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<document id="CF336FB988F6750A047BDC655A359E49" ID-DOI="10.5281/zenodo.276438" ID-GBIF-Dataset="9799d3e6-c10a-4bd4-ad09-99369a0ace8b" ID-ISSN="1175-5326" ID-Zenodo-Dep="276438" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460326812181" checkinUser="plazi" docAuthor="Murao, Ryuki, Tadauchi, Osamu, Goubara, Masashi &amp; Maeta, Yasuo" docDate="2010" docId="03846B10FFBCFFC8FC8CDDD7E1E5FA6B" docLanguage="en" docName="zt02669p033.pdf" docOrigin="Zootaxa 2669" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Lasioglossum (Evylaeus) bicornutum Murao, sp. nov." docType="treatment" docVersion="8" lastPageNumber="9" masterDocId="FFBD1368FFBAFFC0FC1BDF75E468FFA7" masterDocTitle="Taxonomy of the carinaless group of Lasioglossum (Evylaeus) (Hymenoptera, Halictidae) from the Ryukyu Islands, Japan, with description of seven new species" masterLastPageNumber="33" masterPageNumber="1" pageNumber="7" updateTime="1698257709375" updateUser="plazi">
<mods:mods id="EB5B7858E0DE529C17A3654A58EA3040" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CCD79CF42B32CB91D5B1EE733D722CAD">
<mods:title id="6B38BC1FC628F3BB2500CB205D15992C">Taxonomy of the carinaless group of Lasioglossum (Evylaeus) (Hymenoptera, Halictidae) from the Ryukyu Islands, Japan, with description of seven new species</mods:title>
</mods:titleInfo>
<mods:name id="8F2A306C3C5E19A7452B7017830F2E71" type="personal">
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<mods:namePart id="C208AF5F1C80AF6DEC4A00CBB4B5E931">Murao, Ryuki</mods:namePart>
</mods:name>
<mods:name id="24BB1D40426106295F122D9AAB85C004" type="personal">
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</mods:role>
<mods:namePart id="16AEA2842F14D019898F0A307585BDA0">Tadauchi, Osamu</mods:namePart>
</mods:name>
<mods:name id="D858EC571C06405124EB91284BC15761" type="personal">
<mods:role id="B1DA0346225020C4F0FB9C51B4533031">
<mods:roleTerm id="D6C930C5488FC75474D9F200F5557582">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="427D3E24443D842CA69ED705CEEC5819">Goubara, Masashi</mods:namePart>
</mods:name>
<mods:name id="61CA9B02C700705AD081E14459BB292C" type="personal">
<mods:role id="A5DF0E81E16CC5048A292432D1CD3EBA">
<mods:roleTerm id="E1A5C2896D1BF7E82322E9AC3485C3C7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="643550BC03490BFBC0AE45D41F5DAF8B">Maeta, Yasuo</mods:namePart>
</mods:name>
<mods:typeOfResource id="BC4418D9C058BF56BE6D38E78C80BCCF">text</mods:typeOfResource>
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<mods:title id="67E48282B0E5AEE813D302E84D625576">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0F88E3C4A4198E948ECD92B30C7D77FE">
<mods:date id="96427EA79DE8DAA3EEBA2299AE75C587">2010</mods:date>
<mods:detail id="25B888FEE265E57ABBD0C2EF575D090F" type="volume">
<mods:number id="83BB5A38E888E97ECB7FC8496DA3242C">2669</mods:number>
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<mods:start id="C179348935CF7CE5F45BC36589B70E41">1</mods:start>
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<mods:classification id="29E912FC52D95499F7F47EA335F43615">journal article</mods:classification>
<mods:identifier id="27AF5FA33F0987843064DFD1E97B774D" type="DOI">10.5281/zenodo.276438</mods:identifier>
<mods:identifier id="E9B06423715A3308DACB0E5493E39C24" type="GBIF-Dataset">9799d3e6-c10a-4bd4-ad09-99369a0ace8b</mods:identifier>
<mods:identifier id="58F271A6E7B40866E34627F8A47C10A1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EF76C31BD91DC586C8C4BF41BD5AA5F0" type="Zenodo-Dep">276438</mods:identifier>
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<treatment id="03846B10FFBCFFC8FC8CDDD7E1E5FA6B" ID-DOI="http://doi.org/10.5281/zenodo.6210207" ID-GBIF-Taxon="119403864" ID-Zenodo-Dep="6210207" LSID="urn:lsid:plazi:treatment:03846B10FFBCFFC8FC8CDDD7E1E5FA6B" httpUri="http://treatment.plazi.org/id/03846B10FFBCFFC8FC8CDDD7E1E5FA6B" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="C337898DFFBCFFC6FC8CDDD7E536FD78" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8B92DA06FFBCFFC6FC8CDDD7E774FD1B" blockId="6.[151,796,674,735]" box="[151,796,674,700]" pageId="6" pageNumber="7">
<heading id="D0DA6D6AFFBCFFC6FC8CDDD7E774FD1B" bold="true" box="[151,796,674,700]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="B9590614FFBCFFC6FC8CDDD7E774FD1B" bold="true" box="[151,796,674,700]" pageId="6" pageNumber="7">
<taxonomicName id="4C2DA185FFBCFFC6FC8CDDD7E6DFFD1B" authority="Murao" authorityName="Murao" box="[151,695,674,700]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bicornutum" status="sp. nov." subGenus="Evylaeus">
