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<mods:title id="B5BBA81B6905DFF6F2F2D3E49C1F0A47">An annotated checklist of the chondrichthyans of South Africa</mods:title>
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<paragraph id="8B8436B7953DFFDACFF75CE7FDF3276B" blockId="84.[151,1437,1770,2013]" box="[151,522,1770,1797]" pageId="84" pageNumber="85">
<heading id="D0CC81DB953DFFDACFF75CE7FDF3276B" bold="true" box="[151,522,1770,1797]" fontSize="11" level="2" pageId="84" pageNumber="85" reason="4">
<taxonomicName id="4C3B4D34953DFFDACFF75CE7FDF3276B" ID-CoL="4RB3J" ID-ENA="1003795" authority="(Hulley, 1970)" baseAuthorityName="Hulley" baseAuthorityYear="1970" box="[151,522,1770,1797]" class="Elasmobranchii" family="Rajidae" genus="Rajella" kingdom="Animalia" order="Rajiformes" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="dissimilis">
<emphasis id="B94FEAA5953DFFDACFF75CE7FDF3276B" bold="true" box="[151,522,1770,1797]" pageId="84" pageNumber="85">
<emphasis id="B94FEAA5953DFFDACFF75CE7FEA5276A" bold="true" box="[151,348,1770,1797]" italics="true" pageId="84" pageNumber="85">Rajella dissimilis</emphasis>
(
<bibRefCitation id="EFAA4B46953DFFDACE0B5CE7FDF8276B" author="Hulley" box="[363,513,1770,1796]" firstAuthor="Hulley" pageId="84" pageNumber="85" pagination="151 - 220" refId="ref104963" refString="Hulley, P. A. (1970) An investigation of the Rajidae of the west and south coasts of southern Africa. Annals of the South African Museum, 55, 151 - 220." type="journal article" year="1970">Hulley, 1970</bibRefCitation>
)
</emphasis>
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</heading>
</paragraph>
<paragraph id="8B8436B7953DFFDACFF75D03FEE62747" blockId="84.[151,1437,1770,2013]" box="[151,287,1806,1832]" pageId="84" pageNumber="85">Ghost Skate</paragraph>
</subSubSection>
<subSubSection id="C321653C953DFFDBCFF75D3EFC632096" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85" type="reference_group">
<paragraph id="8B8436B7953DFFDACFF75D3EFB96271F" blockId="84.[151,1437,1770,2013]" pageId="84" pageNumber="85">
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<treatmentCitation id="0A9A10A6953DFFDACFF75D3EFDE52723" author="Hulley, P. A." box="[151,540,1842,1868]" page="199" pageId="84" pageNumber="85" year="1970">
<taxonomicName id="4C3B4D34953DFFDACFF75D3EFDE52723" ID-CoL="793N9" authority="Hulley, 1970: 199" authorityName=": Hulley" authorityPageNumber="199" authorityYear="1970" box="[151,540,1842,1868]" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="dissimilis">
<emphasis id="B94FEAA5953DFFDACFF75D3EFEB92723" box="[151,320,1842,1868]" italics="true" pageId="84" pageNumber="85">Raja dissimilis</emphasis>
<materialsCitation id="3B533CEA953DFFDACE2A5D3FFDE52723" ID-GBIF-Occurrence="3072574578" accessionNumber="ZMH 25258" box="[330,540,1842,1868]" collectionCode="ZMH" country="South Africa" county="southeastern Atlantic." latitude="-33.783333" location="West" longLatPrecision="1203" longitude="17.233334" municipality="Town" pageId="84" pageNumber="85" specimenCode="ISH 46-1967" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
<bibRefCitation id="EFAA4B46953DFFDACE2A5D3FFDE52723" author="Hulley" box="[330,540,1842,1868]" firstAuthor="Hulley" page="199" pageId="84" pageNumber="85" pagination="151 - 220" refId="ref104963" refString="Hulley, P. A. (1970) An investigation of the Rajidae of the west and south coasts of southern Africa. Annals of the South African Museum, 55, 151 - 220." type="journal article" year="1970">Hulley, 1970: 199</bibRefCitation>
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, figs. 15, pl. 10 (figs. ab).
