBF090" box="[341,378,2012,2037]" pageId="1" pageNumber="302">VP</collectionCode>
3510, female, 25.6 + 
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, 
<collectingCountry id="F326C793FFD1FA5DFD6DF83BFCB0F090" box="[718,785,2012,2037]" name="Spain" pageId="1" pageNumber="302">Spain</collectingCountry>
, Balearic Islands, Mallorca, 
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, 
<geoCoordinate id="EE05E1C4FFD1FA5DFB7DF83BFACCF090" box="[1246,1389,2012,2037]" direction="east" orientation="longitude" pageId="1" pageNumber="302" precision="9" value="2.9776666">002º58.66’E</geoCoordinate>
, red algae beds, 
<quantity id="4CC92AE6FFD2FA5EFEBDFF70FEF8F7CA" box="[286,345,151,176]" metricMagnitude="1" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="303" unit="m" value="67.0">67 m</quantity>
depth, coll. Instituto Español de Oceanografía-Centre Oceanogràfic de Balears (IEO-COB), 
<date id="FF8FA1C3FFD2FA5EFA23FF7FFEB4F7B0" pageId="2" pageNumber="303" value="2015-04-13">13 April 2015</date>
. The collecting positions for the 
<typeStatus id="548A39A1FFD2FA5EFD27FF5AFD15F7B0" box="[644,692,189,213]" pageId="2" pageNumber="303">type</typeStatus>
material and for the additional material are presented in 
<figureCitation id="130A9B86FFD2FA5EFA93FF5BFAD3F7B1" box="[1328,1394,188,213]" captionStart="FIGURE 1" captionStartId="1.[151,250,1285,1307]" captionTargetBox="[160,1419,851,1259]" captionTargetId="figure@1.[160,1426,844,1264]" captionTargetPageId="1" captionText="FIGURE 1. Map showing the sampling stations where the specimens were collected (black dots). Isobaths of 50, 100 and 200 m are also shown. The holotype was collected at the station numbered 9. The paratypes PMR VP 3511, ZSM 43988, PMR VP 3512, PMR VP 3513 and ZSM 43989, and the specimens included as additional material ZSM 43990, ZSM-PIS-GO- 1065 / 1066, ZSM 43991 and ZSM-PIS-GO- 1067 / 1068, were collected from stations numbered 1, 4, 6, 5, 8, 3, 2, 7 and 9, respectively." httpUri="https://zenodo.org/record/262958/files/figure.png" pageId="2" pageNumber="303">Fig. 1</figureCitation>
.
</paragraph>
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<typeStatus id="548A39A1FFD2FA5EFF64FF38FEE0F79D" box="[199,321,223,248]" pageId="2" pageNumber="303" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED201FC6FFD2FA5EFEF1FF07FE2CF792" box="[338,397,224,247]" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="2" pageNumber="303">PMR</collectionCode>
<collectionCode id="ED201FC6FFD2FA5EFE39FF38FE1EF79D" box="[410,447,223,248]" pageId="2" pageNumber="303">VP</collectionCode>
3511, female, 
<quantity id="4CC92AE6FFD2FA5EFDCAFF07FD72F792" box="[617,723,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="2" pageNumber="303" unit="mm" value="18.7">18.7 mm</quantity>
SL, caudal fin damaged, 
<collectingCountry id="F326C793FFD2FA5EFBACFF38FBF3F79D" box="[1039,1106,223,248]" name="Spain" pageId="2" pageNumber="303">Spain</collectingCountry>
, Balearic Islands, Mallorca, 
<geoCoordinate id="EE05E1C4FFD2FA5EFF34FEE1FEBCF67B" box="[151,285,262,286]" direction="north" orientation="latitude" pageId="2" pageNumber="303" precision="9" value="39.505665">39°30.34’N</geoCoordinate>
, 
<geoCoordinate id="EE05E1C4FFD2FA5EFE89FEE2FE1AF67B" box="[298,443,261,286]" direction="east" orientation="longitude" pageId="2" pageNumber="303" precision="9" value="2.4171667">002°25.03’E</geoCoordinate>
, red algae beds, 
<quantity id="4CC92AE6FFD2FA5EFDD9FEE2FD14F678" box="[634,693,261,286]" metricMagnitude="1" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="303" unit="m" value="67.0">67 m</quantity>
depth, coll. IEO-COB, 
<date id="FF8FA1C3FFD2FA5EFC66FEE1FB04F67B" box="[965,1189,261,286]" pageId="2" pageNumber="303" value="2014-09-12">12 September 2014</date>
; 
<collectionCode id="ED201FC6FFD2FA5EFB11FEE1FB4DF67B" box="[1202,1260,262,286]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="303">ZSM</collectionCode>
43988, female, 
<quantity id="4CC92AE6FFD2FA5EFF34FECCFF5AF627" box="[151,251,299,323]" metricMagnitude="-2" metricUnit="m" metricValue="2.7199999999999998" pageId="2" pageNumber="303" unit="mm" value="27.2">27.2 mm</quantity>
SL, caudal fin damaged, 
<collectingCountry id="F326C793FFD2FA5EFDBEFECDFDC1F626" box="[541,608,298,323]" name="Spain" pageId="2" pageNumber="303">Spain</collectingCountry>
, Balearic Islands, Mallorca, 
<geoCoordinate id="EE05E1C4FFD2FA5EFC02FECCFB86F626" box="[929,1063,299,323]" direction="north" orientation="latitude" pageId="2" pageNumber="303" precision="9" value="39.368168">39°22.09’N</geoCoordinate>
, 
<geoCoordinate id="EE05E1C4FFD2FA5EFB90FECCFB65F626" box="[1075,1220,299,323]" direction="east" orientation="longitude" pageId="2" pageNumber="303" precision="9" value="2.6853333">002°41.12’E</geoCoordinate>
, red algae beds, 
<quantity id="4CC92AE6FFD2FA5EFA23FECDFF0CF602" metricMagnitude="1" metricUnit="m" metricValue="5.1" pageId="2" pageNumber="303" unit="m" value="51.0">51 m</quantity>
depth, coll. IEO-COB, 
<date id="FF8FA1C3FFD2FA5EFE6FFEB7FD13F60D" box="[460,690,335,360]" pageId="2" pageNumber="303" value="2014-09-12">12 September 2014</date>
; 
<collectionCode id="ED201FC6FFD2FA5EFD60FEB7FD5FF602" box="[707,766,336,359]" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="2" pageNumber="303">PMR</collectionCode>
<collectionCode id="ED201FC6FFD2FA5EFCA8FEA8FC91F60D" box="[779,816,335,360]" pageId="2" pageNumber="303">VP</collectionCode>
3512, male, 24.4 + 
<quantity id="4CC92AE6FFD2FA5EFBBFFEA8FBD8F602" box="[1052,1145,335,360]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="303" unit="mm" value="5.5">5.5 mm</quantity>
, 
<collectingCountry id="F326C793FFD2FA5EFB2AFEA8FB6DF60D" box="[1161,1228,335,360]" name="Spain" pageId="2" pageNumber="303">Spain</collectingCountry>
, Balearic Islands, Mallorca, 
<geoCoordinate id="EE05E1C4FFD2FA5EFEACFE92FE37F6EB" box="[271,406,373,398]" direction="north" orientation="latitude" pageId="2" pageNumber="303" precision="9" value="39.351">39°21.06’N</geoCoordinate>
, 
<geoCoordinate id="EE05E1C4FFD2FA5EFE04FE91FD99F6EB" box="[423,568,374,398]" direction="east" orientation="longitude" pageId="2" pageNumber="303" precision="9" value="2.6956668">002°41.74’E</geoCoordinate>
, red algae beds, 
<quantity id="4CC92AE6FFD2FA5EFCABFE92FCE6F6E8" box="[776,839,373,398]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="2" pageNumber="303" unit="m" value="56.0">56 m</quantity>
depth, coll. IEO-COB, 
<date id="FF8FA1C3FFD2FA5EFBCBFE91FAF1F6EB" box="[1128,1360,373,398]" pageId="2" pageNumber="303" value="2014-09-14">14 September 2014</date>
; 
<collectionCode id="ED201FC6FFD2FA5EFAC2FE91FA3DF6E8" box="[1377,1436,374,397]" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="2" pageNumber="303">PMR</collectionCode>
<collectionCode id="ED201FC6FFD2FA5EFF34FE7DFF1DF6D6" box="[151,188,410,435]" pageId="2" pageNumber="303">VP</collectionCode>
3513, female, 24.5 + 
<quantity id="4CC92AE6FFD2FA5EFE04FE7DFE5EF6D7" box="[423,511,410,435]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="2" pageNumber="303" unit="mm" value="5.6">5.6 mm</quantity>
, 
<collectingCountry id="F326C793FFD2FA5EFDA9FE7DFDECF6D6" box="[522,589,410,435]" name="Spain" pageId="2" pageNumber="303">Spain</collectingCountry>
, Balearic Islands, Mallorca, 
<geoCoordinate id="EE05E1C4FFD2FA5EFC29FE7CFBB1F6D6" box="[906,1040,411,435]" direction="north" orientation="latitude" pageId="2" pageNumber="303" precision="9" value="39.505665">39°30.34’N</geoCoordinate>
, 
<geoCoordinate id="EE05E1C4FFD2FA5EFBB8FE7DFB0DF6D6" box="[1051,1196,410,435]" direction="east" orientation="longitude" pageId="2" pageNumber="303" precision="9" value="2.4171667">002°25.03’E</geoCoordinate>
, red algae beds, 
<quantity id="4CC92AE6FFD2FA5EFAC7FE7DFA3DF6D7" box="[1380,1436,410,435]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="2" pageNumber="303" unit="m" value="46.0">46 m</quantity>
depth, coll. IEO-COB, 
<date id="FF8FA1C3FFD2FA5EFE02FE27FD22F6BD" box="[417,643,447,472]" pageId="2" pageNumber="303" value="2014-09-12">12 September 2014</date>
, 
<collectionCode id="ED201FC6FFD2FA5EFD33FE27FD6BF6BD" box="[656,714,448,472]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="303">ZSM</collectionCode>
43989, juvenile male, 14.3 + 4.0 mm, 
<collectingCountry id="F326C793FFD2FA5EFB2CFE58FB73F6BD" box="[1167,1234,447,472]" name="Spain" pageId="2" pageNumber="303">Spain</collectingCountry>
, Balearic Islands, Mallorca, 
<geoCoordinate id="EE05E1C4FFD2FA5EFEA9FE01FE2BF69B" box="[266,394,486,510]" direction="north" orientation="latitude" pageId="2" pageNumber="303" precision="9" value="39.238">39º14.28'N</geoCoordinate>
, 
<geoCoordinate id="EE05E1C4FFD2FA5EFE36FE02FD85F69B" box="[405,548,485,510]" direction="east" orientation="longitude" pageId="2" pageNumber="303" precision="9" value="2.9783335">002º58.70’E</geoCoordinate>
, red algae beds, 
<quantity id="4CC92AE6FFD2FA5EFD7EFE02FCB7F698" box="[733,790,485,510]" metricMagnitude="1" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="303" unit="m" value="67.0">67 m</quantity>
depth, coll. IEO-COB, 
<date id="FF8FA1C3FFD2FA5EFB82FE01FB63F69B" box="[1057,1218,485,510]" pageId="2" pageNumber="303" value="2015-04-13">13 April 2015</date>
.
</paragraph>
<paragraph id="8B8E8703FFD2FA5EFF64FDEDFB17F5BB" blockId="2.[151,1437,151,1928]" pageId="2" pageNumber="303">
<emphasis id="B9455B11FFD2FA5EFF64FDEDFE1EF546" bold="true" box="[199,447,522,547]" pageId="2" pageNumber="303">Additional material.</emphasis>
<collectionCode id="ED201FC6FFD2FA5EFE68FDECFDA4F546" box="[459,517,523,547]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="303">ZSM</collectionCode>
43990, undetermined sex, 17.7 + 
<quantity id="4CC92AE6FFD2FA5EFC00FDECFBA1F547" box="[931,1024,523,547]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="303" unit="mm" value="3.1">3.1 mm</quantity>
, 
<collectingCountry id="F326C793FFD2FA5EFBB2FDEDFBF5F546" box="[1041,1108,522,547]" name="Spain" pageId="2" pageNumber="303">Spain</collectingCountry>
, Balearic Islands, Mallorca, 
<geoCoordinate id="EE05E1C4FFD2FA5EFF34FDC8FEBCF52D" box="[151,285,559,584]" direction="north" orientation="latitude" pageId="2" pageNumber="303" precision="9" value="39.3675">39°22.05’N</geoCoordinate>
, 
<geoCoordinate id="EE05E1C4FFD2FA5EFE89FDD7FE68F52D" box="[298,457,560,584]" direction="east" orientation="longitude" pageId="2" pageNumber="303" precision="1" value="2.6548">002°39.288’E</geoCoordinate>
, red algae beds, 
<quantity id="4CC92AE6FFD2FA5EFD29FDC8FD64F522" box="[650,709,559,584]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="303" unit="m" value="55.0">55 m</quantity>
depth, coll. IEO-COB, 
<date id="FF8FA1C3FFD2FA5EFC75FDC8FB08F52D" box="[982,1193,559,584]" pageId="2" pageNumber="303" value="2014-09-05">5 September 2014</date>
; ZSM-PIS-GO-1065/ 1066, undetermined sex, 17.6 + 
<quantity id="4CC92AE6FFD2FA5EFDB3FDB1FDCBF508" box="[528,618,598,622]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="303" unit="mm" value="3.2">3.2 mm</quantity>
SL, 
<collectingCountry id="F326C793FFD2FA5EFD05FDB2FD48F50B" box="[678,745,597,622]" name="Spain" pageId="2" pageNumber="303">Spain</collectingCountry>
, Balearic Islands, Mallorca, 
<geoCoordinate id="EE05E1C4FFD2FA5EFB96FDB2FB1DF50B" box="[1077,1212,597,622]" direction="north" orientation="latitude" pageId="2" pageNumber="303" precision="9" value="39.4125">39°24.75’N</geoCoordinate>
, 
<geoCoordinate id="EE05E1C4FFD2FA5EFB68FDB2FACBF50B" box="[1227,1386,597,622]" direction="east" orientation="longitude" pageId="2" pageNumber="303" precision="1" value="2.5263">002°31.578’E</geoCoordinate>
, red algae beds, 
<quantity id="4CC92AE6FFD2FA5EFE87FD9DFEC0F5F7" box="[292,353,634,659]" metricMagnitude="1" metricUnit="m" metricValue="6.9" pageId="2" pageNumber="303" unit="m" value="69.0">69 m</quantity>
depth, coll. IEO-COB, 
<date id="FF8FA1C3FFD2FA5EFDDCFD9DFCF6F5F6" box="[639,855,634,659]" pageId="2" pageNumber="303" value="2014-09-05">5 September 2014</date>
; 
<collectionCode id="ED201FC6FFD2FA5EFCC4FD9CFC00F5F6" box="[871,929,635,659]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="303">ZSM</collectionCode>
43991, female, 23.8 + 
<quantity id="4CC92AE6FFD2FA5EFB1FFD9CFAB6F5F7" box="[1212,1303,635,659]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="2" pageNumber="303" unit="mm" value="4.9">4.9 mm</quantity>
SL, 
<collectingCountry id="F326C793FFD2FA5EFAF6FD9DFA39F5F6" box="[1365,1432,634,659]" name="Spain" pageId="2" pageNumber="303">Spain</collectingCountry>
, Balearic Islands, Mallorca, 
<geoCoordinate id="EE05E1C4FFD2FA5EFE6EFD78FDC0F5DD" box="[461,609,671,696]" direction="north" orientation="latitude" pageId="2" pageNumber="303" precision="1" value="39.2747">39°16.482’N</geoCoordinate>
, 
<geoCoordinate id="EE05E1C4FFD2FA5EFDCDFD78FCACF5DD" box="[622,781,671,696]" direction="east" orientation="longitude" pageId="2" pageNumber="303" precision="1" value="2.9513">002°57.078’E</geoCoordinate>
, red algae beds, 
<quantity id="4CC92AE6FFD2FA5EFC6AFD78FBA3F5D2" box="[969,1026,671,696]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="2" pageNumber="303" unit="m" value="56.0">56 m</quantity>
depth, coll. IEO-COB, 
<date id="FF8FA1C3FFD2FA5EFAACFD47FF72F5BB" pageId="2" pageNumber="303" value="2014-09-07">7 September 2014</date>
; ZSM-PIS-GO-1067/1068, undetermined sex, 21.1 + 
<quantity id="4CC92AE6FFD2FA5EFC8EFD21FC22F5B8" box="[813,899,710,734]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="303" unit="mm" value="4.1">4.1 mm</quantity>
SL, same data as 
<typeStatus id="548A39A1FFD2FA5EFBEEFD22FB12F5BB" box="[1101,1203,709,734]" pageId="2" pageNumber="303" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C32BD488FFD2FA5EFF64FD0DFA39F36D" pageId="2" pageNumber="303" type="diagnosis">
<paragraph id="8B8E8703FFD2FA5EFF64FD0DFA39F36D" blockId="2.[151,1437,151,1928]" pageId="2" pageNumber="303">
<emphasis id="B9455B11FFD2FA5EFF64FD0DFE9BF466" bold="true" box="[199,314,746,771]" pageId="2" pageNumber="303">Diagnosis</emphasis>
. (1) Preopercular head canal present with pores 
<emphasis id="B9455B11FFD2FA5EFCF4FD0CFC37F467" box="[855,918,747,771]" italics="true" pageId="2" pageNumber="303">γ, δ, ε</emphasis>
; (2) snout long, equal or longer than eye, 1.0– 
<quantity id="4CC92AE6FFD2FA5EFF34FCF7FF79F44D" box="[151,216,783,808]" metricMagnitude="-2" metricUnit="m" metricValue="2.794" pageId="2" pageNumber="303" unit="in" value="1.1">1.1 in</quantity>
eye length, dorsal profile of snout gently sloping; (3) lower lip ends anteriorly slightly in front of upper lip; (4) scales in lateral series 28 or 29; (5) scales in transverse series 6; (6) head length as percentage of standard length: 28.7–30.0% adults, 32.2% juvenile; (7) greatest body depth, measured in the new species at the pelvic fin origin as percentage of standard length: 15.0–16.8%; (8) eye diameter (as percentage of standard length): 6.9– 7.7%; adult female coloration: (9) head and body to pectoral and pelvic fins orange reddish, the rest of body whitish transparent with a dark reticulate pattern and small amount of orange red pigment; (10) fins mostly transparent and only partially pigmented; (11) caudal peduncle at caudal fin base with a dark orange vertical band.