<emphasis id="B9590614FFBCFFC6FC8CDDD7E555FD1B" bold="true" box="[151,317,674,700]" italics="true" pageId="6" pageNumber="7">Lasioglossum</emphasis>
(
<emphasis id="B9590614FFBCFFC6FD55DDD7E5D3FD1B" bold="true" box="[334,443,674,700]" italics="true" pageId="6" pageNumber="7">Evylaeus</emphasis>
)
<emphasis id="B9590614FFBCFFC6FDD7DDD7E631FD1B" bold="true" box="[460,601,674,700]" italics="true" pageId="6" pageNumber="7">bicornutum</emphasis>
Murao
</taxonomicName>
<taxonomicNameLabel id="A26ABB6FFFBCFFC6FEA4DDD7E774FD1B" box="[703,796,674,700]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B92DA06FFBCFFC6FC8CDDB0E536FD78" blockId="6.[151,796,674,735]" box="[151,350,709,735]" pageId="6" pageNumber="7">
(
<figureCitation id="1316C683FFBCFFC6FC84DDB0E499FD78" box="[159,241,709,735]" captionStart="FIGURE 3" captionStartId="6.[151,257,1883,1907]" captionTargetBox="[157,1430,888,1844]" captionTargetId="figure@6.[155,1432,863,1862]" captionTargetPageId="6" captionText="FIGURE 3. Lasioglossum (Evylaeus) bicornutum Murao, sp. nov. A – F: holotype, female. G – I: paratype, female. A: lateral habitus. B: lower face. C: mesoscutum. D: sculpture on mesoscutum. E: mesoscutellum. F: mesepisternum. G: labrum. H: propodeal dorsum. I: inner hind tibial spur. Scale bars: H, 0.25 mm; G, I, 0.2 mm." httpUri="https://zenodo.org/record/276441/files/figure.png" pageId="6" pageNumber="7">Figs. 3</figureCitation>
,
<figureCitation id="1316C683FFBCFFC6FCE4DDB0E53DFD78" box="[255,341,709,735]" captionStart="FIGURE 4. A, B" captionStartId="7.[151,257,1680,1704]" captionTargetBox="[200,1382,186,1650]" captionTargetId="figure@7.[187,1399,169,1678]" captionTargetPageId="7" captionText="FIGURE 4. A, B: Lasioglossum (Evylaeus) bicornutum Murao, sp. nov. C, D: Lasioglossum (Evylaeus) silvicolum Murao, sp. nov. E, F: Lasioglossum (Evylaeus) latifacies Murao, sp. nov. A – F: holotypes, females. A, C, E: head in frontal view. B, D, F: first metasomal tergum (left half, sculpture; right half, hair pattern). Scale bar: 1 mm." httpUri="https://zenodo.org/record/276442/files/figure.png" pageId="6" pageNumber="7">4A, 4B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C337898DFFBCFFC8FC8CDC7AE5E6FD30" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph id="8B92DA06FFBCFFC6FC8CDC7AE022FC8D" blockId="6.[151,1098,783,810]" box="[151,1098,783,810]" pageId="6" pageNumber="7">
<emphasis id="B9590614FFBCFFC6FC8CDC7AE5F9FC8E" bold="true" box="[151,401,783,809]" pageId="6" pageNumber="7">Description. Female</emphasis>
. Body length
<quantity id="4CD577E3FFBCFFC6FE2CDC65E6A0FC8D" box="[567,712,784,810]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.6" metricValueMin="5.4" pageId="6" pageNumber="7" unit="mm" value="6.0" valueMax="6.6" valueMin="5.4">5.4–6.6 mm</quantity>
, wing length
<quantity id="4CD577E3FFBCFFC6FF7EDC65E79AFC8D" box="[869,1010,784,810]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.7" metricValueMin="4.8" pageId="6" pageNumber="7" unit="mm" value="5.25" valueMax="5.7" valueMin="4.8">4.8–5.7 mm</quantity>
(n= 5).
</paragraph>
<caption id="DF528A8EFFBCFFC6FC8CD82EE01EF814" httpUri="https://zenodo.org/record/276441/files/figure.png" pageId="6" pageNumber="7" targetBox="[157,1430,888,1844]" targetPageId="6">
<paragraph id="8B92DA06FFBCFFC6FC8CD82EE01EF814" blockId="6.[151,1436,1883,1971]" pageId="6" pageNumber="7">
<emphasis id="B9590614FFBCFFC6FC8CD82EE575F8D4" bold="true" box="[151,285,1883,1907]" pageId="6" pageNumber="7">FIGURE 3.</emphasis>
<taxonomicName id="4C2DA185FFBCFFC6FD3FD829E778F8D4" authority="Murao" authorityName="Murao" box="[292,784,1884,1907]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bicornutum" status="sp. nov." subGenus="Evylaeus">
<emphasis id="B9590614FFBCFFC6FD3FD829E5D2F8D4" box="[292,442,1884,1907]" italics="true" pageId="6" pageNumber="7">Lasioglossum</emphasis>
(
<emphasis id="B9590614FFBCFFC6FDD0D829E646F8D4" box="[459,558,1884,1907]" italics="true" pageId="6" pageNumber="7">Evylaeus</emphasis>
)
<emphasis id="B9590614FFBCFFC6FE25D829E6D4F8D4" box="[574,700,1884,1907]" italics="true" pageId="6" pageNumber="7">bicornutum</emphasis>
Murao
</taxonomicName>
,
<taxonomicNameLabel id="A26ABB6FFFBCFFC6FF00D82EE71BF8D4" box="[795,883,1883,1907]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B9590614FFBCFFC6FF00D82EE71BF8D4" bold="true" box="[795,883,1883,1907]" pageId="6" pageNumber="7">sp. nov.</emphasis>
</taxonomicNameLabel>
A–F: holotype, female. G–I: paratype, female. A: lateral habitus. B: lower face. C: mesoscutum. D: sculpture on mesoscutum. E: mesoscutellum. F: mesepisternum. G: labrum. H: propodeal dorsum. I: inner hind tibial spur. Scale bars: H, 0.25 mm; G, I, 0.2 mm.
</paragraph>
</caption>
<caption id="DF528A8EFFBDFFC7FC8CD9E5E165F940" httpUri="https://zenodo.org/record/276442/files/figure.png" pageId="7" pageNumber="8" targetBox="[200,1382,186,1650]" targetPageId="7">
<paragraph id="8B92DA06FFBDFFC7FC8CD9E5E165F940" blockId="7.[151,1436,1680,1767]" pageId="7" pageNumber="8">
<emphasis id="B9590614FFBDFFC7FC8CD9E5E576F90F" bold="true" box="[151,286,1680,1704]" pageId="7" pageNumber="8">FIGURE 4.</emphasis>
A, B:
<taxonomicName id="4C2DA185FFBDFFC7FD71D9E4E733F900" authority="Murao" authorityName="Murao" box="[362,859,1680,1704]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bicornutum" status="sp. nov." subGenus="Evylaeus">
<emphasis id="B9590614FFBDFFC7FD71D9E4E66AF90F" box="[362,514,1681,1704]" italics="true" pageId="7" pageNumber="8">Lasioglossum</emphasis>
(
<emphasis id="B9590614FFBDFFC7FE08D9E4E61FF90F" box="[531,631,1681,1704]" italics="true" pageId="7" pageNumber="8">Evylaeus</emphasis>
)
<emphasis id="B9590614FFBDFFC7FE93D9E4E76FF90F" box="[648,775,1681,1704]" italics="true" pageId="7" pageNumber="8">bicornutum</emphasis>
Murao
</taxonomicName>
,
<emphasis id="B9590614FFBDFFC7FF7CD9E5E7A9F90F" bold="true" box="[871,961,1680,1704]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A26ABB6FFFBDFFC7FF7CD9E5E7A9F90F" box="[871,961,1680,1704]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
C, D:
<taxonomicName id="4C2DA185FFBDFFC7F817D9E4E48BF960" authority="Murao" authorityName="Murao" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="silvicolum" status="sp. nov." subGenus="Evylaeus">
<emphasis id="B9590614FFBDFFC7F817D9E4E0CCF90F" box="[1036,1188,1681,1704]" italics="true" pageId="7" pageNumber="8">Lasioglossum</emphasis>
(
<emphasis id="B9590614FFBDFFC7F8AED9E4E171F90F" box="[1205,1305,1681,1704]" italics="true" pageId="7" pageNumber="8">Evylaeus</emphasis>
)
<emphasis id="B9590614FFBDFFC7F931D9E4E1F4F90F" box="[1322,1436,1681,1704]" italics="true" pageId="7" pageNumber="8">silvicolum</emphasis>
Murao
</taxonomicName>
,
<taxonomicNameLabel id="A26ABB6FFFBDFFC7FCF5D9C5E52FF96F" box="[238,327,1712,1736]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B9590614FFBDFFC7FCF5D9C5E52FF96F" bold="true" box="[238,327,1712,1736]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
E, F:
<taxonomicName id="4C2DA185FFBDFFC7FD97D9C4E708F960" authority="Murao" authorityName="Murao" box="[396,864,1712,1736]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="latifacies" status="sp. nov." subGenus="Evylaeus">
<emphasis id="B9590614FFBDFFC7FD97D9C4E64AF96F" box="[396,546,1713,1736]" italics="true" pageId="7" pageNumber="8">Lasioglossum</emphasis>
(
<emphasis id="B9590614FFBDFFC7FE2FD9C4E6FFF96F" box="[564,663,1713,1736]" italics="true" pageId="7" pageNumber="8">Evylaeus</emphasis>
)
<emphasis id="B9590614FFBDFFC7FEBCD9C4E764F96F" box="[679,780,1713,1736]" italics="true" pageId="7" pageNumber="8">latifacies</emphasis>
Murao
</taxonomicName>
,
<taxonomicNameLabel id="A26ABB6FFFBDFFC7FF70D9C5E7ACF96F" box="[875,964,1712,1736]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B9590614FFBDFFC7FF70D9C5E7ACF96F" bold="true" box="[875,964,1712,1736]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
A–F: holotypes, females. A, C, E: head in frontal view. B, D, F: first metasomal tergum (left half, sculpture; right half, hair pattern). Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8B92DA06FFBDFFC7FCDED863E583F8F2" blockId="7.[151,1437,1814,2027]" pageId="7" pageNumber="8">Color. Tegula yellowish brown or brown translucent. Posterior margins of metasomal terga narrowly yellowish brown translucent.</paragraph>
<paragraph id="8B92DA06FFBDFFC7FCDED814E6F2F807" blockId="7.[151,1437,1814,2027]" pageId="7" pageNumber="8">
Pilosity. T1 (
<figureCitation id="1316C683FFBDFFC7FD7DD814E5AEF8DC" box="[358,454,1889,1915]" captionStart="FIGURE 4. A, B" captionStartId="7.[151,257,1680,1704]" captionTargetBox="[200,1382,186,1650]" captionTargetId="figure@7.[187,1399,169,1678]" captionTargetPageId="7" captionText="FIGURE 4. A, B: Lasioglossum (Evylaeus) bicornutum Murao, sp. nov. C, D: Lasioglossum (Evylaeus) silvicolum Murao, sp. nov. E, F: Lasioglossum (Evylaeus) latifacies Murao, sp. nov. A – F: holotypes, females. A, C, E: head in frontal view. B, D, F: first metasomal tergum (left half, sculpture; right half, hair pattern). Scale bar: 1 mm." httpUri="https://zenodo.org/record/276442/files/figure.png" pageId="7" pageNumber="8">Fig. 4B</figureCitation>
) medially with moderately dense simple and short hairs. Metasomal terga with neither basal hair bands nor apical fimbriae.
</paragraph>
<paragraph id="8B92DA06FFBDFFC7FCDED8DEE1F4F84C" blockId="7.[151,1437,1814,2027]" pageId="7" pageNumber="8">Structure. Head wider than or nearly as wide as long; head length/width ratio 0.95–1.01 (n= 10). Interocellar distance subequal to ocellocular distance. Frons and paraocular area dull, reticulate-punctate.</paragraph>
<paragraph id="8B92DA06FFB2FFC8FC8CDFEDE501FDD5" blockId="8.[151,1437,152,1484]" pageId="8" pageNumber="9">
Supraclypeal area (
<figureCitation id="1316C683FFB2FFC8FD98DFEDE5A3FF15" box="[387,459,152,178]" captionStart="FIGURE 3" captionStartId="6.[151,257,1883,1907]" captionTargetBox="[157,1430,888,1844]" captionTargetId="figure@6.[155,1432,863,1862]" captionTargetPageId="6" captionText="FIGURE 3. Lasioglossum (Evylaeus) bicornutum Murao, sp. nov. A – F: holotype, female. G – I: paratype, female. A: lateral habitus. B: lower face. C: mesoscutum. D: sculpture on mesoscutum. E: mesoscutellum. F: mesepisternum. G: labrum. H: propodeal dorsum. I: inner hind tibial spur. Scale bars: H, 0.25 mm; G, I, 0.2 mm." httpUri="https://zenodo.org/record/276441/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
B) dull, with sparse PP; IS with weak tessellation. Clypeal length 1.2x between lower rim of antennal socket and upper margin of clypeus, with moderately dense PP on upper half, and with larger shallow PP on lower half; IS with weak tessellation on upper half, and smooth on lower half. Basal area of labrum 3.1x as wide as long; lateral projections of distal process small, horn-like; keel of distal process moderately broad, apically bluntly pointed in frontal view. Scape length
<quantity id="4CD577E3FFB2FFC8F817DE58E0F5FEE0" box="[1036,1181,301,327]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="8" pageNumber="9" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.6–0.8 mm</quantity>
(n= 5). Mesoscutum (
<figureCitation id="1316C683FFB2FFC8FCBBDE27E482FECB" box="[160,234,338,364]" captionStart="FIGURE 3" captionStartId="6.[151,257,1883,1907]" captionTargetBox="[157,1430,888,1844]" captionTargetId="figure@6.[155,1432,863,1862]" captionTargetPageId="6" captionText="FIGURE 3. Lasioglossum (Evylaeus) bicornutum Murao, sp. nov. A – F: holotype, female. G – I: paratype, female. A: lateral habitus. B: lower face. C: mesoscutum. D: sculpture on mesoscutum. E: mesoscutellum. F: mesepisternum. G: labrum. H: propodeal dorsum. I: inner hind tibial spur. Scale bars: H, 0.25 mm; G, I, 0.2 mm." httpUri="https://zenodo.org/record/276441/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
C, D) shiny, with dense PP and IS with weak tessellation on the entire surface. Mesoscutellal IS marginally with weak tessellation, medially smooth; other sculptures similar to mesoscutum. Mesepisternum (
<figureCitation id="1316C683FFB2FFC8FCBBDEE8E48EFE10" box="[160,230,413,439]" captionStart="FIGURE 3" captionStartId="6.[151,257,1883,1907]" captionTargetBox="[157,1430,888,1844]" captionTargetId="figure@6.[155,1432,863,1862]" captionTargetPageId="6" captionText="FIGURE 3. Lasioglossum (Evylaeus) bicornutum Murao, sp. nov. A – F: holotype, female. G – I: paratype, female. A: lateral habitus. B: lower face. C: mesoscutum. D: sculpture on mesoscutum. E: mesoscutellum. F: mesepisternum. G: labrum. H: propodeal dorsum. I: inner hind tibial spur. Scale bars: H, 0.25 mm; G, I, 0.2 mm." httpUri="https://zenodo.org/record/276441/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
F) shiny, nearly smooth on upper area, and with weak rugulae on lower area. Propodeal dorsum (
<figureCitation id="1316C683FFB2FFC8F977DEE8E4CDFE7B" captionStart="FIGURE 3" captionStartId="6.[151,257,1883,1907]" captionTargetBox="[157,1430,888,1844]" captionTargetId="figure@6.[155,1432,863,1862]" captionTargetPageId="6" captionText="FIGURE 3. Lasioglossum (Evylaeus) bicornutum Murao, sp. nov. A – F: holotype, female. G – I: paratype, female. A: lateral habitus. B: lower face. C: mesoscutum. D: sculpture on mesoscutum. E: mesoscutellum. F: mesepisternum. G: labrum. H: propodeal dorsum. I: inner hind tibial spur. Scale bars: H, 0.25 mm; G, I, 0.2 mm." httpUri="https://zenodo.org/record/276441/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
H) 0.8x mesoscutellum, and 1.4x as long as metanotum; dorsomedially with longitudinal ridges on basal half, and with distinct tessellation on apical half; lateral slope, propodeal side, and shield with distinct tessellation on the entire surface. Inner hind tibial spur (
<figureCitation id="1316C683FFB2FFC8FF38DD78E700FD80" box="[803,872,525,551]" captionStart="FIGURE 3" captionStartId="6.[151,257,1883,1907]" captionTargetBox="[157,1430,888,1844]" captionTargetId="figure@6.[155,1432,863,1862]" captionTargetPageId="6" captionText="FIGURE 3. Lasioglossum (Evylaeus) bicornutum Murao, sp. nov. A – F: holotype, female. G – I: paratype, female. A: lateral habitus. B: lower face. C: mesoscutum. D: sculpture on mesoscutum. E: mesoscutellum. F: mesepisternum. G: labrum. H: propodeal dorsum. I: inner hind tibial spur. Scale bars: H, 0.25 mm; G, I, 0.2 mm." httpUri="https://zenodo.org/record/276441/files/figure.png" pageId="8" pageNumber="9">
Fig.