<typeStatus id="54808815953DFFDACC095D3FFC2F2723" box="[873,982,1842,1868]" pageId="84" pageNumber="85" type="holotype">Holotype</typeStatus>
: ZMH 25258 [ex
<specimenCode id="DB9D9ECC953DFFDACBCA5D3FFABD2722" box="[1194,1348,1842,1869]" collectionCode="ISH" country="Germany" httpUri="http://grbio.org/cool/9fk0-a162" name="Institut fuer Seefischerei" pageId="84" pageNumber="85">ISH 46-1967</specimenCode>
a]. Type locality:
<location id="8EE4606C953DFFDACE485D5AFEA6271E" LSID="urn:lsid:plazi:treatment:039287A1953DFFDBCFF75CE7FDE921BE:8EE4606C953DFFDACE485D5AFEA6271E" box="[296,351,1879,1905]" country="South Africa" county="southeastern Atlantic." latitude="-33.783333" longLatPrecision="1203" longitude="17.233334" municipality="Town" name="West" pageId="84" pageNumber="85" stateProvince="Western Cape">West</location>
of
<collectingRegion id="49FFF855953DFFDACEE55D5BFE46271F" box="[389,447,1878,1904]" country="South Africa" name="Western Cape" pageId="84" pageNumber="85">Cape</collectingRegion>
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,
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,
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,
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,
<collectingCounty id="62E54E3B953DFFDACC1C5D5BFB96271F" box="[892,1135,1878,1904]" pageId="84" pageNumber="85">southeastern Atlantic.</collectingCounty>
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<paragraph id="8B8436B7953DFFDBCFF75D77FC632096" blockId="84.[151,1437,1770,2013]" lastBlockId="85.[151,1437,150,465]" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85">
<emphasis id="B94FEAA5953DFFDACFF75D77FEA327FB" bold="true" box="[151,346,1914,1940]" pageId="84" pageNumber="85">Local synonymy</emphasis>
:
<taxonomicName id="4C3B4D34953DFFDACE095D76FD1127FB" ID-CoL="793N9" authority=": Hulley, 1970: 199" authorityName=": Hulley" authorityPageNumber="199" authorityYear="1970" box="[361,744,1914,1941]" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="dissimilis">
<emphasis id="B94FEAA5953DFFDACE095D76FDF727FB" box="[361,526,1914,1940]" italics="true" pageId="84" pageNumber="85">Raja dissimilis</emphasis>
:
<bibRefCitation id="EFAA4B46953DFFDACD7D5D77FD1127FB" author="Hulley" box="[541,744,1914,1941]" firstAuthor="Hulley" page="199" pageId="84" pageNumber="85" pagination="151 - 220" refId="ref104963" refString="Hulley, P. A. (1970) An investigation of the Rajidae of the west and south coasts of southern Africa. Annals of the South African Museum, 55, 151 - 220." type="journal article" year="1970">Hulley, 1970: 199</bibRefCitation>
</taxonomicName>
, figs. 15, pl. 10;
<bibRefCitation id="EFAA4B46953DFFDACCC35D77FB9527FB" author="Hulley" box="[931,1132,1914,1940]" firstAuthor="Hulley" page="86" pageId="84" pageNumber="85" pagination="1 - 103" refId="ref105041" refString="Hulley, P. A. (1972 a) The origin, interrelationships and distribution of southern African Rajidae (Chondrichthyes, Batoidei). Annals of the South African Museum, 60, 1 - 103. https: // doi. org / 10.2307 / 1442499" type="journal article" year="1972" yearSuffix="a">Hulley, 1972a: 86</bibRefCitation>
, figs. 5859;
<bibRefCitation id="EFAA4B46953DFFDACA645D77FF0B27D7" author="Hulley" firstAuthor="Hulley" page="121" pageId="84" pageNumber="85" pagination="115 - 128" refId="ref105183" refString="Hulley, P. A. (1986) Family Rajidae, Anacanthobatidae. In: Smith, M. M. &amp; Heemstra, P. C. (Eds.), Smith's Sea Fishes. Macmillan, Johannesburg, pp. 115 - 128." type="book chapter" year="1986">Hulley, 1986: 121</bibRefCitation>
, fig. 25.10;
<bibRefCitation id="EFAA4B46953DFFDACE165D93FDA127D7" author="Stehmann, M." box="[374,600,1950,1977]" pageId="84" pageNumber="85" pagination="1 - 119" refId="ref116339" refString="Stehmann, M. (1995) First and new records of skates (Chondrichthyes, Rajiformes, Rajidae) from the West African continental slope (Morocco to South Africa), with descriptions of two new species. Archiv f ̡ r Fischereiwissenschaft, 43 (1), 1 - 119." type="journal article" year="1995">Stehmann, 1995: 79</bibRefCitation>
.