</paragraph>
</subSubSection>
<subSubSection id="C32BD488FFD2FA58FF64FBF2FAD8F5BD" lastPageId="4" lastPageNumber="305" pageId="2" pageNumber="303" type="description">
<paragraph id="8B8E8703FFD2FA5EFF64FBF2FAB0F256" blockId="2.[151,1437,151,1928]" pageId="2" pageNumber="303">
<emphasis id="B9455B11FFD2FA5EFF64FBF2FEF0F34B" bold="true" box="[199,337,1045,1070]" pageId="2" pageNumber="303">Description</emphasis>
. 
<emphasis id="B9455B11FFD2FA5EFEFCFBF2FDE6F348" box="[351,583,1045,1069]" italics="true" pageId="2" pageNumber="303">General morphology</emphasis>
. Body proportions are given in 
<tableCitation id="C6B3B2B8FFD2FA5EFC08FBF2FC5FF34B" box="[939,1022,1045,1070]" captionStart="TABLE 1" captionStartId="3.[151,239,152,175]" captionTargetBox="[159,1427,234,1991]" captionTargetPageId="3" captionText="TABLE 1. Morphometric characters (as proportional measurements in %) of Speleogobius llorisi sp. nov. Characters are sorted in alphabetic order." httpUri="http://table.plazi.org/id/DF4ED78BFFD3FA5FFF34FF7FFE0DF7AA" pageId="2" pageNumber="303" tableUuid="DF4ED78BFFD3FA5FFF34FF7FFE0DF7AA">Table 1</tableCitation>
. Body moderately elongate, laterally compressed with a slender caudal peduncle. Head long, 28.7–30.0% adults, 32.2% juvenile of SL, slightly depressed, with a nearly horizontal predorsal profile. Snout moderately long and pointed, equal or longer than eye, 1.0–1.1% in eye length, dorsal profile of snout gently sloping. Eyes dorsolateral, extending above dorsal profile. Anterior nostril short, tubular, erect, without process from rim; posterior nostril pore-like, near orbit. Mouth oblique, posterior angle of jaws below anterior edge of eye, anterior tip of mouth in the horizontal level of eye. Lips wide, lower lip ends anteriorly slightly in front of upper lip. Rows of pointed teeth in both jaws, outer row with largest teeth. Tongue truncated. Branchiostegal membrane attached to entire lateral margin of isthmus.
</paragraph>
<paragraph id="8B8E8703FFD2FA5EFF64FAD8FB37F1CD" blockId="2.[151,1437,151,1928]" pageId="2" pageNumber="303">
Fins. D 
<date id="FF8FA1C3FFD2FA5EFE83FAA7FEF3F232" box="[288,338,1344,1368]" pageId="2" pageNumber="303">1 VI</date>
(VI:6); D 
<date id="FF8FA1C3FFD2FA5EFE6CFAA7FE51F23D" box="[463,496,1343,1368]" pageId="2" pageNumber="303">2 I</date>
/6–7 (6: 1, 7: 5); A 
<date id="FF8FA1C3FFD2FA5EFD7EFAD8FCB9F23D" box="[733,792,1343,1368]" bridgedPair="/" pageId="2" pageNumber="303">I/6–7</date>
(6: 5, 7: 1); C 11–12 branched rays (11:1, 12: 2, C of 
<typeStatus id="548A39A1FFD2FA5EFF34FA81FEA3F21B" box="[151,258,1382,1406]" pageId="2" pageNumber="303" type="paratype">paratypes</typeStatus>
PMR VP3511, ZSM 43988 and ZSM 43989 too damaged to count), 14–15 segmented rays (14: 3; 15: 3); P 15 (15: 11, right P of 
<typeStatus id="548A39A1FFD2FA5EFE1CFA6DFD80F2C6" box="[447,545,1418,1443]" pageId="2" pageNumber="303" type="holotype">holotype</typeStatus>
PMR VP3510, too damaged to count), V 
<date id="FF8FA1C3FFD2FA5EFBBDFA6DFB9FF2C6" box="[1054,1086,1418,1443]" bridgedPair="/" pageId="2" pageNumber="303">I/5</date>
+ 
<date id="FF8FA1C3FFD2FA5EFBECFA6DFBD2F2C6" box="[1103,1139,1418,1443]" bridgedPair="/" pageId="2" pageNumber="303">5/I.</date>
Fin-bases and lengths in proportion to standard body length are given in Table I. D 
<date id="FF8FA1C3FFD2FA5EFC89FA57FC02F2A2" box="[810,931,1455,1480]" pageId="2" pageNumber="303">1 II and III</date>
the longest, longest spines of D1 backwards nearly reaching origin of D2 when folded down in male adult and juvenile, not reaching in females. Fin membrane of D 
<date id="FF8FA1C3FFD2FA5EFF6CFA1CFF5EF177" box="[207,255,1531,1555]" pageId="2" pageNumber="303">1 VI</date>
not connected with base of D 
<date id="FF8FA1C3FFD2FA5EFDFBFA1CFDDEF177" box="[600,639,1531,1555]" pageId="2" pageNumber="303">2 I.</date>
D2 commences over vertical of anus to vertical of urogenital papilla, with last ray over vertical of last A ray. A commences below second to third segmented ray of D2. C posterior edge truncate. P uppermost rays within membrane, P rays all branched except uppermost and lowermost rays, ending back before vertical of D2 spine. V emarginated (shortest branches of 
<date id="FF8FA1C3FFD2FA5EFC18F98CFC47F1E6" box="[955,998,1642,1667]" pageId="2" pageNumber="303">V 5</date>
compared to longest branches of 
<date id="FF8FA1C3FFD2FA5EFAC9F98CFA39F1E6" box="[1386,1432,1643,1667]" pageId="2" pageNumber="303">V 4</date>
: 75%–87%), reaching behind anus, to A spine in juvenile, no anterior transverse membrane.
</paragraph>
<paragraph id="8B8E8703FFD2FA5EFF64F952FE68F0ED" blockId="2.[151,1437,151,1928]" pageId="2" pageNumber="303">Squamation. Body covered with ctenoid scales. Predorsal area scaly, scales along dorsal midline 5–7 (5:1, 6:2, 7:1, PMR VP3511, and ZSM 43989 too damaged for count). Opercle and cheek naked. Breast scaly with cycloid scales, scales along ventral midline 3–6 (3:1, 5:3, 6:1, ZSM 43989 too damaged for count). Scales in lateral series 28–29 (left and right side: 28:5, 29:4, PMR VP3511, too damaged to count on one side, ZSM 43989 on both sides), in transverse series 6 (left and right side: 6:8, PMR VP3511, and ZSM 43988 too damaged to count on one side, ZSM 43989 on both sides).</paragraph>
<caption id="DF4ED78BFFD3FA5FFF34FF7FFE0DF7AA" ID-Table-UUID="DF4ED78BFFD3FA5FFF34FF7FFE0DF7AA" httpUri="http://table.plazi.org/id/DF4ED78BFFD3FA5FFF34FF7FFE0DF7AA" pageId="3" pageNumber="304" targetBox="[159,1427,234,1991]" targetIsTable="true" targetPageId="3">
<paragraph id="8B8E8703FFD3FA5FFF34FF7FFE0DF7AA" blockId="3.[151,1436,151,207]" pageId="3" pageNumber="304">
<emphasis id="B9455B11FFD3FA5FFF34FF7FFEA8F7CA" bold="true" box="[151,265,152,175]" pageId="3" pageNumber="304">TABLE 1.</emphasis>
Morphometric characters (as proportional measurements in %) of 
<taxonomicName id="4C31FC80FFD3FA5FFC6EFF70FB03F7CA" box="[973,1186,151,175]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="304" phylum="Chordata" rank="species" species="llorisi" status="sp. nov.">
<emphasis id="B9455B11FFD3FA5FFC6EFF70FB03F7CA" box="[973,1186,151,175]" italics="true" pageId="3" pageNumber="304">Speleogobius llorisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A276E66AFFD3FA5FFB0BFF7FFB5CF7CA" box="[1192,1277,152,175]" pageId="3" pageNumber="304" rank="species">
<emphasis id="B9455B11FFD3FA5FFB0BFF7FFB5CF7CA" bold="true" box="[1192,1277,152,175]" pageId="3" pageNumber="304">sp. nov.</emphasis>
</taxonomicNameLabel>
Characters are sorted in alphabetic order.
</paragraph>
</caption>
<paragraph id="8B8E8703FFD3FA5FFF3CFF0DFAE7F0A2" pageId="3" pageNumber="304">
<table id="F93175A3FFD305A3FF3CFF0DFA32F0A2" box="[159,1427,234,1991]" gridcols="6" gridrows="40" pageId="3" pageNumber="304">
<tr id="35018541FFD305A3FF3CFF0DFA32F639" box="[159,1427,234,348]" gridrow="0" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFF0DFDAAF639" box="[159,523,234,348]" gridcol="0" gridrow="0" pageId="3" pageNumber="304">Specimen</th>
<th id="76D0EC3DFFD305A3FD9EFF0DFD37F639" box="[573,662,234,348]" gridcol="1" gridrow="0" pageId="3" pageNumber="304">P3 PMR VP3510 holotype</th>
<th id="76D0EC3DFFD305A3FD6DFF0DFC6AF639" box="[718,971,234,348]" gridcol="2" gridrow="0" pageId="3" pageNumber="304">P47 P57 PMR ZSM 43988 VP3513 paratype paratype</th>
<th id="76D0EC3DFFD305A3FC55FF0DFBF0F639" box="[1014,1105,234,348]" gridcol="3" gridrow="0" pageId="3" pageNumber="304">P45 PMR VP3511 paratype</th>
<th id="76D0EC3DFFD305A3FB29FF0DFB4CF639" box="[1162,1261,234,348]" gridcol="4" gridrow="0" pageId="3" pageNumber="304">P54 PMR VP3512 paratype</th>
<th id="76D0EC3DFFD305A3FABAFF0DFA32F639" box="[1305,1427,234,348]" gridcol="5" gridrow="0" pageId="3" pageNumber="304">P2 ZSM 43989 paratype</th>
</tr>
<tr id="35018541FFD305A3FF3CFE97FA32F6C1" box="[159,1427,368,420]" gridrow="1" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFE97FDAAF6C1" box="[159,523,368,420]" gridcol="0" gridrow="1" pageId="3" pageNumber="304">sex</th>
<td id="76D0EC3DFFD305A3FD9EFE97FD37F6C1" box="[573,662,368,420]" gridcol="1" gridrow="1" pageId="3" pageNumber="304">female</td>
<td id="76D0EC3DFFD305A3FD6DFE97FC6AF6C1" box="[718,971,368,420]" gridcol="2" gridrow="1" pageId="3" pageNumber="304">female female</td>
<td id="76D0EC3DFFD305A3FC55FE97FBF0F6C1" box="[1014,1105,368,420]" gridcol="3" gridrow="1" pageId="3" pageNumber="304">female</td>
<td id="76D0EC3DFFD305A3FB29FE97FB4CF6C1" box="[1162,1261,368,420]" gridcol="4" gridrow="1" pageId="3" pageNumber="304">male</td>
<td id="76D0EC3DFFD305A3FABAFE97FA32F6C1" box="[1305,1427,368,420]" gridcol="5" gridrow="1" pageId="3" pageNumber="304">male juvenile</td>
</tr>
<tr id="35018541FFD305A3FF3CFE5FFA32F6AB" box="[159,1427,440,462]" gridrow="2" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFE5FFDAAF6AB" box="[159,523,440,462]" gridcol="0" gridrow="2" pageId="3" pageNumber="304">Standard length (Sl) in mm</th>
<td id="76D0EC3DFFD305A3FD9EFE5FFD37F6AB" box="[573,662,440,462]" gridcol="1" gridrow="2" pageId="3" pageNumber="304">25.6</td>
<td id="76D0EC3DFFD305A3FD6DFE5FFC6AF6AB" box="[718,971,440,462]" gridcol="2" gridrow="2" pageId="3" pageNumber="304">27.2 24.5</td>
<td id="76D0EC3DFFD305A3FC55FE5FFBF0F6AB" box="[1014,1105,440,462]" gridcol="3" gridrow="2" pageId="3" pageNumber="304">18.7</td>
<td id="76D0EC3DFFD305A3FB29FE5FFB4CF6AB" box="[1162,1261,440,462]" gridcol="4" gridrow="2" pageId="3" pageNumber="304">24.4</td>
<td id="76D0EC3DFFD305A3FABAFE5FFA32F6AB" box="[1305,1427,440,462]" gridcol="5" gridrow="2" pageId="3" pageNumber="304">14.3</td>
</tr>
<tr id="35018541FFD305A3FF3CFE06FA32F692" box="[159,1427,481,503]" gridrow="3" pageId="3" pageNumber="304" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76D0EC3DFFD305A3FF3CFE06FDAAF692" box="[159,523,481,503]" gridcol="0" gridrow="3" pageId="3" pageNumber="304">% of standard length:</th>
</tr>
<tr id="35018541FFD305A3FF3CFDEDFA32F545" box="[159,1427,522,544]" gridrow="4" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFDEDFDAAF545" box="[159,523,522,544]" gridcol="0" gridrow="4" pageId="3" pageNumber="304">1st spine length of first dorsal fin</th>
<td id="76D0EC3DFFD305A3FD9EFDEDFD37F545" box="[573,662,522,544]" gridcol="1" gridrow="4" pageId="3" pageNumber="304">13.3</td>
<td id="76D0EC3DFFD305A3FD6DFDEDFC6AF545" box="[718,971,522,544]" gridcol="2" gridrow="4" pageId="3" pageNumber="304">12.1 13.9</td>
<td id="76D0EC3DFFD305A3FC55FDEDFBF0F545" box="[1014,1105,522,544]" gridcol="3" gridrow="4" pageId="3" pageNumber="304">11.8</td>
<td id="76D0EC3DFFD305A3FB29FDEDFB4CF545" box="[1162,1261,522,544]" gridcol="4" gridrow="4" pageId="3" pageNumber="304">12.7</td>
<td id="76D0EC3DFFD305A3FABAFDEDFA32F545" box="[1305,1427,522,544]" gridcol="5" gridrow="4" pageId="3" pageNumber="304">13.3</td>
</tr>
<tr id="35018541FFD305A3FF3CFDD3FA32F52F" box="[159,1427,564,586]" gridrow="5" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFDD3FDAAF52F" box="[159,523,564,586]" gridcol="0" gridrow="5" pageId="3" pageNumber="304">2nd spine length of first dorsal fin</th>
<td id="76D0EC3DFFD305A3FD9EFDD3FD37F52F" box="[573,662,564,586]" gridcol="1" gridrow="5" pageId="3" pageNumber="304">14.8</td>
<td id="76D0EC3DFFD305A3FD6DFDD3FC6AF52F" box="[718,971,564,586]" gridcol="2" gridrow="5" pageId="3" pageNumber="304">14.3 damaged</td>
<td id="76D0EC3DFFD305A3FC55FDD3FBF0F52F" box="[1014,1105,564,586]" gridcol="3" gridrow="5" pageId="3" pageNumber="304">damaged</td>
<td id="76D0EC3DFFD305A3FB29FDD3FB4CF52F" box="[1162,1261,564,586]" gridcol="4" gridrow="5" pageId="3" pageNumber="304">14.3</td>
<td id="76D0EC3DFFD305A3FABAFDD3FA32F52F" box="[1305,1427,564,586]" gridcol="5" gridrow="5" pageId="3" pageNumber="304">18.2</td>
</tr>
<tr id="35018541FFD305A3FF3CFDBAFA32F516" box="[159,1427,605,627]" gridrow="6" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFDBAFDAAF516" box="[159,523,605,627]" gridcol="0" gridrow="6" pageId="3" pageNumber="304">3rd spine length of first dorsal fin</th>
<td id="76D0EC3DFFD305A3FD9EFDBAFD37F516" box="[573,662,605,627]" gridcol="1" gridrow="6" pageId="3" pageNumber="304">13.7</td>
