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<document id="45B0A1E3847B7330BE19F1581C4F0D3E" ID-CLB-Dataset="5940" ID-DOI="10.11646/zootaxa.3761.1.1" ID-GBIF-Dataset="5444e27e-3e2a-429b-931b-823a2086e3c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4909057" ID-ZooBank="82E0F1DC-BC98-4E8A-A3D5-21ECB392CC0B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623107451891" checkinUser="felipe" docAuthor="Luk, Stephen P. L. &amp; Marshall, Stephen A." docDate="2014" docId="038487F1FF82FF9AFDC7FB55FF1F0F00" docLanguage="en" docName="zt03761p156.pdf" docOrigin="Zootaxa 3761 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Aptilotella quadrata Luk &amp; Marshall 2014, sp. n." docType="treatment" docVersion="8" lastPageNumber="42" masterDocId="FFBDFF89FFA5FFB3FD50FE2BFFC70D18" masterDocTitle="A revision of the New World genus Aptilotella Duda (Sphaeroceridae: Limosininae)" masterLastPageNumber="156" masterPageNumber="1" pageNumber="40" updateTime="1699002622211" updateUser="ExternalLinkService">
<mods:mods id="CDE88929AB859AE24246CB390191376E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0299F4AD3E1B3DA922E9E0C88516CD1E">
<mods:title id="425A436A979C59900AE3894131D9207E">A revision of the New World genus Aptilotella Duda (Sphaeroceridae: Limosininae)</mods:title>
</mods:titleInfo>
<mods:name id="C856477B715AF7BB9729992949533E28" type="personal">
<mods:role id="DD828443AE794AD599397EE1D396E22E">
<mods:roleTerm id="C9E95C14721F20CE0A607E6734F31D48">Author</mods:roleTerm>
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<mods:namePart id="0FC18264288D1EC03A071FF72044515C">Luk, Stephen P. L.</mods:namePart>
<mods:nameIdentifier id="1840A49E6D2203C9DE76A3BA05AA9E96" type="email">luk@uoguelph.ca</mods:nameIdentifier>
</mods:name>
<mods:name id="6266B4F45FB11F1C92F8DC1D1BB22079" type="personal">
<mods:role id="435F71152CA89DE30186A2BF262516A8">
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<mods:namePart id="D0016624C404B9167775AFA81B09C53B">Marshall, Stephen A.</mods:namePart>
<mods:nameIdentifier id="286E9190EA2E72182FD6AF4F31D5E0A3" type="email">luk@uoguelph.ca</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="94979A3E9C196727FECBA52D82C77B4A">text</mods:typeOfResource>
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<mods:titleInfo id="F408A719ABD43BACF6373DDD8E88DCB2">
<mods:title id="4333D3028B77DE9FA49FCE48B3864273">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8444B91102E544C5D43F48F2404308D4">
<mods:date id="E4626D98F64CA8B692394739E79154B9">2014</mods:date>
<mods:detail id="56AAFDC0DFE944738E8032D53348920C" type="pubDate">
<mods:number id="9A32884462B6447E67C46335E98AAC08">2014-02-05</mods:number>
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<mods:detail id="EF5E89ACD88624DE9ACA6CB4AF627BED" type="volume">
<mods:number id="6989452678B3B83C88DF9CE9FD735D6C">3761</mods:number>
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<mods:detail id="0B3440D661223F6084D2EB9546812A0B" type="issue">
<mods:number id="5CB8AC3FDF486E27C217D33660425266">1</mods:number>
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<mods:start id="A18434BAF8EECEE955144F4126D37348">1</mods:start>
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<mods:classification id="2A2D2B49D8E49B3C931C0B9270A3A743">journal article</mods:classification>
<mods:identifier id="7B09A0D589E9ACFE08676F32745D09F7" type="CLB-Dataset">5940</mods:identifier>
<mods:identifier id="E03DF17E5AED053A12A86D81D85A8D54" type="DOI">10.11646/zootaxa.3761.1.1</mods:identifier>
<mods:identifier id="973FAD1042BEA8D99BB5239863D99DD8" type="GBIF-Dataset">5444e27e-3e2a-429b-931b-823a2086e3c5</mods:identifier>
<mods:identifier id="A211C77C4C0B8EAA71CA2D249245A60D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A6C484E50C363933DE3F676E95B4992F" type="Zenodo-Dep">4909057</mods:identifier>
<mods:identifier id="4313AC3E2D6BBDC25BE74D2CA4DEDE7D" type="ZooBank">82E0F1DC-BC98-4E8A-A3D5-21ECB392CC0B</mods:identifier>
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<treatment id="038487F1FF82FF9AFDC7FB55FF1F0F00" ID-DOI="http://doi.org/10.5281/zenodo.4909147" ID-GBIF-Taxon="183203096" ID-Zenodo-Dep="4909147" LSID="urn:lsid:plazi:treatment:038487F1FF82FF9AFDC7FB55FF1F0F00" httpUri="http://treatment.plazi.org/id/038487F1FF82FF9AFDC7FB55FF1F0F00" lastPageId="41" lastPageNumber="42" pageId="39" pageNumber="40">
<subSubSection id="C337656CFF82FF94FDC7FB55FD6A0880" box="[151,685,1406,1432]" pageId="39" pageNumber="40" type="nomenclature">
<paragraph id="8B9236E7FF82FF94FDC7FB55FD6A0880" blockId="39.[151,685,1406,1466]" box="[151,685,1406,1432]" pageId="39" pageNumber="40">
<heading id="D0DA818BFF82FF94FDC7FB55FD6A0880" bold="true" box="[151,685,1406,1432]" fontSize="11" level="2" pageId="39" pageNumber="40" reason="4">
<emphasis id="B959EAF5FF82FF94FDC7FB55FD6A0880" bold="true" box="[151,685,1406,1432]" pageId="39" pageNumber="40">