<taxonomicName id="4C3B4D34953DFFDACD025D92FB5E27D7" ID-CoL="793N9" authority=": Compagno et al., 1989: 98" authorityName="Compagno" authorityPageNumber="98" authorityYear="1989" box="[610,1191,1950,1977]" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="dissimilis" subGenus="Rajella">
<emphasis id="B94FEAA5953DFFDACD025D92FD6E27D7" box="[610,663,1951,1976]" italics="true" pageId="84" pageNumber="85">Raja</emphasis>
(
<emphasis id="B94FEAA5953DFFDACDC65D93FD0E27D7" box="[678,759,1950,1976]" italics="true" pageId="84" pageNumber="85">Rajella</emphasis>
)
<emphasis id="B94FEAA5953DFFDACC675D93FC8827D7" box="[775,881,1950,1976]" italics="true" pageId="84" pageNumber="85">dissimilis</emphasis>
:
<bibRefCitation id="EFAA4B46953DFFDACC1E5D93FB5E27D7" author="Compagno" box="[894,1191,1950,1977]" etAl="et al." firstAuthor="Compagno" page="98" pageId="84" pageNumber="85" refId="ref96138" refString="Compagno, L. J. V., Ebert, D. A. &amp; Smale, M. J. (1989) Guide to the Sharks and Rays of Southern Africa. Struik Publishers, Cape Town, 158 pp." type="book" year="1989">
Compagno
<emphasis id="B94FEAA5953DFFDACC9F5D92FBC927D7" box="[1023,1072,1950,1976]" italics="true" pageId="84" pageNumber="85">et al</emphasis>
., 1989: 98
</bibRefCitation>
</taxonomicName>
, pl.;
<bibRefCitation id="EFAA4B46953DFFDACBBD5D93FEC027B3" author="Compagno" etAl="et al." firstAuthor="Compagno" page="104" pageId="84" pageNumber="85" pagination="43 - 139" refId="ref96066" refString="Compagno, L. J. V., Ebert, D. A. &amp; Cowley, P. D. (1991) Distribution of offshore demersal cartilaginous fishes (class Chondrichthyes) of the west coast of southern Africa, with notes on their systematics. South African Journal of Marine Science, 11, 43 - 139. https: // doi. org / 10.2989 / 025776191784287664" type="journal article" year="1991">
Compagno
<emphasis id="B94FEAA5953DFFDACA3E5D92FA7727D7" box="[1374,1422,1950,1976]" italics="true" pageId="84" pageNumber="85">et al</emphasis>
., 1991: 104
</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953DFFDACE245DCFFDC827B2" author="Ebert" box="[324,561,1986,2013]" etAl="et al." firstAuthor="Ebert" page="77" pageId="84" pageNumber="85" pagination="71 - 81" refId="ref99172" refString="Ebert, D. A., Cowley, P. D. &amp; Compagno, L. J. V. (1991) A preliminary investigation of the feeding ecology of skates (Batoidea: Rajidae) off the west coast of southern Africa. South African Journal of Marine Science, 10, 71 - 81. https: // doi. org / 10.2989 / 02577619109504621" type="journal article" year="1991">
Ebert
<emphasis id="B94FEAA5953DFFDACEE75DCEFE4127B3" box="[391,440,1986,2012]" italics="true" pageId="84" pageNumber="85">et al</emphasis>
., 1991: 77
</bibRefCitation>
.
