283 lines
37 KiB
XML
283 lines
37 KiB
XML
<document id="CEF99E46F9CBF2192E839D5513E0CA03" ID-CLB-Dataset="187408" ID-DOI="10.1080/00222933.2022.2117107" ID-GBIF-Dataset="25e45e0a-f320-409b-8923-5ab29eabf122" ID-ISSN="1464-5262" ID-Zenodo-Dep="7318178" ID-ZooBank="A6F9B9FB-D3F9-49F2-8FC1-5E6A5E0AB3B1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1668419502508" checkinUser="plazi" docAuthor="Fachin, Diego Aguilar & Hauser, Martin" docDate="2022" docId="561187CDFFF332639B8A4DBFFDF4113C" docLanguage="en" docName="JNATHIST.56.37-40.1513-1562.pdf" docOrigin="Journal of Natural History 56 (37 - 40)" docSource="http://dx.doi.org/10.1080/00222933.2022.2117107" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Formosargus mangoleensis Fachin & Hauser 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="1555" masterDocId="AA28FFB5FFD932489A3F4D31FFA11374" masterDocTitle="Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym" masterLastPageNumber="1562" masterPageNumber="1513" pageNumber="1554" updateTime="1699411179050" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EF849E22FD2E5AE4D65AEA346A2FF423">Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym</mods:title>
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<mods:name id="94117F3366AE71ECAA5A9E70B82E85AD" type="personal">
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<mods:roleTerm id="42FDE71BBBCDBA2ADDCB37D413D2D337">Author</mods:roleTerm>
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<mods:namePart id="6F7BA90E421F3B0772361A95A2964249">Fachin, Diego Aguilar</mods:namePart>
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<mods:name id="E5AD343376441AE516B9F14DEA4C5D0F" type="personal">
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<mods:roleTerm id="5D879C145381B782F96A7DA8B5773504">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6C4BEF1D759F327F749D073DCFD81895">Hauser, Martin</mods:namePart>
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<mods:typeOfResource id="B16EDEBB95ADE039F8BB6CEA45696DB2">text</mods:typeOfResource>
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<mods:title id="04F870057FA5AA4435B14A85AA6626D7">Journal of Natural History</mods:title>
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<mods:part id="54036654E5D76AFD1C22EC617444705C">
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<mods:date id="585084098AE7047C88BAF3B914A6217F">2022</mods:date>
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<mods:number id="17736E45295BDA2F28228B27B7F26E72">2022-10-12</mods:number>
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<mods:detail id="902A848F8D8464C6886B0BD470C808B9" type="volume">
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<mods:number id="AF74DA953A43C4B8911628295FB27F77">56</mods:number>
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<mods:detail id="8ACB4F8114B678F5CA35A3C6CB320BF1" type="issue">
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<mods:number id="7092AC01E1FC4261C74D6CB44B6BA86A">37 - 40</mods:number>
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<mods:start id="87F8D50E65926E06329A23A3A515ED9B">1513</mods:start>
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<mods:url id="BD090F015B06D7453F8C67B4106DBC49">http://dx.doi.org/10.1080/00222933.2022.2117107</mods:url>
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<mods:classification id="FBC9AE191A54290AFBD50DD752A6A721">journal article</mods:classification>
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<mods:identifier id="53AFE49BEE74AB89B7FEFDB297C289FF" type="CLB-Dataset">187408</mods:identifier>
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<mods:identifier id="93C48CD04EFE6902486916F9F891ECDD" type="DOI">10.1080/00222933.2022.2117107</mods:identifier>
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<mods:identifier id="17214FFF6286211938B89A5CD3ADF6DB" type="ISSN">1464-5262</mods:identifier>
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<treatment id="561187CDFFF332639B8A4DBFFDF4113C" ID-DOI="http://doi.org/10.5281/zenodo.7324780" ID-GBIF-Taxon="205007493" ID-Zenodo-Dep="7324780" LSID="urn:lsid:plazi:treatment:561187CDFFF332639B8A4DBFFDF4113C" httpUri="http://treatment.plazi.org/id/561187CDFFF332639B8A4DBFFDF4113C" lastPageId="43" lastPageNumber="1555" pageId="42" pageNumber="1554">
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<subSubSection id="96A26550FFF332629B8A4DBFFCCF13DC" box="[437,878,142,168]" pageId="42" pageNumber="1554" type="nomenclature">
