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<document id="0FF9E8F18C4DF6E29EA16388BF44C85E" ID-CLB-Dataset="30484" ID-DOI="10.11646/zootaxa.4396.1.1" ID-GBIF-Dataset="918f2145-a708-460e-b19b-4f67e6c99b30" ID-ISSN="1175-5326" ID-Zenodo-Dep="1202723" ID-ZooBank="8EEBE6DE-0ECC-4B9C-AD14-01438291782B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1521441296340" checkinUser="plazi" docAuthor="Smith, Patrick M., Paterson, John R. &amp; Brock, Glenn A." docDate="2018" docId="038CC426FFA2FF86FF399EBCFB0AFC2B" docLanguage="en" docName="zootaxa.4396.1.1.pdf" docOrigin="Zootaxa 4396 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hebeia helixensis Jago &amp; Cooper 2005" docType="treatment" docVersion="7" lastPageNumber="16" masterDocId="FFB5BC5EFFAFFF89FFAE9D71FF99FFE3" masterDocTitle="Trilobites and agnostids from the Goyder Formation (Cambrian Series 3, Guzhangian; Mindyallan), Amadeus Basin, central Australia" masterLastPageNumber="67" masterPageNumber="1" pageNumber="14" updateTime="1698532001681" updateUser="ExternalLinkService">
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<mods:title id="5CA5A141BDC2CD8F967481C2067CA1EB">Trilobites and agnostids from the Goyder Formation (Cambrian Series 3, Guzhangian; Mindyallan), Amadeus Basin, central Australia</mods:title>
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<mods:namePart id="59295798DDBBB09B39A0CDBC7A99021A">Smith, Patrick M.</mods:namePart>
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<mods:namePart id="1A54638566724F80AB2EF0A65D6939B2">Paterson, John R.</mods:namePart>
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<mods:namePart id="2EFC6675D7BF118D73BB8A3E3C7F526D">Brock, Glenn A.</mods:namePart>
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<treatment id="038CC426FFA2FF86FF399EBCFB0AFC2B" ID-DOI="http://doi.org/10.5281/zenodo.5980802" ID-GBIF-Taxon="142340466" ID-Zenodo-Dep="5980802" LSID="urn:lsid:plazi:treatment:038CC426FFA2FF86FF399EBCFB0AFC2B" httpUri="http://treatment.plazi.org/id/038CC426FFA2FF86FF399EBCFB0AFC2B" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
<subSubSection id="C33F26BBFFA2FF84FF399EBCFF40FBEB" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8B9A7530FFA2FF84FF399EBCFD57FC04" blockId="13.[151,718,973,1032]" box="[151,718,973,999]" pageId="13" pageNumber="14">
<heading id="D0D2C25CFFA2FF84FF399EBCFD57FC04" bold="true" box="[151,718,973,999]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="B951A922FFA2FF84FF399EBCFD57FC04" bold="true" box="[151,718,973,999]" pageId="13" pageNumber="14">
<emphasis id="B951A922FFA2FF84FF399EBCFEDBFC04" bold="true" box="[151,322,973,999]" italics="true" pageId="13" pageNumber="14">Hadragnostus</emphasis>
<taxonomicName id="4C250EB3FFA2FF84FEE79EBCFD57FC04" authority="Jago &amp; Cooper, 2005" authorityName="Jago &amp; Cooper" authorityYear="2005" box="[329,718,973,999]" class="Trilobita" family="Proasaphiscidae" genus="Hebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="helixensis">
<emphasis id="B951A922FFA2FF84FEE79EBCFE58FC04" bold="true" box="[329,449,973,999]" italics="true" pageId="13" pageNumber="14">helixensis</emphasis>
Jago &amp; Cooper, 2005
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A7530FFA2FF84FF399E9EFF40FBEB" blockId="13.[151,718,973,1032]" box="[151,217,1007,1032]" pageId="13" pageNumber="14">
<figureCitation id="131E69B5FFA2FF84FF399E9EFF40FBEB" box="[151,217,1007,1032]" captionStart="FIGURE 6" captionStartId="14.[151,250,1675,1697]" captionTargetBox="[167,1424,228,1608]" captionTargetId="figure@14.[150,1437,221,1622]" captionTargetPageId="14" captionText="FIGURE 6. Hadragnostus helixensis Jago &amp; Cooper, 2005. All specimens come from sample GOY/97. A, L, CPC42204, partial cephalon; A, dorsal view; L, oblique anterolateral view. B, M, O, CPC42205, partial cephalon; B, dorsal view; M, anterior view; O, lateral view. C, CPC42206, partial cephalon. D, CPC42207, partial cephalon. E, CPC442208, partial cephalon.F,CPC42209, partialcephalon.G,CPC42210,partial cephalon.H, CPC42211, partial cephalon.I, CPC42212, partial cephalon. J, CPC42213, partial cephalon. K, CPC42214, partial cephalon. N, P, X CPC42215, partial pygidium; N, posterior view; P, lateral view; X, dorsal view. Q, CPC42216, partial pygidium. R, CPC42217, partial pygidium. S, CPC42218, partial pygidium. T, CPC42219, partial pygidium. U, CPC42220, partial pygidium. V, CPC42221, partial pygidium. W, CPC42222, partial pygidium.Y, CPC42223, partial pygidium. All scale bars are 1 mm." httpUri="https://zenodo.org/record/1202735/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33F26BBFFA2FF84FF399949FCABFBAD" box="[151,818,1080,1102]" pageId="13" pageNumber="14" type="description">
<paragraph id="8B9A7530FFA2FF84FF399949FCABFBAD" blockId="13.[151,818,1080,1102]" box="[151,818,1080,1102]" pageId="13" pageNumber="14">
2005
<emphasis id="B951A922FFA2FF84FF7E9949FE50FBAD" box="[208,457,1080,1102]" italics="true" pageId="13" pageNumber="14">
Hadragnostus
<taxonomicName id="4C250EB3FFA2FF84FECB9949FE50FBAD" authorityName="Jago &amp; Cooper" authorityYear="2005" box="[357,457,1080,1102]" class="Trilobita" family="Proasaphiscidae" genus="Hebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="helixensis">helixensis</taxonomicName>
</emphasis>
; Jago &amp; Cooper, p. 665, fig. 2AL.
</paragraph>
</subSubSection>
<subSubSection id="C33F26BBFFA2FF84FF39990BFEECFB54" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8B9A7530FFA2FF84FF39990BFEECFB54" blockId="13.[151,1436,1146,2035]" pageId="13" pageNumber="14">
<emphasis id="B951A922FFA2FF84FF39990BFE9FFB70" bold="true" box="[151,262,1146,1171]" pageId="13" pageNumber="14">Material.</emphasis>
Eleven cephala and nine pygidia figured, CPC42204CPC42223. 53 cephala and 36 pygidia not figured (mostly fragments).
</paragraph>
</subSubSection>
<subSubSection id="C33F26BBFFA2FF84FF6999B3FDA4F810" pageId="13" pageNumber="14" type="description">
<paragraph id="8B9A7530FFA2FF84FF6999B3FC1DF9FC" blockId="13.[151,1436,1146,2035]" pageId="13" pageNumber="14">
<emphasis id="B951A922FFA2FF84FF6999B3FEC0FB38" bold="true" box="[199,345,1218,1243]" pageId="13" pageNumber="14">Description.</emphasis>
Cephalon and pygidium each attaining about
<quantity id="4CDDD8D5FFA2FF84FCC799B3FC58FB39" box="[873,961,1218,1243]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="14" unit="mm" value="3.5">3.5 mm</quantity>
in length; nonscrobiculate; surface finely granulose. Cephalon subquadrate to subcircular in outline, about 87% as long as wide, nonspinose, strongly convex; border wide (sag., exsag.), flattened; border furrow well developed, moderately narrow (sag., exsag.); preglabellar median furrow shallow, short (sag.), moderately narrow (tr.). Glabella 71 to 75% (mean 72%, n = 5) as long (sag.) as cephalon. Small anterior lobe clearly defined, subpentagonal to semiovate in shape, obtusely angular anteriorly, occupying 32 to 38% (mean 36%, n = 6) of glabellar length. F3 shallowly incised, bowed moderately rearward; posterior glabellar lobe weakly tapered, strongly convex in its posterior half, less so anteriorly. F2 furrows slightly anterior of glabellar mid-length (sag.), weakly developed as faint notches, gently directed anteriorly for a short distance. F1 furrows effaced to weakly developed; glabellar node indistinct, located slightly posterior of F2 furrows; basal lobes small, subtriangular in outline.
</paragraph>
<paragraph id="8B9A7530FFA2FF84FF699B5BFABCF82C" blockId="13.[151,1436,1146,2035]" pageId="13" pageNumber="14">Pygidium rounded to weakly quadrate in outline, 80% as long as wide, strongly convex, short (exsag.) posterolateral spines; border flattened, wide (sag., exsag.) and narrows distinctly toward anterior; border furrow deep and wide (sag., exsag.). Axis wide (tr.) and long (sag.), reaching posterior border furrow; about 68 to 72% (mean 72%, n = 3) as wide as long and occupying 81 to 88% (mean 86%, n = 3) of pygidial length (sag.); slightly constricted across M2 and expanding to become much wider behind F2; posteriorly rounded and wide (tr.); strongly convex. F1 furrow only faintly developed as weak lateral indentations in axial margin; where visible, furrow is bent strongly forward. M1 lobe broad (tr.) being 38% of pygidial width, lateral margins convex, converging posteriorly. F2 furrow effaced to weakly developed in a few individuals, indentations in axial margin develop into a faint furrow crossing the axis; where visible, F2 slightly curved posteriorly behind axial node. M2 lobe constricted, margins variably convex and converging posteriorly. Axial node well developed, of moderate size, elongate, crossing slightly over F2. Posteroaxis occupying about 63% of axial length, with maximum width near posteroaxis mid-length. Proportions of posterior lobe highly variable. Secondary axial node absent.</paragraph>
<paragraph id="8B9A7530FFA2FF84FF699AAAFDA4F810" blockId="13.[151,1436,1146,2035]" box="[199,573,2010,2035]" pageId="13" pageNumber="14">Hypostome and thorax unknown.</paragraph>
</subSubSection>
<caption id="DF5A25B8FFA1FF87FF399BFAFC17F89B" httpUri="https://zenodo.org/record/1202735/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1675,1697]" targetBox="[167,1424,228,1608]" targetPageId="14">
<paragraph id="8B9A7530FFA1FF87FF399BFAFC17F89B" blockId="14.[151,1436,1675,1912]" pageId="14" pageNumber="15">
<emphasis id="B951A922FFA1FF87FF399BFAFE8CF942" bold="true" box="[151,277,1675,1697]" pageId="14" pageNumber="15">FIGURE 6.</emphasis>
<emphasis id="B951A922FFA1FF87FEB19BFAFE37F942" box="[287,430,1675,1697]" italics="true" pageId="14" pageNumber="15">Hadragnostus</emphasis>
<taxonomicName id="4C250EB3FFA1FF87FE169BFAFC90F942" authority="Jago &amp; Cooper, 2005" authorityName="Jago &amp; Cooper" authorityYear="2005" box="[440,777,1675,1697]" class="Trilobita" family="Proasaphiscidae" genus="Hebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="helixensis">
<emphasis id="B951A922FFA1FF87FE169BFAFD85F942" box="[440,540,1675,1697]" italics="true" pageId="14" pageNumber="15">helixensis</emphasis>
Jago &amp; Cooper, 2005
</taxonomicName>
. All specimens come from sample GOY/97. A, L, CPC42204, partial cephalon; A, dorsal view; L, oblique anterolateral view. B, M, O, CPC42205, partial cephalon; B, dorsal view; M, anterior view; O, lateral view. C, CPC42206, partial cephalon. D, CPC42207, partial cephalon. E, CPC442208, partial cephalon. F, CPC42209, partial cephalon. G, CPC42210, partial cephalon. H, CPC42211, partial cephalon. I, CPC42212, partial cephalon. J, CPC42213, partial cephalon. K, CPC42214, partial cephalon. N, P, X CPC42215, partial pygidium; N, posterior view; P, lateral view; X, dorsal view. Q, CPC42216, partial pygidium. R, CPC42217, partial pygidium. S, CPC42218, partial pygidium. T, CPC42219, partial pygidium. U, CPC42220, partial pygidium. V, CPC42221, partial pygidium. W, CPC42222, partial pygidium. Y, CPC42223, partial pygidium. All scale bars are 1 mm.
</paragraph>
</caption>
<subSubSection id="C33F26BBFFA1FF86FF699AEEFCABFCDB" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8B9A7530FFA1FF86FF699AEEFE8BFDFB" blockId="14.[151,1436,1951,2012]" lastBlockId="15.[151,1437,151,968]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="B951A922FFA1FF87FF699AEEFED5F85B" bold="true" box="[199,332,1951,1976]" pageId="14" pageNumber="15">Discussion.</emphasis>
The Goyder Formation specimens are considered conspecific with
<emphasis id="B951A922FFA1FF87FBE59AEEFB70F85B" box="[1099,1257,1951,1976]" italics="true" pageId="14" pageNumber="15">Hadragnostus</emphasis>
<taxonomicName id="4C250EB3FFA1FF87FB5C9AEEFED3F83F" authority="Jago &amp; Cooper, 2005" authorityName="Jago &amp; Cooper" authorityYear="2005" class="Trilobita" family="Proasaphiscidae" genus="Hebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="helixensis">
<emphasis id="B951A922FFA1FF87FB5C9AEEFAF9F85B" box="[1266,1376,1951,1976]" italics="true" pageId="14" pageNumber="15">helixensis</emphasis>
Jago &amp; Cooper, 2005
</taxonomicName>
from the Mindyallan Spur Formation in
<collectingCountry id="F33235A0FFA1FF87FCBE9AB2FC11F83F" box="[784,904,1987,2012]" name="Antarctica" pageId="14" pageNumber="15">Antarctica</collectingCountry>
. Shared features include: a wide anterior border furrow; an effaced to faintly developed median preglabellar furrow; a large anteroglabella; F3 that is slightly curved posteriorly; a wide (tr.) pygidial axis greater than 50% of pygidial width (tr.); F1 and F2 on the pygidium that are essentially effaced except in exfoliated specimens (e.g.
<figureCitation id="131E69B5FFA0FF86FCC49DAEFC79FF1B" box="[874,992,223,248]" captionStart="FIGURE 6" captionStartId="14.[151,250,1675,1697]" captionTargetBox="[167,1424,228,1608]" captionTargetId="figure@14.[150,1437,221,1622]" captionTargetPageId="14" captionText="FIGURE 6. Hadragnostus helixensis Jago &amp; Cooper, 2005. All specimens come from sample GOY/97. A, L, CPC42204, partial cephalon; A, dorsal view; L, oblique anterolateral view. B, M, O, CPC42205, partial cephalon; B, dorsal view; M, anterior view; O, lateral view. C, CPC42206, partial cephalon. D, CPC42207, partial cephalon. E, CPC442208, partial cephalon.F,CPC42209, partialcephalon.G,CPC42210,partial cephalon.H, CPC42211, partial cephalon.I, CPC42212, partial cephalon. J, CPC42213, partial cephalon. K, CPC42214, partial cephalon. N, P, X CPC42215, partial pygidium; N, posterior view; P, lateral view; X, dorsal view. Q, CPC42216, partial pygidium. R, CPC42217, partial pygidium. S, CPC42218, partial pygidium. T, CPC42219, partial pygidium. U, CPC42220, partial pygidium. V, CPC42221, partial pygidium. W, CPC42222, partial pygidium.Y, CPC42223, partial pygidium. All scale bars are 1 mm." httpUri="https://zenodo.org/record/1202735/files/figure.png" pageId="15" pageNumber="16">Fig. 6P, X</figureCitation>
); a pygidial axis that is constricted (tr.) around M2 and expands (tr.) in the posteroaxis; and a wide posterior border furrow. Only the absence of a node just posterior to F2 on the posteroglabella and the presence of finely granulose ornament differentiate the Goyder Formation material. As the posteroglabellar node and ornament is often difficult to discern in the considerably smaller Antarctic specimens (Jago &amp; Cooper 2005, fig. 2A, B, E, F, H, I), it is likely these differences represent ontogenetic variation. The posteroaxis on the pygidium in the
<typeStatus id="549ECB92FFA0FF86FCF29CE5FC27FE4E" box="[860,958,404,429]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
and a
<typeStatus id="549ECB92FFA0FF86FBA29CE4FBF5FE4E" box="[1036,1132,405,429]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
specimen (Jago &amp; Cooper 2005, fig. 2C, D) appears less rounded posteriorly than those of the Goyder Formation material (
<figureCitation id="131E69B5FFA0FF86FB709CC6FF62FE17" captionStart="FIGURE 6" captionStartId="14.[151,250,1675,1697]" captionTargetBox="[167,1424,228,1608]" captionTargetId="figure@14.[150,1437,221,1622]" captionTargetPageId="14" captionText="FIGURE 6. Hadragnostus helixensis Jago &amp; Cooper, 2005. All specimens come from sample GOY/97. A, L, CPC42204, partial cephalon; A, dorsal view; L, oblique anterolateral view. B, M, O, CPC42205, partial cephalon; B, dorsal view; M, anterior view; O, lateral view. C, CPC42206, partial cephalon. D, CPC42207, partial cephalon. E, CPC442208, partial cephalon.F,CPC42209, partialcephalon.G,CPC42210,partial cephalon.H, CPC42211, partial cephalon.I, CPC42212, partial cephalon. J, CPC42213, partial cephalon. K, CPC42214, partial cephalon. N, P, X CPC42215, partial pygidium; N, posterior view; P, lateral view; X, dorsal view. Q, CPC42216, partial pygidium. R, CPC42217, partial pygidium. S, CPC42218, partial pygidium. T, CPC42219, partial pygidium. U, CPC42220, partial pygidium. V, CPC42221, partial pygidium. W, CPC42222, partial pygidium.Y, CPC42223, partial pygidium. All scale bars are 1 mm." httpUri="https://zenodo.org/record/1202735/files/figure.png" pageId="15" pageNumber="16">Fig. 6Q, S, T, U, W, X, Y</figureCitation>
). Two smaller pygidia figured by Jago &amp; Cooper (2005, fig. 2G, K) demonstrate this feature is variably developed.
</paragraph>
<paragraph id="8B9A7530FFA0FF86FF699F55FCABFCDB" blockId="15.[151,1437,151,968]" pageId="15" pageNumber="16">
<emphasis id="B951A922FFA0FF86FF699F55FE47FDDE" box="[199,478,548,573]" italics="true" pageId="15" pageNumber="16">
Hadragnostus
<taxonomicName id="4C250EB3FFA0FF86FEDE9F55FE47FDDE" authorityName="Jago &amp; Cooper" authorityYear="2005" box="[368,478,548,573]" class="Trilobita" family="Proasaphiscidae" genus="Hebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="helixensis">helixensis</taxonomicName>
</emphasis>
differs from the
<typeStatus id="549ECB92FFA0FF86FD099F54FD4EFDDE" box="[679,727,549,573]" pageId="15" pageNumber="16">type</typeStatus>
species,
<emphasis id="B951A922FFA0FF86FCEA9F54FC13FDDE" box="[836,906,548,573]" italics="true" pageId="15" pageNumber="16">H. las</emphasis>
Öpik, 1967, in having a much wider anterior border, a narrower (tr.) median preglabellar furrow, a pygidium that is longer (sag.) than wide (tr.), a wider (tr.) posteroaxis, and a much wider posterior border. Likewise,
<taxonomicName id="4C250EB3FFA0FF86FC969F1CFC53FD66" authorityName="Jago &amp; Cooper" authorityYear="2005" box="[824,970,620,645]" class="Trilobita" family="Proasaphiscidae" genus="Hebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="helixensis">
<emphasis id="B951A922FFA0FF86FC969F1CFCD5FD67" box="[824,844,621,644]" italics="true" pageId="15" pageNumber="16">H</emphasis>
.
<emphasis id="B951A922FFA0FF86FCF29F1DFC53FD66" box="[860,970,620,645]" italics="true" pageId="15" pageNumber="16">helixensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C250EB3FFA0FF86FBCA9F1CFF4BFD4B" authority="Fortey &amp; Rushton, 1976" authorityName="Fortey &amp; Rushton" authorityYear="1976" class="Trilobita" family="Proasaphiscidae" genus="Hebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="edax">
<emphasis id="B951A922FFA0FF86FBCA9F1CFB27FD66" box="[1124,1214,620,645]" italics="true" pageId="15" pageNumber="16">H. edax</emphasis>
Fortey &amp; Rushton, 1976
</taxonomicName>
, described from the Guzhangian of
<collectingCountry id="F33235A0FFA0FF86FDC09FE1FD07FD44" box="[622,670,656,679]" name="Iran" pageId="15" pageNumber="16">Iran</collectingCountry>
, in having a much wider anterior border, a narrower (tr.) pygidium that is longer sagittally, a greater expansion (tr.) behind M2 of the posteroaxis, and a much wider posterior border.
<emphasis id="B951A922FFA0FF86FF399FA6FE32FD13" box="[151,427,727,752]" italics="true" pageId="15" pageNumber="16">
Hadragnostus
<taxonomicName id="4C250EB3FFA0FF86FE939FA6FE32FD13" authorityName="Jago &amp; Cooper" authorityYear="2005" box="[317,427,727,752]" class="Trilobita" family="Proasaphiscidae" genus="Hebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="helixensis">helixensis</taxonomicName>
</emphasis>
is closest to
<taxonomicName id="4C250EB3FFA0FF86FD939FA8FD5EFD13" baseAuthorityName="Lochman in Lochman &amp; Duncan" baseAuthorityYear="1944" box="[573,711,727,752]" class="Trilobita" family="Proasaphiscidae" genus="Hebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="modestus">
<emphasis id="B951A922FFA0FF86FD939FA8FDC8FD13" box="[573,593,729,752]" italics="true" pageId="15" pageNumber="16">H</emphasis>
.
<emphasis id="B951A922FFA0FF86FDF19FA6FD5EFD13" box="[607,711,727,752]" italics="true" pageId="15" pageNumber="16">modestus</emphasis>
</taxonomicName>
from the Guzhangian of South
<collectingCountry id="F33235A0FFA0FF86FB829FA6FBF6FD13" box="[1068,1135,727,752]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
(
<bibRefCitation id="EFB408C1FFA0FF86FBD09FA9FE92FCF7" pageId="15" pageNumber="16" refId="ref44413" refString="Peng, S. C. &amp; Robison, R. A. (2000) Agnostoid biostratigraphy across the Middle-Upper Cambrian boundary in Hunan, China. Paleontological Society Memoir, 53, 1 - 104. https: // doi. org / 10.1666 / 0022 - 3360 (2000) 53 [1: ABATMC] 2.0. CO; 2" type="journal article">Peng &amp; Robison 2000, p. 32, fig. 23</bibRefCitation>
), but
<taxonomicName id="4C250EB3FFA0FF86FEE79F8CFE40FCF6" authorityName="Jago &amp; Cooper" authorityYear="2005" box="[329,473,764,789]" class="Trilobita" family="Proasaphiscidae" genus="Hebeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="helixensis">
<emphasis id="B951A922FFA0FF86FEE79F8CFE40FCF6" box="[329,473,764,789]" italics="true" pageId="15" pageNumber="16">H. helixensis</emphasis>
</taxonomicName>
differs in possessing a moderately wider anterior border, a narrower (tr.) pygidial axis, and a slightly more posteriorly situated pygidial axial node.
</paragraph>
</subSubSection>
<subSubSection id="C33F26BBFFA0FF86FF699E35FD9CFC63" pageId="15" pageNumber="16" type="description">
<paragraph id="8B9A7530FFA0FF86FF699E35FD9CFC63" blockId="15.[151,1437,151,968]" pageId="15" pageNumber="16">
<emphasis id="B951A922FFA0FF86FF699E35FEC0FCBE" bold="true" box="[199,345,836,861]" pageId="15" pageNumber="16">Occurrence.</emphasis>
GOY section horizons 49.4, 73.2, 83.9 and
<quantity id="4CDDD8D5FFA0FF86FCF69E35FC31FCBF" box="[856,936,836,860]" metricMagnitude="1" metricUnit="m" metricValue="9.19" pageId="15" pageNumber="16" unit="m" value="91.9">91.9 m</quantity>
(
<figureCitation id="131E69B5FFA0FF86FC179E35FC66FCBF" box="[953,1023,836,861]" captionStart="FIGURE 3" captionStartId="9.[151,250,1832,1854]" captionTargetBox="[189,1384,215,1785]" captionTargetId="figure@9.[174,1416,195,1820]" captionTargetPageId="9" captionText="FIGURE 3. Generalised lithostratigraphy and taxon ranges through the Goyder Formation at the GOY section. Sampled horizons are indicated on the right side of the section, with sample numbers representing tape meterage along the section line (see Fig.2). Measurements on the left side of the section are in meters (true thickness) from the base of the Goyder Formation. SF = Shannon Formation, PSS = Pacoota Sandstone. Solid black circles represent the occurrences of taxa present in the GOY section. Hollow circles represent the approximate stratigraphic occurrences of taxa found at nearby spot localities. Ranges of taxa found only atspot localities approximate, and has been inferred using lithological, faunal and geographical information." httpUri="https://zenodo.org/record/1202729/files/figure.png" pageId="15" pageNumber="16">Fig. 3</figureCitation>
). Also recovered at
<collectingCountry id="F33235A0FFA0FF86FB449E35FA89FCBE" box="[1258,1296,836,861]" name="American Samoa" pageId="15" pageNumber="16">AS</collectingCountry>
167,
<collectingCountry id="F33235A0FFA0FF86FAE99E35FAF4FCBE" box="[1351,1389,836,861]" name="American Samoa" pageId="15" pageNumber="16">AS</collectingCountry>
168,
<collectingCountry id="F33235A0FFA0FF86FF399E19FF24FC63" box="[151,189,872,896]" name="American Samoa" pageId="15" pageNumber="16">AS</collectingCountry>
178,
<collectingCountry id="F33235A0FFA0FF86FF5D9E16FE8FFC63" box="[243,278,871,896]" name="American Samoa" pageId="15" pageNumber="16">AS</collectingCountry>
256 and GOYWEST.
</paragraph>
</subSubSection>
<subSubSection id="C33F26BBFFA0FF86FF699EFDFB0AFC2B" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8B9A7530FFA0FF86FF699EFDFB0AFC2B" blockId="15.[151,1437,151,968]" pageId="15" pageNumber="16">
<emphasis id="B951A922FFA0FF86FF699EFDFEF9FC46" bold="true" box="[199,352,908,933]" pageId="15" pageNumber="16">Distribution.</emphasis>
Goyder Formation, Amadeus Basin,
<collectingRegion id="49E1BBD2FFA0FF86FCB79EFDFC69FC46" box="[793,1008,908,933]" country="Australia" name="Northern Territory" pageId="15" pageNumber="16">Northern Territory</collectingRegion>
; Spurs Formation, Northern
<collectingRegion id="49E1BBD2FFA0FF86FAEC9EFDFA05FC46" box="[1346,1436,908,933]" country="Australia" name="Victoria" pageId="15" pageNumber="16">Victoria</collectingRegion>
Land,
<collectingCountry id="F33235A0FFA0FF86FF709EDEFECFFC2B" box="[222,342,943,968]" name="Antarctica" pageId="15" pageNumber="16">Antarctica</collectingCountry>
. All occurrences are Cambrian Series 3, Guzhangian (Mindyallan) in age.
</paragraph>
</subSubSection>
</treatment>
</document>