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<mods:title id="7769CA4B253A07BA9D796E684A828F52">Review of the genus Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae) of Western Palaearctic region</mods:title>
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Key to Western Palaearctic species of
<taxonomicName id="4C0A3ABB90135A6B275FFA022FE9FA0C" ID-CoL="8VRGZ" authority="Leach, 1819" authorityName="Leach" authorityYear="1819" box="[624,761,1481,1508]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="genus">
<emphasis id="B97E9D2A90135A6B275FFA022FE9FA0C" bold="true" box="[624,761,1481,1508]" italics="true" pageId="19" pageNumber="470">Anthobium</emphasis>
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1 Body elongate. Pronotum relatively narrow, about one and a half as broad as long. Elytra distinctly longer than broad. Median lobe and apical portions of parameres significantly broadened (
<figureCitation id="13315DBD90135A6B2660F9E72E86F9A9" box="[847,918,1580,1602]" captionStart="FIGURES 1923" captionStartId="14.[151,264,1852,1876]" captionTargetBox="[163,1421,246,1814]" captionTargetId="figure-56@14.[151,1436,237,1829]" captionTargetPageId="14" captionText="FIGURES 1923. Habitus of Lathrimaeum (types) and Anthobium: 19L. moczarskii (holotype), 20L. metallicum (holotype), 21L. tenue (lectotype), 22L. laterale (holotype), 23A. ganglbaueri (Turkey, Ulu Dað). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240238" httpUri="https://zenodo.org/record/14240238/files/figure.png" pageId="19" pageNumber="470">
Fig.
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in
<bibRefCitation id="EF9B3CC990135A6B2696F9E72971F9A9" author="Shavrin, A. V." box="[953,1121,1580,1602]" pageId="19" pageNumber="470" pagination="301 - 346" refId="ref21525" refString="Shavrin, A. V. (2022 a) A revision of Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). V. Algidum, morosum and tectum groups, a new species of the fusculum group, and faunistic records. Zootaxa, 5104 (3), 301 - 346. https: // doi. org / 10.11646 / zootaxa. 5104.3.1" type="journal article" year="2022">Shavrin (2022a))</bibRefCitation>
. Habitus as in
<figureCitation id="13315DBD90135A6B21DAF9E7283FF9A9" box="[1269,1327,1580,1602]" captionStart="FIGURES 16" captionStartId="5.[151,264,1623,1647]" captionTargetBox="[157,1416,185,1591]" captionTargetId="figure-16@5.[151,1435,179,1597]" captionTargetPageId="5" captionText="FIGURES 16. Habitus of Lathrimaeum (types) and Anthobium: 1L. gracilicorne (holotype), 2L. abantense (holotype), 3L. urgelense (holotype), 4L. hamatum (holotype), 5A. melanocephalum (Georgia: Svaneti), 6L. longicorne (lectotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240228" httpUri="https://zenodo.org/record/14240228/files/figure.png" pageId="19" pageNumber="470">
Fig.
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in
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. Body reddish-brown to dark brown. Body large:
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. Middle Europe.......................
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<emphasis id="B97E9D2A90135A6B2061F982288DF9B6" box="[1358,1437,1609,1630]" italics="true" pageId="19" pageNumber="470">algidum</emphasis>
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<keyLead id="30FBE20D90135A6B25B8F9AC288CF971" pageId="19" pageNumber="470">- Body wide, somewhat shortened. Pronotum transverse, about twice as broad as long or slightly narrower. Median lobe and apical portions of parameres narrower. Body smaller......................................................... 2</keyLead>
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2 Forebody less elevated, somewhat flattened. Middle potion of pronotum evenly elevated, sometimes with indistinct longitudinal impression. Aedeagus as in Fig.
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. Habitus as in
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Fig.
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in
<bibRefCitation id="EF9B3CC990135A6B26E1F974297CF93C" author="Shavrin, A. V." box="[974,1132,1727,1749]" pageId="19" pageNumber="470" pagination="301 - 346" refId="ref21525" refString="Shavrin, A. V. (2022 a) A revision of Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). V. Algidum, morosum and tectum groups, a new species of the fusculum group, and faunistic records. Zootaxa, 5104 (3), 301 - 346. https: // doi. org / 10.11646 / zootaxa. 5104.3.1" type="journal article" year="2022">Shavrin (2022a)</bibRefCitation>
. Body brown to reddish-brown. Body length:
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. Transpalaearctic species....................................................
<taxonomicName id="4C0A3ABB90135A6B2073F916288CF91A" ID-CoL="9GCTW" baseAuthorityName="Paykull" baseAuthorityYear="1789" box="[1372,1436,1757,1778]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="tectum">
<emphasis id="B97E9D2A90135A6B2073F916288CF91A" box="[1372,1436,1757,1778]" italics="true" pageId="19" pageNumber="470">tectum</emphasis>
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<keyLead id="30FBE20D90135A6B25B8F931288CF8C4" pageId="19" pageNumber="470">- Forebody distinctly elevated, convex. Middle portion strongly elevated, usually with distinct longitudinal and/or mediobasal impressions......................................................................................... 3</keyLead>
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<keyLead id="30FBE20D90135A6B25B8F8FF288CF8A1" box="[151,1436,1844,1866]" pageId="19" pageNumber="470">3 Lateral edges of pronotum with distinct crenulation, sometimes forming indistinct mediolateral angles................. 4</keyLead>
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<keyLead id="30FBE20D90135A6B25B8F899288CF88F" box="[151,1436,1873,1895]" pageId="19" pageNumber="470">- Lateral edges of pronotum without crenulation.............................................................. 5</keyLead>
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4 Pronotum about twice as broad as long, with anterior angles relatively strongly protruded anteriad, from widest middle slightly more narrowed anteriad than posteriad, with slightly concave latero-apical margin and lateral edges, sometimes forming indistinct mediolateral angles; median impressions usually indistinct. Aedeagus as in
<figureCitation id="13315DBD90135A6B2138F862299BF857" box="[1047,1163,1961,1983]" captionStart="FIGURES 2430" captionStartId="15.[151,264,1514,1538]" captionTargetBox="[172,1415,200,1482]" captionTargetId="figure-16@15.[164,1423,181,1490]" captionTargetPageId="15" captionText="FIGURES 2430. Aedeagus of Anthobium (Figs 2425: A. moczarskii (Spain, Pinilla del Valle), Figs 2627: A. tenue (Georgia, Surami Pass), Fig. 28: A. ganglbaueri (Turkey, Bademli), Figs 2930: A. ganglbaueri (Turkey, Kastamonu): 24, 26, 2829 parameral view, 25, 27, 30lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14240242" httpUri="https://zenodo.org/record/14240242/files/figure.png" pageId="19" pageNumber="470">Figs. 2930</figureCitation>
. Habitus as in
<figureCitation id="13315DBD90135A6B2037F862284FF857" box="[1304,1375,1961,1983]" captionStart="FIGURES 2430" captionStartId="15.[151,264,1514,1538]" captionTargetBox="[172,1415,200,1482]" captionTargetId="figure-16@15.[164,1423,181,1490]" captionTargetPageId="15" captionText="FIGURES 2430. Aedeagus of Anthobium (Figs 2425: A. moczarskii (Spain, Pinilla del Valle), Figs 2627: A. tenue (Georgia, Surami Pass), Fig. 28: A. ganglbaueri (Turkey, Bademli), Figs 2930: A. ganglbaueri (Turkey, Kastamonu): 24, 26, 2829 parameral view, 25, 27, 30lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14240242" httpUri="https://zenodo.org/record/14240242/files/figure.png" pageId="19" pageNumber="470">Fig. 24</figureCitation>
. Body paler, yellow-brown to brown. Body length:
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. South-eastern Europe,
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, Caucasus.........
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<emphasis id="B97E9D2A90135A6B2008F80C288CF834" box="[1319,1436,1991,2012]" italics="true" pageId="19" pageNumber="470">ganglbaueri</emphasis>
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</keyLead>
</paragraph>
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- Pronotum smaller, distinctly less transverse, with anterior angles slightly protruded anteriad, from widest middle relatively evenly narrowed both anteriad and posteriad, without concave latero-apical margin and mediolateral angles; median impressions deep. Aedeagus as in Figs.
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. Habitus as in
<figureCitation id="13315DBD90145A6C26EBFF7E2EEBFF22" box="[964,1019,181,203]" captionStart="FIGURES 16" captionStartId="5.[151,264,1623,1647]" captionTargetBox="[157,1416,185,1591]" captionTargetId="figure-16@5.[151,1435,179,1597]" captionTargetPageId="5" captionText="FIGURES 16. Habitus of Lathrimaeum (types) and Anthobium: 1L. gracilicorne (holotype), 2L. abantense (holotype), 3L. urgelense (holotype), 4L. hamatum (holotype), 5A. melanocephalum (Georgia: Svaneti), 6L. longicorne (lectotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240228" httpUri="https://zenodo.org/record/14240228/files/figure.png" pageId="20" pageNumber="471">
Fig.
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in
<bibRefCitation id="EF9B3CC990145A6C2138FF7E2809FF22" author="Shavrin, A. V. &amp; Smetana, A." box="[1047,1305,181,203]" pageId="20" pageNumber="471" pagination="451 - 506" refId="ref21783" refString="Shavrin, A. V. &amp; Smetana, A. (2018) A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa, 4508 (4), 451 - 506. https: // doi. org / 10.11646 / zootaxa. 4508.4.1" type="journal article" year="2018">Shavrin &amp; Smetana (2018)</bibRefCitation>
. Body darker, brown to black. Body length:
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. Palaearctic species.......................................
<taxonomicName id="4C0A3ABB90145A6C2018FF05288CFF00" ID-CoL="5VCDR" baseAuthorityName="Erichson" baseAuthorityYear="1839" box="[1335,1436,206,232]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="fusculum">
<emphasis id="B97E9D2A90145A6C2018FF05288CFF00" box="[1335,1436,206,232]" italics="true" pageId="20" pageNumber="471">fusculum</emphasis>
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<keyLead id="30FBE20D90145A6C25B8FF24288CFEED" box="[151,1436,239,261]" pageId="20" pageNumber="471">5 Anterior angles of pronotum strongly protruded anteriad...................................................... 6</keyLead>
</paragraph>
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<keyLead id="30FBE20D90145A6C25B8FEC6288CFECA" box="[151,1436,269,291]" pageId="20" pageNumber="471">- Anterior angles of pronotum slightly protruded anteriad...................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FE599D90145A6C25B8FEE1288CFE3A" pageId="20" pageNumber="471">
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6 Laterobasal margins of pronotum strongly narrowed posteriad; median pronotal impressions deep. Apical part of median lobe moderately narrow; parameres significantly longer than apex of median lobe (
<figureCitation id="13315DBD90145A6C26E8FE8C2926FEB5" box="[967,1078,327,349]" captionStart="FIGURES 712" captionStartId="6.[151,264,1797,1821]" captionTargetBox="[159,1404,189,1766]" captionTargetId="figure-28@6.[151,1436,181,1774]" captionTargetPageId="6" captionText="FIGURES 712. Aedeagus of Lathrimaeum (types) and Anthobium (Fig. 7: L. gracilicorne (holotype), Fig. 8: L. abantense (holotype), Fig. 9: A. atrocephalum (Algeria, Adekar), Figs 1012: A. melanocephalum (Fig. 10: Georgia, Kherkhvashi, Figs 1112: Karachay-Cherkessia, Malaya Khatipara Mt.): 711parameral view, 12lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14240230" httpUri="https://zenodo.org/record/14240230/files/figure.png" pageId="20" pageNumber="471">Figs 1011</figureCitation>
). Habitus as in
<figureCitation id="13315DBD90145A6C21E8FE8C280DFEB5" box="[1223,1309,327,349]" captionStart="FIGURES 16" captionStartId="5.[151,264,1623,1647]" captionTargetBox="[157,1416,185,1591]" captionTargetId="figure-16@5.[151,1435,179,1597]" captionTargetPageId="5" captionText="FIGURES 16. Habitus of Lathrimaeum (types) and Anthobium: 1L. gracilicorne (holotype), 2L. abantense (holotype), 3L. urgelense (holotype), 4L. hamatum (holotype), 5A. melanocephalum (Georgia: Svaneti), 6L. longicorne (lectotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240228" httpUri="https://zenodo.org/record/14240228/files/figure.png" pageId="20" pageNumber="471">Figs 45</figureCitation>
. Body brown to reddish brown. Body length:
<quantity id="4CF2ECDD90145A6C273CFEAE2FB1FE92" box="[531,673,357,378]" metricMagnitude="-3" metricUnit="m" metricValue="3.635" metricValueMax="4.07" metricValueMin="3.2" pageId="20" pageNumber="471" unit="mm" value="3.635" valueMax="4.07" valueMin="3.2">3.204.07 mm</quantity>
. Europe,
<collectingCountry id="F31D01A890145A6C27D4FEAE2E51FE93" box="[763,833,357,379]" name="Turkey" pageId="20" pageNumber="471">Turkey</collectingCountry>
, Caucasus...........................
<taxonomicName id="4C0A3ABB90145A6C21D7FEAE288CFE92" ID-CoL="5VCF5" baseAuthorityName="Illiger" baseAuthorityYear="1794" box="[1272,1436,357,378]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="melanocephalum">
<emphasis id="B97E9D2A90145A6C21D7FEAE288CFE92" box="[1272,1436,357,378]" italics="true" pageId="20" pageNumber="471">melanocephalum</emphasis>
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</keyLead>
</paragraph>
<paragraph id="8BB5413890145A6C25B8FE48288CFE3A" blockId="20.[151,1437,151,877]" pageId="20" pageNumber="471">
<keyLead id="30FBE20D90145A6C25B8FE48288CFE3A" pageId="20" pageNumber="471">
<emphasis id="B97E9D2A90145A6C25B8FE482D8FFE70" box="[151,159,387,408]" italics="true" pageId="20" pageNumber="471">-</emphasis>
Laterobasal margins of pronotum slightly narrowed posteriad; median pronotal impressions shallow.Apical part of median lobe wide; parameres indistinctly exceeding apex of median lobe (
<figureCitation id="13315DBD90145A6C2612FE542EA0FE5D" box="[829,944,415,437]" captionStart="FIGURES 1317" captionStartId="12.[151,264,1813,1837]" captionTargetBox="[159,1428,195,1780]" captionTargetId="figure-28@12.[151,1436,181,1789]" captionTargetPageId="12" captionText="FIGURES 1317. Aedeagus of Anthobium (Figs 1314: A. longicorne (Algeria, Tikjida), Fig. 15: L. longicorne (Algeria, Babor Mt.), Figs 1617: A. metallicum (Turkey, Isparta): 13, 1516parameral view, 14, 17lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14240232" httpUri="https://zenodo.org/record/14240232/files/figure.png" pageId="20" pageNumber="471">Figs 14, 16</figureCitation>
). Habitus as in
<figureCitation id="13315DBD90145A6C2164FE542997FE5D" box="[1099,1159,415,437]" captionStart="FIGURES 16" captionStartId="5.[151,264,1623,1647]" captionTargetBox="[157,1416,185,1591]" captionTargetId="figure-16@5.[151,1435,179,1597]" captionTargetPageId="5" captionText="FIGURES 16. Habitus of Lathrimaeum (types) and Anthobium: 1L. gracilicorne (holotype), 2L. abantense (holotype), 3L. urgelense (holotype), 4L. hamatum (holotype), 5A. melanocephalum (Georgia: Svaneti), 6L. longicorne (lectotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240228" httpUri="https://zenodo.org/record/14240228/files/figure.png" pageId="20" pageNumber="471">Fig. 6</figureCitation>
. Body reddish-brown. Body length:
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. North Africa..............................................................
<taxonomicName id="4C0A3ABB90145A6C201BFE76288CFE3A" ID-CoL="5VCDP" baseAuthorityName="Fauvel" baseAuthorityYear="1886" box="[1332,1436,445,466]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="longicorne">
<emphasis id="B97E9D2A90145A6C201BFE76288CFE3A" box="[1332,1436,445,466]" italics="true" pageId="20" pageNumber="471">longicorne</emphasis>
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7 Body yellowish or yellow-brown. Aedeagus as in Fig. 34h in
<bibRefCitation id="EF9B3CC990145A6C261FFE112EADFE07" author="Zanetti, A." box="[816,957,474,496]" pageId="20" pageNumber="471" pagination="49 - 117" refId="ref22293" refString="Zanetti, A. (2012) Unterfamilie Omaliinae. In: Assing, V. &amp; Schulke, M. (Eds.), Freude-Harde-Lohse-Klausnitzer - Die Kafer Mitteleuropas. Band 4. Staphylinidae I. Zweite neubearbeitete Auflage. Spektrum Akademischer Verlag, Heidelberg, pp. 49 - 117." type="book chapter" year="2012">Zanetti (2012)</bibRefCitation>
. Body length: 3.00
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. Europe, Caucasus..............................................................................................
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Body darker......................................................................................... 8
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<keyLead id="30FBE20D90145A6C25B8FDF9288CFDAF" box="[151,1436,562,584]" pageId="20" pageNumber="471">8 Pronotum without or with indistinct longitudinal impressions. Apical part of median lobe wide....................... 9</keyLead>
</paragraph>
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<emphasis id="B97E9D2A90145A6C25B8FD9B2D8FFD8D" box="[151,159,592,613]" italics="true" pageId="20" pageNumber="471">-</emphasis>
Pronotum with distinct longitudinal impression. Apical part of median lobe moderately narrow...................... 10
</keyLead>
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</keyStep>
<keyStep id="30FE599D90145A6C25B8FDA6288CFD32" pageId="20" pageNumber="471">
<paragraph id="8BB5413890145A6C25B8FDA6288CFD48" blockId="20.[151,1437,151,877]" pageId="20" pageNumber="471">
<keyLead id="30FBE20D90145A6C25B8FDA6288CFD48" pageId="20" pageNumber="471">
9 Punctation of pronotum and elytra moderately fine. Parameres slightly exceeding apex of median lobe, with distinctly broadened preapical parts (
<figureCitation id="13315DBD90145A6C2453FD412CD3FD77" box="[380,451,650,672]" captionStart="FIGURES 2430" captionStartId="15.[151,264,1514,1538]" captionTargetBox="[172,1415,200,1482]" captionTargetId="figure-16@15.[164,1423,181,1490]" captionTargetPageId="15" captionText="FIGURES 2430. Aedeagus of Anthobium (Figs 2425: A. moczarskii (Spain, Pinilla del Valle), Figs 2627: A. tenue (Georgia, Surami Pass), Fig. 28: A. ganglbaueri (Turkey, Bademli), Figs 2930: A. ganglbaueri (Turkey, Kastamonu): 24, 26, 2829 parameral view, 25, 27, 30lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14240242" httpUri="https://zenodo.org/record/14240242/files/figure.png" pageId="20" pageNumber="471">Fig. 27</figureCitation>
). Habitus as in
<figureCitation id="13315DBD90145A6C2777FD412F8FFD77" box="[600,671,650,672]" captionStart="FIGURES 1923" captionStartId="14.[151,264,1852,1876]" captionTargetBox="[163,1421,246,1814]" captionTargetId="figure-56@14.[151,1436,237,1829]" captionTargetPageId="14" captionText="FIGURES 1923. Habitus of Lathrimaeum (types) and Anthobium: 19L. moczarskii (holotype), 20L. metallicum (holotype), 21L. tenue (lectotype), 22L. laterale (holotype), 23A. ganglbaueri (Turkey, Ulu Dað). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240238" httpUri="https://zenodo.org/record/14240238/files/figure.png" pageId="20" pageNumber="471">Fig. 22</figureCitation>
. Body length:
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. Caucasus...........................
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<emphasis id="B97E9D2A90145A6C2047FD40288CFD48" box="[1384,1436,651,672]" italics="true" pageId="20" pageNumber="471">tenue</emphasis>
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<paragraph id="8BB5413890145A6C25B8FD63288CFD32" blockId="20.[151,1437,151,877]" pageId="20" pageNumber="471">
<keyLead id="30FBE20D90145A6C25B8FD63288CFD32" pageId="20" pageNumber="471">
<emphasis id="B97E9D2A90145A6C25B8FD632D8FFD55" box="[151,159,680,701]" italics="true" pageId="20" pageNumber="471">-</emphasis>
Punctation of pronotum and elytra large and deep. Parameres significantly longer than apex of median lobe, with narrow apical parts (
<figureCitation id="13315DBD90145A6C2431FD0E2C76FD33" box="[286,358,709,731]" captionStart="FIGURES 2430" captionStartId="15.[151,264,1514,1538]" captionTargetBox="[172,1415,200,1482]" captionTargetId="figure-16@15.[164,1423,181,1490]" captionTargetPageId="15" captionText="FIGURES 2430. Aedeagus of Anthobium (Figs 2425: A. moczarskii (Spain, Pinilla del Valle), Figs 2627: A. tenue (Georgia, Surami Pass), Fig. 28: A. ganglbaueri (Turkey, Bademli), Figs 2930: A. ganglbaueri (Turkey, Kastamonu): 24, 26, 2829 parameral view, 25, 27, 30lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14240242" httpUri="https://zenodo.org/record/14240242/files/figure.png" pageId="20" pageNumber="471">Fig. 25</figureCitation>
). Habitus as in
<figureCitation id="13315DBD90145A6C24D5FD0E2F51FD32" box="[506,577,709,731]" captionStart="FIGURES 1923" captionStartId="14.[151,264,1852,1876]" captionTargetBox="[163,1421,246,1814]" captionTargetId="figure-56@14.[151,1436,237,1829]" captionTargetPageId="14" captionText="FIGURES 1923. Habitus of Lathrimaeum (types) and Anthobium: 19L. moczarskii (holotype), 20L. metallicum (holotype), 21L. tenue (lectotype), 22L. laterale (holotype), 23A. ganglbaueri (Turkey, Ulu Dað). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240238" httpUri="https://zenodo.org/record/14240238/files/figure.png" pageId="20" pageNumber="471">Fig. 20</figureCitation>
. Body length:
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.
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,
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.........................
<taxonomicName id="4C0A3ABB90145A6C201BFD0E288CFD32" ID-CoL="5VCFC" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1961" box="[1332,1436,709,730]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="moczarskii">
<emphasis id="B97E9D2A90145A6C201BFD0E288CFD32" box="[1332,1436,709,730]" italics="true" pageId="20" pageNumber="471">moczarskii</emphasis>
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10 Apical angles of pronotum not or slightly protruded anteriad. Aedeagus elongate, with relatively narrow apical part of median lobe (
<figureCitation id="13315DBD90145A6C2437FD342C61FCFD" box="[280,369,767,789]" captionStart="FIGURES 712" captionStartId="6.[151,264,1797,1821]" captionTargetBox="[159,1404,189,1766]" captionTargetId="figure-28@6.[151,1436,181,1774]" captionTargetPageId="6" captionText="FIGURES 712. Aedeagus of Lathrimaeum (types) and Anthobium (Fig. 7: L. gracilicorne (holotype), Fig. 8: L. abantense (holotype), Fig. 9: A. atrocephalum (Algeria, Adekar), Figs 1012: A. melanocephalum (Fig. 10: Georgia, Kherkhvashi, Figs 1112: Karachay-Cherkessia, Malaya Khatipara Mt.): 711parameral view, 12lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14240230" httpUri="https://zenodo.org/record/14240230/files/figure.png" pageId="20" pageNumber="471">Figs. 79</figureCitation>
and
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Figs 2,
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</figureCitation>
in
<bibRefCitation id="EF9B3CC990145A6C2709FD342FD1FCFD" author="Shavrin, A. V." box="[550,705,767,789]" pageId="20" pageNumber="471" pagination="401 - 434" refId="ref21454" refString="Shavrin, A. V. (2020) A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). IV. The atrocephalum and convexior groups, and additional species of the morchella, nigrum and reflexum groups. Zootaxa, 4821 (3), 401 - 434. https: // doi. org / 10.11646 / zootaxa. 4821.3.1" type="journal article" year="2020">Shavrin (2020))</bibRefCitation>
. Habitus as in
<figureCitation id="13315DBD90145A6C2664FD342E8EFCFD" box="[843,926,767,789]" captionStart="FIGURES 16" captionStartId="5.[151,264,1623,1647]" captionTargetBox="[157,1416,185,1591]" captionTargetId="figure-16@5.[151,1435,179,1597]" captionTargetPageId="5" captionText="FIGURES 16. Habitus of Lathrimaeum (types) and Anthobium: 1L. gracilicorne (holotype), 2L. abantense (holotype), 3L. urgelense (holotype), 4L. hamatum (holotype), 5A. melanocephalum (Georgia: Svaneti), 6L. longicorne (lectotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240228" httpUri="https://zenodo.org/record/14240228/files/figure.png" pageId="20" pageNumber="471">Figs 13</figureCitation>
and
<figureCitation id="13315DBD90145A6C26E4FD342912FCFD" box="[971,1026,767,789]" captionStart="FIGURES 16" captionStartId="5.[151,264,1623,1647]" captionTargetBox="[157,1416,185,1591]" captionTargetId="figure-16@5.[151,1435,179,1597]" captionTargetPageId="5" captionText="FIGURES 16. Habitus of Lathrimaeum (types) and Anthobium: 1L. gracilicorne (holotype), 2L. abantense (holotype), 3L. urgelense (holotype), 4L. hamatum (holotype), 5A. melanocephalum (Georgia: Svaneti), 6L. longicorne (lectotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240228" httpUri="https://zenodo.org/record/14240228/files/figure.png" pageId="20" pageNumber="471">
Fig.
<quantity id="4CF2ECDD90145A6C26D9FCCB2912FCFD" box="[1014,1026,768,789]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="20" pageNumber="471" unit="in" value="1.0">1</quantity>
</figureCitation>
in
<bibRefCitation id="EF9B3CC990145A6C210FFD3429A3FCFD" author="Shavrin, A. V." box="[1056,1203,767,789]" pageId="20" pageNumber="471" pagination="401 - 434" refId="ref21454" refString="Shavrin, A. V. (2020) A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). IV. The atrocephalum and convexior groups, and additional species of the morchella, nigrum and reflexum groups. Zootaxa, 4821 (3), 401 - 434. https: // doi. org / 10.11646 / zootaxa. 4821.3.1" type="journal article" year="2020">Shavrin (2020)</bibRefCitation>
. Body length:
<quantity id="4CF2ECDD90145A6C2013FD342C1AFCDA" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.6" metricValueMin="2.4" pageId="20" pageNumber="471" unit="mm" value="3.0" valueMax="3.6" valueMin="2.4">2.403.60 mm</quantity>
. Palaearctic species......................................................................
<taxonomicName id="4C0A3ABB90145A6C2037FCD6288CFCDA" ID-CoL="5VCBN" baseAuthorityName="Gyllenhal" baseAuthorityYear="1827" box="[1304,1436,797,818]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="atrocephalum">
<emphasis id="B97E9D2A90145A6C2037FCD6288CFCDA" box="[1304,1436,797,818]" italics="true" pageId="20" pageNumber="471">atrocephalum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB5413890145A6C25B8FCF0288DFC84" blockId="20.[151,1437,151,877]" pageId="20" pageNumber="471">
<keyLead id="30FBE20D90145A6C25B8FCF0288DFC84" pageId="20" pageNumber="471">
<emphasis id="B97E9D2A90145A6C25B8FCF02D8FFCB8" box="[151,159,827,848]" italics="true" pageId="20" pageNumber="471">-</emphasis>
Apical angles of pronotum distinctly protruded anteriad. Aedeagus short, with slightly broadened apical part of median lobe (
<figureCitation id="13315DBD90145A6C25C6FC9C2C20FC85" box="[233,304,855,877]" captionStart="FIGURES 1317" captionStartId="12.[151,264,1813,1837]" captionTargetBox="[159,1428,195,1780]" captionTargetId="figure-28@12.[151,1436,181,1789]" captionTargetPageId="12" captionText="FIGURES 1317. Aedeagus of Anthobium (Figs 1314: A. longicorne (Algeria, Tikjida), Fig. 15: L. longicorne (Algeria, Babor Mt.), Figs 1617: A. metallicum (Turkey, Isparta): 13, 1516parameral view, 14, 17lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14240232" httpUri="https://zenodo.org/record/14240232/files/figure.png" pageId="20" pageNumber="471">Fig. 17</figureCitation>
). Habitus as in
<figureCitation id="13315DBD90145A6C24EAFC9C2F1CFC85" box="[453,524,855,877]" captionStart="FIGURES 1923" captionStartId="14.[151,264,1852,1876]" captionTargetBox="[163,1421,246,1814]" captionTargetId="figure-56@14.[151,1436,237,1829]" captionTargetPageId="14" captionText="FIGURES 1923. Habitus of Lathrimaeum (types) and Anthobium: 19L. moczarskii (holotype), 20L. metallicum (holotype), 21L. tenue (lectotype), 22L. laterale (holotype), 23A. ganglbaueri (Turkey, Ulu Dað). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240238" httpUri="https://zenodo.org/record/14240238/files/figure.png" pageId="20" pageNumber="471">Fig. 21</figureCitation>
. Body length:
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.
<collectingCountry id="F31D01A890145A6C2600FC9C2E74FC85" box="[815,868,855,877]" name="Syria" pageId="20" pageNumber="471">Syria</collectingCountry>
,
<collectingCountry id="F31D01A890145A6C2641FC9C2ED4FC85" box="[878,964,855,877]" name="Lebanon" pageId="20" pageNumber="471">Lebanon</collectingCountry>
,
<collectingCountry id="F31D01A890145A6C26E2FC9C2901FC85" box="[973,1041,855,877]" name="Turkey" pageId="20" pageNumber="471">Turkey</collectingCountry>
.......................
<taxonomicName id="4C0A3ABB90145A6C201DFC9C288DFC84" ID-CoL="3SJ7C" authorityName="Luze" authorityYear="1905" box="[1330,1437,855,876]" class="Insecta" family="Staphylinidae" genus="Lathrimaeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="metallicum">
<emphasis id="B97E9D2A90145A6C201DFC9C288DFC84" box="[1330,1437,855,876]" italics="true" pageId="20" pageNumber="471">metallicum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>