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<paragraph id="DE0736DBFFF332629B8A4DBFFCCF13DC" blockId="42.[437,878,142,168]" box="[437,878,142,168]" pageId="42" pageNumber="1554">
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<heading id="854F81B7FFF332629B8A4DBFFCCF13DC" bold="true" box="[437,878,142,168]" centered="true" fontSize="11" level="2" pageId="42" pageNumber="1554" reason="3">
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<emphasis id="ECCCEAC9FFF332629B8A4DBFFCCF13DC" bold="true" box="[437,878,142,168]" pageId="42" pageNumber="1554">
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<taxonomicName id="19B84D58FFF332629B8A4DBFFCA513DC" ID-CoL="BCY3B" authority="Fachin & Hauser, 2022" authorityName="Fachin & Hauser" authorityYear="2022" box="[437,772,142,168]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="1554" phylum="Arthropoda" rank="species" species="mangoleensis" status="sp. nov.">
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<emphasis id="ECCCEAC9FFF332629B8A4DBFFCA513DC" bold="true" box="[437,772,142,168]" italics="true" pageId="42" pageNumber="1554">Formosargus mangoleensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F7FF57B2FFF3326299334DBFFCCF13DC" box="[780,878,142,168]" pageId="42" pageNumber="1554" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="96A26550FFF332629B474DF0FC0D13AF" box="[376,940,193,219]" pageId="42" pageNumber="1554" type="description">
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<paragraph id="DE0736DBFFF332629B474DF0FC0D13AF" blockId="42.[376,940,193,219]" box="[376,940,193,219]" pageId="42" pageNumber="1554">
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||
(
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<figureCitation id="46832A5EFFF332629BBF4DF0FE4D13AF" box="[384,492,193,219]" captionStart="Figure 4" captionStartId="8.[160,225,1266,1289]" captionTargetBox="[167,1149,145,1232]" captionTargetId="figure-139@8.[163,1153,142,1234]" captionTargetPageId="8" captionText="Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318188" httpUri="https://zenodo.org/record/7318188/files/figure.png" pageId="42" pageNumber="1554">Figures 4</figureCitation>
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(c), 6(c), 7(g), 16(a), 17(i–j), 19(f), 20(a))
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</paragraph>
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</subSubSection>
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<subSubSection id="96A26550FFF332629A9F4C12FD3A129E" pageId="42" pageNumber="1554" type="diagnosis">
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<paragraph id="DE0736DBFFF332629A9F4C12FEDC1249" blockId="42.[160,1156,291,490]" box="[160,381,291,317]" pageId="42" pageNumber="1554">
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<emphasis id="ECCCEAC9FFF332629A9F4C12FEDC1249" bold="true" box="[160,381,291,317]" italics="true" pageId="42" pageNumber="1554">Diagnosis (female)</emphasis>
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</paragraph>
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<paragraph id="DE0736DBFFF332629A9F4C76FD3A129E" blockId="42.[160,1156,291,490]" pageId="42" pageNumber="1554">
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Medial band on scutum narrow, not reaching the extreme anterior end of scutum (
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<figureCitation id="46832A5EFFF332629E034C76FF0E12F6" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="42" pageNumber="1554">Figure 7</figureCitation>
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(g)), wider posteriorly. Vertex at least twice the length of the ocellar tubercle (
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<figureCitation id="46832A5EFFF332629E3E4C58FBFA12F6" box="[1025,1115,361,386]" captionStart="Figure 4" captionStartId="8.[160,225,1266,1289]" captionTargetBox="[167,1149,145,1232]" captionTargetId="figure-139@8.[163,1153,142,1234]" captionTargetPageId="8" captionText="Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318188" httpUri="https://zenodo.org/record/7318188/files/figure.png" pageId="42" pageNumber="1554">Figure 4</figureCitation>
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(c)); upper frons shortened, nearly three times shorter than lower frons (
