414 lines
65 KiB
XML
414 lines
65 KiB
XML
<document id="76CCBBCCB6DF8C01CFA141CDAA0F17D0" ID-CLB-Dataset="31071" ID-DOI="10.11646/zootaxa.4369.3.1" ID-GBIF-Dataset="34826a1d-7146-4d75-9585-b226f798ffdb" ID-ISSN="1175-5326" ID-Zenodo-Dep="1135772" ID-ZooBank="97CAA2FC-F3CC-407B-8768-2BB9DE037C86" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1515148404056" checkinUser="plazi" docAuthor="Mooi, Rich, Martínez, Sergio A., Del Río, Claudia J. & Ramos, Maria Inês Feijó" docDate="2018" docId="0392BB4BFFBBF5497D92F91AFAE4FBB5" docLanguage="en" docName="zootaxa.4369.3.1.pdf" docOrigin="Zootaxa 4369 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Camachoaster maquedensis Mooi & Martínez & Del Río & Ramos 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="316" masterDocId="FFABC333FFB2F5467D05FFF7FFE2FFA5" masterDocTitle="Late Oligocene - Miocene non-lunulate sand dollars of South America: Revision of abertellid taxa and descriptions of two new families, two new genera, and a new species" masterLastPageNumber="326" masterPageNumber="301" pageNumber="310" updateTime="1698512411552" updateUser="ExternalLinkService">
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<mods:title id="BC94BD4928AA0AA78AFF7185FB0F5EA0">Late Oligocene - Miocene non-lunulate sand dollars of South America: Revision of abertellid taxa and descriptions of two new families, two new genera, and a new species</mods:title>
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<mods:namePart id="F4B437D2B658A1743A9590D27A73439F">Mooi, Rich</mods:namePart>
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<mods:namePart id="1ED09D29A7E182B9A08A7604561AC99A">Martínez, Sergio A.</mods:namePart>
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<mods:namePart id="1F712AF666D13139EBA24021A2F9A9C4">Del Río, Claudia J.</mods:namePart>
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<mods:namePart id="A03DC5D3D4818ED3F856A1033C62BC57">Ramos, Maria Inês Feijó</mods:namePart>
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<mods:typeOfResource id="E34FE53A82191092ACC0BD8DC027F98D">text</mods:typeOfResource>
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<mods:title id="8063126B89957F731DDB26A1411498B0">Zootaxa</mods:title>
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<mods:date id="D6242A94EFD4AB1A3A8034CF034C8F26">2018</mods:date>
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<mods:number id="C1FEC79F5A70ED04482E4D47255CA657">2018-01-05</mods:number>
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<treatment id="0392BB4BFFBBF5497D92F91AFAE4FBB5" ID-DOI="http://doi.org/10.5281/zenodo.5961793" ID-GBIF-Taxon="139582394" ID-Zenodo-Dep="5961793" LSID="urn:lsid:plazi:treatment:0392BB4BFFBBF5497D92F91AFAE4FBB5" httpUri="http://treatment.plazi.org/id/0392BB4BFFBBF5497D92F91AFAE4FBB5" lastPageId="15" lastPageNumber="316" pageId="9" pageNumber="310">
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<subSubSection id="C32159D6FFBBF54F7D92F91AFEDDF88C" pageId="9" pageNumber="310" type="nomenclature">
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<paragraph id="8B840A5DFFBBF54F7D92F91AFDD0F8A2" blockId="9.[151,562,1773,1833]" box="[151,562,1773,1799]" pageId="9" pageNumber="310">
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<heading id="D0CCBD31FFBBF54F7D92F91AFDD0F8A2" bold="true" box="[151,562,1773,1799]" fontSize="11" level="1" pageId="9" pageNumber="310" reason="1">
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<emphasis id="B94FD64FFFBBF54F7D92F91AFDD0F8A2" bold="true" box="[151,562,1773,1799]" pageId="9" pageNumber="310">
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<taxonomicName id="4C3B71DEFFBBF54F7D92F91AFE0BF8A2" authority="Mooi & Martínez & Del Río & Ramos, 2018" authorityName="Mooi & Martínez & Del Río & Ramos" authorityYear="2018" box="[151,489,1773,1799]" class="Echinoidea" family="Abertellidae" genus="Camachoaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="9" pageNumber="310" phylum="Echinodermata" rank="species" species="maquedensis" status="sp. nov.">
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<emphasis id="B94FD64FFFBBF54F7D92F91AFE0BF8A2" bold="true" box="[151,489,1773,1799]" italics="true" pageId="9" pageNumber="310">Camachoaster maquedensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27C6B34FFBBF54F7CF5F91AFDD0F8A2" box="[496,562,1773,1799]" pageId="9" pageNumber="310" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B840A5DFFBBF54F7D92F8E7FEDDF88C" blockId="9.[151,562,1773,1833]" box="[151,319,1808,1833]" pageId="9" pageNumber="310">
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<figureCitation id="130016D8FFBBF54F7D92F8E7FEE2F88D" box="[151,256,1808,1833]" captionStart="FIGURE 1" captionStartId="3.[151,250,1776,1798]" captionTargetBox="[162,1426,462,1692]" captionTargetId="figure@3.[151,1436,451,1700]" captionTargetPageId="3" captionText="FIGURE 1. Localities for taxa discussed in the text. Inset at lower left centers on the Santa Cruz region of Patagonia. Numbered occurrences correspond to unequivocal Iheringiella patagonensis localities (inside boxed region in main map): 1. Pampa Castillo, Chile (Andre Wyss, pers. comm.); 2. Cañadón El Lobo, Santa Cruz, Argentina (CPBA 16493-95); 3. Oven Point/Sholl's Tomb, Santa Cruz, Argentina (Ortmann 1902), Punta Cuevas, Santa Cruz, Argentina (MACN 4586 and Cuitiño 2011), Playa La Mina, Santa Cruz, Argentina (Cuitiño 2011); 4. Meseta Chica, Bajo de San Julián, Santa Cruz, Argentina (MACN 5144 and Cuitiño 2011); 5. Upper Rio Chalia, Santa Cruz, Argentina (Ortmann 1902); 6. Puerto Santa Cruz, Santa Cruz, Argentina (ROM 5433M, 5468M, 5469M); 7. Sector Flamenco, Tierra del Fuego, Chile (Larrain 1984)—reported to be from Eocene strata, but this is questionable (Mooi et al. 2000)." httpUri="https://zenodo.org/record/1135774/files/figure.png" pageId="9" pageNumber="310">Figures 1</figureCitation>
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,
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<figureCitation id="130016D8FFBBF54F7C0BF8E7FED9F88C" box="[270,315,1808,1833]" captionStart-0="FIGURE5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="11.[151,250,1912,1934]" captionStartId-1="12.[151,250,1919,1941]" captionStartId-2="13.[151,250,1910,1932]" captionStartId-3="14.[151,250,1883,1905]" captionTargetBox-0="[320,1266,193,1891]" captionTargetBox-1="[301,1285,211,1884]" captionTargetBox-2="[316,1279,208,1872]" captionTargetBox-3="[365,1223,193,1866]" captionTargetId-0="figure@11.[320,1266,193,1891]" captionTargetId-1="figure@12.[301,1285,211,1884]" captionTargetId-2="figure@13.[298,1291,193,1896]" captionTargetId-3="figure@14.[365,1223,193,1866]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="FIGURE5. Camachoaster maquedensisn. sp., photographs of the holotype MACN-Pi 5809.Oral surface shows pencil marks indicating some of the plate architecture.Scale bar 10 mm in length." captionText-1="FIGURE 6. Camachoaster maquedensis n. sp., photographs of the oral surfaces of paratypes. A: MACN-Pi 5859, prepared specimen with pencil marks indicating plate architecture. B: MACN-Pi 5860, partially prepared specimen with pencil marks indicating plate architecture, faint tuberculation, and main branches of food grooves. Scale bars 10 mm in length." captionText-2="FIGURE 7. Comparison of oral plate architecture of Camachoaster maquedensis n. sp. (based on paratypes MACN-Pi 5859 and 5860) and Camachoaster? kewi (= Abertella kewi Durham, 1957) (based on specimens from UCMP locality B8562, Chiapas, Mexico).Food grooves indicated by dotted lines, peristome and periproctin solid black, interambulacra shaded, scale bars 10 mm in length." captionText-3="FIGURE 8. Camachoaster maquedensis n. sp., details of test features. A: Holotype MACN-Pi 5809, pencil marks indicating partialaboral plate architecture in regionwithin andaround petal in ambulacrum II, gonoporesindicated by white arrows, pores of trailing tube feet by black arrows. B: Paratype MACN-Pi 5859, complete aboral ambulacrum IV showing shape of petal. C: Holotype MACN-Pi 5809, details of tuberculation, food groove (black arrow), and peristome (peristomial projection indicated by white arrow).All scale bars 10 mm in length." httpUri-0="https://zenodo.org/record/1135782/files/figure.png" httpUri-1="https://zenodo.org/record/1135784/files/figure.png" httpUri-2="https://zenodo.org/record/1135786/files/figure.png" httpUri-3="https://zenodo.org/record/1135788/files/figure.png" pageId="9" pageNumber="310">5–8</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32159D6FFBBF54F7D92F8AFFE3AF8D4" box="[151,472,1880,1905]" pageId="9" pageNumber="310" type="diagnosis">
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<paragraph id="8B840A5DFFBBF54F7D92F8AFFE3AF8D4" blockId="9.[151,1437,1880,2013]" box="[151,472,1880,1905]" pageId="9" pageNumber="310">