<taxonomicName id="4C3B4D34953DFFDACD5C5DCFFBFF27B3" ID-CoL="4RB3J" authority=": Compagno, 1999: 116" authorityName="Compagno" authorityPageNumber="116" authorityYear="1999" box="[572,1030,1986,2013]" class="Elasmobranchii" family="Rajidae" genus="Rajella" kingdom="Animalia" order="Rajiformes" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="dissimilis">
<emphasis id="B94FEAA5953DFFDACD5C5DCFFD0427B3" box="[572,765,1986,2012]" italics="true" pageId="84" pageNumber="85">Rajella dissimilis</emphasis>
:
<treatmentCitation id="0A9A10A6953DFFDACC6B5DCFFBFF27B3" box="[779,1030,1986,2012]" pageId="84" pageNumber="85">
<bibRefCitation id="EFAA4B46953DFFDACC6B5DCFFBFF27B3" author="Compagno" box="[779,1030,1986,2012]" firstAuthor="Compagno" page="116" pageId="84" pageNumber="85" pagination="75 - 120" refId="ref95720" refString="Compagno, L. J. V. (1999) An overview of chondrichthyan systematics and biodiversity in southern Africa. Transactions of the Royal Society of South Africa, 54, 75 - 120. https: // doi. org / 10.1080 / 00359199909520406" type="journal article" year="1999">Compagno, 1999: 116</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<bibRefCitation id="EFAA4B46953DFFDACB715DCFFA9227B2" author="Compagno &amp; Ebert" box="[1041,1387,1986,2013]" firstAuthor="Compagno" page="135" pageId="84" pageNumber="85" pagination="125 - 145" refId="ref96013" refString="Compagno, L. J. V. &amp; Ebert, D. A. (2007) Southern African skate biodiversity and distribution. Environmental Biology of Fishes, 80, 125 - 145. https: // doi. org / 10.1007 / s 10641 - 007 - 9243 - 4" type="journal article" year="2007">Compagno &amp; Ebert, 2007: 135</bibRefCitation>
, fig. 5b;
<bibRefCitation id="EFAA4B46953CFFDBCF915A9AFDB720DE" author="Ebert &amp; Compagno" box="[241,590,151,177]" firstAuthor="Ebert" page="121" pageId="85" pageNumber="86" pagination="125 - 145" refId="ref96013" refString="Compagno, L. J. V. &amp; Ebert, D. A. (2007) Southern African skate biodiversity and distribution. Environmental Biology of Fishes, 80, 125 - 145. https: // doi. org / 10.1007 / s 10641 - 007 - 9243 - 4" type="journal article" year="2007">Ebert &amp; Compagno, 2007: 121</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953CFFDBCD395A9AFCB120DE" author="Ebert" box="[601,840,150,177]" etAl="et al." firstAuthor="Ebert" page="95" pageId="85" pageNumber="86" pagination="81 - 102" refId="ref98910" refString="Ebert, D. A., Compagno, L. J. V. &amp; Cowley, P. D. (2008) Aspects on the reproductive biology of skates (Chondrichthyes: Rajiformes: Rajoidei) from southern African waters. ICES Journal of Marine Science, 65, 81 - 102. https: // doi. org / 10.1093 / icesjms / fsm 169" type="journal article" year="2008">
Ebert
<emphasis id="B94FEAA5953CFFDBCDFD5A9AFD3720DE" box="[669,718,151,177]" italics="true" pageId="85" pageNumber="86">et al</emphasis>
., 2008: 95
</bibRefCitation>
;