<td id="76D0EC3DFFD305A3FD6DFDBAFC6AF516" box="[718,971,605,627]" gridcol="2" gridrow="6" pageId="3" pageNumber="304">13.6 15.9</td>
<td id="76D0EC3DFFD305A3FC55FDBAFBF0F516" box="[1014,1105,605,627]" gridcol="3" gridrow="6" pageId="3" pageNumber="304">16.0</td>
<td id="76D0EC3DFFD305A3FB29FDBAFB4CF516" box="[1162,1261,605,627]" gridcol="4" gridrow="6" pageId="3" pageNumber="304">13.5</td>
<td id="76D0EC3DFFD305A3FABAFDBAFA32F516" box="[1305,1427,605,627]" gridcol="5" gridrow="6" pageId="3" pageNumber="304">16.8</td>
</tr>
<tr id="35018541FFD305A3FF3CFD61FA32F5F9" box="[159,1427,646,668]" gridrow="7" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFD61FDAAF5F9" box="[159,523,646,668]" gridcol="0" gridrow="7" pageId="3" pageNumber="304">Anal fin base</th>
<td id="76D0EC3DFFD305A3FD9EFD61FD37F5F9" box="[573,662,646,668]" gridcol="1" gridrow="7" pageId="3" pageNumber="304">11.7</td>
<td id="76D0EC3DFFD305A3FD6DFD61FC6AF5F9" box="[718,971,646,668]" gridcol="2" gridrow="7" pageId="3" pageNumber="304">9.6 11.8</td>
<td id="76D0EC3DFFD305A3FC55FD61FBF0F5F9" box="[1014,1105,646,668]" gridcol="3" gridrow="7" pageId="3" pageNumber="304">10.7</td>
<td id="76D0EC3DFFD305A3FB29FD61FB4CF5F9" box="[1162,1261,646,668]" gridcol="4" gridrow="7" pageId="3" pageNumber="304">11.9</td>
<td id="76D0EC3DFFD305A3FABAFD61FA32F5F9" box="[1305,1427,646,668]" gridcol="5" gridrow="7" pageId="3" pageNumber="304">9.1</td>
</tr>
<tr id="35018541FFD305A3FF3CFD57FA32F5A3" box="[159,1427,688,710]" gridrow="8" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFD57FDAAF5A3" box="[159,523,688,710]" gridcol="0" gridrow="8" pageId="3" pageNumber="304">Anal fin spine length</th>
<td id="76D0EC3DFFD305A3FD9EFD57FD37F5A3" box="[573,662,688,710]" gridcol="1" gridrow="8" pageId="3" pageNumber="304">9.4</td>
<td id="76D0EC3DFFD305A3FD6DFD57FC6AF5A3" box="[718,971,688,710]" gridcol="2" gridrow="8" pageId="3" pageNumber="304">9.6 11.0</td>
<td id="76D0EC3DFFD305A3FC55FD57FBF0F5A3" box="[1014,1105,688,710]" gridcol="3" gridrow="8" pageId="3" pageNumber="304">9.1</td>
<td id="76D0EC3DFFD305A3FB29FD57FB4CF5A3" box="[1162,1261,688,710]" gridcol="4" gridrow="8" pageId="3" pageNumber="304">9.4</td>
<td id="76D0EC3DFFD305A3FABAFD57FA32F5A3" box="[1305,1427,688,710]" gridcol="5" gridrow="8" pageId="3" pageNumber="304">10.5</td>
</tr>
<tr id="35018541FFD305A3FF3CFD3EFA32F58A" box="[159,1427,729,751]" gridrow="9" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFD3EFDAAF58A" box="[159,523,729,751]" gridcol="0" gridrow="9" pageId="3" pageNumber="304">Body depth at anal fin origin</th>
<td id="76D0EC3DFFD305A3FD9EFD3EFD37F58A" box="[573,662,729,751]" gridcol="1" gridrow="9" pageId="3" pageNumber="304">14.1</td>
<td id="76D0EC3DFFD305A3FD6DFD3EFC6AF58A" box="[718,971,729,751]" gridcol="2" gridrow="9" pageId="3" pageNumber="304">15.1 14.3</td>
<td id="76D0EC3DFFD305A3FC55FD3EFBF0F58A" box="[1014,1105,729,751]" gridcol="3" gridrow="9" pageId="3" pageNumber="304">13.9</td>
<td id="76D0EC3DFFD305A3FB29FD3EFB4CF58A" box="[1162,1261,729,751]" gridcol="4" gridrow="9" pageId="3" pageNumber="304">13.5</td>
<td id="76D0EC3DFFD305A3FABAFD3EFA32F58A" box="[1305,1427,729,751]" gridcol="5" gridrow="9" pageId="3" pageNumber="304">14.0</td>
</tr>
<tr id="35018541FFD305A3FF3CFCE5FA32F47D" box="[159,1427,770,792]" gridrow="10" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFCE5FDAAF47D" box="[159,523,770,792]" gridcol="0" gridrow="10" pageId="3" pageNumber="304">Body depth at pelvic fin origin</th>
<td id="76D0EC3DFFD305A3FD9EFCE5FD37F47D" box="[573,662,770,792]" gridcol="1" gridrow="10" pageId="3" pageNumber="304">16.8</td>
<td id="76D0EC3DFFD305A3FD6DFCE5FC6AF47D" box="[718,971,770,792]" gridcol="2" gridrow="10" pageId="3" pageNumber="304">16.2 15.5</td>
<td id="76D0EC3DFFD305A3FC55FCE5FBF0F47D" box="[1014,1105,770,792]" gridcol="3" gridrow="10" pageId="3" pageNumber="304">15.0</td>
<td id="76D0EC3DFFD305A3FB29FCE5FB4CF47D" box="[1162,1261,770,792]" gridcol="4" gridrow="10" pageId="3" pageNumber="304">16.0</td>
<td id="76D0EC3DFFD305A3FABAFCE5FA32F47D" box="[1305,1427,770,792]" gridcol="5" gridrow="10" pageId="3" pageNumber="304">16.8</td>
</tr>
<tr id="35018541FFD305A3FF3CFCCBFA32F427" box="[159,1427,812,834]" gridrow="11" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFCCBFDAAF427" box="[159,523,812,834]" gridcol="0" gridrow="11" pageId="3" pageNumber="304">Body width at anal fin origin</th>
<td id="76D0EC3DFFD305A3FD9EFCCBFD37F427" box="[573,662,812,834]" gridcol="1" gridrow="11" pageId="3" pageNumber="304">9.8</td>
<td id="76D0EC3DFFD305A3FD6DFCCBFC6AF427" box="[718,971,812,834]" gridcol="2" gridrow="11" pageId="3" pageNumber="304">9.2 10.2</td>
<td id="76D0EC3DFFD305A3FC55FCCBFBF0F427" box="[1014,1105,812,834]" gridcol="3" gridrow="11" pageId="3" pageNumber="304">10.2</td>
<td id="76D0EC3DFFD305A3FB29FCCBFB4CF427" box="[1162,1261,812,834]" gridcol="4" gridrow="11" pageId="3" pageNumber="304">10.2</td>
<td id="76D0EC3DFFD305A3FABAFCCBFA32F427" box="[1305,1427,812,834]" gridcol="5" gridrow="11" pageId="3" pageNumber="304">9.1</td>
</tr>
<tr id="35018541FFD305A3FF3CFCB2FA32F40E" box="[159,1427,853,875]" gridrow="12" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFCB2FDAAF40E" box="[159,523,853,875]" gridcol="0" gridrow="12" pageId="3" pageNumber="304">Body width at pelvic fin origin</th>
<td id="76D0EC3DFFD305A3FD9EFCB2FD37F40E" box="[573,662,853,875]" gridcol="1" gridrow="12" pageId="3" pageNumber="304">15.2</td>
<td id="76D0EC3DFFD305A3FD6DFCB2FC6AF40E" box="[718,971,853,875]" gridcol="2" gridrow="12" pageId="3" pageNumber="304">14.7 15.5</td>
<td id="76D0EC3DFFD305A3FC55FCB2FBF0F40E" box="[1014,1105,853,875]" gridcol="3" gridrow="12" pageId="3" pageNumber="304">13.9</td>
<td id="76D0EC3DFFD305A3FB29FCB2FB4CF40E" box="[1162,1261,853,875]" gridcol="4" gridrow="12" pageId="3" pageNumber="304">13.9</td>
<td id="76D0EC3DFFD305A3FABAFCB2FA32F40E" box="[1305,1427,853,875]" gridcol="5" gridrow="12" pageId="3" pageNumber="304">16.1</td>
</tr>
<tr id="35018541FFD305A3FF3CFC99FA32F4F1" box="[159,1427,894,916]" gridrow="13" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFC99FDAAF4F1" box="[159,523,894,916]" gridcol="0" gridrow="13" pageId="3" pageNumber="304">Caudal fin length</th>
<td id="76D0EC3DFFD305A3FD9EFC99FD37F4F1" box="[573,662,894,916]" gridcol="1" gridrow="13" pageId="3" pageNumber="304">21.9</td>
<td id="76D0EC3DFFD305A3FD6DFC99FC6AF4F1" box="[718,971,894,916]" gridcol="2" gridrow="13" pageId="3" pageNumber="304">damaged 22.9</td>
<td id="76D0EC3DFFD305A3FC55FC99FBF0F4F1" box="[1014,1105,894,916]" gridcol="3" gridrow="13" pageId="3" pageNumber="304">damaged</td>
<td id="76D0EC3DFFD305A3FB29FC99FB4CF4F1" box="[1162,1261,894,916]" gridcol="4" gridrow="13" pageId="3" pageNumber="304">22.5</td>
<td id="76D0EC3DFFD305A3FABAFC99FA32F4F1" box="[1305,1427,894,916]" gridcol="5" gridrow="13" pageId="3" pageNumber="304">28.0</td>
</tr>
<tr id="35018541FFD305A3FF3CFC4FFA32F4DB" box="[159,1427,936,958]" gridrow="14" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFC4FFDAAF4DB" box="[159,523,936,958]" gridcol="0" gridrow="14" pageId="3" pageNumber="304">Caudal peduncle depth</th>
<td id="76D0EC3DFFD305A3FD9EFC4FFD37F4DB" box="[573,662,936,958]" gridcol="1" gridrow="14" pageId="3" pageNumber="304">8.6</td>
<td id="76D0EC3DFFD305A3FD6DFC4FFC6AF4DB" box="[718,971,936,958]" gridcol="2" gridrow="14" pageId="3" pageNumber="304">8.1 8.2</td>
<td id="76D0EC3DFFD305A3FC55FC4FFBF0F4DB" box="[1014,1105,936,958]" gridcol="3" gridrow="14" pageId="3" pageNumber="304">7.5</td>
<td id="76D0EC3DFFD305A3FB29FC4FFB4CF4DB" box="[1162,1261,936,958]" gridcol="4" gridrow="14" pageId="3" pageNumber="304">7.8</td>
<td id="76D0EC3DFFD305A3FABAFC4FFA32F4DB" box="[1305,1427,936,958]" gridcol="5" gridrow="14" pageId="3" pageNumber="304">8.4</td>
</tr>
<tr id="35018541FFD305A3FF3CFC36FA32F482" box="[159,1427,977,999]" gridrow="15" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFC36FDAAF482" box="[159,523,977,999]" gridcol="0" gridrow="15" pageId="3" pageNumber="304">Caudal peduncle length</th>
<td id="76D0EC3DFFD305A3FD9EFC36FD37F482" box="[573,662,977,999]" gridcol="1" gridrow="15" pageId="3" pageNumber="304">26.6</td>
<td id="76D0EC3DFFD305A3FD6DFC36FC6AF482" box="[718,971,977,999]" gridcol="2" gridrow="15" pageId="3" pageNumber="304">25.4 26.9</td>
<td id="76D0EC3DFFD305A3FC55FC36FBF0F482" box="[1014,1105,977,999]" gridcol="3" gridrow="15" pageId="3" pageNumber="304">28.3</td>
<td id="76D0EC3DFFD305A3FB29FC36FB4CF482" box="[1162,1261,977,999]" gridcol="4" gridrow="15" pageId="3" pageNumber="304">28.3</td>
<td id="76D0EC3DFFD305A3FABAFC36FA32F482" box="[1305,1427,977,999]" gridcol="5" gridrow="15" pageId="3" pageNumber="304">28.0</td>
</tr>
<tr id="35018541FFD305A3FF3CFC1DFA32F375" box="[159,1427,1018,1040]" gridrow="16" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFC1DFDAAF375" box="[159,523,1018,1040]" gridcol="0" gridrow="16" pageId="3" pageNumber="304">First dorsal fin base</th>
<td id="76D0EC3DFFD305A3FD9EFC1DFD37F375" box="[573,662,1018,1040]" gridcol="1" gridrow="16" pageId="3" pageNumber="304">10.5</td>
<td id="76D0EC3DFFD305A3FD6DFC1DFC6AF375" box="[718,971,1018,1040]" gridcol="2" gridrow="16" pageId="3" pageNumber="304">9.9 9.8</td>
<td id="76D0EC3DFFD305A3FC55FC1DFBF0F375" box="[1014,1105,1018,1040]" gridcol="3" gridrow="16" pageId="3" pageNumber="304">9.1</td>
<td id="76D0EC3DFFD305A3FB29FC1DFB4CF375" box="[1162,1261,1018,1040]" gridcol="4" gridrow="16" pageId="3" pageNumber="304">9.0</td>
<td id="76D0EC3DFFD305A3FABAFC1DFA32F375" box="[1305,1427,1018,1040]" gridcol="5" gridrow="16" pageId="3" pageNumber="304">9.1</td>
</tr>
<tr id="35018541FFD305A3FF3CFBC3FA32F35F" box="[159,1427,1060,1082]" gridrow="17" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFBC3FDAAF35F" box="[159,523,1060,1082]" gridcol="0" gridrow="17" pageId="3" pageNumber="304">Head length</th>
<td id="76D0EC3DFFD305A3FD9EFBC3FD37F35F" box="[573,662,1060,1082]" gridcol="1" gridrow="17" pageId="3" pageNumber="304">29.3</td>
<td id="76D0EC3DFFD305A3FD6DFBC3FC6AF35F" box="[718,971,1060,1082]" gridcol="2" gridrow="17" pageId="3" pageNumber="304">28.7 29.4</td>
<td id="76D0EC3DFFD305A3FC55FBC3FBF0F35F" box="[1014,1105,1060,1082]" gridcol="3" gridrow="17" pageId="3" pageNumber="304">29.9</td>
<td id="76D0EC3DFFD305A3FB29FBC3FB4CF35F" box="[1162,1261,1060,1082]" gridcol="4" gridrow="17" pageId="3" pageNumber="304">28.7</td>
<td id="76D0EC3DFFD305A3FABAFBC3FA32F35F" box="[1305,1427,1060,1082]" gridcol="5" gridrow="17" pageId="3" pageNumber="304">32.2</td>
</tr>
<tr id="35018541FFD305A3FF3CFBAAFA32F306" box="[159,1427,1101,1123]" gridrow="18" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFBAAFDAAF306" box="[159,523,1101,1123]" gridcol="0" gridrow="18" pageId="3" pageNumber="304">Head width</th>
<td id="76D0EC3DFFD305A3FD9EFBAAFD37F306" box="[573,662,1101,1123]" gridcol="1" gridrow="18" pageId="3" pageNumber="304">17.2</td>
<td id="76D0EC3DFFD305A3FD6DFBAAFC6AF306" box="[718,971,1101,1123]" gridcol="2" gridrow="18" pageId="3" pageNumber="304">15.4 15.9</td>
<td id="76D0EC3DFFD305A3FC55FBAAFBF0F306" box="[1014,1105,1101,1123]" gridcol="3" gridrow="18" pageId="3" pageNumber="304">14.4</td>
<td id="76D0EC3DFFD305A3FB29FBAAFB4CF306" box="[1162,1261,1101,1123]" gridcol="4" gridrow="18" pageId="3" pageNumber="304">15.6</td>
<td id="76D0EC3DFFD305A3FABAFBAAFA32F306" box="[1305,1427,1101,1123]" gridcol="5" gridrow="18" pageId="3" pageNumber="304">16.1</td>
</tr>
<tr id="35018541FFD305A3FF3CFB91FA32F3E9" box="[159,1427,1142,1164]" gridrow="19" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFB91FDAAF3E9" box="[159,523,1142,1164]" gridcol="0" gridrow="19" pageId="3" pageNumber="304">Pectoral fin length</th>
<td id="76D0EC3DFFD305A3FD9EFB91FD37F3E9" box="[573,662,1142,1164]" gridcol="1" gridrow="19" pageId="3" pageNumber="304">23.4</td>
<td id="76D0EC3DFFD305A3FD6DFB91FC6AF3E9" box="[718,971,1142,1164]" gridcol="2" gridrow="19" pageId="3" pageNumber="304">21.3 22.4</td>
<td id="76D0EC3DFFD305A3FC55FB91FBF0F3E9" box="[1014,1105,1142,1164]" gridcol="3" gridrow="19" pageId="3" pageNumber="304">23.5</td>