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<emphasis id="B94FD64FFFBBF54F7D92F8AFFEF3F8D4" bold="true" box="[151,273,1880,1905]" pageId="9" pageNumber="310">Diagnosis.</emphasis>
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As for the genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32159D6FFBBF54C7DC2F88BFA6DFF5D" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310" type="etymology">
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<paragraph id="8B840A5DFFBBF54C7DC2F88BFA6DFF5D" blockId="9.[151,1437,1880,2013]" lastBlockId="10.[151,1437,151,2013]" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310">
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<emphasis id="B94FD64FFFBBF54F7DC2F88BFEACF830" bold="true" box="[199,334,1916,1941]" pageId="9" pageNumber="310">Etymology.</emphasis>
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Named for the
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<typeStatus id="5480B4FFFFBBF54F7F06F88AFDD1F830" box="[515,563,1917,1941]" pageId="9" pageNumber="310">type</typeStatus>
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locality of Punta Maqueda,
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<collectingRegion id="49FFC4BFFFBBF54F7E7CF88BFB84F830" box="[889,1126,1916,1941]" country="Argentina" name="Santa Cruz" pageId="9" pageNumber="310">Santa Cruz Province</collectingRegion>
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,
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<collectingCountry id="F32C4ACDFFBBF54F7974F88BFB06F830" box="[1137,1252,1916,1941]" name="Argentina" pageId="9" pageNumber="310">Argentina</collectingCountry>
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, where the sand dollars occur in the Chenque Formation, which is exposed about
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<quantity id="4CC3A7B8FFBBF54F7E75F857FC4BF81C" box="[880,937,1952,1977]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="310" unit="km" value="2.0">2 km</quantity>
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south of the point.
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<emphasis id="B94FD64FFFBBF54F7DC2F833FE00F878" bold="true" box="[199,482,1988,2013]" pageId="9" pageNumber="310">
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<typeStatus id="5480B4FFFFBBF54F7DC2F833FEE0F878" box="[199,258,1988,2013]" pageId="9" pageNumber="310">Type</typeStatus>
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material studied.
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</emphasis>
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Material is housed at the Museo Argentino de Ciencias Naturales Bernardino Rivadavia (MACN).
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<typeStatus id="5480B4FFFFB8F54C7C8CFF60FE13FF15" box="[393,497,151,176]" pageId="10" pageNumber="311" type="holotype">Holotype</typeStatus>
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is MACN-Pi 5809, from shoreline deposits about
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<quantity id="4CC3A7B8FFB8F54C793CFF6FFB96FF15" box="[1081,1140,151,176]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="311" unit="km" value="2.0">2 km</quantity>
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south of Punta Maqueda,
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<collectingRegion id="49FFC4BFFFB8F54C7D92FF4BFE65FF70" box="[151,391,188,213]" country="Argentina" name="Santa Cruz" pageId="10" pageNumber="311">Santa Cruz Province</collectingRegion>
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, southern
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<collectingCountry id="F32C4ACDFFB8F54C7CFAFF4BFD90FF70" box="[511,626,188,213]" name="Argentina" pageId="10" pageNumber="311">Argentina</collectingCountry>
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. These deposits form part of the Chenque Formation, lower Miocene. There are two
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<typeStatus id="5480B4FFFFB8F54C7C3CFF17FE45FF5D" box="[313,423,224,248]" pageId="10" pageNumber="311" type="paratype">paratypes</typeStatus>
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, MACN-Pi 5859 and MACN-Pi 5860. These have the same provenience as the
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<typeStatus id="5480B4FFFFB8F54C7820FF28FA69FF5D" box="[1317,1419,223,248]" pageId="10" pageNumber="311" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32159D6FFB8F54C7DC2FEF3FAF0F878" pageId="10" pageNumber="311" type="description">
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<paragraph id="8B840A5DFFB8F54C7DC2FEF3FE87FE75" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="311">
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<emphasis id="B94FD64FFFB8F54C7DC2FEF3FEBBFEB8" bold="true" box="[199,345,260,285]" pageId="10" pageNumber="311">Description.</emphasis>
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<typeStatus id="5480B4FFFFB8F54C7C67FEF3FE28FEB8" box="[354,458,260,285]" pageId="10" pageNumber="311" type="holotype">Holotype</typeStatus>
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and largest specimen (
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<figureCitation id="130016D8FFB8F54C7FD2FEF3FCFEFEB8" box="[727,796,260,285]" captionStart="FIGURE5" captionStartId="11.[151,250,1912,1934]" captionTargetBox="[320,1266,193,1891]" captionTargetId="figure@11.[320,1266,193,1891]" captionTargetPageId="11" captionText="FIGURE5. Camachoaster maquedensisn. sp., photographs of the holotype MACN-Pi 5809.Oral surface shows pencil marks indicating some of the plate architecture.Scale bar 10 mm in length." httpUri="https://zenodo.org/record/1135782/files/figure.png" pageId="10" pageNumber="311">Fig. 5</figureCitation>
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) 59.3 mm TL. Measurements for all known specimens given in
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<tableCitation id="C6B93FE6FFB8F54C7DF9FED0FEB3FEE5" box="[252,337,295,320]" captionStart="TABLE 1" captionStartId="19.[151,239,152,175]" captionText="TABLE 1. Camachoaster maquedensis n. gen., n. sp.: measurements (in mm) of holotype and both paratypes from the Chenque Formation, Punta Maqueda, Santa Cruz Province, Argentina." pageId="10" pageNumber="311">Table 1</tableCitation>
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. All ensuing percentages, calculated to facilitate comparisons with other descriptions herein, are from
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<typeStatus id="5480B4FFFFB8F54C7DD2FEBBFEDFFEC0" box="[215,317,332,357]" pageId="10" pageNumber="311" type="holotype">holotype</typeStatus>
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. Aboral surface slightly domed, oral surface flat. Highest point of test 11% TL, located at apical system. Very shallow but distinct posterior notch, depth just over 5% TL, width 16% TL, notch widening significantly near ambitus. Broad, shallow marginal indentations present where perradial suture meets ambitus in each ambulacrum.
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</paragraph>
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<paragraph id="8B840A5DFFB8F54C7DC2FE2BFEE4FDC5" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="311">
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Apical system monobasal, pentagonal (not star-shaped), 49% TL from ocular III to anterior edge of test, length 8% TL, numerous hydropores scattered over madreporic plate. Four gonopores, one in each of paired interambulacra and located at suture between madreporic plate and first adapical plates of interambulacral column (
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<figureCitation id="130016D8FFB8F54C7D9AFDB0FF1BFDC5" box="[159,249,583,608]" captionStart="FIGURE 8" captionStartId="14.[151,250,1883,1905]" captionTargetBox="[365,1223,193,1866]" captionTargetId="figure@14.[365,1223,193,1866]" captionTargetPageId="14" captionText="FIGURE 8. Camachoaster maquedensis n. sp., details of test features. A: Holotype MACN-Pi 5809, pencil marks indicating partialaboral plate architecture in regionwithin andaround petal in ambulacrum II, gonoporesindicated by white arrows, pores of trailing tube feet by black arrows. B: Paratype MACN-Pi 5859, complete aboral ambulacrum IV showing shape of petal. C: Holotype MACN-Pi 5809, details of tuberculation, food groove (black arrow), and peristome (peristomial projection indicated by white arrow).All scale bars 10 mm in length." httpUri="https://zenodo.org/record/1135788/files/figure.png" pageId="10" pageNumber="311">Fig. 8A</figureCitation>
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).
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</paragraph>
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<paragraph id="8B840A5DFFB8F54C7DC2FD9BFE48FB91" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="311">
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Ambulacra petaloid adapically. Posterior paired petals (I and V) not noticeably longer than any other petals, each extending 56% of corresponding test radius, but 30% TL; anterior paired petals (II and IV) 51% of corresponding test radius, but 29% TL; anterior unpaired (III) 56% of corresponding test radius, but 29% TL. Petal V width at widest point 12% TL, interporiferous zone 4% TL; petal IV width 12% TL, interporiferous zone 5% TL; petal III width 13% TL, interporiferous zone 6% TL. Petals lyrate, almost closed distally (
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<figureCitation id="130016D8FFB8F54C7980FD0BFB31FCB0" box="[1157,1235,764,789]" captionStart="FIGURE5" captionStartId="11.[151,250,1912,1934]" captionTargetBox="[320,1266,193,1891]" captionTargetId="figure@11.[320,1266,193,1891]" captionTargetPageId="11" captionText="FIGURE5. Camachoaster maquedensisn. sp., photographs of the holotype MACN-Pi 5809.Oral surface shows pencil marks indicating some of the plate architecture.Scale bar 10 mm in length." httpUri="https://zenodo.org/record/1135782/files/figure.png" pageId="10" pageNumber="311">Figs. 5</figureCitation>
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,
|
||
<figureCitation id="130016D8FFB8F54C79E5FD0BFAC7FCB1" box="[1248,1317,764,788]" captionStart="FIGURE 8" captionStartId="14.[151,250,1883,1905]" captionTargetBox="[365,1223,193,1866]" captionTargetId="figure@14.[365,1223,193,1866]" captionTargetPageId="14" captionText="FIGURE 8. Camachoaster maquedensis n. sp., details of test features. A: Holotype MACN-Pi 5809, pencil marks indicating partialaboral plate architecture in regionwithin andaround petal in ambulacrum II, gonoporesindicated by white arrows, pores of trailing tube feet by black arrows. B: Paratype MACN-Pi 5859, complete aboral ambulacrum IV showing shape of petal. C: Holotype MACN-Pi 5809, details of tuberculation, food groove (black arrow), and peristome (peristomial projection indicated by white arrow).All scale bars 10 mm in length." httpUri="https://zenodo.org/record/1135788/files/figure.png" pageId="10" pageNumber="311">8A, B</figureCitation>
|
||
), with two or perhaps as many as three trailing tube feet (
|
||
<emphasis id="B94FD64FFFB8F54C7FAFFCD6FD0AFC9D" box="[682,744,801,824]" italics="true" pageId="10" pageNumber="311">sensu</emphasis>
|
||
<bibRefCitation id="EFAA77ACFFB8F54C7FF4FCE8FC8DFC9D" author="Mooi" box="[753,879,799,824]" pageId="10" pageNumber="311" refString="Mooi, R. (1989) Living and fossil genera of the Clypeasteroida (Echinoidea: Echinodermata): An illustrated key and annotated checklist. Smithsonian Contributions to Zoology, 488, 1 - 51. https: // doi. org / 10.5479 / si. 00810282.488" type="journal article" year="1989">Mooi 1989</bibRefCitation>
|
||
) at distal end of each column of respiratory tube feet (
|
||
<figureCitation id="130016D8FFB8F54C7DD0FCB3FED1FCF9" box="[213,307,836,861]" captionStart="FIGURE 8" captionStartId="14.[151,250,1883,1905]" captionTargetBox="[365,1223,193,1866]" captionTargetId="figure@14.[365,1223,193,1866]" captionTargetPageId="14" captionText="FIGURE 8. Camachoaster maquedensis n. sp., details of test features. A: Holotype MACN-Pi 5809, pencil marks indicating partialaboral plate architecture in regionwithin andaround petal in ambulacrum II, gonoporesindicated by white arrows, pores of trailing tube feet by black arrows. B: Paratype MACN-Pi 5859, complete aboral ambulacrum IV showing shape of petal. C: Holotype MACN-Pi 5809, details of tuberculation, food groove (black arrow), and peristome (peristomial projection indicated by white arrow).All scale bars 10 mm in length." httpUri="https://zenodo.org/record/1135788/files/figure.png" pageId="10" pageNumber="311">Fig. 8A</figureCitation>
|
||
). Respiratory tube foot pore pairs strongly conjugated, inner pore slightly elongated, outer pore extremely elongated, comprising about half length of pore pair, apparently subdivided by stereom septae (
|
||
<figureCitation id="130016D8FFB8F54C783BFC90FF4EFC01" captionStart="FIGURE 8" captionStartId="14.[151,250,1883,1905]" captionTargetBox="[365,1223,193,1866]" captionTargetId="figure@14.[365,1223,193,1866]" captionTargetPageId="14" captionText="FIGURE 8. Camachoaster maquedensis n. sp., details of test features. A: Holotype MACN-Pi 5809, pencil marks indicating partialaboral plate architecture in regionwithin andaround petal in ambulacrum II, gonoporesindicated by white arrows, pores of trailing tube feet by black arrows. B: Paratype MACN-Pi 5859, complete aboral ambulacrum IV showing shape of petal. C: Holotype MACN-Pi 5809, details of tuberculation, food groove (black arrow), and peristome (peristomial projection indicated by white arrow).All scale bars 10 mm in length." httpUri="https://zenodo.org/record/1135788/files/figure.png" pageId="10" pageNumber="311">Fig. 8A, B</figureCitation>
|
||
). Five or six occluded plates present at tips of petals (
|
||
<figureCitation id="130016D8FFB8F54C7E0FFC7BFC86FC01" box="[778,868,908,933]" captionStart="FIGURE 8" captionStartId="14.[151,250,1883,1905]" captionTargetBox="[365,1223,193,1866]" captionTargetId="figure@14.[365,1223,193,1866]" captionTargetPageId="14" captionText="FIGURE 8. Camachoaster maquedensis n. sp., details of test features. A: Holotype MACN-Pi 5809, pencil marks indicating partialaboral plate architecture in regionwithin andaround petal in ambulacrum II, gonoporesindicated by white arrows, pores of trailing tube feet by black arrows. B: Paratype MACN-Pi 5859, complete aboral ambulacrum IV showing shape of petal. C: Holotype MACN-Pi 5809, details of tuberculation, food groove (black arrow), and peristome (peristomial projection indicated by white arrow).All scale bars 10 mm in length." httpUri="https://zenodo.org/record/1135788/files/figure.png" pageId="10" pageNumber="311">Fig. 8A</figureCitation>
|
||
). At ambitus, ambulacra greatly widened, forming strip-like ambital plates that follow contour of shallow indentation at each perradius, and curving adapically to form test wall along each side of posterior notch (
|
||
<figureCitation id="130016D8FFB8F54C7FD0FC23FCF9FC49" box="[725,795,980,1005]" captionStart="FIGURE 7" captionStartId="13.[151,250,1910,1932]" captionTargetBox="[316,1279,208,1872]" captionTargetId="figure@13.[298,1291,193,1896]" captionTargetPageId="13" captionText="FIGURE 7. Comparison of oral plate architecture of Camachoaster maquedensis n. sp. (based on paratypes MACN-Pi 5859 and 5860) and Camachoaster? kewi (= Abertella kewi Durham, 1957) (based on specimens from UCMP locality B8562, Chiapas, Mexico).Food grooves indicated by dotted lines, peristome and periproctin solid black, interambulacra shaded, scale bars 10 mm in length." httpUri="https://zenodo.org/record/1135786/files/figure.png" pageId="10" pageNumber="311">Fig. 7</figureCitation>
|
||
). Ambulacra all in agreement with Lovén’s Rule (
|
||
<emphasis id="B94FD64FFFB8F54C785BFC22FA7EFC49" box="[1374,1436,981,1004]" italics="true" pageId="10" pageNumber="311">sensu</emphasis>
|
||
<bibRefCitation id="EFAA77ACFFB8F54C7D92FC00FEB9FBB5" author="David" box="[151,347,1015,1040]" pageId="10" pageNumber="311" refString="David, B., Mooi, R. & Telford, M. (1996) The ontogenetic basis of Loven's Rule clarifies homologies of the echinoid peristome. In: Emson, R., Smith, A. B. & Campbell, A. (Eds.), Echinoderm Research 1995. A. A. Balkema, Rotterdam, pp. 155 - 164." type="journal article" year="1996">
|
||
David
|
||
<emphasis id="B94FD64FFFB8F54C7DE7FC0EFEF0FBB5" box="[226,274,1016,1040]" italics="true" pageId="10" pageNumber="311">et al</emphasis>
|
||
. 1996
|
||
</bibRefCitation>
|
||
). Ambulacral basicoronal plates all similar, narrow and straight with almost parallel radial sutures on each side (
|
||
<figureCitation id="130016D8FFB8F54C7C34FBEBFE62FB90" box="[305,384,1052,1077]" captionStart="FIGURE 6" captionStartId="12.[151,250,1919,1941]" captionTargetBox="[301,1285,211,1884]" captionTargetId="figure@12.[301,1285,211,1884]" captionTargetPageId="12" captionText="FIGURE 6. Camachoaster maquedensis n. sp., photographs of the oral surfaces of paratypes. A: MACN-Pi 5859, prepared specimen with pencil marks indicating plate architecture. B: MACN-Pi 5860, partially prepared specimen with pencil marks indicating plate architecture, faint tuberculation, and main branches of food grooves. Scale bars 10 mm in length." httpUri="https://zenodo.org/record/1135784/files/figure.png" pageId="10" pageNumber="311">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="130016D8FFB8F54C7C89FBEBFE7EFB91" box="[396,412,1052,1076]" captionStart="FIGURE 7" captionStartId="13.[151,250,1910,1932]" captionTargetBox="[316,1279,208,1872]" captionTargetId="figure@13.[298,1291,193,1896]" captionTargetPageId="13" captionText="FIGURE 7. Comparison of oral plate architecture of Camachoaster maquedensis n. sp. (based on paratypes MACN-Pi 5859 and 5860) and Camachoaster? kewi (= Abertella kewi Durham, 1957) (based on specimens from UCMP locality B8562, Chiapas, Mexico).Food grooves indicated by dotted lines, peristome and periproctin solid black, interambulacra shaded, scale bars 10 mm in length." httpUri="https://zenodo.org/record/1135786/files/figure.png" pageId="10" pageNumber="311">7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B840A5DFFB8F54C7DC2FBC8FDBCFADD" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="311">