<date id="FF93FCC6FFB2FFC8FF41DD78E700FD80" box="[858,872,525,551]" pageId="8" pageNumber="9">3</date>
</figureCitation>
I) with 3–4 teeth (n= 5). T1 (
<figureCitation id="1316C683FFB2FFC8F8A2DD78E17BFD80" box="[1209,1299,525,551]" captionStart="FIGURE 4. A, B" captionStartId="7.[151,257,1680,1704]" captionTargetBox="[200,1382,186,1650]" captionTargetId="figure@7.[187,1399,169,1678]" captionTargetPageId="7" captionText="FIGURE 4. A, B: Lasioglossum (Evylaeus) bicornutum Murao, sp. nov. C, D: Lasioglossum (Evylaeus) silvicolum Murao, sp. nov. E, F: Lasioglossum (Evylaeus) latifacies Murao, sp. nov. A – F: holotypes, females. A, C, E: head in frontal view. B, D, F: first metasomal tergum (left half, sculpture; right half, hair pattern). Scale bar: 1 mm." httpUri="https://zenodo.org/record/276442/files/figure.png" pageId="8" pageNumber="9">Fig. 4B</figureCitation>
) with weak lineolation on the entire surface, and medially with moderately dense fine PP. T2–T4 with weak lineolation on the entire surface.
</paragraph>
<paragraph id="8B92DA06FFB2FFC8FCDEDD08E5E6FD30" blockId="8.[151,1437,152,1484]" box="[197,398,637,663]" pageId="8" pageNumber="9">
<emphasis id="B9590614FFB2FFC8FCDEDD08E56CFD30" bold="true" box="[197,260,637,663]" pageId="8" pageNumber="9">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C337898DFFB2FFC8FCDEDDD7E4AFFC8B" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8B92DA06FFB2FFC8FCDEDDD7E4AFFC8B" blockId="8.[151,1437,152,1484]" pageId="8" pageNumber="9">
<emphasis id="B9590614FFB2FFC8FCDEDDD7E529FD1B" bold="true" box="[197,321,674,700]" pageId="8" pageNumber="9">Remarks.</emphasis>
The species code sp. Y–3 reported by
<bibRefCitation id="EFBCA7F7FFB2FFC8FF04DDD7E062FD1B" box="[799,1034,674,700]" pageId="8" pageNumber="9" refString="Maeta, Y., Hanan, M. A., Miyanaga, R. &amp; Goukon, K. (2010) Relationship between entomophilous plants and their flower visitors in an island of Japan. Bulletin of the Hoshizaki Green Foundation, (13), 1 - 34. (In Japanese with English summary)" type="journal article">
Maeta
<emphasis id="B9590614FFB2FFC8FF68DDD7E7C1FD1C" box="[883,937,674,699]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. (2010)
</bibRefCitation>
corresponds to this new species. Within the
<emphasis id="B9590614FFB2FFC8FD07DDBDE5DBFD46" box="[284,435,712,737]" italics="true" pageId="8" pageNumber="9">sexstrigatum</emphasis>
-group, this species is similar to
<taxonomicName id="4C2DA185FFB2FFC8FF2CDDBDE089FD46" box="[823,1249,712,738]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="silvicolum" status="sp. nov." subGenus="Evylaeus">
<emphasis id="B9590614FFB2FFC8FF2CDDBDE7B1FD46" box="[823,985,712,737]" italics="true" pageId="8" pageNumber="9">Lasioglossum</emphasis>
(
<emphasis id="B9590614FFB2FFC8FFF7DDBDE03FFD46" box="[1004,1111,712,737]" italics="true" pageId="8" pageNumber="9">Evylaeus</emphasis>
)
<emphasis id="B9590614FFB2FFC8F873DDBDE089FD46" box="[1128,1249,712,737]" italics="true" pageId="8" pageNumber="9">silvicolum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26ABB6FFFB2FFC8F8F1DDB2E121FD46" box="[1258,1353,711,737]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B9590614FFB2FFC8F8F1DDB2E121FD46" bold="true" box="[1258,1353,711,737]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C2DA185FFB2FFC8F99FDDBDE675FCA0" authority="Ebmer &amp; Sakagami" authorityName="Ebmer &amp; Sakagami" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zipangu" subGenus="Evylaeus">
<emphasis id="B9590614FFB2FFC8F99FDDBDE1FCFD46" box="[1412,1428,712,737]" italics="true" pageId="8" pageNumber="9">L</emphasis>
. (
<emphasis id="B9590614FFB2FFC8FCBADD98E4DBFCA1" box="[161,179,749,774]" italics="true" pageId="8" pageNumber="9">E</emphasis>
.)
<emphasis id="B9590614FFB2FFC8FCD7DD98E541FCA1" box="[204,297,749,774]" italics="true" pageId="8" pageNumber="9">zipangu</emphasis>
Ebmer &amp; Sakagami
</taxonomicName>
from the southern Ryukyus. For the differences among the species, see the key.
</paragraph>
</subSubSection>
<subSubSection id="C337898DFFB2FFC8FCDEDC42E555FCD0" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8B92DA06FFB2FFC8FCDEDC42E555FCD0" blockId="8.[151,1437,152,1484]" pageId="8" pageNumber="9">
<emphasis id="B9590614FFB2FFC8FCDEDC42E539FCF6" bold="true" box="[197,337,823,849]" pageId="8" pageNumber="9">Etymology.</emphasis>
The specific name is derived from the female labrum with horn-like lateral projections on the distal process.
</paragraph>
</subSubSection>
<subSubSection id="C337898DFFB2FFC8FCDEDCF7E562FBF0" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8B92DA06FFB2FFC8FCDEDCF7E06BFC3B" blockId="8.[151,1437,152,1484]" box="[197,1027,898,924]" pageId="8" pageNumber="9">
<emphasis id="B9590614FFB2FFC8FCDEDCF7E50DFC3B" bold="true" box="[197,357,898,924]" pageId="8" pageNumber="9">Distribution.</emphasis>
<collectingCountry id="F33A9A96FFB2FFC8FD77DCF7E5C7FC3B" box="[364,431,898,924]" name="Japan" pageId="8" pageNumber="9">Japan</collectingCountry>
(southern Ryukyus: Ishigaki-jima, Iriomote-jima).
</paragraph>
<paragraph id="8B92DA06FFB2FFC8FCDEDCD2E6A8FC65" blockId="8.[151,1437,152,1484]" box="[197,704,935,962]" pageId="8" pageNumber="9">
<emphasis id="B9590614FFB2FFC8FCDEDCD2E514FC66" bold="true" box="[197,380,935,961]" pageId="8" pageNumber="9">Flight records.</emphasis>
Female: March to October.
</paragraph>
<paragraph id="8B92DA06FFB2FFC8FCDEDCB8E562FBF0" blockId="8.[151,1437,152,1484]" pageId="8" pageNumber="9">
<emphasis id="B9590614FFB2FFC8FCDEDCB8E5E4FC40" bold="true" box="[197,396,973,999]" pageId="8" pageNumber="9">Flower records.</emphasis>
The flowering plants visited by this species were five species in five families listed as follows.
<taxonomicName id="4C2DA185FFB2FFC8FD17DC87E5CAFBAB" box="[268,418,1010,1036]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
:
<taxonomicName id="4C2DA185FFB2FFC8FDB4DC87E60BFBAC" box="[431,611,1010,1035]" class="Magnoliopsida" family="Asteraceae" genus="Bidens" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="pilosa">
<emphasis id="B9590614FFB2FFC8FDB4DC87E60BFBAC" box="[431,611,1010,1035]" italics="true" pageId="8" pageNumber="9">Bidens pilosa</emphasis>
</taxonomicName>
<taxonomicName id="4C2DA185FFB2FFC8FE77DC87E69BFBAC" box="[620,755,1010,1036]" pageId="8" pageNumber="9" rank="variety" variety="minor">
var.
<emphasis id="B9590614FFB2FFC8FEBCDC87E69BFBAC" box="[679,755,1010,1035]" italics="true" pageId="8" pageNumber="9">minor</emphasis>
</taxonomicName>
.
<taxonomicName id="4C2DA185FFB2FFC8FF1EDC87E7ECFBAB" box="[773,900,1010,1036]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
:
<taxonomicName id="4C2DA185FFB2FFC8FF89DC87E0F6FBAC" box="[914,1182,1010,1035]" class="Magnoliopsida" family="Fabaceae" genus="Melilotus" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="officinalis">
<emphasis id="B9590614FFB2FFC8FF89DC87E0F6FBAC" box="[914,1182,1010,1035]" italics="true" pageId="8" pageNumber="9">Melilotus officinalis</emphasis>
</taxonomicName>
subsp. s
<emphasis id="B9590614FFB2FFC8F909DC87E1FDFBAC" box="[1298,1429,1010,1035]" italics="true" pageId="8" pageNumber="9">uaveolens</emphasis>
.
<taxonomicName id="4C2DA185FFB2FFC8FC8CDB6DE51DFB95" box="[151,373,1048,1074]" class="Magnoliopsida" family="Melastomataceae" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Melastomataceae</taxonomicName>
:
<taxonomicName id="4C2DA185FFB2FFC8FD99DB6DE6E7FB96" box="[386,655,1048,1073]" class="Magnoliopsida" family="Melastomataceae" genus="Melastoma" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="candidum">
<emphasis id="B9590614FFB2FFC8FD99DB6DE6E7FB96" box="[386,655,1048,1073]" italics="true" pageId="8" pageNumber="9">Melastoma candidum</emphasis>
</taxonomicName>
.
<taxonomicName id="4C2DA185FFB2FFC8FEBADB6DE729FB95" box="[673,833,1048,1074]" class="Magnoliopsida" family="Primulaceae" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Primulaceae</taxonomicName>
:
<taxonomicName id="4C2DA185FFB2FFC8FF56DB6DE035FB96" box="[845,1117,1048,1073]" class="Magnoliopsida" family="Primulaceae" genus="Lysimachia" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="sikokiana">
<emphasis id="B9590614FFB2FFC8FF56DB6DE035FB96" box="[845,1117,1048,1073]" italics="true" pageId="8" pageNumber="9">Lysimachia sikokiana</emphasis>
</taxonomicName>
.
<taxonomicName id="4C2DA185FFB2FFC8F874DB6DE09EFB95" box="[1135,1270,1048,1074]" class="Magnoliopsida" family="Rubiaceae" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Rubiaceae</taxonomicName>
:
<taxonomicName id="4C2DA185FFB2FFC8F918DB6DE56BFBF1" class="Magnoliopsida" family="Rubiaceae" genus="Ophiorrhiza" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="kuroiwae">
<emphasis id="B9590614FFB2FFC8F918DB6DE56BFBF1" italics="true" pageId="8" pageNumber="9">Ophiorrhiza kuroiwae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C337898DFFB2FFC8FCDEDB17E1E5FA6B" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8B92DA06FFB2FFC8FCDEDB17E51DFA00" blockId="8.[151,1437,152,1484]" pageId="8" pageNumber="9">
<emphasis id="B9590614FFB2FFC8FCDEDB17E5E9FBDB" bold="true" box="[197,385,1122,1148]" pageId="8" pageNumber="9">
<typeStatus id="549664A4FFB2FFC8FCDEDB17E56CFBDB" box="[197,260,1122,1148]" pageId="8" pageNumber="9">Type</typeStatus>
material.
</emphasis>
<typeStatus id="549664A4FFB2FFC8FD91DB17E592FBDB" box="[394,506,1122,1148]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
(
<typeStatus id="549664A4FFB2FFC8FE10DB17E621FBDB" box="[523,585,1122,1148]" pageId="8" pageNumber="9">Type</typeStatus>
No.
<collectionCode id="ED3C42C3FFB2FFC8FE9DDB17E6BEFBDB" box="[646,726,1122,1148]" pageId="8" pageNumber="9">ELKU</collectionCode>
3305): female, Mt. Kawara-dake, Ishigaki-jima, Okinawa Pref., Ryukyus,
<collectingCountry id="F33A9A96FFB2FFC8FD48DBFDE5C7FB05" box="[339,431,1160,1186]" name="Japan" pageId="8" pageNumber="9">JAPAN</collectingCountry>
,
<date id="FF93FCC6FFB2FFC8FDA1DBFDE62FFB05" box="[442,583,1160,1186]" pageId="8" pageNumber="9" value="1964-03-18">18. iii. 1964</date>
(Y. Miyatake leg.,
<collectionCode id="ED3C42C3FFB2FFC8FF33DBFDE714FB05" box="[808,892,1160,1186]" pageId="8" pageNumber="9">ELKU</collectionCode>
).
<typeStatus id="549664A4FFB2FFC8FF95DBFDE06CFB05" box="[910,1028,1160,1186]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
: [Ryukyus:
<collectingCountry id="F33A9A96FFB2FFC8F894DBFDE083FB05" box="[1167,1259,1160,1186]" name="Japan" pageId="8" pageNumber="9">JAPAN</collectingCountry>
] Ishigaki-jima, Okinawa Pref.:
<specimenCount id="9D2B118FFFB2FFC8FD4DDBD8E5ABFB60" box="[342,451,1197,1223]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
, same locality as the
<typeStatus id="549664A4FFB2FFC8FED3DBD8E75DFB60" box="[712,821,1197,1223]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
,
<date id="FF93FCC6FFB2FFC8FF59DBD8E7A3FB60" box="[834,971,1197,1223]" pageId="8" pageNumber="9" value="1963-10-28">28. x. 1963</date>
(Y. Hirashima,
<collectionCode id="ED3C42C3FFB2FFC8F896DBD8E08AFB60" box="[1165,1250,1197,1223]" pageId="8" pageNumber="9">ELKU</collectionCode>
);
<specimenCount id="9D2B118FFFB2FFC8F8E3DBD8E10CFB60" box="[1272,1380,1197,1223]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
, Mt. Omoto-dake,
<date id="FF93FCC6FFB2FFC8FD21DBA7E5DFFB4B" box="[314,439,1234,1260]" pageId="8" pageNumber="9" value="1979-04-09">9. iv. 1979</date>
(H. Makihara,
<collectionCode id="ED3C42C3FFB2FFC8FE75DBA7E6AAFB4B" box="[622,706,1234,1260]" pageId="8" pageNumber="9">ELKU</collectionCode>
). Iriomote-jima, Okinawa Pref.:
<specimenCount id="9D2B118FFFB2FFC8F85FDBA7E0D2FB4B" box="[1092,1210,1234,1260]" pageId="8" pageNumber="9" type="female">3 females</specimenCount>
,
<date id="FF93FCC6FFB2FFC8F8DDDBA7E119FB4B" box="[1222,1393,1234,1260]" pageId="8" pageNumber="9" value="2004-06-07" valueMax="2004-06-11" valueMin="2004-06-07">7–11. vi. 2004</date>
(Y. Maeta,
<collectionCode id="ED3C42C3FFB2FFC8FCF4DB8DE52FFAB5" box="[239,327,1272,1298]" pageId="8" pageNumber="9">MCDS</collectionCode>
),
<date id="FF93FCC6FFB2FFC8FD40DB8DE5B5FAB5" box="[347,477,1272,1298]" pageId="8" pageNumber="9" value="2007-05-23">23. v. 2007</date>
(Y. Maeta,
<collectionCode id="ED3C42C3FFB2FFC8FE73DB8DE6A8FAB5" box="[616,704,1272,1298]" pageId="8" pageNumber="9">MCDS</collectionCode>
),
<date id="FF93FCC6FFB2FFC8FECFDB8DE7EBFAB5" box="[724,899,1272,1298]" pageId="8" pageNumber="9" value="2007-05-23" valueMax="2007-05-27" valueMin="2007-05-23">23–27. v. 2007</date>
(Y. Maeta,
<collectionCode id="ED3C42C3FFB2FFC8F816DB8DE00DFAB5" box="[1037,1125,1272,1298]" pageId="8" pageNumber="9">MCDS</collectionCode>
);
<specimenCount id="9D2B118FFFB2FFC8F862DB8DE08BFAB5" box="[1145,1251,1272,1298]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
, Itokawa-rindô,
<date id="FF93FCC6FFB2FFC8FC8CDA68E520FA90" box="[151,328,1309,1335]" pageId="8" pageNumber="9" value="2004-05-10" valueMax="2004-05-13" valueMin="2004-05-10">10–13. v. 2004</date>
(T. Mita,
<collectionCode id="ED3C42C3FFB2FFC8FDD9DA68E671FA90" box="[450,537,1309,1335]" pageId="8" pageNumber="9">RMPC</collectionCode>
);
<specimenCount id="9D2B118FFFB2FFC8FE35DA68E6F1FA90" box="[558,665,1309,1335]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
, Ôtomi,
<date id="FF93FCC6FFB2FFC8FF1ADA68E7FAFA90" box="[769,914,1309,1335]" pageId="8" pageNumber="9" value="2005-03-16">16. iii. 2005</date>
(R. Murao,
<collectionCode id="ED3C42C3FFB2FFC8F83DDA68E015FA90" box="[1062,1149,1309,1335]" pageId="8" pageNumber="9">RMPC</collectionCode>
);
<specimenCount id="9D2B118FFFB2FFC8F888DA68E161FA90" box="[1171,1289,1309,1335]" pageId="8" pageNumber="9" type="female">2 females</specimenCount>
, Shirahama, Shirine,
<date id="FF93FCC6FFB2FFC8FCE0DA37E5DBFAFB" box="[251,435,1346,1372]" pageId="8" pageNumber="9" value="2004-04-23" valueMax="2004-04-27" valueMin="2004-04-23">23–27. iv. 2004</date>
(T. Mita,
<collectionCode id="ED3C42C3FFB2FFC8FE30DA37E6EAFAFB" box="[555,642,1346,1372]" pageId="8" pageNumber="9">RMPC</collectionCode>
);
<specimenCount id="9D2B118FFFB2FFC8FE8CDA37E763FAFB" box="[663,779,1346,1372]" pageId="8" pageNumber="9" type="female">2 females</specimenCount>
, Shirahama-rindô,
<date id="FF93FCC6FFB2FFC8FFF7DA37E011FAFB" box="[1004,1145,1346,1372]" pageId="8" pageNumber="9" value="2004-04-25">25. iv. 2004</date>
(T. Mita,
<collectionCode id="ED3C42C3FFB2FFC8F8EBDA37E12EFAFB" box="[1264,1350,1346,1372]" pageId="8" pageNumber="9">RMPC</collectionCode>
),
<date id="FF93FCC6FFB2FFC8F940DA37E4B9FA25" pageId="8" pageNumber="9" value="2007-05-21">21. v. 2007</date>
(R. Murao,
<collectionCode id="ED3C42C3FFB2FFC8FD7ADA1DE5D0FA25" box="[353,440,1384,1410]" pageId="8" pageNumber="9">RMPC</collectionCode>
);
<specimenCount id="9D2B118FFFB2FFC8FDD0DA1DE65CFA25" box="[459,564,1384,1410]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
, Sonai,
<date id="FF93FCC6FFB2FFC8FE8BDA1DE774FA25" box="[656,796,1384,1410]" pageId="8" pageNumber="9" value="2005-03-15">15. iii. 2005</date>
(R. Murao,
<collectionCode id="ED3C42C3FFB2FFC8FFB7DA1DE06BFA25" box="[940,1027,1384,1410]" pageId="8" pageNumber="9">RMPC</collectionCode>
);
<specimenCount id="9D2B118FFFB2FFC8F80CDA1DE017FA25" box="[1047,1151,1384,1410]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
, Uehara,
<date id="FF93FCC6FFB2FFC8F8F6DA1DE107FA25" box="[1261,1391,1384,1410]" pageId="8" pageNumber="9" value="1975-05-25">25. v. 1975</date>
(H. Makihara,
<collectionCode id="ED3C42C3FFB2FFC8FD0DDAF8E501FA00" box="[278,361,1421,1447]" pageId="8" pageNumber="9">ELKU</collectionCode>
).
</paragraph>
<paragraph id="8B92DA06FFB2FFC8FCDEDAC7E1E5FA6B" blockId="8.[151,1437,152,1484]" box="[197,1421,1458,1484]" pageId="8" pageNumber="9">
<emphasis id="B9590614FFB2FFC8FCDEDAC7E5FBFA6B" bold="true" box="[197,403,1458,1484]" pageId="8" pageNumber="9">
<typeStatus id="549664A4FFB2FFC8FCDEDAC7E56AFA6B" box="[197,258,1458,1484]" pageId="8" pageNumber="9">Type</typeStatus>
depository.
</emphasis>
The
<typeStatus id="549664A4FFB2FFC8FDD4DAC7E65DFA6B" box="[463,565,1458,1484]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
and 10
<typeStatus id="549664A4FFB2FFC8FE88DAC7E76BFA6B" box="[659,771,1458,1484]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
are deposited in
<collectionCode id="ED3C42C3FFB2FFC8FFD7DAC7E075FA6B" box="[972,1053,1458,1484]" pageId="8" pageNumber="9">ELKU</collectionCode>
, and three
<typeStatus id="549664A4FFB2FFC8F887DAC7E164FA6B" box="[1180,1292,1458,1484]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
in
<collectionCode id="ED3C42C3FFB2FFC8F929DAC7E1E1FA6B" box="[1330,1417,1458,1484]" pageId="8" pageNumber="9">MCDS</collectionCode>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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