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<figureCitation id="131E3FFFFF9A776BFC9091ECBD8ECB9E" box="[834,991,1230,1254]" captionStart-0="Figure 33" captionStart-1="Figures 34–38" captionStart-2="Figures 39–40" captionStart-3="Figures 41–44" captionStart-4="Figures 45–48" captionStart-5="Figures 49–54" captionStart-6="Figures 55–61" captionStart-7="Figures 62–64" captionStartId-0="14.[113,181,1799,1821]" captionStartId-1="15.[116,193,1986,2008]" captionStartId-2="16.[113,190,1317,1339]" captionStartId-3="17.[112,189,1982,2004]" captionStartId-4="18.[115,192,1574,1596]" captionStartId-5="19.[113,190,1296,1318]" captionStartId-6="20.[115,192,1933,1955]" captionStartId-7="21.[115,192,1351,1373]" captionTargetBox-0="[113,1474,234,1785]" captionTargetBox-1="[167,1429,241,1953]" captionTargetBox-2="[113,1474,234,1309]" captionTargetBox-3="[126,1457,335,1922]" captionTargetBox-4="[113,1474,234,1565]" captionTargetBox-5="[113,1474,234,1286]" captionTargetBox-6="[125,1465,236,1909]" captionTargetBox-7="[113,1474,234,1341]" captionTargetId-0="figure-155@14.[113,1476,234,1785]" captionTargetId-1="figure-44@15.[113,1474,234,1956]" captionTargetId-2="figure-390@16.[113,1474,234,1309]" captionTargetId-3="figure-41@17.[113,1474,299,1922]" captionTargetId-4="figure-259@18.[113,1474,234,1565]" captionTargetId-5="figure-392@19.[113,1474,234,1286]" captionTargetId-6="figure-111@20.[113,1474,226,1909]" captionTargetId-7="figure-305@21.[113,1474,234,1341]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionTargetPageId-3="17" captionTargetPageId-4="18" captionTargetPageId-5="19" captionTargetPageId-6="20" captionTargetPageId-7="21" captionText-0="Figure 33. Scorpiops nagphani sp. n., female, paratype BNHS SC 256 in vivo habitus." captionText-1="Figures 34–38. Scorpiops nagphani sp. n., male, holotype, dorsal (34) and ventral (35) views, sternopectinal area (36), metasomal segment V and telson in lateral view (37) and chelicera in dorsal view (38)." captionText-2="Figures 39–40. Scorpiops nagphani sp. n., male, holotype in dorsal (39) and ventral (40) views under UV light." captionText-3="Figures 41–44. Scorpiops nagphani sp. n., female, paratype, BNHS SC 256, in dorsal (41) ventral (42) views, sternopectinal area (43) and metasomal segment V and telson in lateral view (44)." captionText-4="Figures 45–48: Scorpiops nagphani sp. n. Figures 45–46. Male, holotype, carapace under white light (45) and UV fluorescence (46). Figures 47–48. Female, paratype, BHNS SC 256, carapace under white light (47) and UV fluorescence (48)." captionText-5="Figures 49–54. Scorpiops nagphani sp. n., male, holotype. Figures 49–54. Pedipalp chela dorsal (49), ventral (50) and dorsoexternal (51), patella external (52), dorsal (53) and ventral (54) views under UV fluorescence. Trichobothrial pattern indicated by yellow circles." captionText-6="Figures 55–61. Scorpiops nagphani sp. n. Figures 55–58. Scorpiops nagphani sp. n., male, holotype, left legs I-IV, retrolateral aspect. Figures 59–61. Scorpiops nagphani sp. n., male, paratype, INHER-259, hemispermatophore internal (59), external (60) views and capsular region in internal view (61). Abbrevations: distal lamina (dl), capsule (c), trunk (t), pedicel (p), distal posterior lobe (dpl), lateral hook (lh), terminal membrane of the sperm duct (tmsd) and basal carina (bc)." captionText-7="Figures 62–64. Type locality of Scorpiops nagphani sp. n., aerial view of semi-evergreen forest inside Bhimashankar wildlife sanctuary (BWLS), very close to the type locality (62), view of Kolvan waterfalls inside BWLS where the new species was observed (63) and Scorpiops nagphani sp. n., female (specimen not collected) in vivo habitus with juveniles (64)." figureDoi-0="http://doi.org/10.5281/zenodo.5742559" figureDoi-1="http://doi.org/10.5281/zenodo.5742561" figureDoi-2="http://doi.org/10.5281/zenodo.5742563" figureDoi-3="http://doi.org/10.5281/zenodo.5742565" figureDoi-4="http://doi.org/10.5281/zenodo.5742567" figureDoi-5="http://doi.org/10.5281/zenodo.5742569" figureDoi-6="http://doi.org/10.5281/zenodo.5742571" figureDoi-7="http://doi.org/10.5281/zenodo.5742573" httpUri-0="https://zenodo.org/record/5742559/files/figure.png" httpUri-1="https://zenodo.org/record/5742561/files/figure.png" httpUri-2="https://zenodo.org/record/5742563/files/figure.png" httpUri-3="https://zenodo.org/record/5742565/files/figure.png" httpUri-4="https://zenodo.org/record/5742567/files/figure.png" httpUri-5="https://zenodo.org/record/5742569/files/figure.png" httpUri-6="https://zenodo.org/record/5742571/files/figure.png" httpUri-7="https://zenodo.org/record/5742573/files/figure.png" pageId="13" pageNumber="12">Figures 33–64</figureCitation>
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<figureCitation id="131E3FFFFF9A776BFC3891ECBA2ECB9E" box="[1002,1151,1230,1254]" captionStart="Figures 127–138" captionStartId="41.[116,193,1928,1950]" captionTargetBox="[113,1474,245,1897]" captionTargetId="figure-102@41.[113,1474,234,1897]" captionTargetPageId="41" captionText="Figures 127–138: Figures 127–130: Comparison of left pedipalps. Figures 131–134: Comparison of metasomal segments II-IV in lateral view. Figures 135–138: Comparison of pedipalp patella dorsal view with internal aspect tubercles. Figures 127, 131, 135. Scorpiops vrushchik sp. n., male, holotype. Figures 128, 132, 136. Scorpiops nagphani sp. n., male, holotype. Figures 129, 133, 137. Scorpiops neera sp. n., male, holotype. Figures 130, 134, 138. Scorpiops lioneli sp. n., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.5742617" httpUri="https://zenodo.org/record/5742617/files/figure.png" pageId="13" pageNumber="12">128, 132, 136</figureCitation>
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<figureCitation id="131E3FFFFF9A776BFB5891ECBAB8CB9E" box="[1162,1257,1230,1254]" captionStart="Figures 139–152" captionStartId="43.[115,192,1183,1205]" captionTargetBox="[113,1474,234,1167]" captionTargetId="figure-329@43.[113,1474,234,1167]" captionTargetPageId="43" captionText="Figures 139–152: Figures 139–142: Comparison of dorsolateral granules on anal rim of metasomal segment V in dorsal view. Figures 143–152: Comparison of dentition on movable finger of chela with field emission scanning electron microscopy (FESEM) images in dorsal view. Figures 139, 149. Scorpiops vrushchik sp. n., male, holotype. Figures 140, 150. Scorpiops nagphani sp. n., male, holotype. Figures 141, 151. Scorpiops neera sp. n., male, holotype. Figures 142, 152. Scorpiops lioneli sp. n., male, holotype. Figure 143. S. tenuicauda, male, INHER-214. Figure 144. S. satarensis, male, INHER-213. Figure 145. S. deccanensis, male, INHER-86. Figure 146. S. maharashtraensis, male, INHER-179. Figure 147. S. phaltanensis, male, paratype, INHER-64. Figure 148. S. telbaila, male, paratype, INHER-123." figureDoi="http://doi.org/10.5281/zenodo.5742621" httpUri="https://zenodo.org/record/5742621/files/figure.png" pageId="13" pageNumber="12">140, 150</figureCitation>
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<figureCitation id="131E3FFFFF9A776BFB2791ECBB04CB9E" box="[1269,1365,1230,1254]" captionStart-0="Figure 153" captionStart-1="Figure 154" captionStart-2="Figure 155" captionStart-3="Figure 156" captionStart-4="Figure 157" captionStartId-0="45.[113,181,1216,1238]" captionStartId-1="45.[117,185,1992,2014]" captionStartId-2="46.[113,181,942,964]" captionStartId-3="47.[119,187,1584,1606]" captionStartId-4="48.[115,183,1359,1381]" captionTargetBox-0="[114,1474,234,1198]" captionTargetBox-1="[135,1438,1286,1971]" captionTargetBox-2="[117,1465,232,922]" captionTargetBox-3="[145,1432,286,1493]" captionTargetBox-4="[113,1474,257,1330]" captionTargetId-0="figure-7@45.[113,1474,234,1198]" captionTargetId-1="figure-9@45.[113,1474,1260,1983]" captionTargetId-2="figure-631@46.[113,1474,229,925]" captionTargetId-3="figure-220@47.[115,1476,232,1570]" captionTargetId-4="figure-208@48.[113,1474,257,1330]" captionTargetPageId-0="45" captionTargetPageId-1="45" captionTargetPageId-2="46" captionTargetPageId-3="47" captionTargetPageId-4="48" captionText-0="Figure 153. Distribution of Scorpiops species from Peninsular India. Localities with a star indicate new species described in this study." captionText-1="Figure 154. Boxplots showing variations in ratios of pedipalp chela length to width in males for all Scorpiops species from Peninsular India (except S. pachmarhicus due to unavailability of sufficient males)." captionText-2="Figure 155. Projection of first two Principal Component factors explaining 81.89% variation among the nine species of the genus Scorpiops from Peninsular India (except S. pachmarhicus and S. satarensis due to unavailability of sufficient males)." captionText-3="Figure 156. Maximum Likelihood phylogenetic tree (ML) for Scorpiops. Values along the nodes are bootstraps for 1000 iterations." captionText-4="Figure 157. Bayesian phylogenetic tree for Scorpiops. Values along the nodes are Bayesian posterior probabilities for Bayesian Inference and Bayesian Poisson Tree Process (bPTP) respectively. Vertical bars represent delimitation analyses results." figureDoi-0="http://doi.org/10.5281/zenodo.5742623" figureDoi-1="http://doi.org/10.5281/zenodo.5742625" figureDoi-2="http://doi.org/10.5281/zenodo.5742627" figureDoi-3="http://doi.org/10.5281/zenodo.5742631" figureDoi-4="http://doi.org/10.5281/zenodo.5742633" httpUri-0="https://zenodo.org/record/5742623/files/figure.png" httpUri-1="https://zenodo.org/record/5742625/files/figure.png" httpUri-2="https://zenodo.org/record/5742627/files/figure.png" httpUri-3="https://zenodo.org/record/5742631/files/figure.png" httpUri-4="https://zenodo.org/record/5742633/files/figure.png" pageId="13" pageNumber="12">153–157</figureCitation>
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<tableCitation id="C6A716C1FF9A776BFA8D91ECBBEBCB9E" box="[1375,1466,1230,1254]" captionStart="Table 2" captionStartId="22.[113,169,1998,2019]" captionText="Table 2. Morphometric data for Scorpiops nagphani sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to median width), depth (D)." pageId="13" pageNumber="12">Tables 2</tableCitation>
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<tableCitation id="C6A716C1FF9A776BFB8191CCBA33CA7E" box="[1107,1122,1262,1286]" captionStart="Table 3" captionStartId="23.[113,169,1982,2003]" captionText="Table 3. Morphometric data for Scorpiops neera sp. n. and a male paratype of S. nagphani sp. n.Abbreviations: length (L), width (W, in carapace it corresponds to median width), depth (D)." pageId="13" pageNumber="12">3</tableCitation>
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<tableCitation id="C6A716C1FF9A776BFBBC91CCBAC6CA7E" box="[1134,1175,1262,1286]" captionStart-0="Table 5" captionStart-1="Table 6" captionStart-2="Table 7" captionStartId-0="44.[115,171,1066,1087]" captionStartId-1="44.[115,171,1470,1491]" captionStartId-2="44.[115,171,1970,1991]" captionText-0="Table 5. Pairwise uncorrected raw distances (%) expressed as minimum–maximum based on COI gene sequence for Scorpiops species from Maharashtra, India. Values in brackets are intra-clade distances." captionText-1="Table 6: Primers used for PCR amplification and sequencing of (COI) mitochondrial gene." captionText-2="Table 7: Voucher numbers and GenBank accession numbers for the sequence data used for the phylogenetic analysis." pageId="13" pageNumber="12">5–7</tableCitation>
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<collectingCountry id="F33263EAFF9A776BFB26904CBB60CAFE" box="[1268,1329,1390,1414]" name="India" pageId="13" pageNumber="12">India</collectingCountry>
</emphasis>
, 
<collectingRegion id="49E1ED98FF9A776BFAEF904CBD39CADE" country="India" name="Maharashtra" pageId="13" pageNumber="12">Maharashtra State</collectingRegion>
, 
<collectingCounty id="62FB5BF6FF9A776BFCA990ACBA42CADE" box="[891,1043,1422,1446]" pageId="13" pageNumber="12">Pune District</collectingCounty>
, 
<collectingMunicipality id="6BFEB900FF9A776BFBF790ACBA89CADE" box="[1061,1240,1422,1446]" pageId="13" pageNumber="12">Nigdale Village</collectingMunicipality>
, near 
<location id="8EFA75A1FF9A776BFAFA90ACBDAACABE" LSID="urn:lsid:plazi:treatment:038C926CFF9A7773FC09918EBAA2C891:8EFA75A1FF9A776BFAFA90ACBDAACABE" country="India" county="Pune District" latitude="19.077778" longLatPrecision="21" longitude="73.55889" municipality="Nigdale Village" name="Bhimashankar Wildlife Sanctuary" pageId="13" pageNumber="12" stateProvince="Maharashtra">Bhimashankar Wildlife Sanctuary</location>
, 
<geoCoordinate id="EE1145BDFF9A776BFBD7908CBAD3CABE" box="[1029,1154,1454,1478]" degrees="19" direction="north" minutes="04" orientation="latitude" pageId="13" pageNumber="12" precision="15" seconds="40" value="19.077778">19°04'40&quot;N</geoCoordinate>
<geoCoordinate id="EE1145BDFF9A776BFB5B908CBB56CABE" box="[1161,1287,1454,1478]" degrees="73" direction="east" minutes="33" orientation="longitude" pageId="13" pageNumber="12" precision="15" seconds="32" value="73.55889">73°33'32&quot;E</geoCoordinate>
</materialsCitation>
; 
<collectionCode id="ED34BBBFFF9A776BFAC0908CBB0CCABE" box="[1298,1373,1454,1478]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="13" pageNumber="12">BNHS</collectionCode>
.