<td id="76D0EC3DFFD305A3FB29FB91FB4CF3E9" box="[1162,1261,1142,1164]" gridcol="4" gridrow="19" pageId="3" pageNumber="304">21.3</td>
<td id="76D0EC3DFFD305A3FABAFB91FA32F3E9" box="[1305,1427,1142,1164]" gridcol="5" gridrow="19" pageId="3" pageNumber="304">21.7</td>
</tr>
<tr id="35018541FFD305A3FF3CFB47FA32F3D3" box="[159,1427,1184,1206]" gridrow="20" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFB47FDAAF3D3" box="[159,523,1184,1206]" gridcol="0" gridrow="20" pageId="3" pageNumber="304">Pelvic fin length</th>
<td id="76D0EC3DFFD305A3FD9EFB47FD37F3D3" box="[573,662,1184,1206]" gridcol="1" gridrow="20" pageId="3" pageNumber="304">27.0</td>
<td id="76D0EC3DFFD305A3FD6DFB47FC6AF3D3" box="[718,971,1184,1206]" gridcol="2" gridrow="20" pageId="3" pageNumber="304">25.4 26.9</td>
<td id="76D0EC3DFFD305A3FC55FB47FBF0F3D3" box="[1014,1105,1184,1206]" gridcol="3" gridrow="20" pageId="3" pageNumber="304">28.3</td>
<td id="76D0EC3DFFD305A3FB29FB47FB4CF3D3" box="[1162,1261,1184,1206]" gridcol="4" gridrow="20" pageId="3" pageNumber="304">27.0</td>
<td id="76D0EC3DFFD305A3FABAFB47FA32F3D3" box="[1305,1427,1184,1206]" gridcol="5" gridrow="20" pageId="3" pageNumber="304">32.9</td>
</tr>
<tr id="35018541FFD305A3FF3CFB2EFA32F3BA" box="[159,1427,1225,1247]" gridrow="21" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFB2EFDAAF3BA" box="[159,523,1225,1247]" gridcol="0" gridrow="21" pageId="3" pageNumber="304">Pelvic to anus</th>
<td id="76D0EC3DFFD305A3FD9EFB2EFD37F3BA" box="[573,662,1225,1247]" gridcol="1" gridrow="21" pageId="3" pageNumber="304">23.0</td>
<td id="76D0EC3DFFD305A3FD6DFB2EFC6AF3BA" box="[718,971,1225,1247]" gridcol="2" gridrow="21" pageId="3" pageNumber="304">23.9 22.0</td>
<td id="76D0EC3DFFD305A3FC55FB2EFBF0F3BA" box="[1014,1105,1225,1247]" gridcol="3" gridrow="21" pageId="3" pageNumber="304">25.7</td>
<td id="76D0EC3DFFD305A3FB29FB2EFB4CF3BA" box="[1162,1261,1225,1247]" gridcol="4" gridrow="21" pageId="3" pageNumber="304">21.7</td>
<td id="76D0EC3DFFD305A3FABAFB2EFA32F3BA" box="[1305,1427,1225,1247]" gridcol="5" gridrow="21" pageId="3" pageNumber="304">24.5</td>
</tr>
<tr id="35018541FFD305A3FF3CFB15FA32F26D" box="[159,1427,1266,1288]" gridrow="22" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFB15FDAAF26D" box="[159,523,1266,1288]" gridcol="0" gridrow="22" pageId="3" pageNumber="304">Second dorsal fin base</th>
<td id="76D0EC3DFFD305A3FD9EFB15FD37F26D" box="[573,662,1266,1288]" gridcol="1" gridrow="22" pageId="3" pageNumber="304">14.5</td>
<td id="76D0EC3DFFD305A3FD6DFB15FC6AF26D" box="[718,971,1266,1288]" gridcol="2" gridrow="22" pageId="3" pageNumber="304">12.9 13.9</td>
<td id="76D0EC3DFFD305A3FC55FB15FBF0F26D" box="[1014,1105,1266,1288]" gridcol="3" gridrow="22" pageId="3" pageNumber="304">15.5</td>
<td id="76D0EC3DFFD305A3FB29FB15FB4CF26D" box="[1162,1261,1266,1288]" gridcol="4" gridrow="22" pageId="3" pageNumber="304">14.8</td>
<td id="76D0EC3DFFD305A3FABAFB15FA32F26D" box="[1305,1427,1266,1288]" gridcol="5" gridrow="22" pageId="3" pageNumber="304">13.3</td>
</tr>
<tr id="35018541FFD305A3FF3CFAFBFA32F257" box="[159,1427,1308,1330]" gridrow="23" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFAFBFDAAF257" box="[159,523,1308,1330]" gridcol="0" gridrow="23" pageId="3" pageNumber="304">Second dorsal fin spine length</th>
<td id="76D0EC3DFFD305A3FD9EFAFBFD37F257" box="[573,662,1308,1330]" gridcol="1" gridrow="23" pageId="3" pageNumber="304">13.3</td>
<td id="76D0EC3DFFD305A3FD6DFAFBFC6AF257" box="[718,971,1308,1330]" gridcol="2" gridrow="23" pageId="3" pageNumber="304">11.8 13.9</td>
<td id="76D0EC3DFFD305A3FC55FAFBFBF0F257" box="[1014,1105,1308,1330]" gridcol="3" gridrow="23" pageId="3" pageNumber="304">11.8</td>
<td id="76D0EC3DFFD305A3FB29FAFBFB4CF257" box="[1162,1261,1308,1330]" gridcol="4" gridrow="23" pageId="3" pageNumber="304">13.1</td>
<td id="76D0EC3DFFD305A3FABAFAFBFA32F257" box="[1305,1427,1308,1330]" gridcol="5" gridrow="23" pageId="3" pageNumber="304">14.0</td>
</tr>
<tr id="35018541FFD305A3FF3CFAA2FA32F23E" box="[159,1427,1349,1371]" gridrow="24" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFAA2FDAAF23E" box="[159,523,1349,1371]" gridcol="0" gridrow="24" pageId="3" pageNumber="304">Snout to origin of first dorsal fin</th>
<td id="76D0EC3DFFD305A3FD9EFAA2FD37F23E" box="[573,662,1349,1371]" gridcol="1" gridrow="24" pageId="3" pageNumber="304">37.5</td>
<td id="76D0EC3DFFD305A3FD6DFAA2FC6AF23E" box="[718,971,1349,1371]" gridcol="2" gridrow="24" pageId="3" pageNumber="304">36.8 38.4</td>
<td id="76D0EC3DFFD305A3FC55FAA2FBF0F23E" box="[1014,1105,1349,1371]" gridcol="3" gridrow="24" pageId="3" pageNumber="304">38.0</td>
<td id="76D0EC3DFFD305A3FB29FAA2FB4CF23E" box="[1162,1261,1349,1371]" gridcol="4" gridrow="24" pageId="3" pageNumber="304">38.1</td>
<td id="76D0EC3DFFD305A3FABAFAA2FA32F23E" box="[1305,1427,1349,1371]" gridcol="5" gridrow="24" pageId="3" pageNumber="304">38.5</td>
</tr>
<tr id="35018541FFD305A3FF3CFA89FA32F2E1" box="[159,1427,1390,1412]" gridrow="25" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFA89FDAAF2E1" box="[159,523,1390,1412]" gridcol="0" gridrow="25" pageId="3" pageNumber="304">Snout to origin of second dorsal fin</th>
<td id="76D0EC3DFFD305A3FD9EFA89FD37F2E1" box="[573,662,1390,1412]" gridcol="1" gridrow="25" pageId="3" pageNumber="304">56.6</td>
<td id="76D0EC3DFFD305A3FD6DFA89FC6AF2E1" box="[718,971,1390,1412]" gridcol="2" gridrow="25" pageId="3" pageNumber="304">55.1 57.1</td>
<td id="76D0EC3DFFD305A3FC55FA89FBF0F2E1" box="[1014,1105,1390,1412]" gridcol="3" gridrow="25" pageId="3" pageNumber="304">55.6</td>
<td id="76D0EC3DFFD305A3FB29FA89FB4CF2E1" box="[1162,1261,1390,1412]" gridcol="4" gridrow="25" pageId="3" pageNumber="304">55.7</td>
<td id="76D0EC3DFFD305A3FABAFA89FA32F2E1" box="[1305,1427,1390,1412]" gridcol="5" gridrow="25" pageId="3" pageNumber="304">57.3</td>
</tr>
<tr id="35018541FFD305A3FF3CFA7FFA32F2CB" box="[159,1427,1432,1454]" gridrow="26" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFA7FFDAAF2CB" box="[159,523,1432,1454]" gridcol="0" gridrow="26" pageId="3" pageNumber="304">Snout to vertical of anal fin origin</th>
<td id="76D0EC3DFFD305A3FD9EFA7FFD37F2CB" box="[573,662,1432,1454]" gridcol="1" gridrow="26" pageId="3" pageNumber="304">60.5</td>
<td id="76D0EC3DFFD305A3FD6DFA7FFC6AF2CB" box="[718,971,1432,1454]" gridcol="2" gridrow="26" pageId="3" pageNumber="304">61.8 61.2</td>
<td id="76D0EC3DFFD305A3FC55FA7FFBF0F2CB" box="[1014,1105,1432,1454]" gridcol="3" gridrow="26" pageId="3" pageNumber="304">62.0</td>
<td id="76D0EC3DFFD305A3FB29FA7FFB4CF2CB" box="[1162,1261,1432,1454]" gridcol="4" gridrow="26" pageId="3" pageNumber="304">59.0</td>
<td id="76D0EC3DFFD305A3FABAFA7FFA32F2CB" box="[1305,1427,1432,1454]" gridcol="5" gridrow="26" pageId="3" pageNumber="304">62.9</td>
</tr>
<tr id="35018541FFD305A3FF3CFA26FA32F2B2" box="[159,1427,1473,1495]" gridrow="27" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFA26FDAAF2B2" box="[159,523,1473,1495]" gridcol="0" gridrow="27" pageId="3" pageNumber="304">Snout to vertical of anus</th>
<td id="76D0EC3DFFD305A3FD9EFA26FD37F2B2" box="[573,662,1473,1495]" gridcol="1" gridrow="27" pageId="3" pageNumber="304">53.5</td>
<td id="76D0EC3DFFD305A3FD6DFA26FC6AF2B2" box="[718,971,1473,1495]" gridcol="2" gridrow="27" pageId="3" pageNumber="304">54.0 53.1</td>
<td id="76D0EC3DFFD305A3FC55FA26FBF0F2B2" box="[1014,1105,1473,1495]" gridcol="3" gridrow="27" pageId="3" pageNumber="304">54.5</td>
<td id="76D0EC3DFFD305A3FB29FA26FB4CF2B2" box="[1162,1261,1473,1495]" gridcol="4" gridrow="27" pageId="3" pageNumber="304">52.0</td>
<td id="76D0EC3DFFD305A3FABAFA26FA32F2B2" box="[1305,1427,1473,1495]" gridcol="5" gridrow="27" pageId="3" pageNumber="304">56.6</td>
</tr>
<tr id="35018541FFD305A3FF3CFA0DFA32F165" box="[159,1427,1514,1536]" gridrow="28" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CFA0DFDAAF165" box="[159,523,1514,1536]" gridcol="0" gridrow="28" pageId="3" pageNumber="304">Snout to vertical of pelvic fin origin</th>
<td id="76D0EC3DFFD305A3FD9EFA0DFD37F165" box="[573,662,1514,1536]" gridcol="1" gridrow="28" pageId="3" pageNumber="304">31.3</td>
<td id="76D0EC3DFFD305A3FD6DFA0DFC6AF165" box="[718,971,1514,1536]" gridcol="2" gridrow="28" pageId="3" pageNumber="304">30.9 34.3</td>
<td id="76D0EC3DFFD305A3FC55FA0DFBF0F165" box="[1014,1105,1514,1536]" gridcol="3" gridrow="28" pageId="3" pageNumber="304">31.0</td>
<td id="76D0EC3DFFD305A3FB29FA0DFB4CF165" box="[1162,1261,1514,1536]" gridcol="4" gridrow="28" pageId="3" pageNumber="304">33.2</td>
<td id="76D0EC3DFFD305A3FABAFA0DFA32F165" box="[1305,1427,1514,1536]" gridcol="5" gridrow="28" pageId="3" pageNumber="304">34.3</td>
</tr>
<tr id="35018541FFD305A3FF3CF9F3FA32F14F" box="[159,1427,1556,1578]" gridrow="29" pageId="3" pageNumber="304" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76D0EC3DFFD305A3FF3CF9F3FDAAF14F" box="[159,523,1556,1578]" gridcol="0" gridrow="29" pageId="3" pageNumber="304">% of caudal peduncle:</th>
</tr>
<tr id="35018541FFD305A3FF3CF9DAFA32F136" box="[159,1427,1597,1619]" gridrow="30" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CF9DAFDAAF136" box="[159,523,1597,1619]" gridcol="0" gridrow="30" pageId="3" pageNumber="304">Caudal peduncle depth</th>
<td id="76D0EC3DFFD305A3FD9EF9DAFD37F136" box="[573,662,1597,1619]" gridcol="1" gridrow="30" pageId="3" pageNumber="304">32.4</td>
<td id="76D0EC3DFFD305A3FD6DF9DAFC6AF136" box="[718,971,1597,1619]" gridcol="2" gridrow="30" pageId="3" pageNumber="304">31.9 30.3</td>
<td id="76D0EC3DFFD305A3FC55F9DAFBF0F136" box="[1014,1105,1597,1619]" gridcol="3" gridrow="30" pageId="3" pageNumber="304">26.4</td>
<td id="76D0EC3DFFD305A3FB29F9DAFB4CF136" box="[1162,1261,1597,1619]" gridcol="4" gridrow="30" pageId="3" pageNumber="304">27.5</td>
<td id="76D0EC3DFFD305A3FABAF9DAFA32F136" box="[1305,1427,1597,1619]" gridcol="5" gridrow="30" pageId="3" pageNumber="304">30.0</td>
</tr>
<tr id="35018541FFD305A3FF3CF981FA32F119" box="[159,1427,1638,1660]" gridrow="31" pageId="3" pageNumber="304" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76D0EC3DFFD305A3FF3CF981FDAAF119" box="[159,523,1638,1660]" gridcol="0" gridrow="31" pageId="3" pageNumber="304">% of head length</th>
</tr>
<tr id="35018541FFD305A3FF3CF977FA32F1C3" box="[159,1427,1680,1702]" gridrow="32" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CF977FDAAF1C3" box="[159,523,1680,1702]" gridcol="0" gridrow="32" pageId="3" pageNumber="304">Cheek depth</th>
<td id="76D0EC3DFFD305A3FD9EF977FD37F1C3" box="[573,662,1680,1702]" gridcol="1" gridrow="32" pageId="3" pageNumber="304">13.3</td>
<td id="76D0EC3DFFD305A3FD6DF977FC6AF1C3" box="[718,971,1680,1702]" gridcol="2" gridrow="32" pageId="3" pageNumber="304">11.5 13.9</td>
<td id="76D0EC3DFFD305A3FC55F977FBF0F1C3" box="[1014,1105,1680,1702]" gridcol="3" gridrow="32" pageId="3" pageNumber="304">12.5</td>
<td id="76D0EC3DFFD305A3FB29F977FB4CF1C3" box="[1162,1261,1680,1702]" gridcol="4" gridrow="32" pageId="3" pageNumber="304">11.4</td>
<td id="76D0EC3DFFD305A3FABAF977FA32F1C3" box="[1305,1427,1680,1702]" gridcol="5" gridrow="32" pageId="3" pageNumber="304">13.0</td>
</tr>
<tr id="35018541FFD305A3FF3CF95EFA32F1AA" box="[159,1427,1721,1743]" gridrow="33" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CF95EFDAAF1AA" box="[159,523,1721,1743]" gridcol="0" gridrow="33" pageId="3" pageNumber="304">Eye diameter</th>
<td id="76D0EC3DFFD305A3FD9EF95EFD37F1AA" box="[573,662,1721,1743]" gridcol="1" gridrow="33" pageId="3" pageNumber="304">25.3</td>
<td id="76D0EC3DFFD305A3FD6DF95EFC6AF1AA" box="[718,971,1721,1743]" gridcol="2" gridrow="33" pageId="3" pageNumber="304">24.4 23.6</td>
<td id="76D0EC3DFFD305A3FC55F95EFBF0F1AA" box="[1014,1105,1721,1743]" gridcol="3" gridrow="33" pageId="3" pageNumber="304">25.0</td>
<td id="76D0EC3DFFD305A3FB29F95EFB4CF1AA" box="[1162,1261,1721,1743]" gridcol="4" gridrow="33" pageId="3" pageNumber="304">24.3</td>
<td id="76D0EC3DFFD305A3FABAF95EFA32F1AA" box="[1305,1427,1721,1743]" gridcol="5" gridrow="33" pageId="3" pageNumber="304">23.9</td>
</tr>
