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<mods:title id="EE8C9A7B9520E3A68FB035E7E46582AE">Two new species of Lepidiella Enderlein, 1937 (Diptera: Psychodidae) from the Neotropical Region with taxonomic comments about the species of the genus</mods:title>
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<heading id="D0D881C9FFCEFF9DFF71158D3D862030" bold="true" box="[151,676,1522,1549]" fontSize="11" level="1" pageId="4" pageNumber="491" reason="1">
<emphasis id="B95BEAB7FFCEFF9DFF71158D3D862030" bold="true" box="[151,676,1522,1549]" pageId="4" pageNumber="491">
Key to males of
<taxonomicName id="4C2F4D26FFCEFF9DFEB9158C3D862030" ID-CoL="5CF3" ID-ENA="1144717" authority="Enderlein, 1937" authorityName="Enderlein" authorityYear="1937" box="[351,676,1522,1549]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="491" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEAB7FFCEFF9DFEB9158C3EF52031" bold="true" box="[351,471,1522,1548]" italics="true" pageId="4" pageNumber="491">Lepidiella</emphasis>
<bibRefCitation id="EFBE4B54FFCEFF9DFE39158D3D862030" author="Enderlein, G." box="[479,676,1523,1549]" pageId="4" pageNumber="491" pagination="81 - 112" refId="ref4708" refString="Enderlein, G. (1937) Klassifikation der Psychodiden (Dipt.). Deutsche Entomologische Zeitschrift, 1936, 81 - 112." type="journal article" year="1937">Enderlein, 1937</bibRefCitation>
</taxonomicName>
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<paragraph id="8B9036A5FFCEFF9DFF7116463AB92073" blockId="4.[151,1436,1592,2025]" box="[151,1435,1592,1614]" pageId="4" pageNumber="491">
<keyLead id="30DE9590FFCEFF9DFF7116463AB92073" box="[151,1435,1592,1614]" pageId="4" pageNumber="491">1 Head without cornicula............................................................................... 2</keyLead>
</paragraph>
<paragraph id="8B9036A5FFCEFF9DFF7116293AB92051" blockId="4.[151,1436,1592,2025]" box="[151,1435,1622,1644]" pageId="4" pageNumber="491">
<keyLead id="30DE9590FFCEFF9DFF7116293AB92051" box="[151,1435,1622,1644]" pageId="4" pageNumber="491">- Head with multilobed cornicula......................................................................... 6</keyLead>
</paragraph>
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<keyStep id="30DB2E00FFCEFF9DFF71160D3AB9209B" pageId="4" pageNumber="491">
<paragraph id="8B9036A5FFCEFF9DFF71160D3AB920B4" blockId="4.[151,1436,1592,2025]" box="[151,1435,1651,1673]" pageId="4" pageNumber="491">
<keyLead id="30DE9590FFCEFF9DFF71160D3AB920B4" box="[151,1435,1651,1673]" pageId="4" pageNumber="491">
2 Internal paramere with bifurcated apex..................................
<taxonomicName id="4C2F4D26FFCEFF9DFC18160A3B5620B4" ID-CoL="8CH6T" authorityName="Bravo &amp; Santos" authorityYear="2011" box="[1022,1140,1651,1673]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="491" phylum="Arthropoda" rank="species" species="flabellata">
<emphasis id="B95BEAB7FFCEFF9DFC18160A3B5620B4" box="[1022,1140,1651,1673]" italics="true" pageId="4" pageNumber="491">L. flabellata</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFBE4B54FFCEFF9DFB66160D3AB720B4" author="Bravo, F. &amp; Santos, C. B." box="[1152,1429,1651,1673]" pageId="4" pageNumber="491" pagination="264 - 268" refId="ref4301" refString="Bravo, F. &amp; Santos, C. B. (2011) Two new species of Lepidiella (Diptera: Psychodidae: Psychodinae) from the Atlantic Rainforest of southeastern Brazil, Zoologia, 28, 264 - 268. https: // doi. org / 10.1590 / s 1984 - 46702011000200017" type="journal article" year="2011">Bravo &amp; Santos 2011, Fig. 6</bibRefCitation>
)
</keyLead>
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<paragraph id="8B9036A5FFCEFF9DFF7116EF3AB9209B" blockId="4.[151,1436,1592,2025]" box="[151,1435,1680,1702]" pageId="4" pageNumber="491">
<keyLead id="30DE9590FFCEFF9DFF7116EF3AB9209B" box="[151,1435,1680,1702]" pageId="4" pageNumber="491">- Internal paramere not bifurcated, blade shaped............................................................. 3</keyLead>
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<keyLead id="30DE9590FFCEFF9DFF7116D03ABE20F9" box="[151,1436,1710,1732]" pageId="4" pageNumber="491">
3 Ejaculatory apodeme narrow in dorsal view........................................
<taxonomicName id="4C2F4D26FFCEFF9DFB8916D13BF320F9" ID-CoL="6P9M2" authorityName="Bravo" authorityYear="2005" box="[1135,1233,1711,1732]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="491" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="B95BEAB7FFCEFF9DFB8916D13BF320F9" box="[1135,1233,1711,1732]" italics="true" pageId="4" pageNumber="491">L. spinosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFBE4B54FFCEFF9DFB3816D03AB620F9" author="Bravo, F." box="[1246,1428,1710,1732]" pageId="4" pageNumber="491" pagination="490 - 493" refId="ref4246" refString="Bravo, F. (2005) Primeiro registro de Lepidiella Enderlein (Diptera, Psychodidae, Psychodinae) no Brasil e descricao de uma especie nova. Revista Brasileira de Zoologia, 22 (2), 490 - 493. https: // doi. org / 10.1590 / S 0101 - 81752005000200027" type="journal article" year="2005">Bravo 2005, Fig. 5</bibRefCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B9036A5FFCEFF9DFF7116B23AB920DC" blockId="4.[151,1436,1592,2025]" box="[151,1435,1739,1761]" pageId="4" pageNumber="491">
<keyLead id="30DE9590FFCEFF9DFF7116B23AB920DC" box="[151,1435,1739,1761]" pageId="4" pageNumber="491">- Ejaculatory apodeme broad in dorsal view, wider than gonocoxite............................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB2E00FFCEFF9DFF7116963AB92121" pageId="4" pageNumber="491">
<paragraph id="8B9036A5FFCEFF9DFF7116963ABE20C3" blockId="4.[151,1436,1592,2025]" box="[151,1436,1768,1790]" pageId="4" pageNumber="491">
<keyLead id="30DE9590FFCEFF9DFF7116963ABE20C3" box="[151,1436,1768,1790]" pageId="4" pageNumber="491">
4 Ejaculatory apodeme short, 0.7 times the length of gonocoxite..........................................
<taxonomicName id="4C2F4D26FFCEFF9DFAD016973ABE20C3" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[1334,1436,1768,1790]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="491" phylum="Arthropoda" rank="species" species="hansoni">
<emphasis id="B95BEAB7FFCEFF9DFAD016973ABE20C3" box="[1334,1436,1768,1790]" italics="true" pageId="4" pageNumber="491">L. hansoni</emphasis>
</taxonomicName>
</keyLead>
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<paragraph id="8B9036A5FFCEFF9DFF7117793AB92121" blockId="4.[151,1436,1592,2025]" box="[151,1435,1798,1820]" pageId="4" pageNumber="491">
<keyLead id="30DE9590FFCEFF9DFF7117793AB92121" box="[151,1435,1798,1820]" pageId="4" pageNumber="491">- Ejaculatory apodeme long, 2 times the length of gonocoxite.................................................. 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB2E00FFCEFF9DFF71175D3AB421AC" pageId="4" pageNumber="491">
<paragraph id="8B9036A5FFCEFF9DFF71175D3ABE216B" blockId="4.[151,1436,1592,2025]" pageId="4" pageNumber="491">
<keyLead id="30DE9590FFCEFF9DFF71175D3ABE216B" pageId="4" pageNumber="491">
5 Margins of pedicel divergent distally larger than base; hypandrium subrectangular with microtrichia sparrow uniformly, not arranged in circular groups.............................................
<taxonomicName id="4C2F4D26FFCEFF9DFBF6173F3B7D216B" ID-CoL="9HYCZ" authorityName="Bravo &amp; Araujo" authorityYear="2013" box="[1040,1119,1856,1878]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="491" phylum="Arthropoda" rank="species" species="olgae">
<emphasis id="B95BEAB7FFCEFF9DFBF6173F3B7D216B" box="[1040,1119,1856,1878]" italics="true" pageId="4" pageNumber="491">L. olgae</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFBE4B54FFCEFF9DFB8D173E3AB6216B" author="Bravo, F. &amp; Araujo, M. X." box="[1131,1428,1856,1878]" pageId="4" pageNumber="491" pagination="85 - 90" refId="ref4356" refString="Bravo, F. &amp; Araujo, M. X. (2013) Two new species of moth flies (Diptera, Psychodidae) from the semi-arid region of Brazil. Zootaxa, 3693 (1), 85 - 90. https: // doi. org / 10.11646 / zootaxa. 3693.1.6" type="journal article" year="2013">
Bravo &amp; Araújo 2013,
<figureCitation id="13142A20FFCEFF9DFAAC173E3AB6216B" box="[1354,1428,1856,1878]" captionStart="FIGURE 2" captionStartId="3.[151,250,1877,1899]" captionTargetBox="[166,1416,411,1852]" captionTargetId="figure@3.[166,1426,411,1852]" captionTargetPageId="3" captionText="FIGURE 2(AJ). Lepidiella wagneri sp. nov. (A) head; (B) oval gland of cornicula; (C) base of antenna: scape, pedicel and two basal flagellomeres; (D) last flagellomeres and apiculus; (E) last flagellomeres and apiculus (other format); (F) wing; (G) male terminalia, dorsal; (H) male terminalia, ventral: epandrium, cerci and hypoproct; (I) cercus; (J) female terminalia. (Abbreviations: ae = aedeagus, ep = external paramere, ip = internal paramere)." httpUri="https://zenodo.org/record/2623147/files/figure.png" pageId="4" pageNumber="491">Fig. 2H</figureCitation>
</bibRefCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B9036A5FFCEFF9DFF7117213AB421AC" blockId="4.[151,1436,1592,2025]" pageId="4" pageNumber="491">
<keyLead id="30DE9590FFCEFF9DFF7117213AB421AC" pageId="4" pageNumber="491">
- Margins of pedicel parallel; hypandrium wider in the middle portion with microtrichia arranged in circular groups............................................................................................
<taxonomicName id="4C2F4D26FFCEFF9DFB2817023A6121AC" authority="Araújo &amp; Bravo, 2019" authorityName="Araújo &amp; Bravo" authorityYear="2019" box="[1230,1347,1915,1937]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="491" phylum="Arthropoda" rank="species" species="maculosa" status="sp. nov.">
<emphasis id="B95BEAB7FFCEFF9DFB2817023A6121AC" box="[1230,1347,1915,1937]" italics="true" pageId="4" pageNumber="491">L. maculosa</emphasis>
</taxonomicName>
<emphasis id="B95BEAB7FFCEFF9DFAAF17053AB421AC" bold="true" box="[1353,1430,1915,1937]" pageId="4" pageNumber="491">
<taxonomicNameLabel id="A26857CCFFCEFF9DFAAF17053AB421AC" box="[1353,1430,1915,1937]" pageId="4" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<keyStep id="30DB2E00FFCEFF9DFF7117E63AB921F1" pageId="4" pageNumber="491">
<paragraph id="8B9036A5FFCEFF9DFF7117E63AB92193" blockId="4.[151,1436,1592,2025]" box="[151,1435,1944,1966]" pageId="4" pageNumber="491">
<keyLead id="30DE9590FFCEFF9DFF7117E63AB92193" box="[151,1435,1944,1966]" pageId="4" pageNumber="491">6 Scape with a pronounced internal distal lobe............................................................... 7</keyLead>
</paragraph>
<paragraph id="8B9036A5FFCEFF9DFF7117C93AB921F1" blockId="4.[151,1436,1592,2025]" box="[151,1435,1974,1996]" pageId="4" pageNumber="491">
<keyLead id="30DE9590FFCEFF9DFF7117C93AB921F1" box="[151,1435,1974,1996]" pageId="4" pageNumber="491">- Scape cylindrical or with a short internal distal expansion................................................... 12</keyLead>
</paragraph>
</keyStep>
<paragraph id="8B9036A5FFCEFF9CFF7117AA3AB92690" blockId="4.[151,1436,1592,2025]" lastBlockId="5.[151,1436,151,671]" lastPageId="5" lastPageNumber="492" pageId="4" pageNumber="491">
<keyStep id="30DB2E00FFCEFF9CFF7117AA3AB92690" lastPageId="5" lastPageNumber="492" pageId="4" pageNumber="491">
<keyLead id="30DE9590FFCEFF9CFF7117AA3AB92690" lastPageId="5" lastPageNumber="492" pageId="4" pageNumber="491">
7 First flagellomere 3.0 X as long as the second.......................................................
<taxonomicName id="4C2F4D26FFCEFF9DFAD217AA3AB721D4" ID-CoL="8CH6X" authorityName="Bravo &amp; Santos" authorityYear="2011" box="[1332,1429,2003,2025]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="491" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B95BEAB7FFCEFF9DFAD217AA3AB721D4" box="[1332,1429,2003,2025]" italics="true" pageId="4" pageNumber="491">L. robusta</emphasis>
</taxonomicName>
- First flagellomere nearly the same length of second......................................................... 8
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30DB2E00FFCFFF9CFF7110CA3AB926DA" pageId="5" pageNumber="492">
<paragraph id="8B9036A5FFCFFF9CFF7110CA3AB426F7" blockId="5.[151,1436,151,671]" box="[151,1430,180,202]" pageId="5" pageNumber="492">
<keyLead id="30DE9590FFCFFF9CFF7110CA3AB426F7" box="[151,1430,180,202]" pageId="5" pageNumber="492">
8 Scape 3.5 X as long as pedicel..............................................................
<taxonomicName id="4C2F4D26FFCFFF9CFB1D10CB3AB426F7" ID-CoL="6P9LT" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[1275,1430,180,202]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="492" phylum="Arthropoda" rank="species" species="monteverdica">
<emphasis id="B95BEAB7FFCFFF9CFB1D10CB3AB426F7" box="[1275,1430,180,202]" italics="true" pageId="5" pageNumber="492">L. monteverdica</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9036A5FFCFFF9CFF7110AC3AB926DA" blockId="5.[151,1436,151,671]" box="[151,1435,209,231]" pageId="5" pageNumber="492">
<keyLead id="30DE9590FFCFFF9CFF7110AC3AB926DA" box="[151,1435,209,231]" pageId="5" pageNumber="492">- Scape 2.0 2.5 times the length of pedicel................................................................ 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB2E00FFCFFF9CFF7110913ABE271F" pageId="5" pageNumber="492">
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<keyLead id="30DE9590FFCFFF9CFF7110913AB92738" box="[151,1435,239,261]" pageId="5" pageNumber="492">
9 Aedeagal apodeme narrow in dorsal view, narrower than the gonocoxite................
<taxonomicName id="4C2F4D26FFCFFF9CFB83108E3B8D2738" ID-CoL="6P9M7" baseAuthorityName="Satchell" baseAuthorityYear="1955" box="[1125,1199,240,261]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="492" phylum="Arthropoda" rank="species" species="cervi">
<emphasis id="B95BEAB7FFCFFF9CFB83108E3B8D2738" box="[1125,1199,240,261]" italics="true" pageId="5" pageNumber="492">L. cervi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFBE4B54FFCFFF9CFB5D10913AB62738" author="Satchell, G. H." box="[1211,1428,239,261]" pageId="5" pageNumber="492" pagination="85 - 93" refId="ref5161" refString="Satchell, G. H. (1955) XI. - Two new subgenera of Psychodidae (Diptera) from Jamaica, with descriptions of five new species. Annals and Magazine of Natural History, Series 12, 8 (86), 85 - 93. https: // doi. org / 10.1080 / 00222935508651831" type="journal article" year="1955">Satchell 1955, Fig. 5C</bibRefCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B9036A5FFCFFF9CFF7111733ABE271F" blockId="5.[151,1436,151,671]" box="[151,1436,268,290]" pageId="5" pageNumber="492">
<keyLead id="30DE9590FFCFFF9CFF7111733ABE271F" box="[151,1436,268,290]" pageId="5" pageNumber="492">- Aedeagal apodeme broad in dorsal view, with approximately the same width as the gonocoxite..................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB2E00FFCFFF9CFF7111573AB92760" pageId="5" pageNumber="492">
<paragraph id="8B9036A5FFCFFF9CFF7111573ABE2702" blockId="5.[151,1436,151,671]" box="[151,1436,297,319]" pageId="5" pageNumber="492">
<keyLead id="30DE9590FFCFFF9CFF7111573ABE2702" box="[151,1436,297,319]" pageId="5" pageNumber="492">
10 Eyes separated by 3.5 facet diameters.............................................................
<taxonomicName id="4C2F4D26FFCFFF9CFAC811543ABE2702" ID-CoL="8H4QC" baseAuthorityName="Rapp" baseAuthorityYear="1945" box="[1326,1436,297,319]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="492" phylum="Arthropoda" rank="species" species="albipeda">
<emphasis id="B95BEAB7FFCFFF9CFAC811543ABE2702" box="[1326,1436,297,319]" italics="true" pageId="5" pageNumber="492">L. albipeda</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9036A5FFCFFF9CFF7111393AB92760" blockId="5.[151,1436,151,671]" box="[151,1435,327,349]" pageId="5" pageNumber="492">
<keyLead id="30DE9590FFCFFF9CFF7111393AB92760" box="[151,1435,327,349]" pageId="5" pageNumber="492">- Eyes separated by 1.5 facet diameters................................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB2E00FFCFFF9CFF71111A3ABE2788" pageId="5" pageNumber="492">
<paragraph id="8B9036A5FFCFFF9CFF71111A3ABE2747" blockId="5.[151,1436,151,671]" box="[151,1436,356,378]" pageId="5" pageNumber="492">
<keyLead id="30DE9590FFCFFF9CFF71111A3ABE2747" box="[151,1436,356,378]" pageId="5" pageNumber="492">
11 Aedeagus digitiform; ejaculatory apodeme 3 times the length of gonocoxite; hypandrium triangular.....
<taxonomicName id="4C2F4D26FFCFFF9CFB07111B3A6B2747" authority="Araújo &amp; Bravo, 2019" authorityName="Araújo &amp; Bravo" authorityYear="2019" box="[1249,1353,357,378]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="492" phylum="Arthropoda" rank="species" species="wagneri" status="sp. nov.">
<emphasis id="B95BEAB7FFCFFF9CFB07111B3A6B2747" box="[1249,1353,357,378]" italics="true" pageId="5" pageNumber="492">L. wagneri</emphasis>
</taxonomicName>
<emphasis id="B95BEAB7FFCFFF9CFAA9111A3ABE2747" bold="true" box="[1359,1436,356,378]" pageId="5" pageNumber="492">
<taxonomicNameLabel id="A26857CCFFCFFF9CFAA9111A3ABE2747" box="[1359,1436,356,378]" pageId="5" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9036A5FFCFFF9CFF7111FC3ABE2788" blockId="5.[151,1436,151,671]" pageId="5" pageNumber="492">
<keyLead id="30DE9590FFCFFF9CFF7111FC3ABE2788" pageId="5" pageNumber="492">
- Aedeagus with ending in a dorsal spiraliform tip; ejaculatory apodeme 1.2 times the length of gonocoxite; hypandrium subrect- angular, narrow...........................................................
<taxonomicName id="4C2F4D26FFCFFF9CFBAE11DE3BE12788" ID-CoL="6P9LQ" baseAuthorityName="Quate" baseAuthorityYear="1999" box="[1096,1219,415,437]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="492" phylum="Arthropoda" rank="species" species="pickeringi">
<emphasis id="B95BEAB7FFCFFF9CFBAE11DE3BE12788" box="[1096,1219,415,437]" italics="true" pageId="5" pageNumber="492">L. pickeringi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFBE4B54FFCFFF9CFB2911E13AB72788" author="Quate, L. W." box="[1231,1429,415,437]" pageId="5" pageNumber="492" pagination="1 - 648" refId="ref4976" refString="Quate, L. W. (1999) Taxonomy of Neotropical Psychodidae (Diptera) 3. Psychodines of Barro Colorado Island and San Blas, Panama. In: Burger, J. F. (Ed.), Contributions to the Knowledge of Diptera. Memoirs on Entomology, International, 14, pp. 1 - 648." type="journal article" year="1999">
Quate 1999,
<figureCitation id="13142A20FFCFFF9CFAAA11E13AB72788" box="[1356,1429,415,437]" captionStart="FIGURE 2" captionStartId="3.[151,250,1877,1899]" captionTargetBox="[166,1416,411,1852]" captionTargetId="figure@3.[166,1426,411,1852]" captionTargetPageId="3" captionText="FIGURE 2(AJ). Lepidiella wagneri sp. nov. (A) head; (B) oval gland of cornicula; (C) base of antenna: scape, pedicel and two basal flagellomeres; (D) last flagellomeres and apiculus; (E) last flagellomeres and apiculus (other format); (F) wing; (G) male terminalia, dorsal; (H) male terminalia, ventral: epandrium, cerci and hypoproct; (I) cercus; (J) female terminalia. (Abbreviations: ae = aedeagus, ep = external paramere, ip = internal paramere)." httpUri="https://zenodo.org/record/2623147/files/figure.png" pageId="5" pageNumber="492">Fig. 2C</figureCitation>
</bibRefCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB2E00FFCFFF9CFF7111C23AB927D2" pageId="5" pageNumber="492">
<paragraph id="8B9036A5FFCFFF9CFF7111C23ABE27EF" blockId="5.[151,1436,151,671]" box="[151,1436,444,466]" pageId="5" pageNumber="492">
<keyLead id="30DE9590FFCFFF9CFF7111C23ABE27EF" box="[151,1436,444,466]" pageId="5" pageNumber="492">
12 Eyes separated by 1 facet diameters.............................................................
<taxonomicName id="4C2F4D26FFCFFF9CFAF811C33ABE27EF" ID-CoL="8CH6W" baseAuthorityName="Enderlein" baseAuthorityYear="1937" box="[1310,1436,445,466]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="492" phylum="Arthropoda" rank="species" species="niveitarsis">
<emphasis id="B95BEAB7FFCFFF9CFAF811C33ABE27EF" box="[1310,1436,445,466]" italics="true" pageId="5" pageNumber="492">L. niveitarsis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9036A5FFCFFF9CFF7111A43AB927D2" blockId="5.[151,1436,151,671]" box="[151,1435,473,495]" pageId="5" pageNumber="492">
<keyLead id="30DE9590FFCFFF9CFF7111A43AB927D2" box="[151,1435,473,495]" pageId="5" pageNumber="492">- Eyes separated by 2 or three facet diameters.............................................................. 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB2E00FFCFFF9CFF7111893ABE2417" pageId="5" pageNumber="492">
<paragraph id="8B9036A5FFCFFF9CFF7111893AB92430" blockId="5.[151,1436,151,671]" box="[151,1435,503,525]" pageId="5" pageNumber="492">
<keyLead id="30DE9590FFCFFF9CFF7111893AB92430" box="[151,1435,503,525]" pageId="5" pageNumber="492">
13 Radial and medial forks at the same level; first flagellomere 2.0 X longer than second one................
<taxonomicName id="4C2F4D26FFCFFF9CFAFE11863AB92430" ID-CoL="8CH6S" authorityName="Enderlein" authorityYear="1937" box="[1304,1435,503,525]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="492" phylum="Arthropoda" rank="species" species="lanuginosa">
<emphasis id="B95BEAB7FFCFFF9CFAFE11863AB92430" box="[1304,1435,503,525]" italics="true" pageId="5" pageNumber="492">L. lanuginosa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9036A5FFCFFF9CFF71126B3ABE2417" blockId="5.[151,1436,151,671]" box="[151,1436,532,554]" pageId="5" pageNumber="492">
<keyLead id="30DE9590FFCFFF9CFF71126B3ABE2417" box="[151,1436,532,554]" pageId="5" pageNumber="492">- Radial fork apical to medial fork; first flagellomere with the same length or 1.3 X longer than the second one.......... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB2E00FFCFFF9CFF71124F3AB92458" pageId="5" pageNumber="492">
<paragraph id="8B9036A5FFCFFF9CFF71124F3AB9247A" blockId="5.[151,1436,151,671]" box="[151,1435,561,583]" pageId="5" pageNumber="492">
<keyLead id="30DE9590FFCFFF9CFF71124F3AB9247A" box="[151,1435,561,583]" pageId="5" pageNumber="492">
14 Costal area convex, expanded...........................
<taxonomicName id="4C2F4D26FFCFFF9CFCA9124C3CD1247A" ID-CoL="6P9M4" baseAuthorityName="Collantes &amp; Martinez-Ortega" baseAuthorityYear="1998" box="[847,1011,561,583]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="492" phylum="Arthropoda" rank="species" species="matagalpensis">
<emphasis id="B95BEAB7FFCFFF9CFCA9124C3CD1247A" box="[847,1011,561,583]" italics="true" pageId="5" pageNumber="492">L. matagalpensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFBE4B54FFCFFF9CFC19124F3AB7247A" author="Collantes, F. &amp; Martinez-Ortega, E." box="[1023,1429,561,583]" pageId="5" pageNumber="492" pagination="209 - 213" refId="ref4518" refString="Collantes, F. &amp; Martinez-Ortega, E. (1998) Syntomoza matagalpensis, a new Moth-Fly from Nicaragua (Diptera, Psychodidae). Aquatic Insects, 20 (4), 209 - 213. https: // doi. org / 10.1076 / aqin. 20.4.209.4468" type="journal article" year="1998">Collantes &amp; Martínez-Ortega 1998, Fig. 4</bibRefCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B9036A5FFCFFF9CFF7112313AB92458" blockId="5.[151,1436,151,671]" box="[151,1435,591,613]" pageId="5" pageNumber="492">
<keyLead id="30DE9590FFCFFF9CFF7112313AB92458" box="[151,1435,591,613]" pageId="5" pageNumber="492">- Costal area narrow................................................................................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB2E00FFCFFF9CFF7112123AB924A2" pageId="5" pageNumber="492">
<paragraph id="8B9036A5FFCFFF9CFF7112123ABE24BF" blockId="5.[151,1436,151,671]" box="[151,1436,620,642]" pageId="5" pageNumber="492">
<keyLead id="30DE9590FFCFFF9CFF7112123ABE24BF" box="[151,1436,620,642]" pageId="5" pageNumber="492">
15 Aedeagus and parameres asymmetrical........................................
<taxonomicName id="4C2F4D26FFCFFF9CFBAD12133BE324BF" ID-CoL="8H2M8" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[1099,1217,620,642]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="492" phylum="Arthropoda" rank="species" species="zumbadoi">
<emphasis id="B95BEAB7FFCFFF9CFBAD12133BE324BF" box="[1099,1217,620,642]" italics="true" pageId="5" pageNumber="492">L. zumbadoi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFBE4B54FFCFFF9CFB2812123AB724BF" author="Quate, L. W." box="[1230,1429,620,642]" pageId="5" pageNumber="492" pagination="1 - 81" refId="ref4937" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae, Revista de Biologia Tropical, 44 (Supl. 1), 1 - 81." type="journal article" year="1996">Quate 1996, Fig. 5D</bibRefCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B9036A5FFCFFF9CFF7112F43AB924A2" blockId="5.[151,1436,151,671]" box="[151,1435,649,671]" pageId="5" pageNumber="492">
<keyLead id="30DE9590FFCFFF9CFF7112F43AB924A2" box="[151,1435,649,671]" pageId="5" pageNumber="492">
- Aedeagus and parameres symmetrical........................................
<taxonomicName id="4C2F4D26FFCFFF9CFBD212F43BA624A2" ID-CoL="8CH6V" authorityName="Ibanez-Bernal" authorityYear="2010" box="[1076,1156,649,671]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="492" phylum="Arthropoda" rank="species" species="larryi">
<emphasis id="B95BEAB7FFCFFF9CFBD212F43BA624A2" box="[1076,1156,649,671]" italics="true" pageId="5" pageNumber="492">L. larryi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFBE4B54FFCFFF9CFB7712F73AB724A2" author="Ibanez-Bernal, S." box="[1169,1429,649,671]" pageId="5" pageNumber="492" pagination="35 - 40" refId="ref4793" refString="Ibanez-Bernal, S. (2010) A new species of the genus Lepidiella Enderlein (Diptera: Psychodidae) from Colombia. Entomological News, 121, 35 - 40. https: // doi. org / 10.3157 / 021.121.0107" type="journal article" year="2010">Ibañez-Bernal 2010, Fig. 6</bibRefCitation>
)
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>