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<mods:title id="F3BEA60A039B494D989803439DDE36DC">A phylogenetic analysis and systematic revision of the cryptobranch dorids (Mollusca, Nudibranchia, Anthobranchia)</mods:title>
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<paragraph id="8B84CE18FFF66050FCF7FB676A2DD7FC" blockId="25.[880,1374,1193,1217]" box="[880,1374,1193,1217]" pageId="25" pageNumber="560">
<heading id="D0CC7974FFF66050FCF7FB676A2DD7FC" box="[880,1374,1193,1217]" centered="true" fontSize="9" level="2" pageId="25" pageNumber="560" reason="5">
GENUS
<taxonomicName id="4C3BB59BFFF66050FC4FFB646A2DD7FC" ID-CoL="7P56T" ID-ENA="190923" authority="EHRENBERG, 1831" authorityName="Ehrenberg" authorityYear="1831" box="[968,1374,1193,1217]" class="Gastropoda" family="Hexabranchidae" genus="Hexabranchus" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="560" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF66050FC4FFB646BF1D7FC" box="[968,1154,1194,1217]" italics="true" pageId="25" pageNumber="560">HEXABRANCHUS</emphasis>
EHRENBERG, 1831
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3219D93FFF66050FCB4FB1C6AF7D504" pageId="25" pageNumber="560" type="reference_group">
<paragraph id="8B84CE18FFF66050FCB4FB1C6A1DD618" blockId="25.[819,1435,1233,1593]" pageId="25" pageNumber="560">
<taxonomicName id="4C3BB59BFFF66050FCB4FB1C6BD0D7DA" ID-CoL="7P56T" authority="Ehrenberg, 1828" authorityName="Ehrenberg" authorityYear="1828" box="[819,1187,1234,1256]" class="Gastropoda" family="Hexabranchidae" genus="Hexabranchus" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="560" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF66050FCB4FB1C6CA1D7DA" box="[819,978,1234,1255]" italics="true" pageId="25" pageNumber="560">Hexabranchus</emphasis>
<bibRefCitation id="EFAAB3E9FFF66050FC58FB1C6BD0D7DA" author="Ehrenberg" box="[991,1187,1234,1256]" firstAuthor="Ehrenberg" pageId="25" pageNumber="560" refId="ref55258" refString="Ehrenberg CG. 1828 - 1831. Symbolae physicae animalia evertebrata exclusis insectis. Series prima cum tabularum decade prima continent animalia Africana et Asiatica. Decas Prima. In: Hemprich FG, Ehrenberg CG, eds. Symbolae physicae, seu Icones adhue ineditae corporum naturalium novorum aut minus cognitorum, quae ex itineribus per Libyam, Aegyptum, Nubiam, Dengalam, Syriam, Arabiam et Habessiniam Pars Zoologica, 4. Berlin: Officina Academica [pages unnumbered]. Dates of publication: plates 1 - 2 [1828], text [1831]." type="book" year="1828">Ehrenberg, 1828</bibRefCitation>
</taxonomicName>
31 [1831]: 30. Type species:
<taxonomicName id="4C3BB59BFFF66050FC2AFB3F6AE5D63B" ID-CoL="3LFXF" authority="Ehrenberg, 1828" authorityName="Ehrenberg" authorityYear="1828" box="[941,1430,1265,1287]" class="Gastropoda" family="Hexabranchidae" genus="Hexabranchus" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="560" phylum="Mollusca" rank="species" species="praetextus">
<emphasis id="B94F120AFFF66050FC2AFB3F6BBAD63B" box="[941,1225,1265,1286]" italics="true" pageId="25" pageNumber="560">Hexabranchus praetextus</emphasis>
<bibRefCitation id="EFAAB3E9FFF66050FB54FB3F6AE5D63B" author="Ehrenberg" box="[1235,1430,1265,1287]" firstAuthor="Ehrenberg" pageId="25" pageNumber="560" refId="ref55258" refString="Ehrenberg CG. 1828 - 1831. Symbolae physicae animalia evertebrata exclusis insectis. Series prima cum tabularum decade prima continent animalia Africana et Asiatica. Decas Prima. In: Hemprich FG, Ehrenberg CG, eds. Symbolae physicae, seu Icones adhue ineditae corporum naturalium novorum aut minus cognitorum, quae ex itineribus per Libyam, Aegyptum, Nubiam, Dengalam, Syriam, Arabiam et Habessiniam Pars Zoologica, 4. Berlin: Officina Academica [pages unnumbered]. Dates of publication: plates 1 - 2 [1828], text [1831]." type="book" year="1828">Ehrenberg, 1828</bibRefCitation>
</taxonomicName>
, by subsequent designation of J. E.
<bibRefCitation id="EFAAB3E9FFF66050FB5AFAC16A1AD618" author="Gray" box="[1245,1385,1295,1317]" firstAuthor="Gray" pageId="25" pageNumber="560" pagination="129 - 219" refId="ref56529" refString="Gray JE. 1847. A list of the genera of recent Mollusca, their synonyma and types. Proceedings of the Zoological Society of London 15: 129 - 219." type="journal article" year="1847">Gray (1847)</bibRefCitation>
.
</paragraph>
<paragraph id="8B84CE18FFF66050FCB4FAE06B4BD6BC" blockId="25.[819,1435,1233,1593]" pageId="25" pageNumber="560">
<treatmentCitationGroup id="AB2BE936FFF66050FCB4FAE06B4BD6BC" pageId="25" pageNumber="560">
<treatmentCitation id="0A9AE809FFF66050FCB4FAE06A6AD679" author="Adams A" box="[819,1305,1326,1348]" page="495" pageId="25" pageNumber="560" year="1848">
<taxonomicName id="4C3BB59BFFF66050FCB4FAE06A6AD679" ID-CoL="dc37dfde-d028-40a7-bdfa-d02cfebc486d" authority="A. Adams, 1848: 494 - 495" authorityName="A. Adams" authorityPageNumber="494 - 495" authorityYear="1848" box="[819,1305,1326,1348]" class="Gastropoda" family="Hexabranchidae" genus="Heptabranchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="560" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF66050FCB4FAE06CAFD67E" box="[819,988,1326,1347]" italics="true" pageId="25" pageNumber="560">Heptabranchus</emphasis>
A.
<bibRefCitation id="EFAAB3E9FFF66050FB8BFAE06A6AD679" author="Adams" box="[1036,1305,1326,1348]" firstAuthor="Adams" page="494 - 495" pageId="25" pageNumber="560" pagination="1843 - 46" refId="ref52855" refString="Adams A. 1848. Notes from a journal of research into the natural history of the countries visited during the voyage of H. M. S. Samarang, under the command of captain Sir E. Belcher, C. B. In: Belcher E, ed. Narrative of the Voyage of H. M. S. Samarang During the Years 1843 - 46, Vol. 2. London: Reeve and Reeve, 223 - 252." type="book chapter" year="1848">Adams, 1848: 494495</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Type species:
<taxonomicName id="4C3BB59BFFF66050FC0DFA836A1DD65F" authority="A. Adams, 1848" authorityName="A. Adams" authorityYear="1848" box="[906,1390,1357,1379]" class="Gastropoda" family="Hexabranchidae" genus="Heptabranchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="560" phylum="Mollusca" rank="species" species="burnettii">
<emphasis id="B94F120AFFF66050FC0DFA836BD2D65F" box="[906,1185,1357,1378]" italics="true" pageId="25" pageNumber="560">Heptabranchus burnettii</emphasis>
A.
<bibRefCitation id="EFAAB3E9FFF66050FB54FA836A1DD65F" author="Adams" box="[1235,1390,1357,1379]" firstAuthor="Adams" pageId="25" pageNumber="560" pagination="1843 - 46" refId="ref52855" refString="Adams A. 1848. Notes from a journal of research into the natural history of the countries visited during the voyage of H. M. S. Samarang, under the command of captain Sir E. Belcher, C. B. In: Belcher E, ed. Narrative of the Voyage of H. M. S. Samarang During the Years 1843 - 46, Vol. 2. London: Reeve and Reeve, 223 - 252." type="book chapter" year="1848">Adams, 1848</bibRefCitation>
</taxonomicName>
, by original designation.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B84CE18FFF66050FCB4FA446A56D682" blockId="25.[819,1435,1233,1593]" pageId="25" pageNumber="560">
<treatmentCitationGroup id="AB2BE936FFF66050FCB4FA446A56D682" pageId="25" pageNumber="560">
<treatmentCitation id="0A9AE809FFF66050FCB4FA446B0FD6A2" author="Morch OAL" box="[819,1148,1418,1440]" page="34" pageId="25" pageNumber="560" year="1863">
<taxonomicName id="4C3BB59BFFF66050FCB4FA446B0FD6A2" authority="Morch, 1863: 34" authorityName="Morch" authorityPageNumber="34" authorityYear="1863" box="[819,1148,1418,1440]" class="Gastropoda" family="Hexabranchidae" genus="Rhacodoris" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="560" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF66050FCB4FA446CC3D6A2" box="[819,944,1418,1439]" italics="true" pageId="25" pageNumber="560">Rhacodoris</emphasis>
<bibRefCitation id="EFAAB3E9FFF66050FC3CFA446B0FD6A2" author="Morch" box="[955,1148,1418,1440]" firstAuthor="Morch" page="34" pageId="25" pageNumber="560" pagination="21 - 43" refId="ref57535" refString="Morch OAL. 1863. Contributions a la Faune malacologique des Antilles Danoises. Journal de Conchyliologie 11: 21 - 43." type="journal article" year="1863">Mörch, 1863: 34</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Type species:
<taxonomicName id="4C3BB59BFFF66050FAB7FA456B7AD683" authority="Cuvier, 1804" authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="560" phylum="Mollusca" rank="species" species="lacera">
<emphasis id="B94F120AFFF66050FAB7FA456C1CD683" italics="true" pageId="25" pageNumber="560">Doris lacera</emphasis>
<bibRefCitation id="EFAAB3E9FFF66050FCF0FA676B7AD683" author="Cuvier" box="[887,1033,1449,1471]" firstAuthor="Cuvier" pageId="25" pageNumber="560" pagination="447 - 473" refId="ref55151" refString="Cuvier GL. 1804. Memoire sur le genre Doris. Annales du Museum National d'Histoire Naturelle 4: 447 - 473, plates 73 - 74." type="journal article" year="1804">Cuvier, 1804</bibRefCitation>
</taxonomicName>
, by original designation.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B84CE18FFF66050FCB4FA096A5CD6C6" blockId="25.[819,1435,1233,1593]" pageId="25" pageNumber="560">
