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<document id="B6756ADB0B9CEF0C817CBDFFE350FD7D" ID-DOI="10.11646/zootaxa.5567.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14703735" ID-ZooBank="AF6254F6-CE26-41FE-BB85-A3FFD16FAA39" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737371447294" checkinUser="plazi" docAuthor="VIRAKTAMATH, C. A., WEBB, M. D. &amp; YESHWANTH, H. M." docDate="2025" docId="03B75F43F044FF92FF4872B833D12547" docLanguage="en" docName="zootaxa.5567.1.1.pdf" docOrigin="Zootaxa 5567 (1)" docSource="https://doi.org/10.11646/zootaxa.5567.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Cicurbanus VIRAKTAMATH, WEBB &amp; YESHWANTH, 2025, gen. nov." docType="treatment" docVersion="2" lastPageNumber="15" masterDocId="FF8E273BF049FF9CFFDF7756364D2343" masterDocTitle="Review of the Monobazus-group leafhopper genera (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent with description of five new genera and twenty-eight new species" masterLastPageNumber="105" masterPageNumber="1" pageNumber="14" updateTime="1737389185710" updateUser="ExternalLinkService">
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<mods:title id="530401B6CE4418F5677B47EC930DEF41">Review of the Monobazus-group leafhopper genera (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent with description of five new genera and twenty-eight new species</mods:title>
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<mods:namePart id="74ACDEA49DD66ED17AE29022E6D54B0D">VIRAKTAMATH, C. A.</mods:namePart>
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<mods:namePart id="FDA9F1CEABD274DFCBA89C97FC4597BA">WEBB, M. D.</mods:namePart>
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<subSubSection id="C304BDDEF044FF91FF4872B837D7254A" box="[151,410,1518,1545]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BA1EE55F044FF91FF4872B837D7254A" blockId="13.[151,410,1518,1545]" box="[151,410,1518,1545]" pageId="13" pageNumber="14">
<heading id="D0E95939F044FF91FF4872B837D7254A" bold="true" box="[151,410,1518,1545]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="4">
<emphasis id="B96A3247F044FF91FF4872B837D7254A" bold="true" box="[151,410,1518,1545]" pageId="13" pageNumber="14">
<taxonomicName id="4C1E95D6F044FF91FF4872B83768254A" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[151,293,1518,1545]" class="Insecta" family="Cicadellidae" genus="Cicurbanus" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96A3247F044FF91FF4872B83768254A" bold="true" box="[151,293,1518,1545]" italics="true" pageId="13" pageNumber="14">Cicurbanus</emphasis>
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<taxonomicNameLabel id="A2598F3CF044FF91FEF372B937D7254A" box="[300,410,1519,1545]" pageId="13" pageNumber="14" rank="genus">gen. nov.</taxonomicNameLabel>
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<subSubSection id="C304BDDEF044FF92FF48716232E12776" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BA1EE55F044FF91FF48716235482508" blockId="13.[151,773,1587,1612]" box="[151,773,1587,1612]" pageId="13" pageNumber="14">
<typeStatus id="54A550F7F044FF91FF4871623684250F" box="[151,201,1588,1612]" pageId="13" pageNumber="14">Type</typeStatus>
species:
<taxonomicName id="4C1E95D6F044FF91FEF9716534182508" authority="Melichar" authorityName="Melichar" box="[294,597,1587,1611]" class="Insecta" family="Cicadellidae" genus="Thamnotettix" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="greeni">
<emphasis id="B96A3247F044FF91FEF9716537BF2508" box="[294,498,1587,1611]" italics="true" pageId="13" pageNumber="14">Thamnotettix greeni</emphasis>
Melichar
</taxonomicName>
, here designated.
</paragraph>
<paragraph id="8BA1EE55F044FF91FF48712C346B2428" blockId="13.[151,1437,1657,2008]" pageId="13" pageNumber="14">
<emphasis id="B96A3247F044FF91FF48712C374425D0" box="[151,265,1658,1683]" italics="true" pageId="13" pageNumber="14">Diagnosis</emphasis>
. Often with two prominent black spots on crown in addition to anterior marginal small brown spot on either side of median line. Pygofer with blade-like ventral process directed posteriorly either twisted or serrated, style lacking subapical process, aedeagus short with prominent dorsal apodeme that is longer than aedeagal shaft, shaft abruptly anteriorly curved near apex and with one apical spine on ventral surface, gonopore apical. It differs from
<taxonomicName id="4C1E95D6F044FF91FF0A705F371B2460" box="[213,342,1801,1827]" class="Insecta" family="Cicadellidae" genus="Monobazus" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F044FF91FF0A705F371B2460" box="[213,342,1801,1827]" italics="true" pageId="13" pageNumber="14">Monobazus</emphasis>
</taxonomicName>
in not having ventral processes on the aedeagal shaft and from
<taxonomicName id="4C1E95D6F044FF91FBFD705C329A2460" authorityName="Dlabola" authorityYear="1967" box="[1058,1239,1802,1827]" class="Insecta" genus="Mavromoustaca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F044FF91FBFD705C329A2460" box="[1058,1239,1802,1827]" italics="true" pageId="13" pageNumber="14">Mavromoustaca</emphasis>
</taxonomicName>
in the absence of basilateral processes on the aedeagal shaft and from both these genera it can be distinguished by the aedeagus not having a well-developed preatrium.
</paragraph>
<paragraph id="8BA1EE55F044FF91FF18702032DD24F0" blockId="13.[151,1437,1657,2008]" pageId="13" pageNumber="14">
<emphasis id="B96A3247F044FF91FF187020370624CC" box="[199,331,1910,1935]" italics="true" pageId="13" pageNumber="14">Description</emphasis>
. Yellowish ochraceous with prominent pair of black spots on crown and a pair of smaller brown spots on fore margin, basal triangles of exposed mesonotum with black or brown triangles.
</paragraph>
<paragraph id="8BA1EE55F044FF92FF1870EB32302122" blockId="13.[151,1437,1657,2008]" lastBlockId="14.[151,1437,150,1221]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
Medium sized species measuring
<quantity id="4CE643B0F044FF91FD9470EB3498249B" box="[587,725,1981,2008]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="13" pageNumber="14" unit="mm" value="5.0" valueMax="5.5" valueMin="4.5">4.55.5 mm</quantity>
long. Head distinctly narrower than pronotum, crown surface with apical half shagreen, posterior half polished; median sulcus about 0.5 as long as crown; fore margin rounded to face. Eyes large, their outer margin in line with outer margin of crown, obliquely extended on anterolateral angles of pronotum. Ocelli prominent placed a distance less than own diameter away from eyes. Antennae long, almost reaching half-length of body. Face including eyes slightly longer than wide or as wide as long; frontoclypeus somewhat narrow; clypellus broad apically. Pronotum slightly convex dorsally, about 1.51.8× as long as crown; with lateral margins carinate, surface polished, posterior margin almost straight. Exposed mesonotum slightly longer than pronotum. Forewing with claval veins separate, inner claval vein connected to claval suture by cross vein, three anteapical and four apical cells, inner anteapical cell open behind, appendix narrow not extending beyond middle of second apical cell. Pro-tibia comparatively long and slender compared to meso tibia; AV row of setae composed of 15 conical short peg-like in basal 0.66, AM row of setae represented by hair-like AM1, IC row of setae 15, slender hair-like. Metafemur with 2+2+1 apical macrosetae. Meta tibia with PD row macrosetae long, as long or longer than half length of fore tibia; Metabasitarsomere with 3 platellae and one stout tapered seta on either side. Female sternite VII half as long laterally as wide, with broadly V-shaped excavation on posterior margin.
</paragraph>
<paragraph id="8BA1EE55F047FF92FF18753D36AE208A" blockId="14.[151,1437,150,1221]" pageId="14" pageNumber="15">
<emphasis id="B96A3247F047FF92FF18753D372221C6" box="[199,367,619,645]" italics="true" pageId="14" pageNumber="15">Male genitalia</emphasis>
. Pygofer with or without anterior apodemes, tergum short, lobes well produced posteriorly, making tergum deeply U-shaped in dorsal view, well-developed ventral process present and exceeding outer margin of pygofer, with numerous macrosetae in distal 0.50.75 length, ventral margin convex, posterodorsal angle subconically rounded. Valve with posterior margin angularly produced medially. Subgenital plates triangular, elongate, about 2.22.5× as long as basal width, with marginal row of uniseriate macrosetae in basal 0.75 region and numerous hair-like setae in distal half. Style without well-developed subapical lobe, with smooth surface and hair-like setae on lateral surface before apex. Connective Y-shaped, with stem longer than arms. Aedeagus symmetrical, with well-developed dorsal apodeme much longer than shaft, shaft comparatively shorter, sharply anteriorly curved near apex, with spine-like apical process on ventral surface, gonopore apical, shaft with lateral lamellate flanges or ridges.
</paragraph>
<paragraph id="8BA1EE55F047FF92FF18748535A720AE" blockId="14.[151,1437,150,1221]" box="[199,1002,979,1005]" pageId="14" pageNumber="15">
<emphasis id="B96A3247F047FF92FF18748537CB20AE" box="[199,390,979,1005]" italics="true" pageId="14" pageNumber="15">Female genitalia</emphasis>
. First and second pair of valvulae as in
<taxonomicName id="4C1E95D6F047FF92FC9B748535A720AE" authority="Distant." authorityName="Distant." box="[836,1002,979,1005]" class="Insecta" family="Cicadellidae" genus="Banus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F047FF92FC9B748535C720AF" box="[836,906,979,1004]" italics="true" pageId="14" pageNumber="15">Banus</emphasis>
Distant.
</taxonomicName>
</paragraph>
<paragraph id="8BA1EE55F047FF92FF1874A132E12776" blockId="14.[151,1437,150,1221]" pageId="14" pageNumber="15">
<emphasis id="B96A3247F047FF92FF1874A137722752" box="[199,319,1015,1041]" italics="true" pageId="14" pageNumber="15">Etymology</emphasis>
. The generic name is a combination of the Latin adjective cicur, -uris (= tame, mild) and
<taxonomicName id="4C1E95D6F047FF92FACE74A1331A2753" box="[1297,1367,1015,1040]" class="Insecta" family="Cicadellidae" genus="Banus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F047FF92FACE74A1331A2753" box="[1297,1367,1015,1040]" italics="true" pageId="14" pageNumber="15">Banus</emphasis>
</taxonomicName>
(firstcentury desert-dwelling Jewish ascetic, teacher of Titus Flavius Josephus), gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C304BDDEF047FF92FF18736935FA2786" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BA1EE55F047FF92FF18736935FA2786" blockId="14.[151,1437,150,1221]" pageId="14" pageNumber="15">
<emphasis id="B96A3247F047FF92FF1873693779271A" bold="true" box="[199,308,1087,1113]" pageId="14" pageNumber="15">Remarks</emphasis>
. Genus
<taxonomicName id="4C1E95D6F047FF92FE517369345C271A" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[398,529,1087,1113]" class="Insecta" family="Cicadellidae" genus="Cicurbanus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F047FF92FE517369345C271A" box="[398,529,1087,1113]" italics="true" pageId="14" pageNumber="15">Cicurbanus</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C1E95D6F047FF92FD697369357A271A" box="[694,823,1087,1113]" class="Insecta" family="Cicadellidae" genus="Monobazus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F047FF92FD697369357A271A" box="[694,823,1087,1113]" italics="true" pageId="14" pageNumber="15">Monobazus</emphasis>
</taxonomicName>
group of genera and can be recognized by the markings on the crown (see key), male pygofer with ventral process directed posteriorly, style without subapical lobe and aedeagus short, with elongate dorsal apodeme, and shaft with lateral ridges or flanges, with apical spine-like process on ventral surface but lacking a ventral process in the basal half-length.
</paragraph>
</subSubSection>
<subSubSection id="C304BDDEF047FF92FF48724133D12547" pageId="14" pageNumber="15" type="key">
<paragraph id="8BA1EE55F047FF92FF48724137B12672" blockId="14.[151,508,1302,1329]" box="[151,508,1302,1329]" pageId="14" pageNumber="15">
<emphasis id="B96A3247F047FF92FF48724137B12672" bold="true" box="[151,508,1302,1329]" pageId="14" pageNumber="15">
Key to species of
<taxonomicName id="4C1E95D6F047FF92FEB1724037B12672" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[366,508,1302,1329]" class="Insecta" family="Cicadellidae" genus="Cicurbanus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B96A3247F047FF92FEB1724037B12672" bold="true" box="[366,508,1302,1329]" italics="true" pageId="14" pageNumber="15">Cicurbanus</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<key id="FF8FB964F047FF92FF48720A33D12547" pageId="14" pageNumber="15">
<keyStep id="30EAF6F0F047FF92FF48720A33D026CC" pageId="14" pageNumber="15">
<paragraph id="8BA1EE55F047FF92FF48720A33D12632" blockId="14.[151,1437,1372,1540]" box="[151,1436,1372,1394]" pageId="14" pageNumber="15">
<keyLead id="30EF4D60F047FF92FF48720A33D12632" box="[151,1436,1372,1394]" pageId="14" pageNumber="15">1. Crown with two prominent black spots (1MO).............................................................. 2</keyLead>
</paragraph>
<paragraph id="8BA1EE55F047FF92FF48722F33D026CC" blockId="14.[151,1437,1372,1540]" box="[151,1437,1401,1423]" pageId="14" pageNumber="15">
<keyLead id="30EF4D60F047FF92FF48722F33D026CC" box="[151,1437,1401,1423]" pageId="14" pageNumber="15">
- Crown with transverse pale reddish brown stripe margined by a few dark brown spots (
<figureCitation id="1325F2D0F047FF92FBF9722F322326CD" box="[1062,1134,1401,1423]" captionStart="FIGURES 7" captionStartId="48.[151,264,1909,1933]" captionTargetBox="[201,1388,184,1880]" captionTargetId="figure-26@48.[195,1393,181,1884]" captionTargetPageId="48" captionText="FIGURES 7AL. Scaphoideini genera, habitus, dorsal and lateral view and female terminal segments (L, N).AD, Longicornus grandis (Pruthi), male (AB) and female (CD); EG, Monobazus himlayensis Distant syntype female, dorsal (E), lateral view (F), face (G) and ventral view of abdomen (H); IJ, Banus oblatus Distant, type female from Burma, dorsal view (I) and ventral view of abdomen (J); K Monobazus fletcheri (Distant), type female; L, Cicurbanus notatus (Distant), type male." figureDoi="http://doi.org/10.5281/zenodo.14703763" httpUri="https://zenodo.org/record/14703763/files/figure.png" pageId="14" pageNumber="15">Fig. 7L</figureCitation>
)........
<taxonomicName id="4C1E95D6F047FF92FB01722F33D026CC" authority="(Distant)" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" baseAuthorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" baseAuthorityYear="2025" box="[1246,1437,1401,1423]" class="Insecta" family="Cicadellidae" genus="Cicurbanus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="notatus">
<emphasis id="B96A3247F047FF92FB01722F330C26CC" box="[1246,1345,1401,1423]" italics="true" pageId="14" pageNumber="15">C. notatus</emphasis>
(Distant)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EAF6F0F047FF92FF4872C033D12547" pageId="14" pageNumber="15">
<paragraph id="8BA1EE55F047FF92FF4872C033D12689" blockId="14.[151,1437,1372,1540]" pageId="14" pageNumber="15">
<keyLead id="30EF4D60F047FF92FF4872C033D12689" pageId="14" pageNumber="15">
2. Black spots on crown round, mesonotum with basal triangles black (
<figureCitation id="1325F2D0F047FF92FCB572C035F726E8" box="[874,954,1430,1452]" captionStart="FIGURES 1" captionStartId="42.[151,264,1870,1894]" captionTargetBox="[180,1388,186,1841]" captionTargetId="figure-26@42.[175,1415,181,1846]" captionTargetPageId="42" captionText="FIGURES 1AR. Monobazus group of genera, habitus, dorsal and lateral views. AB, Banus curvatus sp. nov., male; CF, B. spiculatus sp. nov., male (CD) and female (EF); GJ, B. umbratus (Melichar), male (GH) and female (IJ); KL, Chandibanus dorsospinosus gen. et sp. nov., holotype male; MN, Cicurbanus greeni (Melichar), male; OR, C. serratus sp. nov., holotype male (OP), paratype female (QR)." figureDoi="http://doi.org/10.5281/zenodo.14703739" httpUri="https://zenodo.org/record/14703739/files/figure.png" pageId="14" pageNumber="15">Fig. 1M</figureCitation>
); male pygofer ventral process smooth and twisted (
<figureCitation id="1325F2D0F047FF92FF3672E2377D268A" box="[233,304,1460,1482]" captionStart="FIGURES 18" captionStartId="59.[151,264,1892,1916]" captionTargetBox="[156,1430,186,1863]" captionTargetId="figure-17@59.[151,1435,181,1868]" captionTargetPageId="59" captionText="FIGURES 18AI. Cicurbanus greeni (Melichar), male genitalia: A, pygofer, lateral view; B, pygofer ventral process, lateral view; C, valve and subgenital plate, ventral view; D, connective, dorsal view; EF, style, dorsal views at different angles; G, aedeagus posteroventral view; H aedeagus, anterodorsal view; I, aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14703795" httpUri="https://zenodo.org/record/14703795/files/figure.png" pageId="14" pageNumber="15">Figs 18</figureCitation>
AB)..........................................................................
<taxonomicName id="4C1E95D6F047FF92FB0872E233D12689" ID-CoL="563PW" authority="(Melichar)" baseAuthorityName="Melichar" box="[1239,1436,1460,1482]" class="Insecta" family="Cicadellidae" genus="Thamnotettix" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="greeni">
<emphasis id="B96A3247F047FF92FB0872E23362268A" box="[1239,1327,1460,1481]" italics="true" pageId="14" pageNumber="15">C. greeni</emphasis>
(Melichar)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA1EE55F047FF92FF48728733D12547" blockId="14.[151,1437,1372,1540]" pageId="14" pageNumber="15">
<keyLead id="30EF4D60F047FF92FF48728733D12547" pageId="14" pageNumber="15">
- Black spots on crown rectangular, mesonotum without black basal triangles (
<figureCitation id="1325F2D0F047FF92FC1D728735B726A5" box="[962,1018,1489,1511]" captionStart="FIGURES 1" captionStartId="42.[151,264,1870,1894]" captionTargetBox="[180,1388,186,1841]" captionTargetId="figure-26@42.[175,1415,181,1846]" captionTargetPageId="42" captionText="FIGURES 1AR. Monobazus group of genera, habitus, dorsal and lateral views. AB, Banus curvatus sp. nov., male; CF, B. spiculatus sp. nov., male (CD) and female (EF); GJ, B. umbratus (Melichar), male (GH) and female (IJ); KL, Chandibanus dorsospinosus gen. et sp. nov., holotype male; MN, Cicurbanus greeni (Melichar), male; OR, C. serratus sp. nov., holotype male (OP), paratype female (QR)." figureDoi="http://doi.org/10.5281/zenodo.14703739" httpUri="https://zenodo.org/record/14703739/files/figure.png" pageId="14" pageNumber="15">Figs 1</figureCitation>
OQ); male pygofer ventral process serrated and not twisted (
<figureCitation id="1325F2D0F047FF92FE5E72B837862540" box="[385,459,1518,1540]" captionStart="FIGURES 20" captionStartId="61.[151,264,1942,1966]" captionTargetBox="[156,1430,186,1913]" captionTargetId="figure-17@61.[151,1435,181,1918]" captionTargetPageId="61" captionText="FIGURES 20AJ. Cicurbanus serratus sp. nov., male genitalia: A, pygofer, lateral view; B, pygofer ventral process, lateral view; C, subgenital plate, ventral view; D, connective, dorsal view; E, style, dorsal view; F, aedeagus dorsal view; G, aedeagus anterodorsal view; H, aedeagal shaft apex, dorsal view; IJ, aedeagus, lateral views." figureDoi="http://doi.org/10.5281/zenodo.14703799" httpUri="https://zenodo.org/record/14703799/files/figure.png" pageId="14" pageNumber="15">Figs 20</figureCitation>
AB)..............................................................
<taxonomicName id="4C1E95D6F047FF92FB0272B933042547" ID-CoL="8R66V" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[1245,1353,1519,1540]" class="Insecta" family="Cicadellidae" genus="Cicurbanus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="serratus" status="sp. nov.">
<emphasis id="B96A3247F047FF92FB0272B933042547" box="[1245,1353,1519,1540]" italics="true" pageId="14" pageNumber="15">C. serratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2598F3CF047FF92FA9172B833D12547" box="[1358,1436,1518,1540]" pageId="14" pageNumber="15" rank="species">
<emphasis id="B96A3247F047FF92FA9172B833DB2540" bold="true" box="[1358,1430,1518,1539]" pageId="14" pageNumber="15">sp. nov</emphasis>
.
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>