<bibRefCitation id="EFAA4B46953CFFDBCC345A9AFBFA20DE" author="Ebert D. A." box="[852,1027,150,177]" pageId="85" pageNumber="86" refId="ref98383" refString="Ebert D. A. (2014) Deep-sea cartilaginous fishes of the Indian Ocean. Vol. 2. Batoids and chimaeras. FAO Species Catalogue for Fishery Purposes No. 8. Vol. 2. FAO, Rome, 129 pp." type="book" year="2014">Ebert, 2014: 75</bibRefCitation>
, fig. 120;
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, fig. 211;
<bibRefCitation id="EFAA4B46953CFFDBCFA65AB6FDF420BA" author="Weigmann, S. &amp; Stehmann, M. F. W. &amp; Thiel, R." box="[198,525,187,213]" pageId="85" pageNumber="86" pagination="359 - 387" refId="ref118816" refString="Weigmann, S., Stehmann, M. F. W. &amp; Thiel, R. (2014 a) Rajella paucispinosa n. sp., a new deep-water skate (Elasmobranchii, Rajidae) from the western Indian Ocean off South Mozambique, and a revised generic diagnosis. Zootaxa, 3847 (3), 359 - 387. https: // doi. org / 10.11646 / zootaxa. 3847.3.3" type="journal article" year="2014">
Weigmann
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., 2014a: 384
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;
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;
<bibRefCitation id="EFAA4B46953CFFDBCC115AB6FBA720BA" author="Last" box="[881,1118,187,213]" etAl="et al." firstAuthor="Last" page="19" pageId="85" pageNumber="86" pagination="11 - 34" refId="ref107873" refString="Last, P. R., Weigmann, S. &amp; Yang, L. (2016 f) Changes to the nomenclature of the skates (Chondrichthyes: Rajiformes). In: Last, P. R. &amp; Yearsley, G. K. (Eds.), Rays of the World: Supplementary Information. CSIRO Australian National Fish Collection, Hobart, pp. 11 - 34." type="book chapter" year="2016" yearSuffix="f">
Last
<emphasis id="B94FEAA5953CFFDBCCC85AB6FC2320BA" box="[936,986,187,213]" italics="true" pageId="85" pageNumber="86">et al</emphasis>
., 2016f: 19
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;
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Last
<emphasis id="B94FEAA5953CFFDBCBC25AB6FB2D20BA" box="[1186,1236,187,213]" italics="true" pageId="85" pageNumber="86">et al</emphasis>
.,
<quantity id="4CC39B52953CFFDBCB895AB6FACA20BA" box="[1257,1331,187,213]" metricMagnitude="0" metricUnit="kg" metricValue="2.016" pageId="85" pageNumber="86" unit="g" value="2016.0">2016g</quantity>
: 339
</bibRefCitation>
, fig. 19.130; McEachran &amp;
<bibRefCitation id="EFAA4B46953CFFDBCEA75AD2FD692096" author="Seret, B." box="[455,656,223,249]" pageId="85" pageNumber="86" pagination="1404 - 1433" refId="ref113064" refString="Seret, B. (2016) Batoid fishes. Multiple families. In: Carpenter, K. E. &amp; De Angelis, N. (Eds.), The Living Marine Resources of the Eastern Central Atlantic. Vol. 2. Bivalves, Gastropods, Hagfishes, Sharks, Batoid Fishes, and Chimaeras. FAO Species Identification Guide for Fishery Purposes. FAO, Rome, pp. 1337 - 1369, 1404 - 1433." type="journal article" year="2016">Séret, 2016: 1402</bibRefCitation>
;
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.
