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<document id="E335963738B57F9D3F6CC9896F948722" ID-DOI="10.21248/contrib.entomol.67.2.325-352" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742448" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638267809870" checkinUser="felipe" docAuthor="Zorn, Carsten, Kobayashi, Hirokazu &amp; Wada, Kaoru" docDate="2017" docId="03826012FFDEFFAD2B5900D1CE37F78B" docLanguage="en" docName="ContribEntomol.67.2.325-352.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Anomala asaitoae Zorn &amp; Kobayashi &amp; Wada 2017, spec. nov." docType="treatment" docUuid="829A3E69-938F-4F9C-998A-C4FE117C5740" docUuidSource="ZooBank" docVersion="3" lastPageNumber="335" masterDocId="FFBB186AFFD7FFA7280F027BCF12FFB3" masterDocTitle="Notes on the genus Anomala SAMOUELLE, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records" masterLastPageNumber="352" masterPageNumber="325" pageNumber="334" updateTime="1699436963292" updateUser="plazi">
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<mods:title id="F36548629496A8A07ED7D3DE1A6D3D16">Notes on the genus Anomala SAMOUELLE, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records</mods:title>
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<mods:namePart id="D9B8D5A66DB2CB392D7D768873B2EACE">Zorn, Carsten</mods:namePart>
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<mods:namePart id="D9178B2F511BAE10FD9812356C6BA67A">Kobayashi, Hirokazu</mods:namePart>
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<mods:namePart id="5AA5ACF34CAA44F77806DC5BA516042E">Wada, Kaoru</mods:namePart>
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<mods:title id="7802227B27F50DEA077DC6F53C6F9FF5">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:date id="1BF2D8E6033F77A497A15AC41671AEF2">2017</mods:date>
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<mods:number id="FB08A885DED86D37F26BF1BF211FD29B">67</mods:number>
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<treatment id="03826012FFDEFFAD2B5900D1CE37F78B" LSID="urn:lsid:zoobank.org:act:829A3E69-938F-4F9C-998A-C4FE117C5740" httpUri="http://treatment.plazi.org/id/03826012FFDEFFAD2B5900D1CE37F78B" lastPageId="10" lastPageNumber="335" pageId="9" pageNumber="334">
<subSubSection id="C331828FFFDEFFAE2B5900D1CA44FD55" pageId="9" pageNumber="334" type="nomenclature">
<paragraph id="8B94D104FFDEFFAE2B5900D1CB6DFD76" blockId="9.[854,1366,682,774]" box="[854,1151,682,709]" pageId="9" pageNumber="334">
<heading id="D0DC6668FFDEFFAE2B5900D1CB6DFD76" box="[854,1151,682,709]" fontSize="12" level="2" pageId="9" pageNumber="334" reason="2">
<taxonomicName id="4C2BAA87FFDEFFAE2B5900D1CB1DFD76" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[854,1039,682,709]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="334" phylum="Arthropoda" rank="species" species="asaitoae" status="sp. nov.">
<emphasis id="B95F0D16FFDEFFAE2B5900D1CB1DFD76" box="[854,1039,682,709]" italics="true" pageId="9" pageNumber="334">Anomala asaitoae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFDEFFAE2C1900D1CB6DFD76" box="[1046,1151,682,709]" pageId="9" pageNumber="334" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
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<subSubSection id="C331828FFFDEFFAE2B590089CCA9FCB5" box="[854,955,754,774]" pageId="9" pageNumber="334" type="description">
<paragraph id="8B94D104FFDEFFAE2B590089CCA9FCB5" blockId="9.[854,1366,682,774]" box="[854,955,754,774]" pageId="9" pageNumber="334">
(
<figureCitation id="1310CD81FFDEFFAE2B550089CC8EFCB5" box="[858,924,754,774]" captionStart="Figs 19" captionStartId="7.[174,214,2023,2045]" captionTargetBox="[214,1362,260,1985]" captionTargetId="figure-122@7.[203,1392,255,2011]" captionTargetPageId="7" captionText="Figs 19: Anomala spp., habitus 1. A. okushimai spec. nov., holotype. 2. A. immeliorata spec. nov., holotype. 3. A. confrater spec. nov., holotype.4. A. imperspicabilis spec.nov., holotype.5. A. ordinata spec.nov., holotype. 6. A. recordata spec.nov., holotype. 7. A. asaitoae spec. nov., holotype. 8. A. asaitoae spec. nov., paratype (female). 9. A. herbacea spec. nov., holotype. Scale = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5742452" httpUri="https://zenodo.org/record/5742452/files/figure.png" pageId="9" pageNumber="334">Figs 7, 8</figureCitation>
,
<figureCitation id="1310CD81FFDEFFAE2BAC0089CCA4FCB5" box="[931,950,754,774]" captionStart="Figs 1417" captionStartId="11.[174,214,2052,2074]" captionTargetBox="[179,1404,258,2008]" captionTargetId="figure-99@11.[174,1421,251,2026]" captionTargetPageId="11" captionText="Figs 1417: Anomala spp., aedeagus dorsal (a), ventral (b) and lateral (c). 14. A. ordinata spec. nov., holotype. 15. A. recordata" figureDoi="http://doi.org/10.5281/zenodo.5742456" httpUri="https://zenodo.org/record/5742456/files/figure.png" pageId="9" pageNumber="334">16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C331828FFFDEFFAE2B3F015BCA28FCCB" pageId="9" pageNumber="334" type="materials_examined">
<paragraph id="8B94D104FFDEFFAE2B3F015BCA28FCCB" blockId="9.[816,1421,799,888]" pageId="9" pageNumber="334">
<emphasis id="B95F0D16FFDEFFAE2B3F015BCCBFFC84" bold="true" box="[816,941,799,824]" pageId="9" pageNumber="334">Type series</emphasis>
:
<materialsCitation id="3B43DB59FFDEFFAE2BB1015BCBCEFCEB" collectionCode="NSMT" collectorName="A. Saito" location="Son La" pageId="9" pageNumber="334" specimenCount="1" stateProvince="Son La" typeStatus="holotype">
<emphasis id="B95F0D16FFDEFFAE2BB1015BCB36FC8B" bold="true" box="[958,1060,800,824]" pageId="9" pageNumber="334">
<typeStatus id="54906FA6FFDEFFAE2BB1015BCB36FC8B" box="[958,1060,800,824]" pageId="9" pageNumber="334" type="holotype">Holotype</typeStatus>
</emphasis>
: , Truong Yen,
<collectingRegion id="49EF1FE6FFDEFFAE2CE4015BCA2EFC84" box="[1259,1340,800,824]" country="Vietnam" name="Son La" pageId="9" pageNumber="334">Son La</collectingRegion>
, VIET- NAM, 1 v, 1995
<collectorName id="26DEB4D2FFDEFFAE2BFF013BCB56FCEB" box="[1008,1092,832,856]" pageId="9" pageNumber="334">A. Saito</collectorName>
leg. (
<collectionCode id="ED3A49C1FFDEFFAE2C89013BCBC7FCEB" box="[1158,1237,832,856]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="9" pageNumber="334">NSMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B43DB59FFDEFFAE2CE5013BCA24FCCB" collectionCode="HKPC" collectorName="A. Saito" location="Son La" pageId="9" pageNumber="334" specimenCount="1" stateProvince="Son La" typeStatus="paratype">
<typeStatus id="54906FA6FFDEFFAE2CE5013BCA46FCEB" box="[1258,1364,832,856]" pageId="9" pageNumber="334" type="paratype">Paratypes</typeStatus>
: 1 , 2 , the same data as for the holotype (
<collectionCode id="ED3A49C1FFDEFFAE2CEE011BCA3CFCCB" box="[1249,1326,864,888]" pageId="9" pageNumber="334">HKPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C331828FFFDEFFAD2B3F01E4CC02FA4B" lastPageId="10" lastPageNumber="335" pageId="9" pageNumber="334" type="description">
<paragraph id="8B94D104FFDEFFAE2B3F01E4CBFEFC64" blockId="9.[816,1422,927,2104]" pageId="9" pageNumber="334">
<emphasis id="B95F0D16FFDEFFAE2B3F01E4CCA0FC04" bold="true" box="[816,946,927,951]" pageId="9" pageNumber="334">Description</emphasis>
: Body shape elongate ovoid, convex; body length: 14.0-16.0 mm, width: 7.0-
<quantity id="4CD37CE1FFDEFFAE2C9C01BBCBF5FC64" box="[1171,1255,960,984]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="9" pageNumber="334" unit="mm" value="8.3">8.3 mm</quantity>
.
</paragraph>
<paragraph id="8B94D104FFDEFFAE2B3F01A4CCC8FB0B" blockId="9.[816,1422,927,2104]" pageId="9" pageNumber="334">
<emphasis id="B95F0D16FFDEFFAE2B3F01A4CC7DFC44" bold="true" box="[816,879,991,1015]" pageId="9" pageNumber="334">Color</emphasis>
. Dorsal surface except antennae black; head and legs with faint bluish-green metallic luster; pronotum, elytra and pygidium with iridescent shine, antennal club dark brown, footstalk reddish brown; elytra with a small orange spot before the middle on each second primary costa; ventral face black with greenish shine and iridescent reflections.
</paragraph>
<paragraph id="8B94D104FFDEFFAE2B3F06BBCA08F9A4" blockId="9.[816,1422,927,2104]" pageId="9" pageNumber="334">
<emphasis id="B95F0D16FFDEFFAE2B3F06BBCC78FB6B" bold="true" box="[816,874,1216,1240]" pageId="9" pageNumber="334">Head</emphasis>
with sparse erect pale yellow setae (ca.
<quantity id="4CD37CE1FFDEFFAE2D1106BBCA7DFB64" box="[1310,1391,1216,1240]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="334" unit="mm" value="0.4">0.4 mm</quantity>
in length) along eyes; interocular distance equals 0.69 times the maximum transverse head width. Antennal club 0.8 times as long as footstalk; antennomeres 8 and 9 somewhat thickened. Clypeus subtrapezoidal with broadly rounded corners and curved anterior margin; surface rugoso-punctate; anterior margin distinctly reflexed; fronto-clypeal suture weakly sinuate. Frons confluently rugoso-punctate in the middle; the punctures becoming sparser and separate laterad and posteriad. Vertex with sparse, small, somewhat transverse punctures.
</paragraph>
<paragraph id="8B94D104FFDEFFAE2B3F045BCB6EF82B" blockId="9.[816,1422,927,2104]" pageId="9" pageNumber="334">
<emphasis id="B95F0D16FFDEFFAE2B3F045BCCB3F98B" bold="true" box="[816,929,1568,1592]" pageId="9" pageNumber="334">Pronotum</emphasis>
1.6 times as wide as long; sides evenly arcuate, subparallel basad, convergent anteriorly; widest at base; lateral margin hidden in dorsal view before the middle by a small protuberance; lateral marginal line almost reaching the hind angles; basal marginal line very weakly indicated medially of humerus; front angles produced and acute, hind angles obtuse, rounded off; surface evenly covered with transverse moderately large punctures; punctures larger and partly confluent towards the front angles, somewhat smaller on disc; each puncture bearing a microscopic seta; with erect yellow setae (
<quantity id="4CD37CE1FFDEFFAE2D0B051BCA9FF8C4" box="[1284,1421,1888,1912]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="9" pageNumber="334" unit="mm" value="0.375" valueMax="0.4" valueMin="0.35">0.35-0.4 mm</quantity>
in length) along lateral margin.
</paragraph>
<paragraph id="8B94D104FFDEFFAE2B3F05E4CBA3F864" blockId="9.[816,1422,927,2104]" pageId="9" pageNumber="334">
<emphasis id="B95F0D16FFDEFFAE2B3F05E4CC8CF804" bold="true" box="[816,926,1951,1975]" pageId="9" pageNumber="334">Scutellum</emphasis>
1.6 times as wide as long, apex rather acute; with irregular, transverse punctures.
</paragraph>
<paragraph id="8B94D104FFDEFFAD2B3F059BCDC0FDA4" blockId="9.[816,1422,927,2104]" lastBlockId="10.[212,817,256,1368]" lastPageId="10" lastPageNumber="335" pageId="9" pageNumber="334">
<emphasis id="B95F0D16FFDEFFAE2B3F059BCC63F84B" bold="true" box="[816,881,2016,2040]" pageId="9" pageNumber="334">Elytra</emphasis>
regularly striate-sulcate; primary costae slightly more convex than interstices; 5th primary costa narrow and slightly more elevated than the others; striae not marked by a regular row of punctures; entire surface covered with large, transverse, microscopic-seta-bearing punctures, concentrating in the sulci, leaving out the ridges of primary costae and being sparser in the interstices; punctures adjacent to each other, partly coalescing and forming small transverse striae; humerus with rather simple punctation; epipleura strongly developed, abruptly ending at the level of second abdominal ventrite; marginal membrane starting approximately at level of middle of metacoxa.
</paragraph>
<paragraph id="8B94D104FFDDFFAD28DB0064CC0AFD0B" blockId="10.[212,817,256,1368]" pageId="10" pageNumber="335">
<emphasis id="B95F0D16FFDDFFAD28DB0064CE52FD84" bold="true" box="[212,320,543,567]" pageId="10" pageNumber="335">Pygidium</emphasis>
triangular, rounded at apex, somewhat tumid; disc with reticulate striolation which is arranged concentrically around the tumidity; with evenly distributed microscopic setae; longer sparse yellow setae (
<quantity id="4CD37CE1FFDDFFAD2AF600FBCE20FD04" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="7.5" metricValueMin="2.5" pageId="10" pageNumber="335" unit="mm" value="0.5" valueMax="0.75" valueMin="0.25">0.25- 0.75 mm</quantity>
in length) in apical and baso-lateral portions.
</paragraph>
<paragraph id="8B94D104FFDDFFAD28DB00C4CD62FD44" blockId="10.[212,817,256,1368]" pageId="10" pageNumber="335">
<emphasis id="B95F0D16FFDDFFAD28DB00C4CE71FD64" bold="true" box="[212,355,703,727]" pageId="10" pageNumber="335">Mesoventrite</emphasis>
very narrow and somewhat concave between the approximated mesocoxae.
</paragraph>
<paragraph id="8B94D104FFDDFFAD28DB0084CDCEFCCB" blockId="10.[212,817,256,1368]" pageId="10" pageNumber="335">
<emphasis id="B95F0D16FFDDFFAD28DB0084CE72FCA4" bold="true" box="[212,352,767,791]" pageId="10" pageNumber="335">Metaventrite</emphasis>
with dense, transversely confluent punctures at the sides, sparsely punctate in the middle; with pale yellow setae (
<quantity id="4CD37CE1FFDDFFAD29AD013BCD3EFCE4" box="[418,556,832,856]" metricMagnitude="-4" metricUnit="m" metricValue="3.15" metricValueMax="5.0" metricValueMin="1.3" pageId="10" pageNumber="335" unit="mm" value="0.315" valueMax="0.5" valueMin="0.13">0.13-0.5 mm</quantity>
in length) at the sides which become sparser and shorter in the middle.
</paragraph>
<paragraph id="8B94D104FFDDFFAD28DB0104CEC9FB8B" blockId="10.[212,817,256,1368]" pageId="10" pageNumber="335">
<emphasis id="B95F0D16FFDDFFAD28DB0104CEACFC24" bold="true" box="[212,446,895,919]" pageId="10" pageNumber="335">Abdominal ventrites</emphasis>
with very transverse punctures which are partly confluent at the sides and sparser in the middle; each puncture with microscopic seta; each ventrite with a transverse row of semierect yellow setae (
<quantity id="4CD37CE1FFDDFFAD2917067BCEBDFBA4" box="[280,431,1024,1048]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" metricValueMax="6.3" metricValueMin="1.3" pageId="10" pageNumber="335" unit="mm" value="0.38" valueMax="0.63" valueMin="0.13">0.13-0.63 mm</quantity>
in length); sides of first abdominal ventrite slightly carinate.
</paragraph>
<paragraph id="8B94D104FFDDFFAD28DB063BCDE6FAE4" blockId="10.[212,817,256,1368]" pageId="10" pageNumber="335">
<emphasis id="B95F0D16FFDDFFAD28DB063BCE17FBEB" bold="true" box="[212,261,1088,1112]" pageId="10" pageNumber="335">Legs</emphasis>
slender and long; protibia bidentate; terminal tooth prolonged and acute at apex in male; protarsal segment 5 thickened, ventrally with a little tooth approximately in the middle and a deep concavity in the apical half; inner protarsal claw and outer mesotarsal claw apically incised; the upper branch of the inner protarsal claw small and slender, the lower branch broad, somewhat enlarged ventrally with little tooth; the upper branch of outer mesotarsal claw slightly more slender than the lower one.
</paragraph>
<paragraph id="8B94D104FFDDFFAD28DB07FBCC02FA4B" blockId="10.[212,817,1408,1528]" pageId="10" pageNumber="335">
<emphasis id="B95F0D16FFDDFFAD28DB07FBCE39FA2B" bold="true" box="[212,299,1408,1432]" pageId="10" pageNumber="335">Females</emphasis>
: Body shape stouter; antennal club not thickened; pronotum broader, more convex; lateral carina of elytra broadened behind middle; pygidium less tumid; terminal tooth of protibia spatulate; inner protarsal claw slender.
</paragraph>
</subSubSection>
<subSubSection id="C331828FFFDDFFAD28DB0464CD6EF82B" pageId="10" pageNumber="335" type="diagnosis">
<paragraph id="8B94D104FFDDFFAD28DB0464CD6EF82B" blockId="10.[212,817,1567,1944]" pageId="10" pageNumber="335">
<emphasis id="B95F0D16FFDDFFAD28DB0464CE53F984" bold="true" box="[212,321,1567,1591]" pageId="10" pageNumber="335">Diagnosis</emphasis>
: Like the previous species,
<taxonomicName id="4C2BAA87FFDDFFAD2A6B045BCDCAF984" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[612,728,1568,1591]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="asaitoae">
<emphasis id="B95F0D16FFDDFFAD2A6B045BCDCAF984" box="[612,728,1568,1591]" italics="true" pageId="10" pageNumber="335">A. asaitoae</emphasis>
</taxonomicName>
belongs to the group containing
<taxonomicName id="4C2BAA87FFDDFFAD29DB043BCD45F9EB" authorityName="BURMEISTER" authorityYear="1855" box="[468,599,1600,1624]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="spiloptera">
<emphasis id="B95F0D16FFDDFFAD29DB043BCD45F9EB" box="[468,599,1600,1624]" italics="true" pageId="10" pageNumber="335">A. spiloptera</emphasis>
</taxonomicName>
, recognizable by the peculiar dorsal connective membrane between the parameres (for details see “Diagnosis” of
<taxonomicName id="4C2BAA87FFDDFFAD2A5F04FBCDC2F92B" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[592,720,1664,1688]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="recordata">
<emphasis id="B95F0D16FFDDFFAD2A5F04FBCDC2F92B" box="[592,720,1664,1688]" italics="true" pageId="10" pageNumber="335">A. recordata</emphasis>
</taxonomicName>
). Within this group,
<taxonomicName id="4C2BAA87FFDDFFAD295B04DBCEDAF904" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[340,456,1696,1719]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="asaitoae">
<emphasis id="B95F0D16FFDDFFAD295B04DBCEDAF904" box="[340,456,1696,1719]" italics="true" pageId="10" pageNumber="335">A. asaitoae</emphasis>
</taxonomicName>
is differentiated by the enlarged two last antennal segments in males, the carinate sides of the first abdominal ventrite, the enlarged lateral margin of the elytra, the small body size and the unique, asymmetric shape of the aedeagus with spine-like apices of the parameres. The ventral plate has no apical process (fig. 16). None of the other species related to
<taxonomicName id="4C2BAA87FFDDFFAD2AA2051BCC22F8CB" authorityName="BURMEISTER" authorityYear="1855" box="[685,816,1888,1912]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="spiloptera">
<emphasis id="B95F0D16FFDDFFAD2AA2051BCC22F8CB" box="[685,816,1888,1912]" italics="true" pageId="10" pageNumber="335">A. spiloptera</emphasis>
</taxonomicName>
has a similar combination of characters.
</paragraph>
</subSubSection>
<subSubSection id="C331828FFFDDFFAD28DB05BBCE37F78B" pageId="10" pageNumber="335" type="etymology">
<paragraph id="8B94D104FFDDFFAD28DB05BBCE37F78B" blockId="10.[212,817,1984,2104]" pageId="10" pageNumber="335">
<emphasis id="B95F0D16FFDDFFAD28DB05BBCE59F86B" bold="true" box="[212,331,1984,2008]" pageId="10" pageNumber="335">Etymology</emphasis>
: The new species is named after Dr. Akiko Saito, entomologist at the Natural History Museum and Institute,
<collectingRegion id="49EF1FE6FFDDFFAD29340A7BCE6CF7AB" box="[315,382,2048,2072]" country="Japan" name="Chiba" pageId="10" pageNumber="335">Chiba</collectingRegion>
, who collected the
<typeStatus id="54906FA6FFDDFFAD2A5A0A7BCDA1F7AB" box="[597,691,2048,2072]" pageId="10" pageNumber="335" type="holotype">holotype</typeStatus>
of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>