171 lines
18 KiB
XML
171 lines
18 KiB
XML
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<mods:title id="2323036DD8BBF56C3BAE19BC13DB876C">Late Oligocene - Miocene non-lunulate sand dollars of South America: Revision of abertellid taxa and descriptions of two new families, two new genera, and a new species</mods:title>
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<mods:namePart id="03BF22C16E2B3E0D03E041FA874BE86D">Mooi, Rich</mods:namePart>
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<mods:namePart id="ABEB07858017B4A6C98217A3535A7DD8">Martínez, Sergio A.</mods:namePart>
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<mods:namePart id="7EEF7CE153B302E37513FD1C49A77BB2">Del Río, Claudia J.</mods:namePart>
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<mods:namePart id="9DEE5D782F1D08935C81C4E4A0D6E883">Ramos, Maria Inês Feijó</mods:namePart>
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<mods:date id="2876CAF0CB06AE09DD329F66C77CB494">2018</mods:date>
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<treatment id="0392BB4BFFA3F5577D92FE39FDB5FBCD" ID-DOI="http://doi.org/10.5281/zenodo.5961804" ID-GBIF-Taxon="139582396" ID-Zenodo-Dep="5961804" LSID="urn:lsid:plazi:treatment:0392BB4BFFA3F5577D92FE39FDB5FBCD" httpUri="http://treatment.plazi.org/id/0392BB4BFFA3F5577D92FE39FDB5FBCD" lastPageNumber="318" pageId="17" pageNumber="318">
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<subSubSection id="C32159D6FFA3F5577D92FE39FD36FD8E" pageId="17" pageNumber="318" type="nomenclature">
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<paragraph id="8B840A5DFFA3F5577D92FE39FDAFFE4D" blockId="17.[151,589,462,488]" box="[151,589,462,488]" pageId="17" pageNumber="318">
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<heading id="D0CCBD31FFA3F5577D92FE39FDAFFE4D" bold="true" box="[151,589,462,488]" fontSize="11" level="1" pageId="17" pageNumber="318" reason="1">
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<taxonomicName id="4C3B71DEFFA3F5577D92FE39FDAFFE4D" authority="Kroh et al., 2013" authorityName="Kroh et al." authorityYear="2013" box="[151,589,462,488]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="17" pageNumber="318" phylum="Echinodermata" rank="species" species="miskellyi">
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<emphasis id="B94FD64FFFA3F5577D92FE39FDAFFE4D" bold="true" box="[151,589,462,488]" pageId="17" pageNumber="318">
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<emphasis id="B94FD64FFFA3F5577D92FE39FE9BFE4D" bold="true" box="[151,377,462,488]" italics="true" pageId="17" pageNumber="318">Abertella miskellyi</emphasis>
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<bibRefCitation id="EFAA77ACFFA3F5577C85FE39FDAFFE4D" author="Kroh" box="[384,589,462,488]" pageId="17" pageNumber="318" refString="Kroh, A., Mooi, R., del Rio, C. & Neumann, C. (2013) A new late Cenozoic species of Abertella (Echinoidea: Clypeasteroida) from Patagonia. Zootaxa, 3608 (5), 369 - 378. https: // doi. org / 10.11646 / zootaxa. 3608.5.5" type="journal article" year="2013">
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Kroh
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<emphasis id="B94FD64FFFA3F5577CCCFE39FDE6FE4D" bold="true" box="[457,516,462,488]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
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, 2013
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</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B840A5DFFA3F5577D92FDE2FD36FD8E" blockId="17.[151,724,533,555]" box="[151,724,533,555]" pageId="17" pageNumber="318">
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<bibRefCitation id="EFAA77ACFFA3F5577D92FDE2FF28FD8E" author="Kroh" box="[151,202,533,555]" pageId="17" pageNumber="318" refString="Kroh, A., Mooi, R., del Rio, C. & Neumann, C. (2013) A new late Cenozoic species of Abertella (Echinoidea: Clypeasteroida) from Patagonia. Zootaxa, 3608 (5), 369 - 378. https: // doi. org / 10.11646 / zootaxa. 3608.5.5" type="journal article" year="2013">2013</bibRefCitation>
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<taxonomicName id="4C3B71DEFFA3F5577DD5FDE2FDE0FD8E" authority="Kroh et al." authorityName="Kroh et al." box="[208,514,533,555]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="17" pageNumber="318" phylum="Echinodermata" rank="species" species="miskellyi">
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<emphasis id="B94FD64FFFA3F5577DD5FDE2FE6CFD8E" box="[208,398,533,555]" italics="true" pageId="17" pageNumber="318">Abertella miskellyi</emphasis>
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Kroh
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<emphasis id="B94FD64FFFA3F5577CCAFDE1FDE0FD8E" box="[463,514,533,555]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
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||
</taxonomicName>
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||
: 371–376, figs. 2–6.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C32159D6FFA3F5577D92FDA0FDD5FD79" pageId="17" pageNumber="318" type="diagnosis">
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<paragraph id="8B840A5DFFA3F5577D92FDA0FDD5FD79" blockId="17.[151,1437,599,1128]" pageId="17" pageNumber="318">
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<emphasis id="B94FD64FFFA3F5577D92FDA0FEE8FDD5" bold="true" box="[151,266,599,624]" pageId="17" pageNumber="318">Diagnosis</emphasis>
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.
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<taxonomicName id="4C3B71DEFFA3F5577C1DFDAFFE62FDD5" authorityName="Durham" authorityYear="1953" box="[280,384,600,624]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="17" pageNumber="318" phylum="Echinodermata" rank="genus">
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<emphasis id="B94FD64FFFA3F5577C1DFDAFFE62FDD5" box="[280,384,600,624]" italics="true" pageId="17" pageNumber="318">Abertella</emphasis>
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</taxonomicName>
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with strongly heterogeneous interambulacral basicoronal plates (small in interambulacrum 5, largest in interambulacra 2 and 3); oral interambulacra discontinuous by involvement of two adjacent ambulacral plates (rather than one as in other
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<taxonomicName id="4C3B71DEFFA3F5577F2EFD57FD71FD1D" authorityName="Durham" authorityYear="1953" box="[555,659,672,696]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="17" pageNumber="318" phylum="Echinodermata" rank="genus">
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<emphasis id="B94FD64FFFA3F5577F2EFD57FD71FD1D" box="[555,659,672,696]" italics="true" pageId="17" pageNumber="318">Abertella</emphasis>
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</taxonomicName>
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) in at least one of the oral interambulacra; apparent "violation" of Lovén's Rule in oral ambulacrum III.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32159D6FFA3F5577DC2FD10FC29FCA5" box="[199,971,743,768]" pageId="17" pageNumber="318" type="materials_examined">
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<paragraph id="8B840A5DFFA3F5577DC2FD10FC29FCA5" blockId="17.[151,1437,599,1128]" box="[199,971,743,768]" pageId="17" pageNumber="318">
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<emphasis id="B94FD64FFFA3F5577DC2FD10FE31FCA5" bold="true" box="[199,467,743,768]" pageId="17" pageNumber="318">
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<typeStatus id="5480B4FFFFA3F5577DC2FD10FEE0FCA5" box="[199,258,743,768]" pageId="17" pageNumber="318">Type</typeStatus>
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material studied.
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||
</emphasis>
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<typeStatus id="5480B4FFFFA3F5577CDFFD10FDA0FCA5" box="[474,578,743,768]" pageId="17" pageNumber="318" type="holotype">Holotype</typeStatus>
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MB E.7463,
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<typeStatus id="5480B4FFFFA3F5577FDCFD1FFCD8FCA5" box="[729,826,744,768]" pageId="17" pageNumber="318" type="paratype">paratype</typeStatus>
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MB E.7462.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32159D6FFA3F5577DC2FCFBFD4DFCC9" pageId="17" pageNumber="318" type="description">
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<paragraph id="8B840A5DFFA3F5577DC2FCFBFD82FC81" blockId="17.[151,1437,599,1128]" box="[199,608,780,805]" pageId="17" pageNumber="318">
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<emphasis id="B94FD64FFFA3F5577DC2FCFBFEBBFC80" bold="true" box="[199,345,780,805]" pageId="17" pageNumber="318">Description.</emphasis>
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See
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<bibRefCitation id="EFAA77ACFFA3F5577C8BFCFBFDBEFC81" box="[398,604,780,805]" pageId="17" pageNumber="318" refString="Kroh, A., Mooi, R., del Rio, C. & Neumann, C. (2013) A new late Cenozoic species of Abertella (Echinoidea: Clypeasteroida) from Patagonia. Zootaxa, 3608 (5), 369 - 378. https: // doi. org / 10.11646 / zootaxa. 3608.5.5" type="journal article">
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Kroh
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<emphasis id="B94FD64FFFA3F5577CCAFCFAFDE5FC81" box="[463,519,780,804]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
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(2013)
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</bibRefCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8B840A5DFFA3F5577DC2FCD8FD4DFCC9" blockId="17.[151,1437,599,1128]" pageId="17" pageNumber="318">
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<materialsCitation id="3B530000FFA3F5577DC2FCD8FD4DFCC9" ID-GBIF-Occurrence="1802482791" country="Argentina" location="Chubut Province" pageId="17" pageNumber="318" specimenCount="1" stateProvince="Chubut Province">
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<emphasis id="B94FD64FFFA3F5577DC2FCD8FEBBFCED" bold="true" box="[199,345,815,840]" pageId="17" pageNumber="318">Occurrence.</emphasis>
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<taxonomicName id="4C3B71DEFFA3F5577C66FCC7FE0BFCED" box="[355,489,816,840]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="17" pageNumber="318" phylum="Echinodermata" rank="species" species="miskellyi">
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<emphasis id="B94FD64FFFA3F5577C66FCC7FE0BFCED" box="[355,489,816,840]" italics="true" pageId="17" pageNumber="318">A. miskellyi</emphasis>
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</taxonomicName>
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is recorded only from the (possibly early) Miocene of
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<collectingRegion id="49FFC4BFFFA3F557796EFCD8FACFFCED" box="[1131,1325,815,840]" country="Argentina" name="Chubut" pageId="17" pageNumber="318">Chubut Province</collectingRegion>
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, southern
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<collectingCountry id="F32C4ACDFFA3F5577D92FCA3FEE4FCC8" box="[151,262,852,877]" name="Argentina" pageId="17" pageNumber="318">Argentina</collectingCountry>
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in the Camarones Formation (
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<figureCitation id="130016D8FFA3F5577F59FCA3FD7CFCC9" box="[604,670,852,877]" captionStart="FIGURE 1" captionStartId="3.[151,250,1776,1798]" captionTargetBox="[162,1426,462,1692]" captionTargetId="figure@3.[151,1436,451,1700]" captionTargetPageId="3" captionText="FIGURE 1. Localities for taxa discussed in the text. Inset at lower left centers on the Santa Cruz region of Patagonia. Numbered occurrences correspond to unequivocal Iheringiella patagonensis localities (inside boxed region in main map): 1. Pampa Castillo, Chile (Andre Wyss, pers. comm.); 2. Cañadón El Lobo, Santa Cruz, Argentina (CPBA 16493-95); 3. Oven Point/Sholl's Tomb, Santa Cruz, Argentina (Ortmann 1902), Punta Cuevas, Santa Cruz, Argentina (MACN 4586 and Cuitiño 2011), Playa La Mina, Santa Cruz, Argentina (Cuitiño 2011); 4. Meseta Chica, Bajo de San Julián, Santa Cruz, Argentina (MACN 5144 and Cuitiño 2011); 5. Upper Rio Chalia, Santa Cruz, Argentina (Ortmann 1902); 6. Puerto Santa Cruz, Santa Cruz, Argentina (ROM 5433M, 5468M, 5469M); 7. Sector Flamenco, Tierra del Fuego, Chile (Larrain 1984)—reported to be from Eocene strata, but this is questionable (Mooi et al. 2000)." httpUri="https://zenodo.org/record/1135774/files/figure.png" pageId="17" pageNumber="318">Fig. 1</figureCitation>
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).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32159D6FFA3F5577DC2FC80FDB5FBCD" pageId="17" pageNumber="318" type="discussion">
|
||
<paragraph id="8B840A5DFFA3F5577DC2FC80FDB5FBCD" blockId="17.[151,1437,599,1128]" pageId="17" pageNumber="318">
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||
<emphasis id="B94FD64FFFA3F5577DC2FC80FED9FC35" bold="true" box="[199,315,887,912]" pageId="17" pageNumber="318">Remarks.</emphasis>
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This species is unique among
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<taxonomicName id="4C3B71DEFFA3F5577FA1FC8FFCEEFC35" authorityName="Durham" authorityYear="1953" box="[676,780,888,912]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="17" pageNumber="318" phylum="Echinodermata" rank="genus">
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<emphasis id="B94FD64FFFA3F5577FA1FC8FFCEEFC35" box="[676,780,888,912]" italics="true" pageId="17" pageNumber="318">Abertella</emphasis>
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</taxonomicName>
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in that the posterior interambulacral basicoronal plate is much smaller than that in each of the paired interambulacral basicoronals. It is also the only
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<taxonomicName id="4C3B71DEFFA3F55779C2FC6BFACDFC11" authorityName="Durham" authorityYear="1953" box="[1223,1327,924,948]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="17" pageNumber="318" phylum="Echinodermata" rank="genus">
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||
<emphasis id="B94FD64FFFA3F55779C2FC6BFACDFC11" box="[1223,1327,924,948]" italics="true" pageId="17" pageNumber="318">Abertella</emphasis>
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</taxonomicName>
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in which specimens are known to have a discontinuity involving more than one postbasicoronal plate in a given halfambulacrum. An unusual feature of one of the known specimens of
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<taxonomicName id="4C3B71DEFFA3F5577E8CFC13FBE9FC59" box="[905,1035,996,1020]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="17" pageNumber="318" phylum="Echinodermata" rank="species" species="miskellyi">
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<emphasis id="B94FD64FFFA3F5577E8CFC13FBE9FC59" box="[905,1035,996,1020]" italics="true" pageId="17" pageNumber="318">A. miskellyi</emphasis>
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</taxonomicName>
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is that ambulacral basicoronal III at first seemed longer in the "a" column than in the "b" column, suggesting violation of Lovén's Rule (
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<emphasis id="B94FD64FFFA3F55779F2FBFEFAD7FB85" box="[1271,1333,1033,1056]" italics="true" pageId="17" pageNumber="318">sensu</emphasis>
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<bibRefCitation id="EFAA77ACFFA3F5577839FBF0FF16FBE0" author="David" pageId="17" pageNumber="318" refString="David, B., Mooi, R. & Telford, M. (1996) The ontogenetic basis of Loven's Rule clarifies homologies of the echinoid peristome. In: Emson, R., Smith, A. B. & Campbell, A. (Eds.), Echinoderm Research 1995. A. A. Balkema, Rotterdam, pp. 155 - 164." type="journal article" year="1996">
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||
David
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<emphasis id="B94FD64FFFA3F557788DFBFEFF4FFBE1" italics="true" pageId="17" pageNumber="318">et al</emphasis>
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||
. 1996
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||
</bibRefCitation>
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||
) until a small plate was detected, wedged between the basicoronal plates of ambulacrum III to re-establish the Lovénian pattern (
|
||
<bibRefCitation id="EFAA77ACFFA3F5577C88FBB8FDA9FBCD" author="Kroh" box="[397,587,1103,1128]" pageId="17" pageNumber="318" refString="Kroh, A., Mooi, R., del Rio, C. & Neumann, C. (2013) A new late Cenozoic species of Abertella (Echinoidea: Clypeasteroida) from Patagonia. Zootaxa, 3608 (5), 369 - 378. https: // doi. org / 10.11646 / zootaxa. 3608.5.5" type="journal article" year="2013">
|
||
Kroh
|
||
<emphasis id="B94FD64FFFA3F5577CCAFBA6FDEAFBCD" box="[463,520,1104,1128]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |