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<mods:title id="294C07037CBEA257B70CCCE6124B2198">Revision of Scissurellidae, Anatomidae and Fissurellidae (Gastropoda: Vetigastropoda) from the Plio-Pleistocene of the Philippines</mods:title>
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<mods:namePart id="275D34F6D9FAAB1AFCFBA9C4C05F5EC9">Helwerda, Renate A.</mods:namePart>
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<mods:date id="0435BE89720A20DC154E992DCE39517C">2014</mods:date>
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<treatment id="039287E3FFF9FFC5FF41FD3CFE92FA81" ID-DOI="http://doi.org/10.5281/zenodo.5625973" ID-GBIF-Taxon="119511863" ID-Zenodo-Dep="5625973" LSID="urn:lsid:plazi:treatment:039287E3FFF9FFC5FF41FD3CFE92FA81" httpUri="http://treatment.plazi.org/id/039287E3FFF9FFC5FF41FD3CFE92FA81" lastPageNumber="189" pageId="6" pageNumber="189">
<subSubSection id="C321657EFFF9FFC5FF41FD3CFD23FC9B" pageId="6" pageNumber="189" type="nomenclature">
<paragraph id="8B8436F5FFF9FFC5FF41FD3CFDF6FCEC" blockId="6.[151,543,758,819]" box="[151,543,758,783]" pageId="6" pageNumber="189">
<heading id="D0CC8199FFF9FFC5FF41FD3CFDF6FCEC" bold="true" box="[151,543,758,783]" fontSize="10" level="2" pageId="6" pageNumber="189" reason="2">
<taxonomicName id="4C3B4D76FFF9FFC5FF41FD3CFDF6FCEC" ID-CoL="39FDJ" authority="Habe, 1953" authorityName="Habe" authorityYear="1953" box="[151,543,758,783]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="6" pageNumber="189" phylum="Mollusca" rank="species" species="tosaensis">
<emphasis id="B94FEAE7FFF9FFC5FF41FD3CFDF6FCEC" bold="true" box="[151,543,758,783]" pageId="6" pageNumber="189">
<emphasis id="B94FEAE7FFF9FFC5FF41FD3CFE7BFCED" bold="true" box="[151,402,758,783]" italics="true" pageId="6" pageNumber="189">Emarginula tosaensis</emphasis>
<bibRefCitation id="EFAA4B04FFF9FFC5FE4FFD3CFDF6FCEC" author="Habe" box="[409,543,758,783]" pageId="6" pageNumber="189" refString="Habe, T. (1953) Fissurellidae in Japan (2). Publications of the Seto Marine Biological Laboratory, 3, 33 - 51." type="journal article" year="1953">Habe, 1953</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8B8436F5FFF9FFC5FF41FCD0FEFCFCD0" blockId="6.[151,543,758,819]" box="[151,277,794,819]" pageId="6" pageNumber="189">
(
<figureCitation id="13002A70FFF9FFC5FF48FCD0FEE4FCD0" box="[158,269,794,819]" captionStart="FIGURES 10 12. 10" captionStartId="7.[151,264,1858,1880]" captionTargetBox="[167,1410,193,1800]" captionTargetId="figure@7.[167,1411,193,1802]" captionTargetPageId="7" captionText="FIGURES 10 12. 10. Emarginula tosaensis Habe, 1953. RGM 961.722. Locality Anda 3. (a) left view, (b) right view, (c) basal view, (d) apical view. 11. Emarginula aff. souverbiana Pilsbry, 1890. RGM 961.723. Locality Anda 6. (a) left view, (b) right view, (c) basal view, (d) apical view. 12. Zeidora geigeri nov. spec. RGM 961.724. Locality Anda 2. (a) left view, (b) right view, (c) basal view, (d) apical view, (e) SEM apical view, (f) SEM detail of protoconch." httpUri="https://zenodo.org/record/229065/files/figure.png" pageId="6" pageNumber="189">Figure 10</figureCitation>
)
</paragraph>
<paragraph id="8B8436F5FFF9FFC5FF41FCA8FD23FC9B" blockId="6.[151,892,866,950]" box="[151,714,866,888]" pageId="6" pageNumber="189">
1953
<taxonomicName id="4C3B4D76FFF9FFC5FF39FCA8FE23FC9B" box="[239,458,866,888]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="6" pageNumber="189" phylum="Mollusca" rank="species" species="tosaensis">
<emphasis id="B94FEAE7FFF9FFC5FF39FCA8FE23FC9B" box="[239,458,866,888]" italics="true" pageId="6" pageNumber="189">Emarginula tosaensis</emphasis>
</taxonomicName>
nov. spec.—Habe: p. 39.
</paragraph>
</subSubSection>
<subSubSection id="C321657EFFF9FFC5FF41FC4BFC95FC55" pageId="6" pageNumber="189" type="reference_group">
<paragraph id="8B8436F5FFF9FFC5FF41FC4BFCF8FC74" blockId="6.[151,892,866,950]" box="[151,785,897,919]" pageId="6" pageNumber="189">
1999
<taxonomicName id="4C3B4D76FFF9FFC5FF39FC4BFDAAFC74" ID-CoL="39FDJ" authority="Habe, 1953" authorityName="Habe" authorityYear="1953" box="[239,579,897,919]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="6" pageNumber="189" phylum="Mollusca" rank="species" species="tosaensis">
<emphasis id="B94FEAE7FFF9FFC5FF39FC4BFE23FC74" box="[239,458,897,919]" italics="true" pageId="6" pageNumber="189">Emarginula tosaensis</emphasis>
<bibRefCitation id="EFAA4B04FFF9FFC5FE06FC4BFDAAFC74" author="Habe" box="[464,579,897,919]" pageId="6" pageNumber="189" refString="Habe, T. (1953) Fissurellidae in Japan (2). Publications of the Seto Marine Biological Laboratory, 3, 33 - 51." type="journal article" year="1953">Habe, 1953</bibRefCitation>
</taxonomicName>
—Higo
<emphasis id="B94FEAE7FFF9FFC5FD41FC48FD22FC74" box="[663,715,897,919]" italics="true" pageId="6" pageNumber="189">et al.</emphasis>
: p. 39.
</paragraph>
<paragraph id="8B8436F5FFF9FFC5FF41FC6AFC95FC55" blockId="6.[151,892,866,950]" box="[151,892,928,950]" pageId="6" pageNumber="189">
2000
<taxonomicName id="4C3B4D76FFF9FFC5FF39FC6AFDADFC55" ID-CoL="39FDJ" authority="Habe, 1953" authorityName="Habe" authorityYear="1953" box="[239,580,928,950]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="6" pageNumber="189" phylum="Mollusca" rank="species" species="tosaensis">
<emphasis id="B94FEAE7FFF9FFC5FF39FC6AFE23FC55" box="[239,458,928,950]" italics="true" pageId="6" pageNumber="189">Emarginula tosaensis</emphasis>
<bibRefCitation id="EFAA4B04FFF9FFC5FE06FC6AFDADFC55" author="Habe" box="[464,580,928,950]" pageId="6" pageNumber="189" refString="Habe, T. (1953) Fissurellidae in Japan (2). Publications of the Seto Marine Biological Laboratory, 3, 33 - 51." type="journal article" year="1953">Habe, 1953</bibRefCitation>
</taxonomicName>
—Sasaki: p. 45, pl. 22, fig. 10.
</paragraph>
</subSubSection>
<subSubSection id="C321657EFFF9FFC5FF41FC28FC90FB69" pageId="6" pageNumber="189" type="materials_examined">
<paragraph id="8B8436F5FFF9FFC5FF41FC28FDC6FC18" blockId="6.[151,1436,994,1379]" box="[151,559,994,1019]" pageId="6" pageNumber="189">
<emphasis id="B94FEAE7FFF9FFC5FF41FC28FEEFFC18" bold="true" box="[151,262,994,1019]" pageId="6" pageNumber="189">Material.</emphasis>
Anda3 (1)
<collectionCode id="ED2AAE30FFF9FFC5FE50FC29FE2FFC18" box="[390,454,995,1019]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="189">RGM</collectionCode>
961.722.
</paragraph>
<paragraph id="8B8436F5FFF9FFC5FF11FBCCFC90FB69" blockId="6.[151,1436,994,1379]" pageId="6" pageNumber="189">
<emphasis id="B94FEAE7FFF9FFC5FF11FBCCFE73FBFC" bold="true" box="[199,410,1030,1055]" pageId="6" pageNumber="189">Characterization.</emphasis>
Shell patelliform; L
<quantity id="4CC39B10FFF9FFC5FD5AFBCDFD1CFBFD" box="[652,757,1031,1055]" metricMagnitude="-3" metricUnit="m" metricValue="8.97" pageId="6" pageNumber="189" unit="mm" value="8.97">8.97 mm</quantity>
; H
<quantity id="4CC39B10FFF9FFC5FCC9FBCDFC61FBFD" box="[799,904,1031,1055]" metricMagnitude="-3" metricUnit="m" metricValue="4.07" pageId="6" pageNumber="189" unit="mm" value="4.07">4.07 mm</quantity>
;
<geoCoordinate id="EE0F5032FFF9FFC5FC40FBCDFC03FBFC" box="[918,1002,1030,1055]" direction="west" orientation="longitude" pageId="6" pageNumber="189" precision="555" value="-6.23">
W
<quantity id="4CC39B10FFF9FFC5FC6FFBCCFC03FBFC" box="[953,1002,1030,1055]" metricMagnitude="-3" metricUnit="m" metricValue="6.23" pageId="6" pageNumber="189" unit="mm" value="6.23">6.23</quantity>
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mm; apex at 0.75 of shell length, not strongly involute; sculpture of about 40 radial ribs, primary and secondary ribs difficult to distinguish, commarginal ribs forming dense knobs on radial ribs; selenizone filled with sediment, demarcated by moderately strong keels; slit well developed; posterior slope slightly concave.
</paragraph>
</subSubSection>
<subSubSection id="C321657EFFF9FFC5FF11FB5CFB94FB30" pageId="6" pageNumber="189" type="distribution">
<paragraph id="8B8436F5FFF9FFC5FF11FB5CFB94FB30" blockId="6.[151,1436,994,1379]" pageId="6" pageNumber="189">
<emphasis id="B94FEAE7FFF9FFC5FF11FB5CFE89FB4C" bold="true" box="[199,352,1174,1199]" pageId="6" pageNumber="189">Distribution.</emphasis>
Sagami Bay to Western Kyushu,
<collectingCountry id="F32C7665FFF9FFC5FD33FB5DFCC1FB4D" box="[741,808,1175,1198]" name="Japan" pageId="6" pageNumber="189">Japan</collectingCountry>
,
<quantity id="4CC39B10FFF9FFC5FCE2FB5CFC4EFB4D" box="[820,935,1174,1199]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="6" pageNumber="189" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50150 m</quantity>
(
<bibRefCitation id="EFAA4B04FFF9FFC5FC6EFB5CFB91FB4C" author="Higo" box="[952,1144,1174,1199]" pageId="6" pageNumber="189" refString="Higo, S., Callomon, P. &amp; Goto, Y. (1999) Catalogue and Bibliography of the Marine Shell-Bearing Mollusca of Japan. Elle Scientific Publications, Osaka, pp. 749." type="book chapter" year="1999">
Higo
<emphasis id="B94FEAE7FFF9FFC5FC2CFB5DFBDDFB4D" box="[1018,1076,1174,1198]" italics="true" pageId="6" pageNumber="189">et al.</emphasis>
1999
</bibRefCitation>
;
<bibRefCitation id="EFAA4B04FFF9FFC5FB53FB5CFAFDFB4C" author="Sasaki" box="[1157,1300,1174,1199]" pageId="6" pageNumber="189" refString="Sasaki, T. (2000) Family Fissurellidae. In: Okutani, T. (Ed.), Marine Mollusks in Japan. Tokai University Press, Tokyo, pp. 45 - 53." type="book chapter" year="2000">Sasaki 2000</bibRefCitation>
). Also from the Pleistocene of the Moeshima shell bed, Kagoshima Bay, Kyushu,
<collectingCountry id="F32C7665FFF9FFC5FC75FB71FC0AFB31" box="[931,995,1211,1234]" name="Japan" pageId="6" pageNumber="189">Japan</collectingCountry>
(
<bibRefCitation id="EFAA4B04FFF9FFC5FC27FB70FB98FB30" author="Habe" box="[1009,1137,1210,1235]" pageId="6" pageNumber="189" refString="Habe, T. (1953) Fissurellidae in Japan (2). Publications of the Seto Marine Biological Laboratory, 3, 33 - 51." type="journal article" year="1953">Habe 1953</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C321657EFFF9FFC5FF11FB14FE92FA81" pageId="6" pageNumber="189" type="discussion">
<paragraph id="8B8436F5FFF9FFC5FF11FB14FE92FA81" blockId="6.[151,1436,994,1379]" pageId="6" pageNumber="189">
<emphasis id="B94FEAE7FFF9FFC5FF11FB14FED2FB14" bold="true" box="[199,315,1246,1271]" pageId="6" pageNumber="189">Remarks.</emphasis>
<taxonomicName id="4C3B4D76FFF9FFC5FE92FB14FC85FB14" authority="Chapman &amp; Gabriel, 1923" authorityName="Chapman &amp; Gabriel" authorityYear="1923" box="[324,876,1246,1271]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="6" pageNumber="189" phylum="Mollusca" rank="species" species="dennanti">
<emphasis id="B94FEAE7FFF9FFC5FE92FB14FDDDFB15" box="[324,564,1246,1270]" italics="true" pageId="6" pageNumber="189">Emarginula dennanti</emphasis>
<bibRefCitation id="EFAA4B04FFF9FFC5FDEBFB14FC85FB14" author="Chapman" box="[573,876,1246,1271]" pageId="6" pageNumber="189" refString="Chapman, F. &amp; Gabriel, C. J. (1923) A revision and description of the Australian Tertiary Patellidae, Patelloididae, Cocculinidae and Fissurellidae. Proceedings of the Royal Society of Victoria, 36, 22 - 40." type="journal article" year="1923">Chapman &amp; Gabriel, 1923</bibRefCitation>
</taxonomicName>
from the Oligocene and Miocene of
<collectingCountry id="F32C7665FFF9FFC5FACCFB14FA69FB14" box="[1306,1408,1246,1271]" name="Australia" pageId="6" pageNumber="189">Australia</collectingCountry>
is quite similar with respect to its shape and radial ribbing, but its commarginal ribs are spaced farther apart.
<taxonomicName id="4C3B4D76FFF9FFC5FA52FAC9FE56FADC" authority="Pilsbry, 1890" authorityName="Pilsbry" authorityYear="1890" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="6" pageNumber="189" phylum="Mollusca" rank="species" species="souverbiana">
<emphasis id="B94FEAE7FFF9FFC5FA52FAC9FECBFADD" italics="true" pageId="6" pageNumber="189">E. souverbiana</emphasis>
<bibRefCitation id="EFAA4B04FFF9FFC5FEFCFAECFE56FADC" author="Pilsbry" box="[298,447,1318,1343]" pageId="6" pageNumber="189" refString="Pilsbry, H. A. (1890) Manual of Conchology; Structural and Systematic. Vol. 12. Academy of Natural Sciences of Philadelphia, Philadelphia, 323 pp." type="book" year="1890">Pilsbry, 1890</bibRefCitation>
</taxonomicName>
is also similar in its shape and radial ribbing, but its commarginal ribs are weaker and spaced farther apart.
</paragraph>
</subSubSection>
</treatment>
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