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<document id="90EB0E207952D7C14D04DF9E3A4F6499" ID-DOI="10.3853/j.0067-1975.63.2011.1581" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239547" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629575327011" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2011" docId="0384503AFFB83C5BFC1FF9ACFE696979" docLanguage="en" docName="RecAustMus.63.2.167-202.pdf" docOrigin="Records of the Australian Museum 63 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.63.2011.1581" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada atlantis Köhler 2011, n.sp." docType="treatment" docVersion="3" lastPageNumber="172" masterDocId="FFBD2842FFBB3C5EFFB8FFD2FFED6E4A" masterDocTitle="Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae)" masterLastPageNumber="202" masterPageNumber="167" pageNumber="170" updateTime="1699078749944" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="17B2FED150781F1DD0DFD85F7B9F473E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2656B66DBC0C00C8805BE20810396943">Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae)</mods:title>
</mods:titleInfo>
<mods:name id="A24A29EF4086D1C570361AC6960DB02C" type="personal">
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<mods:namePart id="C437B7A258B4D08895FC67FC51765791">Köhler, Frank</mods:namePart>
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<mods:typeOfResource id="0DD43CD093532DEC8E2BE59B299520FA">text</mods:typeOfResource>
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<mods:title id="7A269FB5D15B2BC4299A5CA6E6DC6E34">Records of the Australian Museum</mods:title>
</mods:titleInfo>
<mods:part id="A81FA8CD01A79F1D06C96514BE99D30D">
<mods:date id="36B51C748772EDB030657DBD283A038C">2011</mods:date>
<mods:detail id="EE92BEC5CCCEAC3367A4D75E8FFF11A7" type="pubDate">
<mods:number id="ACAA5438CCAA0B57E77A35BA401D7248">2011-11-30</mods:number>
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<mods:number id="DAA9DE034BD63E18A640CFA0E89A2127">63</mods:number>
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<mods:url id="4D13DA31C35357BA36895308B789B118">http://dx.doi.org/10.3853/j.0067-1975.63.2011.1581</mods:url>
</mods:location>
<mods:classification id="9675EC312500CF6E558ADC7F2A9B0C42">journal article</mods:classification>
<mods:identifier id="B5B3E633B370D28EB3630C0D7C3AD975" type="DOI">10.3853/j.0067-1975.63.2011.1581</mods:identifier>
<mods:identifier id="84FE102C2E1E73A0249D7FDDD9F57913" type="ISSN">2201-4349</mods:identifier>
<mods:identifier id="DEF2F4CE10420D7C278352F16F1718A7" type="Zenodo-Dep">5239547</mods:identifier>
<mods:identifier id="2EAB1020B9F3EA865B6D924F2D42B461" type="ZooBank">1BCD4085-D2B9-400D-B504-8C85C30303D6</mods:identifier>
</mods:mods>
<treatment id="0384503AFFB83C5BFC1FF9ACFE696979" LSID="urn:lsid:plazi:treatment:0384503AFFB83C5BFC1FF9ACFE696979" httpUri="http://treatment.plazi.org/id/0384503AFFB83C5BFC1FF9ACFE696979" lastPageId="5" lastPageNumber="172" pageId="3" pageNumber="170">
<subSubSection id="C337B2A7FFB83C5DFC1FF9ACFAED68D2" box="[935,1280,1662,1688]" pageId="3" pageNumber="170" type="nomenclature">
<paragraph id="8B92E12CFFB83C5DFC1FF9ACFAED68D2" blockId="3.[935,1280,1662,1688]" box="[935,1280,1662,1688]" pageId="3" pageNumber="170">
<heading id="D0DA5640FFB83C5DFC1FF9ACFAED68D2" box="[935,1280,1662,1688]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="170" reason="2">
<emphasis id="B9593D3EFFB83C5DFC1FF9ACFAED68D2" bold="true" box="[935,1280,1662,1688]" pageId="3" pageNumber="170">
<taxonomicName id="4C2D9AAFFFB83C5DFC1FF9ACFB5168D2" ID-CoL="D7Q3" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[935,1212,1662,1688]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="170" phylum="Mollusca" rank="species" species="atlantis" status="sp. nov.">
<emphasis id="B9593D3EFFB83C5DFC1FF9ACFB5168D2" bold="true" box="[935,1212,1662,1688]" italics="true" pageId="3" pageNumber="170">Amplirhagada atlantis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A8045FFB83C5DFB7CF9ACFAED68D2" box="[1220,1280,1662,1688]" pageId="3" pageNumber="170" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C337B2A7FFB83C5AFCA2F968FDC76DDA" lastPageId="4" lastPageNumber="171" pageId="3" pageNumber="170" type="materials_examined">
<paragraph id="8B92E12CFFB83C5DFCA2F968FA606946" blockId="3.[794,1421,1722,1804]" pageId="3" pageNumber="170">
<materialsCitation id="3B45EB71FFB83C5DFCA2F968FA656946" collectingDate="1987-06-04" collectionCode="V" collectorName="NW Kimberley" latitude="-14.077778" location="Cape Bernier" longLatPrecision="21" longitude="127.49027" pageId="3" pageNumber="170" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B9593D3EFFB83C5DFCA2F968FC436898" bold="true" box="[794,942,1722,1746]" pageId="3" pageNumber="170">
<typeStatus id="54965F8EFFB83C5DFCA2F968FCBE6898" box="[794,851,1722,1746]" pageId="3" pageNumber="170">Type</typeStatus>
locality
</emphasis>
.