</paragraph>
<paragraph id="8B9A237AFF9A776BFCE290CCBA82C99E" blockId="13.[816,1474,1518,1766]" pageId="13" pageNumber="12">
<materialsCitation id="3B4D2927FF9A776BFCE290CCBA82C99E" collectingDate="2020-07-02" collectingDateMax="2020-07-21" collectingDateMin="2020-07-02" collectionCode="TYPE, BNHS" collectorName="All &amp; S. Sulakhe &amp; Sulakhe, S &amp; Deshpande, M &amp; Ketkar, A &amp; G. Gowande." country="India" county="Pune District" elevation="963" latitude="19.077778" location="Bhimashankar Wildlife Sanctuary" longLatPrecision="21" longitude="73.55889" municipality="Nigdale Village" pageId="13" pageNumber="12" specimenCode="SC 254, SC 255, SC 257, SC 256" specimenCount="9" specimenCount-female="4" specimenCount-male="5" stateProvince="Maharashtra" typeStatus="holotype">
<collectionCode id="ED34BBBFFF9A776BFCE290CCBD37C97D" box="[816,870,1518,1541]" pageId="13" pageNumber="12">TYPE</collectionCode>
MATERIAL. 
<emphasis id="B951FF68FF9A776BFC3C90CCBA7AC97E" bold="true" box="[1006,1067,1518,1542]" pageId="13" pageNumber="12">
<collectingCountry id="F33263EAFF9A776BFC3C90CCBA7AC97E" box="[1006,1067,1518,1542]" name="India" pageId="13" pageNumber="12">India</collectingCountry>
</emphasis>
, 
<emphasis id="B951FF68FF9A776BFB9390CCBB40C97D" box="[1089,1297,1518,1542]" italics="true" pageId="13" pageNumber="12">
<collectingRegion id="49E1ED98FF9A776BFB9390CCBB40C97D" box="[1089,1297,1518,1542]" country="India" name="Maharashtra" pageId="13" pageNumber="12">Maharashtra State</collectingRegion>
</emphasis>
, 
<collectingCounty id="62FB5BF6FF9A776BFAF590CCBBEFC97E" box="[1319,1470,1518,1542]" pageId="13" pageNumber="12">Pune District</collectingCounty>
, 
<collectingMunicipality id="6BFEB900FF9A776BFCE2932CBDB6C95E" box="[816,999,1550,1574]" pageId="13" pageNumber="12">Nigdale Village</collectingMunicipality>
, near 
<location id="8EFA75A1FF9A776BFBEF932CBBEFC95E" LSID="urn:lsid:plazi:treatment:038C926CFF9A7773FC09918EBAA2C891:8EFA75A1FF9A776BFBEF932CBBEFC95E" box="[1085,1470,1550,1574]" country="India" county="Pune District" latitude="19.077778" longLatPrecision="21" longitude="73.55889" municipality="Nigdale Village" name="Bhimashankar Wildlife Sanctuary" pageId="13" pageNumber="12" stateProvince="Maharashtra">Bhimashankar Wildlife Sanctuary</location>
, 
<geoCoordinate id="EE1145BDFF9A776BFCE2930CBDFFC93E" box="[816,942,1582,1606]" degrees="19" direction="north" minutes="04" orientation="latitude" pageId="13" pageNumber="12" precision="15" seconds="40" value="19.077778">19°04'40&quot;N</geoCoordinate>
<geoCoordinate id="EE1145BDFF9A776BFC64930CBA62C93E" box="[950,1075,1582,1606]" degrees="73" direction="east" minutes="33" orientation="longitude" pageId="13" pageNumber="12" precision="15" seconds="32" value="73.55889">73°33'32&quot;E</geoCoordinate>
, 
<elevation id="0008C449FF9A776BFBED930CBA9EC93E" box="[1087,1231,1582,1606]" metricMagnitude="2" metricUnit="m" metricValue="9.629999999999999" pageId="13" pageNumber="12" unit="m" value="963.0">
<quantity id="4CDD8E9FFF9A776BFBED930CBAD5C93D" box="[1087,1156,1582,1606]" metricMagnitude="2" metricUnit="m" metricValue="9.629999999999999" pageId="13" pageNumber="12" unit="m" value="963.0">963 m</quantity>
a. s. l.
</elevation>
, 
<specimenCount id="9D23E8F3FF9A776BFB0F930CBAAFC93E" box="[1245,1278,1582,1606]" pageId="13" pageNumber="12" type="male">1♂</specimenCount>
(
<typeStatus id="549E9DD8FF9A776BFADF930CBB3EC93E" box="[1293,1391,1582,1606]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
, 
<collectionCode id="ED34BBBFFF9A776BFAA9930CBB93C93E" box="[1403,1474,1582,1606]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="13" pageNumber="12">BNHS</collectionCode>
<specimenCode id="DB838B01FF9A776BFCE2936CBDD9C91E" box="[816,904,1614,1638]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="13" pageNumber="12">SC 254</specimenCode>
), 
<specimenCount id="9D23E8F3FF9A776BFC4C936CBDEEC91E" box="[926,959,1614,1638]" pageId="13" pageNumber="12" type="male">4♂</specimenCount>
(
<typeStatus id="549E9DD8FF9A776BFC03936CBA6AC91E" box="[977,1083,1614,1638]" pageId="13" pageNumber="12" type="paratype">paratypes</typeStatus>
, INHER-257, 259, 263, 
<collectionCode id="ED34BBBFFF9A776BFA82936CBBC6C91E" box="[1360,1431,1614,1638]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="13" pageNumber="12">BNHS</collectionCode>
<specimenCode id="DB838B01FF9A776BFA73936CBD0AC9FE" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="13" pageNumber="12">SC 255</specimenCode>
), 
<specimenCount id="9D23E8F3FF9A776BFCA0934CBDC2C9FE" box="[882,915,1646,1670]" pageId="13" pageNumber="12" type="female">2♀</specimenCount>
(
<typeStatus id="549E9DD8FF9A776BFC77934CBA54C9FE" box="[933,1029,1646,1670]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
, INHER- 260, 
<collectionCode id="ED34BBBFFF9A776BFB60934CBAA8C9FE" box="[1202,1273,1646,1670]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="13" pageNumber="12">BNHS</collectionCode>
<specimenCode id="DB838B01FF9A776BFAD1934CBB08C9FE" box="[1283,1369,1646,1670]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="13" pageNumber="12">SC 257</specimenCode>
), 
<date id="FF9B05BAFF9A776BFAA2934CBD39C9DE" pageId="13" pageNumber="12" value="2020-07-21">
<collectingDate id="EFDFFC52FF9A776BFAA2934CBD39C9DE" pageId="13" pageNumber="12" value="2020-07-21">21 July 2020</collectingDate>
</date>
, 
<specimenCount id="9D23E8F3FF9A776BFCA493ACBDC6C9DE" box="[886,919,1678,1702]" pageId="13" pageNumber="12" type="female">2♀</specimenCount>
(
<typeStatus id="549E9DD8FF9A776BFC7B93ACBA58C9DE" box="[937,1033,1678,1702]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
, INHER- 338, 
<collectionCode id="ED34BBBFFF9A776BFB6693ACBAAAC9DE" box="[1204,1275,1678,1702]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="13" pageNumber="12">BNHS</collectionCode>
<specimenCode id="DB838B01FF9A776BFAD793ACBB0AC9DE" box="[1285,1371,1678,1702]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="13" pageNumber="12">SC 256</specimenCode>
), 
<date id="FF9B05BAFF9A776BFAA393ACBD38C9BE" pageId="13" pageNumber="12" value="2020-07-02">
<collectingDate id="EFDFFC52FF9A776BFAA393ACBD38C9BE" pageId="13" pageNumber="12" value="2020-07-02">02 July 2020</collectingDate>
</date>
. 
<collectorName id="26D046ACFF9A776BFCA6938CBDC7C9BE" box="[884,918,1710,1734]" pageId="13" pageNumber="12">All</collectorName>
specimens collected by 
<collectorName id="26D046ACFF9A776BFB71938CBB4BC9BE" box="[1187,1306,1710,1734]" pageId="13" pageNumber="12">S. Sulakhe</collectorName>
, S. 
<collectorName id="26D046ACFF9A776BFA97938CBD1AC99D" pageId="13" pageNumber="12">Deshpande, M</collectorName>
. 
<collectorName id="26D046ACFF9A776BFC8793ECBDEEC99E" box="[853,959,1742,1766]" pageId="13" pageNumber="12">Ketkar, A</collectorName>
. Marathe &amp; 
<collectorName id="26D046ACFF9A776BFB9693ECBA82C99E" box="[1092,1235,1742,1766]" pageId="13" pageNumber="12">G. Gowande.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33F70F1FF9A776BFCE2922CBBF9C89E" pageId="13" pageNumber="12" type="etymology">
<paragraph id="8B9A237AFF9A776BFCE2922CBBF9C89E" blockId="13.[816,1475,1806,2022]" pageId="13" pageNumber="12">
ETYMOLOGY. The specific epithet is noun in opposition, named after a steep hill top with a rocky cliff inside Bhimashankar Wildlife Sanctuary, which is popularly known as “Nagphani” in Marathi language, meaning Cobra (nag) hood (phani), which is very close to the 
<typeStatus id="549E9DD8FF9A776BFC2F92ACBA7BC8DE" box="[1021,1066,1934,1958]" pageId="13" pageNumber="12">type</typeStatus>
locality. The hiking route to Nagphani passes through thick semi-evergreen forest with basaltic boulders where abundance of the new species was observed.