|
||
Interambulacra narrow and straight on oral surface, narrowing towards ambitus, but containing paired, zig-zag plates right up to madreporic plate. On oral surface, three or four postbasicoronal plates in each halfinterambulacrum in interambulacrum 5, four or five in in other interambulacra. Widest point of each interambulacrum about one third of way from basicoronal to ambitus, narrowing distally so that paired interambulacra about 22% width of adjacent ambulacra at ambitus. In each paired interambulacrum, first postbasicoronal greatly elongated, about four times as long as wide, about twice length of corresponding second postbasicoronal. In interambulacra 1 to 4, basicoronals broadly in contact with both corresponding first postbasicoronals. Unpaired, posterior interambulacrum 5 discontinuous, separated from basicoronal by adjacent ambulacral postbasicoronals (
|
||
<figureCitation id="130016D8FFB8F54C7CE1FAA8FDD1FADD" box="[484,563,1375,1400]" captionStart="FIGURE 6" captionStartId="12.[151,250,1919,1941]" captionTargetBox="[301,1285,211,1884]" captionTargetId="figure@12.[301,1285,211,1884]" captionTargetPageId="12" captionText="FIGURE 6. Camachoaster maquedensis n. sp., photographs of the oral surfaces of paratypes. A: MACN-Pi 5859, prepared specimen with pencil marks indicating plate architecture. B: MACN-Pi 5860, partially prepared specimen with pencil marks indicating plate architecture, faint tuberculation, and main branches of food grooves. Scale bars 10 mm in length." httpUri="https://zenodo.org/record/1135784/files/figure.png" pageId="10" pageNumber="311">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="130016D8FFB8F54C7F3AFA97FDADFADD" box="[575,591,1376,1400]" captionStart="FIGURE 7" captionStartId="13.[151,250,1910,1932]" captionTargetBox="[316,1279,208,1872]" captionTargetId="figure@13.[298,1291,193,1896]" captionTargetPageId="13" captionText="FIGURE 7. Comparison of oral plate architecture of Camachoaster maquedensis n. sp. (based on paratypes MACN-Pi 5859 and 5860) and Camachoaster? kewi (= Abertella kewi Durham, 1957) (based on specimens from UCMP locality B8562, Chiapas, Mexico).Food grooves indicated by dotted lines, peristome and periproctin solid black, interambulacra shaded, scale bars 10 mm in length." httpUri="https://zenodo.org/record/1135786/files/figure.png" pageId="10" pageNumber="311">7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B840A5DFFB8F54C7DC2FA73FEBBF9AD" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="311">
|
||
Peristome circular, relatively small, about 5% TL, with distinct perradial process in each ambulacrum extending into peristome beyond slight bulge containing sphaeridium (
|
||
<figureCitation id="130016D8FFB8F54C7EB8FA50FBF5FA65" box="[957,1047,1447,1472]" captionStart="FIGURE 8" captionStartId="14.[151,250,1883,1905]" captionTargetBox="[365,1223,193,1866]" captionTargetId="figure@14.[365,1223,193,1866]" captionTargetPageId="14" captionText="FIGURE 8. Camachoaster maquedensis n. sp., details of test features. A: Holotype MACN-Pi 5809, pencil marks indicating partialaboral plate architecture in regionwithin andaround petal in ambulacrum II, gonoporesindicated by white arrows, pores of trailing tube feet by black arrows. B: Paratype MACN-Pi 5859, complete aboral ambulacrum IV showing shape of petal. C: Holotype MACN-Pi 5809, details of tuberculation, food groove (black arrow), and peristome (peristomial projection indicated by white arrow).All scale bars 10 mm in length." httpUri="https://zenodo.org/record/1135788/files/figure.png" pageId="10" pageNumber="311">Fig. 8C</figureCitation>
|
||
). Anterior edge of peristome 51% TL from anterior edge of test. Periproct small, about 3% TL, on oral surface between second and third pair of postbasicoronals.
|
||
</paragraph>
|
||
<paragraph id="8B840A5DFFB8F54C7DC2F9E3FDB3F9D1" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="311">
|
||
Aboral tuberculation homogeneous. Very slight enlargement of tubercles in oral interambulacral regions relative to those in ambulacral regions. In specimens with best preservation of surface detail, distinct tube foot pores visible in food grooves (
|
||
<figureCitation id="130016D8FFB8F54C7CEEF9ABFDA1F9D1" box="[491,579,1628,1653]" captionStart="FIGURE 8" captionStartId="14.[151,250,1883,1905]" captionTargetBox="[365,1223,193,1866]" captionTargetId="figure@14.[365,1223,193,1866]" captionTargetPageId="14" captionText="FIGURE 8. Camachoaster maquedensis n. sp., details of test features. A: Holotype MACN-Pi 5809, pencil marks indicating partialaboral plate architecture in regionwithin andaround petal in ambulacrum II, gonoporesindicated by white arrows, pores of trailing tube feet by black arrows. B: Paratype MACN-Pi 5859, complete aboral ambulacrum IV showing shape of petal. C: Holotype MACN-Pi 5809, details of tuberculation, food groove (black arrow), and peristome (peristomial projection indicated by white arrow).All scale bars 10 mm in length." httpUri="https://zenodo.org/record/1135788/files/figure.png" pageId="10" pageNumber="311">Fig. 8C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B840A5DFFB8F54C7DC2F988FE79F8A0" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="311">
|
||
Food grooves well developed (
|
||
<figureCitation id="130016D8FFB8F54C7F24F988FD92F93D" box="[545,624,1663,1688]" captionStart="FIGURE5" captionStartId="11.[151,250,1912,1934]" captionTargetBox="[320,1266,193,1891]" captionTargetId="figure@11.[320,1266,193,1891]" captionTargetPageId="11" captionText="FIGURE5. Camachoaster maquedensisn. sp., photographs of the holotype MACN-Pi 5809.Oral surface shows pencil marks indicating some of the plate architecture.Scale bar 10 mm in length." httpUri="https://zenodo.org/record/1135782/files/figure.png" pageId="10" pageNumber="311">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="130016D8FFB8F54C7F79F976FD6EF93D" box="[636,652,1665,1688]" captionStart="FIGURE 7" captionStartId="13.[151,250,1910,1932]" captionTargetBox="[316,1279,208,1872]" captionTargetId="figure@13.[298,1291,193,1896]" captionTargetPageId="13" captionText="FIGURE 7. Comparison of oral plate architecture of Camachoaster maquedensis n. sp. (based on paratypes MACN-Pi 5859 and 5860) and Camachoaster? kewi (= Abertella kewi Durham, 1957) (based on specimens from UCMP locality B8562, Chiapas, Mexico).Food grooves indicated by dotted lines, peristome and periproctin solid black, interambulacra shaded, scale bars 10 mm in length." httpUri="https://zenodo.org/record/1135786/files/figure.png" pageId="10" pageNumber="311">7</figureCitation>
|
||
,
|
||
<figureCitation id="130016D8FFB8F54C7F9DF977FD59F93D" box="[664,699,1664,1688]" captionStart="FIGURE 8" captionStartId="14.[151,250,1883,1905]" captionTargetBox="[365,1223,193,1866]" captionTargetId="figure@14.[365,1223,193,1866]" captionTargetPageId="14" captionText="FIGURE 8. Camachoaster maquedensis n. sp., details of test features. A: Holotype MACN-Pi 5809, pencil marks indicating partialaboral plate architecture in regionwithin andaround petal in ambulacrum II, gonoporesindicated by white arrows, pores of trailing tube feet by black arrows. B: Paratype MACN-Pi 5859, complete aboral ambulacrum IV showing shape of petal. C: Holotype MACN-Pi 5809, details of tuberculation, food groove (black arrow), and peristome (peristomial projection indicated by white arrow).All scale bars 10 mm in length." httpUri="https://zenodo.org/record/1135788/files/figure.png" pageId="10" pageNumber="311">8C</figureCitation>
|
||
), restricted to oral surface, with primary bifurcation near adapical ends of ambulacral basicoronal plates. After this branch point, food grooves continuously diverging as they approach ambitus. Secondary branching well developed (
|
||
<figureCitation id="130016D8FFB8F54C7E2DF930FC8DF945" box="[808,879,1735,1760]" captionStart="FIGURE5" captionStartId="11.[151,250,1912,1934]" captionTargetBox="[320,1266,193,1891]" captionTargetId="figure@11.[320,1266,193,1891]" captionTargetPageId="11" captionText="FIGURE5. Camachoaster maquedensisn. sp., photographs of the holotype MACN-Pi 5809.Oral surface shows pencil marks indicating some of the plate architecture.Scale bar 10 mm in length." httpUri="https://zenodo.org/record/1135782/files/figure.png" pageId="10" pageNumber="311">Fig. 5</figureCitation>
|
||
). Extremely shallow depressions along perradial sutures on oral surface.