<collectingRegion id="49E92FCEFFB83C5DFC06F969FB6C6898" box="[958,1153,1723,1746]" country="Australia" name="Western Australia" pageId="3" pageNumber="170">Western Australia</collectingRegion>
,
<collectorName id="26D884FAFFB83C5DFB37F969FADA6898" box="[1167,1335,1723,1746]" pageId="3" pageNumber="170">NW Kimberley</collectorName>
, Joseph Bonaparte Gulf,
<quantity id="4CD54CC9FFB83C5DFC76F90AFBF468A5" box="[974,1049,1752,1775]" metricMagnitude="3" metricUnit="m" metricValue="8.6" pageId="3" pageNumber="170" unit="km" value="8.6">8.6 km</quantity>
SSE of
<location id="8EF2B7F7FFB83C5DFBC1F90AFAE468A5" LSID="urn:lsid:plazi:treatment:0384503AFFB83C5BFC1FF9ACFE696979:8EF2B7F7FFB83C5DFBC1F90AFAE468A5" box="[1145,1289,1752,1775]" latitude="-14.077778" longLatPrecision="21" longitude="127.49027" name="Cape Bernier" pageId="3" pageNumber="170" stateProvince="Western Australia">Cape Bernier</location>
;
<geoCoordinate id="EE1987EBFFB83C5DFAAFF90AFA6068A5" box="[1303,1421,1752,1775]" degrees="14" direction="south" minutes="04" orientation="latitude" pageId="3" pageNumber="170" precision="15" seconds="40" value="-14.077778">14°04'40&quot;S</geoCoordinate>
<geoCoordinate id="EE1987EBFFB83C5DFCA2F927FC766946" box="[794,923,1781,1804]" degrees="127" direction="east" minutes="29" orientation="longitude" pageId="3" pageNumber="170" precision="15" seconds="25" value="127.49027">127°29'25&quot;E</geoCoordinate>
(RFS-04-1; coll.
<collectionCode id="ED3C79E9FFB83C5DFBFDF927FBBA6946" box="[1093,1111,1781,1804]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="170" type="Museum">V</collectionCode>
. Kessner,
<date id="FF93C7ECFFB83C5DFB01F927FADF6946" box="[1209,1330,1781,1804]" pageId="3" pageNumber="170" value="1987-06-04">
<collectingDate id="EFD73E04FFB83C5DFB01F927FADF6946" box="[1209,1330,1781,1804]" pageId="3" pageNumber="170" value="1987-06-04">04 Jun 1987</collectingDate>
</date>
) (
<figureCitation id="1316FDA9FFB83C5DFAFDF927FA906946" box="[1349,1405,1781,1804]" captionStart="Figure 1" captionStartId="1.[121,184,1877,1898]" captionTargetBox="[149,1389,975,1848]" captionTargetId="figure-423@1.[136,1403,961,1856]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species described in the present work. (A) A. atlantis n.sp. (B) A. carsoniana n.sp. (C) A. alicunda n.sp. (D) A. davidsoniana n.sp. (E) A. vialae n.sp. and A. discoidea n.sp. (F) A. forrestiana n.sp. (G) A. inusitata n.sp. and A. epiphallica n.sp. (H) A. lindsayae n.sp. (I) A. angustocauda n.sp. (K) A. moraniana n.sp. (L) A. gardneriana n.sp. (M) A. bendraytoni n.sp. (N) A. coffea n.sp. (P) A. carinata Solem, 1981. When species occur at more than one locality, the type locality of newly described taxa is marked with an asterisk." figureDoi="http://doi.org/10.5281/zenodo.5239549" httpUri="https://zenodo.org/record/5239549/files/figure.png" pageId="3" pageNumber="170">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B92E12CFFB83C5DFCA2F8E2FB946990" blockId="3.[794,1422,1840,2010]" pageId="3" pageNumber="170">
<emphasis id="B9593D3EFFB83C5DFCA2F8E2FC236902" bold="true" box="[794,974,1840,1864]" pageId="3" pageNumber="170">Type material</emphasis>
.
<materialsCitation id="3B45EB71FFB83C5DFC5AF8E2FB8E692E" accessionNumber="S34704" collectionCode="WAM" pageId="3" pageNumber="170" specimenCount="1" typeStatus="holotype">
<typeStatus id="54965F8EFFB83C5DFC5AF8E2FBBD690D" box="[994,1104,1840,1863]" pageId="3" pageNumber="170" type="holotype">Holotype</typeStatus>
<collectionCode id="ED3C79E9FFB83C5DFBE3F8E2FB4D690D" box="[1115,1184,1840,1863]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="3" pageNumber="170" type="Museum">WAM</collectionCode>
<accessionNumber id="947E7CCFFFB83C5DFB14F8E2FAEB690D" box="[1196,1286,1840,1863]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34704" pageId="3" pageNumber="170" type="EnaNcbi">S34704</accessionNumber>
(preserved specimen) (Pl. 1.1;
<tableCitation id="C6AFD497FFB83C5DFBB9F89FFBBB692E" box="[1025,1110,1869,1892]" captionStart="Table 1" captionStartId="33.[121,184,1071,1095]" captionText="Table 1. Shell measurements of the type series. Given are maximumminimum (mean±standard deviation) of N measured adult shells. Abbreviations used: HT = holotype." pageId="3" pageNumber="170">Table 1</tableCitation>
)
</materialsCitation>
.
<materialsCitation id="3B45EB71FFB83C5DFBCBF89FFC7969D5" accessionNumber="S34705" collectionCode="AM, WAM" pageId="3" pageNumber="170" specimenCount="1" typeStatus="paratype">
<typeStatus id="54965F8EFFB83C5DFBCBF89FFB0D692E" box="[1139,1248,1869,1892]" pageId="3" pageNumber="170" type="paratype">Paratypes</typeStatus>
<collectionCode id="ED3C79E9FFB83C5DFB51F89FFAF8692E" box="[1257,1301,1869,1892]" country="Australia" name="Australian Museum" pageId="3" pageNumber="170" type="Museum">AM</collectionCode>
C.472918 (4 preserved specimens),
<collectionCode id="ED3C79E9FFB83C5DFBF2F8B9FB6369C8" box="[1098,1166,1899,1922]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="3" pageNumber="170" type="Museum">WAM</collectionCode>
<accessionNumber id="947E7CCFFFB83C5DFB21F8B9FB1D69C8" box="[1177,1264,1899,1922]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34705" pageId="3" pageNumber="170" type="EnaNcbi">S34705</accessionNumber>
(8 preserved specimens)
</materialsCitation>
,
<materialsCitation id="3B45EB71FFB83C5DFC26F85AFAB469D5" accessionNumber="S34705" box="[926,1369,1928,1951]" collectionCode="FMNH" isTailEnumeration="true" pageId="3" pageNumber="170" specimenCode="FMNH 220006" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB8B4957FFB83C5DFC26F85AFBAF69D5" box="[926,1090,1928,1951]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="170" type="Museum">FMNH 220006</specimenCode>
(12 preserved specimens)
</materialsCitation>
,
<materialsCitation id="3B45EB71FFB83C5DFADAF85AFA6469F6" accessionNumber="S34706" collectionCode="AM, WAM" pageId="3" pageNumber="170" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED3C79E9FFB83C5DFADAF85AFA6069D5" box="[1378,1421,1928,1951]" country="Australia" name="Australian Museum" pageId="3" pageNumber="170" type="Museum">AM</collectionCode>
C.472919 (3 dried shells),
<collectionCode id="ED3C79E9FFB83C5DFB8FF877FB9569F6" box="[1079,1144,1957,1980]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="3" pageNumber="170" type="Museum">WAM</collectionCode>
<accessionNumber id="947E7CCFFFB83C5DFBC7F877FB3C69F6" box="[1151,1233,1957,1980]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34706" pageId="3" pageNumber="170" type="EnaNcbi">S34706</accessionNumber>
(10 dried shells)
</materialsCitation>
,
<materialsCitation id="3B45EB71FFB83C5DFCA2F811FB986990" accessionNumber="S34706" box="[794,1141,1987,2010]" collectionCode="FMNH" isTailEnumeration="true" pageId="3" pageNumber="170" specimenCode="FMNH 220005" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB8B4957FFB83C5DFCA2F811FC506990" box="[794,957,1987,2010]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="170" type="Museum">FMNH 220005</specimenCode>
(13 dried shells)
</materialsCitation>
.
</paragraph>
<caption id="DF52B1A4FFBF3C5AFF1EFDFAFF156CE2" ID-DOI="http://doi.org/10.5281/zenodo.5239553" ID-Zenodo-Dep="5239553" httpUri="https://zenodo.org/record/5239553/files/figure.png" pageId="4" pageNumber="171" startId="4.[166,228,552,573]" targetBox="[170,786,160,527]" targetPageId="4">
<paragraph id="8B92E12CFFBF3C5AFF1EFDFAFF156CE2" blockId="4.[166,795,552,680]" pageId="4" pageNumber="171">
Figure 3. Genitalia of
<taxonomicName id="4C2D9AAFFFBF3C5AFECAFDFAFDAA6C77" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[370,583,552,573]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="171" phylum="Mollusca" rank="species" species="atlantis" status="sp. nov.">
<emphasis id="B9593D3EFFBF3C5AFECAFDFAFDAA6C77" box="[370,583,552,573]" italics="true" pageId="4" pageNumber="171">Amplirhagada atlantis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A8045FFBF3C5AFDF3FDFAFD956C77" box="[587,632,552,573]" pageId="4" pageNumber="171" rank="species">n.sp.</taxonomicNameLabel>
(AM C.472918). Abbreviations:
<emphasis id="B9593D3EFFBF3C5AFE85FD91FEB86C12" box="[317,341,579,600]" italics="true" pageId="4" pageNumber="171">ag</emphasis>
, albumen gland;
<emphasis id="B9593D3EFFBF3C5AFDBAFD91FDF46C12" box="[514,537,579,600]" italics="true" pageId="4" pageNumber="171">bc</emphasis>
, bursa copulatrix;
<emphasis id="B9593D3EFFBF3C5AFD6BFD91FD0B6C12" box="[723,742,579,600]" italics="true" pageId="4" pageNumber="171">fo</emphasis>
, free oviduct;
<emphasis id="B9593D3EFFBF3C5AFF46FD8CFEFB6C39" box="[254,278,606,627]" italics="true" pageId="4" pageNumber="171">hd</emphasis>
, hermaphroditic duct;
<emphasis id="B9593D3EFFBF3C5AFE44FD8CFDE56C39" box="[508,520,606,627]" italics="true" pageId="4" pageNumber="171">p</emphasis>
, penis;
<emphasis id="B9593D3EFFBF3C5AFDE2FD8CFD986C39" box="[602,629,606,627]" italics="true" pageId="4" pageNumber="171">rm</emphasis>
, penial retractor muscle;
<emphasis id="B9593D3EFFBF3C5AFF4FFDAAFEFF6CC7" box="[247,274,632,653]" italics="true" pageId="4" pageNumber="171">so,</emphasis>
sperm oviduct;
<emphasis id="B9593D3EFFBF3C5AFE15FDAAFE296CC7" box="[429,452,632,653]" italics="true" pageId="4" pageNumber="171">va</emphasis>
, vagina;
<emphasis id="B9593D3EFFBF3C5AFDA4FDAAFDDE6CC7" box="[540,563,632,653]" italics="true" pageId="4" pageNumber="171">vd</emphasis>
, vas deferens. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="8B92E12CFFBF3C5AFF1EFD37FDC76DDA" blockId="4.[166,794,741,912]" pageId="4" pageNumber="171">
<emphasis id="B9593D3EFFBF3C5AFF1EFD37FE7E6CB7" bold="true" box="[166,403,741,765]" pageId="4" pageNumber="171">Additional material</emphasis>
(not dissected).
<materialsCitation id="3B45EB71FFBF3C5AFDEFFD34FDCB6DDA" accessionNumber="S34751, S34752" collectingDate="1987-06-10" collectionCode="WAM, V" latitude="-14.115278" location="Evelyn Island" longLatPrecision="21" longitude="127.51945" pageId="4" pageNumber="171" specimenCode="FMNH 220327, FMNH 220328, RFS 10-2" specimenCount="1">
<collectionCode id="ED3C79E9FFBF3C5AFDEFFD34FD746CB7" box="[599,665,742,765]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="4" pageNumber="171" type="Museum">WAM</collectionCode>
<accessionNumber id="947E7CCFFFBF3C5AFD1CFD34FD156CB7" box="[676,760,742,765]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34751" pageId="4" pageNumber="171" type="EnaNcbi">S34751</accessionNumber>
(5 preserved specimens),
<specimenCode id="DB8B4957FFBF3C5AFE01FCD1FD816D50" box="[441,620,771,794]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="171" type="Museum">FMNH 220327</specimenCode>
(10 preserved specimens),
<collectionCode id="ED3C79E9FFBF3C5AFE9DFCF3FE886D72" box="[293,357,801,824]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="4" pageNumber="171" type="Museum">WAM</collectionCode>
<accessionNumber id="947E7CCFFFBF3C5AFED2FCF3FE546D72" box="[362,441,801,824]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34752" pageId="4" pageNumber="171" type="EnaNcbi">S34752</accessionNumber>
(8 dried shells),
<specimenCode id="DB8B4957FFBF3C5AFDD9FCF3FD126D72" box="[609,767,801,824]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="171" type="Museum">FMNH 220328</specimenCode>
(7 dried shells), all from
<quantity id="4CD54CC9FFBF3C5AFE34FCECFE386D1F" box="[396,469,830,853]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="171" unit="km" value="5.6">5.6 km</quantity>
W of
<location id="8EF2B7F7FFBF3C5AFDABFCECFD4A6D1F" LSID="urn:lsid:plazi:treatment:0384503AFFB83C5BFC1FF9ACFE696979:8EF2B7F7FFBF3C5AFDABFCECFD4A6D1F" box="[531,679,830,853]" latitude="-14.115278" longLatPrecision="21" longitude="127.51945" name="Evelyn Island" pageId="4" pageNumber="171">Evelyn Island</location>
, mainland, behind beach and mangroves;
<geoCoordinate id="EE1987EBFFBF3C5AFE64FC89FDBE6D38" box="[476,595,859,882]" degrees="14" direction="south" minutes="06" orientation="latitude" pageId="4" pageNumber="171" precision="15" seconds="55" value="-14.115278">14°06'55&quot;S</geoCoordinate>
<geoCoordinate id="EE1987EBFFBF3C5AFDE0FC89FD306D38" box="[600,733,859,882]" degrees="127" direction="east" minutes="31" orientation="longitude" pageId="4" pageNumber="171" precision="15" seconds="10" value="127.51945">127°31'10&quot;E</geoCoordinate>
(
<specimenCode id="DB8B4957FFBF3C5AFD51FC89FF376DDA" collectionCode="RFS" pageId="4" pageNumber="171">RFS 10-2</specimenCode>
; coll.
<collectionCode id="ED3C79E9FFBF3C5AFEA1FCABFEC16DDA" box="[281,300,889,912]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="171" type="Museum">V</collectionCode>
. Kessner,
<date id="FF93C7ECFFBF3C5AFE20FCABFDF36DDA" box="[408,542,889,912]" pageId="4" pageNumber="171" value="1987-06-10">
<collectingDate id="EFD73E04FFBF3C5AFE20FCABFDF36DDA" box="[408,542,889,912]" pageId="4" pageNumber="171" value="1987-06-10">10 Jun 1987</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C337B2A7FFBF3C5AFF1EFC61FD4C6AD2" pageId="4" pageNumber="171" type="etymology">
<paragraph id="8B92E12CFFBF3C5AFF1EFC61FD4C6AD2" blockId="4.[166,794,947,1176]" pageId="4" pageNumber="171">
<emphasis id="B9593D3EFFBF3C5AFF1EFC61FECE6D81" bold="true" box="[166,291,947,971]" pageId="4" pageNumber="171">Etymology</emphasis>
. Specific name referring to the name of the seaplane “Atlantis 1925” of the two German aviators Hans Bertram and Adolph Klausmann, who in 1932 undertook a goodwill flight from Cologne,
<collectingCountry id="F33AA1BCFFBF3C5AFE5FFBD9FDA66A68" box="[487,587,1035,1058]" name="Germany" pageId="4" pageNumber="171">Germany</collectingCountry>
, to
<collectingCountry id="F33AA1BCFFBF3C5AFDD4FBD9FD3C6A68" box="[620,721,1035,1058]" name="Australia" pageId="4" pageNumber="171">Australia</collectingCountry>
. On
<date id="FF93C7ECFFBF3C5AFD47FBD9FEE26A0A" pageId="4" pageNumber="171" value="1932-05-14">14 May 1932</date>
the aviators took off from Timor bound for Darwin but were stranded due to bad weather near Bertram Cove, where they survived for 40 days before being rescued. Being a noun the species epithet keeps its male suffix.
</paragraph>
</subSubSection>
<subSubSection id="C337B2A7FFBF3C5AFE2EFB6EFB736985" pageId="4" pageNumber="171" type="description">
<paragraph id="8B92E12CFFBF3C5AFE2EFB6EFDC76A9C" blockId="4.[406,554,1212,1238]" box="[406,554,1212,1238]" pageId="4" pageNumber="171">
<emphasis id="B9593D3EFFBF3C5AFE2EFB6EFDC76A9C" bold="true" box="[406,554,1212,1238]" pageId="4" pageNumber="171">Description</emphasis>
</paragraph>
<paragraph id="8B92E12CFFBF3C5AFF1EFB2BFE4E68C7" blockId="4.[166,794,1273,2000]" pageId="4" pageNumber="171">
<emphasis id="B9593D3EFFBF3C5AFF1EFB2BFF366B5A" box="[166,219,1273,1296]" italics="true" pageId="4" pageNumber="171">Shell</emphasis>
(Pl. 1.1,
<figureCitation id="1316FDA9FFBF3C5AFE82FB2BFE436B5A" box="[314,430,1273,1296]" captionStart="Figure 2" captionStartId="3.[121,184,1515,1536]" captionTargetBox="[121,1421,151,1498]" captionTargetId="figure-264@3.[121,1421,151,1498]" captionTargetPageId="3" captionText="Figure 2. Amplirhagada atlantis n.sp. SEM of shell (AM C.472919), radula and jaw (AM C.472918). (A) Protoconch viewed from above. (B) Detail of first teleoconch whorls viewed from above. (C) Close-up showing sculpture on last whorl. (D) Jaw. (E) Close-up showing morphology of central and inner lateral teeth.(F) Central and lateral teeth viewed from above.(G) Inner and central marginal teeth viewed from above. Scale bars: AC, 200 µm; D, 100 µm; EF, 20 µm; G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5239551" httpUri="https://zenodo.org/record/5239551/files/figure.png" pageId="4" pageNumber="171">Fig. 2AC</figureCitation>
). Broadly conical with moderately elevated spire. Rather thin but solid, translucent. Periphery well rounded to slightly angulate; upper and basal sectors rounded. Umbilicus 8090% concealed by columellar reflection. Background colour horn to almost crème; no spiral bands visible; outer and inner lip colour whitish. Protoconch
<quantity id="4CD54CC9FFBF3C5AFF1EFA7BFF106B8A" box="[166,253,1449,1472]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="4" pageNumber="171" unit="mm" value="2.6">2.6 mm</quantity>
in diameter, comprising about 1.5 whorls, with weak radially elongated pustulations. Teleoconch with faint, regular axial growth lines, evenly distributed across whorls of shell and across whorl diameter. Angle of aperture 4560 degrees; outer lip thin, well rounded, well expanded, not or slightly reflected; basal node absent or weak. Parietal wall of inner lip inconspicuous. Average shell size 11.8±0.8 × 16.2±
<quantity id="4CD54CC9FFBF3C5AFF5BF9A4FED868C7" box="[227,309,1654,1677]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="171" unit="mm" value="0.9">0.9 mm</quantity>
(
<tableCitation id="C6AFD497FFBF3C5AFEFBF9A4FE7E68C7" box="[323,403,1654,1677]" captionStart="Table 1" captionStartId="33.[121,184,1071,1095]" captionText="Table 1. Shell measurements of the type series. Given are maximumminimum (mean±standard deviation) of N measured adult shells. Abbreviations used: HT = holotype." pageId="4" pageNumber="171">Table 1</tableCitation>
).
</paragraph>
<caption id="DF52B1A4FFBF3C5AFCFFFABAFB036B87" ID-DOI="http://doi.org/10.5281/zenodo.5239555" ID-Zenodo-Dep="5239555" httpUri="https://zenodo.org/record/5239555/files/figure.png" pageId="4" pageNumber="171" startId="4.[839,903,1384,1405]" targetBox="[864,1401,151,1362]" targetPageId="4">
<paragraph id="8B92E12CFFBF3C5AFCFFFABAFB036B87" blockId="4.[839,1467,1384,1486]" pageId="4" pageNumber="171">
Figure 4. Penial anatomy of
<taxonomicName id="4C2D9AAFFFBF3C5AFBD4FABBFAA66B34" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[1132,1355,1385,1406]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="171" phylum="Mollusca" rank="species" species="atlantis" status="sp. nov.">
<emphasis id="B9593D3EFFBF3C5AFBD4FABBFAA66B34" box="[1132,1355,1385,1406]" italics="true" pageId="4" pageNumber="171">Amplirhagada atlantis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A8045FFBF3C5AFAECFABAFA6E6B37" box="[1364,1411,1384,1405]" pageId="4" pageNumber="171" rank="species">n.sp.</taxonomicNameLabel>
(AM C.472918). Abbreviations:
<emphasis id="B9593D3EFFBF3C5AFBEAFA51FB826BD2" box="[1106,1135,1411,1432]" italics="true" pageId="4" pageNumber="171">mp</emphasis>
, main pilaster;
<emphasis id="B9593D3EFFBF3C5AFAB6FA51FACE6BD2" box="[1294,1315,1411,1432]" italics="true" pageId="4" pageNumber="171">ps</emphasis>
, penial sheath;
<emphasis id="B9593D3EFFBF3C5AFCFFFA4CFCB26BF9" box="[839,863,1438,1459]" italics="true" pageId="4" pageNumber="171">pu</emphasis>
, penis wall pustulation;
<emphasis id="B9593D3EFFBF3C5AFBEBFA4CFB876BF9" box="[1107,1130,1438,1459]" italics="true" pageId="4" pageNumber="171">pv</emphasis>
, penial verge;
<emphasis id="B9593D3EFFBF3C5AFB46FA4CFAF46BF9" box="[1278,1305,1438,1459]" italics="true" pageId="4" pageNumber="171">rm</emphasis>
, penial retractor muscle;
<emphasis id="B9593D3EFFBF3C5AFC20FA6BFC426B84" box="[920,943,1465,1486]" italics="true" pageId="4" pageNumber="171">vd</emphasis>
, vas deferens. Scale bar = 3 mm.
</paragraph>
</caption>
<paragraph id="8B92E12CFFBF3C5AFF7DF941FE6B68AF" blockId="4.[166,794,1273,2000]" pageId="4" pageNumber="171">
<emphasis id="B9593D3EFFBF3C5AFF7DF941FE1E68E0" box="[197,499,1683,1706]" italics="true" pageId="4" pageNumber="171">Radular and jaw morphology</emphasis>
(
<figureCitation id="1316FDA9FFBF3C5AFE47F941FD9A68E0" box="[511,631,1683,1706]" captionStart="Figure 2" captionStartId="3.[121,184,1515,1536]" captionTargetBox="[121,1421,151,1498]" captionTargetId="figure-264@3.[121,1421,151,1498]" captionTargetPageId="3" captionText="Figure 2. Amplirhagada atlantis n.sp. SEM of shell (AM C.472919), radula and jaw (AM C.472918). (A) Protoconch viewed from above. (B) Detail of first teleoconch whorls viewed from above. (C) Close-up showing sculpture on last whorl. (D) Jaw. (E) Close-up showing morphology of central and inner lateral teeth.(F) Central and lateral teeth viewed from above.(G) Inner and central marginal teeth viewed from above. Scale bars: AC, 200 µm; D, 100 µm; EF, 20 µm; G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5239551" httpUri="https://zenodo.org/record/5239551/files/figure.png" pageId="4" pageNumber="171">Fig. 2 DG</figureCitation>
). Tooth formula C + 14 + 4 +?; average number of rows of teeth 130 (n = 2). Jaw with nine plates.
</paragraph>
<paragraph id="8B92E12CFFBF3C5AFF7DF939FA8D69DE" blockId="4.[166,794,1273,2000]" lastBlockId="4.[839,1468,1594,1999]" pageId="4" pageNumber="171">
<emphasis id="B9593D3EFFBF3C5AFF7DF939FE736948" box="[197,414,1771,1794]" italics="true" pageId="4" pageNumber="171">Genital morphology</emphasis>
(
<figureCitation id="1316FDA9FFBF3C5AFE15F939FDFD6948" box="[429,528,1771,1794]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStartId-0="4.[166,228,552,573]" captionStartId-1="4.[839,903,1384,1405]" captionTargetBox-0="[170,786,160,527]" captionTargetBox-1="[864,1401,151,1362]" captionTargetId-0="figure-58@4.[166,793,151,535]" captionTargetId-1="figure-102@4.[839,1466,148,1373]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionText-0="Figure 3. Genitalia of Amplirhagada atlantis n.sp. (AM C.472918). Abbreviations: ag, albumen gland; bc, bursa copulatrix; fo, free oviduct; hd, hermaphroditic duct; p, penis; rm, penial retractor muscle; so, sperm oviduct; va, vagina; vd, vas deferens. Scale bar = 5 mm." captionText-1="Figure 4. Penial anatomy of Amplirhagada atlantis n.sp. (AM C.472918). Abbreviations: mp, main pilaster; ps, penial sheath; pu, penis wall pustulation; pv, penial verge; rm, penial retractor muscle; vd, vas deferens. Scale bar = 3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5239553" figureDoi-1="http://doi.org/10.5281/zenodo.5239555" httpUri-0="https://zenodo.org/record/5239553/files/figure.png" httpUri-1="https://zenodo.org/record/5239555/files/figure.png" pageId="4" pageNumber="171">Figs 34</figureCitation>
). Penis straight, more or less of same length as anterior part of oviduct. Penial sheath delicate. Length of penial retractor muscle equivalent to about ¼ of length of penial complex. Penial verge extending about ¼ of length of penial chamber, slender with rounded tip. Almost entire inner penial wall covered by strongly developed pustules, which are arranged to form several corrugated longitudinal pilasters. Main stimulatory pilaster well developed, elongate, comprising proximal third of inner penial wall; sculptured by smooth lateral ridges being formed by fused and flattened pustules. Vas deferens entering penial sheath near apex of penial complex. Vagina long, tubular, posteriorly slightly inflated; inner vaginal wall and inner wall of spermathecal duct support continuous, welldeveloped, smooth longitudinal pilasters. Bursa copulatrix short, reaching base of spermoviduct; head elongately inflated, connected with spermoviduct by connective tissue, wall of head delicate, smooth. Free oviduct rather straight comprising less than half of length of anterior part of oviduct. Spermoviduct longer than anterior part of oviduct.
</paragraph>
<paragraph id="8B92E12CFFBF3C5AFCDDF849FB736985" blockId="4.[839,1468,1594,1999]" pageId="4" pageNumber="171">
<emphasis id="B9593D3EFFBF3C5AFCDDF849FBBE69F8" box="[869,1107,1947,1970]" italics="true" pageId="4" pageNumber="171">Aestivation strategy</emphasis>
. Free or rock sealer, in talus throughout vine thicket patches.
</paragraph>
</subSubSection>
<caption id="DF52B1A4FFBE3C5BFFC1F9C4FD8C6831" ID-DOI="http://doi.org/10.5281/zenodo.5239557" ID-Zenodo-Dep="5239557" httpUri="https://zenodo.org/record/5239557/files/figure.png" pageId="5" pageNumber="172" startId="5.[121,184,1558,1579]" targetBox="[121,1421,151,1538]" targetPageId="5">
<paragraph id="8B92E12CFFBE3C5BFFC1F9C4FD8C6831" blockId="5.[120,1422,1558,1660]" pageId="5" pageNumber="172">
Figure 5.