</paragraph>
</subSubSection>
<caption id="DF5A73F2FF997768FFA39225BDE6C865" ID-DOI="http://doi.org/10.5281/zenodo.5742559" ID-Zenodo-Dep="5742559" box="[113,951,1799,1821]" httpUri="https://zenodo.org/record/5742559/files/figure.png" pageId="14" pageNumber="13" startId="14.[113,181,1799,1821]" targetBox="[113,1474,234,1785]" targetPageId="14">
<paragraph id="8B9A237AFF997768FFA39225BDE6C865" blockId="14.[113,951,1799,1821]" box="[113,951,1799,1821]" pageId="14" pageNumber="13">
<emphasis id="B951FF68FF997768FFA39225BE82C864" bold="true" box="[113,211,1799,1821]" pageId="14" pageNumber="13">Figure 33</emphasis>
. 
<taxonomicName id="4C2558F9FF997768FF0D922ABFF1C864" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[223,416,1799,1821]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FF997768FF0D922ABFF1C864" box="[223,416,1799,1821]" italics="true" pageId="14" pageNumber="13">Scorpiops nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FF997768FE74922ABF8AC865" box="[422,475,1800,1821]" pageId="14" pageNumber="13" rank="species">
<emphasis id="B951FF68FF997768FE74922ABFECC865" bold="true" box="[422,445,1800,1821]" pageId="14" pageNumber="13">sp</emphasis>
. 
<emphasis id="B951FF68FF997768FE1A922ABF84C865" bold="true" box="[456,469,1800,1821]" pageId="14" pageNumber="13">n</emphasis>
.
</taxonomicNameLabel>
, female, paratype BNHS SC 256 in vivo habitus.
</paragraph>
</caption>
<subSubSection id="C33F70F1FF997776FFA0921EBCD3C948" lastPageId="16" lastPageNumber="15" pageId="14" pageNumber="13" type="diagnosis">
<paragraph id="8B9A237AFF997776FFA0921EBCD3C948" blockId="14.[114,772,1852,2036]" lastBlockId="16.[113,772,1368,1584]" lastPageId="16" lastPageNumber="15" pageId="14" pageNumber="13">
DIAGNOSIS (
<specimenCount id="9D23E8F3FF997768FED2921EBF49C82C" box="[256,280,1852,1876]" pageId="14" pageNumber="13" type="male">♂</specimenCount>
<specimenCount id="9D23E8F3FF997768FECA921EBF78C82C" box="[280,297,1852,1876]" pageId="14" pageNumber="13" type="female">♀</specimenCount>
). Total length 
<quantity id="4CDD8E9FFF997768FE39921EBC3FC82C" box="[491,622,1852,1876]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="14" pageNumber="13" unit="mm" value="40.0" valueMax="42.0" valueMin="38.0">38–42 mm</quantity>
. Base color uniformly blackish. Pectinal teeth number 
<quantity id="4CDD8E9FFF997768FDA1927EBC93C80C" box="[627,706,1884,1908]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="1.778" metricValueMin="1.524" pageId="14" pageNumber="13" unit="in" value="6.5" valueMax="7.0" valueMin="6.0">6–7 in</quantity>
both sexes, fulcra reduced to absent. Pectine morphology of 
<typeStatus id="549E9DD8FF997768FD04925FBD55C8ED" box="[726,772,1917,1941]" pageId="14" pageNumber="13">type</typeStatus>
P3. Patella of pedipalp with 18–23 (5 
<emphasis id="B951FF68FF997768FDCD92BEBC69C8CC" box="[543,568,1948,1972]" italics="true" pageId="14" pageNumber="13">eb</emphasis>
, 2 
<emphasis id="B951FF68FF997768FDB292BEBCD5C8CC" box="[608,644,1948,1972]" italics="true" pageId="14" pageNumber="13">esb</emphasis>
, 2 
<emphasis id="B951FF68FF997768FD7E92BFBC9AC8CC" box="[684,715,1949,1972]" italics="true" pageId="14" pageNumber="13">em</emphasis>
, 4–8 
<emphasis id="B951FF68FF997768FFA0929FBEC1C8AC" box="[114,144,1981,2004]" italics="true" pageId="14" pageNumber="13">est</emphasis>
, 5–6 
<emphasis id="B951FF68FF997768FF01929FBEB7C8AC" box="[211,230,1981,2004]" italics="true" pageId="14" pageNumber="13">et</emphasis>
) external and 12–14 ventral trichobothria. Chela of pedipalp with 4 ventral trichobothria located on ventral surface. Chelal trichobothrium 
<emphasis id="B951FF68FF997768FB41921EBAE8C823" box="[1171,1209,1852,1883]" italics="true" pageId="14" pageNumber="13">Eb 3</emphasis>
is located in proximal half of manus between trichobothria Dt and Db. Fingers of pedipalps strongly undulate in male and margins undulate in female. Chela length to width ratio 3.7–4.0 in males. Pedipalp movable finger with ca 45–50 IAD, which form second row, parallel with MD (ca 
<quantity id="4CDD8E9FFF997768FBC492FEBA24C88C" box="[1046,1141,2012,2036]" metricMagnitude="0" metricUnit="m" metricValue="1.5875" metricValueMax="1.6509999999999998" metricValueMin="1.524" pageId="14" pageNumber="13" unit="in" value="62.5" valueMax="65.0" valueMin="60.0">60–65 in</quantity>
number); there are also 3–4 ID and 10–11 OD present. Total pedipalp length to mesosomal tergite length in males 1.6–1.9. Tarsomere II of legs with 4–6 stout median ventral spinules. Metasoma I, II and III with ten, metasoma IV with eight and metasoma V with 7 carinae. Dorsal lateral carina on II-IV metasomal segments ending posteriorly into a short spine. Telson elongate and smooth, length to depth ratio 2.5–2.9; annular ring absent.
</paragraph>
</subSubSection>
<caption id="DF5A73F2FF987769FFA692E0BC8AC88D" ID-DOI="http://doi.org/10.5281/zenodo.5742561" ID-Zenodo-Dep="5742561" httpUri="https://zenodo.org/record/5742561/files/figure.png" pageId="15" pageNumber="14" startId="15.[116,193,1986,2008]" targetBox="[167,1429,241,1953]" targetPageId="15">
<paragraph id="8B9A237AFF987769FFA692E0BC8AC88D" blockId="15.[116,1474,1986,2037]" pageId="15" pageNumber="14">
<emphasis id="B951FF68FF987769FFA692E0BF52C8AF" bold="true" box="[116,259,1986,2008]" pageId="15" pageNumber="14">Figures 34–38</emphasis>
. 
<taxonomicName id="4C2558F9FF987769FEDC92E0BF9EC8AF" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[270,463,1986,2007]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FF987769FEDC92E0BF9EC8AF" box="[270,463,1986,2007]" italics="true" pageId="15" pageNumber="14">Scorpiops nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FF987769FE0692E0BC58C8AF" box="[468,521,1986,2007]" pageId="15" pageNumber="14" rank="species">
<emphasis id="B951FF68FF987769FE0692E0BFBAC8AF" bold="true" box="[468,491,1986,2007]" pageId="15" pageNumber="14">sp</emphasis>
. 
<emphasis id="B951FF68FF987769FE2592E0BC55C8AF" bold="true" box="[503,516,1986,2007]" pageId="15" pageNumber="14">n</emphasis>
.
</taxonomicNameLabel>
, male, holotype, dorsal (34) and ventral (35) views, sternopectinal area (36), metasomal segment V and telson in lateral view (37) and chelicera in dorsal view (38).
</paragraph>
</caption>
<caption id="DF5A73F2FF877776FFA39007BAF7CA43" ID-DOI="http://doi.org/10.5281/zenodo.5742563" ID-Zenodo-Dep="5742563" box="[113,1190,1317,1340]" httpUri="https://zenodo.org/record/5742563/files/figure.png" pageId="16" pageNumber="15" startId="16.[113,190,1317,1339]" targetBox="[113,1474,234,1309]" targetPageId="16">
<paragraph id="8B9A237AFF877776FFA39007BAF7CA43" blockId="16.[113,1190,1317,1340]" box="[113,1190,1317,1340]" pageId="16" pageNumber="15">
<emphasis id="B951FF68FF877776FFA39007BF50CA42" bold="true" box="[113,257,1317,1339]" pageId="16" pageNumber="15">Figures 39–40</emphasis>
. 
<taxonomicName id="4C2558F9FF877776FEDE9004BF9CCA42" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[268,461,1317,1339]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FF877776FEDE9004BF9CCA42" box="[268,461,1317,1339]" italics="true" pageId="16" pageNumber="15">Scorpiops nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FF877776FE019004BC59CA43" box="[467,520,1318,1339]" pageId="16" pageNumber="15" rank="species">
<emphasis id="B951FF68FF877776FE019004BFBBCA43" bold="true" box="[467,490,1318,1339]" pageId="16" pageNumber="15">sp</emphasis>
. 
<emphasis id="B951FF68FF877776FE249004BC52CA43" bold="true" box="[502,515,1318,1339]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
, male, holotype in dorsal (39) and ventral (40) views under UV light.
</paragraph>
</caption>
<subSubSection id="C33F70F1FF877773FFA3937BBEA8C8F1" lastPageId="21" lastPageNumber="20" pageId="16" pageNumber="15" type="description">
<paragraph id="8B9A237AFF877776FFA3937BBC9EC909" blockId="16.[113,772,1624,2033]" box="[113,719,1624,1649]" pageId="16" pageNumber="15">
DESCRIPTION (
<specimenCount id="9D23E8F3FF877776FED9937BBF4EC909" box="[267,287,1625,1649]" pageId="16" pageNumber="15" type="male">♂</specimenCount>
<typeStatus id="549E9DD8FF877776FEF4937ABFD7C908" box="[294,390,1624,1648]" pageId="16" pageNumber="15" type="holotype">holotype</typeStatus>
, measurements in 
<tableCitation id="C6A716C1FF877776FD99937ABC92C909" box="[587,707,1624,1649]" captionStart-0="Table 2" captionStart-1="Table 3" captionStartId-0="22.[113,169,1998,2019]" captionStartId-1="23.[113,169,1982,2003]" captionText-0="Table 2. Morphometric data for Scorpiops nagphani sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to median width), depth (D)." captionText-1="Table 3. Morphometric data for Scorpiops neera sp. n. and a male paratype of S. nagphani sp. n.Abbreviations: length (L), width (W, in carapace it corresponds to median width), depth (D)." pageId="16" pageNumber="15">Tables 2–3</tableCitation>
).
</paragraph>
<paragraph id="8B9A237AFF877776FFA3935ABA1ECA88" blockId="16.[113,772,1624,2033]" lastBlockId="16.[816,1475,1368,2032]" pageId="16" pageNumber="15">
<emphasis id="B951FF68FF877776FFA3935ABEBDC9E8" bold="true" box="[113,236,1656,1680]" pageId="16" pageNumber="15">Coloration</emphasis>
(in preservation) (
<figureCitation id="131E3FFFFF877776FE68935ABC16C9E8" box="[442,583,1656,1681]" captionStart-0="Figure 33" captionStart-1="Figures 34–38" captionStartId-0="14.[113,181,1799,1821]" captionStartId-1="15.[116,193,1986,2008]" captionTargetBox-0="[113,1474,234,1785]" captionTargetBox-1="[167,1429,241,1953]" captionTargetId-0="figure-155@14.[113,1476,234,1785]" captionTargetId-1="figure-44@15.[113,1474,234,1956]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="Figure 33. Scorpiops nagphani sp. n., female, paratype BNHS SC 256 in vivo habitus." captionText-1="Figures 34–38. Scorpiops nagphani sp. n., male, holotype, dorsal (34) and ventral (35) views, sternopectinal area (36), metasomal segment V and telson in lateral view (37) and chelicera in dorsal view (38)." figureDoi-0="http://doi.org/10.5281/zenodo.5742559" figureDoi-1="http://doi.org/10.5281/zenodo.5742561" httpUri-0="https://zenodo.org/record/5742559/files/figure.png" httpUri-1="https://zenodo.org/record/5742561/files/figure.png" pageId="16" pageNumber="15">Figs. 33– 35</figureCitation>
, 
<figureCitation id="131E3FFFFF877776FD87935BBCCEC9E9" box="[597,671,1657,1681]" captionStart="Figures 41–44" captionStartId="17.[112,189,1982,2004]" captionTargetBox="[126,1457,335,1922]" captionTargetId="figure-41@17.[113,1474,299,1922]" captionTargetPageId="17" captionText="Figures 41–44. Scorpiops nagphani sp. n., female, paratype, BNHS SC 256, in dorsal (41) ventral (42) views, sternopectinal area (43) and metasomal segment V and telson in lateral view (44)." figureDoi="http://doi.org/10.5281/zenodo.5742565" httpUri="https://zenodo.org/record/5742565/files/figure.png" pageId="16" pageNumber="15">41, 42</figureCitation>
). Overall body color blackish. Legs blackish brown. Telson orangish brown on vesicle and dark brown on aculeus. Ventral portion of body dark brown. Carapace and fingers of manus blackish. Pedipalps blackish brown, darker on carinae. Chelicera basal segment blackish brown. Fingers of chelicera brownish black. 
