166 lines
18 KiB
XML
166 lines
18 KiB
XML
<document id="D3BEF014218CB886DA90114FD549155B" ID-DOI="10.5281/zenodo.189132" ID-GBIF-Dataset="86143b99-7b3c-492d-9494-d645fc9c647b" ID-ISSN="1175-5326" ID-Zenodo-Dep="189132" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461099427313" checkinUser="plazi" docAuthor="Hutchinson, Mark N., Doughty, Paul & Oliver, Paul M." docDate="2009" docId="038687D23B65755B65990A213D8913F3" docLanguage="en" docName="zt02167p046.pdf" docOrigin="Zootaxa 2167" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Diplodactylus Gray 1832" docType="treatment" docVersion="9" lastPageNumber="29" masterDocId="FFBFFFAA3B66755F650E0E353D7A115B" masterDocTitle="Taxonomic revision of the stone geckos (Squamata: Diplodactylidae: Diplodactylus) of southern Australia" masterLastPageNumber="46" masterPageNumber="25" pageNumber="28" updateTime="1698590674586" updateUser="plazi">
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<mods:title id="F8C2945074F3D5C4D58149C45E43D12A">Taxonomic revision of the stone geckos (Squamata: Diplodactylidae: Diplodactylus) of southern Australia</mods:title>
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<mods:namePart id="70BAA301017F736B0103793A0F2F5E1E">Hutchinson, Mark N.</mods:namePart>
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<mods:namePart id="67A169C767D5BAA88B100EBA887BDA48">Doughty, Paul</mods:namePart>
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<mods:namePart id="E3AA1BF59B932DC8099CE1D83AB5EA0F">Oliver, Paul M.</mods:namePart>
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<mods:title id="8B17C59F443160EFE8DB288A5C2B4020">Zootaxa</mods:title>
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<mods:date id="F216DB0D507F96A13E47E74BC10819C9">2009</mods:date>
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<mods:number id="EB605484913C72CFBB04EC2C86BFAF39">2167</mods:number>
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<mods:identifier id="75AC86AEE42E8C7A5853210CFB3B1F9F" type="DOI">10.5281/zenodo.189132</mods:identifier>
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<treatment id="038687D23B65755B65990A213D8913F3" ID-DOI="http://doi.org/10.5281/zenodo.6221886" ID-GBIF-Taxon="119621284" ID-Zenodo-Dep="6221886" LSID="urn:lsid:plazi:treatment:038687D23B65755B65990A213D8913F3" httpUri="http://treatment.plazi.org/id/038687D23B65755B65990A213D8913F3" lastPageId="4" lastPageNumber="29" pageId="3" pageNumber="28">
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<subSubSection id="C335654F3B65755C65990A213CB61576" box="[151,460,1043,1070]" pageId="3" pageNumber="28" type="nomenclature">
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<paragraph id="8B9036C43B65755C65990A213CB61576" blockId="3.[151,460,1043,1070]" box="[151,460,1043,1070]" pageId="3" pageNumber="28">
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<heading id="D0D881A83B65755C65990A213CB61576" bold="true" box="[151,460,1043,1070]" fontSize="11" level="1" pageId="3" pageNumber="28" reason="1">
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<taxonomicName id="4C2F4D473B65755C65990A213CB61576" authority="Gray, 1832" authorityName="Gray" authorityYear="1832" box="[151,460,1043,1070]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="28" phylum="Chordata" rank="genus">
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<emphasis id="B95BEAD63B65755C65990A213CB61576" bold="true" box="[151,460,1043,1070]" pageId="3" pageNumber="28">
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<emphasis id="B95BEAD63B65755C65990A213C471575" bold="true" box="[151,317,1044,1070]" italics="true" pageId="3" pageNumber="28">Diplodactylus</emphasis>
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<bibRefCitation id="EFBE4B353B65755C644A0A263CB61576" author="Gray" box="[324,460,1043,1069]" pageId="3" pageNumber="45" refString="Gray, J. E. (1832) Three new animals brought from New Holland by Mr Cunningham. Proceedings of the Zoological Society of London, 1832, 39 - 40." type="proceedings paper" year="1832">Gray, 1832</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C335654F3B65755C65990A683F8F152C" box="[151,757,1117,1143]" pageId="3" pageNumber="28" type="materials_examined">
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<paragraph id="8B9036C43B65755C65990A683F8F152C" blockId="3.[151,1437,1117,2033]" box="[151,757,1117,1143]" pageId="3" pageNumber="28">
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<emphasis id="B95BEAD63B65755C65990A683C48152C" bold="true" box="[151,306,1117,1143]" pageId="3" pageNumber="28">
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<typeStatus id="549488663B65755C65990A683DAE152C" box="[151,212,1117,1143]" pageId="3" pageNumber="28">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName id="4C2F4D473B65755C644F0A6B3F3D152C" authority="Gray, 1832" authorityName="Gray" authorityYear="1832" box="[321,583,1117,1143]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="28" phylum="Chordata" rank="species" species="vittatus">
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<emphasis id="B95BEAD63B65755C644F0A6B3CC6152C" box="[321,444,1118,1143]" italics="true" pageId="3" pageNumber="28">D. vittatus</emphasis>
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<bibRefCitation id="EFBE4B353B65755C64CD0A683F3D152C" author="Gray" box="[451,583,1117,1143]" pageId="3" pageNumber="45" refString="Gray, J. E. (1832) Three new animals brought from New Holland by Mr Cunningham. Proceedings of the Zoological Society of London, 1832, 39 - 40." type="proceedings paper" year="1832">Gray, 1832</bibRefCitation>
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</taxonomicName>
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, by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection id="C335654F3B65755C65C80AB13EDA1404" pageId="3" pageNumber="28" type="diagnosis">
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<paragraph id="8B9036C43B65755C65C80AB13EDA1404" blockId="3.[151,1437,1117,2033]" pageId="3" pageNumber="28">
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<emphasis id="B95BEAD63B65755C65C80AB13C3C15C5" bold="true" box="[198,326,1156,1182]" pageId="3" pageNumber="28">Diagnosis.</emphasis>
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A genus of the
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<taxonomicName id="4C2F4D473B65755C671F0AB13ED215C5" authority="Han et al. 2004" authorityName="Han et al." authorityYear="2004" box="[529,936,1156,1182]" class="Reptilia" family="Diplodactylidae" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="28" phylum="Chordata" rank="family">
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Diplodactylidae (
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<bibRefCitation id="EFBE4B353B65755C67D00AB13EDB15C5" author="Han" box="[734,929,1156,1182]" pageId="3" pageNumber="45" refString="Han, D., Zhou, K. & Bauer, A. M. (2004) Phylogenetic relationships among gekkotan lizards inferred from C-mos nuclear DNA sequences and a new classification of the Gekkota. Biological Journal of the Linnean Society, 83, 353 - 368." type="journal article" year="2004">
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Han
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<emphasis id="B95BEAD63B65755C66120AB13E2315C6" box="[796,857,1156,1181]" italics="true" pageId="3" pageNumber="28">et al.</emphasis>
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2004
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</bibRefCitation>
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)
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</taxonomicName>
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distinguished from all but
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<taxonomicName id="4C2F4D473B65755C61FB0AB1381D15C6" box="[1269,1383,1156,1181]" class="Reptilia" family="Diplodactylidae" genus="Lucasium" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="28" phylum="Chordata" rank="genus">
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<emphasis id="B95BEAD63B65755C61FB0AB1381D15C6" box="[1269,1383,1156,1181]" italics="true" pageId="3" pageNumber="28">Lucasium</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C2F4D473B65755C65990A9E3C38159F" box="[151,322,1195,1220]" class="Reptilia" family="Diplodactylidae" genus="Rhynchoedura" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="28" phylum="Chordata" rank="genus">
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<emphasis id="B95BEAD63B65755C65990A9E3C38159F" box="[151,322,1195,1220]" italics="true" pageId="3" pageNumber="28">Rhynchoedura</emphasis>
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</taxonomicName>
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by having both lateral and medial pairs of cloacal bones present. Distinguished from
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<taxonomicName id="4C2F4D473B65755C60240A9E38E6159F" box="[1322,1436,1195,1220]" class="Reptilia" family="Diplodactylidae" genus="Lucasium" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="28" phylum="Chordata" rank="genus">
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<emphasis id="B95BEAD63B65755C60240A9E38E6159F" box="[1322,1436,1195,1220]" italics="true" pageId="3" pageNumber="28">Lucasium</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C2F4D473B65755C65C50AE43C0615B1" box="[203,380,1233,1258]" class="Reptilia" family="Diplodactylidae" genus="Rhynchoedura" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="28" phylum="Chordata" rank="genus">
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<emphasis id="B95BEAD63B65755C65C50AE43C0615B1" box="[203,380,1233,1258]" italics="true" pageId="3" pageNumber="28">Rhynchoedura</emphasis>
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</taxonomicName>
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by the anteriorly enlarged jugal bone that enters the floor of the lacrimal foramen, by having relatively high numbers of preanal spinose scales (generally> 5), absence of preanal pores and shorter, stouter proportions of the body and tail; fourth toe on hind foot approximately four times as long as wide, tail generally swollen and less than 80% of SVL (
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<bibRefCitation id="EFBE4B353B65755C67A10B703EEE1404" author="Oliver" box="[687,916,1349,1375]" pageId="3" pageNumber="45" refString="Oliver, P. M., Hutchinson, M. N. & Cooper, S. J. B. (2007 b) Phylogenetic relationships in the lizard genus Diplodactylus Gray and resurrection of Lucasium Wermuth (Gekkota, Diplodactylidae). Australian Journal of Zoology, 55, 197 - 210." type="journal article" year="2007" yearSuffix="b">
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Oliver
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<emphasis id="B95BEAD63B65755C660A0B733E451404" box="[772,831,1350,1375]" italics="true" pageId="3" pageNumber="28">et al.</emphasis>
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2007b
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C335654F3B65755B65C80B593D9A1028" lastPageId="4" lastPageNumber="29" pageId="3" pageNumber="28" type="reference_group">
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<paragraph id="8B9036C43B65755C65C80B593C421652" blockId="3.[151,1437,1117,2033]" pageId="3" pageNumber="28">
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<emphasis id="B95BEAD63B65755C65C80B593F1B14DD" bold="true" box="[198,609,1388,1414]" pageId="3" pageNumber="28">
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The
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<taxonomicName id="4C2F4D473B65755C640D0B593CF814DD" box="[259,386,1388,1414]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="28" phylum="Chordata" rank="species" species="vittatus">
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<emphasis id="B95BEAD63B65755C640D0B593CF814DD" bold="true" box="[259,386,1388,1414]" italics="true" pageId="3" pageNumber="28">D. vittatus</emphasis>
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</taxonomicName>
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species complex.
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</emphasis>
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<bibRefCitation id="EFBE4B353B65755C67630B593E1014DD" box="[621,874,1388,1414]" pageId="3" pageNumber="45" refString="Oliver, P., Hugall, A., Adams, M., Cooper, S. J. B. & Hutchinson, M. (2007 a) Genetic elucidation of ancient and cryptic diversity in a group of Australian geckos: the Diplodactylus vittatus complex. Molecular Phylogenetics and Evolution, 44, 77 - 88." type="journal article">
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Oliver
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<emphasis id="B95BEAD63B65755C67CA0B593E7914DE" box="[708,771,1388,1413]" italics="true" pageId="3" pageNumber="28">et al.</emphasis>
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(2007a)
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</bibRefCitation>
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referred to lizards described below as the
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<taxonomicName id="4C2F4D473B65755C60710B593D8B14F7" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="28" phylum="Chordata" rank="species" species="vittatus">
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<emphasis id="B95BEAD63B65755C60710B593D8B14F7" italics="true" pageId="3" pageNumber="28">D. vittatus</emphasis>
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</taxonomicName>
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species complex, and the data presented there and in
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<bibRefCitation id="EFBE4B353B65755C66860BA739FD14F7" box="[904,1159,1426,1452]" pageId="3" pageNumber="45" refString="Oliver, P. M., Hutchinson, M. N. & Cooper, S. J. B. (2007 b) Phylogenetic relationships in the lizard genus Diplodactylus Gray and resurrection of Lucasium Wermuth (Gekkota, Diplodactylidae). Australian Journal of Zoology, 55, 197 - 210." type="journal article">
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Oliver
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<emphasis id="B95BEAD63B65755C66D10BA6396414F7" box="[991,1054,1427,1452]" italics="true" pageId="3" pageNumber="28">et al.</emphasis>
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(2007b)
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</bibRefCitation>
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show that they form a paraphyletic assemblage within
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<emphasis id="B95BEAD63B65755C671A0B8C3FBB1489" box="[532,705,1465,1490]" italics="true" pageId="3" pageNumber="28">
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<taxonomicName id="4C2F4D473B65755C671A0B8C3FC71489" box="[532,701,1465,1490]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="28" phylum="Chordata" rank="genus">Diplodactylus</taxonomicName>
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.
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</emphasis>
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They share a general similarity in habitus and colour pattern, sufficiently similar that all were formerly combined as a single species,
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<taxonomicName id="4C2F4D473B65755C66EF0BD5382914A1" authority="Kluge (1967)" authorityName="Kluge" authorityYear="1967" box="[993,1363,1504,1530]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="28" phylum="Chordata" rank="species" species="vittatus">
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<emphasis id="B95BEAD63B65755C66EF0BD5392114A2" box="[993,1115,1504,1529]" italics="true" pageId="3" pageNumber="28">D. vittatus</emphasis>
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,
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<emphasis id="B95BEAD63B65755C61640BD539D114A2" box="[1130,1195,1504,1529]" italics="true" pageId="3" pageNumber="28">sensu</emphasis>
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<bibRefCitation id="EFBE4B353B65755C61BC0BD5382914A1" author="Kluge" box="[1202,1363,1504,1530]" pageId="3" pageNumber="45" refString="Kluge, A. G. (1967) Systematics, phylogeny and zoogeography of the lizard genus Diplodactylus Gray (Gekkonidae). Australian Journal of Zoology, 15, 1007 - 1108." type="journal article" year="1967">Kluge (1967)</bibRefCitation>
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</taxonomicName>
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. The head and body are moderately stout in general proportions, limbs are of moderate length and build, the tail shows little taper until near the tip and frequently has its maximum diameter at about the midpoint, producing a spindle-shape. The undersides of the digits typically have a single series of enlarged, hemispherical subdigital ‘lamellae’ which tend to break up into smaller scales towards the base of the toes. The labial scales are differentiated from adjacent scales and the nostril is generally surrounded by the rostral, two supranasals, two or three small postnasals, and the first supralabial. There are no tubercular or spinose caudal scales.
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</paragraph>
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<paragraph id="8B9036C43B65755B65C809203D9A1028" blockId="3.[151,1437,1117,2033]" lastBlockId="4.[151,1437,152,680]" lastPageId="4" lastPageNumber="29" pageId="3" pageNumber="28">The species are coloured in shades of grey, brown or reddish-brown and black, with a dorsal pattern that includes a pale vertebral stripe usually scalloped to produce a zigzag shape, but which may be straight-edged, or broken into a series of angular or irregular dorsal blotches. The vertebral stripe may fork on the neck with a process extending to each eye, or may broaden to form a pale cap over the nape and dorsal surface of the occipital and parietal areas. The edges of the vertebral stripe are weakly to boldly edged with blackish paravertebral margins, the colour fading laterally to a medium shade of grey or brown. The light dorsal stripe continues on to the tail when original, but is often less continuous on the tail than on the body. Colouring of the flanks can be divided into an upper lateral zone of a medium to dark shade of the basic body colouring, and a lower lateral zone that is an extension of the ventral colouring. The upper lateral zone may be immaculate, with or without a peppering of paler and darker scales, or may have a few to many small to large circular spots. Limbs are coloured and patterned similarly to the body. The venter is unmarked light grey to white.</paragraph>
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</subSubSection>
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<subSubSection id="C335654F3B62755B65C80FB53D8913F3" pageId="4" pageNumber="29" type="description">
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<paragraph id="8B9036C43B62755B65C80FB53C341355" blockId="4.[151,1437,152,680]" pageId="4" pageNumber="29">
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The only member of the complex to depart from this general scheme is
|
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<taxonomicName id="4C2F4D473B62755B611D0FB5398810C2" box="[1043,1266,384,409]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="29" phylum="Chordata" rank="species" species="polyophthalmus">
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||
<emphasis id="B95BEAD63B62755B611D0FB5398810C2" box="[1043,1266,384,409]" italics="true" pageId="4" pageNumber="29">D. polyophthalmus</emphasis>
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</taxonomicName>
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, which has an indistinct pale vertebral zone and numerous small, blurry pale dorsal spots. The tendency of all of the markings of
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<taxonomicName id="4C2F4D473B62755B64250FF83F7010BD" box="[299,522,461,486]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="29" phylum="Chordata" rank="species" species="polyophthalmus">
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||
<emphasis id="B95BEAD63B62755B64250FF83F7010BD" box="[299,522,461,486]" italics="true" pageId="4" pageNumber="29">D. polyophthalmus</emphasis>
|
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</taxonomicName>
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||
to ‘bleed’ into the surrounding colour distinguishes it from all other members of the complex.
|
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</paragraph>
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<paragraph id="8B9036C43B62755B65C80C2F3D8913F3" blockId="4.[151,1437,152,680]" pageId="4" pageNumber="29">The similar species section is based on areas of sympatry and parapatry, and is designed to draw attention to distinguishing features of species that might be found together. As most species show extensive overlap in morphology, this focuses attention on to those characters that are most useful for particular combinations of species.</paragraph>
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</subSubSection>
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</treatment>
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</document> |