<taxonomicName id="4C2D4D64FF82FF94FDC7FB55FD980880" authority="Luk &amp; Marshall" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[151,607,1406,1432]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="quadrata" status="sp. nov.">
<emphasis id="B959EAF5FF82FF94FDC7FB55FE4F0880" bold="true" box="[151,392,1406,1432]" italics="true" pageId="39" pageNumber="40">Aptilotella quadrata</emphasis>
Luk &amp; Marshall
</taxonomicName>
,
<taxonomicNameLabel id="A26A578EFF82FF94FF3BFB55FD6A0880" box="[619,685,1406,1432]" pageId="39" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C337656CFF82FF9BFDC7FB8AFE450BFE" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="description">
<paragraph id="8B9236E7FF82FF94FDC7FB8AFE5F08A2" blockId="39.[151,685,1406,1466]" box="[151,408,1441,1466]" pageId="39" pageNumber="40">
<figureCitation id="13162A62FF82FF94FDC7FB8AFEC708A2" box="[151,256,1441,1466]" captionStart="FIGURE 3" captionStartId="40.[151,249,884,906]" captionTargetId="figure-472@40.[369,1218,195,862]" captionTargetPageId="40" captionText="FIGURE 3. Aptilotella quadrata Luk &amp; Marshall, sp. n., holotype male." figureDoi="http://doi.org/10.5281/zenodo.5042241" httpUri="https://zenodo.org/record/5042241/files/figure.png" pageId="39" pageNumber="40">Figures 3</figureCitation>
and
<figureCitation id="13162A62FF82FF94FC66FB8AFE5F08A2" box="[310,408,1441,1466]" captionStart-0="FIGURES 288–291" captionStart-1="FIGURES 292–294" captionStartId-0="141.[151,264,1258,1281]" captionStartId-1="142.[151,264,1143,1166]" captionTargetBox-0="[394,1191,201,1229]" captionTargetBox-1="[426,1161,196,1118]" captionTargetId-0="figure-42@141.[386,1200,193,1237]" captionTargetId-1="figure-55@142.[420,1167,193,1123]" captionTargetPageId-0="141" captionTargetPageId-1="142" captionText-0="FIGURES 288–291. Aptilotella quadrata Luk &amp; Marshall, sp. n.: 288—male terminalia, posterior; 289—male terminalia, left lateral; 290—male synsternite 6+7; 291—male sternite 5." captionText-1="FIGURES 292–294. Aptilotella quadrata Luk &amp; Marshall, sp. n.: 292—aedeagus with postgonites removed; 293—left postgonite; 294—hypandrium. Abbreviations: BP = basiphallus, DS = dorsal sclerite, LFS = lateral flanking sclerite, VFS = ventral flanking sclerite." figureDoi-0="http://doi.org/10.5281/zenodo.4909298" figureDoi-1="http://doi.org/10.5281/zenodo.4909300" httpUri-0="https://zenodo.org/record/4909298/files/figure.png" httpUri-1="https://zenodo.org/record/4909300/files/figure.png" pageId="39" pageNumber="40">288–294</figureCitation>
</paragraph>
<paragraph id="8B9236E7FF82FF94FDC7FBC2FF2C0AAA" blockId="39.[151,1436,1513,1970]" pageId="39" pageNumber="40">
<emphasis id="B959EAF5FF82FF94FDC7FBC2FEE40B1A" bold="true" box="[151,291,1513,1538]" italics="true" pageId="39" pageNumber="40">Description.</emphasis>
Habitus as in
<figureCitation id="13162A62FF82FF94FC94FBC2FDE30B1A" box="[452,548,1513,1538]" captionStart="FIGURE 3" captionStartId="40.[151,249,884,906]" captionTargetId="figure-472@40.[369,1218,195,862]" captionTargetPageId="40" captionText="FIGURE 3. Aptilotella quadrata Luk &amp; Marshall, sp. n., holotype male." figureDoi="http://doi.org/10.5281/zenodo.5042241" httpUri="https://zenodo.org/record/5042241/files/figure.png" pageId="39" pageNumber="40">Figure 3</figureCitation>
. Body length
<quantity id="4CD59B02FF82FF94FF90FBC2FC8D0B19" box="[704,842,1513,1538]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" pageId="39" pageNumber="40" unit="mm" value="1.35" valueMax="1.6" valueMin="1.1">1.1–1.6 mm</quantity>
. Head ground color yellow. Frons finely rugose; pale areas attaining the occiput but not the anterior margin; brown medial stripes each about one-seventh the width of frons, with narrow orange borders; dark brown orbital stripes confined to posterior corners of frons, their orange borders confluent with the medial stripes; ocular emargination with pruinose patch. Ocellar tubercle scarcely raised; ocellar bristle two-thirds the length of frons. Orbital bristle absent; orbital setulae inconspicuous, in two pairs. Interfrontal setae inconspicuous, in three pairs. Face shining; facial tubercle bulbous; gena with black ventral margin. Antenna brown, first flagellomere paler in basal half. Occiput with dark brown margin. Scutum and scutellum black, shining. Scutum with only a few short setae. Scutellum bare; 2.4 times wider than long, 0.6 times the width of scutum. Apical scutellar bristles 2.5 times as long as basal. Pleuron black; anepisternum and anepimeron with pale pruinose stripe. Legs orange; coxa black; femur dark orange, basal half of hind femur brown; mid tibia dark brown, with a single proximal anterodorsal bristle. Wing rudiment dark brown. Abdomen black, shining; tergites distally with a row of evenly-spaced, short setae; sternites microtrichose. Cercus and surstylus orange.
</paragraph>
<caption id="DF52666FFF8DFF9BFDC7FD5FFCBE0E92" ID-DOI="http://doi.org/10.5281/zenodo.5042241" ID-Zenodo-Dep="5042241" box="[151,889,884,906]" httpUri="https://zenodo.org/record/5042241/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,249,884,906]" targetBox="[375,1210,209,856]" targetPageId="40">
<paragraph id="8B9236E7FF8DFF9BFDC7FD5FFCBE0E92" blockId="40.[151,889,884,906]" box="[151,889,884,906]" pageId="40" pageNumber="41">
<emphasis id="B959EAF5FF8DFF9BFDC7FD5FFED50E91" bold="true" box="[151,274,884,906]" pageId="40" pageNumber="41">FIGURE 3.</emphasis>
<taxonomicName id="4C2D4D64FF8DFF9BFC49FD5FFD540E92" authority="Luk &amp; Marshall" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[281,659,884,906]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="quadrata" status="sp. nov.">
<emphasis id="B959EAF5FF8DFF9BFC49FD5FFE200E92" box="[281,487,884,906]" italics="true" pageId="40" pageNumber="41">Aptilotella quadrata</emphasis>
Luk &amp; Marshall
</taxonomicName>
,
<emphasis id="B959EAF5FF8DFF9BFFCDFD5FFD110E91" bold="true" box="[669,726,884,905]" pageId="40" pageNumber="41">
<taxonomicNameLabel id="A26A578EFF8DFF9BFFCDFD5FFD110E91" box="[669,726,884,905]" pageId="40" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype male.
</paragraph>
</caption>
<paragraph id="8B9236E7FF8DFF9BFD97FD9DFC420B2B" blockId="40.[151,1436,950,2020]" pageId="40" pageNumber="41">
<emphasis id="B959EAF5FF8DFF9BFD97FD9DFE400ED7" bold="true" box="[199,391,950,975]" italics="true" pageId="40" pageNumber="41">Male terminalia.</emphasis>
Sternite 5 (
<figureCitation id="13162A62FF8DFF9BFF57FD9DFDA10ED7" box="[519,614,950,975]" captionStart="FIGURES 288–291" captionStartId="141.[151,264,1258,1281]" captionTargetBox="[394,1191,201,1229]" captionTargetId="figure-42@141.[386,1200,193,1237]" captionTargetPageId="141" captionText="FIGURES 288–291. Aptilotella quadrata Luk &amp; Marshall, sp. n.: 288—male terminalia, posterior; 289—male terminalia, left lateral; 290—male synsternite 6+7; 291—male sternite 5." figureDoi="http://doi.org/10.5281/zenodo.4909298" httpUri="https://zenodo.org/record/4909298/files/figure.png" pageId="40" pageNumber="41">Fig. 291</figureCitation>
) narrow and asymmetrical; posteromedial half shallowly emarginate; two long, apically rounded lobes arising in posteromedial quarter, their interior margin setaeous. Synsternite 6+7 (
<figureCitation id="13162A62FF8DFF9BF83EFDF1FF04090F" captionStart="FIGURES 288–291" captionStartId="141.[151,264,1258,1281]" captionTargetBox="[394,1191,201,1229]" captionTargetId="figure-42@141.[386,1200,193,1237]" captionTargetPageId="141" captionText="FIGURES 288–291. Aptilotella quadrata Luk &amp; Marshall, sp. n.: 288—male terminalia, posterior; 289—male terminalia, left lateral; 290—male synsternite 6+7; 291—male sternite 5." figureDoi="http://doi.org/10.5281/zenodo.4909298" httpUri="https://zenodo.org/record/4909298/files/figure.png" pageId="40" pageNumber="41">Fig. 290</figureCitation>
) with humped margin before medial bridge; interior surface of medial bridge with free, saddle-shaped sclerite. Cercus (
<figureCitation id="13162A62FF8DFF9BFDA8FA09FE640923" box="[248,419,1058,1083]" captionStart="FIGURES 288–291" captionStartId="141.[151,264,1258,1281]" captionTargetBox="[394,1191,201,1229]" captionTargetId="figure-42@141.[386,1200,193,1237]" captionTargetPageId="141" captionText="FIGURES 288–291. Aptilotella quadrata Luk &amp; Marshall, sp. n.: 288—male terminalia, posterior; 289—male terminalia, left lateral; 290—male synsternite 6+7; 291—male sternite 5." figureDoi="http://doi.org/10.5281/zenodo.4909298" httpUri="https://zenodo.org/record/4909298/files/figure.png" pageId="40" pageNumber="41">Figs. 288, 289</figureCitation>
) curved, twice as long as basal width; base triangular, with four long setae near outer margin, and a much longer marginal seta subequal to the length of cercus; distal three-fifths slender, bearing two small, well-separated sensory setae. Surstylus (
<figureCitation id="13162A62FF8DFF9BFFE2FA41FC9D099B" box="[690,858,1130,1155]" captionStart="FIGURES 288–291" captionStartId="141.[151,264,1258,1281]" captionTargetBox="[394,1191,201,1229]" captionTargetId="figure-42@141.[386,1200,193,1237]" captionTargetPageId="141" captionText="FIGURES 288–291. Aptilotella quadrata Luk &amp; Marshall, sp. n.: 288—male terminalia, posterior; 289—male terminalia, left lateral; 290—male synsternite 6+7; 291—male sternite 5." figureDoi="http://doi.org/10.5281/zenodo.4909298" httpUri="https://zenodo.org/record/4909298/files/figure.png" pageId="40" pageNumber="41">Figs. 288, 289</figureCitation>
) horseshoe-shaped, basally enlarged; anterior lobe apically dilated and truncate, with scattered sensory setae; posterior lobe with setaceous outer margin, apically bearing three large, articulated, claw-like spines. Postgonite (
<figureCitation id="13162A62FF8DFF9BFE19FA99FC6E09D3" box="[841,937,1202,1227]" captionStart="FIGURES 292–294" captionStartId="142.[151,264,1143,1166]" captionTargetBox="[426,1161,196,1118]" captionTargetId="figure-55@142.[420,1167,193,1123]" captionTargetPageId="142" captionText="FIGURES 292–294. Aptilotella quadrata Luk &amp; Marshall, sp. n.: 292—aedeagus with postgonites removed; 293—left postgonite; 294—hypandrium. Abbreviations: BP = basiphallus, DS = dorsal sclerite, LFS = lateral flanking sclerite, VFS = ventral flanking sclerite." figureDoi="http://doi.org/10.5281/zenodo.4909300" httpUri="https://zenodo.org/record/4909300/files/figure.png" pageId="40" pageNumber="41">Fig. 293</figureCitation>
) sinuate; descending arm 0.6 times the total length, of uniform width and slightly bent midway, with two distal sensory setae; articulatory process for pregonite broad, rounded; articulatory process for basiphallus knobbed. Hypandrium (
<figureCitation id="13162A62FF8DFF9BFEBEFAD1FB89080B" box="[1006,1102,1274,1299]" captionStart="FIGURES 292–294" captionStartId="142.[151,264,1143,1166]" captionTargetBox="[426,1161,196,1118]" captionTargetId="figure-55@142.[420,1167,193,1123]" captionTargetPageId="142" captionText="FIGURES 292–294. Aptilotella quadrata Luk &amp; Marshall, sp. n.: 292—aedeagus with postgonites removed; 293—left postgonite; 294—hypandrium. Abbreviations: BP = basiphallus, DS = dorsal sclerite, LFS = lateral flanking sclerite, VFS = ventral flanking sclerite." figureDoi="http://doi.org/10.5281/zenodo.4909300" httpUri="https://zenodo.org/record/4909300/files/figure.png" pageId="40" pageNumber="41">Fig. 294</figureCitation>
) with medial rod straight and apically truncate; hypandrial arms apparently fused to broad base, very slender, with the apex truncate and bent back; pregonite minute and squared. Aedeagus as in
<figureCitation id="13162A62FF8DFF9BFE5FFB69FC560843" box="[783,913,1346,1371]" captionStart="FIGURES 292–294" captionStartId="142.[151,264,1143,1166]" captionTargetBox="[426,1161,196,1118]" captionTargetId="figure-55@142.[420,1167,193,1123]" captionTargetPageId="142" captionText="FIGURES 292–294. Aptilotella quadrata Luk &amp; Marshall, sp. n.: 292—aedeagus with postgonites removed; 293—left postgonite; 294—hypandrium. Abbreviations: BP = basiphallus, DS = dorsal sclerite, LFS = lateral flanking sclerite, VFS = ventral flanking sclerite." figureDoi="http://doi.org/10.5281/zenodo.4909300" httpUri="https://zenodo.org/record/4909300/files/figure.png" pageId="40" pageNumber="41">Figure 292</figureCitation>
. Basiphallus compressed, arched, basal lobe projecting into distiphallus; articulatory process for postgonite pointed, erect. Ejaculatory apodeme short. Lateral flanking sclerites with dark dorsal and ventral margins, broadly fused both ventro- and dorsobasally; apicodorsally armed with minute sclerotized teeth. Dorsal sclerite constricted medially, its sides forming a channel; apex laterally lobed and bearing minute sclerotized teeth. Ventral flanking sclerites with the basal article fused along ventrodistal margin of lateral flanking sclerite; the spatulate medial article sharply pointed beneath the basal article; the concave distal article fused to the medial article and sharply pointed over it.
</paragraph>
<paragraph id="8B9236E7FF8DFF9BFD97F815FB570B4F" blockId="40.[151,1436,950,2020]" box="[199,1168,1598,1623]" pageId="40" pageNumber="41">
<emphasis id="B959EAF5FF8DFF9BFD97F815FE630B4F" bold="true" box="[199,420,1598,1623]" italics="true" pageId="40" pageNumber="41">Female terminalia.</emphasis>
Not examined; retracted from view in the single female specimen.
</paragraph>
<paragraph id="8B9236E7FF8DFF9BFD97F849FE450BFE" blockId="40.[151,1436,950,2020]" pageId="40" pageNumber="41">
<emphasis id="B959EAF5FF8DFF9BFD97F849FEFE0B63" bold="true" box="[199,313,1634,1659]" pageId="40" pageNumber="41">
<emphasis id="B959EAF5FF8DFF9BFD97F849FEF50B63" bold="true" box="[199,306,1634,1659]" italics="true" pageId="40" pageNumber="41">Variation</emphasis>
.
</emphasis>
The
<specimenCount id="9D2BFD6EFF8DFF9BFC24F848FDDE0B63" box="[372,537,1634,1659]" pageId="40" pageNumber="41" type="generic">two specimens</specimenCount>
from the “trail to Cerro Chirripó” have the dark genal band continuing through the clypeus, a thick dark band across the facial excavation, dark brown legs, and two anterodorsal bristles on the mid tibia. Although otherwise identical to the
<typeStatus id="54968845FF8DFF9BFFCBF880FD0C0BDB" box="[667,715,1707,1731]" pageId="40" pageNumber="41">type</typeStatus>
material, both are females and cannot be positively identified as
<taxonomicName id="4C2D4D64FF8DFF9BFDC7F8E4FEBD0BFF" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[151,378,1743,1767]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="quadrata">
<emphasis id="B959EAF5FF8DFF9BFDC7F8E4FEBD0BFF" box="[151,378,1743,1767]" italics="true" pageId="40" pageNumber="41">Aptilotella quadrata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C337656CFF8DFF9BFD97F8D9FE7E0A37" pageId="40" pageNumber="41" type="etymology">
<paragraph id="8B9236E7FF8DFF9BFD97F8D9FE7E0A37" blockId="40.[151,1436,950,2020]" pageId="40" pageNumber="41">
<emphasis id="B959EAF5FF8DFF9BFD97F8D9FE800A13" bold="true" box="[199,327,1778,1803]" italics="true" pageId="40" pageNumber="41">Etymology.</emphasis>
The species epithet describes the roughly squared tergite 1 plate, a defining character of this species and sister species.
</paragraph>
</subSubSection>
<subSubSection id="C337656CFF8DFF9AFD97F910FDF90DCD" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="8B9236E7FF8DFF9BFD97F910FDA00A6E" blockId="40.[151,1436,950,2020]" pageId="40" pageNumber="41">
<emphasis id="B959EAF5FF8DFF9BFD97F910FEA80A4B" bold="true" box="[199,367,1850,1875]" italics="true" pageId="40" pageNumber="41">Type material.</emphasis>
<materialsCitation id="3B453CBAFF8DFF9BFC29F911FDA00A6E" ID-GBIF-Occurrence="3311767380" collectingDate="1985-04-07" collectingDateMax="1985-04-13" collectingDateMin="1985-04-07" collectionCode="INBC" collectorName="H. Goulet &amp; L. Masner." country="Costa Rica" elevation="3200" location="Highway" pageId="40" pageNumber="41" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54968845FF8DFF9BFC29F911FE260A4B" box="[377,481,1850,1875]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
<specimenCount id="9D2BFD6EFF8DFF9BFCBCF910FDC40A4A" box="[492,515,1851,1874]" pageId="40" pageNumber="41" type="male">♂</specimenCount>
,
<collectionCode id="ED3CAE22FF8DFF9BFF42F911FD9F0A4B" box="[530,600,1850,1875]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="40" pageNumber="41">INBC</collectionCode>
.
<collectingCountry id="F33A7677FF8DFF9BFF35F911FCD70A4B" box="[613,784,1850,1875]" name="Costa Rica" pageId="40" pageNumber="41">COSTA RICA</collectingCountry>
:
<location id="8EF2603CFF8DFF9BFE4EF911FC430A4B" LSID="urn:lsid:plazi:treatment:038487F1FF82FF9AFDC7FB55FF1F0F00:8EF2603CFF8DFF9BFE4EF911FC430A4B" box="[798,900,1850,1875]" country="Costa Rica" name="Highway" pageId="40" pageNumber="41">Highway</location>
2, km 96,
<quantity id="4CD59B02FF8DFF9BF958F911FBA50A4A" box="[1032,1122,1850,1875]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="40" pageNumber="41" unit="m" value="3200.0">
<elevation id="0000D1D4FF8DFF9BF958F911FBA50A4A" box="[1032,1122,1850,1875]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="40" pageNumber="41" unit="m" value="3200.0">3200 m</elevation>
</quantity>
,
<date id="FF931027FF8DFF9BF921F911FAC10A4B" box="[1137,1286,1850,1875]" pageId="40" pageNumber="41" value="1985-04-07" valueMax="1985-04-13" valueMin="1985-04-07">
<collectingDate id="EFD7E9CFFF8DFF9BF921F911FAC10A4B" box="[1137,1286,1850,1875]" pageId="40" pageNumber="41" value="1985-04-07" valueMax="1985-04-13" valueMin="1985-04-07">7–13.iv.1985</collectingDate>
</date>
, cloud forest (oak),
<collectingMethod id="526C4EF0FF8DFF9BFDB0F974FEFA0A6E" box="[224,317,1887,1910]" pageId="40" pageNumber="41">pan trap</collectingMethod>
,
<collectorName id="26D85331FF8DFF9BFC19F974FE700A6F" box="[329,439,1886,1911]" pageId="40" pageNumber="41">H. Goulet</collectorName>
and
<collectorName id="26D85331FF8DFF9BFCBEF974FDA00A6E" box="[494,615,1887,1910]" pageId="40" pageNumber="41">L. Masner.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8B9236E7FF8DFF9BFD97F9A8FADB0AFC" blockId="40.[151,1436,950,2020]" pageId="40" pageNumber="41">
<materialsCitation id="3B453CBAFF8DFF9BFD97F9A8FD1C0AA7" ID-GBIF-Occurrence="3311767430" collectingDate="1997-06-26" collectionCode="DEBU" collectorName="R. S. Anderson" country="Costa Rica" elevation="2800" latitude="9.558333" location="Villa Mills" longLatPrecision="912" longitude="-83.7" pageId="40" pageNumber="41" specimenCount="1" specimenCount-female="1" stateProvince="San Jose" typeStatus="paratype">
<emphasis id="B959EAF5FF8DFF9BFD97F9A8FEF80A82" bold="true" box="[199,319,1923,1946]" italics="true" pageId="40" pageNumber="41">
<typeStatus id="54968845FF8DFF9BFD97F9A8FEFC0A82" box="[199,315,1923,1946]" pageId="40" pageNumber="41" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="F33A7677FF8DFF9BFC19F9A9FE330A83" box="[329,500,1922,1947]" name="Costa Rica" pageId="40" pageNumber="41">COSTA RICA</collectingCountry>
:
<collectingRegion id="49E9F805FF8DFF9BFF50F9A9FDA10A83" box="[512,614,1922,1947]" country="Costa Rica" name="San Jose" pageId="40" pageNumber="41">San José</collectingRegion>
,
<quantity id="4CD59B02FF8DFF9BFF24F9A9FD680A83" box="[628,687,1922,1947]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="40" pageNumber="41" unit="km" value="2.0">2 km</quantity>
E
<location id="8EF2603CFF8DFF9BFF83F9A9FC960A83" LSID="urn:lsid:plazi:treatment:038487F1FF82FF9AFDC7FB55FF1F0F00:8EF2603CFF8DFF9BFF83F9A9FC960A83" box="[723,849,1922,1947]" country="Costa Rica" latitude="9.558333" longLatPrecision="912" longitude="-83.7" name="Villa Mills" pageId="40" pageNumber="41" stateProvince="San Jose">Villa Mills</location>
,
<geoCoordinate id="EE195020FF8DFF9BFE0EF9A9FC1C0A83" box="[862,987,1922,1947]" degrees="9" direction="north" minutes="33" orientation="latitude" pageId="40" pageNumber="41" precision="15" seconds="30" value="9.558333">9°33’30”N</geoCoordinate>
,
<geoCoordinate id="EE195020FF8DFF9BFEBAF9A9FB960A83" box="[1002,1105,1922,1947]" degrees="83" direction="west" minutes="42" orientation="longitude" pageId="40" pageNumber="41" precision="925" value="-83.7">83°42’W</geoCoordinate>
,
<quantity id="4CD59B02FF8DFF9BF90FF9A9FB7F0A82" box="[1119,1208,1922,1947]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="40" pageNumber="41" unit="m" value="2800.0">
<elevation id="0000D1D4FF8DFF9BF90FF9A9FB7F0A82" box="[1119,1208,1922,1947]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="40" pageNumber="41" unit="m" value="2800.0">2800 m</elevation>
</quantity>
,
<date id="FF931027FF8DFF9BF996F9A9FA870A83" box="[1222,1344,1922,1947]" pageId="40" pageNumber="41" value="1997-06-26">
<collectingDate id="EFD7E9CFFF8DFF9BF996F9A9FA870A83" box="[1222,1344,1922,1947]" pageId="40" pageNumber="41" value="1997-06-26">26.vi.1997</collectingDate>
</date>
, second growth oak forest litter,
<collectorName id="26D85331FF8DFF9BFCF8F98DFD8A0AA7" box="[424,589,1958,1983]" pageId="40" pageNumber="41">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFF8DFF9BFF30F98DFDB30AA7" box="[608,628,1958,1983]" pageId="40" pageNumber="41" type="female">♀</specimenCount>
,
<collectionCode id="ED3CAE22FF8DFF9BFFD1F98DFD150AA7" box="[641,722,1958,1983]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="40" pageNumber="41">DEBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B453CBAFF8DFF9BFFB6F98CFA5F0AA7" ID-GBIF-Occurrence="3311767314" box="[742,1432,1958,1983]" collectingDate="1998-02-15" collectionCode="DEBU" country="Costa Rica" elevation="2750" pageId="40" pageNumber="41" specimenCount="1" specimenCount-male="1" stateProvince="San Jose" typeStatus="paratype">
same as previous label but at
<quantity id="4CD59B02FF8DFF9BF964F98DFB4B0AA6" box="[1076,1164,1958,1983]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="40" pageNumber="41" unit="m" value="2750.0">
<elevation id="0000D1D4FF8DFF9BF964F98DFB4B0AA6" box="[1076,1164,1958,1983]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="40" pageNumber="41" unit="m" value="2750.0">2750 m</elevation>
</quantity>
,
<date id="FF931027FF8DFF9BF9C8F98DFACD0AA7" box="[1176,1290,1958,1983]" pageId="40" pageNumber="41" value="1998-02-15">
<collectingDate id="EFD7E9CFFF8DFF9BF9C8F98DFACD0AA7" box="[1176,1290,1958,1983]" pageId="40" pageNumber="41" value="1998-02-15">15.ii.1998</collectingDate>
</date>
(
<specimenCount id="9D2BFD6EFF8DFF9BF84BF98DFAF50AA7" box="[1307,1330,1958,1983]" pageId="40" pageNumber="41" type="male">♂</specimenCount>
,
<collectionCode id="ED3CAE22FF8DFF9BF86FF98DFA570AA7" box="[1343,1424,1958,1983]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="40" pageNumber="41">DEBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B453CBAFF8DFF9BFDC7F9E0FADF0AFC" ID-GBIF-Occurrence="3311767354" box="[151,1304,1995,2020]" collectingDate="1999-06-25" collectionCode="DEBU" collectorName="R. S. Anderson" country="Costa Rica" elevation="3350" location="Base Crestones" municipality="Cerro Chirripo" pageId="40" pageNumber="41" specimenCount="1" specimenCount-male="1" stateProvince="San Jose" typeStatus="paratype">
<collectingMunicipality id="6BF6AC9DFF8DFF9BFDC7F9E0FEF80AFC" box="[151,319,1995,2020]" pageId="40" pageNumber="41">Cerro Chirripó</collectingMunicipality>
,
<location id="8EF2603CFF8DFF9BFC1BF9E0FE3E0AFC" LSID="urn:lsid:plazi:treatment:038487F1FF82FF9AFDC7FB55FF1F0F00:8EF2603CFF8DFF9BFC1BF9E0FE3E0AFC" box="[331,505,1995,2020]" country="Costa Rica" municipality="Cerro Chirripo" name="Base Crestones" pageId="40" pageNumber="41" stateProvince="San Jose">Base Crestones</location>
,
<quantity id="4CD59B02FF8DFF9BFF54F9E0FD9C0AFA" box="[516,603,1995,2020]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="40" pageNumber="41" unit="m" value="3350.0">
<elevation id="0000D1D4FF8DFF9BFF54F9E0FD9C0AFA" box="[516,603,1995,2020]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="40" pageNumber="41" unit="m" value="3350.0">3350 m</elevation>
</quantity>
,
<date id="FF931027FF8DFF9BFF37F9E0FD260AFC" box="[615,737,1995,2020]" pageId="40" pageNumber="41" value="1999-06-25">
<collectingDate id="EFD7E9CFFF8DFF9BFF37F9E0FD260AFC" box="[615,737,1995,2020]" pageId="40" pageNumber="41" value="1999-06-25">25.vi.1999</collectingDate>
</date>
, elfin forest, leaf litter,
<collectorName id="26D85331FF8DFF9BFEB8F9E0FB4B0AFC" box="[1000,1164,1995,2020]" pageId="40" pageNumber="41">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFF8DFF9BF9CCF9E0FB740AFC" box="[1180,1203,1995,2020]" pageId="40" pageNumber="41" type="male">♂</specimenCount>
,
<collectionCode id="ED3CAE22FF8DFF9BF9EFF9E0FAD70AFC" box="[1215,1296,1995,2020]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="40" pageNumber="41">DEBU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9236E7FF8CFF9AFD97FEBCFDF90DCD" blockId="41.[151,1436,151,536]" pageId="41" pageNumber="42">
<emphasis id="B959EAF5FF8CFF9AFD97FEBCFE330DA8" bold="true" box="[199,500,151,176]" italics="true" pageId="41" pageNumber="42">Other material examined.</emphasis>
<materialsCitation id="3B453CBAFF8CFF9AFCAEFEBCFDFD0DCD" ID-GBIF-Occurrence="3311767379" collectingDate="1999-06-27" collectorName="R. S. Anderson" country="Costa Rica" elevation="2800" location="Cerro Chirripo" pageId="41" pageNumber="42" specimenCount="2" specimenCount-female="2" stateProvince="Cartago">
<collectingCountry id="F33A7677FF8CFF9AFCAEFEBCFD6E0DA8" box="[510,681,151,176]" name="Costa Rica" pageId="41" pageNumber="42">COSTA RICA</collectingCountry>
:
<collectingRegion id="49E9F805FF8CFF9AFFE5FEBCFCD70DA8" box="[693,784,151,176]" country="Costa Rica" name="Cartago" pageId="41" pageNumber="42">Cartago</collectingRegion>
,
<location id="8EF2603CFF8CFF9AFE4EFEBCFC0E0DA8" LSID="urn:lsid:plazi:treatment:038487F1FF82FF9AFDC7FB55FF1F0F00:8EF2603CFF8CFF9AFE4EFEBCFC0E0DA8" box="[798,969,151,176]" country="Costa Rica" name="Cerro Chirripo" pageId="41" pageNumber="42" stateProvince="Cartago">Cerro Chirripó</location>
, trail to,
<quantity id="4CD59B02FF8CFF9AF964FEBCFB4A0DB7" box="[1076,1165,151,176]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="41" pageNumber="42" unit="m" value="2800.0">
<elevation id="0000D1D4FF8CFF9AF964FEBCFB4A0DB7" box="[1076,1165,151,176]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="41" pageNumber="42" unit="m" value="2800.0">2800 m</elevation>
</quantity>
,
<date id="FF931027FF8CFF9AF9CCFEBCFAD10DA8" box="[1180,1302,151,176]" pageId="41" pageNumber="42" value="1999-06-27">
<collectingDate id="EFD7E9CFFF8CFF9AF9CCFEBCFAD10DA8" box="[1180,1302,151,176]" pageId="41" pageNumber="42" value="1999-06-27">27.vi.1999</collectingDate>
</date>
, mixed oak forest, leaf litter,
<collectorName id="26D85331FF8CFF9AFC07FE97FE3C0DCD" box="[343,507,188,213]" pageId="41" pageNumber="42">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFF8CFF9AFF5DFE97FDF70DCD" box="[525,560,188,213]" pageId="41" pageNumber="42" type="female">2♀</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C337656CFF8CFF9AFD97FEF4FF1F0F00" pageId="41" pageNumber="42" type="discussion">
<paragraph id="8B9236E7FF8CFF9AFD97FEF4FBC40C90" blockId="41.[151,1436,151,536]" pageId="41" pageNumber="42">
<emphasis id="B959EAF5FF8CFF9AFD97FEF4FE8F0DE0" bold="true" box="[199,328,223,248]" pageId="41" pageNumber="42">
<emphasis id="B959EAF5FF8CFF9AFD97FEF4FE850DE0" bold="true" box="[199,322,223,248]" italics="true" pageId="41" pageNumber="42">Comments</emphasis>
.
</emphasis>
Externally,
<taxonomicName id="4C2D4D64FF8CFF9AFC85FECBFD7B0DE0" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[469,700,224,248]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="quadrata">
<emphasis id="B959EAF5FF8CFF9AFC85FECBFD7B0DE0" box="[469,700,224,248]" italics="true" pageId="41" pageNumber="42">Aptilotella quadrata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2D4D64FF8CFF9AFFA8FECBFC590DE0" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[760,926,224,248]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="umbracatus">
<emphasis id="B959EAF5FF8CFF9AFFA8FECBFC590DE0" box="[760,926,224,248]" italics="true" pageId="41" pageNumber="42">A. umbracatus</emphasis>
</taxonomicName>
are recognizable by their very shiny, nearly hairless body, completely separated lateral remnant of tergite 1, and unicolorous legs with one or two long anterodorsal bristles on the mid tibia. They are also distinguished from all other
<taxonomicName id="4C2D4D64FF8CFF9AF949FF03FB480C58" authorityName="Duda" authorityYear="1924" box="[1049,1167,296,320]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAF5FF8CFF9AF949FF03FB480C58" box="[1049,1167,296,320]" italics="true" pageId="41" pageNumber="42">Aptilotella</emphasis>
</taxonomicName>
by the distiphallus with a prominent dorsal sclerite and articulated medial and distal articles of the spoon-shaped ventral flanking sclerites, the loss of the ventrobasal sclerite, and a spindle-shaped ejaculatory apodeme.
</paragraph>
<paragraph id="8B9236E7FF8CFF9AFD97FFBFFF1F0F00" blockId="41.[151,1436,151,536]" pageId="41" pageNumber="42">
Few members of the genus are as large and robust as
<taxonomicName id="4C2D4D64FF8CFF9AFE7DFFBFFBD40CB4" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[813,1043,404,428]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="quadrata">
<emphasis id="B959EAF5FF8CFF9AFE7DFFBFFBD40CB4" box="[813,1043,404,428]" italics="true" pageId="41" pageNumber="42">Aptilotella quadrata</emphasis>
</taxonomicName>
, and not even the more slender
<taxonomicName id="4C2D4D64FF8CFF9AF8D4FFBFFEC00CC8" authorityName="Luk &amp; Marshall" authorityYear="2014" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="involucris">
<emphasis id="B959EAF5FF8CFF9AF8D4FFBFFEC00CC8" italics="true" pageId="41" pageNumber="42">A. involucris</emphasis>
</taxonomicName>
possesses such a prominently bilobed male sternite. A female specimen of an undescribed species, collected at carrion with
<taxonomicName id="4C2D4D64FF8CFF9AFCFBFFF7FD890CEC" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[427,590,476,500]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="umbracatus">
<emphasis id="B959EAF5FF8CFF9AFCFBFFF7FD890CEC" box="[427,590,476,500]" italics="true" pageId="41" pageNumber="42">A. umbracatus</emphasis>
</taxonomicName>
in
<collectingCountry id="F33A7677FF8CFF9AFF20FFF7FD0B0CEB" box="[624,716,476,499]" name="Panama" pageId="41" pageNumber="42">Panama</collectingCountry>
, is superficially similar to
<taxonomicName id="4C2D4D64FF8CFF9AFEA3FFF7FBBF0CEC" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[1011,1144,476,500]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="quadrata">
<emphasis id="B959EAF5FF8CFF9AFEA3FFF7FBBF0CEC" box="[1011,1144,476,500]" italics="true" pageId="41" pageNumber="42">A. quadrata</emphasis>
</taxonomicName>
but lacks markings on the frons.
</paragraph>
</subSubSection>
</treatment>
</document>
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