<emphasis id="B951FF68FF877776FFA3921ABE8EC828" bold="true" box="[113,223,1848,1872]" pageId="16" pageNumber="15">Carapace</emphasis>
(
<figureCitation id="131E3FFFFF877776FF3E921ABF3FC828" box="[236,366,1848,1872]" captionStart="Figures 45–48" captionStartId="18.[115,192,1574,1596]" captionTargetBox="[113,1474,234,1565]" captionTargetId="figure-259@18.[113,1474,234,1565]" captionTargetPageId="18" captionText="Figures 45–48: Scorpiops nagphani sp. n. Figures 45–46. Male, holotype, carapace under white light (45) and UV fluorescence (46). Figures 47–48. Female, paratype, BHNS SC 256, carapace under white light (47) and UV fluorescence (48)." figureDoi="http://doi.org/10.5281/zenodo.5742567" httpUri="https://zenodo.org/record/5742567/files/figure.png" pageId="16" pageNumber="15">Figs. 45–48</figureCitation>
). Anterior margin of carapace strongly granular with deep U shaped anterior median notch. Anterior sub margin coarsely granular. Anterior margin of carapace between lateral eyes and U shaped anterior median notch slightly curved. Entire surface of carapace with fine granules. Anterio-lateral ocular tubercles granular provided with 
<typeStatus id="549E9DD8FF877776FD0792FBBD52C889" box="[725,771,2009,2033]" pageId="16" pageNumber="15">type</typeStatus>
3A lateral ocelli. Two pairs of large major ocelli and one pair of minor ocelli. Median ocular tubercle granular on dorsal portion with a pair of median ocelli situated in the ratio of 1:2.1 (ratio of median ocelli to anterior margin and median ocelli to posterior margin).
</paragraph>
<paragraph id="8B9A237AFF877776FCE290DABA37C9E8" blockId="16.[816,1475,1368,2032]" pageId="16" pageNumber="15">
<emphasis id="B951FF68FF877776FCE290DABDF8C968" bold="true" box="[816,937,1528,1552]" pageId="16" pageNumber="15">Chelicerae</emphasis>
(
<figureCitation id="131E3FFFFF877776FC6490DABA55C969" box="[950,1028,1528,1553]" captionStart="Figures 34–38" captionStartId="15.[116,193,1986,2008]" captionTargetBox="[167,1429,241,1953]" captionTargetId="figure-44@15.[113,1474,234,1956]" captionTargetPageId="15" captionText="Figures 34–38. Scorpiops nagphani sp. n., male, holotype, dorsal (34) and ventral (35) views, sternopectinal area (36), metasomal segment V and telson in lateral view (37) and chelicera in dorsal view (38)." figureDoi="http://doi.org/10.5281/zenodo.5742561" httpUri="https://zenodo.org/record/5742561/files/figure.png" pageId="16" pageNumber="15">Fig. 38</figureCitation>
). Proximal portion with reticulated mosaic design. Fixed finger of chelicera with 3 large triangular teeth on inner margin. Ventral fang of movable finger with a row of 5 minute teeth on inner margin. Dorsal fang of movable finger with 4 teeth on inner margin.
</paragraph>
<paragraph id="8B9A237AFF877774FCE293BABC1FC835" blockId="16.[816,1475,1368,2032]" lastBlockId="18.[115,773,1653,2029]" lastPageId="18" lastPageNumber="17" pageId="16" pageNumber="15">
<emphasis id="B951FF68FF877776FCE293BABDC4C9C8" bold="true" box="[816,917,1688,1712]" pageId="16" pageNumber="15">Pedipalp</emphasis>
(
<figureCitation id="131E3FFFFF877776FC7B93BABA7EC9C8" box="[937,1071,1688,1712]" captionStart="Figures 49–54" captionStartId="19.[113,190,1296,1318]" captionTargetBox="[113,1474,234,1286]" captionTargetId="figure-392@19.[113,1474,234,1286]" captionTargetPageId="19" captionText="Figures 49–54. Scorpiops nagphani sp. n., male, holotype. Figures 49–54. Pedipalp chela dorsal (49), ventral (50) and dorsoexternal (51), patella external (52), dorsal (53) and ventral (54) views under UV fluorescence. Trichobothrial pattern indicated by yellow circles." figureDoi="http://doi.org/10.5281/zenodo.5742569" httpUri="https://zenodo.org/record/5742569/files/figure.png" pageId="16" pageNumber="15">Figs. 49–54</figureCitation>
, 
<figureCitation id="131E3FFFFF877776FBED93BBBAF5C9C8" box="[1087,1188,1688,1713]" captionStart="Figures 127–138" captionStartId="41.[116,193,1928,1950]" captionTargetBox="[113,1474,245,1897]" captionTargetId="figure-102@41.[113,1474,234,1897]" captionTargetPageId="41" captionText="Figures 127–138: Figures 127–130: Comparison of left pedipalps. Figures 131–134: Comparison of metasomal segments II-IV in lateral view. Figures 135–138: Comparison of pedipalp patella dorsal view with internal aspect tubercles. Figures 127, 131, 135. Scorpiops vrushchik sp. n., male, holotype. Figures 128, 132, 136. Scorpiops nagphani sp. n., male, holotype. Figures 129, 133, 137. Scorpiops neera sp. n., male, holotype. Figures 130, 134, 138. Scorpiops lioneli sp. n., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.5742617" httpUri="https://zenodo.org/record/5742617/files/figure.png" pageId="16" pageNumber="15">128, 136</figureCitation>
, 
<figureCitation id="131E3FFFFF877776FB6693BABAB1C9C8" box="[1204,1248,1688,1712]" captionStart="Figures 139–152" captionStartId="43.[115,192,1183,1205]" captionTargetBox="[113,1474,234,1167]" captionTargetId="figure-329@43.[113,1474,234,1167]" captionTargetPageId="43" captionText="Figures 139–152: Figures 139–142: Comparison of dorsolateral granules on anal rim of metasomal segment V in dorsal view. Figures 143–152: Comparison of dentition on movable finger of chela with field emission scanning electron microscopy (FESEM) images in dorsal view. Figures 139, 149. Scorpiops vrushchik sp. n., male, holotype. Figures 140, 150. Scorpiops nagphani sp. n., male, holotype. Figures 141, 151. Scorpiops neera sp. n., male, holotype. Figures 142, 152. Scorpiops lioneli sp. n., male, holotype. Figure 143. S. tenuicauda, male, INHER-214. Figure 144. S. satarensis, male, INHER-213. Figure 145. S. deccanensis, male, INHER-86. Figure 146. S. maharashtraensis, male, INHER-179. Figure 147. S. phaltanensis, male, paratype, INHER-64. Figure 148. S. telbaila, male, paratype, INHER-123." figureDoi="http://doi.org/10.5281/zenodo.5742621" httpUri="https://zenodo.org/record/5742621/files/figure.png" pageId="16" pageNumber="15">150</figureCitation>
). Femur and patella dorsoventrally flattened. Femur with 6 carinae (dorsal retrolateral, dorsal prolateral, retrolateral dorsosubmedian, prolateral ventrosubmedian, ventral retrolateral and ventral prolateral). Ventral prolateral carina weak and present only on 1/3rd proximal portion. Intercarinal space finely and almost evenly granular. Patella with 5 carinae (dorsal retrolateral, dorsal prolateral, retrolateral dorsosubmedian, ventral retrolateral and ventral prolateral). Internal surface of patella with one large posterio-ventral tubercle with one small, thick adjacent bulge and one adjacent small posterio-dorsal tubercle and anterio-ventral tubercle absent (
<figureCitation id="131E3FFFFF857774FE349357BC10C9F5" box="[486,577,1653,1677]" captionStart="Figures 127–138" captionStartId="41.[116,193,1928,1950]" captionTargetBox="[113,1474,245,1897]" captionTargetId="figure-102@41.[113,1474,234,1897]" captionTargetPageId="41" captionText="Figures 127–138: Figures 127–130: Comparison of left pedipalps. Figures 131–134: Comparison of metasomal segments II-IV in lateral view. Figures 135–138: Comparison of pedipalp patella dorsal view with internal aspect tubercles. Figures 127, 131, 135. Scorpiops vrushchik sp. n., male, holotype. Figures 128, 132, 136. Scorpiops nagphani sp. n., male, holotype. Figures 129, 133, 137. Scorpiops neera sp. n., male, holotype. Figures 130, 134, 138. Scorpiops lioneli sp. n., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.5742617" httpUri="https://zenodo.org/record/5742617/files/figure.png" pageId="18" pageNumber="17">Fig. 136</figureCitation>
). Manus elongated with 5 carinae (dorsal retrolateral, retrolateral median, ventral retrolateral, dorsal prolateral and ventral prolateral). Dorsal retrolateral and dorsal prolateral carinae running anteriorly up to the tip of fixed finger. Intercarinal space finely granular on inner and outer surface. Both fingers scalloped deeply at the base. Trichobothrial pattern neobothriotaxic.
</paragraph>
<caption id="DF5A73F2FF867777FFA2929CBC20C88A" ID-DOI="http://doi.org/10.5281/zenodo.5742565" ID-Zenodo-Dep="5742565" httpUri="https://zenodo.org/record/5742565/files/figure.png" pageId="17" pageNumber="16" startId="17.[112,189,1982,2004]" targetBox="[126,1457,335,1922]" targetPageId="17">
<paragraph id="8B9A237AFF867777FFA2929CBC20C88A" blockId="17.[112,1471,1982,2034]" pageId="17" pageNumber="16">
<emphasis id="B951FF68FF867777FFA2929CBF51C8AC" bold="true" box="[112,256,1982,2004]" pageId="17" pageNumber="16">Figures 41–44</emphasis>
. 
<taxonomicName id="4C2558F9FF867777FEDF929DBF81C8AC" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[269,464,1983,2004]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FF867777FEDF929DBF81C8AC" box="[269,464,1983,2004]" italics="true" pageId="17" pageNumber="16">Scorpiops nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FF867777FE04929DBC5CC8AC" box="[470,525,1983,2004]" pageId="17" pageNumber="16" rank="species">
<emphasis id="B951FF68FF867777FE04929DBFBCC8AC" bold="true" box="[470,493,1983,2004]" pageId="17" pageNumber="16">sp</emphasis>
. 
<emphasis id="B951FF68FF867777FE28929DBC56C8AC" bold="true" box="[506,519,1983,2004]" pageId="17" pageNumber="16">n</emphasis>
.
</taxonomicNameLabel>
, female, paratype, BNHS SC 256, in dorsal (41) ventral (42) views, sternopectinal area (43) and metasomal segment V and telson in lateral view (44).
</paragraph>
</caption>
<caption id="DF5A73F2FF857774FFA19304BA64C921" ID-DOI="http://doi.org/10.5281/zenodo.5742567" ID-Zenodo-Dep="5742567" httpUri="https://zenodo.org/record/5742567/files/figure.png" pageId="18" pageNumber="17" startId="18.[115,192,1574,1596]" targetBox="[113,1474,234,1565]" targetPageId="18">
<paragraph id="8B9A237AFF857774FFA19304BA64C921" blockId="18.[115,1473,1574,1625]" pageId="18" pageNumber="17">
<emphasis id="B951FF68FF857774FFA19304BF50C943" bold="true" box="[115,257,1574,1596]" pageId="18" pageNumber="17">Figures 45–48</emphasis>
: 
<taxonomicName id="4C2558F9FF857774FEDF9305BF9FC943" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[269,462,1574,1596]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FF857774FEDF9305BF9FC943" box="[269,462,1574,1596]" italics="true" pageId="18" pageNumber="17">Scorpiops nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FF857774FE019305BC59C944" box="[467,520,1575,1596]" pageId="18" pageNumber="17" rank="species">
<emphasis id="B951FF68FF857774FE019305BFBBC944" bold="true" box="[467,490,1575,1596]" pageId="18" pageNumber="17">sp</emphasis>
. 