</paragraph>
</subSubSection>
<subSubSection id="C321653C953CFFDBCFF75B0FFC00210A" pageId="85" pageNumber="86" type="materials_examined">
<paragraph id="8B8436B7953CFFDBCFF75B0FFE97212E" blockId="85.[151,1437,150,465]" pageId="85" pageNumber="86">
<materialsCitation id="3B533CEA953CFFDBCFF75B0FFE97212E" ID-GBIF-Occurrence="3072574496" accessionNumber="ZMH 25258" collectionCode="ZMH" country="South Africa" location="South Africa" pageId="85" pageNumber="86" specimenCode="SAIAB 27140, SAIAB 27141, SAIAB 27142, SAIAB 53231" specimenCount="1" typeStatus="holotype">
<emphasis id="B94FEAA5953CFFDBCFF75B0FFE062173" bold="true" box="[151,511,258,285]" pageId="85" pageNumber="86">
<collectingCountry id="F32C7627953CFFDBCFF75B0FFED72173" box="[151,302,258,284]" name="South Africa" pageId="85" pageNumber="86">South Africa</collectingCountry>
voucher material
</emphasis>
:
<typeStatus id="54808815953CFFDBCD6C5B0EFD802172" box="[524,633,259,285]" pageId="85" pageNumber="86" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED2AAE72953CFFDBCDE25B0EFD392172" box="[642,704,259,285]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="85" pageNumber="86" type="Museum">ZMH</collectionCode>
25258. Non-types:
<specimenCode id="DB9D9ECC953CFFDBCCFC5B0EFBC62172" box="[924,1087,259,285]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="85" pageNumber="86" type="Other type of research institution/biorepository">SAIAB 27140</specimenCode>
,
<specimenCode id="DB9D9ECC953CFFDBCB285B0EFB122172" box="[1096,1259,259,285]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="85" pageNumber="86" type="Other type of research institution/biorepository">SAIAB 27141</specimenCode>
,
<specimenCode id="DB9D9ECC953CFFDBCB955B0EFA612172" box="[1269,1432,259,285]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="85" pageNumber="86" type="Other type of research institution/biorepository">SAIAB 27142</specimenCode>
,
<specimenCode id="DB9D9ECC953CFFDBCFA65B2AFE92212E" box="[198,363,294,321]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="85" pageNumber="86" type="Other type of research institution/biorepository">SAIAB 53231</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8436B7953CFFDBCFF75B47FC00210A" blockId="85.[151,1437,150,465]" box="[151,1017,330,357]" pageId="85" pageNumber="86">
<emphasis id="B94FEAA5953CFFDBCFF75B47FE2A210B" bold="true" box="[151,467,330,356]" pageId="85" pageNumber="86">South African distribution</emphasis>
: The Orange River (NC) to St Francis
<collectingRegion id="49FFF855953CFFDBCCE85B46FC4C210A" box="[904,949,331,357]" country="Somalia" name="Bay" pageId="85" pageNumber="86">Bay</collectingRegion>
(EC).
</paragraph>
</subSubSection>
<subSubSection id="C321653C953CFFDBCFF75B62FDE921BE" pageId="85" pageNumber="86" type="discussion">
<paragraph id="8B8436B7953CFFDBCFF75B62FCD221C2" blockId="85.[151,1437,150,465]" pageId="85" pageNumber="86">
<emphasis id="B94FEAA5953CFFDBCFF75B62FEFD21E6" bold="true" box="[151,260,367,393]" pageId="85" pageNumber="86">Remarks</emphasis>
: A little known, deep-sea skate that is often misidentified with other similar “grey” skates, but its distribution is now known to include northwest Africa.
</paragraph>
<paragraph id="8B8436B7953CFFDBCFF75BBBFDE921BE" blockId="85.[151,1437,150,465]" box="[151,528,438,465]" pageId="85" pageNumber="86">
<emphasis id="B94FEAA5953CFFDBCFF75BBBFE7D21BE" bold="true" box="[151,388,438,465]" pageId="85" pageNumber="86">Conservation status</emphasis>
: LC (2004).
</paragraph>
</subSubSection>
</treatment>
</document>