<treatmentCitationGroup id="AB2BE936FFF66050FCB4FA096A5CD6C6" pageId="25" pageNumber="560">
<treatmentCitation id="0A9AE809FFF66050FCB4FA096BD7D6E1" author="Abraham PS" box="[819,1188,1479,1501]" page="237" pageId="25" pageNumber="560" year="1877">
<taxonomicName id="4C3BB59BFFF66050FCB4FA096BD7D6E1" ID-CoL="7c47bcb1-debd-49d3-9bf3-0fca145c1cce" authority="Abraham, 1877: 237" authorityName="Abraham" authorityPageNumber="237" authorityYear="1877" box="[819,1188,1479,1501]" class="Gastropoda" family="Hexabranchidae" genus="Aethedoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="560" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF66050FCB4FA096CDBD6E1" box="[819,936,1479,1500]" italics="true" pageId="25" pageNumber="560">Aethedoris</emphasis>
<bibRefCitation id="EFAAB3E9FFF66050FC34FA096BD7D6E1" author="Abraham" box="[947,1188,1479,1501]" firstAuthor="Abraham" page="237" pageId="25" pageNumber="560" pagination="196 - 269" refId="ref52815" refString="Abraham PS. 1877. Revision of the anthobranchiate nudibranchiate Mollusca, with descriptions or notices of forty-one hitherto undescribed species. Proceedings of the Zoological Society of London 1877: 196 - 269, plates 27 - 30." type="journal article" year="1877">Abraham, 1877: 237</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Type species:
<taxonomicName id="4C3BB59BFFF66050FAD1FA096BF8D6C6" authority="Abraham, 1877" authorityName="Abraham" authorityYear="1877" class="Gastropoda" family="Hexabranchidae" genus="Aethedoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="560" phylum="Mollusca" rank="species" species="indica">
<emphasis id="B94F120AFFF66050FAD1FA096CA1D6C6" italics="true" pageId="25" pageNumber="560">Aethedoris indica</emphasis>
<bibRefCitation id="EFAAB3E9FFF66050FC5EFA286BF8D6C6" author="Abraham" box="[985,1163,1510,1532]" firstAuthor="Abraham" pageId="25" pageNumber="560" pagination="196 - 269" refId="ref52815" refString="Abraham PS. 1877. Revision of the anthobranchiate nudibranchiate Mollusca, with descriptions or notices of forty-one hitherto undescribed species. Proceedings of the Zoological Society of London 1877: 196 - 269, plates 27 - 30." type="journal article" year="1877">Abraham, 1877</bibRefCitation>
</taxonomicName>
, by monotypy.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B84CE18FFF66050FCB4F9CB6AF7D504" blockId="25.[819,1435,1233,1593]" pageId="25" pageNumber="560">
<treatmentCitationGroup id="AB2BE936FFF66050FCB4F9CB6AF7D504" pageId="25" pageNumber="560">
<treatmentCitation id="0A9AE809FFF66050FCB4F9CB6B98D527" author="Collingwood" box="[819,1259,1541,1563]" page="132" pageId="25" pageNumber="560" year="1881">
<taxonomicName id="4C3BB59BFFF66050FCB4F9CB6B98D527" authority="Collingwood, 1881" authorityName="Collingwood" authorityYear="1881" box="[819,1259,1541,1563]" class="Gastropoda" family="Archidorididae" genus="Albania" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="560" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF66050FCB4F9CB6CF8D527" box="[819,907,1541,1562]" italics="true" pageId="25" pageNumber="560">
<collectingCountry id="F32C8E88FFF66050FCB4F9CB6CF8D527" box="[819,907,1541,1562]" name="Albania" pageId="25" pageNumber="560">Albania</collectingCountry>
</emphasis>
<bibRefCitation id="EFAAB3E9FFF66050FC1DF9CB6B98D527" author="Collingwood" box="[922,1259,1541,1563]" firstAuthor="Collingwood" page="132 - 133" pageId="25" pageNumber="560" pagination="123 - 140" refId="ref55113" refString="Collingwood C. 1881. On some new species of nudibranchiate mollusca from the eastern seas. Transactions of the Linnean Society of London 2 (2): 123 - 140, plates 9 - 10." type="journal article" year="1881">Collingwood, 1881: 132133</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Type species:
<taxonomicName id="4C3BB59BFFF66050FCCBF9ED6B93D505" authority="Collingwood, 1881" authorityName="Collingwood" authorityYear="1881" box="[844,1248,1571,1593]" class="Gastropoda" family="Archidorididae" genus="Albania" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="560" phylum="Mollusca" rank="species" species="formosa">
<emphasis id="B94F120AFFF66050FCCBF9ED6B77D505" box="[844,1028,1571,1592]" italics="true" pageId="25" pageNumber="560">Albania formosa</emphasis>
<bibRefCitation id="EFAAB3E9FFF66050FB8BF9ED6B93D505" author="Collingwood" box="[1036,1248,1571,1593]" firstAuthor="Collingwood" pageId="25" pageNumber="560" pagination="123 - 140" refId="ref55113" refString="Collingwood C. 1881. On some new species of nudibranchiate mollusca from the eastern seas. Transactions of the Linnean Society of London 2 (2): 123 - 140, plates 9 - 10." type="journal article" year="1881">Collingwood, 1881</bibRefCitation>
</taxonomicName>
, by monotypy.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3219D93FFF66053FCB4F94A6E46D22A" lastPageId="26" lastPageNumber="561" pageId="25" pageNumber="560" type="diagnosis">
<paragraph id="8B84CE18FFF66050FCB4F94A6CD3D5A4" blockId="25.[819,1435,1668,1904]" box="[819,928,1668,1689]" pageId="25" pageNumber="560">
<emphasis id="B94F120AFFF66050FCB4F94A6CD3D5A4" box="[819,928,1668,1689]" italics="true" pageId="25" pageNumber="560">Diagnosis</emphasis>
</paragraph>
<paragraph id="8B84CE18FFF66053FCB4F96D6E46D22A" blockId="25.[819,1435,1668,1904]" lastBlockId="26.[142,758,196,279]" lastPageId="26" lastPageNumber="561" pageId="25" pageNumber="560">Dorsum smooth, lacking tubercles. Head with two large, flattened and lobate oral tentacles. Anterior border of the foot simple. Gill contractile, not retractile. Radula composed of simple, hamate teeth. Labial cuticle completely covered with rodlets and having several transverse grooves. Buccal mass with numerous and strong muscles attached. Reproductive system with a tubular, non differentiated prostate. Penis and vagina devoid of hooks. Vestibular or accessory glands absent.</paragraph>
</subSubSection>
<subSubSection id="C3219D93FFF56053FF09FE936CBFD2D0" pageId="26" pageNumber="561" type="discussion">
<paragraph id="8B84CE18FFF56053FF09FE936F83D24F" blockId="26.[142,759,349,1904]" box="[142,240,349,370]" pageId="26" pageNumber="561">
<emphasis id="B94F120AFFF56053FF09FE936F83D24F" box="[142,240,349,370]" italics="true" pageId="26" pageNumber="561">Remarks</emphasis>
</paragraph>
<paragraph id="8B84CE18FFF56053FF09FEB26FA7D198" blockId="26.[142,759,349,1904]" pageId="26" pageNumber="561">
The genus
<taxonomicName id="4C3BB59BFFF56053FE9EFEB26ECBD2AC" authorityName="Ehrenberg" authorityYear="1828" box="[281,440,380,401]" class="Gastropoda" family="Hexabranchidae" genus="Hexabranchus" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF56053FE9EFEB26ECBD2AC" box="[281,440,380,401]" italics="true" pageId="26" pageNumber="561">Hexabranchus</emphasis>
</taxonomicName>
was originally introduced by
<bibRefCitation id="EFAAB3E9FFF56053FF3BFE546EFBD28C" author="Ehrenberg" box="[188,392,410,433]" firstAuthor="Ehrenberg" pageId="26" pageNumber="561" refId="ref55258" refString="Ehrenberg CG. 1828 - 1831. Symbolae physicae animalia evertebrata exclusis insectis. Series prima cum tabularum decade prima continent animalia Africana et Asiatica. Decas Prima. In: Hemprich FG, Ehrenberg CG, eds. Symbolae physicae, seu Icones adhue ineditae corporum naturalium novorum aut minus cognitorum, quae ex itineribus per Libyam, Aegyptum, Nubiam, Dengalam, Syriam, Arabiam et Habessiniam Pars Zoologica, 4. Berlin: Officina Academica [pages unnumbered]. Dates of publication: plates 1 - 2 [1828], text [1831]." type="book" year="1828">Ehrenberg (1828</bibRefCitation>
31) based on three species:
<taxonomicName id="4C3BB59BFFF56053FF09FE776D07D2F3" authority="Ehrenberg, 1831" authorityName="Ehrenberg" authorityYear="1831" box="[142,628,441,463]" class="Gastropoda" family="Hexabranchidae" genus="Hexabranchus" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="species" species="praetextus">
<emphasis id="B94F120AFFF56053FF09FE776EDAD2F2" box="[142,425,441,463]" italics="true" pageId="26" pageNumber="561">Hexabranchus praetextus</emphasis>
Ehrenberg, 1831
</taxonomicName>
,
<taxonomicName id="4C3BB59BFFF56053FD05FE746D7DD2D0" authority="Ruppell &amp; Leuckart, 1830" authorityName="Ruppell &amp; Leuckart" authorityYear="1830" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="species" species="sanguinea">
<emphasis id="B94F120AFFF56053FD05FE746FAAD2D0" italics="true" pageId="26" pageNumber="561">Doris sanguinea</emphasis>
Rüppell &amp; Leuckart, 1830
</taxonomicName>
and
<taxonomicName id="4C3BB59BFFF56053FDC8FE166D85D2D0" box="[591,758,472,493]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="species" species="laciniata">
<emphasis id="B94F120AFFF56053FDC8FE166D85D2D0" box="[591,758,472,493]" italics="true" pageId="26" pageNumber="561">Doris laciniata</emphasis>
</taxonomicName>
Cuvier (error for
<taxonomicName id="4C3BB59BFFF56053FED4FE396D03D130" authority="Cuvier, 1804" authorityName="Cuvier" authorityYear="1804" box="[339,624,502,525]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="species" species="lacera">
<emphasis id="B94F120AFFF56053FED4FE396EAAD136" box="[339,473,502,524]" italics="true" pageId="26" pageNumber="561">Doris lacera</emphasis>
<bibRefCitation id="EFAAB3E9FFF56053FE58FE386D03D130" author="Cuvier" box="[479,624,502,525]" firstAuthor="Cuvier" pageId="26" pageNumber="561" pagination="447 - 473" refId="ref55151" refString="Cuvier GL. 1804. Memoire sur le genre Doris. Annales du Museum National d'Histoire Naturelle 4: 447 - 473, plates 73 - 74." type="journal article" year="1804">Cuvier, 1804</bibRefCitation>
</taxonomicName>
).
<taxonomicName id="4C3BB59BFFF56053FD06FE386E33D116" authorityName="Ehrenberg" authorityYear="1831" class="Gastropoda" family="Hexabranchidae" genus="Hexabranchus" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="species" species="praetextus">
<emphasis id="B94F120AFFF56053FD06FE386E33D116" italics="true" pageId="26" pageNumber="561">Hexabranchus praetextus</emphasis>
</taxonomicName>
was subsequently selected by
<bibRefCitation id="EFAAB3E9FFF56053FD3AFDDB6FA5D177" author="Gray" firstAuthor="Gray" pageId="26" pageNumber="561" pagination="129 - 219" refId="ref56529" refString="Gray JE. 1847. A list of the genera of recent Mollusca, their synonyma and types. Proceedings of the Zoological Society of London 15: 129 - 219." type="journal article" year="1847">Gray (1847)</bibRefCitation>
as the
<typeStatus id="548070BAFFF56053FEB5FDFA6E10D174" box="[306,355,564,585]" pageId="26" pageNumber="561">type</typeStatus>
species. This species was detailed described and illustrated by Ehrenberg, (182831), and its features agree with the current usage of the name.
</paragraph>
<paragraph id="8B84CE18FFF56053FF20FD606DC8D74C" blockId="26.[142,759,349,1904]" pageId="26" pageNumber="561">
<bibRefCitation id="EFAAB3E9FFF56053FF20FD606E3BD1F8" author="Adams" box="[167,328,686,709]" firstAuthor="Adams" pageId="26" pageNumber="561" pagination="1843 - 46" refId="ref52855" refString="Adams A. 1848. Notes from a journal of research into the natural history of the countries visited during the voyage of H. M. S. Samarang, under the command of captain Sir E. Belcher, C. B. In: Belcher E, ed. Narrative of the Voyage of H. M. S. Samarang During the Years 1843 - 46, Vol. 2. London: Reeve and Reeve, 223 - 252." type="book chapter" year="1848">Adams (1848)</bibRefCitation>
described the genus
<taxonomicName id="4C3BB59BFFF56053FDC1FD606D9CD1FE" authorityName="A. Adams" authorityYear="1848" box="[582,751,686,707]" class="Gastropoda" family="Hexabranchidae" genus="Heptabranchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF56053FDC1FD606D9CD1FE" box="[582,751,686,707]" italics="true" pageId="26" pageNumber="561">Heptabranchus</emphasis>
</taxonomicName>
,
<typeStatus id="548070BAFFF56053FF09FD006FCCD1DE" box="[142,191,718,739]" pageId="26" pageNumber="561">type</typeStatus>
species by original designation
<taxonomicName id="4C3BB59BFFF56053FDCAFD036EB5D03C" authority="A. Adams, 1848" authorityName="A. Adams" authorityYear="1848" class="Gastropoda" family="Hexabranchidae" genus="Heptabranchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="species" species="burnettii">
<emphasis id="B94F120AFFF56053FDCAFD036F9CD03C" italics="true" pageId="26" pageNumber="561">Heptabranchus burnettii</emphasis>
A.
<bibRefCitation id="EFAAB3E9FFF56053FEA0FD226EB5D03C" author="Adams" box="[295,454,748,770]" firstAuthor="Adams" pageId="26" pageNumber="561" pagination="1843 - 46" refId="ref52855" refString="Adams A. 1848. Notes from a journal of research into the natural history of the countries visited during the voyage of H. M. S. Samarang, under the command of captain Sir E. Belcher, C. B. In: Belcher E, ed. Narrative of the Voyage of H. M. S. Samarang During the Years 1843 - 46, Vol. 2. London: Reeve and Reeve, 223 - 252." type="book chapter" year="1848">Adams, 1848</bibRefCitation>
</taxonomicName>
, as being very close to
<taxonomicName id="4C3BB59BFFF56053FF09FCC46E5ED022" authorityName="Ehrenberg" authorityYear="1828" box="[142,301,778,799]" class="Gastropoda" family="Hexabranchidae" genus="Hexabranchus" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF56053FF09FCC46E5ED022" box="[142,301,778,799]" italics="true" pageId="26" pageNumber="561">Hexabranchus</emphasis>
</taxonomicName>
, but showing several differences in the number of gills and mantle widtH. In his opinion, these differences supported the separation of two different genera. Nowadays it is known that species of
<taxonomicName id="4C3BB59BFFF56053FF09FC4B6E5ED0A7" authorityName="Ehrenberg" authorityYear="1828" box="[142,301,901,922]" class="Gastropoda" family="Hexabranchidae" genus="Hexabranchus" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF56053FF09FC4B6E5ED0A7" box="[142,301,901,922]" italics="true" pageId="26" pageNumber="561">Hexabranchus</emphasis>
</taxonomicName>
can contract and spread out the mantle margin (
<bibRefCitation id="EFAAB3E9FFF56053FF77FC6A6EDCD087" author="Thompson" box="[240,431,932,954]" firstAuthor="Thompson" pageId="26" pageNumber="561" pagination="1 - 5" refId="ref59584" refString="Thompson TE. 1972. Observations on Hexabranchus from the Australian Great Barrier Reef (Gastropoda: Opisthobranchia). The Veliger 15: 1 - 5." type="journal article" year="1972">Thompson, 1972</bibRefCitation>
), so the same animal is able to show a narrow mantle margin with the foot extending beyond it (as described by
<bibRefCitation id="EFAAB3E9FFF56053FE6DFC2F6D0DD0CA" author="Adams" box="[490,638,993,1015]" firstAuthor="Adams" pageId="26" pageNumber="561" pagination="1843 - 46" refId="ref52855" refString="Adams A. 1848. Notes from a journal of research into the natural history of the countries visited during the voyage of H. M. S. Samarang, under the command of captain Sir E. Belcher, C. B. In: Belcher E, ed. Narrative of the Voyage of H. M. S. Samarang During the Years 1843 - 46, Vol. 2. London: Reeve and Reeve, 223 - 252." type="book chapter" year="1848">Adams, 1848</bibRefCitation>
) or a wide mantle completely covering the foot. In addition, the number of branchial leaves is variable among the same species. Therefore, there is no doubt that
<taxonomicName id="4C3BB59BFFF56053FD3EFBF06E71D74C" authorityName="A. Adams" authorityYear="1848" class="Gastropoda" family="Hexabranchidae" genus="Heptabranchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF56053FD3EFBF06E71D74C" italics="true" pageId="26" pageNumber="561">Heptabranchus</emphasis>
</taxonomicName>
is a junior synonym of
<taxonomicName id="4C3BB59BFFF56053FD90FB926DC6D74C" authorityName="Ehrenberg" authorityYear="1828" box="[535,693,1116,1137]" class="Gastropoda" family="Hexabranchidae" genus="Hexabranchus" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF56053FD90FB926DC6D74C" box="[535,693,1116,1137]" italics="true" pageId="26" pageNumber="561">Hexabranchus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B84CE18FFF56053FF20FBB46DDDD6DC" blockId="26.[142,759,349,1904]" pageId="26" pageNumber="561">
<bibRefCitation id="EFAAB3E9FFF56053FF20FBB46E30D7AD" author="Morch" box="[167,323,1146,1168]" firstAuthor="Morch" pageId="26" pageNumber="561" pagination="21 - 43" refId="ref57535" refString="Morch OAL. 1863. Contributions a la Faune malacologique des Antilles Danoises. Journal de Conchyliologie 11: 21 - 43." type="journal article" year="1863">Mörch (1863)</bibRefCitation>
introduced the name
<taxonomicName id="4C3BB59BFFF56053FDC8FBB46DBFD7B2" authorityName="Morch" authorityYear="1863" box="[591,716,1146,1167]" class="Gastropoda" family="Hexabranchidae" genus="Rhacodoris" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF56053FDC8FBB46DBFD7B2" box="[591,716,1146,1167]" italics="true" pageId="26" pageNumber="561">Rhacodoris</emphasis>
</taxonomicName>
for
<taxonomicName id="4C3BB59BFFF56053FF09FB576EC6D793" authority="sensu Gray" authorityName="Gray" box="[142,437,1177,1199]" class="Gastropoda" family="Hexabranchidae" genus="Hexabranchus" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF56053FF09FB576E00D792" box="[142,371,1177,1199]" italics="true" pageId="26" pageNumber="561">Hexabranchus sensu</emphasis>
Gray
</taxonomicName>
non Ehrenberg, with
<taxonomicName id="4C3BB59BFFF56053FD3DFB546F82D7F0" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="species" species="laciniata">
<emphasis id="B94F120AFFF56053FD3DFB546F82D7F0" italics="true" pageId="26" pageNumber="561">Doris laciniata</emphasis>
</taxonomicName>
Cuvier (error for
<taxonomicName id="4C3BB59BFFF56053FE4CFB766D9DD7F3" authority="Cuvier, 1804" authorityName="Cuvier" authorityYear="1804" box="[459,750,1208,1230]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="species" species="lacera">
<emphasis id="B94F120AFFF56053FE4CFB766D27D7F0" box="[459,596,1208,1229]" italics="true" pageId="26" pageNumber="561">Doris lacera</emphasis>
<bibRefCitation id="EFAAB3E9FFF56053FDDAFB766D9DD7F3" author="Cuvier" box="[605,750,1208,1230]" firstAuthor="Cuvier" pageId="26" pageNumber="561" pagination="447 - 473" refId="ref55151" refString="Cuvier GL. 1804. Memoire sur le genre Doris. Annales du Museum National d'Histoire Naturelle 4: 447 - 473, plates 73 - 74." type="journal article" year="1804">Cuvier, 1804</bibRefCitation>
</taxonomicName>
) as the
<typeStatus id="548070BAFFF56053FF64FB196E67D7D1" box="[227,276,1239,1260]" pageId="26" pageNumber="561">type</typeStatus>
species by original designation. He also stated that
<taxonomicName id="4C3BB59BFFF56053FE9CFB386ED6D637" authorityName="Cuvier" authorityYear="1804" box="[283,421,1269,1291]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="species" species="lacera">
<emphasis id="B94F120AFFF56053FE9CFB386ED6D637" box="[283,421,1269,1291]" italics="true" pageId="26" pageNumber="561">Doris lacera</emphasis>
</taxonomicName>
was mistakenly reported as belonging to the genus
<taxonomicName id="4C3BB59BFFF56053FE21FADA6D36D614" box="[422,581,1300,1321]" class="Gastropoda" family="Hexabranchidae" genus="Hexabranchus" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF56053FE21FADA6D36D614" box="[422,581,1300,1321]" italics="true" pageId="26" pageNumber="561">Hexabranchus</emphasis>