|
||
</paragraph>
|
||
<paragraph id="8B840A5DFFB8F54C7DC2F8F8FD08F88D" blockId="10.[151,1437,151,2013]" box="[199,746,1807,1833]" pageId="10" pageNumber="311">
|
||
<emphasis id="B94FD64FFFB8F54C7DC2F8F8FEBBF88D" bold="true" box="[199,345,1807,1832]" pageId="10" pageNumber="311">Occurrence.</emphasis>
|
||
Known only from the type locality.
|
||
</paragraph>
|
||
<paragraph id="8B840A5DFFB8F54C7DC2F8C3FAF0F878" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="311">
|
||
<emphasis id="B94FD64FFFB8F54C7DC2F8C3FE6DF8E8" bold="true" box="[199,399,1844,1869]" pageId="10" pageNumber="311">Geologic setting.</emphasis>
|
||
The shallow marine deposits in which the new species was found have been described in
|
||
<bibRefCitation id="EFAA77ACFFB8F54C7D92F8A0FE8CF8D5" box="[151,366,1879,1904]" pageId="10" pageNumber="311" refString="Mooi, R., Martinez, S. & del Rio, C. J. (2016) A new South American Miocene species of ' one-holed' sand dollar (Echinoidea: Clypeasteroida: Monophorasteridae). Zootaxa, 4173 (1), 45 - 54. https: // doi. org / 10.11646 / zootaxa. 4173.1.4" type="journal article">
|
||
Mooi
|
||
<emphasis id="B94FD64FFFB8F54C7DD9F8AEFEF5F8D5" box="[220,279,1880,1904]" italics="true" pageId="10" pageNumber="311">et al.</emphasis>
|
||
(2016)
|
||
</bibRefCitation>
|
||
. In summary, the rocks of the Chenque Formation in the Golfo San Jorge Basin (Chubut and Santa Cruz Provinces, Argentina) exposed near Punta Maqueda have been described as early Miocene (del Río 2004). The type specimens of
|
||
<taxonomicName id="4C3B71DEFFB8F54C7CEEF857FC59F81C" authority="Mooi et al., 2016" authorityName="Mooi et al." authorityYear="2016" box="[491,955,1952,1977]" class="Echinoidea" family="Monophorasteridae" genus="Monophoraster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="10" pageNumber="311" phylum="Echinodermata" rank="species" species="telfordi">
|
||
<emphasis id="B94FD64FFFB8F54C7CEEF857FD13F81D" box="[491,753,1952,1976]" italics="true" pageId="10" pageNumber="311">Monophoraster telfordi</emphasis>
|
||
<bibRefCitation id="EFAA77ACFFB8F54C7FFCF857FC59F81C" author="Mooi" box="[761,955,1952,1977]" pageId="10" pageNumber="311" refString="Mooi, R., Martinez, S. & del Rio, C. J. (2016) A new South American Miocene species of ' one-holed' sand dollar (Echinoidea: Clypeasteroida: Monophorasteridae). Zootaxa, 4173 (1), 45 - 54. https: // doi. org / 10.11646 / zootaxa. 4173.1.4" type="journal article" year="2016">
|
||
Mooi
|
||
<emphasis id="B94FD64FFFB8F54C7E38F856FC97F81D" box="[829,885,1952,1976]" italics="true" pageId="10" pageNumber="311">et al.</emphasis>
|
||
, 2016
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
were collected from the same thin bed (up to
|
||
<quantity id="4CC3A7B8FFB8F54C7DB1F833FF1BF879" box="[180,249,1988,2013]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="311" unit="cm" value="15.0">15 cm</quantity>
|
||
thick) of fine-grained sandstones from which the new species of non-lunulate was recovered.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF445AD5FFB9F54D7D92F88FFCAEF809" httpUri="https://zenodo.org/record/1135782/files/figure.png" pageId="11" pageNumber="312" targetBox="[320,1266,193,1891]" targetPageId="11">
|
||
<paragraph id="8B840A5DFFB9F54D7D92F88FFCAEF809" blockId="11.[151,1436,1912,1964]" pageId="11" pageNumber="312">
|
||
<emphasis id="B94FD64FFFB9F54D7D92F88FFEF0F828" bold="true" box="[151,274,1912,1934]" pageId="11" pageNumber="312">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C3B71DEFFB9F54D7C1CF88FFDD4F82B" authority="Mooi & Martínez & Del Río & Ramos, 2018" authorityName="Mooi & Martínez & Del Río & Ramos" authorityYear="2018" box="[281,566,1912,1934]" class="Echinoidea" family="Abertellidae" genus="Camachoaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="11" pageNumber="312" phylum="Echinodermata" rank="species" species="maquedensis" status="sp. nov.">
|
||
<emphasis id="B94FD64FFFB9F54D7C1CF88FFDD4F82B" box="[281,566,1912,1934]" italics="true" pageId="11" pageNumber="312">Camachoaster maquedensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C6B34FFB9F54D7F3EF88FFD96F828" box="[571,628,1912,1933]" pageId="11" pageNumber="312" rank="species">
|
||
<emphasis id="B94FD64FFFB9F54D7F3EF88FFD96F828" bold="true" box="[571,628,1912,1933]" pageId="11" pageNumber="312">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, photographs of the holotype MACN-Pi 5809. Oral surface shows pencil marks indicating some of the plate architecture. Scale bar 10 mm in length.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF445AD5FFBEF54A7D92F888FAF3F877" httpUri="https://zenodo.org/record/1135784/files/figure.png" pageId="12" pageNumber="313" targetBox="[301,1285,211,1884]" targetPageId="12">
|
||
<paragraph id="8B840A5DFFBEF54A7D92F888FAF3F877" blockId="12.[151,1436,1919,2002]" pageId="12" pageNumber="313">
|
||
<emphasis id="B94FD64FFFBEF54A7D92F888FEF6F831" bold="true" box="[151,276,1919,1941]" pageId="12" pageNumber="313">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C3B71DEFFBEF54A7C18F888FDDEF830" authority="Mooi & Martínez & Del Río & Ramos, 2018" authorityName="Mooi & Martínez & Del Río & Ramos" authorityYear="2018" box="[285,572,1919,1941]" class="Echinoidea" family="Abertellidae" genus="Camachoaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="12" pageNumber="313" phylum="Echinodermata" rank="species" species="maquedensis" status="sp. nov.">
|
||
<emphasis id="B94FD64FFFBEF54A7C18F888FDDEF830" box="[285,572,1919,1941]" italics="true" pageId="12" pageNumber="313">Camachoaster maquedensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C6B34FFBEF54A7F41F888FD9DF831" box="[580,639,1919,1940]" pageId="12" pageNumber="313" rank="species">
|
||
<emphasis id="B94FD64FFFBEF54A7F41F888FD9DF831" bold="true" box="[580,639,1919,1940]" pageId="12" pageNumber="313">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, photographs of the oral surfaces of paratypes. A: MACN-Pi 5859, prepared specimen with pencil marks indicating plate architecture. B: MACN-Pi 5860, partially prepared specimen with pencil marks indicating plate architecture, faint tuberculation, and main branches of food grooves. Scale bars 10 mm in length.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF445AD5FFBFF54B7D92F881FE97F84D" httpUri="https://zenodo.org/record/1135786/files/figure.png" pageId="13" pageNumber="314" targetBox="[316,1279,208,1872]" targetPageId="13">
|
||
<paragraph id="8B840A5DFFBFF54B7D92F881FE97F84D" blockId="13.[151,1436,1910,2024]" pageId="13" pageNumber="314">
|
||
<emphasis id="B94FD64FFFBFF54B7D92F881FEF6F829" bold="true" box="[151,276,1910,1932]" pageId="13" pageNumber="314">FIGURE 7.</emphasis>
|
||
Comparison of oral plate architecture of
|
||
<taxonomicName id="4C3B71DEFFBFF54B7FBBF881FC3EF829" authority="Mooi & Martínez & Del Río & Ramos, 2018" authorityName="Mooi & Martínez & Del Río & Ramos" authorityYear="2018" box="[702,988,1910,1932]" class="Echinoidea" family="Abertellidae" genus="Camachoaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="13" pageNumber="314" phylum="Echinodermata" rank="species" species="maquedensis" status="sp. nov.">
|
||
<emphasis id="B94FD64FFFBFF54B7FBBF881FC3EF829" box="[702,988,1910,1932]" italics="true" pageId="13" pageNumber="314">Camachoaster maquedensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C6B34FFBFF54B7EE1F880FBFCF829" box="[996,1054,1911,1932]" pageId="13" pageNumber="314" rank="species">
|
||
<emphasis id="B94FD64FFFBFF54B7EE1F880FBFCF829" bold="true" box="[996,1054,1911,1932]" pageId="13" pageNumber="314">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(based on paratypes MACN-Pi 5859 and 5860) and
|
||
<taxonomicName id="4C3B71DEFFBFF54B7C3CF862FE2FF80E" authority="Mooi & Martínez & Del Río & Ramos, 2018" authorityName="Mooi & Martínez & Del Río & Ramos" authorityYear="2018" box="[313,461,1941,1963]" class="Echinoidea" family="Abertellidae" genus="Camachoaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="13" pageNumber="310" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94FD64FFFBFF54B7C3CF862FE2FF80E" box="[313,461,1941,1963]" italics="true" pageId="13" pageNumber="314">Camachoaster</emphasis>
|
||
</taxonomicName>
|
||
?
|
||
<taxonomicName id="4C3B71DEFFBFF54B7CEEF862FDFBF80E" authorityName="Durham" authorityYear="1957" box="[491,537,1941,1963]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="13" pageNumber="314" phylum="Echinodermata" rank="species" species="kewi">
|
||
<emphasis id="B94FD64FFFBFF54B7CEEF862FDFBF80E" box="[491,537,1941,1963]" italics="true" pageId="13" pageNumber="314">kewi</emphasis>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="4C3B71DEFFBFF54B7F41F862FC9EF80E" authority="Durham, 1957" authorityName="Durham" authorityYear="1957" box="[580,892,1941,1963]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="13" pageNumber="314" phylum="Echinodermata" rank="species" species="kewi">
|
||
<emphasis id="B94FD64FFFBFF54B7F41F862FD38F80E" box="[580,730,1941,1963]" italics="true" pageId="13" pageNumber="314">Abertella kewi</emphasis>
|
||
Durham, 1957
|
||
</taxonomicName>
|
||
) (based on specimens from UCMP locality B8562, Chiapas, Mexico). Food grooves indicated by dotted lines, peristome and periproct in solid black, interambulacra shaded, scale bars 10 mm in length.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF445AD5FFBCF5487D92F8ACFD61F84E" httpUri="https://zenodo.org/record/1135788/files/figure.png" pageId="14" pageNumber="315" targetBox="[365,1223,193,1866]" targetPageId="14">
|
||
<paragraph id="8B840A5DFFBCF5487D92F8ACFD61F84E" blockId="14.[151,1436,1883,2027]" pageId="14" pageNumber="315">
|
||
<emphasis id="B94FD64FFFBCF5487D92F8ACFEF1F8D5" bold="true" box="[151,275,1883,1905]" pageId="14" pageNumber="315">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C3B71DEFFBCF5487C1EF8ACFDDBF8D4" authority="Mooi & Martínez & Del Río & Ramos, 2018" authorityName="Mooi & Martínez & Del Río & Ramos" authorityYear="2018" box="[283,569,1883,1905]" class="Echinoidea" family="Abertellidae" genus="Camachoaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="14" pageNumber="315" phylum="Echinodermata" rank="species" species="maquedensis" status="sp. nov.">
|
||
<emphasis id="B94FD64FFFBCF5487C1EF8ACFDDBF8D4" box="[283,569,1883,1905]" italics="true" pageId="14" pageNumber="315">Camachoaster maquedensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C6B34FFBCF5487F45F8ACFD98F8D5" box="[576,634,1883,1904]" pageId="14" pageNumber="315" rank="species">
|
||
<emphasis id="B94FD64FFFBCF5487F45F8ACFD98F8D5" bold="true" box="[576,634,1883,1904]" pageId="14" pageNumber="315">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, details of test features. A: Holotype MACN-Pi 5809, pencil marks indicating partial aboral plate architecture in region within and around petal in ambulacrum II, gonopores indicated by white arrows, pores of trailing tube feet by black arrows. B: Paratype MACN-Pi 5859, complete aboral ambulacrum IV showing shape of petal. C: Holotype MACN-Pi 5809, details of tuberculation, food groove (black arrow), and peristome (peristomial projection indicated by white arrow). All scale bars 10 mm in length.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32159D6FFBDF5497DC2FF60FAE4FBB5" pageId="15" pageNumber="316" type="discussion">
|
||
<paragraph id="8B840A5DFFBDF5497DC2FF60FD66FEC0" blockId="15.[151,1437,151,1040]" pageId="15" pageNumber="316">
|
||
<emphasis id="B94FD64FFFBDF5497DC2FF60FED9FF15" bold="true" box="[199,315,151,176]" pageId="15" pageNumber="316">Remarks.</emphasis>
|
||
Although
|
||
<taxonomicName id="4C3B71DEFFBDF5497CB0FF6FFD13FF15" authority="Mooi & Martínez & Del Río & Ramos, 2018" authorityName="Mooi & Martínez & Del Río & Ramos" authorityYear="2018" box="[437,753,152,176]" class="Echinoidea" family="Abertellidae" genus="Camachoaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="species" species="maquedensis" status="sp. nov.">
|
||
<emphasis id="B94FD64FFFBDF5497CB0FF6FFD13FF15" box="[437,753,152,176]" italics="true" pageId="15" pageNumber="316">Camachoaster maquedensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C6B34FFBDF5497FFCFF60FCDBFF15" box="[761,825,151,176]" pageId="15" pageNumber="316" rank="species">
|
||
<emphasis id="B94FD64FFFBDF5497FFCFF60FCDBFF15" bold="true" box="[761,825,151,176]" pageId="15" pageNumber="316">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is found in the same strata as
|
||
<taxonomicName id="4C3B71DEFFBDF5497991FF6FFEBFFF70" authority="Mooi et al., 2016" authorityName="Mooi et al." authorityYear="2016" class="Echinoidea" family="Monophorasteridae" genus="Monophoraster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="species" species="telfordi">
|
||
<emphasis id="B94FD64FFFBDF5497991FF6FFA7EFF15" box="[1172,1436,152,176]" italics="true" pageId="15" pageNumber="316">Monophoraster telfordi</emphasis>
|
||
<bibRefCitation id="EFAA77ACFFBDF5497D92FF4BFEBFFF70" author="Mooi" box="[151,349,188,213]" pageId="15" pageNumber="316" refString="Mooi, R., Martinez, S. & del Rio, C. J. (2016) A new South American Miocene species of ' one-holed' sand dollar (Echinoidea: Clypeasteroida: Monophorasteridae). Zootaxa, 4173 (1), 45 - 54. https: // doi. org / 10.11646 / zootaxa. 4173.1.4" type="journal article" year="2016">
|
||
Mooi
|
||
<emphasis id="B94FD64FFFBDF5497DDEFF4AFEEEFF71" box="[219,268,188,212]" italics="true" pageId="15" pageNumber="316">et al</emphasis>
|
||
., 2016
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
, the former is immediately recognizable as a different species, as well as representing a different major scutelliform clade, by its complete lack of an anal lunule.
|
||
<taxonomicName id="4C3B71DEFFBDF5497E6BFF17FBF3FF5D" authority="Mooi & Martínez & Del Río & Ramos, 2018" authorityName="Mooi & Martínez & Del Río & Ramos" authorityYear="2018" box="[878,1041,224,248]" class="Echinoidea" family="Abertellidae" genus="Camachoaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="310" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94FD64FFFBDF5497E6BFF17FBF3FF5D" box="[878,1041,224,248]" italics="true" pageId="15" pageNumber="316">Camachoaster</emphasis>
|
||
</taxonomicName>
|
||
resembles the
|
||
<taxonomicName id="4C3B71DEFFBDF54979BEFF28FAA4FF5D" authorityName="Durham" authorityYear="1955" box="[1211,1350,223,248]" class="Echinoidea" family="Abertellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="family">Abertellidae</taxonomicName>
|
||
only in the possession of a shallow notch, and in the discontinuity of the posterior interambulacrum. However, the continuity of the paired interambulacra immediately invites favorable comparison with
|
||
<taxonomicName id="4C3B71DEFFBDF5497994FEDFFAFCFEE5" authority="Mooi & Martínez & Del Río & Ramos, 2018" authorityName="Mooi & Martínez & Del Río & Ramos" authorityYear="2018" box="[1169,1310,296,320]" class="Echinoidea" family="Eoscutellidae" genus="Placatenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="306" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94FD64FFFBDF5497994FEDFFAFCFEE5" box="[1169,1310,296,320]" italics="true" pageId="15" pageNumber="316">Placatenella</emphasis>
|
||
</taxonomicName>
|
||
, as do the similar shapes and dimensions of the petals.
|
||
</paragraph>
|
||
<paragraph id="8B840A5DFFBDF5497DC2FE98FAE4FBB5" blockId="15.[151,1437,151,1040]" pageId="15" pageNumber="316">
|
||
Nevertheless,
|
||
<taxonomicName id="4C3B71DEFFBDF5497C60FE87FDEBFE2D" authority="Mooi & Martínez & Del Río & Ramos, 2018" authorityName="Mooi & Martínez & Del Río & Ramos" authorityYear="2018" box="[357,521,368,392]" class="Echinoidea" family="Abertellidae" genus="Camachoaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="310" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94FD64FFFBDF5497C60FE87FDEBFE2D" box="[357,521,368,392]" italics="true" pageId="15" pageNumber="316">Camachoaster</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C3B71DEFFBDF5497F99FE87FCC8FE2D" authority="Mooi & Martínez & Del Río & Ramos, 2018" authorityName="Mooi & Martínez & Del Río & Ramos" authorityYear="2018" box="[668,810,368,392]" class="Echinoidea" family="Eoscutellidae" genus="Placatenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="306" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94FD64FFFBDF5497F99FE87FCC8FE2D" box="[668,810,368,392]" italics="true" pageId="15" pageNumber="316">Placatenella</emphasis>
|
||
</taxonomicName>
|
||
in having the posterior interambulacrum discontinuous.
|
||
<taxonomicName id="4C3B71DEFFBDF5497D92FE63FED8FE09" authority="Mooi & Martínez & Del Río & Ramos, 2018" authorityName="Mooi & Martínez & Del Río & Ramos" authorityYear="2018" box="[151,314,404,428]" class="Echinoidea" family="Abertellidae" genus="Camachoaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="310" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94FD64FFFBDF5497D92FE63FED8FE09" box="[151,314,404,428]" italics="true" pageId="15" pageNumber="316">Camachoaster</emphasis>
|
||
</taxonomicName>
|
||
's combination of continuous paired interambulacra with a discontinuous posterior, unpaired interambulacrum, is unique among South American forms. A small species of
|
||
<taxonomicName id="4C3B71DEFFBDF549790CFE4FFB78FE75" authorityName="Durham" authorityYear="1955" box="[1033,1178,440,464]" class="Echinoidea" family="Echinarachniidae" genus="Vaquerosella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B94FD64FFFBDF549790CFE4FFB78FE75" box="[1033,1178,440,464]" italics="true" pageId="15" pageNumber="316">Vaquerosella</emphasis>
|
||
</taxonomicName>
|
||
from California seems to have a similar oral plate pattern. However, among large scutelliforms with a posterior notch, this condition seems to be shared only with a Mexican species previously ascribed to
|
||
<taxonomicName id="4C3B71DEFFBDF5497EDAFDF7FBA5FDBD" authorityName="Durham" authorityYear="1953" box="[991,1095,512,536]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B94FD64FFFBDF5497EDAFDF7FBA5FDBD" box="[991,1095,512,536]" italics="true" pageId="15" pageNumber="316">Abertella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B71DEFFBDF549795CFDF7FA80FDBD" authority="Durham, 1957" authorityName="Durham" authorityYear="1957" box="[1113,1378,511,536]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="species" species="kewi">
|
||
<emphasis id="B94FD64FFFBDF549795CFDF7FB4DFDBD" box="[1113,1199,512,536]" italics="true" pageId="15" pageNumber="316">A. kewi</emphasis>
|
||
<bibRefCitation id="EFAA77ACFFBDF54979BFFE08FA80FDBD" author="Durham" box="[1210,1378,511,536]" pageId="15" pageNumber="316" refString="Durham, J. W. (1957) Notes on echinoids. Journal of Paleontology, 31 (3), 625 - 631." type="journal article" year="1957">Durham, 1957</bibRefCitation>
|
||
</taxonomicName>
|
||
. The original description of
|
||
<taxonomicName id="4C3B71DEFFBDF5497CA0FDD3FE1FFD99" authorityName="Durham" authorityYear="1957" box="[421,509,548,572]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="species" species="kewi">
|
||
<emphasis id="B94FD64FFFBDF5497CA0FDD3FE1FFD99" box="[421,509,548,572]" italics="true" pageId="15" pageNumber="316">A. kewi</emphasis>
|
||
</taxonomicName>
|
||
lacks figures of, or any reference to, oral plate architecture, likely due to the condition of the specimens. However, access to material from UCMP locality B8562, collected from the same locality as the types (Simojovel, Chiapas, Mexico) allows reconstruction of this architecture for comparison with
|
||
<taxonomicName id="4C3B71DEFFBDF5497D92FD67FE30FD0D" authority="Mooi & Martínez & Del Río & Ramos, 2018" authorityName="Mooi & Martínez & Del Río & Ramos" authorityYear="2018" box="[151,466,656,680]" class="Echinoidea" family="Abertellidae" genus="Camachoaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="species" species="maquedensis" status="sp. nov.">
|
||
<emphasis id="B94FD64FFFBDF5497D92FD67FE30FD0D" box="[151,466,656,680]" italics="true" pageId="15" pageNumber="316">Camachoaster maquedensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C6B34FFBDF5497CDCFD78FDFBFD0D" box="[473,537,655,680]" pageId="15" pageNumber="316" rank="species">
|
||
<emphasis id="B94FD64FFFBDF5497CDCFD78FDFBFD0D" bold="true" box="[473,537,655,680]" pageId="15" pageNumber="316">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="130016D8FFBDF5497F2DFD78FD8FFD0D" box="[552,621,655,680]" captionStart="FIGURE 7" captionStartId="13.[151,250,1910,1932]" captionTargetBox="[316,1279,208,1872]" captionTargetId="figure@13.[298,1291,193,1896]" captionTargetPageId="13" captionText="FIGURE 7. Comparison of oral plate architecture of Camachoaster maquedensis n. sp. (based on paratypes MACN-Pi 5859 and 5860) and Camachoaster? kewi (= Abertella kewi Durham, 1957) (based on specimens from UCMP locality B8562, Chiapas, Mexico).Food grooves indicated by dotted lines, peristome and periproctin solid black, interambulacra shaded, scale bars 10 mm in length." httpUri="https://zenodo.org/record/1135786/files/figure.png" pageId="15" pageNumber="316">Fig. 7</figureCitation>
|
||
). This suggests that
|
||
<taxonomicName id="4C3B71DEFFBDF5497E56FD67FC44FD0D" authorityName="Durham" authorityYear="1957" box="[851,934,656,680]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="species" species="kewi">
|
||
<emphasis id="B94FD64FFFBDF5497E56FD67FC44FD0D" box="[851,934,656,680]" italics="true" pageId="15" pageNumber="316">A. kewi</emphasis>
|
||
</taxonomicName>
|
||
is not an
|
||
<taxonomicName id="4C3B71DEFFBDF5497911FD67FB9EFD0D" authorityName="Durham" authorityYear="1953" box="[1044,1148,656,680]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B94FD64FFFBDF5497911FD67FB9EFD0D" box="[1044,1148,656,680]" italics="true" pageId="15" pageNumber="316">Abertella</emphasis>
|
||
</taxonomicName>
|
||
. Aspects of the oral plate pattern are reminiscent of some abertellid species, notably
|
||
<taxonomicName id="4C3B71DEFFBDF5497E57FD43FAE5FD69" authority="Kroh et al., 2013" authorityName="Kroh et al." authorityYear="2013" box="[850,1287,692,717]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="species" species="miskellyi">
|
||
<emphasis id="B94FD64FFFBDF5497E57FD43FBC8FD69" box="[850,1066,692,716]" italics="true" pageId="15" pageNumber="316">Abertella miskellyi</emphasis>
|
||
<bibRefCitation id="EFAA77ACFFBDF5497933FD43FAE5FD69" author="Kroh" box="[1078,1287,692,717]" pageId="15" pageNumber="316" refString="Kroh, A., Mooi, R., del Rio, C. & Neumann, C. (2013) A new late Cenozoic species of Abertella (Echinoidea: Clypeasteroida) from Patagonia. Zootaxa, 3608 (5), 369 - 378. https: // doi. org / 10.11646 / zootaxa. 3608.5.5" type="journal article" year="2013">
|
||
Kroh
|
||
<emphasis id="B94FD64FFFBDF5497979FD42FB5BFD69" box="[1148,1209,692,716]" italics="true" pageId="15" pageNumber="316">et al.</emphasis>
|
||
, 2013
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
, such as the periproct placement, the reduced posterior interambulacral basicoronal, and discontinuity of the posterior interambulacrum. However,
|
||
<taxonomicName id="4C3B71DEFFBDF5497CDCFD0BFDC9FCB1" authorityName="Durham" authorityYear="1957" box="[473,555,764,788]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="species" species="kewi">
|
||
<emphasis id="B94FD64FFFBDF5497CDCFD0BFDC9FCB1" box="[473,555,764,788]" italics="true" pageId="15" pageNumber="316">A. kewi</emphasis>
|
||
</taxonomicName>
|
||
is unlike any abertellid in the continuity of the paired interambulacra, and the insertion of very small first postbasicoronals in either the "a" or the "b" column of each of the ambulacra. The latter situation is seen in the Pliocene
|
||
<taxonomicName id="4C3B71DEFFBDF5497CFEFCB3FCFBFCF8" authority="Cragin, 1895" authorityName="Cragin" authorityYear="1895" box="[507,793,836,861]" class="Echinoidea" family="Echinarachniidae" genus="Scutellaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B94FD64FFFBDF5497CFEFCB3FD9DFCF9" box="[507,639,836,860]" italics="true" pageId="15" pageNumber="316">Scutellaster</emphasis>
|
||
<bibRefCitation id="EFAA77ACFFBDF5497F80FCB3FCFBFCF8" author="Cragin" box="[645,793,836,861]" pageId="15" pageNumber="316" refString="Cragin, F. W. (1895) A new Cretaceous genus of Clypeasteridae. American Geologist, 15, 90 - 91." type="journal article" year="1895">Cragin, 1895</bibRefCitation>
|
||
</taxonomicName>
|
||
, the Miocene and Pliocene
|
||
<taxonomicName id="4C3B71DEFFBDF549794EFCB2FAAFFCF8" authority="Nisiyama, 1935" authorityName="Nisiyama" authorityYear="1935" box="[1099,1357,836,861]" class="Echinoidea" family="Echinarachniidae" genus="Kewia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B94FD64FFFBDF549794EFCB2FB71FCF9" box="[1099,1171,837,860]" italics="true" pageId="15" pageNumber="316">Kewia</emphasis>
|
||
Nisiyama, 1935
|
||
</taxonomicName>
|
||
, and in the Eocene
|
||
<taxonomicName id="4C3B71DEFFBDF5497C24FC9FFE75FC25" authorityName="Grant & Hertlein" authorityYear="1938" box="[289,407,872,896]" class="Echinoidea" family="Eoscutellidae" genus="Eoscutella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B94FD64FFFBDF5497C24FC9FFE75FC25" box="[289,407,872,896]" italics="true" pageId="15" pageNumber="316">Eoscutella</emphasis>
|
||
</taxonomicName>
|
||
, all from the northwest Pacific coasts of North America. The occurrence of these small postbasicoronals has not yet been analyzed in a phylogenetic context in order to determine its overall significance. The aforementioned differences between
|
||
<taxonomicName id="4C3B71DEFFBDF5497F61FC47FC7DFC6D" authority="Mooi & Martínez & Del Río & Ramos, 2018" authorityName="Mooi & Martínez & Del Río & Ramos" authorityYear="2018" box="[612,927,944,968]" class="Echinoidea" family="Abertellidae" genus="Camachoaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="species" species="maquedensis" status="sp. nov.">
|
||
<emphasis id="B94FD64FFFBDF5497F61FC47FC7DFC6D" box="[612,927,944,968]" italics="true" pageId="15" pageNumber="316">Camachoaster maquedensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C6B34FFBDF5497EA0FC58FC01FC6D" box="[933,995,943,968]" pageId="15" pageNumber="316" rank="species">
|
||
<emphasis id="B94FD64FFFBDF5497EA0FC58FC01FC6D" bold="true" box="[933,995,943,968]" pageId="15" pageNumber="316">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C3B71DEFFBDF549791CFC47FB88FC6D" authorityName="Durham" authorityYear="1957" box="[1049,1130,944,968]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="species" species="kewi">
|
||
<emphasis id="B94FD64FFFBDF549791CFC47FB88FC6D" box="[1049,1130,944,968]" italics="true" pageId="15" pageNumber="316">A. kewi</emphasis>
|
||
</taxonomicName>
|
||
might suggest that they are not congeneric, but we here provisionally place
|
||
<taxonomicName id="4C3B71DEFFBDF5497FC1FC23FC04FC49" authority="in Camachoaster" authorityName="in Camachoaster" box="[708,998,980,1005]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="species" species="kewi">
|
||
<emphasis id="B94FD64FFFBDF5497FC1FC23FCFBFC49" box="[708,793,980,1004]" italics="true" pageId="15" pageNumber="316">A. kewi</emphasis>
|
||
in
|
||
<emphasis id="B94FD64FFFBDF5497E46FC23FC04FC49" box="[835,998,980,1004]" italics="true" pageId="15" pageNumber="316">Camachoaster</emphasis>
|
||
</taxonomicName>
|
||
as
|
||
<taxonomicName id="4C3B71DEFFBDF5497914FC23FB57FC49" authority="Mooi & Martínez & Del Río & Ramos, 2018" authorityName="Mooi & Martínez & Del Río & Ramos" authorityYear="2018" box="[1041,1205,980,1004]" class="Echinoidea" family="Abertellidae" genus="Camachoaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="310" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94FD64FFFBDF5497914FC23FB57FC49" box="[1041,1205,980,1004]" italics="true" pageId="15" pageNumber="316">Camachoaster</emphasis>
|
||
</taxonomicName>
|
||
?
|
||
<taxonomicName id="4C3B71DEFFBDF54979D1FC23FAE5FC49" box="[1236,1287,980,1004]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="species" species="kewi">
|
||
<emphasis id="B94FD64FFFBDF54979D1FC23FAE5FC49" box="[1236,1287,980,1004]" italics="true" pageId="15" pageNumber="316">kewi</emphasis>
|
||
</taxonomicName>
|
||
pending full revision of all the relevant North American taxa associated with the
|
||
<taxonomicName id="4C3B71DEFFBDF5497E96FC00FBFFFBB5" authorityName="Durham" authorityYear="1955" box="[915,1053,1015,1040]" class="Echinoidea" family="Abertellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="316" phylum="Echinodermata" rank="family">Abertellidae</taxonomicName>
|
||
and Placatenellidae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |