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<document id="0D70CE720C54938B5367B54EECB24712" ID-CLB-Dataset="46097" ID-DOI="10.11646/zootaxa.3784.4.4" ID-GBIF-Dataset="16f488d9-eb77-4ff5-a7ce-99eeb7875371" ID-ISSN="1175-5326" ID-Zenodo-Dep="252218" ID-ZooBank="D5E03502-7BD3-41F4-A4CF-5537B1462A23" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460404217522" checkinUser="plazi" docAuthor="Makarkin, Vladimir N. &amp; Archibald, S. Bruce" docDate="2014" docId="039287A3FE15370623C76D60FB2DFB07" docLanguage="en" docName="zt03784p444.pdf" docOrigin="Zootaxa 3784 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Archiraphidia Handlirsch 1910" docType="treatment" docVersion="9" lastPageNumber="421" masterDocId="FFABFFDBFE06371223506A3EFFE0FF82" masterDocTitle="A revision of the late Eocene snakeflies (Raphidioptera) of the Florissant Formation, Colorado, with special reference to the wing venation of the Raphidiomorpha" masterLastPageNumber="444" masterPageNumber="401" pageNumber="420" updateTime="1698299590966" updateUser="ExternalLinkService">
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<mods:title id="5C958B1FEBF8200B72B63937EE934DC7">A revision of the late Eocene snakeflies (Raphidioptera) of the Florissant Formation, Colorado, with special reference to the wing venation of the Raphidiomorpha</mods:title>
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<mods:namePart id="6CD806F7EF2BF5E17B70E9D18D3C7F74">Makarkin, Vladimir N.</mods:namePart>
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<mods:namePart id="7C18401654E44291423708DBCA7913B9">Archibald, S. Bruce</mods:namePart>
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<mods:date id="9F2D69FD17D4A39C3025F0E36A6DBBDC">2014</mods:date>
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<treatment id="039287A3FE15370623C76D60FB2DFB07" ID-DOI="http://doi.org/10.5281/zenodo.6131157" ID-GBIF-Taxon="119420866" ID-Zenodo-Dep="6131157" LSID="urn:lsid:plazi:treatment:039287A3FE15370623C76D60FB2DFB07" httpUri="http://treatment.plazi.org/id/039287A3FE15370623C76D60FB2DFB07" lastPageId="20" lastPageNumber="421" pageId="19" pageNumber="420">
<subSubSection id="C321653EFE15370123C76D60FD96F8FA" box="[151,630,1886,1912]" pageId="19" pageNumber="420" type="nomenclature">
<paragraph id="8B8436B5FE15370123C76D60FD96F8FA" blockId="19.[151,630,1886,1912]" box="[151,630,1886,1912]" pageId="19" pageNumber="420">
<heading id="D0CC81D9FE15370123C76D60FD96F8FA" bold="true" box="[151,630,1886,1912]" fontSize="11" level="1" pageId="19" pageNumber="420" reason="1">
<emphasis id="B94FEAA7FE15370123C76D60FD96F8FA" bold="true" box="[151,630,1886,1912]" pageId="19" pageNumber="420">
Genus
<taxonomicName id="4C3B4D36FE15370123BE6D60FD96F8FA" ID-CoL="327F" authority="Handlirsch, 1910" authorityName="Handlirsch" authorityYear="1910" box="[238,630,1886,1912]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE15370123BE6D60FE79F8FA" bold="true" box="[238,409,1886,1912]" italics="true" pageId="19" pageNumber="420">Archiraphidia</emphasis>
<bibRefCitation id="EFAA4B44FE15370122F06D60FD96F8FA" author="Handlirsch" box="[416,630,1886,1912]" pageId="19" pageNumber="420" refString="Handlirsch, A. (1910) Canadian fossil insects. 5. Insects from the Tertiary lake deposits of the Southern Interior of British Columbia collected by Mr. Lawrence Lambe, in 1906. In: Contributions to Canadian Palaeontology. Vol. II. Part III. Memoirs of the Geological Survey of Canada, 12 - P, pp. 93 - 129." type="book chapter" year="1910">Handlirsch, 1910</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C321653EFE15370123C76DA0FCAAF870" pageId="19" pageNumber="420" type="reference_group">
<paragraph id="8B8436B5FE15370123C76DA0FCAAF870" blockId="19.[151,1436,1950,2034]" pageId="19" pageNumber="420">
<treatmentCitationGroup id="AB2B119BFE15370123C76DA0FCAAF870" pageId="19" pageNumber="420">
<taxonomicName id="4C3B4D36FE15370123C76DA0FDF8F836" ID-CoL="327F" authority="Handlirsch, 1910: 103" authorityName="Handlirsch" authorityPageNumber="103" authorityYear="1910" box="[151,536,1950,1972]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE15370123C76DA0FEC6F836" box="[151,294,1950,1972]" italics="true" pageId="19" pageNumber="420">Archiraphidia</emphasis>
<treatmentCitation id="0A9A10A4FE153701227E6DA0FDF8F836" author="Handlirsch" box="[302,536,1950,1972]" page="103" pageId="19" pageNumber="420" year="1910">
<bibRefCitation id="EFAA4B44FE153701227E6DA0FE02F836" author="Handlirsch" box="[302,482,1950,1972]" pageId="19" pageNumber="420" refString="Handlirsch, A. (1910) Canadian fossil insects. 5. Insects from the Tertiary lake deposits of the Southern Interior of British Columbia collected by Mr. Lawrence Lambe, in 1906. In: Contributions to Canadian Palaeontology. Vol. II. Part III. Memoirs of the Geological Survey of Canada, 12 - P, pp. 93 - 129." type="book chapter" year="1910">Handlirsch, 1910</bibRefCitation>
: 103
</treatmentCitation>
</taxonomicName>
;
<bibRefCitation id="EFAA4B44FE15370121746DA0FD2FF836" author="Handlirsch" box="[548,719,1950,1972]" pageId="19" pageNumber="420" refString="Handlirsch, A. (1920 - 1921) Siebentes Kapitel. Palaeontologie. In: Schroder, C. (Ed.), Handbuch der Entomologie. 3. Band. Gustav Fischer, Jena, pp. 17 - 306. [issued in 1920 (pp. 113 - 208), 1921 (pp. 209 - 306)]." type="journal article" year="1920">
Handlirsch
<date id="FF851075FE15370121CC6DA0FD2FF836" box="[668,719,1950,1972]" pageId="19" pageNumber="420" value="1920" valueMax="1921">1920</date>
</bibRefCitation>
–1921 [1921]: 255;
<treatmentCitation id="0A9A10A4FE15370120CF6DA0FB3DF836" author="Cockerell" box="[927,1245,1950,1972]" page="295" pageId="19" pageNumber="420" year="1925">
<bibRefCitation id="EFAA4B44FE15370120CF6DA0FB48F836" author="Cockerell" box="[927,1192,1950,1972]" pageId="19" pageNumber="420" refString="Cockerell, T. D. A. &amp; Custer, C. (1925) A new fossil Inocellia (Neuroptera) from Florissant. The Entomologist, 58, 295 - 297." type="journal article" year="1925">Cockerell &amp; Custer 1925</bibRefCitation>
: 295
</treatmentCitation>
;
<treatmentCitation id="0A9A10A4FE15370127BA6DA1FF10F851" author="Martynov" page="244" pageId="19" pageNumber="420" year="1925">
<bibRefCitation id="EFAA4B44FE15370127BA6DA1FA78F836" author="Martynov" box="[1258,1432,1950,1972]" pageId="19" pageNumber="420" refString="Martynov, A. V. (1925 a) To the knowledge of fossil insects from Jurassic beds in Turkestan. 1. Raphidioptera. Izvestia Rossiiskoi Akademii Nauk, (VI), 19, 233 - 246." type="journal article" year="1925" yearSuffix="a">Martynov 1925a</bibRefCitation>
: 244
</treatmentCitation>
;
<treatmentCitation id="0A9A10A4FE15370123AC6D83FE33F851" author="Carpenter" box="[252,467,1981,2003]" page="145" pageId="19" pageNumber="420" year="1936">
<bibRefCitation id="EFAA4B44FE15370123AC6D83FE7EF851" author="Carpenter" box="[252,414,1981,2003]" pageId="19" pageNumber="420" refString="Carpenter, F. M. (1936) Revision of the Nearctic Raphidiodea (recent and fossil). Proceedings of the American Academy of Arts and Science, 71, 89 - 157, 2 pls." type="journal article" year="1936">Carpenter 1936</bibRefCitation>
: 145
</treatmentCitation>
;
<treatmentCitation id="0A9A10A4FE153701228F6D80FD79F851" author="Metzger" box="[479,665,1981,2003]" page="41" pageId="19" pageNumber="420" year="1960">
<bibRefCitation id="EFAA4B44FE153701228F6D80FD91F851" author="Metzger" box="[479,625,1981,2003]" pageId="19" pageNumber="420" refString="Metzger, R. (1960) Die Kamelhalsfliegen (Neuroptera, Raphidiidae). Die Neue Brehm-Bucherei. Bd. 254 A. Ziemsen Verlag, Wittenberg Lutheerstadt, 47 pp." type="book" year="1960">Metzger 1960</bibRefCitation>
: 41
</treatmentCitation>
;
<treatmentCitation id="0A9A10A4FE15370121F66D83FC83F851" author="Oswald" box="[678,867,1981,2003]" page="160" pageId="19" pageNumber="420" year="1990">
<bibRefCitation id="EFAA4B44FE15370121F66D83FCD1F851" author="Oswald" box="[678,817,1981,2003]" pageId="19" pageNumber="420" refString="Oswald, J. D. (1990) Raphidioptera. In: Grimaldi, D. A. (Ed.), Insects from the Santana Formation, Lower Cretaceous, of Brazil. Bulletin of the American Museum of Natural History, 195, pp. 154 - 163." type="journal article" year="1990">Oswald 1990</bibRefCitation>
: 160
</treatmentCitation>
(as a synonym of
<taxonomicName id="4C3B4D36FE15370127776D83FB64F851" ID-CoL="775X" box="[1063,1156,1981,2003]" class="Insecta" family="Raphidiidae" genus="Raphidia" kingdom="Animalia" order="Raphidioptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE15370127776D83FB64F851" box="[1063,1156,1981,2003]" italics="true" pageId="19" pageNumber="420">Raphidia</emphasis>
</taxonomicName>
);
<treatmentCitation id="0A9A10A4FE15370127CB6D83FA78F851" author="Aspock" box="[1179,1432,1981,2003]" page="536" pageId="19" pageNumber="420" year="1991">
<bibRefCitation id="EFAA4B44FE15370127CB6D83FA84F851" author="Aspock" box="[1179,1380,1981,2003]" pageId="19" pageNumber="420" refString="Aspock, H., Aspock, U. &amp; Rausch, H. (1991) Die Raphidiopteren der Erde. Eine monographische Darstellung der Systematik, Taxonomie, Biologie, Okologie und Chorologie der rezenten Raphidiopteren der Erde, mit einer zusammenfassenden Ubersicht der fossilen Raphidiopteren (Insecta: Neuropteroidea). 2 Vols. Goecke &amp; Evers, Krefeld, 730 &amp; 550 pp." type="book" year="1991">
Aspöck
<emphasis id="B94FEAA7FE15370127A16D80FAC6F851" box="[1265,1318,1981,2003]" italics="true" pageId="19" pageNumber="420">et al.</emphasis>
1991
</bibRefCitation>
: 536
</treatmentCitation>
, 538, 665;
<treatmentCitation id="0A9A10A4FE153701227D6DE2FE27F870" author="Engel" box="[301,455,2012,2034]" page="21" pageId="19" pageNumber="420" year="2002">
<bibRefCitation id="EFAA4B44FE153701227D6DE2FE44F870" author="Engel" box="[301,420,2012,2034]" pageId="19" pageNumber="420" refString="Engel, M. S. (2002) The smallest snakefly (Raphidioptera: Mesoraphidiidae): a new species in Cretaceous amber from Myanmar, with a catalog of fossil snakeflies. American Museum Novitates, 3363, 1 - 22. http: // dx. doi. org / 10.1206 / 0003 - 0082 (2002) 363 &lt;0001: tssrma&gt; 2.0. co; 2" type="journal article" year="2002">Engel 2002</bibRefCitation>
: 21
</treatmentCitation>
(as a tentative synonym of
<taxonomicName id="4C3B4D36FE153701218F6DE2FCDCF870" ID-CoL="775X" box="[735,828,2012,2034]" class="Insecta" family="Raphidiidae" genus="Raphidia" kingdom="Animalia" order="Raphidioptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE153701218F6DE2FCDCF870" box="[735,828,2012,2034]" italics="true" pageId="19" pageNumber="420">Raphidia</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C321653EFE12370623C76AA9FBE7FF32" box="[151,1031,151,176]" pageId="20" pageNumber="421" type="materials_examined">
<paragraph id="8B8436B5FE12370623C76AA9FBE7FF32" blockId="20.[151,1436,151,1157]" box="[151,1031,151,176]" pageId="20" pageNumber="421">
<emphasis id="B94FEAA7FE12370623C76AA9FED2FF32" bold="true" box="[151,306,151,176]" pageId="20" pageNumber="421">
<typeStatus id="54808817FE12370623C76AA9FF32FF32" box="[151,210,151,176]" pageId="20" pageNumber="421">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C3B4D36FE12370622696AA9FD51FF32" authority="Scudder, 1890" authorityName="Scudder" authorityYear="1890" box="[313,689,151,176]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="tumulata">
<emphasis id="B94FEAA7FE12370622696AA9FDE7FF32" box="[313,519,151,176]" italics="true" pageId="20" pageNumber="421">Inocellia tumulata</emphasis>
<bibRefCitation id="EFAA4B44FE123706215E6AA9FD51FF32" author="Scudder" box="[526,689,151,176]" pageId="20" pageNumber="421" refString="Scudder, S. H. (1890) The Tertiary Insects of North America. Report of the United States Geological Survey of the Territories, 13, 1 - 734, 28 pls." type="journal article" year="1890">Scudder, 1890</bibRefCitation>
</taxonomicName>
, by monotypy (
<bibRefCitation id="EFAA4B44FE12370620306AA9FC1AFF32" author="Oswald" box="[864,1018,151,176]" pageId="20" pageNumber="421" refString="Oswald, J. D. (1990) Raphidioptera. In: Grimaldi, D. A. (Ed.), Insects from the Santana Formation, Lower Cretaceous, of Brazil. Bulletin of the American Museum of Natural History, 195, pp. 154 - 163." type="journal article" year="1990">Oswald 1990</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C321653EFE12370623976A82FC47FE75" pageId="20" pageNumber="421" type="diagnosis">
<paragraph id="8B8436B5FE12370623976A82FC56FE2D" blockId="20.[151,1436,151,1157]" pageId="20" pageNumber="421">
<emphasis id="B94FEAA7FE12370623976A82FEA1FF57" bold="true" box="[199,321,188,213]" pageId="20" pageNumber="421">Diagnosis.</emphasis>
Very small raphidiids (forewing &lt;
<quantity id="4CC39B50FE12370621BC6A82FCA3FF56" box="[748,835,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="421" unit="mm" value="10.0">10 mm</quantity>
long). Both wings: relatively short ScP [long in
<taxonomicName id="4C3B4D36FE12370623C76AE1FED7FF7A" box="[151,311,223,248]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE12370623C76AE1FED7FF7A" box="[151,311,223,248]" italics="true" pageId="20" pageNumber="421">Megaraphidia</emphasis>
</taxonomicName>
]; short, dark pterostigma in which incorporated branch of RA not or poorly visible [incorporated branch of RA clearly visible in
<taxonomicName id="4C3B4D36FE12370621436B3AFD53FE9F" box="[531,691,260,285]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE12370621436B3AFD53FE9F" box="[531,691,260,285]" italics="true" pageId="20" pageNumber="421">Megaraphidia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3B4D36FE12370621956B3AFC47FE9F" box="[709,935,260,285]" class="Insecta" family="Raphidiidae" genus="Florissantoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94FEAA7FE12370621956B3AFC47FE9F" box="[709,935,260,285]" italics="true" pageId="20" pageNumber="421">Florissantoraphidia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57DCFE12370620E26B3BFBC2FE9E" box="[946,1058,261,285]" pageId="20" pageNumber="421" rank="genus">gen. nov.</taxonomicNameLabel>
]. Forewing: additional crossvein 2b scp-r sometimes present [not detected in
<taxonomicName id="4C3B4D36FE123706212D6B14FCFDFEC1" box="[637,797,298,323]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE123706212D6B14FCFDFEC1" box="[637,797,298,323]" italics="true" pageId="20" pageNumber="421">Megaraphidia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3B4D36FE123706207C6B14FBEEFEC1" box="[812,1038,298,323]" class="Insecta" family="Raphidiidae" genus="Florissantoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94FEAA7FE123706207C6B14FBEEFEC1" box="[812,1038,298,323]" italics="true" pageId="20" pageNumber="421">Florissantoraphidia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57DCFE12370627466B15FB62FEC0" box="[1046,1154,299,323]" pageId="20" pageNumber="421" rank="genus">gen. nov.</taxonomicNameLabel>
]; MP strongly zigzagged [moderately zigzagged in
<taxonomicName id="4C3B4D36FE12370622906B70FD80FEE5" box="[448,608,334,359]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE12370622906B70FD80FEE5" box="[448,608,334,359]" italics="true" pageId="20" pageNumber="421">Megaraphidia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3B4D36FE12370621206B70FCB2FEE5" box="[624,850,334,359]" class="Insecta" family="Raphidiidae" genus="Florissantoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94FEAA7FE12370621206B70FCB2FEE5" box="[624,850,334,359]" italics="true" pageId="20" pageNumber="421">Florissantoraphidia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57DCFE123706200B6B6EFC28FEE5" box="[859,968,336,360]" pageId="20" pageNumber="421" rank="genus">gen. nov.</taxonomicNameLabel>
]; branches of CuA, MP short, not longer then width of
<emphasis id="B94FEAA7FE123706226C6B4CFE8DFE09" box="[316,365,370,395]" italics="true" pageId="20" pageNumber="421">2doi</emphasis>
[distinctly longer than width of
<emphasis id="B94FEAA7FE12370621B96B4CFCFCFE09" box="[745,796,370,395]" italics="true" pageId="20" pageNumber="421">2doi</emphasis>
in
<taxonomicName id="4C3B4D36FE12370620146B4CFC04FE09" box="[836,996,370,395]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE12370620146B4CFC04FE09" box="[836,996,370,395]" italics="true" pageId="20" pageNumber="421">Megaraphidia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3B4D36FE12370620A66B4CFB38FE09" box="[1014,1240,370,395]" class="Insecta" family="Raphidiidae" genus="Florissantoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94FEAA7FE12370620A66B4CFB38FE09" box="[1014,1240,370,395]" italics="true" pageId="20" pageNumber="421">Florissantoraphidia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57DCFE12370627B16B4DFAAFFE08" box="[1249,1359,371,395]" pageId="20" pageNumber="421" rank="genus">gen. nov.</taxonomicNameLabel>
].
<collectingCountry id="F32C7625FE12370626346B4CFA7CFE09" box="[1380,1436,370,395]" name="India" pageId="20" pageNumber="421">Hind</collectingCountry>
wing:
<emphasis id="B94FEAA7FE123706238D6BA8FEE1FE2D" box="[221,257,406,431]" italics="true" pageId="20" pageNumber="421">doi</emphasis>
single [two in
<taxonomicName id="4C3B4D36FE12370622F96BA8FDA9FE2D" box="[425,585,406,431]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE12370622F96BA8FDA9FE2D" box="[425,585,406,431]" italics="true" pageId="20" pageNumber="421">Megaraphidia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3B4D36FE12370621076BA8FCD9FE2D" box="[599,825,406,431]" class="Insecta" family="Raphidiidae" genus="Florissantoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94FEAA7FE12370621076BA8FCD9FE2D" box="[599,825,406,431]" italics="true" pageId="20" pageNumber="421">Florissantoraphidia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57DCFE12370620106BA6FC4BFE2D" box="[832,939,408,432]" pageId="20" pageNumber="421" rank="genus">gen. nov.</taxonomicNameLabel>
].
</paragraph>
<paragraph id="8B8436B5FE12370623976B84FC47FE75" blockId="20.[151,1436,151,1157]" pageId="20" pageNumber="421">
<emphasis id="B94FEAA7FE12370623976B84FE75FE51" bold="true" box="[199,405,442,467]" pageId="20" pageNumber="421">Species included.</emphasis>
Three species from Florissant,
<taxonomicName id="4C3B4D36FE12370620506B84FBEAFE51" box="[768,1034,442,467]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="tumulata">
<emphasis id="B94FEAA7FE12370620506B84FBEAFE51" box="[768,1034,442,467]" italics="true" pageId="20" pageNumber="421">Archiraphidia tumulata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3B4D36FE123706274B6B85FA8AFE51" authority="Scudder, 1890" authorityName="Scudder" authorityYear="1890" box="[1051,1386,442,467]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="tranquilla">
<emphasis id="B94FEAA7FE123706274B6B85FB4CFE51" box="[1051,1196,442,467]" italics="true" pageId="20" pageNumber="421">A. tranquilla</emphasis>
(
<bibRefCitation id="EFAA4B44FE12370627EC6B84FA82FE51" author="Scudder" box="[1212,1378,442,467]" pageId="20" pageNumber="421" refString="Scudder, S. H. (1890) The Tertiary Insects of North America. Report of the United States Geological Survey of the Territories, 13, 1 - 734, 28 pls." type="journal article" year="1890">Scudder, 1890</bibRefCitation>
)
</taxonomicName>
and
<emphasis id="B94FEAA7FE12370623C76BE1FF5DFE74" box="[151,189,479,502]" italics="true" pageId="20" pageNumber="421">A.?</emphasis>
<taxonomicName id="4C3B4D36FE12370623946BE0FDE2FE75" authority="(Scudder, 1890)" authorityName="Scudder" authorityYear="1890" baseAuthorityName="Scudder" baseAuthorityYear="1890" box="[196,514,478,503]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="somnolenta">
<emphasis id="B94FEAA7FE12370623946BE0FEA5FE75" box="[196,325,478,503]" italics="true" pageId="20" pageNumber="421">somnolenta</emphasis>
(
<bibRefCitation id="EFAA4B44FE12370622036BE0FE1AFE75" author="Scudder" box="[339,506,478,503]" pageId="20" pageNumber="421" refString="Scudder, S. H. (1890) The Tertiary Insects of North America. Report of the United States Geological Survey of the Territories, 13, 1 - 734, 28 pls." type="journal article" year="1890">Scudder, 1890</bibRefCitation>
)
</taxonomicName>
. The latter assignment is preliminary.
</paragraph>
</subSubSection>
<subSubSection id="C321653EFE1237062397683CFB2DFB07" pageId="20" pageNumber="421" type="discussion">
<paragraph id="8B8436B5FE1237062397683CFCBAFD29" blockId="20.[151,1436,151,1157]" pageId="20" pageNumber="421">
<emphasis id="B94FEAA7FE1237062397683CFEDBFD99" bold="true" box="[199,315,514,539]" pageId="20" pageNumber="421">Remarks.</emphasis>
This genus is characterized by its pterostigma appearing very dark (
<figureCitation id="13002A30FE1237062702683CFB79FD99" box="[1106,1177,514,539]" captionStart="FIGURE 9" captionStartId="19.[151,249,1674,1696]" captionTargetBox="[151,1436,519,1662]" captionTargetId="figure@19.[151,1436,519,1662]" captionTargetPageId="19" captionText="FIGURE 9. Pterostigma of Archiraphidia species. A – C, A. tumulata (Scudder, 1890): A, left wings. B, right forewing. C, right hind wing. D – F, A. tranquilla (Scudder, 1890): D, four wings of the holotype MCZ 446. E, four wings of MCZ 4137 (lectotype of A. eventa (Scudder, 1890 )); F, left wings of MCZ 253 (paralectotype of A. eventa). G, three wings of A.? somnolenta (Scudder, 1890). H, right forewing of A.? somnolenta. HW, hind wing; LFW, left forewing; RFW, right forewing. All wings are shown with apex to right and not to scale." httpUri="https://zenodo.org/record/252227/files/figure.png" pageId="20" pageNumber="421">Fig. 9</figureCitation>
), and the incorporated pterostigmal RA branch being hardly detectable. The paler pterostigma in some wings, however, is probably due to poor preservation of pigmentation (
<figureCitation id="13002A30FE12370621676874FD63FDE1" box="[567,643,586,611]" captionStart="FIGURE 9" captionStartId="19.[151,249,1674,1696]" captionTargetBox="[151,1436,519,1662]" captionTargetId="figure@19.[151,1436,519,1662]" captionTargetPageId="19" captionText="FIGURE 9. Pterostigma of Archiraphidia species. A – C, A. tumulata (Scudder, 1890): A, left wings. B, right forewing. C, right hind wing. D – F, A. tranquilla (Scudder, 1890): D, four wings of the holotype MCZ 446. E, four wings of MCZ 4137 (lectotype of A. eventa (Scudder, 1890 )); F, left wings of MCZ 253 (paralectotype of A. eventa). G, three wings of A.? somnolenta (Scudder, 1890). H, right forewing of A.? somnolenta. HW, hind wing; LFW, left forewing; RFW, right forewing. All wings are shown with apex to right and not to scale." httpUri="https://zenodo.org/record/252227/files/figure.png" pageId="20" pageNumber="421">Figs 9</figureCitation>
B, F). A branch of RA incorporated within the pterostigma is only detected in one wing—and then only poorly visible—however, we assume that is likely present in all of these, although await future specimens to strengthen this supposition.
</paragraph>
<paragraph id="8B8436B5FE12370623976888FB2DFB07" blockId="20.[151,1436,151,1157]" pageId="20" pageNumber="421">
The minute size and venation of this genus are most similar to those of the genus
<taxonomicName id="4C3B4D36FE12370627306888FB0DFD4D" box="[1120,1261,694,719]" class="Insecta" family="Raphidiidae" genus="Harraphidia" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE12370627306888FB0DFD4D" box="[1120,1261,694,719]" italics="true" pageId="20" pageNumber="421">Harraphidia</emphasis>
</taxonomicName>
(forewing
<quantity id="4CC39B50FE123706263B6888FF0DFD70" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.5" metricValueMin="5.5" pageId="20" pageNumber="421" unit="mm" value="7.0" valueMax="8.5" valueMin="5.5">5.5– 8.5 mm</quantity>
long; see
<bibRefCitation id="EFAA4B44FE123706220F68E5FD4FFD71" author="Monserrat" box="[351,687,730,755]" pageId="20" pageNumber="421" refString="Monserrat, V. J. &amp; Papenberg, D. (2006) Revision del genero Harraphidia Steinmann, 1963 con la descripcion de dos nuevas especies de la peninsula Iberica y de Marruecos (Insecta, Raphidioptera). Graellsia, 62, 203 - 222. http: // dx. doi. org / 10.3989 / graellsia. 2006. v 62. i 2.67" type="journal article" year="2006">
<collectingCountry id="F32C7625FE123706220F68E5FE32FD70" box="[351,466,731,754]" name="Montserrat" pageId="20" pageNumber="421">Monserrat</collectingCountry>
&amp; Papenberg 2006
</bibRefCitation>
). They share in particular a relatively short ScP; short pterostigma; 3ra-rp located within pterostigma; the same configuration of RA distad pterostigma in both wings; the presence of additional crossvein 2b scp-r; short branches of CuA and MP in the forewing; one
<emphasis id="B94FEAA7FE123706276E691BFB81FCBC" box="[1086,1121,805,830]" italics="true" pageId="20" pageNumber="421">doi</emphasis>
in the hind wing. In extant
<taxonomicName id="4C3B4D36FE12370623C76977FEC1FCE0" box="[151,289,841,866]" class="Insecta" family="Raphidiidae" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="family">Raphidiidae</taxonomicName>
, the single
<emphasis id="B94FEAA7FE12370622F26977FE26FCE0" box="[418,454,841,866]" italics="true" pageId="20" pageNumber="421">doi</emphasis>
is a generic feature only of
<taxonomicName id="4C3B4D36FE12370620556977FC72FCE0" box="[773,914,841,866]" class="Insecta" family="Raphidiidae" genus="Harraphidia" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE12370620556977FC72FCE0" box="[773,914,841,866]" italics="true" pageId="20" pageNumber="421">Harraphidia</emphasis>
</taxonomicName>
, although this condition is known, rarely, in a few other species, where it is considered anomalous (see above). The additional crossvein 2b scp-r is also characteristic of only this genus (in extant Raphiididae).
<taxonomicName id="4C3B4D36FE123706203169AAFC1EFC2F" box="[865,1022,916,941]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE123706203169AAFC1EFC2F" box="[865,1022,916,941]" italics="true" pageId="20" pageNumber="421">Archiraphidia</emphasis>
</taxonomicName>
may only be distinguished from
<taxonomicName id="4C3B4D36FE12370623C76986FEC5FC53" box="[151,293,952,977]" class="Insecta" family="Raphidiidae" genus="Harraphidia" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE12370623C76986FEC5FC53" box="[151,293,952,977]" italics="true" pageId="20" pageNumber="421">Harraphidia</emphasis>
</taxonomicName>
in that MP+CuA never has the long stem characteristic of the latter. These genera are undoubtedly closely related, but the occurrence of this condition (i.e., MP+CuA) and the poorly preserved male terminalia in specimens of
<taxonomicName id="4C3B4D36FE12370622116E3EFE3FFB9B" box="[321,479,1024,1049]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE12370622116E3EFE3FFB9B" box="[321,479,1024,1049]" italics="true" pageId="20" pageNumber="421">Archiraphidia</emphasis>
</taxonomicName>
(the senior name) prevent their synonymization. Moreover, although the male terminalia in these genera are rather similar in their general appearance, the ectoproct of all
<taxonomicName id="4C3B4D36FE12370627CC6E1AFACAFBBF" box="[1180,1322,1060,1085]" class="Insecta" family="Raphidiidae" genus="Harraphidia" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE12370627CC6E1AFACAFBBF" box="[1180,1322,1060,1085]" italics="true" pageId="20" pageNumber="421">Harraphidia</emphasis>
</taxonomicName>
species is clearly caudally elongate, whereas that of
<taxonomicName id="4C3B4D36FE12370621D76E76FCC5FBE3" box="[647,805,1096,1121]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE12370621D76E76FCC5FBE3" box="[647,805,1096,1121]" italics="true" pageId="20" pageNumber="421">Archiraphidia</emphasis>
</taxonomicName>
clearly not (
<figureCitation id="13002A30FE12370620EE6E76FBF6FBE3" box="[958,1046,1096,1121]" captionStart="FIGURE 15" captionStartId="26.[151,249,1459,1481]" captionTargetBox="[151,1436,481,1437]" captionTargetId="figure@26.[151,1436,481,1437]" captionTargetPageId="26" captionText="FIGURE 15. Archiraphidia tranquilla (Scudder, 1890) (lectotype of A. eventa (Scudder, 1890), MCZ 4137). Apex of abdomen (lateral view). gp 9, gonapophyses of segment 9 (hypovalva); gs 9, gonostyli of gonocoxites 9; gx 9, gonocoxites of segment 9; S 8, S 9, sternites 8 and 9; T 8, T 9, T 10, tergites 8 to 10. Scale bar = 400 µm." httpUri="https://zenodo.org/record/252233/files/figure.png" pageId="20" pageNumber="421">Fig. 15</figureCitation>
). Also, 1r-m in the hind wing of
<taxonomicName id="4C3B4D36FE12370623C76E52FEC5FB07" box="[151,293,1132,1157]" class="Insecta" family="Raphidiidae" genus="Harraphidia" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE12370623C76E52FEC5FB07" box="[151,293,1132,1157]" italics="true" pageId="20" pageNumber="421">Harraphidia</emphasis>
</taxonomicName>
is crossvein-like, but this is not detected in
<taxonomicName id="4C3B4D36FE12370620406E52FC4EFB07" box="[784,942,1132,1157]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA7FE12370620406E52FC4EFB07" box="[784,942,1132,1157]" italics="true" pageId="20" pageNumber="421">Archiraphidia</emphasis>
</taxonomicName>
due to poor preservation.
</paragraph>
</subSubSection>
</treatment>
</document>
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