<tr id="35018541FFD305A3FF3CF905FA32F19D" box="[159,1427,1762,1784]" gridrow="34" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CF905FDAAF19D" box="[159,523,1762,1784]" gridcol="0" gridrow="34" pageId="3" pageNumber="304">Head depth</th>
<td id="76D0EC3DFFD305A3FD9EF905FD37F19D" box="[573,662,1762,1784]" gridcol="1" gridrow="34" pageId="3" pageNumber="304">42.7</td>
<td id="76D0EC3DFFD305A3FD6DF905FC6AF19D" box="[718,971,1762,1784]" gridcol="2" gridrow="34" pageId="3" pageNumber="304">42.3 43.1</td>
<td id="76D0EC3DFFD305A3FC55F905FBF0F19D" box="[1014,1105,1762,1784]" gridcol="3" gridrow="34" pageId="3" pageNumber="304">39.3</td>
<td id="76D0EC3DFFD305A3FB29F905FB4CF19D" box="[1162,1261,1762,1784]" gridcol="4" gridrow="34" pageId="3" pageNumber="304">42.9</td>
<td id="76D0EC3DFFD305A3FABAF905FA32F19D" box="[1305,1427,1762,1784]" gridcol="5" gridrow="34" pageId="3" pageNumber="304">43.5</td>
</tr>
<tr id="35018541FFD305A3FF3CF8EBFA32F047" box="[159,1427,1804,1826]" gridrow="35" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CF8EBFDAAF047" box="[159,523,1804,1826]" gridcol="0" gridrow="35" pageId="3" pageNumber="304">Head width</th>
<td id="76D0EC3DFFD305A3FD9EF8EBFD37F047" box="[573,662,1804,1826]" gridcol="1" gridrow="35" pageId="3" pageNumber="304">58.7</td>
<td id="76D0EC3DFFD305A3FD6DF8EBFC6AF047" box="[718,971,1804,1826]" gridcol="2" gridrow="35" pageId="3" pageNumber="304">53.8 54.2</td>
<td id="76D0EC3DFFD305A3FC55F8EBFBF0F047" box="[1014,1105,1804,1826]" gridcol="3" gridrow="35" pageId="3" pageNumber="304">48.2</td>
<td id="76D0EC3DFFD305A3FB29F8EBFB4CF047" box="[1162,1261,1804,1826]" gridcol="4" gridrow="35" pageId="3" pageNumber="304">54.3</td>
<td id="76D0EC3DFFD305A3FABAF8EBFA32F047" box="[1305,1427,1804,1826]" gridcol="5" gridrow="35" pageId="3" pageNumber="304">50.0</td>
</tr>
<tr id="35018541FFD305A3FF3CF8D2FA32F02E" box="[159,1427,1845,1867]" gridrow="36" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CF8D2FDAAF02E" box="[159,523,1845,1867]" gridcol="0" gridrow="36" pageId="3" pageNumber="304">Postorbital length</th>
<td id="76D0EC3DFFD305A3FD9EF8D2FD37F02E" box="[573,662,1845,1867]" gridcol="1" gridrow="36" pageId="3" pageNumber="304">52.0</td>
<td id="76D0EC3DFFD305A3FD6DF8D2FC6AF02E" box="[718,971,1845,1867]" gridcol="2" gridrow="36" pageId="3" pageNumber="304">48.7 51.4</td>
<td id="76D0EC3DFFD305A3FC55F8D2FBF0F02E" box="[1014,1105,1845,1867]" gridcol="3" gridrow="36" pageId="3" pageNumber="304">51.8</td>
<td id="76D0EC3DFFD305A3FB29F8D2FB4CF02E" box="[1162,1261,1845,1867]" gridcol="4" gridrow="36" pageId="3" pageNumber="304">50.0</td>
<td id="76D0EC3DFFD305A3FABAF8D2FA32F02E" box="[1305,1427,1845,1867]" gridcol="5" gridrow="36" pageId="3" pageNumber="304">52.2</td>
</tr>
<tr id="35018541FFD305A3FF3CF8B9FA32F011" box="[159,1427,1886,1908]" gridrow="37" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CF8B9FDAAF011" box="[159,523,1886,1908]" gridcol="0" gridrow="37" pageId="3" pageNumber="304">Snout length</th>
<td id="76D0EC3DFFD305A3FD9EF8B9FD37F011" box="[573,662,1886,1908]" gridcol="1" gridrow="37" pageId="3" pageNumber="304">25.3</td>
<td id="76D0EC3DFFD305A3FD6DF8B9FC6AF011" box="[718,971,1886,1908]" gridcol="2" gridrow="37" pageId="3" pageNumber="304">24.4 26.4</td>
<td id="76D0EC3DFFD305A3FC55F8B9FBF0F011" box="[1014,1105,1886,1908]" gridcol="3" gridrow="37" pageId="3" pageNumber="304">26.8</td>
<td id="76D0EC3DFFD305A3FB29F8B9FB4CF011" box="[1162,1261,1886,1908]" gridcol="4" gridrow="37" pageId="3" pageNumber="304">25.7</td>
<td id="76D0EC3DFFD305A3FABAF8B9FA32F011" box="[1305,1427,1886,1908]" gridcol="5" gridrow="37" pageId="3" pageNumber="304">26.1</td>
</tr>
<tr id="35018541FFD305A3FF3CF86FFA32F0FB" box="[159,1427,1928,1950]" gridrow="38" pageId="3" pageNumber="304" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76D0EC3DFFD305A3FF3CF86FFDAAF0FB" box="[159,523,1928,1950]" gridcol="0" gridrow="38" pageId="3" pageNumber="304">% of eye diameter</th>
</tr>
<tr id="35018541FFD305A3FF3CF856FA32F0A2" box="[159,1427,1969,1991]" gridrow="39" pageId="3" pageNumber="304">
<th id="76D0EC3DFFD305A3FF3CF856FDAAF0A2" box="[159,523,1969,1991]" gridcol="0" gridrow="39" pageId="3" pageNumber="304">Interorbital width</th>
<td id="76D0EC3DFFD305A3FD9EF856FD37F0A2" box="[573,662,1969,1991]" gridcol="1" gridrow="39" pageId="3" pageNumber="304">31.6</td>
<td id="76D0EC3DFFD305A3FD6DF856FC6AF0A2" box="[718,971,1969,1991]" gridcol="2" gridrow="39" pageId="3" pageNumber="304">36.8 35.3</td>
<td id="76D0EC3DFFD305A3FC55F856FBF0F0A2" box="[1014,1105,1969,1991]" gridcol="3" gridrow="39" pageId="3" pageNumber="304">35.7</td>
<td id="76D0EC3DFFD305A3FB29F856FB4CF0A2" box="[1162,1261,1969,1991]" gridcol="4" gridrow="39" pageId="3" pageNumber="304">29.4</td>
<td id="76D0EC3DFFD305A3FABAF856FA32F0A2" box="[1305,1427,1969,1991]" gridcol="5" gridrow="39" pageId="3" pageNumber="304">36.4</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8E8703FFD4FA58FF64FF7FFAD8F5BD" blockId="4.[151,1436,151,728]" pageId="4" pageNumber="305">
LL system (
<figureCitation id="130A9B86FFD4FA58FEEFFF70FE2AF7D5" box="[332,395,151,176]" captionStart="FIGURE 4" captionStartId="5.[151,250,1628,1650]" captionTargetBox="[179,1414,1022,1594]" captionTargetId="figure@5.[173,1414,1012,1605]" captionTargetPageId="5" captionText="FIGURE 4. Speleogobius llorisi sp. nov., head LL sensory papillae and canal pores: paratype, PMR VP 3512 (P 54), male, 24.4 + 5.5 mm, Spain, Balearic Islands, Mallorca. Terminology in text. Drawing by M. Kovačić." httpUri="https://zenodo.org/record/262961/files/figure.png" pageId="4" pageNumber="305">Fig 4</figureCitation>
). Head with anterior oculoscapular and preopercular canals, carrying pores 
<emphasis id="B9455B11FFD4FA58FB74FF7EFACDF7D5" box="[1239,1388,152,176]" italics="true" pageId="4" pageNumber="305">σ, λ, κ, α, β, ρ</emphasis>
and 
<emphasis id="B9455B11FFD4FA58FF34FF5AFF74F7B1" box="[151,213,188,212]" italics="true" pageId="4" pageNumber="305">γ, δ, ε</emphasis>
respectively (ZSM 43989 with surface too dried and damaged to see head canals and pores clearly). Posterior oculoscapular canal absent. Rows and number of sensory papillae as follows (all 
<typeStatus id="548A39A1FFD4FA58FBBCFF07FBEEF79D" box="[1055,1103,224,248]" pageId="4" pageNumber="305">type</typeStatus>
material checked, but usually only part of LL system visible on specimens due to the skin damage, entire LL system visible in 
<typeStatus id="548A39A1FFD4FA58FB5BFEE2FAF9F678" box="[1272,1368,261,285]" pageId="4" pageNumber="305" type="paratype">paratype</typeStatus>
PMR VP3512): (1) 
<emphasis id="B9455B11FFD4FA58FE97FECFFE05F625" box="[308,420,296,320]" italics="true" pageId="4" pageNumber="305">preorbital</emphasis>
: snout with single row 
<emphasis id="B9455B11FFD4FA58FD12FECEFD66F625" box="[689,711,297,320]" italics="true" pageId="4" pageNumber="305">rs</emphasis>
as 2 papillae close to pore 
<emphasis id="B9455B11FFD4FA58FC5EFECEFBAAF625" box="[1021,1035,297,320]" italics="true" pageId="4" pageNumber="305">σ</emphasis>
and posterior nostril. Lateral series 
<emphasis id="B9455B11FFD4FA58FF34FEAAFF17F60C" box="[151,182,333,361]" italics="true" pageId="4" pageNumber="305">c1,2</emphasis>
as two or three papillae in longitudinal row starting behind anterior nostril and going backwards above upper lip. (2) 
<emphasis id="B9455B11FFD4FA58FF49FE94FEFCF6EE" box="[234,349,371,395]" italics="true" pageId="4" pageNumber="305">suborbital</emphasis>
: rows without transverse proliferation; 
<emphasis id="B9455B11FFD4FA58FCB8FE94FC88F6EF" box="[795,809,371,394]" italics="true" pageId="4" pageNumber="305">a</emphasis>
below rear part of eye starting posteriorly at pore 
<emphasis id="B9455B11FFD4FA58FAFEFE94FACDF6EF" box="[1373,1388,371,394]" italics="true" pageId="4" pageNumber="305">α</emphasis>
(3); 
<emphasis id="B9455B11FFD4FA58FF34FE7FFF02F6CA" box="[151,163,408,431]" italics="true" pageId="4" pageNumber="305">c</emphasis>
below eye (4–5), anteriorly beginning below front border of eye, posteriorly ending before 
<emphasis id="B9455B11FFD4FA58FB61FE7FFB71F6CA" box="[1218,1232,408,431]" italics="true" pageId="4" pageNumber="305">a</emphasis>
row; papillae 
<emphasis id="B9455B11FFD4FA58FADBFE70FA3DF6CB" box="[1400,1436,407,430]" italics="true" pageId="4" pageNumber="305">cp’</emphasis>
and 
<emphasis id="B9455B11FFD4FA58FF64FE5CFF43F6B7" box="[199,226,443,466]" italics="true" pageId="4" pageNumber="305">cp</emphasis>
longitudinally arranged in front of 
<emphasis id="B9455B11FFD4FA58FDD1FE5CFD27F6B6" box="[626,646,443,467]" italics="true" pageId="4" pageNumber="305">b.</emphasis>
Row 
<emphasis id="B9455B11FFD4FA58FD68FE5CFD78F6B6" box="[715,729,443,467]" italics="true" pageId="4" pageNumber="305">b</emphasis>
single papilla behind rear border of eye and bellow level of it. Longitudinal row 
<emphasis id="B9455B11FFD4FA58FEC4FE38FED4F692" box="[359,373,479,503]" italics="true" pageId="4" pageNumber="305">d</emphasis>
as two papillae above upper lip posterior part. (3) 
<emphasis id="B9455B11FFD4FA58FC69FE38FB7CF692" box="[970,1245,479,503]" italics="true" pageId="4" pageNumber="305">preoperculo-mandibular</emphasis>
: anterior part of external row 
<emphasis id="B9455B11FFD4FA58FE91FDE4FE9FF57F" box="[306,318,515,538]" italics="true" pageId="4" pageNumber="305">e</emphasis>
not visible in all specimens, anterior part of internal row 
<emphasis id="B9455B11FFD4FA58FC4DFDE4FC57F57F" box="[1006,1014,515,538]" italics="true" pageId="4" pageNumber="305">i</emphasis>
(5) and row 
<emphasis id="B9455B11FFD4FA58FB35FDE4FB3FF57E" box="[1174,1182,515,539]" italics="true" pageId="4" pageNumber="305">f</emphasis>
(2) present. Posterior sections of row 
<emphasis id="B9455B11FFD4FA58FEEBFDCDFEF5F524" box="[328,340,554,577]" italics="true" pageId="4" pageNumber="305">e</emphasis>
(8–12) and row 
<emphasis id="B9455B11FFD4FA58FDAEFDCDFDB4F524" box="[525,533,554,577]" italics="true" pageId="4" pageNumber="305">i</emphasis>
(12–13). (4) 
<emphasis id="B9455B11FFD4FA58FD0AFDCEFCE8F524" box="[681,841,553,577]" italics="true" pageId="4" pageNumber="305">oculoscapular</emphasis>
: anterior longitudinal row 
<date id="FF8FA1C3FFD4FA58FBD6FDCDFB2BF551" box="[1141,1162,550,577]" pageId="4" pageNumber="305">
<emphasis id="B9455B11FFD4FA58FBD6FDCDFB2BF551" box="[1141,1162,550,577]" italics="true" pageId="4" pageNumber="305">x1</emphasis>
</date>
(2–3) above pore 
<emphasis id="B9455B11FFD4FA58FAF6FDCDFAC3F524" box="[1365,1378,554,577]" italics="true" pageId="4" pageNumber="305">ρ</emphasis>
, row 
<emphasis id="B9455B11FFD4FA58FF34FDB6FF03F50D" box="[151,162,593,616]" italics="true" pageId="4" pageNumber="305">z</emphasis>
(2) in the middle between pores 
<emphasis id="B9455B11FFD4FA58FDB1FDB6FD9DF50D" box="[530,572,593,617]" italics="true" pageId="4" pageNumber="305">β, ρ</emphasis>
and 
<emphasis id="B9455B11FFD4FA58FDD1FDB6FDDCF50D" box="[626,637,593,616]" italics="true" pageId="4" pageNumber="305">γ</emphasis>
, row 
<emphasis id="B9455B11FFD4FA58FD1EFDB6FD6AF50D" box="[701,715,593,616]" italics="true" pageId="4" pageNumber="305">q</emphasis>
(2–3) behind pore 
<emphasis id="B9455B11FFD4FA58FC02FDB6FC0FF50D" box="[929,942,593,616]" italics="true" pageId="4" pageNumber="305">ρ</emphasis>
, posterior longitudinal row 
<date id="FF8FA1C3FFD4FA58FB4BFDB6FB5DF53E" box="[1256,1276,589,616]" pageId="4" pageNumber="305">
<emphasis id="B9455B11FFD4FA58FB4BFDB6FB5DF53E" box="[1256,1276,589,616]" italics="true" pageId="4" pageNumber="305">x2</emphasis>
</date>
and row 
<emphasis id="B9455B11FFD4FA58FAC6FDB6FAD0F50D" box="[1381,1393,593,616]" italics="true" pageId="4" pageNumber="305">y</emphasis>
not visible. Axillary rows 
<emphasis id="B9455B11FFD4FA58FE36FD9FFE17F5E7" box="[405,438,628,655]" italics="true" pageId="4" pageNumber="305">as1</emphasis>
(3), 
<emphasis id="B9455B11FFD4FA58FE45FD9FFDA6F5E7" box="[486,519,628,655]" italics="true" pageId="4" pageNumber="305">as2</emphasis>
(3), 
<emphasis id="B9455B11FFD4FA58FD98FD9FFDFDF5E7" box="[571,604,628,655]" italics="true" pageId="4" pageNumber="305">as3</emphasis>
(2–4), 
<emphasis id="B9455B11FFD4FA58FD0BFD90FD67F5E7" box="[680,710,628,655]" italics="true" pageId="4" pageNumber="305">la1</emphasis>
(1) and 
<emphasis id="B9455B11FFD4FA58FC8BFD90FCE4F5E7" box="[808,837,628,655]" italics="true" pageId="4" pageNumber="305">la2</emphasis>
(1) present. (5) 
<emphasis id="B9455B11FFD4FA58FBA3FD90FBCCF5EA" box="[1024,1133,631,655]" italics="true" pageId="4" pageNumber="305">opercular</emphasis>
: transverse row 
<emphasis id="B9455B11FFD4FA58FA88FD9FFAE0F5EA" box="[1323,1345,632,655]" italics="true" pageId="4" pageNumber="305">ot</emphasis>
(6–10); superior longitudinal row 
<emphasis id="B9455B11FFD4FA58FE18FD7BFE75F5D6" box="[443,468,668,691]" italics="true" pageId="4" pageNumber="305">os</emphasis>
(3); inferior longitudinal row 
<emphasis id="B9455B11FFD4FA58FC84FD7BFC9CF5D6" box="[807,829,668,691]" italics="true" pageId="4" pageNumber="305">oi</emphasis>
(2). (6) 
<emphasis id="B9455B11FFD4FA58FC39FD7BFBE4F5D6" box="[922,1093,667,691]" italics="true" pageId="4" pageNumber="305">anterior dorsal</emphasis>
: row 
<emphasis id="B9455B11FFD4FA58FB25FD7BFB35F5D6" box="[1158,1172,668,691]" italics="true" pageId="4" pageNumber="305">n</emphasis>
as single papila behind eye, rows 
<emphasis id="B9455B11FFD4FA58FEAAFD27FEB6F5B2" box="[265,279,704,727]" italics="true" pageId="4" pageNumber="305">g</emphasis>
and 
<emphasis id="B9455B11FFD4FA58FEEEFD27FEC0F5B2" box="[333,353,704,727]" italics="true" pageId="4" pageNumber="305">m</emphasis>
just as single papilla, row 
<emphasis id="B9455B11FFD4FA58FD2EFD27FD3AF5B2" box="[653,667,704,727]" italics="true" pageId="4" pageNumber="305">o</emphasis>
absent and longitudinal row 
<emphasis id="B9455B11FFD4FA58FC42FD58FC4EF5B2" box="[993,1007,703,727]" italics="true" pageId="4" pageNumber="305">h</emphasis>
(3–4) poorly visible among scales.
</paragraph>
</subSubSection>
<caption id="DF4ED78BFFD4FA58FF34F9F2FB7FF12F" httpUri="https://zenodo.org/record/262959/files/figure.png" pageId="4" pageNumber="305" targetBox="[151,1436,780,1535]" targetPageId="4">
<paragraph id="8B8E8703FFD4FA58FF34F9F2FB7FF12F" blockId="4.[151,1436,1557,1610]" pageId="4" pageNumber="305">
<emphasis id="B9455B11FFD4FA58FF34F9F2FEB7F14E" bold="true" box="[151,278,1557,1579]" pageId="4" pageNumber="305">FIGURE 2.</emphasis>
<taxonomicName id="4C31FC80FFD4FA58FE83F9F1FE4FF14E" box="[288,494,1558,1579]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="305" phylum="Chordata" rank="species" species="llorisi" status="sp. nov.">
<emphasis id="B9455B11FFD4FA58FE83F9F1FE4FF14E" box="[288,494,1558,1579]" italics="true" pageId="4" pageNumber="305">Speleogobius llorisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A276E66AFFD4FA58FE5BF9F1FDECF14E" box="[504,589,1558,1579]" pageId="4" pageNumber="305" rank="species">
<emphasis id="B9455B11FFD4FA58FE5BF9F1FDECF14E" bold="true" box="[504,589,1558,1579]" pageId="4" pageNumber="305">sp. nov.</emphasis>
</taxonomicNameLabel>
Holotype: PMR VP3510 (P3), female, 25.6 + 5.6 mm, Spain, Balearic Islands, Mallorca. 
<emphasis id="B9455B11FFD4FA58FF5CF9D3FEB0F12C" bold="true" box="[255,273,1588,1609]" pageId="4" pageNumber="305">A</emphasis>
: freshly collected specimen; 
<emphasis id="B9455B11FFD4FA58FD99F9D2FDEAF12F" bold="true" box="[570,587,1589,1610]" pageId="4" pageNumber="305">B</emphasis>
: preserved specimen. Photos by (A) F. Ordines, (B) M. Kovačić.
</paragraph>
</caption>
<subSubSection id="C32BD488FFD4FA5AFF64F990FE08F6ED" lastPageId="6" lastPageNumber="307" pageId="4" pageNumber="305" type="materials_examined">
<paragraph id="8B8E8703FFD4FA59FF64F990FD7CF6B5" blockId="4.[151,1437,1654,2004]" lastBlockId="5.[151,1436,151,464]" lastPageId="5" lastPageNumber="306" pageId="4" pageNumber="305">
<emphasis id="B9455B11FFD4FA58FF64F990FDF8F1EA" box="[199,601,1655,1679]" italics="true" pageId="4" pageNumber="305">Colour of freshly collected material</emphasis>
. Freshly collected female (
<figureCitation id="130A9B86FFD4FA58FC28F991FC67F1EA" box="[907,966,1654,1679]" captionStart="FIGURE 2" captionStartId="4.[151,250,1557,1579]" captionTargetBox="[151,1436,780,1535]" captionTargetId="figure@4.[151,1436,780,1536]" captionTargetPageId="4" captionText="FIGURE 2. Speleogobius llorisi sp. nov. Holotype: PMR VP 3510 (P 3), female, 25.6 + 5.6 mm, Spain, Balearic Islands, Mallorca. A: freshly collected specimen; B: preserved specimen. Photos by (A) F. Ordines, (B) M. Kovačić." httpUri="https://zenodo.org/record/262959/files/figure.png" pageId="4" pageNumber="305">Fig 2</figureCitation>
A)—most scales had already been lost on the freshly collected specimens, so coloration may have been influenced. Head and body to V and P bases orange red, rest of body whitish-transparent with a pattern of melanophores concentrated at posterior edges of scales, resulting in an imbricated pattern. Predorsal area pink whitish, underside of head and cheek mottled orange red and whitish, anterior chin with two partially contingent blotches of melanophore; a transverse orange-red band on chin posterior part of chin below anterior eye margin; opercle and prepelvic area orange red. On body melanophores more concentrated along lateral midline forming four blotches. Reddish to grey silhouette of vertebral column visible behind. Caudal peduncle with dark blotch below midlateral line, expanding vertically backwards into dark orange vertical stripe with a narrow dark and well defined margin at C base. Small amount of orange red to pink pigment also visible scattered on body, mostly along lateral midline. Dorsum with three broad, transverse dark bands of similar intensity and on flank contiguous with lateral side blotches: the first over D1 base, the second over D2 base and the third over caudal peduncle; dark bands alternating with four whitish saddles. Prepelvic area and V base orange red. Ventral midline and belly whitish. Silvery peritoneum visible bellow skin on belly. Fins mostly transparent and partially pigmented. D1 and D2 with melanophores arranged to an ill-defined band in the middle part and at the fin base and with erythrophores distributed mainly in the upper part of the fins. C at base contingent with posterior dark edge of dark orange vertical caudal peduncle stripe; main part of C with orange yellow dots arranged in ill-defined narrow bands over the rest of the fin. A transparent. P with orange mark in anterior part and dark mark in the middle of the fin, the rest of the fin transparent with scattered erythrophores. V with red orange base, dark middle part and white or transparent tips.
</paragraph>
<caption id="DF4ED78BFFD5FA59FF34FC69FD14F4A6" httpUri="https://zenodo.org/record/262960/files/figure.png" pageId="5" pageNumber="306" targetBox="[151,1435,517,888]" targetPageId="5">
<paragraph id="8B8E8703FFD5FA59FF34FC69FD14F4A6" blockId="5.[151,1436,910,963]" pageId="5" pageNumber="306">
<emphasis id="B9455B11FFD5FA59FF34FC69FEB6F4C6" bold="true" box="[151,279,910,932]" pageId="5" pageNumber="306">FIGURE 3.</emphasis>
<taxonomicName id="4C31FC80FFD5FA59FE81FC69FE50F4C6" box="[290,497,910,931]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="306" phylum="Chordata" rank="species" species="llorisi" status="sp. nov.">
<emphasis id="B9455B11FFD5FA59FE81FC69FE50F4C6" box="[290,497,910,931]" italics="true" pageId="5" pageNumber="306">Speleogobius llorisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A276E66AFFD5FA59FE58FC68FDF1F4C1" box="[507,592,911,932]" pageId="5" pageNumber="306" rank="species">
<emphasis id="B9455B11FFD5FA59FE58FC68FDF1F4C1" bold="true" box="[507,592,911,932]" pageId="5" pageNumber="306">sp. nov.</emphasis>
</taxonomicNameLabel>
Paratype: PMR VP3512 (P54), male, 24.4 + 5.5 mm, Spain, Balearic Islands, Mallorca. Preserved specimen. Photo by M. Kovačić.
</paragraph>
</caption>
<caption id="DF4ED78BFFD5FA59FF34F9BBFB94F1F5" httpUri="https://zenodo.org/record/262961/files/figure.png" pageId="5" pageNumber="306" targetBox="[179,1414,1022,1594]" targetPageId="5">
<paragraph id="8B8E8703FFD5FA59FF34F9BBFB94F1F5" blockId="5.[151,1436,1628,1680]" pageId="5" pageNumber="306">
<emphasis id="B9455B11FFD5FA59FF34F9BBFEB3F114" bold="true" box="[151,274,1628,1650]" pageId="5" pageNumber="306">FIGURE 4.</emphasis>
<taxonomicName id="4C31FC80FFD5FA59FEBAF9BBFE42F114" box="[281,483,1628,1649]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="306" phylum="Chordata" rank="species" species="llorisi" status="sp. nov.">
<emphasis id="B9455B11FFD5FA59FEBAF9BBFE42F114" box="[281,483,1628,1649]" italics="true" pageId="5" pageNumber="306">Speleogobius llorisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A276E66AFFD5FA59FE4BF9BBFD98F114" box="[488,569,1628,1649]" pageId="5" pageNumber="306" rank="species">
<emphasis id="B9455B11FFD5FA59FE4BF9BBFD98F114" bold="true" box="[488,569,1628,1649]" pageId="5" pageNumber="306">sp. nov.</emphasis>
</taxonomicNameLabel>
, head LL sensory papillae and canal pores: paratype, PMR VP3512 (P54), male, 24.4 + 5.5 mm, Spain, Balearic Islands, Mallorca. Terminology in text. Drawing by M. Kovačić.
</paragraph>
</caption>
<paragraph id="8B8E8703FFD5FA5AFF64F95FFF5BF79D" blockId="5.[151,1437,1719,2032]" lastBlockId="6.[151,1437,151,1473]" lastPageId="6" lastPageNumber="307" pageId="5" pageNumber="306">
<emphasis id="B9455B11FFD5FA59FF64F95FFDDCF1B5" box="[199,637,1720,1744]" italics="true" pageId="5" pageNumber="306">Colour preserved specimens in alcohol</emphasis>
(
<figureCitation id="130A9B86FFD5FA59FD2FF950FD78F1B5" box="[652,729,1719,1744]" captionStart="FIGURE 2" captionStartId="4.[151,250,1557,1579]" captionTargetBox="[151,1436,780,1535]" captionTargetId="figure@4.[151,1436,780,1536]" captionTargetPageId="4" captionText="FIGURE 2. Speleogobius llorisi sp. nov. Holotype: PMR VP 3510 (P 3), female, 25.6 + 5.6 mm, Spain, Balearic Islands, Mallorca. A: freshly collected specimen; B: preserved specimen. Photos by (A) F. Ordines, (B) M. Kovačić." httpUri="https://zenodo.org/record/262959/files/figure.png" pageId="5" pageNumber="306">Figs. 2</figureCitation>
B and 3 — most scales missing on the preserved specimens, so preserved coloration may have been influenced). Body yellowish white. Melanophores on body concentrated on posterior edges of scales (or scale positions when scales are lost) resulting in an imbricated pattern on lateral and upper sides of body. Melanophores more concentrated along lateral midline forming elongate blotches in adults, in juvenile as a continuous longitudinal stripe along posterior part of lateral midline, and on dorsal side forming dark bands. Caudal peduncle with larger dark blotch in lower part. Predorsal area, opercle and cheek mottled to dotted with pigments. Head underside mostly whitish with various quantities of pigments present. D1 pigmented with melanophores, but no melanophores visible in the juvenile male. D2 and A pigmented in adult males, not in the juvenile male. D2 poorly pigmented and A transparent in females. C transparent, except for dark pigments at the fin base. P transparent, with dark blotch at lower part of the fin center, not visible in the juvenile. Most of the V pigmented in adult male, females with pigmentation only at anterior half of the fin, the juvenile almost without pigment.
</paragraph>
<paragraph id="8B8E8703FFD6FA5AFF64FEE3FE08F6ED" blockId="6.[151,1437,151,1473]" pageId="6" pageNumber="307">
Vertebral column and pterygiophore insertion pattern (pty). 12 precaudal and 17 caudal vertebrae (including urostyle); total count 29. One epural; pty 3–2211000; two pre-haemal pterygiophores (based on four non-type specimens listed under “Additional Material“). These counts are identical with those of four specimens of 
<taxonomicName id="4C31FC80FFD6FA5AFA24FEABFEA2F6ED" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="307" phylum="Chordata" rank="species" species="trigloides">
<emphasis id="B9455B11FFD6FA5AFA24FEABFEA2F6ED" italics="true" pageId="6" pageNumber="307">S. trigloides</emphasis>
</taxonomicName>
(
<collectionCode id="ED201FC6FFD6FA5AFEB2FE97FEEDF6ED" box="[273,332,368,392]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="6" pageNumber="307">ZSM</collectionCode>
35457).
</paragraph>
</subSubSection>
<subSubSection id="C32BD488FFD6FA5AFF64FE73FC1EF489" pageId="6" pageNumber="307" type="etymology">
<paragraph id="8B8E8703FFD6FA5AFF64FE73FADAF6B5" blockId="6.[151,1437,151,1473]" pageId="6" pageNumber="307">
<emphasis id="B9455B11FFD6FA5AFF64FE73FEE7F6C8" bold="true" box="[199,326,404,429]" pageId="6" pageNumber="307">Etymology</emphasis>
. The species is named 
<taxonomicName id="4C31FC80FFD6FA5AFDE8FE73FD2FF6C9" box="[587,654,404,428]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="307" phylum="Chordata" rank="species" species="llorisi">
<emphasis id="B9455B11FFD6FA5AFDE8FE73FD2FF6C9" box="[587,654,404,428]" italics="true" pageId="6" pageNumber="307">llorisi</emphasis>
</taxonomicName>
in honor of Dr. Domenec Lloris, an ichthyologist from the Institut de Ciències del Mar, in Barcelona, in recognition for his outstanding contribution to the knowledge of fish species.
</paragraph>
<paragraph id="8B8E8703FFD6FA5AFF64FE3BFC1EF489" blockId="6.[151,1437,151,1473]" pageId="6" pageNumber="307">
<emphasis id="B9455B11FFD6FA5AFF64FE3BFE82F690" bold="true" box="[199,291,476,501]" pageId="6" pageNumber="307">Habitat</emphasis>
. All individuals of the new species of 
<taxonomicName id="4C31FC80FFD6FA5AFD74FE3BFCCDF691" box="[727,876,476,500]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="307" phylum="Chordata" rank="genus">
<emphasis id="B9455B11FFD6FA5AFD74FE3BFCCDF691" box="[727,876,476,500]" italics="true" pageId="6" pageNumber="307">Speleogobius</emphasis>
</taxonomicName>
were collected from sampling stations located on red algae beds. These beds are one of the most widespread 
<typeStatus id="548A39A1FFD6FA5AFC91FDE7FCCCF57D" box="[818,877,512,536]" pageId="6" pageNumber="307">types</typeStatus>
of habitats of the shallow continental shelf of the Balearic Islands in depths between 50 and 
<quantity id="4CC92AE6FFD6FA5AFDD1FDC3FD05F559" box="[626,676,548,572]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="307" unit="m" value="80.0">80m</quantity>
(
<bibRefCitation id="EFA0FAF2FFD6FA5AFD11FDC3FCD2F559" author="Ballesteros" box="[690,883,548,573]" pageId="6" pageNumber="310" refString="Ballesteros, E. (1994) The deep-water Peyssonnelia beds from the Balearic Islands (western Mediterranean). P. S. Z. N. I. Marine Ecology, 15, 233 - 253. http: // dx. doi. org / 10.1111 / j. 1439 - 0485.1994. tb 00055. x" type="journal article" year="1994">Ballesteros 1994</bibRefCitation>
; 
<bibRefCitation id="EFA0FAF2FFD6FA5AFCDEFDC3FB34F559" author="Ordines" box="[893,1173,548,573]" pageId="6" pageNumber="310" refString="Ordines, F. &amp; Massuti, E. (2009) Relationships between macro-epibenthic communities and fish on the shelf grounds of the western Mediterranean. Aquatic Conservation: Marine and Freshwater Ecosystems, 19, 370 - 383. http: // dx. doi. org / 10.1002 / aqc. 969" type="journal article" year="2009">Ordines &amp; Massutí 2009</bibRefCitation>
). The sampling stations where the specimens of the new species occurred presented very similar epibenthic communities in which predominant algae species on the basal stratum belong to the families 
<taxonomicName id="4C31FC80FFD6FA5AFBA0FD8BFB3CF5E0" box="[1027,1181,620,645]" class="Florideophyceae" family="Corallinaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Corallinales" pageId="6" pageNumber="307" phylum="Rhodophyta" rank="family">Corallinaceae</taxonomicName>
(with 
<taxonomicName id="4C31FC80FFD6FA5AFB54FD8BFEBDF5CD" class="Florideophyceae" family="Corallinaceae" genus="Lithothamnion" higherTaxonomySource="GBIF" kingdom="Plantae" order="Corallinales" pageId="6" pageNumber="307" phylum="Rhodophyta" rank="species" species="corallioides">
<emphasis id="B9455B11FFD6FA5AFB54FD8BFEBDF5CD" italics="true" pageId="6" pageNumber="307">Lithothamnion corallioides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C31FC80FFD6FA5AFE88FD77FDBEF5CD" box="[299,543,656,680]" class="Florideophyceae" family="Corallinaceae" genus="Lithothamnion" higherTaxonomySource="GBIF" kingdom="Plantae" order="Corallinales" pageId="6" pageNumber="307" phylum="Rhodophyta" rank="species" species="valens">
<emphasis id="B9455B11FFD6FA5AFE88FD77FDBEF5CD" box="[299,543,656,680]" italics="true" pageId="6" pageNumber="307">Lithothamnion valens</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C31FC80FFD6FA5AFD8DFD77FCF1F5CD" box="[558,848,656,680]" class="Florideophyceae" family="Corallinaceae" genus="Phymatolithon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Corallinales" pageId="6" pageNumber="307" phylum="Rhodophyta" rank="species" species="calcareum">
<emphasis id="B9455B11FFD6FA5AFD8DFD77FCF1F5CD" box="[558,848,656,680]" italics="true" pageId="6" pageNumber="307">Phymatolithon calcareum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C31FC80FFD6FA5AFC2BFD77FBC9F5CD" box="[904,1128,656,680]" class="Florideophyceae" family="Corallinaceae" genus="Spongites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Corallinales" pageId="6" pageNumber="307" phylum="Rhodophyta" rank="species" species="fruticolus">
<emphasis id="B9455B11FFD6FA5AFC2BFD77FBC9F5CD" box="[904,1128,656,680]" italics="true" pageId="6" pageNumber="307">Spongites fruticolus</emphasis>
</taxonomicName>
among the most abundant) and to Peyssonneliacea (with 
<taxonomicName id="4C31FC80FFD6FA5AFE4AFD53FD64F5A9" box="[489,709,692,716]" class="Florideophyceae" family="Peyssonneliaceae" genus="Peyssonnelia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cryptonemiales" pageId="6" pageNumber="307" phylum="Rhodophyta" rank="species" species="rubra">
<emphasis id="B9455B11FFD6FA5AFE4AFD53FD64F5A9" box="[489,709,692,716]" italics="true" pageId="6" pageNumber="307">Peyssonnelia rubra</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C31FC80FFD6FA5AFD5DFD53FBABF5A9" box="[766,1034,692,716]" class="Florideophyceae" family="Peyssonneliaceae" genus="Peyssonnelia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cryptonemiales" pageId="6" pageNumber="307" phylum="Rhodophyta" rank="species" species="inamoena">
<emphasis id="B9455B11FFD6FA5AFD5DFD53FBABF5A9" box="[766,1034,692,716]" italics="true" pageId="6" pageNumber="307">Peyssonnelia inamoena</emphasis>
</taxonomicName>
among the most abundant), and on the erect stratum to 
<taxonomicName id="4C31FC80FFD6FA5AFEDEFD30FD99F595" box="[381,568,727,752]" class="Florideophyceae" family="Phyllophoraceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gigartinales" pageId="6" pageNumber="307" phylum="Rhodophyta" rank="family">Phyllophoraceae</taxonomicName>
(with 
<taxonomicName id="4C31FC80FFD6FA5AFD27FD3FFCFEF58A" box="[644,863,728,752]" class="Insecta" family="Phaneropteridae" genus="Phyllophora" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="crispa">
<emphasis id="B9455B11FFD6FA5AFD27FD3FFCFEF58A" box="[644,863,728,752]" italics="true" pageId="6" pageNumber="307">Phyllophora crispa</emphasis>
</taxonomicName>
). 
<collectingCountry id="F326C793FFD6FA5AFCDAFD3FFC39F595" box="[889,920,728,752]" name="American Samoa" pageId="6" pageNumber="307">As</collectingCountry>
a result of the high biomass of 
<taxonomicName id="4C31FC80FFD6FA5AFAABFD3FFA3DF595" box="[1288,1436,728,752]" class="Florideophyceae" family="Peyssonneliaceae" genus="Peyssonnelia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cryptonemiales" pageId="6" pageNumber="307" phylum="Rhodophyta" rank="genus">
<emphasis id="B9455B11FFD6FA5AFAABFD3FFA3DF595" box="[1288,1436,728,752]" italics="true" pageId="6" pageNumber="307">Peyssonnelia</emphasis>
</taxonomicName>
spp., the habitat present at the sampling stations can be classified as 
<taxonomicName id="4C31FC80FFD6FA5AFC19FD1BFBEFF471" box="[954,1102,764,788]" class="Florideophyceae" family="Peyssonneliaceae" genus="Peyssonnelia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cryptonemiales" pageId="6" pageNumber="307" phylum="Rhodophyta" rank="genus">
<emphasis id="B9455B11FFD6FA5AFC19FD1BFBEFF471" box="[954,1102,764,788]" italics="true" pageId="6" pageNumber="307">Peyssonnelia</emphasis>
</taxonomicName>
beds, a frequently occurring habitat in the south west of Mallorca, and the south of 
<collectingCountry id="F326C793FFD6FA5AFD58FCC7FCC3F452" box="[763,866,800,823]" name="Spain" pageId="6" pageNumber="307">Menorca</collectingCountry>
, that presents the highest biomasses of algae in the circalittoral sea bed of the Balearic Islands. The 
<taxonomicName id="4C31FC80FFD6FA5AFD7EFCA3FCD0F439" box="[733,881,836,860]" class="Florideophyceae" family="Peyssonneliaceae" genus="Peyssonnelia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cryptonemiales" pageId="6" pageNumber="307" phylum="Rhodophyta" rank="genus">
<emphasis id="B9455B11FFD6FA5AFD7EFCA3FCD0F439" box="[733,881,836,860]" italics="true" pageId="6" pageNumber="307">Peyssonnelia</emphasis>
</taxonomicName>
beds are frequently associated with open bays characterized by fine sediments, even mud, where calm periods alternate with periods of eddy forming currents (
<bibRefCitation id="EFA0FAF2FFD6FA5AFF3DFC6BFEBEF4C0" author="Peres" box="[158,287,908,933]" pageId="6" pageNumber="310" refString="Peres, J. M. (1985) History of the Mediterranean biota and the colonization of the depths. In: Margalef, R. (Ed.), Western Mediterranean. Pergamon Press, London, pp. 198 - 232." type="book chapter" year="1985">Pérès 1985</bibRefCitation>
; 
<bibRefCitation id="EFA0FAF2FFD6FA5AFE8FFC6BFDA6F4C1" author="Ordines" box="[300,519,908,933]" pageId="6" pageNumber="310" refString="Ordines, F., Jorda, G., Quetglas, A., Flexas, M., Moranta, J. &amp; Massuti, E. (2011) Connections between hydrodynamics, benthic landscape and associated fauna in the Balearic Islands, western Mediterranean. Continental Shelf Research, 31, 1835 - 1844. http: // dx. doi. org / 10.1016 / j. csr. 2011.08.007" type="journal article" year="2011">
Ordines 
<emphasis id="B9455B11FFD6FA5AFE2FFC6AFE67F4C1" box="[396,454,908,932]" italics="true" pageId="6" pageNumber="307">et al.</emphasis>
2011
</bibRefCitation>
; 
<bibRefCitation id="EFA0FAF2FFD6FA5AFDB7FC6BFD77F4C1" author="Joher" box="[532,726,908,933]" pageId="6" pageNumber="310" refString="Joher, S., Ballesteros, E., Cebrian, E., Sanchez, N. &amp; Rodriguez-Prieto, C. (2012) Deep-water macroalgal-dominated coastal detritic assemblages on the continental shelf off Mallorca and Menorca (Balearic Islands, Western Mediterranean). Botanica Marina, 55 (5), 485 - 497. http: // dx. doi. org / 10.1515 / bot- 2012 - 0113" type="journal article" year="2012">
Joher 
<emphasis id="B9455B11FFD6FA5AFDFBFC6AFD30F4C1" box="[600,657,908,932]" italics="true" pageId="6" pageNumber="307">et al.</emphasis>
2012
</bibRefCitation>
). This was the location of the sampling stations where the new 
<taxonomicName id="4C31FC80FFD6FA5AFF34FC57FE8DF4AD" box="[151,300,944,968]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="307" phylum="Chordata" rank="genus">
<emphasis id="B9455B11FFD6FA5AFF34FC57FE8DF4AD" box="[151,300,944,968]" italics="true" pageId="6" pageNumber="307">Speleogobius</emphasis>
</taxonomicName>
species was found: a sheltered area, without significant currents, and at the lee of the north winds, which cause the most important storms in the area (
<bibRefCitation id="EFA0FAF2FFD6FA5AFD74FC33FC12F489" author="Ordines" box="[727,947,980,1005]" pageId="6" pageNumber="310" refString="Ordines, F., Jorda, G., Quetglas, A., Flexas, M., Moranta, J. &amp; Massuti, E. (2011) Connections between hydrodynamics, benthic landscape and associated fauna in the Balearic Islands, western Mediterranean. Continental Shelf Research, 31, 1835 - 1844. http: // dx. doi. org / 10.1016 / j. csr. 2011.08.007" type="journal article" year="2011">
Ordines 
<emphasis id="B9455B11FFD6FA5AFC9BFC32FCD0F489" box="[824,881,980,1004]" italics="true" pageId="6" pageNumber="307">et al.</emphasis>
2011
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32BD488FFD6FA54FF64FC10FCD8F79D" lastPageId="8" lastPageNumber="309" pageId="6" pageNumber="307" type="description">
<paragraph id="8B8E8703FFD6FA5AFF64FC10FDE9F2A5" blockId="6.[151,1437,151,1473]" pageId="6" pageNumber="307">
<emphasis id="B9455B11FFD6FA5AFF64FC10FD93F375" bold="true" box="[199,562,1015,1040]" pageId="6" pageNumber="307">
Comparison with 
<taxonomicName id="4C31FC80FFD6FA5AFE03FC10FD8AF375" box="[416,555,1015,1040]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="307" phylum="Chordata" rank="species" species="trigloides">
<emphasis id="B9455B11FFD6FA5AFE03FC10FD8AF375" bold="true" box="[416,555,1015,1040]" italics="true" pageId="6" pageNumber="307">S. trigloides</emphasis>
</taxonomicName>
.
</emphasis>
<taxonomicName id="4C31FC80FFD6FA5AFD99FC1FFCBAF375" box="[570,795,1016,1040]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="307" phylum="Chordata" rank="species" species="llorisi" status="sp. nov.">
<emphasis id="B9455B11FFD6FA5AFD99FC1FFCBAF375" box="[570,795,1016,1040]" italics="true" pageId="6" pageNumber="307">Speleogobius llorisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A276E66AFFD6FA5AFC87FC10FCDFF375" box="[804,894,1015,1040]" pageId="6" pageNumber="307" rank="species">
<emphasis id="B9455B11FFD6FA5AFC87FC10FCDFF375" bold="true" box="[804,894,1015,1040]" pageId="6" pageNumber="307">sp. nov.</emphasis>
</taxonomicNameLabel>
is easily separated from 
<taxonomicName id="4C31FC80FFD6FA5AFB3EFC1FFA86F375" box="[1181,1319,1016,1040]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="307" phylum="Chordata" rank="species" species="trigloides">
<emphasis id="B9455B11FFD6FA5AFB3EFC1FFA86F375" box="[1181,1319,1016,1040]" italics="true" pageId="6" pageNumber="307">S. trigloides</emphasis>
</taxonomicName>
(
<figureCitation id="130A9B86FFD6FA5AFA95FC10FA21F375" box="[1334,1408,1015,1040]" captionStart="FIGURE 2" captionStartId="4.[151,250,1557,1579]" captionTargetBox="[151,1436,780,1535]" captionTargetId="figure@4.[151,1436,780,1536]" captionTargetPageId="4" captionText="FIGURE 2. Speleogobius llorisi sp. nov. Holotype: PMR VP 3510 (P 3), female, 25.6 + 5.6 mm, Spain, Balearic Islands, Mallorca. A: freshly collected specimen; B: preserved specimen. Photos by (A) F. Ordines, (B) M. Kovačić." httpUri="https://zenodo.org/record/262959/files/figure.png" pageId="6" pageNumber="307">Figs 2</figureCitation>
, 
<figureCitation id="130A9B86FFD6FA5AFA2DFC1FFA3DF375" box="[1422,1436,1016,1040]" captionStart="FIGURE 3" captionStartId="5.[151,250,910,932]" captionTargetBox="[151,1435,517,888]" captionTargetId="figure@5.[151,1436,517,889]" captionTargetPageId="5" captionText="FIGURE 3. Speleogobius llorisi sp. nov. Paratype: PMR VP 3512 (P 54), male, 24.4 + 5.5 mm, Spain, Balearic Islands, Mallorca. Preserved specimen. Photo by M. Kovačić." httpUri="https://zenodo.org/record/262960/files/figure.png" pageId="6" pageNumber="307">3</figureCitation>
and 
<figureCitation id="130A9B86FFD6FA5AFF65FBFBFF75F350" box="[198,212,1052,1077]" captionStart="FIGURE 5" captionStartId="6.[151,250,1893,1915]" captionTargetBox="[151,1436,1525,1871]" captionTargetId="figure@6.[151,1436,1525,1872]" captionTargetPageId="6" captionText="FIGURE 5. Speleogobius llorisi sp. nov. X-ray of specimen ZSM-PIS-GO- 1067 / 1068, undetermined sex, 21.1 + 4.1 mm, Spain, Balearic Islands, Mallorca. X-ray by Z. Gholami." httpUri="https://zenodo.org/record/262962/files/figure.png" pageId="6" pageNumber="307">5</figureCitation>
) by several morphological characters: preopercular head canal present with pores 
<emphasis id="B9455B11FFD6FA5AFBCAFBFAFB6CF351" box="[1129,1229,1052,1076]" italics="true" pageId="6" pageNumber="307">γ, δ, ε vs.</emphasis>
preopercular head canal absent; snout long, equal or longer than eye, 1.0– 
<quantity id="4CC92AE6FFD6FA5AFCA9FBA7FCE9F33D" box="[778,840,1087,1112]" metricMagnitude="-2" metricUnit="m" metricValue="2.794" pageId="6" pageNumber="307" unit="in" value="1.1">1.1 in</quantity>
eye length, dorsal profile of snout gently sloping 
<emphasis id="B9455B11FFD6FA5AFADEFBA6FA3DF33D" box="[1405,1436,1089,1112]" italics="true" pageId="6" pageNumber="307">vs.</emphasis>
snout short, shorter than eye, 
<quantity id="4CC92AE6FFD6FA5AFE41FB83FDF2F318" box="[482,595,1124,1149]" metricMagnitude="-2" metricUnit="m" metricValue="1.7779999999999998" metricValueMax="2.032" metricValueMin="1.524" pageId="6" pageNumber="307" unit="in" value="0.7" valueMax="0.8" valueMin="0.6">0.6–0.8 in</quantity>
eye length (measured on comparative material), snout dorsal profile steep; lower lip slightly protruding upper lip (prognathous) 
<emphasis id="B9455B11FFD6FA5AFD5DFB6EFCBCF3C5" box="[766,797,1161,1184]" italics="true" pageId="6" pageNumber="307">vs.</emphasis>
upper lip slightly protruding lower lip (retrognathous); scales in lateral series 28 or 29 
<emphasis id="B9455B11FFD6FA5AFE51FB4AFDB0F3A1" box="[498,529,1197,1220]" italics="true" pageId="6" pageNumber="307">vs.</emphasis>
26; scales in transverse series 6 
<emphasis id="B9455B11FFD6FA5AFCD9FB4AFC39F3A1" box="[890,920,1197,1220]" italics="true" pageId="6" pageNumber="307">vs.</emphasis>
7–8 (
<quantity id="4CC92AE6FFD6FA5AFC71FB4BFC5CF3A0" box="[978,1021,1196,1221]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="6" pageNumber="307" unit="in" value="7.0">7 in</quantity>
<typeStatus id="548A39A1FFD6FA5AFBA0FB4BFBCBF3A0" box="[1027,1130,1196,1221]" pageId="6" pageNumber="307" type="holotype">holotype</typeStatus>
); head length as percentage of standard length: 28.7–30.0% adults, 32.2% juvenile 
<emphasis id="B9455B11FFD6FA5AFC90FB36FCEBF38D" box="[819,842,1233,1256]" italics="true" pageId="6" pageNumber="307">vs</emphasis>
. 25.5%–29.1% in comparative material, 
<typeStatus id="548A39A1FFD6FA5AFA99FB28FA3DF38D" box="[1338,1436,1231,1256]" pageId="6" pageNumber="307" type="holotype">holotype</typeStatus>
recalculated to percentages 27.9% (overlap present only with 
<taxonomicName id="4C31FC80FFD6FA5AFCEBFB13FC71F269" box="[840,976,1268,1292]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="307" phylum="Chordata" rank="species" species="trigloides">
<emphasis id="B9455B11FFD6FA5AFCEBFB13FC71F269" box="[840,976,1268,1292]" italics="true" pageId="6" pageNumber="307">S. trigloides</emphasis>
</taxonomicName>
of standard length 
<quantity id="4CC92AE6FFD6FA5AFB04FB13FB57F269" box="[1191,1270,1268,1293]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="307" unit="mm" value="15.0">15 mm</quantity>
or smaller due to allometry), i.e. the new species has longer head compared to body length; body depth at the pelvic fin origin as percentage of standard length: 15.0–16.8% 
<emphasis id="B9455B11FFD6FA5AFD38FADAFD13F231" box="[667,690,1341,1364]" italics="true" pageId="6" pageNumber="307">vs</emphasis>
. 19.3–20.6% in comparative material, 
<typeStatus id="548A39A1FFD6FA5AFB20FADBFB44F230" box="[1155,1253,1340,1365]" pageId="6" pageNumber="307" type="holotype">holotype</typeStatus>
recalculated to percentages 25.1% i.e. body of new species is more slender; eye diameter (as percentage of standard length): 6.9– 7.7% 
<emphasis id="B9455B11FFD6FA5AFF7AFA62FF51F2F9" box="[217,240,1413,1436]" italics="true" pageId="6" pageNumber="307">vs</emphasis>
. 8.1%–9.7% in comparative material, 
<typeStatus id="548A39A1FFD6FA5AFD06FA63FCA6F2F8" box="[677,775,1412,1437]" pageId="6" pageNumber="307" type="holotype">holotype</typeStatus>
recalculated to percentages 9.3%, i.e. the new species has smaller eyes compared to body length.
</paragraph>
<caption id="DF4ED78BFFD6FA5AFF34F882FD70F0FF" httpUri="https://zenodo.org/record/262962/files/figure.png" pageId="6" pageNumber="307" targetBox="[151,1436,1525,1871]" targetPageId="6">
<paragraph id="8B8E8703FFD6FA5AFF34F882FD70F0FF" blockId="6.[151,1436,1893,1946]" pageId="6" pageNumber="307">
<emphasis id="B9455B11FFD6FA5AFF34F882FEB4F01E" bold="true" box="[151,277,1893,1915]" pageId="6" pageNumber="307">FIGURE 5.</emphasis>
<taxonomicName id="4C31FC80FFD6FA5AFEBCF882FE4CF01F" box="[287,493,1893,1914]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="307" phylum="Chordata" rank="species" species="llorisi" status="sp. nov.">
<emphasis id="B9455B11FFD6FA5AFEBCF882FE4CF01F" box="[287,493,1893,1914]" italics="true" pageId="6" pageNumber="307">Speleogobius llorisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A276E66AFFD6FA5AFE56F881FDE8F01E" box="[501,585,1894,1915]" pageId="6" pageNumber="307" rank="species">
<emphasis id="B9455B11FFD6FA5AFE56F881FDE8F01E" bold="true" box="[501,585,1894,1915]" pageId="6" pageNumber="307">sp. nov.</emphasis>
</taxonomicNameLabel>
X-ray of specimen ZSM-PIS-GO-1067/1068, undetermined sex, 21.1 + 4.1 mm, Spain, Balearic Islands, Mallorca. X-ray by Z. Gholami.
</paragraph>
</caption>
<caption id="DF4ED78BFFD7FA5BFF34F987FD8BF194" httpUri="https://zenodo.org/record/262963/files/figure.png" pageId="7" pageNumber="308" targetBox="[311,1275,193,1609]" targetPageId="7">
<paragraph id="8B8E8703FFD7FA5BFF34F987FD8BF194" blockId="7.[151,1436,1632,1777]" pageId="7" pageNumber="308">
<emphasis id="B9455B11FFD7FA5BFF34F987FEB5F110" bold="true" box="[151,276,1632,1654]" pageId="7" pageNumber="308">FIGURE 6.</emphasis>
<taxonomicName id="4C31FC80FFD7FA5BFEBFF987FDACF110" box="[284,525,1632,1653]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="trigloides">
<emphasis id="B9455B11FFD7FA5BFEBFF987FDACF110" box="[284,525,1632,1653]" italics="true" pageId="7" pageNumber="308">Speleogobius trigloides</emphasis>
</taxonomicName>
. Preserved specimens. 
<emphasis id="B9455B11FFD7FA5BFD5CF987FCB0F110" bold="true" box="[767,785,1632,1653]" pageId="7" pageNumber="308">A</emphasis>
: PMR VP3103, female, 19.6 + 4.9 mm, 6 June 2013, Adriatic Sea, Croatia, the Kvarner area, Oštro; 
<emphasis id="B9455B11FFD7FA5BFD81F998FD92F1F1" bold="true" box="[546,563,1663,1684]" pageId="7" pageNumber="308">B</emphasis>
: PMR VP3331, male, 16.5 + 4.1 mm, 11 June 2013, Adriatic Sea, Croatia, the Hvar island, Smočiguzica. 
<emphasis id="B9455B11FFD7FA5BFED6F97AFE26F1D6" bold="true" box="[373,391,1693,1715]" pageId="7" pageNumber="308">C</emphasis>
: ZSM PIS-GO-552, female, 15 May 2013, Adriatic Sea, Croatia, Kvarner area, Bay of Žurkovo at Kostrena; 
<emphasis id="B9455B11FFD7FA5BFF5CF95BFEB0F1B4" bold="true" box="[255,273,1724,1745]" pageId="7" pageNumber="308">D</emphasis>
: ZSM PIS-GO-883, male, 3 Sept 2013, Adriatic Sea, Croatia, Kvarner area, Cres Island at Pod Beli. Photos (A, B) by M. Kovačić, (C,D) by U. Schliewen.
</paragraph>
</caption>
<paragraph id="8B8E8703FFD7FA54FF64F8FFFCD8F79D" blockId="7.[151,1436,1816,2021]" lastBlockId="8.[151,1436,151,248]" lastPageId="8" lastPageNumber="309" pageId="7" pageNumber="308">
<taxonomicName id="4C31FC80FFD7FA5BFF64F8FFFE8FF055" box="[199,302,1816,1840]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="llorisi" status="sp. nov.">
<emphasis id="B9455B11FFD7FA5BFF64F8FFFE8FF055" box="[199,302,1816,1840]" italics="true" pageId="7" pageNumber="308">S. llorisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A276E66AFFD7FA5BFE96F8FFFE2EF054" box="[309,399,1816,1841]" pageId="7" pageNumber="308" rank="species">
<emphasis id="B9455B11FFD7FA5BFE96F8FFFE2EF054" bold="true" box="[309,399,1816,1841]" pageId="7" pageNumber="308">sp. nov.</emphasis>
</taxonomicNameLabel>
is also easily separated from 
<taxonomicName id="4C31FC80FFD7FA5BFD7DF8FFFCC7F055" box="[734,870,1816,1840]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="trigloides">
<emphasis id="B9455B11FFD7FA5BFD7DF8FFFCC7F055" box="[734,870,1816,1840]" italics="true" pageId="7" pageNumber="308">S. trigloides</emphasis>
</taxonomicName>
by several coloration characters (
<figureCitation id="130A9B86FFD7FA5BFB41F8FFFA8DF055" box="[1250,1324,1816,1841]" captionStart="FIGURE 2" captionStartId="4.[151,250,1557,1579]" captionTargetBox="[151,1436,780,1535]" captionTargetId="figure@4.[151,1436,780,1536]" captionTargetPageId="4" captionText="FIGURE 2. Speleogobius llorisi sp. nov. Holotype: PMR VP 3510 (P 3), female, 25.6 + 5.6 mm, Spain, Balearic Islands, Mallorca. A: freshly collected specimen; B: preserved specimen. Photos by (A) F. Ordines, (B) M. Kovačić." httpUri="https://zenodo.org/record/262959/files/figure.png" pageId="7" pageNumber="308">Figs 2</figureCitation>
, 
<figureCitation id="130A9B86FFD7FA5BFA9BF8FFFAE7F055" box="[1336,1350,1816,1840]" captionStart="FIGURE 3" captionStartId="5.[151,250,910,932]" captionTargetBox="[151,1435,517,888]" captionTargetId="figure@5.[151,1436,517,889]" captionTargetPageId="5" captionText="FIGURE 3. Speleogobius llorisi sp. nov. Paratype: PMR VP 3512 (P 54), male, 24.4 + 5.5 mm, Spain, Balearic Islands, Mallorca. Preserved specimen. Photo by M. Kovačić." httpUri="https://zenodo.org/record/262960/files/figure.png" pageId="7" pageNumber="308">3</figureCitation>
and 
<figureCitation id="130A9B86FFD7FA5BFADDF8FFFA2CF054" box="[1406,1421,1816,1841]" captionStart="FIGURE 5" captionStartId="6.[151,250,1893,1915]" captionTargetBox="[151,1436,1525,1871]" captionTargetId="figure@6.[151,1436,1525,1872]" captionTargetPageId="6" captionText="FIGURE 5. Speleogobius llorisi sp. nov. X-ray of specimen ZSM-PIS-GO- 1067 / 1068, undetermined sex, 21.1 + 4.1 mm, Spain, Balearic Islands, Mallorca. X-ray by Z. Gholami." httpUri="https://zenodo.org/record/262962/files/figure.png" pageId="7" pageNumber="308">5</figureCitation>
), at least with regard to adult females (coloration of adult 
<taxonomicName id="4C31FC80FFD7FA5BFC92F8DBFC32F031" box="[817,915,1852,1876]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="llorisi">
<emphasis id="B9455B11FFD7FA5BFC92F8DBFC32F031" box="[817,915,1852,1876]" italics="true" pageId="7" pageNumber="308">S. llorisi</emphasis>
</taxonomicName>
males remains unknown): head and body to pectoral and pelvic fins bright orange reddish, pale orange stripes on C, blackish pelvic fin centers with white margins, and posterior part of body whitish-transparent with a dark reticulate pattern and small amount of orange red pigment 
<emphasis id="B9455B11FFD7FA5BFE8BF84EFE9EF0A5" box="[296,319,1961,1984]" italics="true" pageId="7" pageNumber="308">vs</emphasis>
. female dark red, with several white patches on dorsal and ventral sides, a yellow caudal fin and posterior caudal peduncle; unpaired fins predominantly transparent with orange pigments on distal parts and one dark band 
<emphasis id="B9455B11FFD8FA54FEAEFF7EFE85F7D5" box="[269,292,153,176]" italics="true" pageId="8" pageNumber="309">vs</emphasis>
. D1 and D2 of 
<taxonomicName id="4C31FC80FFD8FA54FE70FF7FFDFBF7D5" box="[467,602,152,176]" class="Actinopterygii" family="Gobiidae" genus="Speleogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="trigloides">
<emphasis id="B9455B11FFD8FA54FE70FF7FFDFBF7D5" box="[467,602,152,176]" italics="true" pageId="8" pageNumber="309">S. trigloides</emphasis>
</taxonomicName>
reddish with white bands (D1) or dots (D2); caudal peduncle at caudal fin base with a dark orange vertical band posteriorly fringed by a narrow dark line 
<emphasis id="B9455B11FFD8FA54FB84FF5AFBE7F7B1" box="[1063,1094,189,212]" italics="true" pageId="8" pageNumber="309">vs.</emphasis>
caudal peduncle at caudal fin base uniformly orange-yellow without orange dark vertical stripe.
</paragraph>
</subSubSection>
</treatment>
</document>