</taxonomicName>
, from which it differs in having a special cavity for each branchial leaf and one common cavity for all the gill. The examination of the
<typeStatus id="548070BAFFF56053FEABFABE6E2ED6B8" box="[300,349,1392,1413]" pageId="26" pageNumber="561">type</typeStatus>
material and original description of
<taxonomicName id="4C3BB59BFFF56053FF09FA416EA1D698" authority="(Cuvier, 1804)" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[142,466,1422,1445]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="species" species="lacera">
<emphasis id="B94F120AFFF56053FF09FA416E6FD69E" box="[142,284,1422,1444]" italics="true" pageId="26" pageNumber="561">Doris lacera</emphasis>
(
<bibRefCitation id="EFAAB3E9FFF56053FEB6FA406EB9D698" author="Cuvier" box="[305,458,1422,1445]" firstAuthor="Cuvier" pageId="26" pageNumber="561" pagination="447 - 473" refId="ref55151" refString="Cuvier GL. 1804. Memoire sur le genre Doris. Annales du Museum National d'Histoire Naturelle 4: 447 - 473, plates 73 - 74." type="journal article" year="1804">Cuvier, 1804</bibRefCitation>
)
</taxonomicName>
, show that this species clearly belongs to the genus
<taxonomicName id="4C3BB59BFFF56053FE53FA636D00D6FF" box="[468,627,1453,1474]" class="Gastropoda" family="Hexabranchidae" genus="Hexabranchus" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF56053FE53FA636D00D6FF" box="[468,627,1453,1474]" italics="true" pageId="26" pageNumber="561">Hexabranchus</emphasis>
</taxonomicName>
, and therefore
<taxonomicName id="4C3BB59BFFF56053FF44FA026E33D6DC" authorityName="Morch" authorityYear="1863" box="[195,320,1484,1505]" class="Gastropoda" family="Hexabranchidae" genus="Rhacodoris" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF56053FF44FA026E33D6DC" box="[195,320,1484,1505]" italics="true" pageId="26" pageNumber="561">Rhacodoris</emphasis>
</taxonomicName>
is a junior subjective synonym.
</paragraph>
<paragraph id="8B84CE18FFF56053FF20FA246A3CD3C5" blockId="26.[142,759,349,1904]" lastBlockId="26.[806,1423,196,493]" pageId="26" pageNumber="561">
The genus
<taxonomicName id="4C3BB59BFFF56053FEB4FA246EDBD6C2" authorityName="Abraham" authorityYear="1877" box="[307,424,1514,1535]" class="Gastropoda" family="Hexabranchidae" genus="Aethedoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF56053FEB4FA246EDBD6C2" box="[307,424,1514,1535]" italics="true" pageId="26" pageNumber="561">Aethedoris</emphasis>
</taxonomicName>
and the species
<taxonomicName id="4C3BB59BFFF56053FD06FA246FA7D523" authorityName="Abraham" authorityYear="1877" class="Gastropoda" family="Hexabranchidae" genus="Aethedoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="species" species="indica">
<emphasis id="B94F120AFFF56053FD06FA246FA7D523" italics="true" pageId="26" pageNumber="561">Aethedoris indica</emphasis>
</taxonomicName>
were erected by
<bibRefCitation id="EFAAB3E9FFF56053FE16F9C76D39D522" author="Abraham" box="[401,586,1545,1567]" firstAuthor="Abraham" pageId="26" pageNumber="561" pagination="196 - 269" refId="ref52815" refString="Abraham PS. 1877. Revision of the anthobranchiate nudibranchiate Mollusca, with descriptions or notices of forty-one hitherto undescribed species. Proceedings of the Zoological Society of London 1877: 196 - 269, plates 27 - 30." type="journal article" year="1877">Abraham (1877)</bibRefCitation>
based on a picture of
<bibRefCitation id="EFAAB3E9FFF56053FF60F9E66E8FD503" author="Alder &amp; Hancock" box="[231,508,1576,1598]" firstAuthor="Alder" pageId="26" pageNumber="561" pagination="113 - 147" refId="ref53398" refString="Alder J, Hancock A. 1864. Notice of a collection of nudibranchiate Mollusca made in India by Water Elliot Esq., with descriptions of several new genera and species. Transactions of the Zoological Society of London 5: 113 - 147, plates 28 - 33." type="journal article" year="1864">Alder &amp; Hancock (1864</bibRefCitation>
; pl. 33, fig. 20) which represents a contracted, probably dead specimen. The two large and lobate oral tentacles shown in the picture clearly identified the specimen as belonging to the genus
<taxonomicName id="4C3BB59BFFF56053FE80F96C6ED6D58A" box="[263,421,1698,1719]" class="Gastropoda" family="Hexabranchidae" genus="Hexabranchus" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF56053FE80F96C6ED6D58A" box="[263,421,1698,1719]" italics="true" pageId="26" pageNumber="561">Hexabranchus</emphasis>
</taxonomicName>
, but they were considered by Abraham as the most striking feature of his new taxa. He interpreted them as a bilobed head, each lobe being semicrescentic, with the apex curving backwards and the margin bearing 1214 conical dentations. The
<typeStatus id="548070BAFFF56053FE8DF8F26E48D46C" box="[266,315,1852,1873]" pageId="26" pageNumber="561">type</typeStatus>
material of
<taxonomicName id="4C3BB59BFFF56053FE41F8F26DFBD46C" authorityName="Abraham" authorityYear="1877" box="[454,648,1852,1873]" class="Gastropoda" family="Hexabranchidae" genus="Aethedoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="species" species="indica">
<emphasis id="B94F120AFFF56053FE41F8F26DFBD46C" box="[454,648,1852,1873]" italics="true" pageId="26" pageNumber="561">Aethedoris indica</emphasis>
</taxonomicName>
could not be located in BMNH and is probably lost. However, it is very likely that the animal figured by Alder &amp; Hancock was a dead specimen of
<taxonomicName id="4C3BB59BFFF56053FB2CFF2C6A3AD3CA" box="[1195,1353,226,247]" class="Gastropoda" family="Hexabranchidae" genus="Hexabranchus" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF56053FB2CFF2C6A3AD3CA" box="[1195,1353,226,247]" italics="true" pageId="26" pageNumber="561">Hexabranchus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B84CE18FFF56053FCB8FECF6CBFD2D0" blockId="26.[806,1423,196,493]" pageId="26" pageNumber="561">
<bibRefCitation id="EFAAB3E9FFF56053FCB8FECF6B69D22A" author="Collingwood" box="[831,1050,257,279]" firstAuthor="Collingwood" pageId="26" pageNumber="561" pagination="123 - 140" refId="ref55113" refString="Collingwood C. 1881. On some new species of nudibranchiate mollusca from the eastern seas. Transactions of the Linnean Society of London 2 (2): 123 - 140, plates 9 - 10." type="journal article" year="1881">Collingwood (1881)</bibRefCitation>
introduced the new genus
<emphasis id="B94F120AFFF56053FAD4FECF6C39D208" italics="true" pageId="26" pageNumber="561">
<collectingCountry id="F32C8E88FFF56053FAD4FECF6C39D208" name="Albania" pageId="26" pageNumber="561">Albania</collectingCountry>
</emphasis>
with
<taxonomicName id="4C3BB59BFFF56053FC1FFEEE6A44D208" authority="Collingwood, 1881" authorityName="Collingwood" authorityYear="1881" box="[920,1335,288,310]" class="Gastropoda" family="Archidorididae" genus="Albania" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="species" species="formosa">
<emphasis id="B94F120AFFF56053FC1FFEEE6B27D208" box="[920,1108,288,309]" italics="true" pageId="26" pageNumber="561">Albania formosa</emphasis>
<bibRefCitation id="EFAAB3E9FFF56053FBE6FEEE6A44D208" author="Collingwood" box="[1121,1335,288,310]" firstAuthor="Collingwood" pageId="26" pageNumber="561" pagination="123 - 140" refId="ref55113" refString="Collingwood C. 1881. On some new species of nudibranchiate mollusca from the eastern seas. Transactions of the Linnean Society of London 2 (2): 123 - 140, plates 9 - 10." type="journal article" year="1881">Collingwood, 1881</bibRefCitation>
</taxonomicName>
as the single included species (
<typeStatus id="548070BAFFF56053FBC4FEF16B06D269" box="[1091,1141,319,340]" pageId="26" pageNumber="561">type</typeStatus>
by monotypy). The features of
<emphasis id="B94F120AFFF56053FC0AFE936C96D24F" box="[909,997,349,370]" italics="true" pageId="26" pageNumber="561">
<collectingCountry id="F32C8E88FFF56053FC0AFE936C96D24F" box="[909,997,349,370]" name="Albania" pageId="26" pageNumber="561">Albania</collectingCountry>
</emphasis>
are identical to those of the genus
<taxonomicName id="4C3BB59BFFF56053FCA1FEB26CB6D2AC" box="[806,965,380,401]" class="Gastropoda" family="Hexabranchidae" genus="Hexabranchus" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF56053FCA1FEB26CB6D2AC" box="[806,965,380,401]" italics="true" pageId="26" pageNumber="561">Hexabranchus</emphasis>
</taxonomicName>
. In this case the
<typeStatus id="548070BAFFF56053FB45FEB26B80D2AC" box="[1218,1267,380,401]" pageId="26" pageNumber="561">type</typeStatus>
material of
<taxonomicName id="4C3BB59BFFF56053FCA1FE546C93D28D" authorityName="Collingwood" authorityYear="1881" box="[806,992,410,432]" class="Gastropoda" family="Archidorididae" genus="Albania" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="species" species="formosa">
<emphasis id="B94F120AFFF56053FCA1FE546C93D28D" box="[806,992,410,432]" italics="true" pageId="26" pageNumber="561">Albania formosa</emphasis>
</taxonomicName>
is also lost but there are not doubts that this nominal species belong to the genus
<taxonomicName id="4C3BB59BFFF56053FCA1FE166CB6D2D0" box="[806,965,472,493]" class="Gastropoda" family="Hexabranchidae" genus="Hexabranchus" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="561" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFF56053FCA1FE166CB6D2D0" box="[806,965,472,493]" italics="true" pageId="26" pageNumber="561">Hexabranchus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>