<taxonomicName id="4C2D9AAFFFBE3C5BFF6FF9C5FE396866" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[215,468,1559,1580]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="172" phylum="Mollusca" rank="species" species="carsoniana" status="sp. nov.">
<emphasis id="B9593D3EFFBE3C5BFF6FF9C5FE396866" box="[215,468,1559,1580]" italics="true" pageId="5" pageNumber="172">Amplirhagada carsoniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A8045FFBE3C5BFE62F9C4FDE56861" box="[474,520,1558,1579]" pageId="5" pageNumber="172" rank="species">n.sp.</taxonomicNameLabel>
SEM of shell (AM C.472920), radula and jaw (WAM
<accessionNumber id="947E7CCFFFBE3C5BFBA3F9C4FB856861" box="[1051,1128,1558,1579]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34708" pageId="5" pageNumber="172" type="EnaNcbi">S34708</accessionNumber>
).
<emphasis id="B9593D3EFFBE3C5BFBC1F9C5FB756866" box="[1145,1176,1559,1580]" italics="true" pageId="5" pageNumber="172">(A)</emphasis>
Protoconch viewed from above.
<emphasis id="B9593D3EFFBE3C5BFF05F9E3FF31680C" box="[189,220,1585,1606]" italics="true" pageId="5" pageNumber="172">(B)</emphasis>
Detail of first teleoconch whorls viewed from above.
<emphasis id="B9593D3EFFBE3C5BFD5BF9E3FCEE680C" box="[739,771,1585,1606]" italics="true" pageId="5" pageNumber="172">(C)</emphasis>
First teleoconch whorls viewed obliquely from above.
<emphasis id="B9593D3EFFBE3C5BFAAFF9E3FAD5680C" box="[1303,1336,1585,1606]" italics="true" pageId="5" pageNumber="172">(D)</emphasis>
Jaw.
<emphasis id="B9593D3EFFBE3C5BFAD6F9E3FA60680C" box="[1390,1421,1585,1606]" italics="true" pageId="5" pageNumber="172">(E)</emphasis>
Central and lateral teeth viewed from above.
<emphasis id="B9593D3EFFBE3C5BFD88F99EFDA2682B" box="[560,591,1612,1633]" italics="true" pageId="5" pageNumber="172">(F)</emphasis>
Close-up of marginal teeth.
<emphasis id="B9593D3EFFBE3C5BFCDEF99EFC6A682B" box="[870,903,1612,1633]" italics="true" pageId="5" pageNumber="172">(G)</emphasis>
Inner and central marginal teeth viewed from above. Scale bars:
<emphasis id="B9593D3EFFBE3C5BFF5FF9B5FEFF6836" box="[231,274,1639,1660]" italics="true" pageId="5" pageNumber="172">AC</emphasis>
, 200 µm;
<emphasis id="B9593D3EFFBE3C5BFECBF9B5FE696836" box="[371,388,1639,1660]" italics="true" pageId="5" pageNumber="172">D</emphasis>
, 100 µm;
<emphasis id="B9593D3EFFBE3C5BFE5EF9B5FDFF6836" box="[486,530,1639,1660]" italics="true" pageId="5" pageNumber="172">EG</emphasis>
, 10 µm.
</paragraph>
</caption>
<subSubSection id="C337B2A7FFBE3C5BFFC1F974FE696979" pageId="5" pageNumber="172" type="discussion">
<paragraph id="8B92E12CFFBE3C5BFFC1F974FE696979" blockId="5.[121,748,1702,1843]" pageId="5" pageNumber="172">
<emphasis id="B9593D3EFFBE3C5BFFC1F974FF0E68F4" bold="true" box="[121,227,1702,1726]" pageId="5" pageNumber="172">Remarks</emphasis>
. Anatomical description based on dissections of
<specimenCount id="9D2B2AA5FFBE3C5BFF23F916FF126891" box="[155,255,1732,1755]" pageId="5" pageNumber="172" type="generic" typeStatus="type">two type</specimenCount>
specimens. The non-type sample contains predominantly immature individuals not suitable for anatomical dissections. Listed by Solem (1991) as “
<taxonomicName id="4C2D9AAFFFBE3C5BFF3DF8CEFEB16979" authority="NSP" authorityName="NSP" box="[133,348,1820,1843]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="172" phylum="Mollusca" rank="genus">
<emphasis id="B9593D3EFFBE3C5BFF3DF8CEFEF26979" box="[133,287,1820,1843]" italics="true" pageId="5" pageNumber="172">Amplirhagada</emphasis>
NSP
</taxonomicName>
17”.
</paragraph>
</subSubSection>
</treatment>
</document>