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<figureCitation id="46832A5EFFF3326299CB4CBDFBFB12D1" box="[1012,1114,396,421]" captionStart="Figure 6" captionStartId="10.[160,225,1285,1308]" captionTargetBox="[167,1149,145,1251]" captionTargetId="figure-99@10.[163,1153,142,1253]" captionTargetPageId="10" captionText="Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi="http://doi.org/10.5281/zenodo.7318193" httpUri="https://zenodo.org/record/7318193/files/figure.png" pageId="42" pageNumber="1554">Figure 6</figureCitation>
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(c)). Mediotergite completely yellow; anepisternum with a small subrectangular dark spot (
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<figureCitation id="46832A5EFFF332629A974CE0FEB3129E" box="[168,274,465,490]" captionStart="Figure 20" captionStartId="32.[160,225,1630,1653]" captionTargetBox="[171,1149,154,1593]" captionTargetId="figure-3@32.[163,1153,142,1598]" captionTargetPageId="32" captionText="Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively." figureDoi="http://doi.org/10.5281/zenodo.7318228" httpUri="https://zenodo.org/record/7318228/files/figure.png" pageId="42" pageNumber="1554">Figure 20</figureCitation>
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(a)). Abdomen entirely pale yellow.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="96A26550FFF332629A9F4F13FCE4119D" pageId="42" pageNumber="1554" type="materials_examined">
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||
<paragraph id="DE0736DBFFF332629A9F4F13FEDE1148" blockId="42.[160,1156,546,745]" box="[160,383,546,572]" pageId="42" pageNumber="1554">
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||
<emphasis id="ECCCEAC9FFF332629A9F4F13FEDE1148" bold="true" box="[160,383,546,572]" italics="true" pageId="42" pageNumber="1554">Material examined</emphasis>
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</paragraph>
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||
<paragraph id="DE0736DBFFF332629A9F4F74FCE4119D" blockId="42.[160,1156,546,745]" pageId="42" pageNumber="1554">
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||
<materialsCitation id="6ED03C86FFF332629A9F4F74FB2111D0" ID-GBIF-Occurrence="3965743330" collectingDate="1995-03-07" collectingDateMax="1995-03-24" collectingDateMin="1995-03-07" collectionCode="RMNH" collectorName="C. v. Achterberg & Y. Yasir" country="Indonesia" elevation="70" location="Mangole" municipality="Sula Isl." pageId="42" pageNumber="1554" specimenCount="1" specimenCount-female="1" stateProvince="North Maluku Province" typeStatus="holotype">
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||
<emphasis id="ECCCEAC9FFF332629A9F4F74FE82112A" bold="true" box="[160,291,581,606]" pageId="42" pageNumber="1554">
|
||
<typeStatus id="01038879FFF332629A9F4F74FE82112A" box="[160,291,581,606]" pageId="42" pageNumber="1554" type="holotype">HOLOTYPE</typeStatus>
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||
</emphasis>
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||
,
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||
<specimenCount id="C8BEFD52FFF332629B104F74FEF61129" box="[303,343,581,606]" pageId="42" pageNumber="1554" type="female">1 ♀</specimenCount>
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||
(genitalia dissected and stored in microvial with glycerine),
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<emphasis id="ECCCEAC9FFF3326299C94F74FBDF112A" bold="true" box="[1014,1150,581,606]" pageId="42" pageNumber="1554">
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<collectingCountry id="A6AF764BFFF3326299C94F74FBDF112A" box="[1014,1150,581,606]" name="Indonesia" pageId="42" pageNumber="1554">INDONESIA</collectingCountry>
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</emphasis>
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, [
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<collectingRegion id="1C7CF839FFF332629A9A4F59FE1C11F5" box="[165,445,616,641]" country="Indonesia" name="Maluku Utara" pageId="42" pageNumber="1554">North Maluku Province</collectingRegion>
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],
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<collectingRegion id="1C7CF839FFF332629BEE4F59FD8B11F5" box="[465,554,616,641]" pageId="42" pageNumber="1554">Sula Isl.</collectingRegion>
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[
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<collectingCountry id="A6AF764BFFF3326298044F59FD2211F5" box="[571,643,616,641]" name="Iceland" pageId="42" pageNumber="1554">Island</collectingCountry>
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]
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<location id="DB676000FFF3326298AB4F59FD5B11F5" LSID="urn:lsid:plazi:treatment:561187CDFFF332639B8A4DBFFDF4113C:DB676000FFF3326298AB4F59FD5B11F5" box="[660,762,616,641]" country="Indonesia" municipality="Sula Isl." name="Mangole" pageId="42" pageNumber="1554" stateProvince="North Maluku Province">Mangole</location>
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,
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<locationDeviation id="CFB52FF9FFF3326299364F59FBE411F5" box="[777,1093,616,641]" pageId="42" pageNumber="1554">
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<quantity id="19409B3EFFF3326299364F59FCE711F5" box="[777,838,616,641]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="1554" unit="km" value="2.0">2 km</quantity>
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||
W. Mandafuhi Camp
|
||
</locationDeviation>
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, Mal. [
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<location id="DB676000FFF332629A9A4FBAFEA111D0" LSID="urn:lsid:plazi:treatment:561187CDFFF332639B8A4DBFFDF4113C:DB676000FFF332629A9A4FBAFEA111D0" box="[165,256,651,676]" country="Indonesia" municipality="Sula Isl." name="Malaise" pageId="42" pageNumber="1554" stateProvince="North Maluku Province">Malaise</location>
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||
] trap 19, c
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<quantity id="19409B3EFFF332629BAF4FBAFE7111D0" box="[400,464,651,676]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="42" pageNumber="1554" unit="m" value="70.0">
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<elevation id="5595D1E8FFF332629BAF4FBAFE7111D0" box="[400,464,651,676]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="42" pageNumber="1554" unit="m" value="70.0">70 m</elevation>
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</quantity>
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,
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<date id="AA06101BFFF332629BDF4FBAFD1F11D0" box="[480,702,651,676]" pageId="42" pageNumber="1554" value="1995-03-07" valueMax="1995-03-24" valueMin="1995-03-07">
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<collectingDate id="BA42E9F3FFF332629BDF4FBAFD1F11D0" box="[480,702,651,676]" pageId="42" pageNumber="1554" value="1995-03-07" valueMax="1995-03-24" valueMin="1995-03-07">07–24 March 1995</collectingDate>
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</date>
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,
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||
<collectorName id="734D530DFFF3326298F24FBAFC2011D0" box="[717,897,651,676]" pageId="42" pageNumber="1554">C.v. Achterberg</collectorName>
|
||
and
|
||
<collectorName id="734D530DFFF3326299FB4FBAFBBA11D0" box="[964,1051,651,676]" pageId="42" pageNumber="1554">Y. Yasir</collectorName>
|
||
(
|
||
<collectionCode id="B8A9AE1EFFF332629E134FBAFBDB11D0" box="[1068,1146,651,676]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="42" pageNumber="1554" type="Museum">RMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="6ED03C86FFF332629A9F4F9CFCE0119D" ID-GBIF-Occurrence="3965743318" collectingDate="1995-03-05" collectionCode="RMNH" collectorName="C. v. Achterberg" country="Indonesia" location="Mandafuhi Camp" municipality="Mangole" pageId="42" pageNumber="1554" specimenCount="1" specimenCount-female="1" stateProvince="North Maluku Province" typeStatus="paratype">
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||
<emphasis id="ECCCEAC9FFF332629A9F4F9CFEBF11B2" bold="true" box="[160,286,685,710]" pageId="42" pageNumber="1554">
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<typeStatus id="01038879FFF332629A9F4F9CFEBF11B2" box="[160,286,685,710]" pageId="42" pageNumber="1554" type="paratype">PARATYPE</typeStatus>
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||
</emphasis>
|
||
,
|
||
<specimenCount id="C8BEFD52FFF332629B114F9CFEF811B0" box="[302,345,685,710]" pageId="42" pageNumber="1554" type="female">1 ♀</specimenCount>
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,
|
||
<emphasis id="ECCCEAC9FFF332629B574F9CFE5111B2" bold="true" box="[360,496,685,710]" pageId="42" pageNumber="1554">
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<collectingCountry id="A6AF764BFFF332629B574F9CFE5111B2" box="[360,496,685,710]" name="Indonesia" pageId="42" pageNumber="1554">INDONESIA</collectingCountry>
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</emphasis>
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, [
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||
<collectingRegion id="1C7CF839FFF33262983B4F9CFCB311B2" box="[516,786,685,710]" country="Indonesia" name="Maluku Utara" pageId="42" pageNumber="1554">North Maluku Province</collectingRegion>
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],
|
||
<collectingMunicipality id="3E63ACA1FFF33262991B4F9CFC2811B2" box="[804,905,685,710]" pageId="42" pageNumber="1554">Mangole</collectingMunicipality>
|
||
,
|
||
<location id="DB676000FFF3326299AA4F9CFBFA11B2" LSID="urn:lsid:plazi:treatment:561187CDFFF332639B8A4DBFFDF4113C:DB676000FFF3326299AA4F9CFBFA11B2" box="[917,1115,685,710]" country="Indonesia" municipality="Mangole" name="Mandafuhi Camp" pageId="42" pageNumber="1554" stateProvince="North Maluku Province">Mandafuhi Camp</location>
|
||
,
|
||
<date id="AA06101BFFF332629E574F9CFE87119D" pageId="42" pageNumber="1554" value="1995-03-05">
|
||
<collectingDate id="BA42E9F3FFF332629E574F9CFE87119D" pageId="42" pageNumber="1554" value="1995-03-05">05 March 1995</collectingDate>
|
||
</date>
|
||
, along road,
|
||
<collectorName id="734D530DFFF332629B874FE1FDC9119D" box="[440,616,720,745]" pageId="42" pageNumber="1554">C.v. Achterberg</collectorName>
|
||
,
|
||
<collectionCode id="B8A9AE1EFFF33262984D4FE1FD18119D" box="[626,697,720,745]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" pageId="42" pageNumber="1554">RMNH</collectionCode>
|
||
95’ (
|
||
<collectionCode id="B8A9AE1EFFF3326298D14FE1FC9A119D" box="[750,827,720,745]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="42" pageNumber="1554" type="Museum">RMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="96A26550FFF332629A9F4E11FEB216C2" pageId="42" pageNumber="1554" type="description">
|
||
<paragraph id="DE0736DBFFF332629A9F4E11FE87104E" blockId="42.[160,1156,800,1346]" box="[160,294,800,826]" pageId="42" pageNumber="1554">
|
||
<emphasis id="ECCCEAC9FFF332629A9F4E11FE87104E" bold="true" box="[160,294,800,826]" italics="true" pageId="42" pageNumber="1554">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF332629A9F4E75FC0D1636" blockId="42.[160,1156,800,1346]" pageId="42" pageNumber="1554">
|
||
<emphasis id="ECCCEAC9FFF332629A9F4E75FF561029" bold="true" box="[160,247,836,861]" pageId="42" pageNumber="1554">Female</emphasis>
|
||
. Length (n = 2): body, 10.0 mm; wing, 9.0 mm.
|
||
<emphasis id="ECCCEAC9FFF3326299224E75FCFA1029" bold="true" box="[797,859,836,861]" pageId="42" pageNumber="1554">Head</emphasis>
|
||
(
|
||
<figureCitation id="46832A5EFFF3326299534E75FC741029" box="[876,981,836,861]" captionStart="Figure 4" captionStartId="8.[160,225,1266,1289]" captionTargetBox="[167,1149,145,1232]" captionTargetId="figure-139@8.[163,1153,142,1234]" captionTargetPageId="8" captionText="Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318188" httpUri="https://zenodo.org/record/7318188/files/figure.png" pageId="42" pageNumber="1554">Figures 4</figureCitation>
|
||
(c), 6(c)). Vertex area twice longer than ocellar tubercle. Upper frons shortened, twice wider than long, nearly three times shorter lower frons; upper frons black lateral area nearly as broad as ocellar tubercle width, with short brownish pilosity; anterior margin nearly straight at the edge with lower frons, only medially pointing towards antennal insertion. Fourth flagellomere rounded, as wide as each previous flagellomere alone or slightly wider. Face three times shorter than lower frons.
|
||
<emphasis id="ECCCEAC9FFF3326298364925FDFC1759" bold="true" box="[521,605,1044,1069]" pageId="42" pageNumber="1554">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="46832A5EFFF3326298524925FD6D1759" box="[621,716,1044,1069]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="42" pageNumber="1554">Figure 7</figureCitation>
|
||
(g)). Medial dark brown band covering less than one-third of scutum width, not reaching extreme anterior end of scutum, wider on posterior half and scutellum; pleuron pale yellow, with only a small subrectangular dark brown spot on anepisternum.
|
||
<emphasis id="ECCCEAC9FFF332629801494DFDDD17E1" bold="true" box="[574,636,1148,1173]" pageId="42" pageNumber="1554">Wing</emphasis>
|
||
. R2+3 originating slightly before r–m, weakly curved on basal third, straight on apical two-thirds.
|
||
<emphasis id="ECCCEAC9FFF3326298E649AFFCEF17C3" bold="true" box="[729,846,1182,1207]" pageId="42" pageNumber="1554">Abdomen</emphasis>
|
||
. Completely yellow; pilosity yellow.
|
||
<emphasis id="ECCCEAC9FFF332629AC849F0FED717AE" bold="true" box="[247,374,1217,1242]" pageId="42" pageNumber="1554">Terminalia</emphasis>
|
||
(
|
||
<figureCitation id="46832A5EFFF332629BB949F0FE5C17AE" box="[390,509,1217,1242]" captionStart="Figure 16" captionStartId="28.[160,226,964,987]" captionTargetBox="[164,1149,144,929]" captionTargetId="figure-182@28.[162,1152,142,932]" captionTargetPageId="28" captionText="Figure 16. Female terminalia of Formosargus species, dorsal view. (a) F. mangoleensis sp. nov, holotype. (b) F. trivittatus sp. nov., paratype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite." figureDoi="http://doi.org/10.5281/zenodo.7318216" httpUri="https://zenodo.org/record/7318216/files/figure.png" pageId="42" pageNumber="1554">Figures 16</figureCitation>
|
||
(a), 17(i, j)). Genital fork wider at posterior half at level of posterior bridge; anterior half gradually narrowing into a long and curved anterior end; posterior bridge bilobed; posterolateral process parallel, slightly wider at base, gradually narrowing towards a rounded apex; genital opening roughly circular.
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF332629A9F4848FF7B16E7" blockId="42.[160,275,1401,1462]" box="[160,218,1401,1427]" pageId="42" pageNumber="1554">
|
||
<emphasis id="ECCCEAC9FFF332629A9F4848FF7B16E7" bold="true" box="[160,218,1401,1427]" italics="true" pageId="42" pageNumber="1554">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF332629A9F48ACFEB216C2" blockId="42.[160,275,1401,1462]" box="[160,275,1437,1462]" pageId="42" pageNumber="1554">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="96A26550FFF332629A9F48DCFCEC1538" pageId="42" pageNumber="1554" type="etymology">
|
||
<paragraph id="DE0736DBFFF332629A9F48DCFEBC1573" blockId="42.[160,1155,1517,1612]" box="[160,285,1517,1543]" pageId="42" pageNumber="1554">
|
||
<emphasis id="ECCCEAC9FFF332629A9F48DCFEBC1573" bold="true" box="[160,285,1517,1543]" italics="true" pageId="42" pageNumber="1554">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF332629A9F4B20FCEC1538" blockId="42.[160,1155,1517,1612]" pageId="42" pageNumber="1554">
|
||
The specific epithet,
|
||
<taxonomicName id="19B84D58FFF332629BB84B20FDBA155E" authorityName="Fachin & Hauser" authorityYear="2022" box="[391,539,1553,1578]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="1554" phylum="Arthropoda" rank="species" species="mangoleensis">
|
||
<emphasis id="ECCCEAC9FFF332629BB84B20FDBA155E" box="[391,539,1553,1578]" italics="true" pageId="42" pageNumber="1554">mangoleensis</emphasis>
|
||
</taxonomicName>
|
||
, refers to the collection site of the
|
||
<typeStatus id="01038879FFF33262999F4B20FBA5155E" box="[928,1028,1553,1578]" pageId="42" pageNumber="1554" type="holotype">holotype</typeStatus>
|
||
of the new species, the island of
|
||
<location id="DB676000FFF332629BAD4B02FE561538" LSID="urn:lsid:plazi:treatment:561187CDFFF332639B8A4DBFFDF4113C:DB676000FFF332629BAD4B02FE561538" box="[402,503,1587,1612]" country="Indonesia" municipality="Sula Islands" name="Mangole" pageId="42" pageNumber="1554">Mangole</location>
|
||
, in the
|
||
<collectingMunicipality id="3E63ACA1FFF3326298754B02FD6E1538" box="[586,719,1587,1612]" pageId="42" pageNumber="1554">Sula Islands</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="A6AF764BFFF3326298E54B02FCE81538" box="[730,841,1587,1612]" name="Indonesia" pageId="42" pageNumber="1554">Indonesia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="96A26550FFF332629A9F4BB5FC6715B4" pageId="42" pageNumber="1554" type="distribution">
|
||
<paragraph id="DE0736DBFFF332629A9F4BB5FE1815EA" blockId="42.[160,966,1668,1728]" box="[160,441,1668,1694]" pageId="42" pageNumber="1554">
|
||
<emphasis id="ECCCEAC9FFF332629A9F4BB5FE1815EA" bold="true" box="[160,441,1668,1694]" italics="true" pageId="42" pageNumber="1554">Geographic distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF332629A9F4B96FC6715B4" blockId="42.[160,966,1668,1728]" box="[160,966,1703,1728]" pageId="42" pageNumber="1554">
|
||
<emphasis id="ECCCEAC9FFF332629A9F4B96FEFC15B4" bold="true" box="[160,349,1703,1728]" pageId="42" pageNumber="1554">Oriental Region</emphasis>
|
||
.
|
||
<collectingCountry id="A6AF764BFFF332629B554B96FE7615B4" box="[362,471,1703,1728]" name="Indonesia" pageId="42" pageNumber="1554">Indonesia</collectingCountry>
|
||
(
|
||
<collectingRegion id="1C7CF839FFF332629BDB4B96FD2315B4" box="[484,642,1703,1728]" country="Indonesia" name="Maluku Utara" pageId="42" pageNumber="1554">North Maluku</collectingRegion>
|
||
: Mangole Island) (
|
||
<figureCitation id="46832A5EFFF3326299704B96FC1815B4" box="[847,953,1703,1728]" captionStart="Figure 26" captionStartId="38.[160,226,870,893]" captionTargetBox="[168,1148,146,834]" captionTargetId="figure-167@38.[163,1153,142,839]" captionTargetPageId="38" captionText="Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line." figureDoi="http://doi.org/10.5281/zenodo.7318240" httpUri="https://zenodo.org/record/7318240/files/figure.png" pageId="42" pageNumber="1554">Figure 26</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="96A26550FFF232639A9F4DBFFDF4113C" pageId="43" pageNumber="1555" type="discussion">
|
||
<paragraph id="DE0736DBFFF232639A9F4DBFFEBF13DC" blockId="43.[160,1156,142,584]" box="[160,286,142,168]" pageId="43" pageNumber="1555">
|
||
<emphasis id="ECCCEAC9FFF232639A9F4DBFFEBF13DC" bold="true" box="[160,286,142,168]" italics="true" pageId="43" pageNumber="1555">Comments</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF232639A9F4D83FDF4113C" blockId="43.[160,1156,142,584]" pageId="43" pageNumber="1555">
|
||
This species, as mentioned, shares with the females of
|
||
<taxonomicName id="19B84D58FFF23263993A4D83FCCC13BF" authorityName="James" authorityYear="1939" box="[773,877,178,203]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="1555" phylum="Arthropoda" rank="species" species="kerteszi">
|
||
<emphasis id="ECCCEAC9FFF23263993A4D83FCCC13BF" box="[773,877,178,203]" italics="true" pageId="43" pageNumber="1555">F. kerteszi</emphasis>
|
||
</taxonomicName>
|
||
a small dark spot on the anepisternum and mediotergite completely yellow. However, it is distinguished from
|
||
<taxonomicName id="19B84D58FFF232639E4E4DE5FF511264" authorityName="James" authorityYear="1939" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="1555" phylum="Arthropoda" rank="species" species="kerteszi">
|
||
<emphasis id="ECCCEAC9FFF232639E4E4DE5FF511264" italics="true" pageId="43" pageNumber="1555">F. kerteszi</emphasis>
|
||
</taxonomicName>
|
||
by a vertex area twice the length of the ocellar tubercle (
|
||
<figureCitation id="46832A5EFFF2326399B54DC6FC491264" box="[906,1000,247,272]" captionStart="Figure 6" captionStartId="10.[160,225,1285,1308]" captionTargetBox="[167,1149,145,1251]" captionTargetId="figure-99@10.[163,1153,142,1253]" captionTargetPageId="10" captionText="Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi="http://doi.org/10.5281/zenodo.7318193" httpUri="https://zenodo.org/record/7318193/files/figure.png" pageId="43" pageNumber="1555">Figure 6</figureCitation>
|
||
(c)); also, in
|
||
<taxonomicName id="19B84D58FFF232639E4E4DC6FE951247" authority="Fachin & Hauser, 2022" authorityName="Fachin & Hauser" authorityYear="2022" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="1555" phylum="Arthropoda" rank="species" species="mangoleensis" status="sp. nov.">
|
||
<emphasis id="ECCCEAC9FFF232639E4E4DC6FE951247" italics="true" pageId="43" pageNumber="1555">F. mangoleensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F7FF57B2FFF232639B014C2BFE3A1247" box="[318,411,282,307]" pageId="43" pageNumber="1555" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, the upper frons is relatively shorter than that in
|
||
<taxonomicName id="19B84D58FFF2326399DF4C2BFBED1247" authorityName="James" authorityYear="1939" box="[992,1100,282,307]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="1555" phylum="Arthropoda" rank="species" species="kerteszi">
|
||
<emphasis id="ECCCEAC9FFF2326399DF4C2BFBED1247" box="[992,1100,282,307]" italics="true" pageId="43" pageNumber="1555">F. kerteszi</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<figureCitation id="46832A5EFFF232639A9F4C0DFEA91221" box="[160,264,316,341]" captionStart="Figure 5" captionStartId="9.[160,225,1289,1312]" captionTargetBox="[167,1149,145,1251]" captionTargetId="figure-113@9.[163,1153,142,1257]" captionTargetPageId="9" captionText="Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi="http://doi.org/10.5281/zenodo.7318191" httpUri="https://zenodo.org/record/7318191/files/figure.png" pageId="43" pageNumber="1555">Figures 5</figureCitation>
|
||
(b, c) and 6(c) for comparison) and it has a medial band slightly wider, that does not reach the extreme apex of the anterior part of scutum (see
|
||
<figureCitation id="46832A5EFFF23263994E4C6EFC78120C" box="[881,985,351,376]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="43" pageNumber="1555">Figures 7</figureCitation>
|
||
(b) and 7(g) for comparison). Besides these features,
|
||
<taxonomicName id="19B84D58FFF2326398774CB3FD5912EF" authority="Fachin & Hauser, 2022" authorityName="Fachin & Hauser" authorityYear="2022" box="[584,760,386,411]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="1555" phylum="Arthropoda" rank="species" species="mangoleensis" status="sp. nov.">
|
||
<emphasis id="ECCCEAC9FFF2326398774CB3FD5912EF" box="[584,760,386,411]" italics="true" pageId="43" pageNumber="1555">F. mangoleensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F7FF57B2FFF23263993D4CB3FCFD12EF" box="[770,860,386,411]" pageId="43" pageNumber="1555" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is so far the only species reported within the limits of Wallacea (
|
||
<figureCitation id="46832A5EFFF2326398674C95FD6312C9" box="[600,706,420,445]" captionStart="Figure 26" captionStartId="38.[160,226,870,893]" captionTargetBox="[168,1148,146,834]" captionTargetId="figure-167@38.[163,1153,142,839]" captionTargetPageId="38" captionText="Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line." figureDoi="http://doi.org/10.5281/zenodo.7318240" httpUri="https://zenodo.org/record/7318240/files/figure.png" pageId="43" pageNumber="1555">Figure 26</figureCitation>
|
||
) – the area between Wallace’s Line and Lydekker’s Line – while
|
||
<taxonomicName id="19B84D58FFF232639B954CF6FDB21294" authorityName="James" authorityYear="1939" box="[426,531,455,480]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="1555" phylum="Arthropoda" rank="species" species="kerteszi">
|
||
<emphasis id="ECCCEAC9FFF232639B954CF6FDB21294" box="[426,531,455,480]" italics="true" pageId="43" pageNumber="1555">F. kerteszi</emphasis>
|
||
</taxonomicName>
|
||
is found more to the north, beyond Wallace’s Line. The nearest species in geographical proximity to
|
||
<taxonomicName id="19B84D58FFF23263988E4CDBFCF61177" authority="Fachin & Hauser, 2022" authorityName="Fachin & Hauser" authorityYear="2022" box="[689,855,490,515]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="1555" phylum="Arthropoda" rank="species" species="mangolensis" status="sp. nov.">
|
||
<emphasis id="ECCCEAC9FFF23263988E4CDBFCF61177" box="[689,855,490,515]" italics="true" pageId="43" pageNumber="1555">F. mangolensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F7FF57B2FFF23263995D4CDBFC1C1177" box="[866,957,490,515]" pageId="43" pageNumber="1555" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is a record of
|
||
<taxonomicName id="19B84D58FFF232639E4E4CDBFF541151" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="1555" phylum="Arthropoda" rank="species" species="lineatus">
|
||
<emphasis id="ECCCEAC9FFF232639E4E4CDBFF541151" italics="true" pageId="43" pageNumber="1555">F. lineatus</emphasis>
|
||
</taxonomicName>
|
||
from the Island of Waigeo,
|
||
<collectingCountry id="A6AF764BFFF23263981A4F3DFD351151" box="[549,660,524,549]" name="Indonesia" pageId="43" pageNumber="1555">Indonesia</collectingCountry>
|
||
, near the north-west boundary of Lydekker’s Line, which is in the Australian Region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |