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<document id="F1849ABAFFCFCD8A16FADE5734C50830" ID-CLB-Dataset="187408" ID-DOI="10.1080/00222933.2022.2117107" ID-GBIF-Dataset="25e45e0a-f320-409b-8923-5ab29eabf122" ID-ISSN="1464-5262" ID-Zenodo-Dep="7318178" ID-ZooBank="A6F9B9FB-D3F9-49F2-8FC1-5E6A5E0AB3B1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1668419502508" checkinUser="plazi" docAuthor="Fachin, Diego Aguilar &amp; Hauser, Martin" docDate="2022" docId="561187CDFFDC32589BD648DBFB25177F" docLanguage="en" docName="JNATHIST.56.37-40.1513-1562.pdf" docOrigin="Journal of Natural History 56 (37 - 40)" docSource="http://dx.doi.org/10.1080/00222933.2022.2117107" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Formosargus James 1939" docType="treatment" docVersion="7" lastPageNumber="1528" masterDocId="AA28FFB5FFD932489A3F4D31FFA11374" masterDocTitle="Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym" masterLastPageNumber="1562" masterPageNumber="1513" pageNumber="1517" updateTime="1699411179050" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="12321D17CC467EC950D5A9618865A0F0">Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym</mods:title>
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<taxonomicName id="19B84D58FFDC324D9BD648DBFC9B1571" ID-CoL="4KTF" authority="James, 1939" authorityName="James" authorityYear="1939" box="[489,826,1514,1541]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="1517" phylum="Arthropoda" rank="genus">
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<bibRefCitation id="BA294B2AFFDC324D98A748DBFC9B1571" author="James MT" box="[664,826,1514,1541]" pageId="5" pageNumber="1517" pagination="31 - 37" refId="ref18969" refString="James MT. 1939. New Formosan Stratiomyidae in the collection of the Deutsches Entomologisches Institut. Arbeiten uber morphologische und taxonomische Entomologie aus Berlin-Dahlem. 6 (1): 31 - 37." type="journal article" year="1939">James, 1939</bibRefCitation>
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captionTargetPageId-5="10" captionTargetPageId-6="12" captionTargetPageId-7="13" captionTargetPageId-8="17" captionTargetPageId-9="18" captionText-0="Figure 1. Formosargus berezovskiyi sp. nov., live specimen, male, Taiwan, Orchid Island (© Yi-Xiang Lin (Gaga)). Available from: http://gaga.biodiv.tw/9501bx/536.htm." captionText-1="Figure 2. Antennae of Formosargus species. (a) F. lineatus, male. (b) F. lineatus, female. (c) F. trivittatus sp. nov., male, holotype. Abbreviations: ar t flgm, arista-like terminal flagellomere; flg, flagellum; ped, pedicel; scp, scape." captionText-10="Figure 11. Male terminalia of Formosargus lineatus. (a) Genitalia, dorsal view; arrow: reduced gonocoxal apodeme. (b) Genitalia, lateral view; arrow: inner projection of synsternite. (c) Genitalia, ventral view; arrow: medial line. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; gonst, gonostylus; ph, phallus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger." captionText-11="Figure 12. Male terminalia of Formosargus sagittocerus. (a) Genitalia,dorsal view;arrow:reduced gonocoxal apodeme. (b) Genitalia, lateral view. (c) Genitalia, ventral view. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; goncx brg, gonocoxal bridge; gonst, gonostylus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger." captionText-12="Figure 13. Male terminalia of Formosargus trivittatus sp. nov., holotype. (a) Genitalia, dorsal view; arrow: reduced gonocoxal apodeme. (b) Genitalia, lateral view. (c) Genitalia, ventral view. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger." captionText-13="Figure 14. Male terminalia of Formosargus woodleyi sp. nov., holotype. (a) Genitalia, dorsal view; arrow: reduced gonocoxal apodeme. (b) Genitalia, lateral view. (c) Genitalia, ventral view. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath; prct, proctiger." captionText-14="Figure 15. Female terminalia of Formosargus species, dorsal view. (a) F. kerteszi. (b) F. lineatus. (c) F. berezovskiyi sp. nov., paratype. (d) F. borneensis sp. nov., holotype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite." captionText-15="Figure 16. Female terminalia of Formosargus species, dorsal view. (a) F. mangoleensis sp. nov, holotype. (b) F. trivittatus sp. nov., paratype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite." captionText-16="Figure 17. Genital fork (sternite 9) of Formosargus species. (a–b). F. kerteszi (a. dorsal view; b. ventral view). (c–d). F. lineatus (c. dorsal view; d. ventral view). (e–f) F. berezovskiyi sp. nov., holotype (e. dorsal view; f. ventral view). (g–h) F. borneensis sp. nov., holotype (g. dorsal view; h. ventral view). (i–j) F. mangoleensis sp. nov., holotype (i. dorsal view; j. ventral view). (k–l) F. trivittatus sp. nov., paratype (k. dorsal view;l. ventral view). Abbreviations: go, genital opening;pb, posterior bridge; pp, posterolateral process." captionText-17="Figure 18. Habitus of Formosargus species. (a) F. kerteszi, female, dorsal. (b–e) F. lineatus (b–c. male, dorsal and lateral, respectively; d–e. female, dorsal and lateral, respectively). (f) F. sagittocerus, male, dorsal." captionText-18="Figure 19. Habitus of Formosargus species. (a) F. sagittocerus, male, lateral. (b–c) F. berezovskiyi sp. nov., female, paratype, dorsal and lateral, respectively. (d–e) F. borneensis sp. nov., female, holotype, dorsal and lateral, respectively. (f) F. mangoleensis sp. nov., female, holotype, dorsal." captionText-19="Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively." captionText-2="Figure 3. Heads of Formosargus species, dorsal view. (a) F. kerteszi, female. (b–c) F. lineatus (b. male; c. female). (d) F. sagittocerus, male." captionText-20="Figure 21. Formosargus kerteszi, male,holotype [DEI]. (a) Habitus, lateral view. (b) Habitus, dorsal view. (c) Wing. (d) Antenna. (e) Male terminalia, ventral view. (f) Labels." captionText-21="Figure 22. Formosargus kerteszi, female, paratype [WSU] and allotype [DEI]. (a–d). Paratype. (e–i). Allotype." captionText-22="Figure 23. Chrysochlora lineata, female, holotype [ZMAN]. (a) Habitus, dorsal view. (b) Habitus, lateral view. (c) Habitus, frontal view." captionText-23="Figure 24. Amsaria sagittocera, female, holotype [MZB]. (a–b) Habitus, dorsal view. (c–d) Habitus, lateral view. (e) Head, frontal view." captionText-24="Figure 25. Formosargus variegatus, male, holotype [UZMC]. (a) Habitus, dorsal view.(b) Habitus, lateral view. Arrow points to a modified structure in the mid leg. (c) Labels (© Natural History Museum of Denmark, University of Copenhagen)." captionText-25="Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line." captionText-3="Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype." captionText-4="Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons." captionText-5="Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons." captionText-6="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." captionText-7="Figure 8. Male mid legs of Formosargus species, male. (a–b) F. kerteszi. (c–e) F. lineatus. (f–h) F. sagittocerus. (i–j) F. woodleyi sp. nov., holotype. Arrow: modified structure at base of basitarsus. Abbreviations: fem, femur; tsm 1, first tarsomere." captionText-8="Figure 9. Wings of Formosargus species. (a) F. lineatus, female. (b) F. berezovskiyi sp. nov., female, paratype. (c) F. woodleyi sp. nov., male, holotype. Abbreviations: A1, first branch of anal vein; al, alula; an lb, anal lobe; bm, basal medial cell; br, basal radial cell; C, costal vein; c, costal cell; Cu, cubital vein or cubitus; CuA, anterior branch of cubital vein; cua, anterior cubital cell; CuA+CuP, anterior branch of cubital vein and posterior branch of cubital vein; CuP, posterior branch of cubital vein; cup, posterior cubital cell; d, discal cell; h, humeral crossvein; M, medial vein or media; M1, first branch of media; m1, first medial cell; M1+2, fused first and second branch of media; M2, second branch of media; m2+3, fused second and third cell of media;M3, third branch of media; M3+4, fused third and fourth branch of media; M4, fourth branch of media; m4, fourth medial cell; m–cu, medial–cubital crossvein; m–m, medial crossvein; R1, anterior branch of radius; r1, first radial cell; R2+3, second branch of radius; r2+3, second and third radial cell; R4, upper branch of third branch of radius; r4, fourth radial cell; R5, lower branch of third branch of radius; r5, fifth radial cell; r–m, radial–medial crossvein; Rs, radial sector; Sc, subcostal vein sc, subcostal cell." captionText-9="Figure 10. Male terminalia of Formosargus species (epandrium, proctiger, and cerci removed). (a–b) F. lineatus (a. dorsal view; b. lateral view). (c–e) F. trivittatus sp. nov., holotype (c. dorsal view; d. ventral view; e. lateral view). (f) F. woodleyi sp. nov., lateral view, holotype. Abbreviations: goncx apod, gonocoxal apodeme; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath. Scale bars: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7318182" figureDoi-1="http://doi.org/10.5281/zenodo.7318184" figureDoi-10="http://doi.org/10.5281/zenodo.7318205" figureDoi-11="http://doi.org/10.5281/zenodo.7318207" figureDoi-12="http://doi.org/10.5281/zenodo.7318209" figureDoi-13="http://doi.org/10.5281/zenodo.7318211" figureDoi-14="http://doi.org/10.5281/zenodo.7318213" figureDoi-15="http://doi.org/10.5281/zenodo.7318216" figureDoi-16="http://doi.org/10.5281/zenodo.7318220" figureDoi-17="http://doi.org/10.5281/zenodo.7318222" figureDoi-18="http://doi.org/10.5281/zenodo.7318226" figureDoi-19="http://doi.org/10.5281/zenodo.7318228" figureDoi-2="http://doi.org/10.5281/zenodo.7318186" figureDoi-20="http://doi.org/10.5281/zenodo.7318230" figureDoi-21="http://doi.org/10.5281/zenodo.7318232" figureDoi-22="http://doi.org/10.5281/zenodo.7318234" figureDoi-23="http://doi.org/10.5281/zenodo.7318236" figureDoi-24="http://doi.org/10.5281/zenodo.7318238" figureDoi-25="http://doi.org/10.5281/zenodo.7318240" figureDoi-3="http://doi.org/10.5281/zenodo.7318188" figureDoi-4="http://doi.org/10.5281/zenodo.7318191" figureDoi-5="http://doi.org/10.5281/zenodo.7318193" figureDoi-6="http://doi.org/10.5281/zenodo.7318195" figureDoi-7="http://doi.org/10.5281/zenodo.7318197" figureDoi-8="http://doi.org/10.5281/zenodo.7318199" figureDoi-9="http://doi.org/10.5281/zenodo.7318203" httpUri-0="https://zenodo.org/record/7318182/files/figure.png" httpUri-1="https://zenodo.org/record/7318184/files/figure.png" httpUri-10="https://zenodo.org/record/7318205/files/figure.png" httpUri-11="https://zenodo.org/record/7318207/files/figure.png" httpUri-12="https://zenodo.org/record/7318209/files/figure.png" httpUri-13="https://zenodo.org/record/7318211/files/figure.png" httpUri-14="https://zenodo.org/record/7318213/files/figure.png" httpUri-15="https://zenodo.org/record/7318216/files/figure.png" httpUri-16="https://zenodo.org/record/7318220/files/figure.png" httpUri-17="https://zenodo.org/record/7318222/files/figure.png" httpUri-18="https://zenodo.org/record/7318226/files/figure.png" httpUri-19="https://zenodo.org/record/7318228/files/figure.png" httpUri-2="https://zenodo.org/record/7318186/files/figure.png" httpUri-20="https://zenodo.org/record/7318230/files/figure.png" httpUri-21="https://zenodo.org/record/7318232/files/figure.png" httpUri-22="https://zenodo.org/record/7318234/files/figure.png" httpUri-23="https://zenodo.org/record/7318236/files/figure.png" httpUri-24="https://zenodo.org/record/7318238/files/figure.png" httpUri-25="https://zenodo.org/record/7318240/files/figure.png" httpUri-3="https://zenodo.org/record/7318188/files/figure.png" httpUri-4="https://zenodo.org/record/7318191/files/figure.png" httpUri-5="https://zenodo.org/record/7318193/files/figure.png" httpUri-6="https://zenodo.org/record/7318195/files/figure.png" httpUri-7="https://zenodo.org/record/7318197/files/figure.png" httpUri-8="https://zenodo.org/record/7318199/files/figure.png" httpUri-9="https://zenodo.org/record/7318203/files/figure.png" pageId="5" pageNumber="1517">Figures 1–26</figureCitation>
)
</paragraph>
</subSubSection>
<treatmentCitationGroup id="FEA811F5FFDC324D9A9F4B0EFE9B150F" pageId="5" pageNumber="1517">
<subSubSection id="96A26550FFDC324D9A9F4B0EFDB1152C" box="[160,528,1599,1624]" pageId="5" pageNumber="1517" type="reference_group">
<paragraph id="DE0736DBFFDC324D9A9F4B0EFDB1152C" blockId="5.[160,1155,1599,1728]" box="[160,528,1599,1624]" pageId="5" pageNumber="1517">
<treatmentCitation id="5F1910CAFFDC324D9A9F4B0EFDAD152C" author="James MT" box="[160,524,1599,1624]" page="35" pageId="5" pageNumber="1517" year="1939">
<taxonomicName id="19B84D58FFDC324D9A9F4B0EFDAD152C" ID-CoL="4KTF" authority="James, 1939: 35" authorityName="James" authorityPageNumber="35" authorityYear="1939" box="[160,524,1599,1624]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="1517" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFDC324D9A9F4B0EFE9D152C" bold="true" box="[160,316,1599,1624]" italics="true" pageId="5" pageNumber="1517">Formosargus</emphasis>
<bibRefCitation id="BA294B2AFFDC324D9B764B0EFDAD152C" author="James MT" box="[329,524,1599,1624]" pageId="5" pageNumber="1517" pagination="31 - 37" refId="ref18969" refString="James MT. 1939. New Formosan Stratiomyidae in the collection of the Deutsches Entomologisches Institut. Arbeiten uber morphologische und taxonomische Entomologie aus Berlin-Dahlem. 6 (1): 31 - 37." type="journal article" year="1939">James, 1939: 35</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="96A26550FFDC324D98234B0EFE9B150F" pageId="5" pageNumber="1517" type="type_taxon">
<paragraph id="DE0736DBFFDC324D98234B0EFE9B150F" blockId="5.[160,1155,1599,1728]" pageId="5" pageNumber="1517">
<typeStatus id="01038879FFDC324D98234B0EFDF5152C" box="[540,596,1599,1624]" pageId="5" pageNumber="1517">Type</typeStatus>
species: 
<taxonomicName id="19B84D58FFDC324D98F94B0EFBF4152C" ID-CoL="6JHMJ" authority="James, 1939" authorityName="James" authorityYear="1939" box="[710,1109,1599,1624]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="1517" phylum="Arthropoda" rank="species" species="kerteszi">
<emphasis id="ECCCEAC9FFDC324D98F94B0EFC16152C" box="[710,951,1599,1624]" italics="true" pageId="5" pageNumber="1517">Formosargus kerteszi</emphasis>
<bibRefCitation id="BA294B2AFFDC324D99FB4B0EFBF4152C" author="James MT" box="[964,1109,1599,1624]" pageId="5" pageNumber="1517" pagination="31 - 37" refId="ref18969" refString="James MT. 1939. New Formosan Stratiomyidae in the collection of the Deutsches Entomologisches Institut. Arbeiten uber morphologische und taxonomische Entomologie aus Berlin-Dahlem. 6 (1): 31 - 37." type="journal article" year="1939">James, 1939</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
</treatmentCitationGroup>
<treatmentCitationGroup id="FEA811F5FFDC324E9A9F4BB4FEB51542" lastPageId="6" lastPageNumber="1518" pageId="5" pageNumber="1517">
<subSubSection id="96A26550FFDC324D9A9F4BB4FDBD15B4" pageId="5" pageNumber="1517" type="reference_group">
<paragraph id="DE0736DBFFDC324D9A9F4BB4FDBD15B4" blockId="5.[160,1155,1599,1728]" pageId="5" pageNumber="1517">
<treatmentCitation id="5F1910CAFFDC324D9A9F4BB4FE5E15EA" author="Adisoemarto S" box="[160,511,1669,1694]" page="69" pageId="5" pageNumber="1517" year="1974">
<taxonomicName id="19B84D58FFDC324D9A9F4BB4FE5E15EA" ID-CoL="62CDB" authority="Adisoemarto, 1974: 69" authorityName="Adisoemarto" authorityPageNumber="69" authorityYear="1974" box="[160,511,1669,1694]" class="Insecta" family="Stratiomyidae" genus="Amsaria" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="1517" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFDC324D9A9F4BB4FF5815EA" box="[160,249,1669,1694]" italics="true" pageId="5" pageNumber="1517">Amsaria</emphasis>
<bibRefCitation id="BA294B2AFFDC324D9B3E4BB4FE5E15EA" author="Adisoemarto S" box="[257,511,1669,1694]" pageId="5" pageNumber="1517" pagination="69 - 71" refId="ref18465" refString="Adisoemarto S. 1974. A new genus of soldier flies from Indonesia (Diptera: Stratiomyidae). Treubia. 28 (3): 69 - 71." type="journal article" year="1974">Adisoemarto, 1974: 69</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. 
<typeStatus id="01038879FFDC324D98354BB4FDE115EA" box="[522,576,1669,1694]" pageId="5" pageNumber="1517">Type</typeStatus>
species: 
<taxonomicName id="19B84D58FFDC324D98994BB4FBFB15EA" ID-CoL="D8CH" authority="Adisoemarto, 1974" authorityName="Adisoemarto" authorityYear="1974" box="[678,1114,1669,1694]" class="Insecta" family="Stratiomyidae" genus="Amsaria" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="1517" phylum="Arthropoda" rank="species" species="sagittocera">
<emphasis id="ECCCEAC9FFDC324D98994BB4FCDE15EA" box="[678,895,1669,1694]" italics="true" pageId="5" pageNumber="1517">Amsaria sagittocera</emphasis>
<bibRefCitation id="BA294B2AFFDC324D99B84BB4FBFB15EA" author="Adisoemarto S" box="[903,1114,1669,1694]" pageId="5" pageNumber="1517" pagination="69 - 71" refId="ref18465" refString="Adisoemarto S. 1974. A new genus of soldier flies from Indonesia (Diptera: Stratiomyidae). Treubia. 28 (3): 69 - 71." type="journal article" year="1974">Adisoemarto, 1974</bibRefCitation>
</taxonomicName>
, by original designation. 
<emphasis id="ECCCEAC9FFDC324D9B924B96FDBD15B4" bold="true" box="[429,540,1703,1728]" pageId="5" pageNumber="1517">Syn. nov.</emphasis>
</paragraph>
</subSubSection>
<caption id="8AC76653FFDF324E9A9F484DFEFF16BA" ID-DOI="http://doi.org/10.5281/zenodo.7318184" ID-Zenodo-Dep="7318184" httpUri="https://zenodo.org/record/7318184/files/figure.png" pageId="6" pageNumber="1518" startId="6.[160,225,1404,1427]" targetBox="[165,1150,144,1369]" targetPageId="6">
<paragraph id="DE0736DBFFDF324E9A9F484DFEFF16BA" blockId="6.[160,1156,1404,1486]" pageId="6" pageNumber="1518">
<emphasis id="ECCCEAC9FFDF324E9A9F484DFF5816E7" bold="true" box="[160,249,1404,1427]" pageId="6" pageNumber="1518">Figure 2.</emphasis>
Antennae of 
<taxonomicName id="19B84D58FFDF324E9B40484DFDEC16E7" box="[383,589,1404,1427]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="1518" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="ECCCEAC9FFDF324E9B40484DFE5B16E7" box="[383,506,1404,1427]" italics="true" pageId="6" pageNumber="1518">Formosargus</emphasis>
species.
</taxonomicName>
(a) 
<taxonomicName id="19B84D58FFDF324E984E484DFD7016E7" box="[625,721,1404,1427]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="1518" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="ECCCEAC9FFDF324E984E484DFD7016E7" box="[625,721,1404,1427]" italics="true" pageId="6" pageNumber="1518">F. lineatus</emphasis>
</taxonomicName>
, male. (b) 
<taxonomicName id="19B84D58FFDF324E9906484DFC3B16E7" box="[825,922,1404,1427]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="1518" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="ECCCEAC9FFDF324E9906484DFC3B16E7" box="[825,922,1404,1427]" italics="true" pageId="6" pageNumber="1518">F. lineatus</emphasis>
</taxonomicName>
, female. (c) 
<taxonomicName id="19B84D58FFDF324E9E2E484DFB2216E7" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[1041,1155,1404,1427]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="1518" phylum="Arthropoda" rank="species" species="trivittatus" status="sp. nov.">
<emphasis id="ECCCEAC9FFDF324E9E2E484DFB2216E7" box="[1041,1155,1404,1427]" italics="true" pageId="6" pageNumber="1518">F. trivittatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFDF324E9A9F48A8FF4E16C4" box="[160,239,1433,1456]" pageId="6" pageNumber="1518" rank="species">sp. nov.</taxonomicNameLabel>
, male, holotype. Abbreviations: ar t flgm, arista-like terminal flagellomere; flg, flagellum; ped, pedicel; scp, scape.
</paragraph>
</caption>
<subSubSection id="96A26550FFDF324E9A9F4B2DFEB51542" box="[160,276,1564,1590]" pageId="6" pageNumber="1518" type="diagnosis">
<paragraph id="DE0736DBFFDF324E9A9F4B2DFEB51542" blockId="6.[160,1155,1564,1694]" box="[160,276,1564,1590]" pageId="6" pageNumber="1518">
<emphasis id="ECCCEAC9FFDF324E9A9F4B2DFEB51542" bold="true" box="[160,276,1564,1590]" italics="true" pageId="6" pageNumber="1518">Diagnosis</emphasis>
</paragraph>
</subSubSection>
</treatmentCitationGroup>
<paragraph id="DE0736DBFFDF324F9A9F4B71FE5C1654" blockId="6.[160,1155,1564,1694]" lastBlockId="7.[160,1156,1045,1312]" lastPageId="7" lastPageNumber="1519" pageId="6" pageNumber="1518">
Dichoptic eyes in both sexes (
<figureCitation id="46832A5EFFDF324E9BD24B71FDCF152D" box="[493,622,1600,1625]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-2="Figure 5" captionStart-3="Figure 6" captionStartId-0="7.[160,225,906,929]" captionStartId-1="8.[160,225,1266,1289]" captionStartId-2="9.[160,225,1289,1312]" captionStartId-3="10.[160,225,1285,1308]" captionTargetBox-0="[167,1149,145,869]" captionTargetBox-1="[167,1149,145,1232]" captionTargetBox-2="[167,1149,145,1251]" captionTargetBox-3="[167,1149,145,1251]" captionTargetId-0="figure-267@7.[163,1153,142,874]" captionTargetId-1="figure-139@8.[163,1153,142,1234]" captionTargetId-2="figure-113@9.[163,1153,142,1257]" captionTargetId-3="figure-99@10.[163,1153,142,1253]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="Figure 3. Heads of Formosargus species, dorsal view. (a) F. kerteszi, female. (b–c) F. lineatus (b. male; c. female). (d) F. sagittocerus, male." captionText-1="Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype." captionText-2="Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons." captionText-3="Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi-0="http://doi.org/10.5281/zenodo.7318186" figureDoi-1="http://doi.org/10.5281/zenodo.7318188" figureDoi-2="http://doi.org/10.5281/zenodo.7318191" figureDoi-3="http://doi.org/10.5281/zenodo.7318193" httpUri-0="https://zenodo.org/record/7318186/files/figure.png" httpUri-1="https://zenodo.org/record/7318188/files/figure.png" httpUri-2="https://zenodo.org/record/7318191/files/figure.png" httpUri-3="https://zenodo.org/record/7318193/files/figure.png" pageId="6" pageNumber="1518">Figures 3–6</figureCitation>
). Ocellar tubercle often reduced in size, as wide as vertex area or much narrower (
<figureCitation id="46832A5EFFDF324E982B4B52FD321508" box="[532,659,1635,1660]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStartId-0="7.[160,225,906,929]" captionStartId-1="8.[160,225,1266,1289]" captionTargetBox-0="[167,1149,145,869]" captionTargetBox-1="[167,1149,145,1232]" captionTargetId-0="figure-267@7.[163,1153,142,874]" captionTargetId-1="figure-139@8.[163,1153,142,1234]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figure 3. Heads of Formosargus species, dorsal view. (a) F. kerteszi, female. (b–c) F. lineatus (b. male; c. female). (d) F. sagittocerus, male." captionText-1="Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype." figureDoi-0="http://doi.org/10.5281/zenodo.7318186" figureDoi-1="http://doi.org/10.5281/zenodo.7318188" httpUri-0="https://zenodo.org/record/7318186/files/figure.png" httpUri-1="https://zenodo.org/record/7318188/files/figure.png" pageId="6" pageNumber="1518">Figures 3–4</figureCitation>
); upper frons short, wider than long, at most only slightly longer than wide; lower frons often longer than upper frons (
<figureCitation id="46832A5EFFDF324E99CC4BB4FBD715EA" box="[1011,1142,1669,1694]" captionStart-0="Figure 5" captionStart-1="Figure 6" captionStartId-0="9.[160,225,1289,1312]" captionStartId-1="10.[160,225,1285,1308]" captionTargetBox-0="[167,1149,145,1251]" captionTargetBox-1="[167,1149,145,1251]" captionTargetId-0="figure-113@9.[163,1153,142,1257]" captionTargetId-1="figure-99@10.[163,1153,142,1253]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons." captionText-1="Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi-0="http://doi.org/10.5281/zenodo.7318191" figureDoi-1="http://doi.org/10.5281/zenodo.7318193" httpUri-0="https://zenodo.org/record/7318191/files/figure.png" httpUri-1="https://zenodo.org/record/7318193/files/figure.png" pageId="6" pageNumber="1518">Figures 5–6</figureCitation>
). Male mid legs with a modified structure at base of first tarsomere (
<figureCitation id="46832A5EFFDE324F998B4924FBB2175A" box="[948,1043,1045,1070]" captionStart="Figure 8" captionStartId="13.[160,225,1395,1418]" captionTargetBox="[167,1149,145,1360]" captionTargetId="figure-51@13.[163,1153,142,1363]" captionTargetPageId="13" captionText="Figure 8. Male mid legs of Formosargus species, male. (a–b) F. kerteszi. (c–e) F. lineatus. (f–h) F. sagittocerus. (i–j) F. woodleyi sp. nov., holotype. Arrow: modified structure at base of basitarsus. Abbreviations: fem, femur; tsm 1, first tarsomere." figureDoi="http://doi.org/10.5281/zenodo.7318197" httpUri="https://zenodo.org/record/7318197/files/figure.png" pageId="7" pageNumber="1519">Figure 8</figureCitation>
). M4 vein branches off from discal cell, with m–cu crossvein connected at M3+4 fork (
<figureCitation id="46832A5EFFDE324F99CD4906FBEE1724" box="[1010,1103,1079,1104]" captionStart="Figure 9" captionStartId="17.[160,225,1285,1308]" captionTargetBox="[171,1149,145,1249]" captionTargetId="figure-3@17.[163,1153,142,1253]" captionTargetPageId="17" captionText="Figure 9. Wings of Formosargus species. (a) F. lineatus, female. (b) F. berezovskiyi sp. nov., female, paratype. (c) F. woodleyi sp. nov., male, holotype. Abbreviations: A1, first branch of anal vein; al, alula; an lb, anal lobe; bm, basal medial cell; br, basal radial cell; C, costal vein; c, costal cell; Cu, cubital vein or cubitus; CuA, anterior branch of cubital vein; cua, anterior cubital cell; CuA+CuP, anterior branch of cubital vein and posterior branch of cubital vein; CuP, posterior branch of cubital vein; cup, posterior cubital cell; d, discal cell; h, humeral crossvein; M, medial vein or media; M1, first branch of media; m1, first medial cell; M1+2, fused first and second branch of media; M2, second branch of media; m2+3, fused second and third cell of media;M3, third branch of media; M3+4, fused third and fourth branch of media; M4, fourth branch of media; m4, fourth medial cell; m–cu, medial–cubital crossvein; m–m, medial crossvein; R1, anterior branch of radius; r1, first radial cell; R2+3, second branch of radius; r2+3, second and third radial cell; R4, upper branch of third branch of radius; r4, fourth radial cell; R5, lower branch of third branch of radius; r5, fifth radial cell; r–m, radial–medial crossvein; Rs, radial sector; Sc, subcostal vein sc, subcostal cell." figureDoi="http://doi.org/10.5281/zenodo.7318199" httpUri="https://zenodo.org/record/7318199/files/figure.png" pageId="7" pageNumber="1519">Figure 9</figureCitation>
); M1 and M3 long, but often fading, only apparent basally or as a fold (
<figureCitation id="46832A5EFFDE324F99A2496BFC5D1707" box="[925,1020,1114,1139]" captionStart="Figure 9" captionStartId="17.[160,225,1285,1308]" captionTargetBox="[171,1149,145,1249]" captionTargetId="figure-3@17.[163,1153,142,1253]" captionTargetPageId="17" captionText="Figure 9. Wings of Formosargus species. (a) F. lineatus, female. (b) F. berezovskiyi sp. nov., female, paratype. (c) F. woodleyi sp. nov., male, holotype. Abbreviations: A1, first branch of anal vein; al, alula; an lb, anal lobe; bm, basal medial cell; br, basal radial cell; C, costal vein; c, costal cell; Cu, cubital vein or cubitus; CuA, anterior branch of cubital vein; cua, anterior cubital cell; CuA+CuP, anterior branch of cubital vein and posterior branch of cubital vein; CuP, posterior branch of cubital vein; cup, posterior cubital cell; d, discal cell; h, humeral crossvein; M, medial vein or media; M1, first branch of media; m1, first medial cell; M1+2, fused first and second branch of media; M2, second branch of media; m2+3, fused second and third cell of media;M3, third branch of media; M3+4, fused third and fourth branch of media; M4, fourth branch of media; m4, fourth medial cell; m–cu, medial–cubital crossvein; m–m, medial crossvein; R1, anterior branch of radius; r1, first radial cell; R2+3, second branch of radius; r2+3, second and third radial cell; R4, upper branch of third branch of radius; r4, fourth radial cell; R5, lower branch of third branch of radius; r5, fifth radial cell; r–m, radial–medial crossvein; Rs, radial sector; Sc, subcostal vein sc, subcostal cell." figureDoi="http://doi.org/10.5281/zenodo.7318199" httpUri="https://zenodo.org/record/7318199/files/figure.png" pageId="7" pageNumber="1519">Figure 9</figureCitation>
(a)). In male genitalia, gonocoxal apodeme reduced, not discernible; gonocoxal bridge at distal third; phallus and parameral sheath very reduced (
<figureCitation id="46832A5EFFDE324F98AB49AEFD5C17CC" box="[660,765,1183,1208]" captionStart="Figure 10" captionStartId="18.[160,225,1626,1649]" captionTargetBox="[173,1142,148,1584]" captionTargetId="figure-3@18.[163,1153,142,1594]" captionTargetPageId="18" captionText="Figure 10. Male terminalia of Formosargus species (epandrium, proctiger, and cerci removed). (a–b) F. lineatus (a. dorsal view; b. lateral view). (c–e) F. trivittatus sp. nov., holotype (c. dorsal view; d. ventral view; e. lateral view). (f) F. woodleyi sp. nov., lateral view, holotype. Abbreviations: goncx apod, gonocoxal apodeme; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7318203" httpUri="https://zenodo.org/record/7318203/files/figure.png" pageId="7" pageNumber="1519">Figure 10</figureCitation>
). Male and female mainly differing in the colouration over thorax and abdomen, with scutal bands slender and less intense and abdomen often completely yellowish in females; frons, frontal callus, and face in females broader than in males.
</paragraph>
<caption id="8AC76653FFDE324F9A9F4EBBFE4510CA" ID-DOI="http://doi.org/10.5281/zenodo.7318186" ID-Zenodo-Dep="7318186" httpUri="https://zenodo.org/record/7318186/files/figure.png" pageId="7" pageNumber="1519" startId="7.[160,225,906,929]" targetBox="[167,1149,145,869]" targetPageId="7">
<paragraph id="DE0736DBFFDE324F9A9F4EBBFE4510CA" blockId="7.[160,1156,906,958]" pageId="7" pageNumber="1519">
<emphasis id="ECCCEAC9FFDE324F9A9F4EBBFF5810D5" bold="true" box="[160,249,906,929]" pageId="7" pageNumber="1519">Figure 3.</emphasis>
Heads of 
<taxonomicName id="19B84D58FFDE324F9B644EBBFD8410D5" box="[347,549,906,929]" pageId="7" pageNumber="1519">
<emphasis id="ECCCEAC9FFDE324F9B644EBBFE7710D5" box="[347,470,906,929]" italics="true" pageId="7" pageNumber="1519">Formosargus</emphasis>
species
</taxonomicName>
, dorsal view. (a) 
<taxonomicName id="19B84D58FFDE324F98F84EBBFC8210D5" authorityName="James" authorityYear="1939" box="[711,803,906,929]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="1519" phylum="Arthropoda" rank="species" species="kerteszi">
<emphasis id="ECCCEAC9FFDE324F98F84EBBFC8210D5" box="[711,803,906,929]" italics="true" pageId="7" pageNumber="1519">F. kerteszi</emphasis>
</taxonomicName>
, female. (b–c) 
<taxonomicName id="19B84D58FFDE324F998C4EBBFBB210D5" box="[947,1043,906,929]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="1519" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="ECCCEAC9FFDE324F998C4EBBFBB210D5" box="[947,1043,906,929]" italics="true" pageId="7" pageNumber="1519">F. lineatus</emphasis>
</taxonomicName>
(b. male; c. female). (d) 
<taxonomicName id="19B84D58FFDE324F9B274E96FE0010CA" authorityName="Fachin &amp; Hauser" authorityYear="2022" baseAuthorityName="Adisoemarto" baseAuthorityYear="1974" box="[280,417,935,958]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="1519" phylum="Arthropoda" rank="species" species="sagittocerus">
<emphasis id="ECCCEAC9FFDE324F9B274E96FE0010CA" box="[280,417,935,958]" italics="true" pageId="7" pageNumber="1519">F. sagittocerus</emphasis>
</taxonomicName>
, male.
</paragraph>
</caption>
<subSubSection id="96A26550FFDE32459A9F487DFEAA15DE" lastPageId="13" lastPageNumber="1525" pageId="7" pageNumber="1519" type="description">
<paragraph id="DE0736DBFFDE324F9A9F487DFEE11612" blockId="7.[160,1156,1356,1728]" box="[160,320,1356,1382]" pageId="7" pageNumber="1519">
<emphasis id="ECCCEAC9FFDE324F9A9F487DFEE11612" bold="true" box="[160,320,1356,1382]" italics="true" pageId="7" pageNumber="1519">Redescription</emphasis>
</paragraph>
<paragraph id="DE0736DBFFDE32439A9F485EFB2511DB" blockId="7.[160,1156,1356,1728]" lastBlockId="11.[160,1156,142,1727]" lastPageId="11" lastPageNumber="1523" pageId="7" pageNumber="1519">
<emphasis id="ECCCEAC9FFDE324F9A9F485EFF7B16FC" bold="true" box="[160,218,1391,1416]" pageId="7" pageNumber="1519">Male</emphasis>
. Length (mm): body, 6.0–6.5; wing, 5.5–6.0. 
<emphasis id="ECCCEAC9FFDE324F98F5485EFCA916FC" bold="true" box="[714,776,1391,1416]" pageId="7" pageNumber="1519">Head</emphasis>
(
<figureCitation id="46832A5EFFDE324F9928485EFC3816FC" box="[791,921,1391,1416]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-2="Figure 5" captionStart-3="Figure 6" captionStartId-0="7.[160,225,906,929]" captionStartId-1="8.[160,225,1266,1289]" captionStartId-2="9.[160,225,1289,1312]" captionStartId-3="10.[160,225,1285,1308]" captionTargetBox-0="[167,1149,145,869]" captionTargetBox-1="[167,1149,145,1232]" captionTargetBox-2="[167,1149,145,1251]" captionTargetBox-3="[167,1149,145,1251]" captionTargetId-0="figure-267@7.[163,1153,142,874]" captionTargetId-1="figure-139@8.[163,1153,142,1234]" captionTargetId-2="figure-113@9.[163,1153,142,1257]" captionTargetId-3="figure-99@10.[163,1153,142,1253]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="Figure 3. Heads of Formosargus species, dorsal view. (a) F. kerteszi, female. (b–c) F. lineatus (b. male; c. female). (d) F. sagittocerus, male." captionText-1="Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype." captionText-2="Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons." captionText-3="Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi-0="http://doi.org/10.5281/zenodo.7318186" figureDoi-1="http://doi.org/10.5281/zenodo.7318188" figureDoi-2="http://doi.org/10.5281/zenodo.7318191" figureDoi-3="http://doi.org/10.5281/zenodo.7318193" httpUri-0="https://zenodo.org/record/7318186/files/figure.png" httpUri-1="https://zenodo.org/record/7318188/files/figure.png" httpUri-2="https://zenodo.org/record/7318191/files/figure.png" httpUri-3="https://zenodo.org/record/7318193/files/figure.png" pageId="7" pageNumber="1519">Figures 3–6</figureCitation>
). Typical of sargines, higher than wide and concave in dorsal view; eyes dichoptic, with very few sparse and minute setae on the eyes, and ommatidia uniform in size, except for a roughly circular area with slightly bigger ommatidia frontally. Frons, frontal callus, and face width nearly one-fifth of head or less, margins parallel-sided. Occiput black. Vertex area (from the posterior ocellus to the margin of the occiput) wide, mostly white to yellowish, except for a thin dark brown line on each side laterally; pilosity often yellowish, short to medium in length. Ocellar tubercle dark brown to black at centre, yellowish or brownish externally, reduced in size, often as wide as vertex or shorter; ocelli in the form of equilateral triangle; pilosity often yellowish, denser than in vertex, short to medium in length. Upper frons (black area between anterior ocellus and white lower frons) often wider than long or at most slightly longer and at least twice shorter than lower frons (whitish part above antenna); medial area of upper frons yellowish or brownish, at least twice as slender as the lateral area; black lateral area width varying from short to as wide as ocellar tubercle width, with short and dense yellowish to darkish pilosity; shape of anterior margin of upper frons at edge with lower frons: inverse-U-shaped (e.g. 
<figureCitation id="46832A5EFFD1324099024B0FFC391523" box="[829,920,1598,1623]" captionStart="Figure 5" captionStartId="9.[160,225,1289,1312]" captionTargetBox="[167,1149,145,1251]" captionTargetId="figure-113@9.[163,1153,142,1257]" captionTargetPageId="9" captionText="Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi="http://doi.org/10.5281/zenodo.7318191" httpUri="https://zenodo.org/record/7318191/files/figure.png" pageId="8" pageNumber="1520">Figure 5</figureCitation>
(a)), straight to nearly straight (
<figureCitation id="46832A5EFFD132409B3B4B50FEC0150E" box="[260,353,1633,1658]" captionStart="Figure 6" captionStartId="10.[160,225,1285,1308]" captionTargetBox="[167,1149,145,1251]" captionTargetId="figure-99@10.[163,1153,142,1253]" captionTargetPageId="10" captionText="Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi="http://doi.org/10.5281/zenodo.7318193" httpUri="https://zenodo.org/record/7318193/files/figure.png" pageId="8" pageNumber="1520">Figure 6</figureCitation>
(d))) or U-shaped, with median triangular extension slightly (
<figureCitation id="46832A5EFFD132409E3B4B50FBFE150E" box="[1028,1119,1633,1658]" captionStart="Figure 6" captionStartId="10.[160,225,1285,1308]" captionTargetBox="[167,1149,145,1251]" captionTargetId="figure-99@10.[163,1153,142,1253]" captionTargetPageId="10" captionText="Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi="http://doi.org/10.5281/zenodo.7318193" httpUri="https://zenodo.org/record/7318193/files/figure.png" pageId="8" pageNumber="1520">Figure 6</figureCitation>
(e)) to strongly (
<figureCitation id="46832A5EFFD132409B1A4BB5FE2115E9" box="[293,384,1668,1693]" captionStart="Figure 5" captionStartId="9.[160,225,1289,1312]" captionTargetBox="[167,1149,145,1251]" captionTargetId="figure-113@9.[163,1153,142,1257]" captionTargetPageId="9" captionText="Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi="http://doi.org/10.5281/zenodo.7318191" httpUri="https://zenodo.org/record/7318191/files/figure.png" pageId="8" pageNumber="1520">Figure 5</figureCitation>
(f)) pointing towards antennal insertion. Frontal callus (= upper part of the lower frons) not distinguishable from basal half of lower frons in colour; latter with short whitish pilosity near antenna insertion. Antennae inserted at basal third of head; scape slightly longer than pedicel; mostly yellowish to reddish brown, except for a dorsoapical arista-like terminal flagellomere often dark brown, longer than remaining antenna, with a few setae at base, but often with only two; basal four flagellomeres compact, with short, relatively dense, yellowish hair-like setae; fourth flagellomere rounded or subtriangular, as long as each previous flagellomere separately (
<figureCitation id="46832A5EFFD0324199184B53FC26150F" box="[807,903,1634,1659]" captionStart="Figure 2" captionStartId="6.[160,225,1404,1427]" captionTargetBox="[165,1150,144,1369]" captionTargetId="figure-53@6.[163,1153,142,1372]" captionTargetPageId="6" captionText="Figure 2. Antennae of Formosargus species. (a) F. lineatus, male. (b) F. lineatus, female. (c) F. trivittatus sp. nov., male, holotype. Abbreviations: ar t flgm, arista-like terminal flagellomere; flg, flagellum; ped, pedicel; scp, scape." figureDoi="http://doi.org/10.5281/zenodo.7318184" httpUri="https://zenodo.org/record/7318184/files/figure.png" pageId="9" pageNumber="1521">Figure 2</figureCitation>
(a, b)) or much longer (
<figureCitation id="46832A5EFFD032419A974BB4FEA315EA" box="[168,258,1669,1694]" captionStart="Figure 2" captionStartId="6.[160,225,1404,1427]" captionTargetBox="[165,1150,144,1369]" captionTargetId="figure-53@6.[163,1153,142,1372]" captionTargetPageId="6" captionText="Figure 2. Antennae of Formosargus species. (a) F. lineatus, male. (b) F. lineatus, female. (c) F. trivittatus sp. nov., male, holotype. Abbreviations: ar t flgm, arista-like terminal flagellomere; flg, flagellum; ped, pedicel; scp, scape." figureDoi="http://doi.org/10.5281/zenodo.7318184" httpUri="https://zenodo.org/record/7318184/files/figure.png" pageId="9" pageNumber="1521">Figure 2</figureCitation>
(c)). Face white to yellow, much shorter than lower frons, pilosity short, yellowish. Proboscis whitish to reddish yellow, labellum well developed; maxillary palpus short, twosegmented, distal segment enlarged and shorter than basal segment. 
<emphasis id="ECCCEAC9FFD33242998848E6FBAA1684" bold="true" box="[951,1035,1495,1520]" pageId="10" pageNumber="1522">Thorax</emphasis>
(
<figureCitation id="46832A5EFFD332429E2548E6FBD71684" box="[1050,1142,1495,1520]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="10" pageNumber="1522">Figure 7</figureCitation>
). Scutum mostly pale yellow in background, but reddish in some species; with a varied pattern of dark brown markings, ranging from a single complete medial band over scutum (e.g. 
<figureCitation id="46832A5EFFD332429B0D4B0EFE2E152C" box="[306,399,1599,1624]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="10" pageNumber="1522">Figure 7</figureCitation>
(a)), with or without additional anterior and posterior lateral bands (e.g. 
<figureCitation id="46832A5EFFD332429AE94B53FE95150F" box="[214,308,1634,1659]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="10" pageNumber="1522">Figure 7</figureCitation>
(h)), to a completely dark scutum (
<figureCitation id="46832A5EFFD3324298874B53FCB6150F" box="[696,791,1634,1659]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="10" pageNumber="1522">Figure 7</figureCitation>
(i)); scutellum often marked with dark brown, at least in the centre. Pleuron pale yellow to reddish yellow in background, with anepisternum, katepisternum, laterotergite, and mediotergite often marked with dark brown; pilosity white to yellow. Legs varying from whitish to reddish yellow, with at least the last three to four hind tarsomeres often dark brown; virtually unmodified, except for hind leg with first tarsomere much longer than tibia, and a modified structure at apex of mid first tarsomere, often hook-like or coiled, unique for each species (
<figureCitation id="46832A5EFFD2324399EA4DC7FB90127B" box="[981,1073,246,271]" captionStart="Figure 8" captionStartId="13.[160,225,1395,1418]" captionTargetBox="[167,1149,145,1360]" captionTargetId="figure-51@13.[163,1153,142,1363]" captionTargetPageId="13" captionText="Figure 8. Male mid legs of Formosargus species, male. (a–b) F. kerteszi. (c–e) F. lineatus. (f–h) F. sagittocerus. (i–j) F. woodleyi sp. nov., holotype. Arrow: modified structure at base of basitarsus. Abbreviations: fem, femur; tsm 1, first tarsomere." figureDoi="http://doi.org/10.5281/zenodo.7318197" httpUri="https://zenodo.org/record/7318197/files/figure.png" pageId="11" pageNumber="1523">Figure 8</figureCitation>
). 
<emphasis id="ECCCEAC9FFD232439E7A4DC7FB22127B" bold="true" box="[1093,1155,246,271]" pageId="11" pageNumber="1523">Wing</emphasis>
(
<figureCitation id="46832A5EFFD232439A974C28FEA71246" box="[168,262,281,306]" captionStart="Figure 9" captionStartId="17.[160,225,1285,1308]" captionTargetBox="[171,1149,145,1249]" captionTargetId="figure-3@17.[163,1153,142,1253]" captionTargetPageId="17" captionText="Figure 9. Wings of Formosargus species. (a) F. lineatus, female. (b) F. berezovskiyi sp. nov., female, paratype. (c) F. woodleyi sp. nov., male, holotype. Abbreviations: A1, first branch of anal vein; al, alula; an lb, anal lobe; bm, basal medial cell; br, basal radial cell; C, costal vein; c, costal cell; Cu, cubital vein or cubitus; CuA, anterior branch of cubital vein; cua, anterior cubital cell; CuA+CuP, anterior branch of cubital vein and posterior branch of cubital vein; CuP, posterior branch of cubital vein; cup, posterior cubital cell; d, discal cell; h, humeral crossvein; M, medial vein or media; M1, first branch of media; m1, first medial cell; M1+2, fused first and second branch of media; M2, second branch of media; m2+3, fused second and third cell of media;M3, third branch of media; M3+4, fused third and fourth branch of media; M4, fourth branch of media; m4, fourth medial cell; m–cu, medial–cubital crossvein; m–m, medial crossvein; R1, anterior branch of radius; r1, first radial cell; R2+3, second branch of radius; r2+3, second and third radial cell; R4, upper branch of third branch of radius; r4, fourth radial cell; R5, lower branch of third branch of radius; r5, fifth radial cell; r–m, radial–medial crossvein; Rs, radial sector; Sc, subcostal vein sc, subcostal cell." figureDoi="http://doi.org/10.5281/zenodo.7318199" httpUri="https://zenodo.org/record/7318199/files/figure.png" pageId="11" pageNumber="1523">Figure 9</figureCitation>
). Wing membrane hyaline, with brownish veins; only microtrichia over membrane, except on alula, basal area of cup and br, bm, and cua cells anteriorly. C ending at wing tip. R2+3 originating slightly before or at level of r–m crossvein; basal third of R2+3 often curved, apical two-thirds straight, parallel to R1 towards wing margin. R4 weakly curved at base, arising from R5 with almost right angle, nearly straight on apical twothirds. R5 slightly curved posteriorly, ending posterior to wing tip. M fading on nearly entire extension between bm and br cells, except on its extreme apical portion; distal branch of M1+2 longer than proximal branch of M1+2; m–m as long as short branch of M3; M1 and M3 weakly developed compared to M2, often discernible proximally or as a fold; M4 connected to discal cell; r–m as long as R4; m–cu crossvein connected to M3+4 fork. Medial branches often not reaching wing margin. Cu vein basal to true cubital fork only setose on its basal half. Alula narrow, without microtrichia. Halter pale yellow to yellow.
</paragraph>
<caption id="8AC76653FFD132409A9F49C3FBA61637" ID-DOI="http://doi.org/10.5281/zenodo.7318188" ID-Zenodo-Dep="7318188" httpUri="https://zenodo.org/record/7318188/files/figure.png" pageId="8" pageNumber="1520" startId="8.[160,225,1266,1289]" targetBox="[167,1149,145,1232]" targetPageId="8">
<paragraph id="DE0736DBFFD132409A9F49C3FBA61637" blockId="8.[160,1157,1266,1347]" pageId="8" pageNumber="1520">
<emphasis id="ECCCEAC9FFD132409A9F49C3FF58167D" bold="true" box="[160,249,1266,1289]" pageId="8" pageNumber="1520">Figure 4.</emphasis>
Heads of 
<taxonomicName id="19B84D58FFD132409B6649C3FD83167D" box="[345,546,1266,1289]" pageId="8" pageNumber="1520">
<emphasis id="ECCCEAC9FFD132409B6649C3FE75167D" box="[345,468,1266,1289]" italics="true" pageId="8" pageNumber="1520">Formosargus</emphasis>
species
</taxonomicName>
, dorsal view. (a) 
<taxonomicName id="19B84D58FFD1324098FD49C3FCE8167D" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[706,841,1266,1289]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="1520" phylum="Arthropoda" rank="species" species="berezovskiyi" status="sp. nov.">
<emphasis id="ECCCEAC9FFD1324098FD49C3FCE8167D" box="[706,841,1266,1289]" italics="true" pageId="8" pageNumber="1520">F. berezovskiyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD13240997049C3FC3C167D" box="[847,925,1266,1289]" pageId="8" pageNumber="1520" rank="species">sp. nov.</taxonomicNameLabel>
, female, paratype. (b) 
<taxonomicName id="19B84D58FFD132409E4C49C3FEA41652" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="1520" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
<emphasis id="ECCCEAC9FFD132409E4C49C3FEA41652" italics="true" pageId="8" pageNumber="1520">F. borneensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD132409B34483EFEFB1652" box="[267,346,1295,1318]" pageId="8" pageNumber="1520" rank="species">sp. nov.</taxonomicNameLabel>
, female, holotype. (c) 
<taxonomicName id="19B84D58FFD13240980E483EFD6B1652" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[561,714,1295,1318]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="1520" phylum="Arthropoda" rank="species" species="mangoleensis" status="sp. nov.">
<emphasis id="ECCCEAC9FFD13240980E483EFD6B1652" box="[561,714,1295,1318]" italics="true" pageId="8" pageNumber="1520">F. mangoleensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD1324098EF483EFCBE1652" box="[720,799,1295,1318]" pageId="8" pageNumber="1520" rank="species">sp. nov.</taxonomicNameLabel>
, female, holotype. (d–e) 
<taxonomicName id="19B84D58FFD132409E2E483EFB221652" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[1041,1155,1295,1318]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="1520" phylum="Arthropoda" rank="species" species="trivittatus" status="sp. nov.">
<emphasis id="ECCCEAC9FFD132409E2E483EFB221652" box="[1041,1155,1295,1318]" italics="true" pageId="8" pageNumber="1520">F. trivittatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD132409A9F481DFF4F1637" box="[160,238,1324,1347]" pageId="8" pageNumber="1520" rank="species">sp. nov.</taxonomicNameLabel>
(d. male, holotype; e. female, paratype). (f) 
<taxonomicName id="19B84D58FFD13240989E481DFCAC1637" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[673,781,1324,1347]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="1520" phylum="Arthropoda" rank="species" species="woodleyi" status="sp. nov.">
<emphasis id="ECCCEAC9FFD13240989E481DFCAC1637" box="[673,781,1324,1347]" italics="true" pageId="8" pageNumber="1520">F. woodleyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD13240992B481DFCC51637" box="[788,868,1324,1347]" pageId="8" pageNumber="1520" rank="species">sp. nov.</taxonomicNameLabel>
, male, holotype.
</paragraph>
</caption>
<caption id="8AC76653FFD032419A9F4838FC05162E" ID-DOI="http://doi.org/10.5281/zenodo.7318191" ID-Zenodo-Dep="7318191" httpUri="https://zenodo.org/record/7318191/files/figure.png" pageId="9" pageNumber="1521" startId="9.[160,225,1289,1312]" targetBox="[167,1149,145,1251]" targetPageId="9">
<paragraph id="DE0736DBFFD032419A9F4838FC05162E" blockId="9.[160,1157,1289,1370]" pageId="9" pageNumber="1521">
<emphasis id="ECCCEAC9FFD032419A9F4838FF5B1654" bold="true" box="[160,250,1289,1312]" pageId="9" pageNumber="1521">Figure 5.</emphasis>
Heads of 
<taxonomicName id="19B84D58FFD032419B604838FD8B1654" box="[351,554,1289,1312]" pageId="9" pageNumber="1521">
<emphasis id="ECCCEAC9FFD032419B604838FE7B1654" box="[351,474,1289,1312]" italics="true" pageId="9" pageNumber="1521">Formosargus</emphasis>
species
</taxonomicName>
, frontal view. (a–c) 
<taxonomicName id="19B84D58FFD0324198D34838FCEB1654" authorityName="James" authorityYear="1939" box="[748,842,1289,1312]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1521" phylum="Arthropoda" rank="species" species="kerteszi">
<emphasis id="ECCCEAC9FFD0324198D34838FCEB1654" box="[748,842,1289,1312]" italics="true" pageId="9" pageNumber="1521">F. kerteszi</emphasis>
</taxonomicName>
(a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) 
<taxonomicName id="19B84D58FFD03241989A4817FCA71649" box="[677,774,1318,1341]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1521" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="ECCCEAC9FFD03241989A4817FCA71649" box="[677,774,1318,1341]" italics="true" pageId="9" pageNumber="1521">F. lineatus</emphasis>
</taxonomicName>
(d. male; e. female). (f) 
<taxonomicName id="19B84D58FFD0324199CA4817FBDF1649" authorityName="Fachin &amp; Hauser" authorityYear="2022" baseAuthorityName="Adisoemarto" baseAuthorityYear="1974" box="[1013,1150,1318,1341]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1521" phylum="Arthropoda" rank="species" species="sagittocerus">
<emphasis id="ECCCEAC9FFD0324199CA4817FBDF1649" box="[1013,1150,1318,1341]" italics="true" pageId="9" pageNumber="1521">F. sagittocerus</emphasis>
</taxonomicName>
, male. Arrow: anterior margin of the upper frons at the edge with lower frons.
</paragraph>
</caption>
<caption id="8AC76653FFD332429A9F4834FCC91600" ID-DOI="http://doi.org/10.5281/zenodo.7318193" ID-Zenodo-Dep="7318193" httpUri="https://zenodo.org/record/7318193/files/figure.png" pageId="10" pageNumber="1522" startId="10.[160,225,1285,1308]" targetBox="[167,1149,145,1251]" targetPageId="10">
<paragraph id="DE0736DBFFD332429A9F4834FCC91600" blockId="10.[160,1157,1285,1396]" pageId="10" pageNumber="1522">
<emphasis id="ECCCEAC9FFD332429A9F4834FF5B1668" bold="true" box="[160,250,1285,1308]" pageId="10" pageNumber="1522">Figure 6.</emphasis>
Heads of 
<taxonomicName id="19B84D58FFD332429B624834FD891668" box="[349,552,1285,1308]" pageId="10" pageNumber="1522">
<emphasis id="ECCCEAC9FFD332429B624834FE791668" box="[349,472,1285,1308]" italics="true" pageId="10" pageNumber="1522">Formosargus</emphasis>
species
</taxonomicName>
, frontal view. (a) 
<taxonomicName id="19B84D58FFD3324298EE4834FCFB1668" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[721,858,1285,1308]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1522" phylum="Arthropoda" rank="species" species="berezovskiyi" status="sp. nov.">
<emphasis id="ECCCEAC9FFD3324298EE4834FCFB1668" box="[721,858,1285,1308]" italics="true" pageId="10" pageNumber="1522">F. berezovskiyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD33242995F4834FC0E1668" box="[864,943,1285,1308]" pageId="10" pageNumber="1522" rank="species">sp. nov.</taxonomicNameLabel>
, female, paratype. (b) 
<taxonomicName id="19B84D58FFD332429A9F4813FE80164D" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[160,289,1314,1337]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1522" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
<emphasis id="ECCCEAC9FFD332429A9F4813FE80164D" box="[160,289,1314,1337]" italics="true" pageId="10" pageNumber="1522">F. borneensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD332429B134813FE20164D" box="[300,385,1314,1337]" pageId="10" pageNumber="1522" rank="species">sp. nov.</taxonomicNameLabel>
, female, holotype. (c) 
<taxonomicName id="19B84D58FFD3324298524813FCAD164D" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[621,780,1314,1337]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1522" phylum="Arthropoda" rank="species" species="mangoleensis" status="sp. nov.">
<emphasis id="ECCCEAC9FFD3324298524813FCAD164D" box="[621,780,1314,1337]" italics="true" pageId="10" pageNumber="1522">F. mangoleensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD3324299284813FCCA164D" box="[791,875,1314,1337]" pageId="10" pageNumber="1522" rank="species">sp. nov.</taxonomicNameLabel>
, female, holotype. (d–e) 
<taxonomicName id="19B84D58FFD332429E4C4813FF5D1623" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1522" phylum="Arthropoda" rank="species" species="trivittatus" status="sp. nov.">
<emphasis id="ECCCEAC9FFD332429E4C4813FF5D1623" italics="true" pageId="10" pageNumber="1522">F. trivittatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD332429B3A4871FEF51623" box="[261,340,1344,1367]" pageId="10" pageNumber="1522" rank="species">sp. nov.</taxonomicNameLabel>
(d. male, holotype; e. female, paratype). (f) 
<taxonomicName id="19B84D58FFD33242992A4871FC221623" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[789,899,1344,1367]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="1522" phylum="Arthropoda" rank="species" species="woodleyi" status="sp. nov.">
<emphasis id="ECCCEAC9FFD33242992A4871FC221623" box="[789,899,1344,1367]" italics="true" pageId="10" pageNumber="1522">F. woodleyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD3324299B34871FC7F1623" box="[908,990,1344,1367]" pageId="10" pageNumber="1522" rank="species">sp. nov.</taxonomicNameLabel>
, male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons.
</paragraph>
</caption>
<paragraph id="DE0736DBFFD232439A9F4F88FF5917AE" blockId="11.[160,1156,142,1727]" pageId="11" pageNumber="1523">
<emphasis id="ECCCEAC9FFD232439A9F4F88FEB411A6" bold="true" box="[160,277,697,722]" pageId="11" pageNumber="1523">Abdomen</emphasis>
. Pale yellow to reddish yellow in background, with at least a pair of roughly subrectangular dark brown spots on tergite 1, and tergites 2–5 mostly yellowish, with a pair of proximal or medial dark brown spots to nearly entirely dark; more than twice longer than broad, wider at about fourth segment; first segment slightly shorter than remaining segments, parallel-sided; divergent margins on segments 2–4, convergent margins on segment 5; pilosity short and yellowish. 
<emphasis id="ECCCEAC9FFD2324398CE4E57FCD1100B" bold="true" box="[753,880,870,895]" pageId="11" pageNumber="1523">Terminalia</emphasis>
(
<figureCitation id="46832A5EFFD2324399BE4E57FBBF100B" box="[897,1054,870,895]" captionStart-0="Figure 10" captionStart-1="Figure 11" captionStart-2="Figure 12" captionStart-3="Figure 13" captionStart-4="Figure 14" captionStartId-0="18.[160,225,1626,1649]" captionStartId-1="19.[160,225,1550,1573]" captionStartId-2="21.[160,225,1549,1572]" captionStartId-3="23.[160,225,1550,1573]" captionStartId-4="25.[160,225,1550,1573]" captionTargetBox-0="[173,1142,148,1584]" captionTargetBox-1="[208,1106,144,1515]" captionTargetBox-2="[207,1108,144,1516]" captionTargetBox-3="[208,1105,143,1516]" captionTargetBox-4="[208,1106,144,1516]" captionTargetId-0="figure-3@18.[163,1153,142,1594]" captionTargetId-1="figure-1@19.[206,1109,142,1518]" captionTargetId-2="figure-1@21.[205,1110,142,1518]" captionTargetId-3="figure-1@23.[207,1108,142,1518]" captionTargetId-4="figure-1@25.[207,1108,142,1518]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionTargetPageId-2="21" captionTargetPageId-3="23" captionTargetPageId-4="25" captionText-0="Figure 10. Male terminalia of Formosargus species (epandrium, proctiger, and cerci removed). (a–b) F. lineatus (a. dorsal view; b. lateral view). (c–e) F. trivittatus sp. nov., holotype (c. dorsal view; d. ventral view; e. lateral view). (f) F. woodleyi sp. nov., lateral view, holotype. Abbreviations: goncx apod, gonocoxal apodeme; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath. Scale bars: 0.1 mm." captionText-1="Figure 11. Male terminalia of Formosargus lineatus. (a) Genitalia, dorsal view; arrow: reduced gonocoxal apodeme. (b) Genitalia, lateral view; arrow: inner projection of synsternite. (c) Genitalia, ventral view; arrow: medial line. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; gonst, gonostylus; ph, phallus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger." captionText-2="Figure 12. Male terminalia of Formosargus sagittocerus. (a) Genitalia,dorsal view;arrow:reduced gonocoxal apodeme. (b) Genitalia, lateral view. (c) Genitalia, ventral view. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; goncx brg, gonocoxal bridge; gonst, gonostylus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger." captionText-3="Figure 13. Male terminalia of Formosargus trivittatus sp. nov., holotype. (a) Genitalia, dorsal view; arrow: reduced gonocoxal apodeme. (b) Genitalia, lateral view. (c) Genitalia, ventral view. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger." captionText-4="Figure 14. Male terminalia of Formosargus woodleyi sp. nov., holotype. (a) Genitalia, dorsal view; arrow: reduced gonocoxal apodeme. (b) Genitalia, lateral view. (c) Genitalia, ventral view. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath; prct, proctiger." figureDoi-0="http://doi.org/10.5281/zenodo.7318203" figureDoi-1="http://doi.org/10.5281/zenodo.7318205" figureDoi-2="http://doi.org/10.5281/zenodo.7318207" figureDoi-3="http://doi.org/10.5281/zenodo.7318209" figureDoi-4="http://doi.org/10.5281/zenodo.7318211" httpUri-0="https://zenodo.org/record/7318203/files/figure.png" httpUri-1="https://zenodo.org/record/7318205/files/figure.png" httpUri-2="https://zenodo.org/record/7318207/files/figure.png" httpUri-3="https://zenodo.org/record/7318209/files/figure.png" httpUri-4="https://zenodo.org/record/7318211/files/figure.png" pageId="11" pageNumber="1523">Figures 10–14</figureCitation>
). Genital capsule subrectangular, slightly taller than wide; distal margin weakly developed dorsally, exceeding ventral margin height; ventral margin of synsternite clearly U-shaped, often with projections medially; synsternite with a pair of well-developed inner projections, ventral to phallus, curved at base, parallel on distal two-thirds. Gonocoxal apodeme very reduced; gonocoxal bridge placed at distal margin of genitalia. Phallus trifid, with very shortened lobes; parameral sheath roughly elliptical or subrectangular, nearly entirely encapsulating the phallus; both very reduced in size, placed at distal half of genitalia. Gonostylus wide, varied in shape, with expansions or bilobed. Epandrium wider than long. Proctiger (tergite 10) well developed, as long as cercus, triangular; cercus slender, rodshaped.
</paragraph>
<paragraph id="DE0736DBFFD232439AFF49D5FB251611" blockId="11.[160,1156,142,1727]" pageId="11" pageNumber="1523">
<emphasis id="ECCCEAC9FFD232439AFF49D5FEB91789" bold="true" box="[192,280,1252,1277]" pageId="11" pageNumber="1523">Female</emphasis>
. Similar to male, except as follows: Length (mm): body, 5.0–10.0; wing, 5.0–9.0. 
<emphasis id="ECCCEAC9FFD232439B3B4837FEE2166B" bold="true" box="[260,323,1286,1311]" pageId="11" pageNumber="1523">Head</emphasis>
(
<figureCitation id="46832A5EFFD232439B664837FE42166B" box="[345,483,1286,1311]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-2="Figure 5" captionStart-3="Figure 6" captionStartId-0="7.[160,225,906,929]" captionStartId-1="8.[160,225,1266,1289]" captionStartId-2="9.[160,225,1289,1312]" captionStartId-3="10.[160,225,1285,1308]" captionTargetBox-0="[167,1149,145,869]" captionTargetBox-1="[167,1149,145,1232]" captionTargetBox-2="[167,1149,145,1251]" captionTargetBox-3="[167,1149,145,1251]" captionTargetId-0="figure-267@7.[163,1153,142,874]" captionTargetId-1="figure-139@8.[163,1153,142,1234]" captionTargetId-2="figure-113@9.[163,1153,142,1257]" captionTargetId-3="figure-99@10.[163,1153,142,1253]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="Figure 3. Heads of Formosargus species, dorsal view. (a) F. kerteszi, female. (b–c) F. lineatus (b. male; c. female). (d) F. sagittocerus, male." captionText-1="Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype." captionText-2="Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons." captionText-3="Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi-0="http://doi.org/10.5281/zenodo.7318186" figureDoi-1="http://doi.org/10.5281/zenodo.7318188" figureDoi-2="http://doi.org/10.5281/zenodo.7318191" figureDoi-3="http://doi.org/10.5281/zenodo.7318193" httpUri-0="https://zenodo.org/record/7318186/files/figure.png" httpUri-1="https://zenodo.org/record/7318188/files/figure.png" httpUri-2="https://zenodo.org/record/7318191/files/figure.png" httpUri-3="https://zenodo.org/record/7318193/files/figure.png" pageId="11" pageNumber="1523">Figures 3–6</figureCitation>
). Frons, frontal callus and face wider than one-fifth of head. Face much shorter than lower frons or at most as long as. 
<emphasis id="ECCCEAC9FFD23243999F4818FC541636" bold="true" box="[928,1013,1321,1346]" pageId="11" pageNumber="1523">Thorax</emphasis>
(
<figureCitation id="46832A5EFFD232439E394818FBC81636" box="[1030,1129,1321,1346]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="11" pageNumber="1523">Figure 7</figureCitation>
(b, e–g)). Scutal bands or maculae over scutum often slender and less intense in colour.
</paragraph>
<paragraph id="DE0736DBFFD232459A9F485FFEAA15DE" blockId="11.[160,1156,142,1727]" lastBlockId="13.[160,1157,1542,1706]" lastPageId="13" lastPageNumber="1525" pageId="11" pageNumber="1523">
<emphasis id="ECCCEAC9FFD232439A9F485FFEB616F3" bold="true" box="[160,279,1390,1415]" pageId="11" pageNumber="1523">Abdomen</emphasis>
. Mostly pale in colouration, varying from completely yellow to well maculated with dark brown on each visible tergite at centre or distally; spots of tergites 2– 5 often weakly or not connected medially; clearly oval, with divergent margins on segments 1–2, parallel margins on segment 3, convergent margins on segments 4–5. 
<emphasis id="ECCCEAC9FFD232439A9F48C8FE821566" bold="true" box="[160,291,1529,1554]" pageId="11" pageNumber="1523">Terminalia</emphasis>
(
<figureCitation id="46832A5EFFD232439B0648C8FE431566" box="[313,482,1529,1554]" captionStart-0="Figure 15" captionStart-1="Figure 16" captionStart-2="Figure 17" captionStartId-0="27.[160,225,1701,1724]" captionStartId-1="28.[160,226,964,987]" captionStartId-2="29.[160,225,1383,1406]" captionTargetBox-0="[210,1104,144,1666]" captionTargetBox-1="[164,1149,144,929]" captionTargetBox-2="[166,1150,145,1348]" captionTargetId-0="figure-1@27.[208,1107,142,1669]" captionTargetId-1="figure-182@28.[162,1152,142,932]" captionTargetId-2="figure-42@29.[163,1153,142,1351]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionTargetPageId-2="29" captionText-0="Figure 15. Female terminalia of Formosargus species, dorsal view. (a) F. kerteszi. (b) F. lineatus. (c) F. berezovskiyi sp. nov., paratype. (d) F. borneensis sp. nov., holotype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite." captionText-1="Figure 16. Female terminalia of Formosargus species, dorsal view. (a) F. mangoleensis sp. nov, holotype. (b) F. trivittatus sp. nov., paratype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite." captionText-2="Figure 17. Genital fork (sternite 9) of Formosargus species. (a–b). F. kerteszi (a. dorsal view; b. ventral view). (c–d). F. lineatus (c. dorsal view; d. ventral view). (e–f) F. berezovskiyi sp. nov., holotype (e. dorsal view; f. ventral view). (g–h) F. borneensis sp. nov., holotype (g. dorsal view; h. ventral view). (i–j) F. mangoleensis sp. nov., holotype (i. dorsal view; j. ventral view). (k–l) F. trivittatus sp. nov., paratype (k. dorsal view;l. ventral view). Abbreviations: go, genital opening;pb, posterior bridge; pp, posterolateral process." figureDoi-0="http://doi.org/10.5281/zenodo.7318213" figureDoi-1="http://doi.org/10.5281/zenodo.7318216" figureDoi-2="http://doi.org/10.5281/zenodo.7318220" httpUri-0="https://zenodo.org/record/7318213/files/figure.png" httpUri-1="https://zenodo.org/record/7318216/files/figure.png" httpUri-2="https://zenodo.org/record/7318220/files/figure.png" pageId="11" pageNumber="1523">Figures 15–17</figureCitation>
). Typical of sargines, mostly differing in the shape of genital fork. Tergite 8 rectangular, nearly twice longer than wide; sternite 8 wider than tergite 8, subtriangular on its distal fourth. Tergite 9 sclerite strongly chitinised laterally. Genital fork (sternite 9) often wider posteriorly, with anterior half gradually (
<figureCitation id="46832A5EFFD232439A974BB5FEBA15E9" box="[168,283,1668,1693]" captionStart="Figure 17" captionStartId="29.[160,225,1383,1406]" captionTargetBox="[166,1150,145,1348]" captionTargetId="figure-42@29.[163,1153,142,1351]" captionTargetPageId="29" captionText="Figure 17. Genital fork (sternite 9) of Formosargus species. (a–b). F. kerteszi (a. dorsal view; b. ventral view). (c–d). F. lineatus (c. dorsal view; d. ventral view). (e–f) F. berezovskiyi sp. nov., holotype (e. dorsal view; f. ventral view). (g–h) F. borneensis sp. nov., holotype (g. dorsal view; h. ventral view). (i–j) F. mangoleensis sp. nov., holotype (i. dorsal view; j. ventral view). (k–l) F. trivittatus sp. nov., paratype (k. dorsal view;l. ventral view). Abbreviations: go, genital opening;pb, posterior bridge; pp, posterolateral process." figureDoi="http://doi.org/10.5281/zenodo.7318220" httpUri="https://zenodo.org/record/7318220/files/figure.png" pageId="11" pageNumber="1523">Figure 17</figureCitation>
(a)) or only basal third abruptly (
<figureCitation id="46832A5EFFD2324398874BB5FC8A15E9" box="[696,811,1668,1693]" captionStart="Figure 17" captionStartId="29.[160,225,1383,1406]" captionTargetBox="[166,1150,145,1348]" captionTargetId="figure-42@29.[163,1153,142,1351]" captionTargetPageId="29" captionText="Figure 17. Genital fork (sternite 9) of Formosargus species. (a–b). F. kerteszi (a. dorsal view; b. ventral view). (c–d). F. lineatus (c. dorsal view; d. ventral view). (e–f) F. berezovskiyi sp. nov., holotype (e. dorsal view; f. ventral view). (g–h) F. borneensis sp. nov., holotype (g. dorsal view; h. ventral view). (i–j) F. mangoleensis sp. nov., holotype (i. dorsal view; j. ventral view). (k–l) F. trivittatus sp. nov., paratype (k. dorsal view;l. ventral view). Abbreviations: go, genital opening;pb, posterior bridge; pp, posterolateral process." figureDoi="http://doi.org/10.5281/zenodo.7318220" httpUri="https://zenodo.org/record/7318220/files/figure.png" pageId="11" pageNumber="1523">Figure 17</figureCitation>
(c)) narrowing into an often straight anterior end; posterior bridge from nearly straight (
<figureCitation id="46832A5EFFD23243995E4B97FC7015CB" box="[865,977,1702,1727]" captionStart="Figure 17" captionStartId="29.[160,225,1383,1406]" captionTargetBox="[166,1150,145,1348]" captionTargetId="figure-42@29.[163,1153,142,1351]" captionTargetPageId="29" captionText="Figure 17. Genital fork (sternite 9) of Formosargus species. (a–b). F. kerteszi (a. dorsal view; b. ventral view). (c–d). F. lineatus (c. dorsal view; d. ventral view). (e–f) F. berezovskiyi sp. nov., holotype (e. dorsal view; f. ventral view). (g–h) F. borneensis sp. nov., holotype (g. dorsal view; h. ventral view). (i–j) F. mangoleensis sp. nov., holotype (i. dorsal view; j. ventral view). (k–l) F. trivittatus sp. nov., paratype (k. dorsal view;l. ventral view). Abbreviations: go, genital opening;pb, posterior bridge; pp, posterolateral process." figureDoi="http://doi.org/10.5281/zenodo.7318220" httpUri="https://zenodo.org/record/7318220/files/figure.png" pageId="11" pageNumber="1523">Figure 17</figureCitation>
(k, l)) to weakly bilobed (e.g. 
<figureCitation id="46832A5EFFD432459B014B37FE12156B" box="[318,435,1542,1567]" captionStart="Figure 17" captionStartId="29.[160,225,1383,1406]" captionTargetBox="[166,1150,145,1348]" captionTargetId="figure-42@29.[163,1153,142,1351]" captionTargetPageId="29" captionText="Figure 17. Genital fork (sternite 9) of Formosargus species. (a–b). F. kerteszi (a. dorsal view; b. ventral view). (c–d). F. lineatus (c. dorsal view; d. ventral view). (e–f) F. berezovskiyi sp. nov., holotype (e. dorsal view; f. ventral view). (g–h) F. borneensis sp. nov., holotype (g. dorsal view; h. ventral view). (i–j) F. mangoleensis sp. nov., holotype (i. dorsal view; j. ventral view). (k–l) F. trivittatus sp. nov., paratype (k. dorsal view;l. ventral view). Abbreviations: go, genital opening;pb, posterior bridge; pp, posterolateral process." figureDoi="http://doi.org/10.5281/zenodo.7318220" httpUri="https://zenodo.org/record/7318220/files/figure.png" pageId="13" pageNumber="1525">Figure 17</figureCitation>
(i, j)); posterolateral process curved at basal third, parallel on apical two-thirds; genital opening reduced, from elliptical to nearly circular-shaped, displaced posteriorly on the genital fork. Tergite 10 well developed, triangular. Cercus two-segmented; first segment of cercus nearly twice as long as second segment.
</paragraph>
</subSubSection>
<caption id="8AC76653FFD532449A9F4B16FB2415C7" ID-DOI="http://doi.org/10.5281/zenodo.7318195" ID-Zenodo-Dep="7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="12" pageNumber="1524" startId="12.[160,225,1575,1598]" targetBox="[167,1149,145,1537]" targetPageId="12">
<paragraph id="DE0736DBFFD532449A9F4B16FB2415C7" blockId="12.[160,1157,1575,1715]" pageId="12" pageNumber="1524">
<emphasis id="ECCCEAC9FFD532449A9F4B16FF5C154A" bold="true" box="[160,253,1575,1598]" pageId="12" pageNumber="1524">Figure 7.</emphasis>
Thorax of 
<taxonomicName id="19B84D58FFD532449B4E4B16FD9F154A" box="[369,574,1575,1598]" pageId="12" pageNumber="1524">
<emphasis id="ECCCEAC9FFD532449B4E4B16FE4D154A" box="[369,492,1575,1598]" italics="true" pageId="12" pageNumber="1524">Formosargus</emphasis>
species
</taxonomicName>
, dorsal view. (a–b) 
<taxonomicName id="19B84D58FFD5324499364B16FCCB154A" authorityName="James" authorityYear="1939" box="[777,874,1575,1598]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="1524" phylum="Arthropoda" rank="species" species="kerteszi">
<emphasis id="ECCCEAC9FFD5324499364B16FCCB154A" box="[777,874,1575,1598]" italics="true" pageId="12" pageNumber="1524">F. kerteszi</emphasis>
</taxonomicName>
(a. male, holotype [DEI]; b. female, Philippines). (c) 
<taxonomicName id="19B84D58FFD532449BB34B75FE4D152F" box="[396,492,1604,1627]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="1524" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="ECCCEAC9FFD532449BB34B75FE4D152F" box="[396,492,1604,1627]" italics="true" pageId="12" pageNumber="1524">F. lineatus</emphasis>
</taxonomicName>
, male; arrow: medial band. (d) 
<taxonomicName id="19B84D58FFD53244991D4B75FC0A152F" authorityName="Fachin &amp; Hauser" authorityYear="2022" baseAuthorityName="Adisoemarto" baseAuthorityYear="1974" box="[802,939,1604,1627]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="1524" phylum="Arthropoda" rank="species" species="sagittocerus">
<emphasis id="ECCCEAC9FFD53244991D4B75FC0A152F" box="[802,939,1604,1627]" italics="true" pageId="12" pageNumber="1524">F. sagittocerus</emphasis>
</taxonomicName>
, male. (e) 
<taxonomicName id="19B84D58FFD532449E2C4B75FF61150C" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="1524" phylum="Arthropoda" rank="species" species="berezovskiyi" status="sp. nov.">
<emphasis id="ECCCEAC9FFD532449E2C4B75FF61150C" italics="true" pageId="12" pageNumber="1524">F. berezovskiyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD532449AF94B50FEB7150C" box="[198,278,1633,1656]" pageId="12" pageNumber="1524" rank="species">sp. nov.</taxonomicNameLabel>
, female, paratype; arrow: mediotergite. (f) 
<taxonomicName id="19B84D58FFD5324498854B50FC97150C" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[698,822,1633,1656]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="1524" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
<emphasis id="ECCCEAC9FFD5324498854B50FC97150C" box="[698,822,1633,1656]" italics="true" pageId="12" pageNumber="1524">F. borneensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD5324499034B50FC2D150C" box="[828,908,1633,1656]" pageId="12" pageNumber="1524" rank="species">sp. nov.</taxonomicNameLabel>
, female, holotype; arrow: notopleural macula. (g) 
<taxonomicName id="19B84D58FFD532449B9F4B4EFDE015E2" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[416,577,1663,1686]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="1524" phylum="Arthropoda" rank="species" species="mangoleensis" status="sp. nov.">
<emphasis id="ECCCEAC9FFD532449B9F4B4EFDE015E2" box="[416,577,1663,1686]" italics="true" pageId="12" pageNumber="1524">F. mangoleensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD5324498734B4EFD0315E2" box="[588,674,1663,1686]" pageId="12" pageNumber="1524" rank="species">sp. nov.</taxonomicNameLabel>
, female, holotype; arrow: mediotergite. (h) 
<taxonomicName id="19B84D58FFD532449E4C4B4EFF5D15C7" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="1524" phylum="Arthropoda" rank="species" species="trivittatus" status="sp. nov.">
<emphasis id="ECCCEAC9FFD532449E4C4B4EFF5D15C7" italics="true" pageId="12" pageNumber="1524">F. trivittatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD532449B3E4BADFEF115C7" box="[257,336,1692,1715]" pageId="12" pageNumber="1524" rank="species">sp. nov.</taxonomicNameLabel>
, male, holotype; arrow: anterior lateral band. (i) 
<taxonomicName id="19B84D58FFD53244991A4BADFC3115C7" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[805,912,1692,1715]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="1524" phylum="Arthropoda" rank="species" species="woodleyi" status="sp. nov.">
<emphasis id="ECCCEAC9FFD53244991A4BADFC3115C7" box="[805,912,1692,1715]" italics="true" pageId="12" pageNumber="1524">F. woodleyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD5324499A94BADFC4515C7" box="[918,996,1692,1715]" pageId="12" pageNumber="1524" rank="species">sp. nov.</taxonomicNameLabel>
, male, holotype.
</paragraph>
</caption>
<caption id="8AC76653FFD432459A9F4842FD2616B0" ID-DOI="http://doi.org/10.5281/zenodo.7318197" ID-Zenodo-Dep="7318197" httpUri="https://zenodo.org/record/7318197/files/figure.png" pageId="13" pageNumber="1525" startId="13.[160,225,1395,1418]" targetBox="[167,1149,145,1360]" targetPageId="13">
<paragraph id="DE0736DBFFD432459A9F4842FD2616B0" blockId="13.[160,1157,1395,1476]" pageId="13" pageNumber="1525">
<emphasis id="ECCCEAC9FFD432459A9F4842FF5F16FE" bold="true" box="[160,254,1395,1418]" pageId="13" pageNumber="1525">Figure 8.</emphasis>
Male mid legs of 
<taxonomicName id="19B84D58FFD432459BFD4842FD3016FE" box="[450,657,1395,1418]" pageId="13" pageNumber="1525">
<emphasis id="ECCCEAC9FFD432459BFD4842FD9C16FE" box="[450,573,1395,1418]" italics="true" pageId="13" pageNumber="1525">Formosargus</emphasis>
species
</taxonomicName>
, male. (a–b) 
<taxonomicName id="19B84D58FFD4324599244842FCDD16FE" authorityName="James" authorityYear="1939" box="[795,892,1395,1418]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="1525" phylum="Arthropoda" rank="species" species="kerteszi">
<emphasis id="ECCCEAC9FFD4324599244842FCDD16FE" box="[795,892,1395,1418]" italics="true" pageId="13" pageNumber="1525">F. kerteszi</emphasis>
</taxonomicName>
. (c–e) 
<taxonomicName id="19B84D58FFD4324599F94842FB8B16FE" box="[966,1066,1395,1418]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="1525" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="ECCCEAC9FFD4324599F94842FB8B16FE" box="[966,1066,1395,1418]" italics="true" pageId="13" pageNumber="1525">F. lineatus</emphasis>
</taxonomicName>
. (f–h) 
<taxonomicName id="19B84D58FFD432459E4C4842FEB316D3" authorityName="Fachin &amp; Hauser" authorityYear="2022" baseAuthorityName="Adisoemarto" baseAuthorityYear="1974" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="1525" phylum="Arthropoda" rank="species" species="sagittocerus">
<emphasis id="ECCCEAC9FFD432459E4C4842FEB316D3" italics="true" pageId="13" pageNumber="1525">F. sagittocerus</emphasis>
</taxonomicName>
. (i–j) 
<taxonomicName id="19B84D58FFD432459B6C48A1FE6216D3" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[339,451,1424,1447]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="1525" phylum="Arthropoda" rank="species" species="woodleyi" status="sp. nov.">
<emphasis id="ECCCEAC9FFD432459B6C48A1FE6216D3" box="[339,451,1424,1447]" italics="true" pageId="13" pageNumber="1525">F. woodleyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD432459BF148A1FD8316D3" box="[462,546,1424,1447]" pageId="13" pageNumber="1525" rank="species">sp. nov.</taxonomicNameLabel>
, holotype. Arrow: modified structure at base of basitarsus. Abbreviations: fem, femur; tsm 1, first tarsomere.
</paragraph>
</caption>
<subSubSection id="96A26550FFD732469A9F4DBFFCD71247" pageId="14" pageNumber="1526" type="distribution">
<paragraph id="DE0736DBFFD732469A9F4DBFFE1813DC" blockId="14.[160,1157,142,307]" box="[160,441,142,168]" pageId="14" pageNumber="1526">
<emphasis id="ECCCEAC9FFD732469A9F4DBFFE1813DC" bold="true" box="[160,441,142,168]" italics="true" pageId="14" pageNumber="1526">Geographic distribution</emphasis>
</paragraph>
<paragraph id="DE0736DBFFD732469A9F4D83FCD71247" blockId="14.[160,1157,142,307]" pageId="14" pageNumber="1526">
<emphasis id="ECCCEAC9FFD732469A9F4D83FEFC13BF" bold="true" box="[160,349,178,203]" pageId="14" pageNumber="1526">Oriental Region</emphasis>
: 
<collectingCountry id="A6AF764BFFD732469B554D83FE1B13BF" box="[362,442,178,203]" name="Taiwan" pageId="14" pageNumber="1526">Taiwan</collectingCountry>
(Pingtung; 
<collectingCounty id="37664E57FFD7324698014D83FD7613BF" box="[574,727,178,203]" pageId="14" pageNumber="1526">Orchid Island</collectingCounty>
), 
<collectingCountry id="A6AF764BFFD7324698D74D83FCC213BF" box="[744,867,178,203]" name="Philippines" pageId="14" pageNumber="1526">Philippines</collectingCountry>
(Benguet; Eastern Samar: 
<collectingMunicipality id="3E63ACA1FFD732469A9F4DE5FEED1399" box="[160,332,212,237]" pageId="14" pageNumber="1526">Calicoan Island</collectingMunicipality>
; Laguna; Palawan), 
<collectingCountry id="A6AF764BFFD7324698174DE5FD291399" box="[552,648,212,237]" name="Thailand" pageId="14" pageNumber="1526">Thailand</collectingCountry>
(Trang), 
<collectingCountry id="A6AF764BFFD7324698D34DE5FCED1399" box="[748,844,212,237]" name="Malaysia" pageId="14" pageNumber="1526">Malaysia</collectingCountry>
(Pahang; Sabah), 
<collectingCountry id="A6AF764BFFD732469E294DE5FB221399" box="[1046,1155,212,237]" name="Indonesia" pageId="14" pageNumber="1526">Indonesia</collectingCountry>
(North Maluku: 
<location id="DB676000FFD732469B744DC6FDB21264" LSID="urn:lsid:plazi:treatment:561187CDFFDC32589BD648DBFB25177F:DB676000FFD732469B744DC6FDB21264" box="[331,531,247,272]" country="Taiwan" county="Orchid Island" municipality="Calicoan Island" name="Island of Mangole" pageId="14" pageNumber="1526" stateProvince="Queensland">Island of Mangole</location>
; West Java); 
<emphasis id="ECCCEAC9FFD7324698A44DC6FCCE1264" bold="true" box="[667,879,247,272]" pageId="14" pageNumber="1526">
<location id="DB676000FFD7324698A44DC6FCCE1264" LSID="urn:lsid:plazi:treatment:561187CDFFDC32589BD648DBFB25177F:DB676000FFD7324698A44DC6FCCE1264" box="[667,879,247,272]" country="Taiwan" county="Orchid Island" municipality="Calicoan Island" name="Australian Region" pageId="14" pageNumber="1526" stateProvince="Queensland">Australian Region</location>
</emphasis>
: 
<collectingCountry id="A6AF764BFFD7324699454DC6FC461264" box="[890,999,247,272]" name="Indonesia" pageId="14" pageNumber="1526">Indonesia</collectingCountry>
(West Papua), 
<collectingCountry id="A6AF764BFFD732469A9F4C2BFED71247" box="[160,374,282,307]" name="Papua New Guinea" pageId="14" pageNumber="1526">Papua New Guinea</collectingCountry>
(Morobe); 
<collectingCountry id="A6AF764BFFD732469BCD4C2BFDF21247" box="[498,595,282,307]" name="Australia" pageId="14" pageNumber="1526">Australia</collectingCountry>
(
<collectingRegion id="1C7CF839FFD73246985E4C2BFD4A1247" box="[609,747,282,307]" country="Australia" name="Queensland" pageId="14" pageNumber="1526">Queensland</collectingRegion>
) (
<figureCitation id="46832A5EFFD7324698C04C2BFCC81247" box="[767,873,282,307]" captionStart="Figure 26" captionStartId="38.[160,226,870,893]" captionTargetBox="[168,1148,146,834]" captionTargetId="figure-167@38.[163,1153,142,839]" captionTargetPageId="38" captionText="Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line." figureDoi="http://doi.org/10.5281/zenodo.7318240" httpUri="https://zenodo.org/record/7318240/files/figure.png" pageId="14" pageNumber="1526">Figure 26</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="96A26550FFD732479A9F4C6FFB201063" lastPageId="15" lastPageNumber="1527" pageId="14" pageNumber="1526" type="discussion">
<paragraph id="DE0736DBFFD732469A9F4C6FFEBF120C" blockId="14.[160,1156,350,1728]" box="[160,286,350,376]" pageId="14" pageNumber="1526">
<emphasis id="ECCCEAC9FFD732469A9F4C6FFEBF120C" bold="true" box="[160,286,350,376]" italics="true" pageId="14" pageNumber="1526">Comments</emphasis>
</paragraph>
<paragraph id="DE0736DBFFD732469A9F4CB3FCF4111F" blockId="14.[160,1156,350,1728]" pageId="14" pageNumber="1526">
The Malagasy 
<taxonomicName id="19B84D58FFD732469B024CB3FDD212EF" authority="Lindner, 1959" authorityName="Lindner" authorityYear="1959" box="[317,627,386,411]" class="Insecta" family="Stratiomyidae" genus="Gongrosargus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD732469B024CB3FE7512EF" box="[317,468,386,411]" italics="true" pageId="14" pageNumber="1526">Gongrosargus</emphasis>
Lindner, 1959
</taxonomicName>
is the only other genus of 
<taxonomicName id="19B84D58FFD7324699A74CB3FC5812EF" authorityName="Walker" authorityYear="1834" box="[920,1017,386,411]" class="Insecta" family="Stratiomyidae" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="subFamily" subFamily="Sarginae">Sarginae</taxonomicName>
that has the M4 vein branching off from the discal cell in the wing, with m–cu crossvein connected to the fork of M3+4 (
<bibRefCitation id="BA294B2AFFD732469B614CF6FD991294" author="Hauser M &amp; Woodley NE &amp; Fachin DA" box="[350,568,455,480]" pageId="14" pageNumber="1526" pagination="919 - 979" refId="ref18907" refString="Hauser M, Woodley NE, Fachin DA. 2017 b. 41. Stratiomyidae (Soldier flies). In: Kirk-Spriggs AH, Sinclair BJ, editors. Manual of Afrotropical Diptera. Vol. 2. Nematocerous Diptera and lower Brachycera. Suricata 5. Pretoria: South African National Biodiversity Institute; p. 919 - 979." type="book chapter" year="2017">Hauser et al. 2017b</bibRefCitation>
, p. 965, fig. 182). These, however, are very large flies and distinct due to the presence of more than five antennal flagellomeres (see 
<bibRefCitation id="BA294B2AFFD732469E274CDBFEA51152" author="Hauser M &amp; Woodley NE &amp; Fachin DA" pageId="14" pageNumber="1526" pagination="919 - 979" refId="ref18907" refString="Hauser M, Woodley NE, Fachin DA. 2017 b. 41. Stratiomyidae (Soldier flies). In: Kirk-Spriggs AH, Sinclair BJ, editors. Manual of Afrotropical Diptera. Vol. 2. Nematocerous Diptera and lower Brachycera. Suricata 5. Pretoria: South African National Biodiversity Institute; p. 919 - 979." type="book chapter" year="2017">Hauser et al. 2017b</bibRefCitation>
, p. 957, fig. 130), while all remaining genera of 
<taxonomicName id="19B84D58FFD7324699384F3CFCC91152" authorityName="Walker" authorityYear="1834" box="[775,872,525,550]" class="Insecta" family="Stratiomyidae" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="subFamily" subFamily="Sarginae">Sarginae</taxonomicName>
have only five. Therefore, this may indicate that the genus does not belong to the 
<taxonomicName id="19B84D58FFD73246990B4F1EFC39113C" authorityName="Walker" authorityYear="1834" box="[820,920,559,584]" class="Insecta" family="Stratiomyidae" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="subFamily" subFamily="Sarginae">Sarginae</taxonomicName>
, but may be instead more closely related to the Chrysochlorinae and 
<taxonomicName id="19B84D58FFD7324698FA4F63FCF0111F" authorityName="Loew" authorityYear="1862" box="[709,849,594,619]" class="Insecta" family="Stratiomyidae" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="subFamily" subFamily="Hermetiinae">Hermetiinae</taxonomicName>
.
</paragraph>
<paragraph id="DE0736DBFFD732469AFF4F44FCB5178A" blockId="14.[160,1156,350,1728]" pageId="14" pageNumber="1526">
<bibRefCitation id="BA294B2AFFD732469AFF4F44FE0111FA" author="Adisoemarto S" box="[192,416,629,654]" pageId="14" pageNumber="1526" pagination="69 - 71" refId="ref18465" refString="Adisoemarto S. 1974. A new genus of soldier flies from Indonesia (Diptera: Stratiomyidae). Treubia. 28 (3): 69 - 71." type="journal article" year="1974">Adisoemarto (1974)</bibRefCitation>
compared his new genus 
<taxonomicName id="19B84D58FFD7324698E34F44FC9411FA" authorityName="Adisoemarto" authorityYear="1974" box="[732,821,629,654]" class="Insecta" family="Stratiomyidae" genus="Amsaria" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD7324698E34F44FC9411FA" box="[732,821,629,654]" italics="true" pageId="14" pageNumber="1526">Amsaria</emphasis>
</taxonomicName>
with the genera 
<taxonomicName id="19B84D58FFD732469E3A4F44FBEF11FA" authorityName="Fabricius" authorityYear="1798" box="[1029,1102,629,654]" class="Insecta" family="Stratiomyidae" genus="Sargus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD732469E3A4F44FBEF11FA" box="[1029,1102,629,654]" italics="true" pageId="14" pageNumber="1526">Sargus</emphasis>
</taxonomicName>
and 
<taxonomicName id="19B84D58FFD732469A9F4FA6FF5E11C4" authorityName="Loew" authorityYear="1855" box="[160,255,663,688]" class="Insecta" family="Stratiomyidae" genus="Ptecticus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD732469A9F4FA6FF5E11C4" box="[160,255,663,688]" italics="true" pageId="14" pageNumber="1526">Ptecticus</emphasis>
</taxonomicName>
and pointed out it could be distinguished from the latter two by the shape of the antenna, wing veins and thorax colouration. This is in part correct. 
<taxonomicName id="19B84D58FFD7324699B24F8BFBBB11A7" authorityName="James" authorityYear="1939" box="[909,1050,698,723]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD7324699B24F8BFBBB11A7" box="[909,1050,698,723]" italics="true" pageId="14" pageNumber="1526">Formosargus</emphasis>
</taxonomicName>
does not have a thumb-like projection of the pedicel towards the inner face of the flagellar complex as seen in 
<taxonomicName id="19B84D58FFD732469B244FCEFEDB106C" authorityName="Loew" authorityYear="1855" box="[283,378,767,792]" class="Insecta" family="Stratiomyidae" genus="Ptecticus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD732469B244FCEFEDB106C" box="[283,378,767,792]" italics="true" pageId="14" pageNumber="1526">Ptecticus</emphasis>
</taxonomicName>
(e.g. 
<bibRefCitation id="BA294B2AFFD732469B844FCEFD42106C" author="Rozkosny R &amp; Kovac D" box="[443,739,767,792]" pageId="14" pageNumber="1526" pagination="75 - 86" refId="ref19757" refString="Rozkosny R, Kovac D. 1994. Adults and larvae of two Ptecticus Loew from Peninsular Malaysia (Diptera, Stratiomyidae). Tijdschrift voor Entomologie. 137: 75 - 86." type="journal article" year="1994">Rozkošný and Kovac 1994</bibRefCitation>
, p. 78, fig. 16); and in some species, including that described by Adisoemarto, the fourth flagellomere alone can be as broad as the remaining flagellomeres (see his p. 70, fig. 4), a feature that excludes it from 
<taxonomicName id="19B84D58FFD732469E224E75FBC71029" authorityName="Fabricius" authorityYear="1798" box="[1053,1126,836,861]" class="Insecta" family="Stratiomyidae" genus="Sargus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD732469E224E75FBC71029" box="[1053,1126,836,861]" italics="true" pageId="14" pageNumber="1526">Sargus</emphasis>
</taxonomicName>
as well. In respect of the scutum colour, the presence of a single medial dark band is indeed rather unique for most species of the genus, but some 
<taxonomicName id="19B84D58FFD7324699194EBBFBAC10D7" box="[806,1037,906,931]" pageId="14" pageNumber="1526">
<emphasis id="ECCCEAC9FFD7324699194EBBFC1210D7" box="[806,947,906,931]" italics="true" pageId="14" pageNumber="1526">Formosargus</emphasis>
species
</taxonomicName>
have also additional lateral markings, bands, or a completely dark scutum. In the Oriental Region, a three-banded scutum can be seen in 
<taxonomicName id="19B84D58FFD7324698764EFEFD09109C" authorityName="Loew" authorityYear="1855" box="[585,680,975,1000]" class="Insecta" family="Stratiomyidae" genus="Ptecticus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD7324698764EFEFD09109C" box="[585,680,975,1000]" italics="true" pageId="14" pageNumber="1526">Ptecticus</emphasis>
</taxonomicName>
, for example in 
<taxonomicName id="19B84D58FFD73246995F4EFEFE8D177F" authority="Rozkosny and Kovac, 1996" authorityName="Rozkosny and Kovac" authorityYear="1996" class="Insecta" family="Stratiomyidae" genus="Ptecticus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="ECCCEAC9FFD73246995F4EFEFC41109C" box="[864,992,975,1000]" italics="true" pageId="14" pageNumber="1526">P. proximus</emphasis>
Rozkošný and Kovac, 1996
</taxonomicName>
. Therefore, this feature, the colouration of the thorax, is also not a good choice for genus identification. Adisoemarto, however, was not aware of James’s description of 
<taxonomicName id="19B84D58FFD732469AD14906FDB31724" authority="(James 1939)" baseAuthorityName="James" baseAuthorityYear="1939" box="[238,530,1079,1104]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD732469AD14906FEDA1724" box="[238,379,1079,1104]" italics="true" pageId="14" pageNumber="1526">Formosargus</emphasis>
(
<bibRefCitation id="BA294B2AFFD732469BBA4906FDA81724" author="James MT" box="[389,521,1079,1104]" pageId="14" pageNumber="1526" pagination="31 - 37" refId="ref18969" refString="James MT. 1939. New Formosan Stratiomyidae in the collection of the Deutsches Entomologisches Institut. Arbeiten uber morphologische und taxonomische Entomologie aus Berlin-Dahlem. 6 (1): 31 - 37." type="journal article" year="1939">James 1939</bibRefCitation>
)
</taxonomicName>
, nor of his second species described in 1969, in which he stated that the genus could be distinguished by the m–cu connected to the M3+4 fork and by veins M1 and M3 mostly fading. After examining photographs of the female 
<typeStatus id="01038879FFD732469E3C494CFBC617E2" box="[1027,1127,1149,1174]" pageId="14" pageNumber="1526" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="19B84D58FFD732469A9F49AEFEDF17CC" authorityName="Adisoemarto" authorityYear="1974" box="[160,382,1183,1208]" class="Insecta" family="Stratiomyidae" genus="Amsaria" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="species" species="sagittocera">
<emphasis id="ECCCEAC9FFD732469A9F49AEFEDF17CC" box="[160,382,1183,1208]" italics="true" pageId="14" pageNumber="1526">Amsaria sagittocera</emphasis>
</taxonomicName>
(
<figureCitation id="46832A5EFFD732469BAE49AEFE5E17CC" box="[401,511,1183,1208]" captionStart="Figure 24" captionStartId="36.[160,225,1620,1643]" captionTargetBox="[167,1148,145,1585]" captionTargetId="figure-3@36.[163,1153,142,1588]" captionTargetPageId="36" captionText="Figure 24. Amsaria sagittocera, female, holotype [MZB]. (a–b) Habitus, dorsal view. (c–d) Habitus, lateral view. (e) Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.7318236" httpUri="https://zenodo.org/record/7318236/files/figure.png" pageId="14" pageNumber="1526">Figure 24</figureCitation>
) housed in the MZB collection and the illustrations in 
<bibRefCitation id="BA294B2AFFD732469A9F49F3FE2617AF" author="Adisoemarto S" box="[160,391,1218,1243]" pageId="14" pageNumber="1526" pagination="69 - 71" refId="ref18465" refString="Adisoemarto S. 1974. A new genus of soldier flies from Indonesia (Diptera: Stratiomyidae). Treubia. 28 (3): 69 - 71." type="journal article" year="1974">Adisoemarto (1974)</bibRefCitation>
, it is clear to us that it belongs to 
<taxonomicName id="19B84D58FFD73246990349F3FC6A17AF" baseAuthorityName="James" baseAuthorityYear="1939" box="[828,971,1218,1243]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD73246990349F3FC6A17AF" box="[828,971,1218,1243]" italics="true" pageId="14" pageNumber="1526">Formosargus</emphasis>
</taxonomicName>
, and therefore, 
<taxonomicName id="19B84D58FFD732469A9F49D4FF58178A" authorityName="Adisoemarto" authorityYear="1974" box="[160,249,1253,1278]" class="Insecta" family="Stratiomyidae" genus="Amsaria" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD732469A9F49D4FF58178A" box="[160,249,1253,1278]" italics="true" pageId="14" pageNumber="1526">Amsaria</emphasis>
</taxonomicName>
is proposed as a new synonym of 
<taxonomicName id="19B84D58FFD7324698BD49D4FCAE178A" baseAuthorityName="James" baseAuthorityYear="1939" box="[642,783,1253,1278]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD7324698BD49D4FCAE178A" box="[642,783,1253,1278]" italics="true" pageId="14" pageNumber="1526">Formosargus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="DE0736DBFFD732479AFF4836FDCC1133" blockId="14.[160,1156,350,1728]" lastBlockId="15.[160,1156,142,791]" lastPageId="15" lastPageNumber="1527" pageId="14" pageNumber="1526">
The male genitalia, illustrated for the first the time in this paper, is rather unique in the 
<taxonomicName id="19B84D58FFD732469A9F481BFEA51637" authorityName="Walker" authorityYear="1834" box="[160,260,1322,1347]" class="Insecta" family="Stratiomyidae" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="subFamily" subFamily="Sarginae">Sarginae</taxonomicName>
, distinct from the genitalia of the other genera worldwide. In some 
<taxonomicName id="19B84D58FFD732469E22481BFB201637" authorityName="Walker" authorityYear="1834" box="[1053,1153,1322,1347]" class="Insecta" family="Stratiomyidae" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="subFamily" subFamily="Sarginae">Sarginae</taxonomicName>
, there is a medial line, sometimes only visible as a subtle scar, over the ventral face of the synsternite, basal to the gonostylus insertion, with different degrees of sclerotisation that may be evidence of the fusion between the gonocoxites. This condition can be seen in 
<taxonomicName id="19B84D58FFD732469A9F4884FEBA16BA" authorityName="Wiedemann" authorityYear="1830" box="[160,283,1461,1486]" class="Insecta" family="Stratiomyidae" genus="Acrochaeta" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD732469A9F4884FEBA16BA" box="[160,283,1461,1486]" italics="true" pageId="14" pageNumber="1526">Acrochaeta</emphasis>
</taxonomicName>
and some 
<taxonomicName id="19B84D58FFD732469BA34884FDBB16BA" authorityName="Loew" authorityYear="1855" box="[412,538,1461,1486]" class="Insecta" family="Stratiomyidae" genus="Merosargus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD732469BA34884FDBB16BA" box="[412,538,1461,1486]" italics="true" pageId="14" pageNumber="1526">Merosargus</emphasis>
</taxonomicName>
, where the medial line can be further modified into a highly sclerotised projection internally (e.g. 
<bibRefCitation id="BA294B2AFFD7324698AD48E6FC101684" author="Fachin DA &amp; Amorim DS" box="[658,945,1495,1520]" pageId="14" pageNumber="1526" pagination="1" refId="ref18648" refString="Fachin DA, Amorim DS. 2015. Taxonomic revision and cladistic analysis of the Neotropical genus Acrochaeta Wiedemann, 1830 (Diptera: Stratiomyidae: Sarginae). Zootaxa. 4050 (1): 001 - 110. doi: 10.11646 / zootaxa. 4050.1.1." type="journal article" year="2015">Fachin and Amorim 2015</bibRefCitation>
, p. 91, fig. 335). In the three genera, this medial line on the synsternite is present, although it is not well marked externally in 
<taxonomicName id="19B84D58FFD732469BB24B2CFDBB1542" baseAuthorityName="James" baseAuthorityYear="1939" box="[397,538,1565,1590]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD732469BB24B2CFDBB1542" box="[397,538,1565,1590]" italics="true" pageId="14" pageNumber="1526">Formosargus</emphasis>
</taxonomicName>
as it can be in 
<taxonomicName id="19B84D58FFD7324698FB4B2CFC9E1542" authorityName="Wiedemann" authorityYear="1830" box="[708,831,1565,1590]" class="Insecta" family="Stratiomyidae" genus="Acrochaeta" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD7324698FB4B2CFC9E1542" box="[708,831,1565,1590]" italics="true" pageId="14" pageNumber="1526">Acrochaeta</emphasis>
</taxonomicName>
and 
<taxonomicName id="19B84D58FFD7324699484B2CFBF51542" box="[887,1108,1565,1590]" class="Insecta" family="Stratiomyidae" genus="Merosargus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="ECCCEAC9FFD7324699484B2CFC541542" box="[887,1013,1565,1590]" italics="true" pageId="14" pageNumber="1526">Merosargus</emphasis>
species.
</taxonomicName>
The medial line is, then, in 
<emphasis id="ECCCEAC9FFD732469B9A4B0EFD1A152C" box="[421,699,1599,1624]" italics="true" pageId="14" pageNumber="1526">
<taxonomicName id="19B84D58FFD732469B9A4B0EFD80152C" authorityName="Wiedemann" authorityYear="1830" box="[421,545,1599,1624]" class="Insecta" family="Stratiomyidae" genus="Acrochaeta" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">Acrochaeta</taxonomicName>
, 
<taxonomicName id="19B84D58FFD7324698114B0EFD1A152C" baseAuthorityName="James" baseAuthorityYear="1939" box="[558,699,1599,1624]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">Formosargus</taxonomicName>
</emphasis>
, and in some 
<taxonomicName id="19B84D58FFD73246995F4B0EFC7F152C" authorityName="Loew" authorityYear="1855" box="[864,990,1599,1624]" class="Insecta" family="Stratiomyidae" genus="Merosargus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD73246995F4B0EFC7F152C" box="[864,990,1599,1624]" italics="true" pageId="14" pageNumber="1526">Merosargus</emphasis>
</taxonomicName>
, connected to the inner walls of the synsternite (e.g. 
<figureCitation id="46832A5EFFD73246987B4B53FD0C150F" box="[580,685,1634,1659]" captionStart="Figure 10" captionStartId="18.[160,225,1626,1649]" captionTargetBox="[173,1142,148,1584]" captionTargetId="figure-3@18.[163,1153,142,1594]" captionTargetPageId="18" captionText="Figure 10. Male terminalia of Formosargus species (epandrium, proctiger, and cerci removed). (a–b) F. lineatus (a. dorsal view; b. lateral view). (c–e) F. trivittatus sp. nov., holotype (c. dorsal view; d. ventral view; e. lateral view). (f) F. woodleyi sp. nov., lateral view, holotype. Abbreviations: goncx apod, gonocoxal apodeme; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7318203" httpUri="https://zenodo.org/record/7318203/files/figure.png" pageId="14" pageNumber="1526">Figure 10</figureCitation>
(b)); 
<bibRefCitation id="BA294B2AFFD7324698E24B53FC55150F" author="Fachin DA &amp; Amorim DS" box="[733,1012,1634,1659]" pageId="14" pageNumber="1526" pagination="1" refId="ref18648" refString="Fachin DA, Amorim DS. 2015. Taxonomic revision and cladistic analysis of the Neotropical genus Acrochaeta Wiedemann, 1830 (Diptera: Stratiomyidae: Sarginae). Zootaxa. 4050 (1): 001 - 110. doi: 10.11646 / zootaxa. 4050.1.1." type="journal article" year="2015">Fachin and Amorim 2015</bibRefCitation>
, p. 19, fig. 40 and p. 90, fig. 331)). Additionally, 
<taxonomicName id="19B84D58FFD7324698264BB4FD0715EA" baseAuthorityName="James" baseAuthorityYear="1939" box="[537,678,1669,1694]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1526" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD7324698264BB4FD0715EA" box="[537,678,1669,1694]" italics="true" pageId="14" pageNumber="1526">Formosargus</emphasis>
</taxonomicName>
shares with these two Neotropical genera the presence of an inner projection in the synsternite, but the condition is slightly different. The inner projection itself in 
<taxonomicName id="19B84D58FFD63247984C4DBFFCA313D3" baseAuthorityName="James" baseAuthorityYear="1939" box="[627,770,142,167]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="1527" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD63247984C4DBFFCA313D3" box="[627,770,142,167]" italics="true" pageId="15" pageNumber="1527">Formosargus</emphasis>
</taxonomicName>
is rather long, parallel and well defined, sometimes with strong setae distally, and free of an innermost connection with the ventral or inner margins of the gonocoxite (
<figureCitation id="46832A5EFFD63247988A4DE5FCBF1399" box="[693,798,212,237]" captionStart="Figure 11" captionStartId="19.[160,225,1550,1573]" captionTargetBox="[208,1106,144,1515]" captionTargetId="figure-1@19.[206,1109,142,1518]" captionTargetPageId="19" captionText="Figure 11. Male terminalia of Formosargus lineatus. (a) Genitalia, dorsal view; arrow: reduced gonocoxal apodeme. (b) Genitalia, lateral view; arrow: inner projection of synsternite. (c) Genitalia, ventral view; arrow: medial line. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; gonst, gonostylus; ph, phallus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger." figureDoi="http://doi.org/10.5281/zenodo.7318205" httpUri="https://zenodo.org/record/7318205/files/figure.png" pageId="15" pageNumber="1527">Figure 11</figureCitation>
), while in some 
<taxonomicName id="19B84D58FFD6324799EB4DE5FBF31399" authorityName="Loew" authorityYear="1855" box="[980,1106,212,237]" class="Insecta" family="Stratiomyidae" genus="Merosargus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="1527" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD6324799EB4DE5FBF31399" box="[980,1106,212,237]" italics="true" pageId="15" pageNumber="1527">Merosargus</emphasis>
</taxonomicName>
and in some members of 
<taxonomicName id="19B84D58FFD632479B9B4DC7FD80127B" authorityName="Wiedemann" authorityYear="1830" box="[420,545,246,271]" class="Insecta" family="Stratiomyidae" genus="Acrochaeta" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="1527" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD632479B9B4DC7FD80127B" box="[420,545,246,271]" italics="true" pageId="15" pageNumber="1527">Acrochaeta</emphasis>
</taxonomicName>
(e.g. 
<taxonomicName id="19B84D58FFD6324798564DC7FC60127B" authority="Wiedemann, 1830" authorityName="Wiedemann" authorityYear="1830" box="[617,961,246,271]" class="Insecta" family="Stratiomyidae" genus="Acrochaeta" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="1527" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="ECCCEAC9FFD6324798564DC7FD40127B" box="[617,737,246,271]" italics="true" pageId="15" pageNumber="1527">A. fasciata</emphasis>
Wiedemann, 1830
</taxonomicName>
; see 
<bibRefCitation id="BA294B2AFFD632479E3D4DC7FE9A1246" author="Fachin DA &amp; Amorim DS" pageId="15" pageNumber="1527" pagination="1" refId="ref18648" refString="Fachin DA, Amorim DS. 2015. Taxonomic revision and cladistic analysis of the Neotropical genus Acrochaeta Wiedemann, 1830 (Diptera: Stratiomyidae: Sarginae). Zootaxa. 4050 (1): 001 - 110. doi: 10.11646 / zootaxa. 4050.1.1." type="journal article" year="2015">Fachin and Amorim 2015</bibRefCitation>
, p. 19, fig. 40), the inner projection is wide and simple, with no signal of reduction around the gonostylus, and in the other members of 
<taxonomicName id="19B84D58FFD6324799614C0DFC781221" authorityName="Wiedemann" authorityYear="1830" box="[862,985,316,341]" class="Insecta" family="Stratiomyidae" genus="Acrochaeta" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="1527" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD6324799614C0DFC781221" box="[862,985,316,341]" italics="true" pageId="15" pageNumber="1527">Acrochaeta</emphasis>
</taxonomicName>
(e.g. 
<taxonomicName id="19B84D58FFD632479E2C4C0DFE811203" authority="Bigot, 1879" authorityName="Bigot" authorityYear="1879" class="Insecta" family="Stratiomyidae" genus="Acrochaeta" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="1527" phylum="Arthropoda" rank="species" species="flaveola">
<emphasis id="ECCCEAC9FFD632479E2C4C0DFB221221" box="[1043,1155,316,341]" italics="true" pageId="15" pageNumber="1527">A. flaveola</emphasis>
Bigot, 1879
</taxonomicName>
; see 
<bibRefCitation id="BA294B2AFFD632479B694C6FFDD11203" author="Fachin DA &amp; Amorim DS" box="[342,624,350,375]" pageId="15" pageNumber="1527" pagination="1" refId="ref18648" refString="Fachin DA, Amorim DS. 2015. Taxonomic revision and cladistic analysis of the Neotropical genus Acrochaeta Wiedemann, 1830 (Diptera: Stratiomyidae: Sarginae). Zootaxa. 4050 (1): 001 - 110. doi: 10.11646 / zootaxa. 4050.1.1." type="journal article" year="2015">Fachin and Amorim 2015</bibRefCitation>
, p. 53, fig. 61), the inner projection is still platelike but reduced laterally and around the gonostylus. In 
<taxonomicName id="19B84D58FFD6324799154CB0FC1612EE" baseAuthorityName="James" baseAuthorityYear="1939" box="[810,951,385,410]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="1527" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD6324799154CB0FC1612EE" box="[810,951,385,410]" italics="true" pageId="15" pageNumber="1527">Formosargus</emphasis>
</taxonomicName>
, both the phallus and the parameral sheath are highly reduced and displaced distally in the genitalia (see e. g. 
<figureCitation id="46832A5EFFD632479A854CF7FE8212AB" box="[186,291,454,479]" captionStart="Figure 12" captionStartId="21.[160,225,1549,1572]" captionTargetBox="[207,1108,144,1516]" captionTargetId="figure-1@21.[205,1110,142,1518]" captionTargetPageId="21" captionText="Figure 12. Male terminalia of Formosargus sagittocerus. (a) Genitalia,dorsal view;arrow:reduced gonocoxal apodeme. (b) Genitalia, lateral view. (c) Genitalia, ventral view. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; goncx brg, gonocoxal bridge; gonst, gonostylus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger." figureDoi="http://doi.org/10.5281/zenodo.7318207" httpUri="https://zenodo.org/record/7318207/files/figure.png" pageId="15" pageNumber="1527">Figure 12</figureCitation>
(a)), which may be related to the reduction of the gonocoxal apodeme as well, recognisable only by an enlarging at the centre of the gonocoxal bridge (see e.g. 
<figureCitation id="46832A5EFFD632479E234CD8FB221176" box="[1052,1155,489,514]" captionStart="Figure 11" captionStartId="19.[160,225,1550,1573]" captionTargetBox="[208,1106,144,1515]" captionTargetId="figure-1@19.[206,1109,142,1518]" captionTargetPageId="19" captionText="Figure 11. Male terminalia of Formosargus lineatus. (a) Genitalia, dorsal view; arrow: reduced gonocoxal apodeme. (b) Genitalia, lateral view; arrow: inner projection of synsternite. (c) Genitalia, ventral view; arrow: medial line. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; gonst, gonostylus; ph, phallus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger." figureDoi="http://doi.org/10.5281/zenodo.7318205" httpUri="https://zenodo.org/record/7318205/files/figure.png" pageId="15" pageNumber="1527">Figure 11</figureCitation>
(a), see black arrow). These features have also not been found in other sargines and may be apomorphies for 
<taxonomicName id="19B84D58FFD632479BB94F1FFDB21133" baseAuthorityName="James" baseAuthorityYear="1939" box="[390,531,558,583]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="1527" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD632479BB94F1FFDB21133" box="[390,531,558,583]" italics="true" pageId="15" pageNumber="1527">Formosargus</emphasis>
</taxonomicName>
as well.
</paragraph>
<paragraph id="DE0736DBFFD632479AFF4F60FB201063" blockId="15.[160,1156,142,791]" pageId="15" pageNumber="1527">
Another important finding is the presence of a modified structure at the base of the first tarsomere in the male mid leg (
<figureCitation id="46832A5EFFD632479BC84F45FDF111F9" box="[503,592,628,653]" captionStart="Figure 8" captionStartId="13.[160,225,1395,1418]" captionTargetBox="[167,1149,145,1360]" captionTargetId="figure-51@13.[163,1153,142,1363]" captionTargetPageId="13" captionText="Figure 8. Male mid legs of Formosargus species, male. (a–b) F. kerteszi. (c–e) F. lineatus. (f–h) F. sagittocerus. (i–j) F. woodleyi sp. nov., holotype. Arrow: modified structure at base of basitarsus. Abbreviations: fem, femur; tsm 1, first tarsomere." figureDoi="http://doi.org/10.5281/zenodo.7318197" httpUri="https://zenodo.org/record/7318197/files/figure.png" pageId="15" pageNumber="1527">Figure 8</figureCitation>
), which is, besides the hind tibial flange of 
<taxonomicName id="19B84D58FFD632479E194F45FDB011DB" authority="Hauser and Rozkosny, 2007" authorityName="Hauser and Rozkosny" authorityYear="2007" class="Insecta" family="Stratiomyidae" genus="Ptecticus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="1527" phylum="Arthropoda" rank="species" species="kovaci">
<emphasis id="ECCCEAC9FFD632479E194F45FF4511DB" italics="true" pageId="15" pageNumber="1527">Ptecticus kovaci</emphasis>
Hauser and 
<bibRefCitation id="BA294B2AFFD632479B564FA7FDB011DB" author="Rozkosny R &amp; Kozanek M" box="[361,529,662,687]" pageId="15" pageNumber="1527" pagination="35 - 50" refId="ref19872" refString="Rozkosny R, Kozanek M. 2007. A review of the Oriental Culcua with descriptions of seven new species (Diptera: Stratiomyidae). Insect Syst Evol. 38 (1): 35 - 50." type="journal article" year="2007">Rozkošný, 2007</bibRefCitation>
</taxonomicName>
, the only case of a modified leg within 
<taxonomicName id="19B84D58FFD63247998C4FA7FBB411DB" authorityName="Walker" authorityYear="1834" box="[947,1045,662,687]" class="Insecta" family="Stratiomyidae" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="1527" phylum="Arthropoda" rank="subFamily" subFamily="Sarginae">Sarginae</taxonomicName>
.Males are indeed rarer than females and only 
<bibRefCitation id="BA294B2AFFD6324798214F88FD0511A6" author="James MT" box="[542,676,697,722]" pageId="15" pageNumber="1527" pagination="31 - 37" refId="ref18969" refString="James MT. 1939. New Formosan Stratiomyidae in the collection of the Deutsches Entomologisches Institut. Arbeiten uber morphologische und taxonomische Entomologie aus Berlin-Dahlem. 6 (1): 31 - 37." type="journal article" year="1939">James (1939</bibRefCitation>
, 
<bibRefCitation id="BA294B2AFFD6324798914F88FD4511A6" author="James MT" box="[686,740,697,722]" pageId="15" pageNumber="1527" pagination="339 - 350" refId="ref19004" refString="James MT. 1969. The soldier flies (Diptera, Stratiomyidae) collected in the Philippine Islands by the Noona Dan Expedition. Entomol Medd. 37 (4): 339 - 350." type="journal article" year="1969">1969</bibRefCitation>
) had males to examine, but he did not notice the structure in the male mid leg. In the species of which we have males,the feature is unique for each species, which indicates it may be associated with some sort of courtship.
</paragraph>
</subSubSection>
<subSubSection id="96A26550FFD632479A9F4E75FD7B1029" box="[160,730,836,861]" pageId="15" pageNumber="1527" type="multiple">
<paragraph id="DE0736DBFFD632479A9F4E75FD7B1029" blockId="15.[160,730,836,861]" box="[160,730,836,861]" pageId="15" pageNumber="1527">
<emphasis id="ECCCEAC9FFD632479A9F4E75FD7B1029" bold="true" box="[160,730,836,861]" pageId="15" pageNumber="1527">
Key to the adults of the species of 
<taxonomicName id="19B84D58FFD63247987D4E75FD7B1029" baseAuthorityName="James" baseAuthorityYear="1939" box="[578,730,836,861]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="1527" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFD63247987D4E75FD7B1029" bold="true" box="[578,730,836,861]" italics="true" pageId="15" pageNumber="1527">Formosargus</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="96A26550FFD632589A804EB7FB25177F" lastPageId="16" lastPageNumber="1528" pageId="15" pageNumber="1527" type="key">
<key id="AA2961EAFFD632589A804EB7FB25177F" lastPageId="16" lastPageNumber="1528" pageId="15" pageNumber="1527">
<keyStep id="654C2E7EFFD632479A804EB7FB2217C1" pageId="15" pageNumber="1527">
<paragraph id="DE0736DBFFD632479A804EB7FB22175E" blockId="15.[191,1157,902,1690]" pageId="15" pageNumber="1527">
<keyLead id="654995EEFFD632479A804EB7FB22175E" pageId="15" pageNumber="1527">
1. Scutum with only a dark brown medial band (
<figureCitation id="46832A5EFFD6324798DC4EB7FCE110EB" box="[739,832,902,927]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="15" pageNumber="1527">Figure 7</figureCitation>
(a–c)) extending onto scutellum, in one species with an additional notopleural macula (not to be confused with a dark macula on anepisternum); fourth flagellomere rounded, shorter than previous flagellomeres combined (
<figureCitation id="46832A5EFFD632479BC74EDFFDF51773" box="[504,596,1006,1031]" captionStart="Figure 2" captionStartId="6.[160,225,1404,1427]" captionTargetBox="[165,1150,144,1369]" captionTargetId="figure-53@6.[163,1153,142,1372]" captionTargetPageId="6" captionText="Figure 2. Antennae of Formosargus species. (a) F. lineatus, male. (b) F. lineatus, female. (c) F. trivittatus sp. nov., male, holotype. Abbreviations: ar t flgm, arista-like terminal flagellomere; flg, flagellum; ped, pedicel; scp, scape." figureDoi="http://doi.org/10.5281/zenodo.7318184" httpUri="https://zenodo.org/record/7318184/files/figure.png" pageId="15" pageNumber="1527">Figure 2</figureCitation>
(a, b)); last four hind tarsomeres completely yellow (
<figureCitation id="46832A5EFFD632479ADB4920FEEF175E" box="[228,334,1041,1066]" captionStart="Figure 18" captionStartId="30.[160,225,1620,1643]" captionTargetBox="[167,1145,145,1584]" captionTargetId="figure-3@30.[163,1153,142,1588]" captionTargetPageId="30" captionText="Figure 18. Habitus of Formosargus species. (a) F. kerteszi, female, dorsal. (b–e) F. lineatus (b–c. male, dorsal and lateral, respectively; d–e. female, dorsal and lateral, respectively). (f) F. sagittocerus, male, dorsal." figureDoi="http://doi.org/10.5281/zenodo.7318222" httpUri="https://zenodo.org/record/7318222/files/figure.png" pageId="15" pageNumber="1527">Figure 18</figureCitation>
(c)) ........................................................................................................................................ 2
</keyLead>
</paragraph>
<paragraph id="DE0736DBFFD632479A804905FB2217C1" blockId="15.[191,1157,902,1690]" pageId="15" pageNumber="1527">
<keyLead id="654995EEFFD632479A804905FB2217C1" pageId="15" pageNumber="1527">
- Scutum with additional dark markings beside the medial band and the notopleural macula (
<figureCitation id="46832A5EFFD632479B024967FE3B171B" box="[317,410,1110,1135]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="15" pageNumber="1527">Figure 7</figureCitation>
(h)), in some species scutum nearly completely dark (
<figureCitation id="46832A5EFFD6324799D54967FBE9171B" box="[1002,1096,1110,1135]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="15" pageNumber="1527">Figure 7</figureCitation>
(d, i)); fourth flagellomere subtriangular, as long as previous flagellomeres combined (
<figureCitation id="46832A5EFFD632479AD949ADFEE317C1" box="[230,322,1180,1205]" captionStart="Figure 2" captionStartId="6.[160,225,1404,1427]" captionTargetBox="[165,1150,144,1369]" captionTargetId="figure-53@6.[163,1153,142,1372]" captionTargetPageId="6" captionText="Figure 2. Antennae of Formosargus species. (a) F. lineatus, male. (b) F. lineatus, female. (c) F. trivittatus sp. nov., male, holotype. Abbreviations: ar t flgm, arista-like terminal flagellomere; flg, flagellum; ped, pedicel; scp, scape." figureDoi="http://doi.org/10.5281/zenodo.7318184" httpUri="https://zenodo.org/record/7318184/files/figure.png" pageId="15" pageNumber="1527">Figure 2</figureCitation>
(c)); last four hind tarsomeres black (
<figureCitation id="46832A5EFFD6324798E349ADFCE717C1" box="[732,838,1180,1205]" captionStart="Figure 19" captionStartId="31.[160,225,1625,1648]" captionTargetBox="[168,1148,145,1587]" captionTargetId="figure-3@31.[163,1153,142,1594]" captionTargetPageId="31" captionText="Figure 19. Habitus of Formosargus species. (a) F. sagittocerus, male, lateral. (b–c) F. berezovskiyi sp. nov., female, paratype, dorsal and lateral, respectively. (d–e) F. borneensis sp. nov., female, holotype, dorsal and lateral, respectively. (f) F. mangoleensis sp. nov., female, holotype, dorsal." figureDoi="http://doi.org/10.5281/zenodo.7318226" httpUri="https://zenodo.org/record/7318226/files/figure.png" pageId="15" pageNumber="1527">Figure 19</figureCitation>
(a)) or completely yellow 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="654C2E7EFFD632479A80498FFB22164B" pageId="15" pageNumber="1527">
<paragraph id="DE0736DBFFD632479A80498FFB25178E" blockId="15.[191,1157,902,1690]" pageId="15" pageNumber="1527">
<keyLead id="654995EEFFD632479A80498FFB25178E" pageId="15" pageNumber="1527">
2. Notopleuron with large black macula (
<figureCitation id="46832A5EFFD632479882498FFC8217A3" box="[701,803,1214,1239]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="15" pageNumber="1527">Figure 7</figureCitation>
(f)); male unknown; 
<collectingCountry id="A6AF764BFFD632479E24498FFB2017A3" box="[1051,1153,1214,1239]" name="Malaysia" pageId="15" pageNumber="1527">Malaysia</collectingCountry>
: Borneo............................................................................................................. 
<taxonomicName id="19B84D58FFD6324799A649D0FB84178E" ID-CoL="BCY38" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[921,1061,1249,1274]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="1527" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
<emphasis id="ECCCEAC9FFD6324799A649D0FB84178E" box="[921,1061,1249,1274]" italics="true" pageId="15" pageNumber="1527">F. borneensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD632479E1249D0FB25178E" box="[1069,1156,1249,1274]" pageId="15" pageNumber="1527" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="DE0736DBFFD632479A804835FB22164B" blockId="15.[191,1157,902,1690]" pageId="15" pageNumber="1527">
<keyLead id="654995EEFFD632479A804835FB22164B" pageId="15" pageNumber="1527">
- Notopleuron without black macula (
<figureCitation id="46832A5EFFD6324798474835FD771669" box="[632,726,1284,1309]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="15" pageNumber="1527">Figure 7</figureCitation>
(g)), pale yellow to yellow-orange, like most of the body ................................................................................................................................ 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="654C2E7EFFD632479A804878FB2216BE" pageId="15" pageNumber="1527">
<paragraph id="DE0736DBFFD632479A804878FB2216F1" blockId="15.[191,1157,902,1690]" pageId="15" pageNumber="1527">
<keyLead id="654995EEFFD632479A804878FB2216F1" pageId="15" pageNumber="1527">
3. Mediotergite completely yellow (
<figureCitation id="46832A5EFFD6324798634878FD1D1616" box="[604,700,1353,1378]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="15" pageNumber="1527">Figure 7</figureCitation>
(g)), conforming with the colour of the body ......................................................................................................................................................... 4
</keyLead>
</paragraph>
<paragraph id="DE0736DBFFD632479A8048BFFB2216BE" blockId="15.[191,1157,902,1690]" pageId="15" pageNumber="1527">
<keyLead id="654995EEFFD632479A8048BFFB2216BE" pageId="15" pageNumber="1527">
- Mediotergite completely dark (
<figureCitation id="46832A5EFFD63247986E48BFFD1216D3" box="[593,691,1422,1447]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="15" pageNumber="1527">Figure 7</figureCitation>
(e)), contrasting with the colour of the body ................................................................................................................................................ 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="654C2E7EFFD632479A8048E5FB22157B" pageId="15" pageNumber="1527">
<paragraph id="DE0736DBFFD632479A8048E5FB221699" blockId="15.[191,1157,902,1690]" box="[191,1155,1492,1517]" pageId="15" pageNumber="1527">
<keyLead id="654995EEFFD632479A8048E5FB221699" box="[191,1155,1492,1517]" pageId="15" pageNumber="1527">
4. Anepisternum with black macula (
<figureCitation id="46832A5EFFD63247985F48E5FD6A1699" box="[608,715,1492,1517]" captionStart="Figure 20" captionStartId="32.[160,225,1630,1653]" captionTargetBox="[171,1149,154,1593]" captionTargetId="figure-3@32.[163,1153,142,1598]" captionTargetPageId="32" captionText="Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively." figureDoi="http://doi.org/10.5281/zenodo.7318228" httpUri="https://zenodo.org/record/7318228/files/figure.png" pageId="15" pageNumber="1527">Figure 20</figureCitation>
(a)) .................................................................... 5
</keyLead>
</paragraph>
<paragraph id="DE0736DBFFD632479A8048C7FB22157B" blockId="15.[191,1157,902,1690]" box="[191,1155,1526,1551]" pageId="15" pageNumber="1527">
<keyLead id="654995EEFFD632479A8048C7FB22157B" box="[191,1155,1526,1551]" pageId="15" pageNumber="1527">
- Anepisternum without black macula (
<figureCitation id="46832A5EFFD6324798B648C7FD52157B" box="[649,755,1526,1551]" captionStart="Figure 18" captionStartId="30.[160,225,1620,1643]" captionTargetBox="[167,1145,145,1584]" captionTargetId="figure-3@30.[163,1153,142,1588]" captionTargetPageId="30" captionText="Figure 18. Habitus of Formosargus species. (a) F. kerteszi, female, dorsal. (b–e) F. lineatus (b–c. male, dorsal and lateral, respectively; d–e. female, dorsal and lateral, respectively). (f) F. sagittocerus, male, dorsal." figureDoi="http://doi.org/10.5281/zenodo.7318222" httpUri="https://zenodo.org/record/7318222/files/figure.png" pageId="15" pageNumber="1527">Figure 18</figureCitation>
(c))............................................................. 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="654C2E7EFFD632479A804B28FB2515EE" pageId="15" pageNumber="1527">
<paragraph id="DE0736DBFFD632479A804B28FB221521" blockId="15.[191,1157,902,1690]" pageId="15" pageNumber="1527">
<keyLead id="654995EEFFD632479A804B28FB221521" pageId="15" pageNumber="1527">
5. Vertex area less than twice the length of the ocellar tubercle (
<figureCitation id="46832A5EFFD6324799AA4B28FC5C1546" box="[917,1021,1561,1586]" captionStart="Figure 3" captionStartId="7.[160,225,906,929]" captionTargetBox="[167,1149,145,869]" captionTargetId="figure-267@7.[163,1153,142,874]" captionTargetPageId="7" captionText="Figure 3. Heads of Formosargus species, dorsal view. (a) F. kerteszi, female. (b–c) F. lineatus (b. male; c. female). (d) F. sagittocerus, male." figureDoi="http://doi.org/10.5281/zenodo.7318186" httpUri="https://zenodo.org/record/7318186/files/figure.png" pageId="15" pageNumber="1527">Figures 3</figureCitation>
(a), 22(a, e)); 
<collectingCountry id="A6AF764BFFD632479AE04B0DFE901521" box="[223,305,1596,1621]" name="Taiwan" pageId="15" pageNumber="1527">Taiwan</collectingCountry>
; 
<collectingCountry id="A6AF764BFFD632479B044B0DFE171521" box="[315,438,1596,1621]" name="Philippines" pageId="15" pageNumber="1527">Philippines</collectingCountry>
.................................. 
<taxonomicName id="19B84D58FFD6324798B24B0DFC291521" ID-CoL="6JHMJ" authority="James, 1939" authorityName="James" authorityYear="1939" box="[653,904,1596,1621]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="1527" phylum="Arthropoda" rank="species" species="kerteszi">
<emphasis id="ECCCEAC9FFD6324798B24B0DFD561521" box="[653,759,1596,1621]" italics="true" pageId="15" pageNumber="1527">F. kerteszi</emphasis>
<bibRefCitation id="BA294B2AFFD6324798C04B0DFC291521" author="James MT" box="[767,904,1596,1621]" pageId="15" pageNumber="1527" pagination="31 - 37" refId="ref18969" refString="James MT. 1939. New Formosan Stratiomyidae in the collection of the Deutsches Entomologisches Institut. Arbeiten uber morphologische und taxonomische Entomologie aus Berlin-Dahlem. 6 (1): 31 - 37." type="journal article" year="1939">James, 1939</bibRefCitation>
</taxonomicName>
[in part, only females]
</keyLead>
</paragraph>
<paragraph id="DE0736DBFFD632479A804B6FFB2515EE" blockId="15.[191,1157,902,1690]" pageId="15" pageNumber="1527">
<keyLead id="654995EEFFD632479A804B6FFB2515EE" pageId="15" pageNumber="1527">
- Vertex area at least twice the length of the ocellar tubercle (
<figureCitation id="46832A5EFFD6324799884B6FFBB91503" box="[951,1048,1630,1655]" captionStart="Figure 4" captionStartId="8.[160,225,1266,1289]" captionTargetBox="[167,1149,145,1232]" captionTargetId="figure-139@8.[163,1153,142,1234]" captionTargetPageId="8" captionText="Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318188" httpUri="https://zenodo.org/record/7318188/files/figure.png" pageId="15" pageNumber="1527">Figure 4</figureCitation>
(c)); male unknown; 
<collectingCountry id="A6AF764BFFD632479B6B4BB0FE6215EE" box="[340,451,1665,1690]" name="Indonesia" pageId="15" pageNumber="1527">Indonesia</collectingCountry>
: Mangole Island .......................................... 
<taxonomicName id="19B84D58FFD6324799474BB0FB8415EE" ID-CoL="BCY3B" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[888,1061,1665,1690]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="1527" phylum="Arthropoda" rank="species" species="mangoleensis" status="sp. nov.">
<emphasis id="ECCCEAC9FFD6324799474BB0FB8415EE" box="[888,1061,1665,1690]" italics="true" pageId="15" pageNumber="1527">F. mangoleensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFD632479E124BB0FB2515EE" box="[1069,1156,1665,1690]" pageId="15" pageNumber="1527" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="654C2E7EFFC932589A804DBEFE6A1221" pageId="16" pageNumber="1528">
<paragraph id="DE0736DBFFC932589A804DBEFB221399" blockId="16.[180,1157,143,1035]" pageId="16" pageNumber="1528">
<keyLead id="654995EEFFC932589A804DBEFB221399" pageId="16" pageNumber="1528">
6. Female abdomen completely yellow, without dark markings (
<figureCitation id="46832A5EFFC9325899964DBEFBB913DC" box="[937,1048,143,168]" captionStart="Figure 18" captionStartId="30.[160,225,1620,1643]" captionTargetBox="[167,1145,145,1584]" captionTargetId="figure-3@30.[163,1153,142,1588]" captionTargetPageId="30" captionText="Figure 18. Habitus of Formosargus species. (a) F. kerteszi, female, dorsal. (b–e) F. lineatus (b–c. male, dorsal and lateral, respectively; d–e. female, dorsal and lateral, respectively). (f) F. sagittocerus, male, dorsal." figureDoi="http://doi.org/10.5281/zenodo.7318222" httpUri="https://zenodo.org/record/7318222/files/figure.png" pageId="16" pageNumber="1528">Figure 18</figureCitation>
(d)); male with a pair of dark markings on the lateral margins of tergite 1 (
<figureCitation id="46832A5EFFC93258999C4D83FBAC13BF" box="[931,1037,178,203]" captionStart="Figure 18" captionStartId="30.[160,225,1620,1643]" captionTargetBox="[167,1145,145,1584]" captionTargetId="figure-3@30.[163,1153,142,1588]" captionTargetPageId="30" captionText="Figure 18. Habitus of Formosargus species. (a) F. kerteszi, female, dorsal. (b–e) F. lineatus (b–c. male, dorsal and lateral, respectively; d–e. female, dorsal and lateral, respectively). (f) F. sagittocerus, male, dorsal." figureDoi="http://doi.org/10.5281/zenodo.7318222" httpUri="https://zenodo.org/record/7318222/files/figure.png" pageId="16" pageNumber="1528">Figure 18</figureCitation>
(b)); 
<collectingCountry id="A6AF764BFFC932589E024D83FEC91399" name="Papua New Guinea" pageId="16" pageNumber="1528">Papua New Guinea</collectingCountry>
................................................................................................ 
<taxonomicName id="19B84D58FFC9325899B84DE5FB221399" authority="(de Meijere)" baseAuthorityName="de Meijere" box="[903,1155,212,237]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="1528" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="ECCCEAC9FFC9325899B84DE5FC571399" box="[903,1014,212,237]" italics="true" pageId="16" pageNumber="1528">F. lineatus</emphasis>
(de Meijere)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DE0736DBFFC932589A804DC6FE6A1221" blockId="16.[180,1157,143,1035]" pageId="16" pageNumber="1528">
<keyLead id="654995EEFFC932589A804DC6FE6A1221" pageId="16" pageNumber="1528">
- Female abdominal tergites 2–5 each with a pair of diffuse dark markings (
<bibRefCitation id="BA294B2AFFC932589E2F4DC6FE931247" author="Lessard BD &amp; Yeates DK &amp; Woodley NE" pageId="16" pageNumber="1528" pagination="23 - 43" refId="ref19207" refString="Lessard BD, Yeates DK, Woodley NE. 2020. Review of Australian Sarginae soldier fly genera (Diptera: Stratiomyidae), with first records of Cephalochrysa, Formosargus and Microchrysa. Rec Aust Mus. 72 (2): 23 - 43. doi: 10.3853 / j. 2201 - 4349.72.2020.1683." type="journal article" year="2020">Lessard et al. 2020</bibRefCitation>
, p. 29, fig. 4); male unknown; 
<collectingCountry id="A6AF764BFFC9325898BB4C2BFD491247" box="[644,744,282,307]" name="Australia" pageId="16" pageNumber="1528">Australia</collectingCountry>
: 
<collectingRegion id="1C7CF839FFC9325898CC4C2BFCD81247" box="[755,889,282,307]" country="Australia" name="Queensland" pageId="16" pageNumber="1528">Queensland</collectingRegion>
...... 
<taxonomicName id="19B84D58FFC93258998F4C2BFE6A1221" authority="Lessard and Woodley" authorityName="Lessard and Woodley" authorityYear="2020" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="1528" phylum="Arthropoda" rank="species" species="melanogrammus">
<emphasis id="ECCCEAC9FFC93258998F4C2BFB221247" box="[944,1155,282,307]" italics="true" pageId="16" pageNumber="1528">F. melanogrammus</emphasis>
Lessard and Woodley
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="654C2E7EFFC932589A804C6EFB25113C" pageId="16" pageNumber="1528">
<paragraph id="DE0736DBFFC932589A804C6EFB2212C9" blockId="16.[180,1157,143,1035]" pageId="16" pageNumber="1528">
<keyLead id="654995EEFFC932589A804C6EFB2212C9" pageId="16" pageNumber="1528">
7. Black upper frons wider than long, anterior margin slightly inverse-U-shaped (
<figureCitation id="46832A5EFFC932589E154C6EFB22120C" box="[1066,1155,351,376]" captionStart="Figure 5" captionStartId="9.[160,225,1289,1312]" captionTargetBox="[167,1149,145,1251]" captionTargetId="figure-113@9.[163,1153,142,1257]" captionTargetPageId="9" captionText="Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi="http://doi.org/10.5281/zenodo.7318191" httpUri="https://zenodo.org/record/7318191/files/figure.png" pageId="16" pageNumber="1528">Figure 5</figureCitation>
(a)); scutal band less than one-third of the scutum width, wider on posterior half (
<figureCitation id="46832A5EFFC932589E004CB3FF4A12C9" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="16" pageNumber="1528">Figure 7</figureCitation>
(a)); 
<collectingCountry id="A6AF764BFFC932589B264C95FEC912C9" box="[281,360,420,445]" name="Taiwan" pageId="16" pageNumber="1528">Taiwan</collectingCountry>
: main island; 
<collectingCountry id="A6AF764BFFC932589BC44C95FDD012C9" box="[507,625,420,445]" name="Philippines" pageId="16" pageNumber="1528">Philippines</collectingCountry>
......... 
<taxonomicName id="19B84D58FFC9325898894C95FC0412C9" ID-CoL="6JHMJ" authority="James, 1939" authorityName="James" authorityYear="1939" box="[694,933,420,445]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="1528" phylum="Arthropoda" rank="species" species="kerteszi">
<emphasis id="ECCCEAC9FFC9325898894C95FCBA12C9" box="[694,795,420,445]" italics="true" pageId="16" pageNumber="1528">F. kerteszi</emphasis>
<bibRefCitation id="BA294B2AFFC93258991D4C95FC0412C9" author="James MT" box="[802,933,420,445]" pageId="16" pageNumber="1528" pagination="31 - 37" refId="ref18969" refString="James MT. 1939. New Formosan Stratiomyidae in the collection of the Deutsches Entomologisches Institut. Arbeiten uber morphologische und taxonomische Entomologie aus Berlin-Dahlem. 6 (1): 31 - 37." type="journal article" year="1939">James, 1939</bibRefCitation>
</taxonomicName>
[in part, only males]
</keyLead>
</paragraph>
<paragraph id="DE0736DBFFC932589A804CF6FB25113C" blockId="16.[180,1157,143,1035]" pageId="16" pageNumber="1528">
<keyLead id="654995EEFFC932589A804CF6FB25113C" pageId="16" pageNumber="1528">
- Black upper frons as long as wide, anterior margin slightly pointed towards the middle (
<figureCitation id="46832A5EFFC932589B074CDBFE321177" box="[312,403,490,515]" captionStart="Figure 6" captionStartId="10.[160,225,1285,1308]" captionTargetBox="[167,1149,145,1251]" captionTargetId="figure-99@10.[163,1153,142,1253]" captionTargetPageId="10" captionText="Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi="http://doi.org/10.5281/zenodo.7318193" httpUri="https://zenodo.org/record/7318193/files/figure.png" pageId="16" pageNumber="1528">Figure 6</figureCitation>
(a)); scutal band approximately one-third of the scutum width, more homogeneous in width, except at transverse suture (
<figureCitation id="46832A5EFFC93258990D4F3DFC2E1151" box="[818,911,524,549]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="16" pageNumber="1528">Figure 7</figureCitation>
(e)); male only known from 
<figureCitation id="46832A5EFFC932589B264F1EFED7113C" box="[281,374,559,584]" captionStart="Figure 1" captionStartId="3.[160,225,645,668]" captionTargetBox="[347,969,144,610]" captionTargetId="figure-252@3.[344,972,142,613]" captionTargetPageId="3" captionText="Figure 1. Formosargus berezovskiyi sp. nov., live specimen, male, Taiwan, Orchid Island (© Yi-Xiang Lin (Gaga)). Available from: http://gaga.biodiv.tw/9501bx/536.htm." figureDoi="http://doi.org/10.5281/zenodo.7318182" httpUri="https://zenodo.org/record/7318182/files/figure.png" pageId="16" pageNumber="1528">Figure 1</figureCitation>
; 
<collectingCountry id="A6AF764BFFC932589BBD4F1EFE75113C" box="[386,468,559,584]" name="Taiwan" pageId="16" pageNumber="1528">Taiwan</collectingCountry>
: Orchid Island .............................................. 
<taxonomicName id="19B84D58FFC9325899B54F1EFB84113C" ID-CoL="BCY37" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[906,1061,559,584]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="1528" phylum="Arthropoda" rank="species" species="berezovskiyi" status="sp. nov.">
<emphasis id="ECCCEAC9FFC9325899B54F1EFB84113C" box="[906,1061,559,584]" italics="true" pageId="16" pageNumber="1528">F. berezovskiyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFC932589E124F1EFB25113C" box="[1069,1156,559,584]" pageId="16" pageNumber="1528" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="654C2E7EFFC932589A804F63FB2211C4" pageId="16" pageNumber="1528">
<paragraph id="DE0736DBFFC932589A804F63FB22111F" blockId="16.[180,1157,143,1035]" box="[191,1155,594,619]" pageId="16" pageNumber="1528">
<keyLead id="654995EEFFC932589A804F63FB22111F" box="[191,1155,594,619]" pageId="16" pageNumber="1528">
8. Scutellum completely yellowish (
<figureCitation id="46832A5EFFC9325898704F63FD0D111F" box="[591,684,594,619]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="16" pageNumber="1528">Figure 7</figureCitation>
(h)).......................................................................... 9
</keyLead>
</paragraph>
<paragraph id="DE0736DBFFC932589A804F45FB2211C4" blockId="16.[180,1157,143,1035]" pageId="16" pageNumber="1528">
<keyLead id="654995EEFFC932589A804F45FB2211C4" pageId="16" pageNumber="1528">
- Scutellum with dark markings (
<figureCitation id="46832A5EFFC9325898774F45FD0A11F9" box="[584,683,628,653]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="16" pageNumber="1528">Figure 7</figureCitation>
(i)), which can sometimes be diffuse and poorly defined................................................................................................................................... 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="654C2E7EFFC932589A804F8BFB25104F" pageId="16" pageNumber="1528">
<paragraph id="DE0736DBFFC932589A804F8BFEB71181" blockId="16.[180,1157,143,1035]" pageId="16" pageNumber="1528">
<keyLead id="654995EEFFC932589A804F8BFEB71181" pageId="16" pageNumber="1528">
9. Hind tarsomeres yellowish (
<figureCitation id="46832A5EFFC93258982B4F8BFDDC11A7" box="[532,637,698,723]" captionStart="Figure 25" captionStartId="37.[160,225,1613,1636]" captionTargetBox="[174,1142,150,1573]" captionTargetId="figure-3@37.[163,1153,142,1581]" captionTargetPageId="37" captionText="Figure 25. Formosargus variegatus, male, holotype [UZMC]. (a) Habitus, dorsal view.(b) Habitus, lateral view. Arrow points to a modified structure in the mid leg. (c) Labels (© Natural History Museum of Denmark, University of Copenhagen)." figureDoi="http://doi.org/10.5281/zenodo.7318238" httpUri="https://zenodo.org/record/7318238/files/figure.png" pageId="16" pageNumber="1528">Figure 25</figureCitation>
); 
<collectingCountry id="A6AF764BFFC9325898AD4F8BFCB111A7" box="[658,784,698,723]" name="Philippines" pageId="16" pageNumber="1528">Philippines</collectingCountry>
: 
<collectingRegion id="1C7CF839FFC9325899254F8BFCD611A7" box="[794,887,698,723]" country="Philippines" name="Palawan" pageId="16" pageNumber="1528">Palawan</collectingRegion>
..... 
<taxonomicName id="19B84D58FFC9325899984F8BFEB71181" ID-CoL="6JHMP" authority="James, 1969" authorityName="James" authorityYear="1969" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="1528" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="ECCCEAC9FFC9325899984F8BFB9011A7" box="[935,1073,698,723]" italics="true" pageId="16" pageNumber="1528">F. variegatus</emphasis>
<bibRefCitation id="BA294B2AFFC932589E064F8BFEB71181" author="James MT" pageId="16" pageNumber="1528" pagination="339 - 350" refId="ref19004" refString="James MT. 1969. The soldier flies (Diptera, Stratiomyidae) collected in the Philippine Islands by the Noona Dan Expedition. Entomol Medd. 37 (4): 339 - 350." type="journal article" year="1969">James, 1969</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DE0736DBFFC932589A804FCEFB25104F" blockId="16.[180,1157,143,1035]" pageId="16" pageNumber="1528">
<keyLead id="654995EEFFC932589A804FCEFB25104F" pageId="16" pageNumber="1528">
- Second hind tarsomere white and last three dark brown (
<figureCitation id="46832A5EFFC9325899404FCEFC4D106C" box="[895,1004,767,792]" captionStart="Figure 20" captionStartId="32.[160,225,1630,1653]" captionTargetBox="[171,1149,154,1593]" captionTargetId="figure-3@32.[163,1153,142,1598]" captionTargetPageId="32" captionText="Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively." figureDoi="http://doi.org/10.5281/zenodo.7318228" httpUri="https://zenodo.org/record/7318228/files/figure.png" pageId="16" pageNumber="1528">Figure 20</figureCitation>
(c)); 
<collectingCountry id="A6AF764BFFC932589E214FCEFB21106C" box="[1054,1152,767,792]" name="Malaysia" pageId="16" pageNumber="1528">Malaysia</collectingCountry>
, 
<collectingCountry id="A6AF764BFFC932589AE34E13FE9D104F" box="[220,316,802,827]" name="Thailand" pageId="16" pageNumber="1528">Thailand</collectingCountry>
........................................................................................................... 
<taxonomicName id="19B84D58FFC93258999B4E13FB84104F" ID-CoL="BCY3D" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[932,1061,802,827]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="1528" phylum="Arthropoda" rank="species" species="trivittatus" status="sp. nov.">
<emphasis id="ECCCEAC9FFC93258999B4E13FB84104F" box="[932,1061,802,827]" italics="true" pageId="16" pageNumber="1528">F. trivittatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFC932589E124E13FB25104F" box="[1069,1156,802,827]" pageId="16" pageNumber="1528" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="654C2E7EFFC932589A8B4E75FB25177F" pageId="16" pageNumber="1528">
<paragraph id="DE0736DBFFC932589A8B4E75FB2510D7" blockId="16.[180,1157,143,1035]" pageId="16" pageNumber="1528">
<keyLead id="654995EEFFC932589A8B4E75FB2510D7" pageId="16" pageNumber="1528">
10. Anterior margin of black frons to yellow lower frons straight (
<figureCitation id="46832A5EFFC9325899B84E75FC451029" box="[903,996,836,861]" captionStart="Figure 6" captionStartId="10.[160,225,1285,1308]" captionTargetBox="[167,1149,145,1251]" captionTargetId="figure-99@10.[163,1153,142,1253]" captionTargetPageId="10" captionText="Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi="http://doi.org/10.5281/zenodo.7318193" httpUri="https://zenodo.org/record/7318193/files/figure.png" pageId="16" pageNumber="1528">Figure 6</figureCitation>
(f)); male frons extremely narrow, only slightly broader than ocellar triangle (
<figureCitation id="46832A5EFFC9325899964E56FBAB10F4" box="[937,1034,871,896]" captionStart="Figure 6" captionStartId="10.[160,225,1285,1308]" captionTargetBox="[167,1149,145,1251]" captionTargetId="figure-99@10.[163,1153,142,1253]" captionTargetPageId="10" captionText="Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi="http://doi.org/10.5281/zenodo.7318193" httpUri="https://zenodo.org/record/7318193/files/figure.png" pageId="16" pageNumber="1528">Figure 6</figureCitation>
(f)); female unknown; 
<collectingCountry id="A6AF764BFFC932589B6D4EBBFE7110D7" box="[338,464,906,931]" name="Philippines" pageId="16" pageNumber="1528">Philippines</collectingCountry>
: 
<collectingRegion id="1C7CF839FFC932589BE54EBBFD8C10D7" box="[474,557,906,931]" country="Philippines" name="Laguna" pageId="16" pageNumber="1528">Laguna</collectingRegion>
................................................................. 
<taxonomicName id="19B84D58FFC9325899944EBBFB8410D7" ID-CoL="BCY3F" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[939,1061,906,931]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="1528" phylum="Arthropoda" rank="species" species="woodleyi" status="sp. nov.">
<emphasis id="ECCCEAC9FFC9325899944EBBFB8410D7" box="[939,1061,906,931]" italics="true" pageId="16" pageNumber="1528">F. woodleyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFC932589E124EBBFB2510D7" box="[1069,1156,906,931]" pageId="16" pageNumber="1528" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="DE0736DBFFC932589A804E9DFB25177F" blockId="16.[180,1157,143,1035]" pageId="16" pageNumber="1528">
<keyLead id="654995EEFFC932589A804E9DFB25177F" pageId="16" pageNumber="1528">
- Anterior margin of black frons with median triangular extension pointing towards antennal insertion (
<figureCitation id="46832A5EFFC932589B834EFEFDBC109C" box="[444,541,975,1000]" captionStart="Figure 5" captionStartId="9.[160,225,1289,1312]" captionTargetBox="[167,1149,145,1251]" captionTargetId="figure-113@9.[163,1153,142,1257]" captionTargetPageId="9" captionText="Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi="http://doi.org/10.5281/zenodo.7318191" httpUri="https://zenodo.org/record/7318191/files/figure.png" pageId="16" pageNumber="1528">Figure 5</figureCitation>
(f)); frons wider, more than twice as broad as ocellar triangle (
<figureCitation id="46832A5EFFC932589B7F4EC3FE3C177F" box="[320,413,1010,1035]" captionStart="Figure 5" captionStartId="9.[160,225,1289,1312]" captionTargetBox="[167,1149,145,1251]" captionTargetId="figure-113@9.[163,1153,142,1257]" captionTargetPageId="9" captionText="Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi="http://doi.org/10.5281/zenodo.7318191" httpUri="https://zenodo.org/record/7318191/files/figure.png" pageId="16" pageNumber="1528">Figure 5</figureCitation>
(f)); 
<collectingCountry id="A6AF764BFFC932589BF84EC3FD97177F" box="[455,566,1010,1035]" name="Indonesia" pageId="16" pageNumber="1528">Indonesia</collectingCountry>
: 
<collectingRegion id="1C7CF839FFC93258987E4EC3FD0E177F" box="[577,687,1010,1035]" country="Indonesia" name="Jawa Barat" pageId="16" pageNumber="1528">West Java</collectingRegion>
.......... 
<taxonomicName id="19B84D58FFC9325898C14EC3FB25177F" authority="(Adisoemarto, 1974)" authorityName="Fachin &amp; Hauser" authorityYear="2022" baseAuthorityName="Adisoemarto" baseAuthorityYear="1974" box="[766,1156,1010,1035]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="1528" phylum="Arthropoda" rank="species" species="sagittocerus">
<emphasis id="ECCCEAC9FFC9325898C14EC3FC38177F" box="[766,921,1010,1035]" italics="true" pageId="16" pageNumber="1528">F. sagittocerus</emphasis>
(
<bibRefCitation id="BA294B2AFFC9325899984EC3FBDD177F" author="Adisoemarto S" box="[935,1148,1010,1035]" pageId="16" pageNumber="1528" pagination="69 - 71" refId="ref18465" refString="Adisoemarto S. 1974. A new genus of soldier flies from Indonesia (Diptera: Stratiomyidae). Treubia. 28 (3): 69 - 71." type="journal article" year="1974">Adisoemarto, 1974</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>