<emphasis id="B951FF68FF857774FE279305BC53C944" bold="true" box="[501,514,1575,1596]" pageId="18" pageNumber="17">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B951FF68FF857774FDDC9304BCCDC943" bold="true" box="[526,668,1574,1596]" pageId="18" pageNumber="17">Figures 45–46</emphasis>
. Male, holotype, carapace under white light (45) and UV fluorescence (46). 
<emphasis id="B951FF68FF857774FAA69304BEFEC920" bold="true" pageId="18" pageNumber="17">Figures 47–48</emphasis>
. Female, paratype, BHNS SC 256, carapace under white light (47) and UV fluorescence (48).
</paragraph>
</caption>
<paragraph id="8B9A237AFF857774FFA19277BF4EC895" blockId="18.[115,773,1653,2029]" pageId="18" pageNumber="17">
<emphasis id="B951FF68FF857774FFA19277BEF9C815" bold="true" box="[115,168,1877,1901]" pageId="18" pageNumber="17">Legs</emphasis>
(
<figureCitation id="131E3FFFFF857774FF109277BF08C815" box="[194,345,1877,1901]" captionStart="Figures 34–38" captionStartId="15.[116,193,1986,2008]" captionTargetBox="[167,1429,241,1953]" captionTargetId="figure-44@15.[113,1474,234,1956]" captionTargetPageId="15" captionText="Figures 34–38. Scorpiops nagphani sp. n., male, holotype, dorsal (34) and ventral (35) views, sternopectinal area (36), metasomal segment V and telson in lateral view (37) and chelicera in dorsal view (38)." figureDoi="http://doi.org/10.5281/zenodo.5742561" httpUri="https://zenodo.org/record/5742561/files/figure.png" pageId="18" pageNumber="17">Figs. 34, 35</figureCitation>
, 
<figureCitation id="131E3FFFFF857774FEBD9277BFE5C815" box="[367,436,1877,1901]" captionStart-0="Figures 39–40" captionStart-1="Figures 41–44" captionStartId-0="16.[113,190,1317,1339]" captionStartId-1="17.[112,189,1982,2004]" captionTargetBox-0="[113,1474,234,1309]" captionTargetBox-1="[126,1457,335,1922]" captionTargetId-0="figure-390@16.[113,1474,234,1309]" captionTargetId-1="figure-41@17.[113,1474,299,1922]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="Figures 39–40. Scorpiops nagphani sp. n., male, holotype in dorsal (39) and ventral (40) views under UV light." captionText-1="Figures 41–44. Scorpiops nagphani sp. n., female, paratype, BNHS SC 256, in dorsal (41) ventral (42) views, sternopectinal area (43) and metasomal segment V and telson in lateral view (44)." figureDoi-0="http://doi.org/10.5281/zenodo.5742563" figureDoi-1="http://doi.org/10.5281/zenodo.5742565" httpUri-0="https://zenodo.org/record/5742563/files/figure.png" httpUri-1="https://zenodo.org/record/5742565/files/figure.png" pageId="18" pageNumber="17">39–42</figureCitation>
, 
<figureCitation id="131E3FFFFF857774FE189277BC40C815" box="[458,529,1877,1901]" captionStart="Figures 55–61" captionStartId="20.[115,192,1933,1955]" captionTargetBox="[125,1465,236,1909]" captionTargetId="figure-111@20.[113,1474,226,1909]" captionTargetPageId="20" captionText="Figures 55–61. Scorpiops nagphani sp. n. Figures 55–58. Scorpiops nagphani sp. n., male, holotype, left legs I-IV, retrolateral aspect. Figures 59–61. Scorpiops nagphani sp. n., male, paratype, INHER-259, hemispermatophore internal (59), external (60) views and capsular region in internal view (61). Abbrevations: distal lamina (dl), capsule (c), trunk (t), pedicel (p), distal posterior lobe (dpl), lateral hook (lh), terminal membrane of the sperm duct (tmsd) and basal carina (bc)." figureDoi="http://doi.org/10.5281/zenodo.5742571" httpUri="https://zenodo.org/record/5742571/files/figure.png" pageId="18" pageNumber="17">55–58</figureCitation>
). Femur and patella carinated, intercarinal space densely granular on femur and sparsely granular on patella. Tarsomere I provided with two to four rows of spinules and tarsomere II with single ventral row of spinules.
</paragraph>
<paragraph id="8B9A237AFF857774FCE09357BA08C8D5" blockId="18.[818,1476,1653,2030]" pageId="18" pageNumber="17">
<emphasis id="B951FF68FF857774FCE09357BDC4C9F5" bold="true" box="[818,917,1653,1677]" pageId="18" pageNumber="17">Sternum</emphasis>
, 
<emphasis id="B951FF68FF857774FC759357BB49C9F5" bold="true" box="[935,1304,1653,1677]" pageId="18" pageNumber="17">genital operculum and pectines</emphasis>
(
<figureCitation id="131E3FFFFF857774FAF99357BBD8C9F5" box="[1323,1417,1653,1677]" captionStart="Figures 34–38" captionStartId="15.[116,193,1986,2008]" captionTargetBox="[167,1429,241,1953]" captionTargetId="figure-44@15.[113,1474,234,1956]" captionTargetPageId="15" captionText="Figures 34–38. Scorpiops nagphani sp. n., male, holotype, dorsal (34) and ventral (35) views, sternopectinal area (36), metasomal segment V and telson in lateral view (37) and chelicera in dorsal view (38)." figureDoi="http://doi.org/10.5281/zenodo.5742561" httpUri="https://zenodo.org/record/5742561/files/figure.png" pageId="18" pageNumber="17">Figs. 36</figureCitation>
, 
<figureCitation id="131E3FFFFF857774FA4B9357BBE6C9F5" box="[1433,1463,1653,1677]" captionStart="Figures 41–44" captionStartId="17.[112,189,1982,2004]" captionTargetBox="[126,1457,335,1922]" captionTargetId="figure-41@17.[113,1474,299,1922]" captionTargetPageId="17" captionText="Figures 41–44. Scorpiops nagphani sp. n., female, paratype, BNHS SC 256, in dorsal (41) ventral (42) views, sternopectinal area (43) and metasomal segment V and telson in lateral view (44)." figureDoi="http://doi.org/10.5281/zenodo.5742565" httpUri="https://zenodo.org/record/5742565/files/figure.png" pageId="18" pageNumber="17">43</figureCitation>
). Broad, pentagonal and finely granular only on anterior middle portion. Genital operculum with a pair of strongly protruding genital papillae. Basal piece with slight depression on middle portion. Pectine morphology of 
<typeStatus id="549E9DD8FF857774FB5293D4BAFFC876" box="[1152,1198,1782,1806]" pageId="18" pageNumber="17">type</typeStatus>
P3 with 7/7 pectinal teeth. 
<emphasis id="B951FF68FF857774FCE09237BDFBC855" bold="true" box="[818,938,1813,1837]" pageId="18" pageNumber="17">Mesosoma</emphasis>
(
<figureCitation id="131E3FFFFF857774FC6B9237BA6DC855" box="[953,1084,1813,1837]" captionStart="Figures 34–38" captionStartId="15.[116,193,1986,2008]" captionTargetBox="[167,1429,241,1953]" captionTargetId="figure-44@15.[113,1474,234,1956]" captionTargetPageId="15" captionText="Figures 34–38. Scorpiops nagphani sp. n., male, holotype, dorsal (34) and ventral (35) views, sternopectinal area (36), metasomal segment V and telson in lateral view (37) and chelicera in dorsal view (38)." figureDoi="http://doi.org/10.5281/zenodo.5742561" httpUri="https://zenodo.org/record/5742561/files/figure.png" pageId="18" pageNumber="17">Figs. 34, 35</figureCitation>
, 
<figureCitation id="131E3FFFFF857774FB9A9237BAC1C855" box="[1096,1168,1813,1837]" captionStart-0="Figures 39–40" captionStart-1="Figures 41–44" captionStartId-0="16.[113,190,1317,1339]" captionStartId-1="17.[112,189,1982,2004]" captionTargetBox-0="[113,1474,234,1309]" captionTargetBox-1="[126,1457,335,1922]" captionTargetId-0="figure-390@16.[113,1474,234,1309]" captionTargetId-1="figure-41@17.[113,1474,299,1922]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="Figures 39–40. Scorpiops nagphani sp. n., male, holotype in dorsal (39) and ventral (40) views under UV light." captionText-1="Figures 41–44. Scorpiops nagphani sp. n., female, paratype, BNHS SC 256, in dorsal (41) ventral (42) views, sternopectinal area (43) and metasomal segment V and telson in lateral view (44)." figureDoi-0="http://doi.org/10.5281/zenodo.5742563" figureDoi-1="http://doi.org/10.5281/zenodo.5742565" httpUri-0="https://zenodo.org/record/5742563/files/figure.png" httpUri-1="https://zenodo.org/record/5742565/files/figure.png" pageId="18" pageNumber="17">39–42</figureCitation>
). All tergites finely granular, with median carina absent on segment I. Tergite VII additionally with two pairs of lateral granular carinae present only on half posterior portion. Sternites III-VI entirely smooth. Sternite VII finely granular.
</paragraph>
<paragraph id="8B9A237AFF857775FCE09297BF01C9A1" blockId="18.[818,1476,1653,2030]" lastBlockId="19.[111,769,1377,2041]" lastPageId="19" lastPageNumber="18" pageId="18" pageNumber="17">
<emphasis id="B951FF68FF857774FCE09297BDFBC8B5" bold="true" box="[818,938,1973,1997]" pageId="18" pageNumber="17">Metasoma</emphasis>
(
<figureCitation id="131E3FFFFF857774FC139297BA07C8B5" box="[961,1110,1973,1997]" captionStart="Figures 34–38" captionStartId="15.[116,193,1986,2008]" captionTargetBox="[167,1429,241,1953]" captionTargetId="figure-44@15.[113,1474,234,1956]" captionTargetPageId="15" captionText="Figures 34–38. Scorpiops nagphani sp. n., male, holotype, dorsal (34) and ventral (35) views, sternopectinal area (36), metasomal segment V and telson in lateral view (37) and chelicera in dorsal view (38)." figureDoi="http://doi.org/10.5281/zenodo.5742561" httpUri="https://zenodo.org/record/5742561/files/figure.png" pageId="18" pageNumber="17">Figs. 34, 35</figureCitation>
, 
<figureCitation id="131E3FFFFF857774FBB89297BAE0C8B5" box="[1130,1201,1973,1997]" captionStart-0="Figures 39–40" captionStart-1="Figures 41–44" captionStartId-0="16.[113,190,1317,1339]" captionStartId-1="17.[112,189,1982,2004]" captionTargetBox-0="[113,1474,234,1309]" captionTargetBox-1="[126,1457,335,1922]" captionTargetId-0="figure-390@16.[113,1474,234,1309]" captionTargetId-1="figure-41@17.[113,1474,299,1922]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="Figures 39–40. Scorpiops nagphani sp. n., male, holotype in dorsal (39) and ventral (40) views under UV light." captionText-1="Figures 41–44. Scorpiops nagphani sp. n., female, paratype, BNHS SC 256, in dorsal (41) ventral (42) views, sternopectinal area (43) and metasomal segment V and telson in lateral view (44)." figureDoi-0="http://doi.org/10.5281/zenodo.5742563" figureDoi-1="http://doi.org/10.5281/zenodo.5742565" httpUri-0="https://zenodo.org/record/5742563/files/figure.png" httpUri-1="https://zenodo.org/record/5742565/files/figure.png" pageId="18" pageNumber="17">39–42</figureCitation>
, 
<figureCitation id="131E3FFFFF857774FB169297BABEC8B5" box="[1220,1263,1973,1997]" captionStart="Figures 127–138" captionStartId="41.[116,193,1928,1950]" captionTargetBox="[113,1474,245,1897]" captionTargetId="figure-102@41.[113,1474,234,1897]" captionTargetPageId="41" captionText="Figures 127–138: Figures 127–130: Comparison of left pedipalps. Figures 131–134: Comparison of metasomal segments II-IV in lateral view. Figures 135–138: Comparison of pedipalp patella dorsal view with internal aspect tubercles. Figures 127, 131, 135. Scorpiops vrushchik sp. n., male, holotype. Figures 128, 132, 136. Scorpiops nagphani sp. n., male, holotype. Figures 129, 133, 137. Scorpiops neera sp. n., male, holotype. Figures 130, 134, 138. Scorpiops lioneli sp. n., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.5742617" httpUri="https://zenodo.org/record/5742617/files/figure.png" pageId="18" pageNumber="17">132</figureCitation>
, 
<figureCitation id="131E3FFFFF857774FAD19297BB61C8B5" box="[1283,1328,1973,1997]" captionStart="Figures 139–152" captionStartId="43.[115,192,1183,1205]" captionTargetBox="[113,1474,234,1167]" captionTargetId="figure-329@43.[113,1474,234,1167]" captionTargetPageId="43" captionText="Figures 139–152: Figures 139–142: Comparison of dorsolateral granules on anal rim of metasomal segment V in dorsal view. Figures 143–152: Comparison of dentition on movable finger of chela with field emission scanning electron microscopy (FESEM) images in dorsal view. Figures 139, 149. Scorpiops vrushchik sp. n., male, holotype. Figures 140, 150. Scorpiops nagphani sp. n., male, holotype. Figures 141, 151. Scorpiops neera sp. n., male, holotype. Figures 142, 152. Scorpiops lioneli sp. n., male, holotype. Figure 143. S. tenuicauda, male, INHER-214. Figure 144. S. satarensis, male, INHER-213. Figure 145. S. deccanensis, male, INHER-86. Figure 146. S. maharashtraensis, male, INHER-179. Figure 147. S. phaltanensis, male, paratype, INHER-64. Figure 148. S. telbaila, male, paratype, INHER-123." figureDoi="http://doi.org/10.5281/zenodo.5742621" httpUri="https://zenodo.org/record/5742621/files/figure.png" pageId="18" pageNumber="17">140</figureCitation>
). Metasomal segments I, II and III with 5 pairs of carinae (dorsal lateral, lateral supramedian, lateral inframedian, ventral lateral and ventral submedian), segment II and III with lateral inframedian carina present only on posterior portion, metasomal segments IV with 4 pairs of carinae (dorsal lateral, lateral supramedian, ventral lateral and ventral submedian), and segment V with 3 pairs (dorsal lateral, lateral inframedian and ventral lateral) and 1 single ventral median carinae. Intercarinal space with mixed granules (fine and coarse). Dorsal lateral carination on segments III and IV ending posteriorly into a short spine. Anal rim of segment V evenly crenulated. A pair of dorsolateral granules of anal rim strongly tuberculate.
</paragraph>
<caption id="DF5A73F2FF847775FFA39032BB05CA3B" ID-DOI="http://doi.org/10.5281/zenodo.5742569" ID-Zenodo-Dep="5742569" httpUri="https://zenodo.org/record/5742569/files/figure.png" pageId="19" pageNumber="18" startId="19.[113,190,1296,1318]" targetBox="[113,1474,234,1286]" targetPageId="19">
<paragraph id="8B9A237AFF847775FFA39032BB05CA3B" blockId="19.[113,1471,1296,1347]" pageId="19" pageNumber="18">
<emphasis id="B951FF68FF847775FFA39032BF53CA5E" bold="true" box="[113,258,1296,1318]" pageId="19" pageNumber="18">Figures 49–54</emphasis>
. 
<taxonomicName id="4C2558F9FF847775FEDD9033BF80CA5E" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[271,465,1297,1318]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FF847775FEDD9033BF80CA5E" box="[271,465,1297,1318]" italics="true" pageId="19" pageNumber="18">Scorpiops nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FF847775FE0A9033BC5ECA5E" box="[472,527,1297,1318]" pageId="19" pageNumber="18" rank="species">
<emphasis id="B951FF68FF847775FE0A9033BFBECA5E" bold="true" box="[472,495,1297,1318]" pageId="19" pageNumber="18">sp</emphasis>
. 
<emphasis id="B951FF68FF847775FE2E9033BC58CA5E" bold="true" box="[508,521,1297,1318]" pageId="19" pageNumber="18">n</emphasis>
.
</taxonomicNameLabel>
, male, holotype. 
<emphasis id="B951FF68FF847775FD689032BD1ACA5E" bold="true" box="[698,843,1296,1318]" pageId="19" pageNumber="18">Figures 49–54</emphasis>
. Pedipalp chela dorsal (49), ventral (50) and dorsoexternal (51), patella external (52), dorsal (53) and ventral (54) views under UV fluorescence. Trichobothrial pattern indicated by yellow circles.
</paragraph>
</caption>
<paragraph id="8B9A237AFF847775FFBD93C3BEB2C841" blockId="19.[111,769,1377,2041]" pageId="19" pageNumber="18">
<emphasis id="B951FF68FF847775FFBD93C3BEE9C981" bold="true" box="[111,184,1761,1785]" pageId="19" pageNumber="18">Telson</emphasis>
(
<figureCitation id="131E3FFFFF847775FF1A93C3BF73C981" box="[200,290,1761,1785]" captionStart="Figures 34–38" captionStartId="15.[116,193,1986,2008]" captionTargetBox="[167,1429,241,1953]" captionTargetId="figure-44@15.[113,1474,234,1956]" captionTargetPageId="15" captionText="Figures 34–38. Scorpiops nagphani sp. n., male, holotype, dorsal (34) and ventral (35) views, sternopectinal area (36), metasomal segment V and telson in lateral view (37) and chelicera in dorsal view (38)." figureDoi="http://doi.org/10.5281/zenodo.5742561" httpUri="https://zenodo.org/record/5742561/files/figure.png" pageId="19" pageNumber="18">Figs. 37</figureCitation>
, 
<figureCitation id="131E3FFFFF847775FEFC93C3BF1CC981" box="[302,333,1761,1785]" captionStart="Figures 41–44" captionStartId="17.[112,189,1982,2004]" captionTargetBox="[126,1457,335,1922]" captionTargetId="figure-41@17.[113,1474,299,1922]" captionTargetPageId="17" captionText="Figures 41–44. Scorpiops nagphani sp. n., female, paratype, BNHS SC 256, in dorsal (41) ventral (42) views, sternopectinal area (43) and metasomal segment V and telson in lateral view (44)." figureDoi="http://doi.org/10.5281/zenodo.5742565" httpUri="https://zenodo.org/record/5742565/files/figure.png" pageId="19" pageNumber="18">44</figureCitation>
). Elongated and almost entirely smooth. A weak depression present in between vesicle and at the base of aculeus.
</paragraph>
<paragraph id="8B9A237AFF847775FFBD9263BA3CC941" blockId="19.[111,769,1377,2041]" lastBlockId="19.[814,1471,1377,1593]" pageId="19" pageNumber="18">
<emphasis id="B951FF68FF847775FFBD9263BF0FC821" bold="true" box="[111,350,1857,1881]" pageId="19" pageNumber="18">Hemispermatophore</emphasis>
<specimenCount id="9D23E8F3FF847775FEB99260BF2EC821" box="[363,383,1858,1881]" pageId="19" pageNumber="18" type="male">♂</specimenCount>
<typeStatus id="549E9DD8FF847775FE599260BFBDC822" box="[395,492,1858,1882]" pageId="19" pageNumber="18" type="paratype">paratype</typeStatus>
, INHER-259 (
<figureCitation id="131E3FFFFF847775FD479263BEDCC801" captionStart="Figures 55–61" captionStartId="20.[115,192,1933,1955]" captionTargetBox="[125,1465,236,1909]" captionTargetId="figure-111@20.[113,1474,226,1909]" captionTargetPageId="20" captionText="Figures 55–61. Scorpiops nagphani sp. n. Figures 55–58. Scorpiops nagphani sp. n., male, holotype, left legs I-IV, retrolateral aspect. Figures 59–61. Scorpiops nagphani sp. n., male, paratype, INHER-259, hemispermatophore internal (59), external (60) views and capsular region in internal view (61). Abbrevations: distal lamina (dl), capsule (c), trunk (t), pedicel (p), distal posterior lobe (dpl), lateral hook (lh), terminal membrane of the sperm duct (tmsd) and basal carina (bc)." figureDoi="http://doi.org/10.5281/zenodo.5742571" httpUri="https://zenodo.org/record/5742571/files/figure.png" pageId="19" pageNumber="18">Figs. 59– 61</figureCitation>
). Lamelliform; distal 
<emphasis id="B951FF68FF847775FE509243BC5CC801" box="[386,525,1889,1913]" italics="true" pageId="19" pageNumber="18">basal carina</emphasis>
(
<emphasis id="B951FF68FF847775FDCD9243BC69C801" box="[543,568,1889,1913]" italics="true" pageId="19" pageNumber="18">bc</emphasis>
) with a plate like structure present, sclerotized, crown-like structure (
<emphasis id="B951FF68FF847775FD7092A3BC91C8E1" box="[674,704,1921,1945]" italics="true" pageId="19" pageNumber="18">cls</emphasis>
) with distal margin with 5–6 denticles whose internal surface is attached to terminal membrane of the sperm duct (
<emphasis id="B951FF68FF847775FD1692E3BCA6C8A1" box="[708,759,1985,2009]" italics="true" pageId="19" pageNumber="18">tmsd</emphasis>
) on the inner concave side of the capsule with numerous spicules; pointed lateral hook (
<emphasis id="B951FF68FF847775FB579043BACBCA01" box="[1157,1178,1377,1401]" italics="true" pageId="19" pageNumber="18">lh</emphasis>
); a long, inwardly twisted distal posterior lobe (
<emphasis id="B951FF68FF847775FBF690A3BA16CAE1" box="[1060,1095,1409,1433]" italics="true" pageId="19" pageNumber="18">dpl</emphasis>
) close to the distal lamina, with denticulate margin; short and stout trunk; long and slender distal lamina (
<emphasis id="B951FF68FF847775FC0390E3BDB7CAA1" box="[977,998,1473,1497]" italics="true" pageId="19" pageNumber="18">dl</emphasis>
) narrow at the base and sharply bent and tapered apex. Total length 
<quantity id="4CDD8E9FFF847775FB8990C3BAE5CA81" box="[1115,1204,1505,1529]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="19" pageNumber="18" unit="mm" value="8.1">8.1 mm</quantity>
; pedicel 
<quantity id="4CDD8E9FFF847775FAC990C3BB2FCA81" box="[1307,1406,1505,1529]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" pageId="19" pageNumber="18" unit="mm" value="1.18">1.18 mm</quantity>
long; trunk 
<quantity id="4CDD8E9FFF847775FCBD9323BD80C961" box="[879,977,1537,1561]" metricMagnitude="-3" metricUnit="m" metricValue="1.83" pageId="19" pageNumber="18" unit="mm" value="1.83">1.83 mm</quantity>
long; capsule 
<quantity id="4CDD8E9FFF847775FBA09323BA83C961" box="[1138,1234,1537,1561]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="19" pageNumber="18" unit="mm" value="1.11">1.11 mm</quantity>
long; 
<emphasis id="B951FF68FF847775FAC89323BB6CC961" box="[1306,1341,1537,1561]" italics="true" pageId="19" pageNumber="18">dpl</emphasis>
length 
<quantity id="4CDD8E9FFF847775FA429323BD0AC941" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="19" pageNumber="18" unit="mm" value="0.31">0.31 mm</quantity>
; 
<emphasis id="B951FF68FF847775FCB49303BD2AC941" box="[870,891,1569,1593]" italics="true" pageId="19" pageNumber="18">dl</emphasis>
length 
<quantity id="4CDD8E9FFF847775FC1C9303BA7EC941" box="[974,1071,1569,1593]" metricMagnitude="-3" metricUnit="m" metricValue="3.98" pageId="19" pageNumber="18" unit="mm" value="3.98">3.98 mm</quantity>
long.
</paragraph>
<paragraph id="8B9A237AFF847775FCFC9343BA3FC9A1" blockId="19.[814,1472,1633,1753]" pageId="19" pageNumber="18">
SEXUAL DIMORPHISM. Male genital operculum with pair of genital papillae. In females, the genital operculum medially sutured. Pedipalp manus slender in males and stouter in females (
<figureCitation id="131E3FFFFF847775FC5D93E3BA40C9A1" box="[911,1041,1729,1753]" captionStart="Figures 34–38" captionStartId="15.[116,193,1986,2008]" captionTargetBox="[167,1429,241,1953]" captionTargetId="figure-44@15.[113,1474,234,1956]" captionTargetPageId="15" captionText="Figures 34–38. Scorpiops nagphani sp. n., male, holotype, dorsal (34) and ventral (35) views, sternopectinal area (36), metasomal segment V and telson in lateral view (37) and chelicera in dorsal view (38)." figureDoi="http://doi.org/10.5281/zenodo.5742561" httpUri="https://zenodo.org/record/5742561/files/figure.png" pageId="19" pageNumber="18">Figs. 34–37</figureCitation>
, 
<figureCitation id="131E3FFFFF847775FBCE93E3BA33C9A1" box="[1052,1122,1729,1753]" captionStart-0="Figures 39–40" captionStart-1="Figures 41–44" captionStart-2="Figures 45–48" captionStartId-0="16.[113,190,1317,1339]" captionStartId-1="17.[112,189,1982,2004]" captionStartId-2="18.[115,192,1574,1596]" captionTargetBox-0="[113,1474,234,1309]" captionTargetBox-1="[126,1457,335,1922]" captionTargetBox-2="[113,1474,234,1565]" captionTargetId-0="figure-390@16.[113,1474,234,1309]" captionTargetId-1="figure-41@17.[113,1474,299,1922]" captionTargetId-2="figure-259@18.[113,1474,234,1565]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionText-0="Figures 39–40. Scorpiops nagphani sp. n., male, holotype in dorsal (39) and ventral (40) views under UV light." captionText-1="Figures 41–44. Scorpiops nagphani sp. n., female, paratype, BNHS SC 256, in dorsal (41) ventral (42) views, sternopectinal area (43) and metasomal segment V and telson in lateral view (44)." captionText-2="Figures 45–48: Scorpiops nagphani sp. n. Figures 45–46. Male, holotype, carapace under white light (45) and UV fluorescence (46). Figures 47–48. Female, paratype, BHNS SC 256, carapace under white light (47) and UV fluorescence (48)." figureDoi-0="http://doi.org/10.5281/zenodo.5742563" figureDoi-1="http://doi.org/10.5281/zenodo.5742565" figureDoi-2="http://doi.org/10.5281/zenodo.5742567" httpUri-0="https://zenodo.org/record/5742563/files/figure.png" httpUri-1="https://zenodo.org/record/5742565/files/figure.png" httpUri-2="https://zenodo.org/record/5742567/files/figure.png" pageId="19" pageNumber="18">39–48</figureCitation>
).
</paragraph>
<paragraph id="8B9A237AFF847775FCFC9223BDF0C881" blockId="19.[814,1472,1793,2041]" pageId="19" pageNumber="18">
AFFINITIES. 
<taxonomicName id="4C2558F9FF847775FC7C9220BAD7C861" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[942,1158,1793,1817]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FF847775FC7C9220BAD7C861" box="[942,1158,1793,1817]" italics="true" pageId="19" pageNumber="18">Scorpiops nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FF847775FB5D9223BA9CC861" box="[1167,1229,1793,1817]" pageId="19" pageNumber="18" rank="species">
<emphasis id="B951FF68FF847775FB5D9223BAF9C861" bold="true" box="[1167,1192,1793,1817]" pageId="19" pageNumber="18">sp</emphasis>
. 
<emphasis id="B951FF68FF847775FB6A9223BA96C861" bold="true" box="[1208,1223,1793,1817]" pageId="19" pageNumber="18">n</emphasis>
.
</taxonomicNameLabel>
is closely related to a species from Pardhadi Village, Nashik District, 
<collectingRegion id="49E1ED98FF847775FAE89203BBEEC841" box="[1338,1471,1825,1849]" country="India" name="Maharashtra" pageId="19" pageNumber="18">Maharashtra</collectingRegion>
(
<collectingCountry id="F33263EAFF847775FCE79263BDD5C821" box="[821,900,1857,1881]" name="India" pageId="19" pageNumber="18">INDIA</collectingCountry>
) and 
<emphasis id="B951FF68FF847775FC159260BAFDC821" box="[967,1196,1857,1881]" italics="true" pageId="19" pageNumber="18">
<taxonomicName id="4C2558F9FF847775FC159260BAF9C821" box="[967,1192,1857,1881]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="maharashtraensis">S. maharashtraensis</taxonomicName>
.
</emphasis>
It is separated by a raw genetic divergence of 6.7% from a species from Pardhadi Village, Nashik District, 
<collectingRegion id="49E1ED98FF847775FB8392A3BA87C8E1" box="[1105,1238,1921,1945]" country="India" name="Maharashtra" pageId="19" pageNumber="18">Maharashtra</collectingRegion>
(
<collectingCountry id="F33263EAFF847775FB3F92A3BB6DC8E1" box="[1261,1340,1921,1945]" name="India" pageId="19" pageNumber="18">INDIA</collectingCountry>
) and 9.6% from 
<emphasis id="B951FF68FF847775FCB89280BA1DC8C1" box="[874,1100,1953,1977]" italics="true" pageId="19" pageNumber="18">
<taxonomicName id="4C2558F9FF847775FCB89280BA19C8C1" box="[874,1096,1953,1977]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="maharashtraensis">S. maharashtraensis</taxonomicName>
.
</emphasis>
The new species also differs from its close congeners based on following set of morphological characters.
</paragraph>
<caption id="DF5A73F2FF837772FFA192AFBCF3C881" ID-DOI="http://doi.org/10.5281/zenodo.5742571" ID-Zenodo-Dep="5742571" httpUri="https://zenodo.org/record/5742571/files/figure.png" pageId="20" pageNumber="19" startId="20.[115,192,1933,1955]" targetBox="[125,1465,236,1909]" targetPageId="20">
<paragraph id="8B9A237AFF837772FFA192AFBCF3C881" blockId="20.[115,1473,1933,2042]" pageId="20" pageNumber="19">
<emphasis id="B951FF68FF837772FFA192AFBF51C8DA" bold="true" box="[115,256,1933,1955]" pageId="20" pageNumber="19">Figures 55–61</emphasis>
. 
<taxonomicName id="4C2558F9FF837772FED992ACBF9AC8DB" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[267,459,1934,1955]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FF837772FED992ACBF9AC8DB" box="[267,459,1934,1955]" italics="true" pageId="20" pageNumber="19">Scorpiops nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FF837772FE1D92ACBC55C8DB" box="[463,516,1934,1955]" pageId="20" pageNumber="19" rank="species">
<emphasis id="B951FF68FF837772FE1D92ACBFB7C8DB" bold="true" box="[463,486,1934,1955]" pageId="20" pageNumber="19">sp</emphasis>
. 
<emphasis id="B951FF68FF837772FE2392ACBFAFC8DB" bold="true" box="[497,510,1934,1955]" pageId="20" pageNumber="19">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B951FF68FF837772FDDB92AFBCC6C8DA" bold="true" box="[521,663,1933,1955]" pageId="20" pageNumber="19">Figures 55–58</emphasis>
. 
<taxonomicName id="4C2558F9FF837772FD7092ACBD30C8DB" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[674,865,1934,1955]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FF837772FD7092ACBD30C8DB" box="[674,865,1934,1955]" italics="true" pageId="20" pageNumber="19">Scorpiops nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FF837772FCB492ACBDCBC8DB" box="[870,922,1934,1955]" pageId="20" pageNumber="19" rank="species">
<emphasis id="B951FF68FF837772FCB492ACBD2CC8DB" bold="true" box="[870,893,1934,1955]" pageId="20" pageNumber="19">sp</emphasis>
. 
<emphasis id="B951FF68FF837772FC5592ACBDC5C8DB" bold="true" box="[903,916,1934,1955]" pageId="20" pageNumber="19">n</emphasis>
.
</taxonomicNameLabel>
, male, holotype, left legs I-IV, retrolateral aspect. 
<emphasis id="B951FF68FF837772FAA692AFBEFFC8C7" bold="true" pageId="20" pageNumber="19">Figures 59–61</emphasis>
. 
<taxonomicName id="4C2558F9FF837772FF699289BF2DC8C7" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[187,380,1962,1984]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FF837772FF699289BF2DC8C7" box="[187,380,1962,1984]" italics="true" pageId="20" pageNumber="19">Scorpiops nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FF837772FE519288BFE9C8B8" box="[387,440,1962,1984]" pageId="20" pageNumber="19" rank="species">
<emphasis id="B951FF68FF837772FE519288BFCBC8C7" bold="true" box="[387,410,1962,1983]" pageId="20" pageNumber="19">sp</emphasis>
. 
<emphasis id="B951FF68FF837772FE749288BFE2C8C7" bold="true" box="[422,435,1962,1983]" pageId="20" pageNumber="19">n</emphasis>
.
</taxonomicNameLabel>
, male, paratype, INHER-259, hemispermatophore internal (59), external (60) views and capsular region in internal view (61). Abbrevations: distal lamina (
<emphasis id="B951FF68FF837772FD9892E5BC0CC8A4" box="[586,605,1991,2012]" italics="true" pageId="20" pageNumber="19">dl</emphasis>
), capsule (
<emphasis id="B951FF68FF837772FD1892EABC84C8A5" box="[714,725,1992,2013]" italics="true" pageId="20" pageNumber="19">c</emphasis>
), trunk (
<emphasis id="B951FF68FF837772FCFF92EABD65C8A5" box="[813,820,1992,2013]" italics="true" pageId="20" pageNumber="19">t</emphasis>
), pedicel (
<emphasis id="B951FF68FF837772FC4C92EABDFBC8A5" box="[926,938,1992,2013]" italics="true" pageId="20" pageNumber="19">p</emphasis>
), distal posterior lobe (
<emphasis id="B951FF68FF837772FB4392E5BAE1C8A4" box="[1169,1200,1991,2012]" italics="true" pageId="20" pageNumber="19">dpl</emphasis>
), lateral hook (
<emphasis id="B951FF68FF837772FA9A92E5BB0AC8A4" box="[1352,1371,1991,2012]" italics="true" pageId="20" pageNumber="19">lh</emphasis>
), terminal membrane of the sperm duct (
<emphasis id="B951FF68FF837772FE4192C6BF91C881" box="[403,448,2020,2041]" italics="true" pageId="20" pageNumber="19">tmsd</emphasis>
) and 
<emphasis id="B951FF68FF837772FE2592C6BC21C882" box="[503,624,2020,2042]" italics="true" pageId="20" pageNumber="19">basal carina</emphasis>
(
<emphasis id="B951FF68FF837772FDAC92C6BCC4C881" box="[638,661,2020,2041]" italics="true" pageId="20" pageNumber="19">bc</emphasis>
).
</paragraph>
</caption>
<caption id="DF5A73F2FF827773FFA19065BDCECAEF" ID-DOI="http://doi.org/10.5281/zenodo.5742573" ID-Zenodo-Dep="5742573" httpUri="https://zenodo.org/record/5742573/files/figure.png" pageId="21" pageNumber="20" startId="21.[115,192,1351,1373]" targetBox="[113,1474,234,1341]" targetPageId="21">
<paragraph id="8B9A237AFF827773FFA19065BDCECAEF" blockId="21.[115,1473,1351,1431]" pageId="21" pageNumber="20">
<emphasis id="B951FF68FF827773FFA19065BF54CA24" bold="true" box="[115,261,1351,1373]" pageId="21" pageNumber="20">Figures 62–64</emphasis>
. Type locality of 
<taxonomicName id="4C2558F9FF827773FE6B906ABC2CCA24" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" box="[441,637,1351,1373]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FF827773FE6B906ABC2CCA24" box="[441,637,1351,1373]" italics="true" pageId="21" pageNumber="20">Scorpiops nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FF827773FD54906ABCEFCA25" box="[646,702,1352,1373]" pageId="21" pageNumber="20" rank="species">
<emphasis id="B951FF68FF827773FD54906ABCCCCA25" bold="true" box="[646,669,1352,1373]" pageId="21" pageNumber="20">sp</emphasis>
. 
<emphasis id="B951FF68FF827773FD79906ABCE9CA25" bold="true" box="[683,696,1352,1373]" pageId="21" pageNumber="20">n</emphasis>
.
</taxonomicNameLabel>
, aerial view of semi-evergreen forest inside Bhimashankar wildlife sanctuary (BWLS), very close to the type locality (62), view of Kolvan waterfalls inside BWLS where the new species was observed (63) and 
<taxonomicName id="4C2558F9FF827773FAB39047BE9FCAEE" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FF827773FAB39047BE9FCAEE" italics="true" pageId="21" pageNumber="20">Scorpiops nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FF827773FF0190A3BF59CAEF" box="[211,264,1409,1431]" pageId="21" pageNumber="20" rank="species">
<emphasis id="B951FF68FF827773FF0190A3BEBBCAEE" bold="true" box="[211,234,1409,1430]" pageId="21" pageNumber="20">sp</emphasis>
. 
<emphasis id="B951FF68FF827773FF2490A3BF52CAEE" bold="true" box="[246,259,1409,1430]" pageId="21" pageNumber="20">n</emphasis>
.
</taxonomicNameLabel>
, female (specimen not collected) in vivo habitus with juveniles (64).
</paragraph>
</caption>
<paragraph id="8B9A237AFF827773FFA39093BF2EC9B0" blockId="21.[113,771,1457,1929]" pageId="21" pageNumber="20">
Trichobothria on patella ventral 12–14 as opposed to 
<quantity id="4CDD8E9FFF827773FD099093BEFECA91" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.572" metricValueMin="3.81" pageId="21" pageNumber="20" unit="in" value="16.5" valueMax="18.0" valueMin="15.0">15– 18 in</quantity>
the species from Pardhadi Village, Nashik District, 
<collectingRegion id="49E1ED98FF827773FFA390D3BEA7C971" box="[113,246,1521,1545]" country="India" name="Maharashtra" pageId="21" pageNumber="20">Maharashtra</collectingRegion>
(
<collectingCountry id="F33263EAFF827773FED090D3BF00C971" box="[258,337,1521,1545]" name="India" pageId="21" pageNumber="20">INDIA</collectingCountry>
) and 
<quantity id="4CDD8E9FFF827773FE5590D3BFB2C971" box="[391,483,1521,1545]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" metricValueMax="4.318" metricValueMin="3.81" pageId="21" pageNumber="20" unit="in" value="16.0" valueMax="17.0" valueMin="15.0">15–17 in</quantity>
<taxonomicName id="4C2558F9FF827773FE3A90D3BCEEC971" box="[488,703,1521,1545]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="maharashtraensis">
<emphasis id="B951FF68FF827773FE3A90D3BCEEC971" box="[488,703,1521,1545]" italics="true" pageId="21" pageNumber="20">S. maharashtraensis</emphasis>
</taxonomicName>
; chela length to width ratio in males 3.7–4.1 as opposed to 
<quantity id="4CDD8E9FFF827773FD629333BCBEC951" box="[688,751,1553,1577]" metricMagnitude="-1" metricUnit="m" metricValue="1.1176" pageId="21" pageNumber="20" unit="in" value="4.4">4.4 in</quantity>
a species from Pardhadi Village, Nashik District, 
<collectingRegion id="49E1ED98FF827773FDAC9313BD52C931" box="[638,771,1585,1609]" country="India" name="Maharashtra" pageId="21" pageNumber="20">Maharashtra</collectingRegion>
(
<collectingCountry id="F33263EAFF827773FFAB9373BE99C911" box="[121,200,1617,1641]" name="India" pageId="21" pageNumber="20">INDIA</collectingCountry>
) and 
<quantity id="4CDD8E9FFF827773FED09373BF3FC911" box="[258,366,1617,1641]" metricMagnitude="-1" metricUnit="m" metricValue="1.1049" metricValueMax="1.1176" metricValueMin="1.0921999999999998" pageId="21" pageNumber="20" unit="in" value="4.35" valueMax="4.4" valueMin="4.3">4.3–4.4 in</quantity>
<taxonomicName id="4C2558F9FF827773FEA69373BC1CC911" box="[372,589,1617,1641]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="maharashtraensis">
<emphasis id="B951FF68FF827773FEA69373BC1CC911" box="[372,589,1617,1641]" italics="true" pageId="21" pageNumber="20">S. maharashtraensis</emphasis>
</taxonomicName>
; body coloration blackish brown as opposed to yellowish brown in a species from Pardhadi Village, Nashik District, 
<collectingRegion id="49E1ED98FF827773FDC893B3BCCEC9D1" box="[538,671,1681,1705]" country="India" name="Maharashtra" pageId="21" pageNumber="20">Maharashtra</collectingRegion>
(
<collectingCountry id="F33263EAFF827773FD7F93B3BCADC9D1" box="[685,764,1681,1705]" name="India" pageId="21" pageNumber="20">INDIA</collectingCountry>
) and 
<taxonomicName id="4C2558F9FF827773FF4D9393BF29C9B1" box="[159,376,1713,1737]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="maharashtraensis">
<emphasis id="B951FF68FF827773FF4D9393BF29C9B1" box="[159,376,1713,1737]" italics="true" pageId="21" pageNumber="20">S. maharashtraensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9A237AFF827773FFA393F3BEA8C8F1" blockId="21.[113,771,1457,1929]" pageId="21" pageNumber="20">
It differs from all the remaining species of 
<taxonomicName id="4C2558F9FF827773FDE593F3BCF3C990" authorityName="Peters" authorityYear="1861" box="[567,674,1745,1768]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B951FF68FF827773FDE593F3BCF3C990" box="[567,674,1745,1768]" italics="true" pageId="21" pageNumber="20">Scorpiops</emphasis>
</taxonomicName>
from the northern Western Ghats and northern 
<collectingRegion id="49E1ED98FF827773FDC493D3BCCAC871" box="[534,667,1777,1801]" country="India" name="Maharashtra" pageId="21" pageNumber="20">Maharashtra</collectingRegion>
(
<collectingCountry id="F33263EAFF827773FD7F93D3BCADC871" box="[685,764,1777,1801]" name="India" pageId="21" pageNumber="20">INDIA</collectingCountry>
) by a raw genetic divergence of 10.2–13.2 % (
<tableCitation id="C6A716C1FF827773FDB99233BC91C851" box="[619,704,1809,1833]" captionStart="Table 5" captionStartId="44.[115,171,1066,1087]" captionText="Table 5. Pairwise uncorrected raw distances (%) expressed as minimum–maximum based on COI gene sequence for Scorpiops species from Maharashtra, India. Values in brackets are intra-clade distances." pageId="21" pageNumber="20">Table 5</tableCitation>
). It is also distinguished from all the species of 
<taxonomicName id="4C2558F9FF827773FD8A9213BC92C830" authorityName="Peters" authorityYear="1861" box="[600,707,1841,1864]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B951FF68FF827773FD8A9213BC92C830" box="[600,707,1841,1864]" italics="true" pageId="21" pageNumber="20">Scorpiops</emphasis>
</taxonomicName>
from peninsular India based on the key of morphological characters (See below).
</paragraph>
</subSubSection>
<subSubSection id="C33F70F1FF827773FFA39293BAA2C891" pageId="21" pageNumber="20" type="distribution">
<paragraph id="8B9A237AFF827773FFA39293BD53C891" blockId="21.[113,770,1969,2025]" pageId="21" pageNumber="20">
DISTRIBUTION, HABITAT AND ECOLOGY. Presently 
<taxonomicName id="4C2558F9FF827773FD469293BE88C891" authority="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade, 2021" authorityName="Sulakhe &amp; Deshpande &amp; Dandekar &amp; Padhye &amp; Bastawade" authorityYear="2021" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="nagphani" status="sp. nov.">
<emphasis id="B951FF68FF827773FD469293BE88C891" italics="true" pageId="21" pageNumber="20">Scorpiops nagphani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624213FF827773FF3792F3BF79C890" box="[229,296,2001,2025]" pageId="21" pageNumber="20" rank="species">
<emphasis id="B951FF68FF827773FF3792F3BEAEC891" bold="true" box="[229,255,2001,2025]" pageId="21" pageNumber="20">sp</emphasis>
. 
<emphasis id="B951FF68FF827773FEC392F3BF71C891" bold="true" box="[273,288,2001,2025]" pageId="21" pageNumber="20">n</emphasis>
.
</taxonomicNameLabel>
is known from its 
<typeStatus id="549E9DD8FF827773FDDE92F3BC6DC891" box="[524,572,2001,2025]" pageId="21" pageNumber="20">type</typeStatus>
locality, Nigdale
</paragraph>
<paragraph id="8B9A237AFF827773FCE29093BAA2C891" blockId="21.[816,1474,1457,2025]" pageId="21" pageNumber="20">
Village, near Bhimashankar Wildlife Sanctuary, Pune District, 
<collectingRegion id="49E1ED98FF827773FC4490F3BA75CA91" box="[918,1060,1489,1513]" country="India" name="Maharashtra" pageId="21" pageNumber="20">Maharashtra</collectingRegion>
, 
<collectingCountry id="F33263EAFF827773FBE190F3BA3DCA91" box="[1075,1132,1489,1513]" name="India" pageId="21" pageNumber="20">India</collectingCountry>
and surrounding areas inside Bhimashankar Wildlife Sanctuary. The 
<typeStatus id="549E9DD8FF827773FACB90D3BB18C971" box="[1305,1353,1521,1545]" pageId="21" pageNumber="20">type</typeStatus>
series is collected from private land around Nigdale Village. Scarce population of this species was observed around the exact 
<typeStatus id="549E9DD8FF827773FCE29373BD31C911" box="[816,864,1617,1641]" pageId="21" pageNumber="20">type</typeStatus>
locality however dense population was observed inside the sanctuary on the hiking route to Nagphani and Sambarshingi. The species was seen in the crevices of basaltic boulders in the semi-evergreen forest patches. Dense population was observed in the boulders adjacent to the streams on the hike route to Nagphani cliff and around the route leading to the forest guest house. Interestingly specimens were also seen below the small rocks placed along the hike route to Sambarshingi. Individuals were found to be active at night sitting at the openings of rock crevices and occasionally in the open. The ecology and morphology of the new species is congruent with the lithophilic scorpions (
<figureCitation id="131E3FFFFF827773FBF492F3BAFCC891" box="[1062,1197,2001,2025]" captionStart="Figures 62–64" captionStartId="21.[115,192,1351,1373]" captionTargetBox="[113,1474,234,1341]" captionTargetId="figure-305@21.[113,1474,234,1341]" captionTargetPageId="21" captionText="Figures 62–64. Type locality of Scorpiops nagphani sp. n., aerial view of semi-evergreen forest inside Bhimashankar wildlife sanctuary (BWLS), very close to the type locality (62), view of Kolvan waterfalls inside BWLS where the new species was observed (63) and Scorpiops nagphani sp. n., female (specimen not collected) in vivo habitus with juveniles (64)." figureDoi="http://doi.org/10.5281/zenodo.5742573" httpUri="https://zenodo.org/record/5742573/files/figure.png" pageId="21" pageNumber="20">Figs. 63, 64</figureCitation>
, 
<figureCitation id="131E3FFFFF827773FB6B92F3BAB7C891" box="[1209,1254,2001,2025]" captionStart="Figure 153" captionStartId="45.[113,181,1216,1238]" captionTargetBox="[114,1474,234,1198]" captionTargetId="figure-7@45.[113,1474,234,1198]" captionTargetPageId="45" captionText="Figure 153. Distribution of Scorpiops species from Peninsular India. Localities with a star indicate new species described in this study." figureDoi="http://doi.org/10.5281/zenodo.5742623" httpUri="https://zenodo.org/record/5742623/files/figure.png" pageId="21" pageNumber="20">153</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>