treatments-xml/data/03/A3/87/03A387C1302A872F1198F9710C45FA32.xml

<document id="8F8ED9161FB0E9FD54FE73D952F97DDB" ID-DOI="10.1093/zoolinnean/zlae023" ID-ISSN="0024-4082" ID-Zenodo-Dep="14802079" ID-ZooBank="BB99D24-3973-4413-B127-BDAA83186FA3" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1738154460006" checkinUser="plazi" docAuthor="Carvalho, Rafael N. &amp; Kury, Adriano B." docDate="2025" docId="03A387C1302A872F1198F9710C45FA32" docLanguage="en" docName="zlae023.pdf" docOrigin="Zoological Journal of the Linnean Society (zlae 023) 203 (1)" docSource="https://doi.org/10.1093/zoolinnean/zlae023" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Iamarinus pontesi Carvalho &amp; Kury 2025, gen. et sp. nov." docType="treatment" docVersion="3" lastPageNumber="49" masterDocId="FF9AFFB93000871F1253FFC8090FFFBA" masterDocTitle="Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species" masterLastPageNumber="65" masterPageNumber="1" pageNumber="43" updateTime="1738675863454" updateUser="ExternalLinkService">
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<mods:title id="C28E42FCC2415C9D18394A1EF0F27E79">Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species</mods:title>
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<mods:namePart id="50C238C3CA6350B5BC1406F303D9B527">Carvalho, Rafael N.</mods:namePart>
<mods:affiliation id="CC036C1F12C4AE1B5B45FC9D7F4D0FFB">Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro (UFRJ), Quinta da Boa Vista s / n, São Cristóvão, 20.940 - 040, Rio de Janeiro, Brazil &amp; Departamento de Zoologia, Instituto de Biologia Roberto de Alcantara Gomes, Universidade do Estado do Rio de Janeiro (UERJ), Rua São Francisco Xavier 524, Maracanã, 20.550 - 900, Rio de Janeiro, Brazil</mods:affiliation>
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<mods:namePart id="597368BFA03D55F5D59B23B05F123B37">Kury, Adriano B.</mods:namePart>
<mods:affiliation id="E23213AF6F1CDD00455ED22ADB8F94B9">Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro (UFRJ), Quinta da Boa Vista s / n, São Cristóvão, 20.940 - 040, Rio de Janeiro, Brazil</mods:affiliation>
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<mods:date id="208084E667EFED8FEADFA95462D291EA">2025</mods:date>
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<paragraph id="8BB536D7302A87351198F9710C3FF969" blockId="42.[825,1474,1721,1856]" box="[971,1328,1721,1747]" pageId="42" pageNumber="43">
<emphasis id="B97EEAC5302A87351198F9710C3FF969" bold="true" box="[971,1328,1721,1747]" pageId="42" pageNumber="43">
<taxonomicName id="4C0A4D54302A87351198F9710D99F969" authority="Carvalho &amp; Kury, 2025" authorityName="Carvalho &amp; Kury" authorityYear="2025" box="[971,1174,1721,1747]" class="Arachnida" family="Gonyleptidae" genus="Iamarinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="pontesi" status="gen. et sp. nov.">
<emphasis id="B97EEAC5302A87351198F9710D99F969" bold="true" box="[971,1174,1721,1747]" italics="true" pageId="42" pageNumber="43">Iamarinus pontesi</emphasis>
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<taxonomicNameLabel id="A24D57BE302A873516C8F9710C3FF969" box="[1179,1328,1721,1747]" pageId="42" pageNumber="43" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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<subSubSection id="C310655C302A8735166EF92B0D8BF8FA" pageId="42" pageNumber="43" type="description">
<emphasis id="B97EEAC5302A8735166EF92B0AE2F89B" italics="true" pageId="42" pageNumber="43">
<paragraph id="8BB536D7302A8735166EF92B0DB1F947" blockId="42.[825,1474,1721,1856]" box="[1085,1214,1763,1789]" pageId="42" pageNumber="43">
(
<figureCitation id="13312A52302A87351614F92C0DBBF947" box="[1095,1204,1763,1789]" captionStart-0="Figure 24" captionStart-1="Figure 25" captionStart-2="15" captionStart-3="Figure 28" captionStart-4="Figure 28" captionStartId-0="44.[130,195,1534,1558]" captionStartId-1="45.[113,178,872,896]" captionStartId-2="5.[113,139,194,218]" captionStartId-3="47.[114,179,1699,1723]" captionStartId-4="47.[114,179,1699,1723]" captionTargetBox-0="[129,1473,144,1496]" captionTargetBox-1="[113,1457,144,834]" captionTargetBox-2="[130,1474,146,1751]" captionTargetBox-3="[117,1455,887,1657]" captionTargetBox-4="[117,1455,887,1657]" captionTargetId-0="figure-270@44.[129,1473,144,1496]" captionTargetId-1="figure-706@45.[113,1457,144,834]" captionTargetId-2="figure-11@46.[127,1477,143,1754]" captionTargetId-3="figure-128@47.[114,1458,884,1660]" captionTargetId-4="figure-128@47.[114,1458,884,1660]" captionTargetPageId-0="44" captionTargetPageId-1="45" captionTargetPageId-2="46" captionTargetPageId-3="47" captionTargetPageId-4="47" captionText-0="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." captionText-1="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." captionText-2="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" captionText-3="Figure 28.Iamarinus pontesi, ♀ paratype (MNRJ 60562).A, ocularium, frontal view.B, habitus, dorsal view.C, right trochanter and femur IV, dorsal view. Scale bars: 1 mm (A), 3 mm (B–C)." captionText-4="Figure 28.Iamarinus pontesi, ♀ paratype (MNRJ 60562).A, ocularium, frontal view.B, habitus, dorsal view.C, right trochanter and femur IV, dorsal view. Scale bars: 1 mm (A), 3 mm (B–C)." figureDoi-0="http://doi.org/10.5281/zenodo.14802132" figureDoi-1="http://doi.org/10.5281/zenodo.14802134" figureDoi-3="http://doi.org/10.5281/zenodo.14802142" httpUri-0="https://zenodo.org/record/14802132/files/figure.png" httpUri-1="https://zenodo.org/record/14802134/files/figure.png" httpUri-3="https://zenodo.org/record/14802142/files/figure.png" pageId="42" pageNumber="43">Figs 24–28</figureCitation>
)
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ZooBank 
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</emphasis>
</subSubSection>
<subSubSection id="C310655C302A87341618F8AE0B82FA87" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="etymology">
<paragraph id="8BB536D7302A87351618F8AE0DBEF83A" blockId="42.[825,1474,1894,1988]" box="[1099,1201,1894,1920]" pageId="42" pageNumber="43">
<emphasis id="B97EEAC5302A87351618F8AE0DBEF83A" box="[1099,1201,1894,1920]" italics="true" pageId="42" pageNumber="43">Etymology</emphasis>
</paragraph>
<paragraph id="8BB536D7302A8734116AF8450B82FA87" blockId="42.[825,1474,1894,1988]" lastBlockId="43.[113,763,1286,1341]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<taxonomicName id="4C0A4D54302A8735116AF8450AEFF81E" authorityName="Carvalho &amp; Kury" authorityYear="2025" box="[825,992,1933,1956]" class="Arachnida" family="Gonyleptidae" genus="Iamarinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="pontesi">
<emphasis id="B97EEAC5302A8735116AF8450AEFF81E" box="[825,992,1933,1956]" italics="true" pageId="42" pageNumber="43">Iamarinus pontesi</emphasis>
</taxonomicName>
is designated in acknowledgment of the substantialcontributionsbyconservationistJoãoPontestothepreservation and conservation of biodiversity in the Reserva Natural Guaricica, situated in 
<taxonomicName id="4C0A4D54302B8734128FFAED0834FA87" authority="Arapoti, Guaraquecaba" authorityName="Arapoti, Guaraquecaba" box="[220,315,1317,1341]" class="Insecta" family="Pseudococcidae" genus="Antonina" kingdom="Animalia" order="Hemiptera" pageId="43" pageNumber="25" phylum="Arthropoda" rank="genus">Antonina</taxonomicName>
, 
<collectingRegion id="49CEF835302B87341310FAED0887FA87" box="[323,392,1317,1341]" country="Brazil" name="Parana" pageId="43" pageNumber="44">Paraná</collectingRegion>
, 
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.Gender masculine.
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<caption id="DF75665F302B87341220FC930D9CFC35" ID-DOI="http://doi.org/10.5281/zenodo.14802130" ID-Zenodo-Dep="14802130" httpUri="https://zenodo.org/record/14802130/files/figure.png" pageId="43" pageNumber="44" startId="43.[115,180,859,883]" targetBox="[116,1453,147,816]" targetPageId="43" targetType="figure">
<paragraph id="8BB536D7302B87341220FC930D9CFC35" blockId="43.[113,1410,859,911]" pageId="43" pageNumber="44">
<emphasis id="B97EEAC5302B87341220FC9309D8FCC9" bold="true" box="[115,215,859,883]" pageId="43" pageNumber="44">Figure 23.</emphasis>
<taxonomicName id="4C0A4D54302B87341289FC940862FCC9" baseAuthorityName="Kury, Pinto-da-Rocha &amp; Carvalho" baseAuthorityYear="2018" box="[218,365,859,883]" class="Arachnida" family="Gonyleptidae" genus="Iamarinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="fenax">
<emphasis id="B97EEAC5302B87341289FC940862FCC9" box="[218,365,859,883]" italics="true" pageId="43" pageNumber="44">Iamarinus fenax</emphasis>
</taxonomicName>
, ♀ (MHNCI 6305). A, armature of scutal area III, posterior view. B, ocularium, frontal view. C, habitus, dorsal view. D, Right trochanter and femur IV, dorsal view. E, same, ventral view. Scale bars: 1 mm (A–B), 3 mm (C–E).
</paragraph>
</caption>
<caption id="DF75665F302B87341222FC0C0BF8FC66" ID-Table-UUID="DF75665F302B87341222FC0C0BF8FC66" box="[113,759,964,989]" httpUri="http://table.plazi.org/id/DF75665F302B87341222FC0C0BF8FC66" pageId="43" pageNumber="44" startId="43.[113,168,964,988]" targetBox="[113,762,1010,1218]" targetIsTable="true" targetPageId="43">
<paragraph id="8BB536D7302B87341222FC0C0BF8FC66" blockId="43.[113,759,964,989]" box="[113,759,964,989]" pageId="43" pageNumber="44">
<emphasis id="B97EEAC5302B87341222FC0C09C3FC66" bold="true" box="[113,204,964,988]" pageId="43" pageNumber="44">Table 13.</emphasis>
Leg measurements of 
<emphasis id="B97EEAC5302B873413CCFC0D0B3DFC66" box="[415,562,964,988]" italics="true" pageId="43" pageNumber="44">Iamarinus fenax</emphasis>
, ♀ (MHNCI 6305)
</paragraph>
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<paragraph id="8BB536D7302B87341283FC3B0BF5FB78" pageId="43" pageNumber="44">
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<th id="76EB5DE9302B78E01283FC3A09F6FBB1" box="[208,249,1010,1035]" gridcol="1" gridrow="0" pageId="43" pageNumber="44">Tr</th>
<th id="76EB5DE9302B78E01344FC3A084FFBB1" box="[279,320,1010,1035]" gridcol="2" gridrow="0" pageId="43" pageNumber="44">Fe</th>
<th id="76EB5DE9302B78E0130DFC3A0888FBB1" box="[350,391,1010,1035]" gridcol="3" gridrow="0" pageId="43" pageNumber="44">Pa</th>
<th id="76EB5DE9302B78E013F6FC3A08C1FBB1" box="[421,462,1010,1035]" gridcol="4" gridrow="0" pageId="43" pageNumber="44">Ti</th>
<th id="76EB5DE9302B78E013BFFC3A0B18FBB1" box="[492,535,1010,1035]" gridcol="5" gridrow="0" pageId="43" pageNumber="44">Mt</th>
<th id="76EB5DE9302B78E01066FC3A0B51FBB1" box="[565,606,1010,1035]" gridcol="6" gridrow="0" pageId="43" pageNumber="44">Ta</th>
<th id="76EB5DE9302B78E0102FFC3A0BA8FBB1" box="[636,679,1010,1035]" gridcol="7" gridrow="0" pageId="43" pageNumber="44">Cl</th>
<th id="76EB5DE9302B78E01096FC3A0BF5FBB1" box="[709,762,1010,1035]" gridcol="8" gridrow="0" pageId="43" pageNumber="44">
<emphasis id="B97EEAC5302B87341096FC3B0BF6FBB1" bold="true" box="[709,761,1011,1035]" pageId="43" pageNumber="44">Total</emphasis>
</th>
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<tr id="353A3495302B78E01222FBEA0BF5FB81" box="[113,762,1058,1083]" gridrow="1" pageId="43" pageNumber="44">
<th id="76EB5DE9302B78E01222FBEA09BCFB81" box="[113,179,1058,1083]" gridcol="0" gridrow="1" pageId="43" pageNumber="44">Pp</th>
<td id="76EB5DE9302B78E01283FBEA09F6FB81" box="[208,249,1058,1083]" gridcol="1" gridrow="1" pageId="43" pageNumber="44">0.48</td>
<td id="76EB5DE9302B78E01344FBEA084FFB81" box="[279,320,1058,1083]" gridcol="2" gridrow="1" pageId="43" pageNumber="44">1.44</td>
<td id="76EB5DE9302B78E0130DFBEA0888FB81" box="[350,391,1058,1083]" gridcol="3" gridrow="1" pageId="43" pageNumber="44">0.66</td>
<td id="76EB5DE9302B78E013F6FBEA08C1FB81" box="[421,462,1058,1083]" gridcol="4" gridrow="1" pageId="43" pageNumber="44">1.18</td>
<td id="76EB5DE9302B78E013BFFBEA0B18FB81" box="[492,535,1058,1083]" gridcol="5" gridrow="1" pageId="43" pageNumber="44">−</td>
<td id="76EB5DE9302B78E01066FBEA0B51FB81" box="[565,606,1058,1083]" gridcol="6" gridrow="1" pageId="43" pageNumber="44">0.96</td>
<td id="76EB5DE9302B78E0102FFBEA0BA8FB81" box="[636,679,1058,1083]" gridcol="7" gridrow="1" pageId="43" pageNumber="44">0.93</td>
<td id="76EB5DE9302B78E01096FBEA0BF5FB81" box="[709,762,1058,1083]" gridcol="8" gridrow="1" pageId="43" pageNumber="44">5.66</td>
</tr>
<tr id="353A3495302B78E01222FB8C0BF5FBE7" box="[113,762,1092,1117]" gridrow="2" pageId="43" pageNumber="44">
<th id="76EB5DE9302B78E01222FB8C09BCFBE7" box="[113,179,1092,1117]" gridcol="0" gridrow="2" pageId="43" pageNumber="44">Leg I</th>
<td id="76EB5DE9302B78E01283FB8C09F6FBE7" box="[208,249,1092,1117]" gridcol="1" gridrow="2" pageId="43" pageNumber="44">0.64</td>
<td id="76EB5DE9302B78E01344FB8C084FFBE7" box="[279,320,1092,1117]" gridcol="2" gridrow="2" pageId="43" pageNumber="44">2.25</td>
<td id="76EB5DE9302B78E0130DFB8C0888FBE7" box="[350,391,1092,1117]" gridcol="3" gridrow="2" pageId="43" pageNumber="44">0.80</td>
<td id="76EB5DE9302B78E013F6FB8C08C1FBE7" box="[421,462,1092,1117]" gridcol="4" gridrow="2" pageId="43" pageNumber="44">1.66</td>
<td id="76EB5DE9302B78E013BFFB8C0B18FBE7" box="[492,535,1092,1117]" gridcol="5" gridrow="2" pageId="43" pageNumber="44">2.57</td>
<td id="76EB5DE9302B78E01066FB8C0B51FBE7" box="[565,606,1092,1117]" gridcol="6" gridrow="2" pageId="43" pageNumber="44">1.46</td>
<td id="76EB5DE9302B78E0102FFB8C0BA8FBE7" box="[636,679,1092,1117]" gridcol="7" gridrow="2" pageId="43" pageNumber="44">–</td>
<td id="76EB5DE9302B78E01096FB8C0BF5FBE7" box="[709,762,1092,1117]" gridcol="8" gridrow="2" pageId="43" pageNumber="44">9.39</td>
</tr>
<tr id="353A3495302B78E01222FBAE0BF5FBC5" box="[113,762,1126,1151]" gridrow="3" pageId="43" pageNumber="44">
<th id="76EB5DE9302B78E01222FBAE09BCFBC5" box="[113,179,1126,1151]" gridcol="0" gridrow="3" pageId="43" pageNumber="44">Leg II</th>
<td id="76EB5DE9302B78E01283FBAE09F6FBC5" box="[208,249,1126,1151]" gridcol="1" gridrow="3" pageId="43" pageNumber="44">0.68</td>
<td id="76EB5DE9302B78E01344FBAE084FFBC5" box="[279,320,1126,1151]" gridcol="2" gridrow="3" pageId="43" pageNumber="44">4.11</td>
<td id="76EB5DE9302B78E0130DFBAE0888FBC5" box="[350,391,1126,1151]" gridcol="3" gridrow="3" pageId="43" pageNumber="44">1.21</td>
<td id="76EB5DE9302B78E013F6FBAE08C1FBC5" box="[421,462,1126,1151]" gridcol="4" gridrow="3" pageId="43" pageNumber="44">3.00</td>
<td id="76EB5DE9302B78E013BFFBAE0B18FBC5" box="[492,535,1126,1151]" gridcol="5" gridrow="3" pageId="43" pageNumber="44">4.48</td>
<td id="76EB5DE9302B78E01066FBAE0B51FBC5" box="[565,606,1126,1151]" gridcol="6" gridrow="3" pageId="43" pageNumber="44">2.69</td>
<td id="76EB5DE9302B78E0102FFBAE0BA8FBC5" box="[636,679,1126,1151]" gridcol="7" gridrow="3" pageId="43" pageNumber="44">−</td>
<td id="76EB5DE9302B78E01096FBAE0BF5FBC5" box="[709,762,1126,1151]" gridcol="8" gridrow="3" pageId="43" pageNumber="44">16.18</td>
</tr>
<tr id="353A3495302B78E01222FB400BF5FB1A" box="[113,762,1160,1184]" gridrow="4" pageId="43" pageNumber="44">
<th id="76EB5DE9302B78E01222FB4009BCFB1A" box="[113,179,1160,1184]" gridcol="0" gridrow="4" pageId="43" pageNumber="44">Leg III</th>
<td id="76EB5DE9302B78E01283FB4009F6FB1A" box="[208,249,1160,1184]" gridcol="1" gridrow="4" pageId="43" pageNumber="44">0.78</td>
<td id="76EB5DE9302B78E01344FB40084FFB1A" box="[279,320,1160,1184]" gridcol="2" gridrow="4" pageId="43" pageNumber="44">3.28</td>
<td id="76EB5DE9302B78E0130DFB400888FB1A" box="[350,391,1160,1184]" gridcol="3" gridrow="4" pageId="43" pageNumber="44">1.13</td>
<td id="76EB5DE9302B78E013F6FB4008C1FB1A" box="[421,462,1160,1184]" gridcol="4" gridrow="4" pageId="43" pageNumber="44">2.27</td>
<td id="76EB5DE9302B78E013BFFB400B18FB1A" box="[492,535,1160,1184]" gridcol="5" gridrow="4" pageId="43" pageNumber="44">3.55</td>
<td id="76EB5DE9302B78E01066FB400B51FB1A" box="[565,606,1160,1184]" gridcol="6" gridrow="4" pageId="43" pageNumber="44">1.64</td>
<td id="76EB5DE9302B78E0102FFB400BA8FB1A" box="[636,679,1160,1184]" gridcol="7" gridrow="4" pageId="43" pageNumber="44">−</td>
<td id="76EB5DE9302B78E01096FB400BF5FB1A" box="[709,762,1160,1184]" gridcol="8" gridrow="4" pageId="43" pageNumber="44">12.66</td>
</tr>
<tr id="353A3495302B78E01222FB620BF5FB78" box="[113,762,1194,1218]" gridrow="5" pageId="43" pageNumber="44">
<th id="76EB5DE9302B78E01222FB6209BCFB78" box="[113,179,1194,1218]" gridcol="0" gridrow="5" pageId="43" pageNumber="44">Leg IV</th>
<td id="76EB5DE9302B78E01283FB6209F6FB78" box="[208,249,1194,1218]" gridcol="1" gridrow="5" pageId="43" pageNumber="44">0.98</td>
<td id="76EB5DE9302B78E01344FB62084FFB78" box="[279,320,1194,1218]" gridcol="2" gridrow="5" pageId="43" pageNumber="44">4.57</td>
<td id="76EB5DE9302B78E0130DFB620888FB78" box="[350,391,1194,1218]" gridcol="3" gridrow="5" pageId="43" pageNumber="44">1.21</td>
<td id="76EB5DE9302B78E013F6FB6208C1FB78" box="[421,462,1194,1218]" gridcol="4" gridrow="5" pageId="43" pageNumber="44">2.98</td>
<td id="76EB5DE9302B78E013BFFB620B18FB78" box="[492,535,1194,1218]" gridcol="5" gridrow="5" pageId="43" pageNumber="44">4.53</td>
<td id="76EB5DE9302B78E01066FB620B51FB78" box="[565,606,1194,1218]" gridcol="6" gridrow="5" pageId="43" pageNumber="44">1.80</td>
<td id="76EB5DE9302B78E0102FFB620BA8FB78" box="[636,679,1194,1218]" gridcol="7" gridrow="5" pageId="43" pageNumber="44">−</td>
<td id="76EB5DE9302B78E01096FB620BF5FB78" box="[709,762,1194,1218]" gridcol="8" gridrow="5" pageId="43" pageNumber="44">16.07</td>
</tr>
</table>
</paragraph>
<subSubSection id="C310655C302B873413D4FAAB08E3F8E2" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="8BB536D7302B873413D4FAAB08EAFAC6" blockId="43.[110,765,1378,1880]" box="[391,485,1378,1404]" pageId="43" pageNumber="44">
<emphasis id="B97EEAC5302B873413D4FAAB08EAFAC6" box="[391,485,1378,1404]" italics="true" pageId="43" pageNumber="44">
<typeStatus id="54B18875302B873413D4FAAB08BAFAC6" box="[391,437,1379,1404]" pageId="43" pageNumber="44">Type</typeStatus>
data
</emphasis>
</paragraph>
<paragraph id="8BB536D7302B87341222FA4208E3F8E2" blockId="43.[110,765,1378,1880]" pageId="43" pageNumber="44">
<emphasis id="B97EEAC5302B87341222FA42082DFA1B" box="[113,290,1418,1441]" italics="true" pageId="43" pageNumber="44">
<taxonomicName id="4C0A4D54302B87341222FA420812FA1B" authorityName="Carvalho &amp; Kury" authorityYear="2025" box="[113,285,1418,1441]" class="Arachnida" family="Gonyleptidae" genus="Iamarinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="pontesi">Iamarinus pontesi</taxonomicName>
:
</emphasis>
<materialsCitation id="3B623C8A302B87341379FA440B4CFA1B" box="[298,579,1417,1441]" collectionCode="MNRJ" pageId="43" pageNumber="44" specimenCode="MNRJ 2789" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="9D0CFD5E302B87341379FA44084BFA1A" box="[298,324,1420,1440]" count="1" pageId="43" pageNumber="44" type="male">♂</specimenCount>
<typeStatus id="54B18875302B8734131FFA4108A9FA1B" box="[332,422,1417,1441]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
(
<specimenCode id="DBAC9EAC302B873413E6FA410B33FA1B" box="[437,572,1417,1441]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="43" pageNumber="44">MNRJ 2789</specimenCode>
)
</materialsCitation>
, 
<specimenCount id="9D0CFD5E302B8734101CFA410B71FA1A" box="[591,638,1417,1441]" count="3" pageId="43" pageNumber="44" type="male">3 ♂</specimenCount>
<typeStatus id="54B18875302B873410D5FA420BEAFA18" box="[646,741,1418,1442]" pageId="43" pageNumber="44" type="paratype">paratypes</typeStatus>
<materialsCitation id="3B623C8A302B873410BEFA410BFBFA45" collectingDate="2019-11-09" collectionCode="R" collectorName="Azara, L. N. &amp; Carvalho, R. N. &amp; Kury, A. B." country="Brazil" elevation="178" location="Parana" pageId="43" pageNumber="44" specimenCode="MNRJ 60562" specimenCount="3" specimenCount-female="3" stateProvince="Parana" typeStatus="paratype">
<specimenCount id="9D0CFD5E302B873410BEFA410984FA7A" count="3" pageId="43" pageNumber="44" type="female">3 ♀</specimenCount>
<typeStatus id="54B18875302B873412C3FA6109E0FA7B" box="[144,239,1449,1473]" pageId="43" pageNumber="44" type="paratype">paratypes</typeStatus>
1 juv (
<specimenCode id="DBAC9EAC302B8734137CFA6008CFFA7A" box="[303,448,1448,1472]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="43" pageNumber="44">MNRJ 60562</specimenCode>
), from 
<collectingCountry id="F31D7647302B87341056FA610B51FA7A" box="[517,606,1449,1472]" name="Brazil" pageId="43" pageNumber="44">BRAZIL</collectingCountry>
, state of 
<collectingRegion id="49CEF835302B873410E1FA600BF7FA7A" box="[690,760,1448,1472]" country="Brazil" name="Parana" pageId="43" pageNumber="44">Paraná</collectingRegion>
, 
<taxonomicName id="4C0A4D54302B87341222FA0009DFFA5A" authority="Arapoti, Guaraquecaba" authorityName="Arapoti, Guaraquecaba" box="[113,208,1480,1504]" class="Insecta" family="Pseudococcidae" genus="Antonina" kingdom="Animalia" order="Hemiptera" pageId="43" pageNumber="25" phylum="Arthropoda" rank="genus">Antonina</taxonomicName>
,ReservaBiológicaBomJesus,-25.296 235°,-48.613 820°, 
<quantity id="4CF29B32302B8734123DFA2F09BEFA45" box="[110,177,1511,1535]" metricMagnitude="2" metricUnit="m" metricValue="1.78" pageId="43" pageNumber="44" unit="m" value="178.0">
<elevation id="0027D1E4302B8734123DFA2F09BEFA45" box="[110,177,1511,1535]" metricMagnitude="2" metricUnit="m" metricValue="1.78" pageId="43" pageNumber="44" unit="m" value="178.0">178 m</elevation>
</quantity>
, 
<date id="FFB41017302B873412EFFA2F0829FA45" box="[188,294,1511,1535]" pageId="43" pageNumber="44" value="2019-11-09">
<collectingDate id="EFF0E9FF302B873412EFFA2F0829FA45" box="[188,294,1511,1535]" pageId="43" pageNumber="44" value="2019-11-09">09.xi.2019</collectingDate>
</date>
, 
<collectorName id="26FF5301302B87341363FA2008ACFA45" box="[304,419,1512,1539]" pageId="43" pageNumber="44">Ázara, L.N.</collectorName>
, 
<collectorName id="26FF5301302B873413FEFA2F0B4BFA45" box="[429,580,1511,1535]" pageId="43" pageNumber="44">
Carvalho, 
<taxonomicNameLabel id="A24D57BE302B87341047FA200B4BFA45" box="[532,580,1512,1535]" pageId="43" pageNumber="44" rank="subSpecies">R.N.</taxonomicNameLabel>
</collectorName>
&amp; 
<collectorName id="26FF5301302B87341037FA200BC2FA45" box="[612,717,1511,1536]" pageId="43" pageNumber="44">Kury, A.B.</collectorName>
<emphasis id="B97EEAC5302B87341084FA2F0BFBFA45" box="[727,756,1511,1535]" italics="true" pageId="43" pageNumber="44">leg.</emphasis>
</materialsCitation>
; 
<specimenCount id="9D0CFD5E302B87341222F9CE0991F9A4" box="[113,158,1542,1566]" count="3" pageId="43" pageNumber="44" type="male">3 ♂</specimenCount>
<typeStatus id="54B18875302B873412F6F9CF080BF9A5" box="[165,260,1543,1567]" pageId="43" pageNumber="44" type="paratype">paratypes</typeStatus>
<materialsCitation id="3B623C8A302B87341358F9C209A6F9C6" collectingDate="2021-11-16" collectionCode="R" collectorName="Carvalho, R. N." country="Brazil" elevation="125" location="Reserva Natural Guaricica" municipality="Antonina" pageId="43" pageNumber="44" specimenCode="MNRJ 378" specimenCount="1" specimenCount-female="1" stateProvince="Parana" typeStatus="paratype">
<specimenCount id="9D0CFD5E302B87341358F9C2082AF9A4" box="[267,293,1546,1566]" count="1" pageId="43" pageNumber="44" type="female">♀</specimenCount>
<typeStatus id="54B18875302B87341379F9CF088FF9A5" box="[298,384,1543,1567]" pageId="43" pageNumber="44" type="paratype">paratype</typeStatus>
(
<specimenCode id="DBAC9EAC302B873413DDF9CE0B09F9A4" box="[398,518,1542,1566]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="43" pageNumber="44">MNRJ 378</specimenCode>
), from 
<collectingCountry id="F31D7647302B8734101CF9CF0BA7F9A4" box="[591,680,1543,1566]" name="Brazil" pageId="43" pageNumber="44">BRAZIL</collectingCountry>
, state of 
<collectingRegion id="49CEF835302B87341220F9EE09B7F984" box="[115,184,1574,1598]" country="Brazil" name="Parana" pageId="43" pageNumber="44">Paraná</collectingRegion>
, 
<taxonomicName id="4C0A4D54302B87341291F9EE0873F987" authority=", Reserva" authorityName="Reserva" box="[194,380,1574,1598]" class="Insecta" family="Pseudococcidae" genus="Antonina" kingdom="Animalia" order="Hemiptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<collectingMunicipality id="6BD1ACAD302B87341291F9EE082EF984" box="[194,289,1574,1598]" pageId="43" pageNumber="44">Antonina</collectingMunicipality>
, 
<location id="8ED5600C302B8734137EF9EE0B38F984" LSID="urn:lsid:plazi:treatment:03A387C1302A872F1198F9710C45FA32:8ED5600C302B8734137EF9EE0B38F984" box="[301,567,1574,1598]" country="Brazil" municipality="Antonina" name="Reserva Natural Guaricica" pageId="43" pageNumber="44" stateProvince="Parana">Reserva Natural Guaricica</location>
</taxonomicName>
, 
<location id="8ED5600C302B87341012F9EE0BF6F987" LSID="urn:lsid:plazi:treatment:03A387C1302A872F1198F9710C45FA32:8ED5600C302B87341012F9EE0BF6F987" box="[577,761,1574,1598]" country="Brazil" municipality="Antonina" name="Trilha dos Fornos" pageId="43" pageNumber="44" stateProvince="Parana">Trilha dos Fornos</location>
, -25,30 252°, -48,66 005°, 
<quantity id="4CF29B32302B87341320F98D08B6F9E7" box="[371,441,1605,1629]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="43" pageNumber="44" unit="m" value="125.0">
<elevation id="0027D1E4302B87341320F98D08B6F9E7" box="[371,441,1605,1629]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="43" pageNumber="44" unit="m" value="125.0">125 m</elevation>
</quantity>
, 
<date id="FFB41017302B87341395F98D0B3FF9E7" box="[454,560,1605,1629]" pageId="43" pageNumber="44" value="2021-11-16">
<collectingDate id="EFF0E9FF302B87341395F98D0B3FF9E7" box="[454,560,1605,1629]" pageId="43" pageNumber="44" value="2021-11-16">16.xi.2021</collectingDate>
</date>
, 
<collectorName id="26FF5301302B8734106EF98D0BEEF9E7" box="[573,737,1605,1629]" pageId="43" pageNumber="44">
Carvalho, 
<taxonomicNameLabel id="A24D57BE302B873410F4F98E0BEEF9E7" box="[679,737,1606,1629]" pageId="43" pageNumber="44" rank="subSpecies">R. N.</taxonomicNameLabel>
</collectorName>
<emphasis id="B97EEAC5302B873410B9F98E09A6F9C6" italics="true" pageId="43" pageNumber="44">et al. leg</emphasis>
</materialsCitation>
<emphasis id="B97EEAC5302B873412F9F9AC09A1F9C6" box="[170,174,1636,1660]" italics="true" pageId="43" pageNumber="44">.</emphasis>
; 
<specimenCount id="9D0CFD5E302B873412E8F9A009DAF9C6" box="[187,213,1640,1660]" count="1" pageId="43" pageNumber="44" type="male">♂</specimenCount>
<typeStatus id="54B18875302B8734128FF9AD083DF9C7" box="[220,306,1637,1661]" pageId="43" pageNumber="44" type="paratype">paratype</typeStatus>
<materialsCitation id="3B623C8A302B8734136FF9A008B2F960" collectingDate="2019-11-07" collectionCode="R" collectorName="Azara, L. N. &amp; Carvalho, R. N. &amp; Kury, A. B." country="Brazil" county="Natural Guaricica" elevation="112" location="Reserva Natural Guaricica" municipality="Antonina" pageId="43" pageNumber="44" specimenCode="MNRJ 60561" specimenCount="1" specimenCount-female="1" stateProvince="Parana" typeStatus="paratype">
<specimenCount id="9D0CFD5E302B8734136FF9A00859F9C6" box="[316,342,1640,1660]" count="1" pageId="43" pageNumber="44" type="female">♀</specimenCount>
<typeStatus id="54B18875302B87341308F9AD08BEF9C7" box="[347,433,1637,1661]" pageId="43" pageNumber="44" type="paratype">paratype</typeStatus>
(
<specimenCode id="DBAC9EAC302B873413ECF9AC0B5DF9C6" box="[447,594,1636,1660]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="43" pageNumber="44">MNRJ 60561</specimenCode>
), from 
<collectingCountry id="F31D7647302B873410CEF9AD0BF8F9C6" box="[669,759,1637,1660]" name="Brazil" pageId="43" pageNumber="44">BRAZIL</collectingCountry>
, state of 
<collectingRegion id="49CEF835302B87341294F94C0803F926" box="[199,268,1668,1692]" country="Brazil" name="Parana" pageId="43" pageNumber="44">Paraná</collectingRegion>
, 
<taxonomicName id="4C0A4D54302B8734134AF94C08DBF921" authority=", Reserva" authorityName="Reserva" box="[281,468,1668,1692]" class="Insecta" family="Pseudococcidae" genus="Antonina" kingdom="Animalia" order="Hemiptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<collectingMunicipality id="6BD1ACAD302B8734134AF94C0877F926" box="[281,376,1668,1692]" pageId="43" pageNumber="44">Antonina</collectingMunicipality>
, 
<location id="8ED5600C302B873413D6F94C0B9CF926" LSID="urn:lsid:plazi:treatment:03A387C1302A872F1198F9710C45FA32:8ED5600C302B873413D6F94C0B9CF926" box="[389,659,1668,1692]" country="Brazil" county="Natural Guaricica" municipality="Antonina" name="Reserva Natural Guaricica" pageId="43" pageNumber="44" stateProvince="Parana">Reserva Natural Guaricica</location>
</taxonomicName>
, 
<location id="8ED5600C302B873410CCF94C09A9F901" LSID="urn:lsid:plazi:treatment:03A387C1302A872F1198F9710C45FA32:8ED5600C302B873410CCF94C09A9F901" country="Brazil" county="Natural Guaricica" municipality="Antonina" name="Trilha da Rede" pageId="43" pageNumber="44" stateProvince="Parana">Trilha da Rede</location>
, -25.302 599°, -48.672 713°, 
<quantity id="4CF29B32302B87341397F96B0B08F901" box="[452,519,1699,1723]" metricMagnitude="2" metricUnit="m" metricValue="1.1199999999999999" pageId="43" pageNumber="44" unit="m" value="112.0">
<elevation id="0027D1E4302B87341397F96B0B08F901" box="[452,519,1699,1723]" metricMagnitude="2" metricUnit="m" metricValue="1.1199999999999999" pageId="43" pageNumber="44" unit="m" value="112.0">112 m</elevation>
</quantity>
, 
<date id="FFB41017302B87341043F96B0B74F901" box="[528,635,1699,1723]" pageId="43" pageNumber="44" value="2019-11-07">
<collectingDate id="EFF0E9FF302B87341043F96B0B74F901" box="[528,635,1699,1723]" pageId="43" pageNumber="44" value="2019-11-07">07.xi.2019</collectingDate>
</date>
, 
<collectorName id="26FF5301302B873410D7F96C0BF8F901" box="[644,759,1700,1727]" pageId="43" pageNumber="44">Ázara, L.N.</collectorName>
, 
<collectorName id="26FF5301302B87341220F90A0804F960" box="[115,267,1730,1754]" pageId="43" pageNumber="44">
Carvalho, 
<taxonomicNameLabel id="A24D57BE302B87341288F90B0804F960" box="[219,267,1731,1754]" pageId="43" pageNumber="44" rank="subSpecies">R.N.</taxonomicNameLabel>
</collectorName>
&amp; 
<collectorName id="26FF5301302B8734137DF90B0897F961" box="[302,408,1731,1755]" pageId="43" pageNumber="44">Kury, A.B.</collectorName>
<emphasis id="B97EEAC5302B873413CDF90A08B2F960" box="[414,445,1730,1754]" italics="true" pageId="43" pageNumber="44">leg.</emphasis>
</materialsCitation>
; 
<materialsCitation id="3B623C8A302B87341398F90E08D5F8A3" collectingDate="1988-03-04" collectionCode="MHNCI" collectorName="Bornschein &amp; Motta" country="Brazil" location="Casa da SEMMA" municipality="Guaraquecaba" pageId="43" pageNumber="44" specimenCode="MHNCI 6429" specimenCount="1" specimenCount-male="1" stateProvince="Parana" typeStatus="paratype">
<specimenCount id="9D0CFD5E302B87341398F90E08EAF960" box="[459,485,1734,1754]" count="1" pageId="43" pageNumber="44" type="male">♂</specimenCount>
<typeStatus id="54B18875302B873413BEF90B0B4CF961" box="[493,579,1731,1755]" pageId="43" pageNumber="44" type="paratype">paratype</typeStatus>
(
<specimenCode id="DBAC9EAC302B87341000F90A0BFFF960" box="[595,752,1730,1754]" collectionCode="MHNCI" country="0" httpUri="http://grbio.org/cool/ej4f-8m62" name="Museu de Historia Natural Capao de Imbuia (Brazil)" pageId="43" pageNumber="44">MHNCI 6429</specimenCode>
), from 
<collectingCountry id="F31D7647302B873412FBF92A080EF943" box="[168,257,1762,1785]" name="Brazil" pageId="43" pageNumber="44">BRAZIL</collectingCountry>
, state of 
<collectingRegion id="49CEF835302B87341308F92A08AFF940" box="[347,416,1762,1786]" country="Brazil" name="Parana" pageId="43" pageNumber="44">Paraná</collectingRegion>
, 
<collectingMunicipality id="6BD1ACAD302B873413F9F92A0B34F940" box="[426,571,1762,1786]" pageId="43" pageNumber="44">Guaraqueçaba</collectingMunicipality>
, 
<location id="8ED5600C302B87341017F92A0BF8F940" LSID="urn:lsid:plazi:treatment:03A387C1302A872F1198F9710C45FA32:8ED5600C302B87341017F92A0BF8F940" box="[580,759,1762,1786]" country="Brazil" municipality="Guaraquecaba" name="Casa da SEMMA" pageId="43" pageNumber="44" stateProvince="Parana">Casa da SEMMA</location>
, 
<date id="FFB41017302B87341220F8C909D2F8A3" box="[115,221,1793,1817]" pageId="43" pageNumber="44" value="1988-03-04">
<collectingDate id="EFF0E9FF302B87341220F8C909D2F8A3" box="[115,221,1793,1817]" pageId="43" pageNumber="44" value="1988-03-04">04.iii.1988</collectingDate>
</date>
, 
<collectorName id="26FF5301302B873412B4F8C90856F8A3" box="[231,345,1793,1817]" pageId="43" pageNumber="44">Bornschein</collectorName>
&amp; 
<collectorName id="26FF5301302B8734132AF8CA08B9F8A3" box="[377,438,1794,1817]" pageId="43" pageNumber="44">Motta</collectorName>
<emphasis id="B97EEAC5302B873413EFF8C908D5F8A3" box="[444,474,1793,1817]" italics="true" pageId="43" pageNumber="44">leg.</emphasis>
</materialsCitation>
; 
<materialsCitation id="3B623C8A302B873413B5F8C908E7F8E2" collectingDate="1991-01-13" collectionCode="R" collectorName="Pinto-da-Rocha, R. &amp; Bernils, R. S." country="Brazil" location="Banhado" municipality="Piraquara" pageId="43" pageNumber="44" specimenCode="MHNCI 6835" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Parana" typeStatus="holotype">
<specimenCount id="9D0CFD5E302B873413B5F8C90B1AF8A2" box="[486,533,1793,1817]" count="2" pageId="43" pageNumber="44" type="male">2 ♂</specimenCount>
<specimenCount id="9D0CFD5E302B87341049F8C90B47F8A2" box="[538,584,1793,1817]" count="2" pageId="43" pageNumber="44" type="female">2 ♀</specimenCount>
(
<specimenCode id="DBAC9EAC302B87341006F8C90BFFF8A3" box="[597,752,1793,1817]" collectionCode="MHNCI" country="0" httpUri="http://grbio.org/cool/ej4f-8m62" name="Museu de Historia Natural Capao de Imbuia (Brazil)" pageId="43" pageNumber="44">MHNCI 6835</specimenCode>
), from 
<collectingCountry id="F31D7647302B873412FCF8E90807F882" box="[175,264,1825,1848]" name="Brazil" pageId="43" pageNumber="44">BRAZIL</collectingCountry>
, state of 
<collectingRegion id="49CEF835302B8734133CF8E808BBF882" box="[367,436,1824,1848]" country="Brazil" name="Parana" pageId="43" pageNumber="44">Paraná</collectingRegion>
, 
<taxonomicName id="4C0A4D54302B87341390F8E80B84F882" authority=", Banhado" authorityName="Banhado" box="[451,651,1824,1848]" class="Arachnida" family="Gonyleptidae" genus="Piraquara" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Opiliones" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<collectingMunicipality id="6BD1ACAD302B87341390F8E80B2BF882" box="[451,548,1824,1848]" pageId="43" pageNumber="44">Piraquara</collectingMunicipality>
, 
<location id="8ED5600C302B87341061F8E80B84F882" LSID="urn:lsid:plazi:treatment:03A387C1302A872F1198F9710C45FA32:8ED5600C302B87341061F8E80B84F882" box="[562,651,1824,1848]" country="Brazil" municipality="Piraquara" name="Banhado" pageId="43" pageNumber="44" stateProvince="Parana">Banhado</location>
</taxonomicName>
, 
<date id="FFB41017302B873410C9F8E80BF7F882" box="[666,760,1824,1848]" pageId="43" pageNumber="44" value="1991-01-13">
<collectingDate id="EFF0E9FF302B873410C9F8E80BF7F882" box="[666,760,1824,1848]" pageId="43" pageNumber="44" value="1991-01-13">13.i.1991</collectingDate>
</date>
, 
<collectorName id="26FF5301302B87341222F888083FF8ED" box="[113,304,1856,1880]" pageId="43" pageNumber="44">Pinto-da-Rocha, R.</collectorName>
&amp; 
<collectorName id="26FF5301302B87341318F88808C6F8E2" box="[331,457,1856,1880]" pageId="43" pageNumber="44">Bérnils, R. S.</collectorName>
<emphasis id="B97EEAC5302B8734139EF88808E7F8E2" box="[461,488,1856,1880]" italics="true" pageId="43" pageNumber="44">leg</emphasis>
</materialsCitation>
<emphasis id="B97EEAC5302B873413BBF88808E3F8E2" box="[488,492,1856,1880]" italics="true" pageId="43" pageNumber="44">.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C310655C302B873413D4F8B60A4CFB17" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="8BB536D7302B873413D4F8B608EBF82D" blockId="43.[113,762,1918,2011]" box="[391,484,1918,1943]" pageId="43" pageNumber="44">
<emphasis id="B97EEAC5302B873413D4F8B608EBF82D" box="[391,484,1918,1943]" italics="true" pageId="43" pageNumber="44">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BB536D7302B87341222F86C0A4CFB17" blockId="43.[113,762,1918,2011]" lastBlockId="43.[810,1459,953,1197]" pageId="43" pageNumber="44">
<taxonomicName id="4C0A4D54302B87341222F86C082BF801" authorityName="Carvalho &amp; Kury" authorityYear="2025" box="[113,292,1956,1979]" class="Arachnida" family="Gonyleptidae" genus="Iamarinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="pontesi">
<emphasis id="B97EEAC5302B87341222F86C082BF801" box="[113,292,1956,1979]" italics="true" pageId="43" pageNumber="44">Iamarinus pontesi</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="4C0A4D54302B87341000F86C0BAEF806" baseAuthorityName="Kury, Pinto-da-Rocha &amp; Carvalho" baseAuthorityYear="2018" box="[595,673,1956,1980]" class="Arachnida" family="Gonyleptidae" genus="Iamarinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="fenax">
<emphasis id="B97EEAC5302B87341000F86C0BAEF806" box="[595,673,1956,1980]" italics="true" pageId="43" pageNumber="44">I. fenax</emphasis>
</taxonomicName>
due to: (1) Scutal area I with three pairs of paramedian conspicuous tubercles (
<figureCitation id="13312A52302B873411C4FC710AFFFC6B" box="[919,1008,953,977]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="43" pageNumber="44">Figs 25A</figureCitation>
, 
<figureCitation id="13312A52302B873411AEFC710D28FC6B" box="[1021,1063,953,977]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="43" pageNumber="44">26A</figureCitation>
); (2) Cx IV prodorsal distal apophysis with distal portion forming an acute angle in relation to the longitudinal axis (
<figureCitation id="13312A52302B873411B9FC300D48FBAA" box="[1002,1095,1016,1040]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="43" pageNumber="44">Figs 25A</figureCitation>
, 
<figureCitation id="13312A52302B87341604FC300D8EFBAA" box="[1111,1153,1016,1040]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="43" pageNumber="44">26A</figureCitation>
); (3) Cx IV prodorsal distal apophysis short, slightly bent distad (
<figureCitation id="13312A52302B873416F4FBDF0C38FB95" box="[1191,1335,1047,1071]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="43" pageNumber="44">Fig. 25A–B, D</figureCitation>
); (4) Cx IV prodorsal apophysis with a second branch on central third (
<figureCitation id="13312A52302B873417DAFBFF0A7AFBD4" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="43" pageNumber="44">Figs 25A–C</figureCitation>
, 
<figureCitation id="13312A52302B873411D2FB9E0AC3FBD4" box="[897,972,1110,1134]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="43" pageNumber="44">26A, D</figureCitation>
); (5) Fe IV sinuous (
<figureCitation id="13312A52302B873416FDFB9E0C08FBD4" box="[1198,1287,1110,1134]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="43" pageNumber="44">Figs 25B</figureCitation>
, 
<figureCitation id="13312A52302B87341740FB9E0C56FBD4" box="[1299,1369,1110,1134]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="43" pageNumber="44">26F, H</figureCitation>
); (6) Ti IV with an apical bifurcated spine on retroventral face (
<figureCitation id="13312A52302B87341708FBBD0A3CFB16" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="43" pageNumber="44">Fig. 26F, I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C310655C302B87341666FB040A85FA90" pageId="43" pageNumber="44" type="distribution">
<paragraph id="8BB536D7302B87341666FB040DA7FB5C" blockId="43.[810,1459,1228,1322]" box="[1077,1192,1228,1254]" pageId="43" pageNumber="44">
<emphasis id="B97EEAC5302B87341666FB040DA7FB5C" box="[1077,1192,1228,1254]" italics="true" pageId="43" pageNumber="44">Distribution</emphasis>
</paragraph>
<paragraph id="8BB536D7302B87341179FB3B0A85FA90" blockId="43.[810,1459,1228,1322]" pageId="43" pageNumber="44">
<collectingCountry id="F31D7647302B87341179FB3B0A87FAB0" box="[810,904,1267,1290]" name="Brazil" pageId="43" pageNumber="44">BRAZIL</collectingCountry>
: state of 
<collectingRegion id="49CEF835302B873411B9FB3B0D3BFAB1" box="[1002,1076,1267,1291]" country="Brazil" name="Parana" pageId="43" pageNumber="44">Paraná</collectingRegion>
: 
<taxonomicName id="4C0A4D54302B87341613FB3B0CBCFAB1" authority=", Guaraquecaba, Piraquara" authorityName="Guaraquecaba, Piraquara" box="[1088,1459,1267,1291]" class="Insecta" family="Pseudococcidae" genus="Antonina" kingdom="Animalia" order="Hemiptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">Antonina, Guaraqueçaba, Piraquara</taxonomicName>
(
<figureCitation id="13312A52302B87341166FADA0A75FA90" box="[821,890,1298,1322]" captionStart="Figure 21" captionStartId="41.[113,178,1926,1950]" captionTargetBox="[118,1456,908,1896]" captionTargetId="figure-344@41.[115,1459,905,1899]" captionTargetPageId="41" captionText="Figure 21. South Brazil,showing distribution of Iamarinus species. Shaded areas in the background are Morrone’s regionalization of the Neotropics (‘provinces’; Morrone et al. 2022): Araucaria Forest province (in light green) and Atlantic province (in dark green)." figureDoi="http://doi.org/10.5281/zenodo.14802126" httpUri="https://zenodo.org/record/14802126/files/figure.png" pageId="43" pageNumber="44">Fig. 21</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C310655C302B872F167CFA820C45FA32" lastPageId="48" lastPageNumber="49" pageId="43" pageNumber="44" type="description">
<paragraph id="8BB536D7302B8734167CFA820DA2FADE" blockId="43.[809,1459,1354,1541]" box="[1071,1197,1354,1380]" pageId="43" pageNumber="44">
<emphasis id="B97EEAC5302B8734167CFA820DA2FADE" box="[1071,1197,1354,1380]" italics="true" pageId="43" pageNumber="44">Redescription</emphasis>
</paragraph>
<paragraph id="8BB536D7302B87341179FAB80D79F9BF" blockId="43.[809,1459,1354,1541]" pageId="43" pageNumber="44">
<emphasis id="B97EEAC5302B87341179FAB80AF1FA32" bold="true" box="[810,1022,1392,1416]" pageId="43" pageNumber="44">
<collectionCode id="ED1BAE12302B87341179FAB80A7EFA32" box="[810,881,1392,1416]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="43" pageNumber="44">MNRJ</collectionCode>
2789 (male)
</emphasis>
for the external body illustrations and description; DS, measurements: CW 3.3, CL 2.4, AW 6.0, AL 3.1; Leg I–IV measurements inTable 14; Right/left tarsal (distitarsal) counts: 6(3)/6(3) - 10(3)/10(3) - 7/7 - 7/7. 
<emphasis id="B97EEAC5302B8734174DFA060A7CF9BF" bold="true" pageId="43" pageNumber="44">
<collectionCode id="ED1BAE12302B8734174DFA060C6AFA5C" box="[1310,1381,1486,1510]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="43" pageNumber="44">MNRJ</collectionCode>
60562 (male)
</emphasis>
for genitalic illustrations.
</paragraph>
<paragraph id="8BB536D7302B87331179F9E00D92F957" blockId="43.[810,1461,1575,2006]" lastBlockId="44.[824,1475,1624,1773]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
<emphasis id="B97EEAC5302B87341179F9E00A8FF985" box="[810,896,1576,1599]" italics="true" pageId="43" pageNumber="44">Dorsum:</emphasis>
DS gamma-pyriform, as long as wide, with 
<collectingCountry id="F31D7647302B8734171DF9EF0C62F985" box="[1358,1389,1575,1599]" name="American Samoa" pageId="43" pageNumber="44">AS</collectingCountry>
lateral margins strongly convex, widest and thickest at scutal area 
<collectionCode id="ED1BAE12302B873417C0F98F0CA0F9E4" box="[1427,1455,1607,1630]" pageId="43" pageNumber="44">III</collectionCode>
, with sinuous posterior margin (
<figureCitation id="13312A52302B8734162AF9AD0DFCF9C4" box="[1145,1267,1637,1662]" captionStart="Figure 24" captionStartId="44.[130,195,1534,1558]" captionTargetBox="[129,1473,144,1496]" captionTargetId="figure-270@44.[129,1473,144,1496]" captionTargetPageId="44" captionText="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." figureDoi="http://doi.org/10.5281/zenodo.14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="43" pageNumber="44">Figs 24A–B</figureCitation>
, 
<figureCitation id="13312A52302B87341752F9AD0C48F9C7" box="[1281,1351,1637,1661]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="43" pageNumber="44">25A–B</figureCitation>
, 
<figureCitation id="13312A52302B87341705F9AD0CACF9C7" box="[1366,1443,1637,1661]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="43" pageNumber="44">26A, D</figureCitation>
). DS anterior margin divided by a small central projection in the center and a pair of shallow cheliceral sockets (
<figureCitation id="13312A52302B8734175CF96C0C65F906" box="[1295,1386,1700,1724]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="43" pageNumber="44">Figs 25A</figureCitation>
, 
<figureCitation id="13312A52302B87341724F96C0CADF906" box="[1399,1442,1700,1724]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="43" pageNumber="44">26A</figureCitation>
). Carapace posterior portion with a paramedian pair of prominent tubercles, surrounded by ordinary tubercles on lateral and posterior portions (
<figureCitation id="13312A52302B87341673F8CA0D75F8A0" box="[1056,1146,1794,1818]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="43" pageNumber="44">Figs 25A</figureCitation>
, 
<figureCitation id="13312A52302B873416D4F8CA0DDEF8A0" box="[1159,1233,1794,1818]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="43" pageNumber="44">26A, D</figureCitation>
). Ocularium elliptical (in dorsal view), high (
<emphasis id="B97EEAC5302B8734164AF8EA0D2DF883" box="[1049,1058,1826,1849]" italics="true" pageId="43" pageNumber="44">c</emphasis>
. 4× the eye diameter), perpendicularly placed in the middle of the carapace (
<figureCitation id="13312A52302B873416CAF8890C02F8E3" box="[1177,1293,1857,1881]" captionStart="Figure 24" captionStartId="44.[130,195,1534,1558]" captionTargetBox="[129,1473,144,1496]" captionTargetId="figure-270@44.[129,1473,144,1496]" captionTargetPageId="44" captionText="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." figureDoi="http://doi.org/10.5281/zenodo.14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="43" pageNumber="44">Figs 24A–B</figureCitation>
, 
<figureCitation id="13312A52302B87341745F8890C52F8E3" box="[1302,1373,1857,1881]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="43" pageNumber="44">25A–B</figureCitation>
, 
<figureCitation id="13312A52302B87341735F8890A32F8C2" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="43" pageNumber="44">26A–B, D</figureCitation>
). Ocularium with a pair of almost parallel spines (
<emphasis id="B97EEAC5302B87341764F8A90C4FF8C2" box="[1335,1344,1889,1912]" italics="true" pageId="43" pageNumber="44">c</emphasis>
. 3× the eye diameter) (
<figureCitation id="13312A52302B873411F2F8B70D16F822" box="[929,1049,1919,1944]" captionStart="Figure 24" captionStartId="44.[130,195,1534,1558]" captionTargetBox="[129,1473,144,1496]" captionTargetId="figure-270@44.[129,1473,144,1496]" captionTargetPageId="44" captionText="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." figureDoi="http://doi.org/10.5281/zenodo.14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="43" pageNumber="44">Figs 24A–B</figureCitation>
, 
<figureCitation id="13312A52302B87341676F8B70D63F82D" box="[1061,1132,1919,1943]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="43" pageNumber="44">25A–B</figureCitation>
, 
<figureCitation id="13312A52302B8734162AF8B70DD0F82D" box="[1145,1247,1919,1943]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="43" pageNumber="44">26A–B, D</figureCitation>
). 
<collectingCountry id="F31D7647302B873416A4F8480C19F822" box="[1271,1302,1920,1944]" name="American Samoa" pageId="43" pageNumber="44">AS</collectingCountry>
lateral margins with two rows of tubercles: one external, composed of four–five prominent subconical tubercles at areas II–IV; another internal one with ordinary tubercles from the posterior corner of the carapace to the posterior margin (
<figureCitation id="13312A52302C8733138DF9BF0B37F935" box="[478,568,1655,1679]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="44" pageNumber="45">Figs 25A</figureCitation>
, 
<figureCitation id="13312A52302C87331016F9BF0B7FF935" box="[581,624,1655,1679]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="44" pageNumber="45">26A</figureCitation>
). Mesotergum is divided into four clearly defined areas (
<figureCitation id="13312A52302C8733107EF95E0B87F914" box="[557,648,1686,1710]" captionStart="Figure 24" captionStartId="44.[130,195,1534,1558]" captionTargetBox="[129,1473,144,1496]" captionTargetId="figure-270@44.[129,1473,144,1496]" captionTargetPageId="44" captionText="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." figureDoi="http://doi.org/10.5281/zenodo.14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="44" pageNumber="45">Figs 24A</figureCitation>
, 
<figureCitation id="13312A52302C873310C5F95E0BCFF914" box="[662,704,1686,1710]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="44" pageNumber="45">25A</figureCitation>
, 
<figureCitation id="13312A52302C8733109DF95E0BF7F914" box="[718,760,1686,1710]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="44" pageNumber="45">26A</figureCitation>
). All scutal areas tuberculate (
<figureCitation id="13312A52302C873313CDF97E08F8F977" box="[414,503,1717,1742]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="44" pageNumber="45">Figs 25A</figureCitation>
, 
<figureCitation id="13312A52302C87331050F97D0B42F977" box="[515,589,1717,1741]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="44" pageNumber="45">26A, D</figureCitation>
). Scutal area I divided into left and right halves by a longitudinal median groove (
<figureCitation id="13312A52302C873312D8F93C09E9F8B6" box="[139,230,1780,1804]" captionStart="Figure 24" captionStartId="44.[130,195,1534,1558]" captionTargetBox="[129,1473,144,1496]" captionTargetId="figure-270@44.[129,1473,144,1496]" captionTargetPageId="44" captionText="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." figureDoi="http://doi.org/10.5281/zenodo.14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="44" pageNumber="45">Figs 24A</figureCitation>
, 
<figureCitation id="13312A52302C873312A0F93C0812F8B6" box="[243,285,1780,1804]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="44" pageNumber="45">25A</figureCitation>
, 
<figureCitation id="13312A52302C87331378F93C085AF8B6" box="[299,341,1780,1804]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="44" pageNumber="45">26A</figureCitation>
). Scutal areas I–II with three pairs of conspicuous tubercles (
<emphasis id="B97EEAC5302C8733130BF8DC086EF891" box="[344,353,1812,1835]" italics="true" pageId="44" pageNumber="45">c</emphasis>
. 2× the ordinary tubercles) (
<figureCitation id="13312A52302C873310F0F8DC0A0DF891" box="[675,770,1811,1836]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="44" pageNumber="45">Figs 25A</figureCitation>
, 
<figureCitation id="13312A52302C873312D2F8FB09C2F8F0" box="[129,205,1843,1867]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="44" pageNumber="45">26A, D</figureCitation>
). Scutal area II posterior-lateral margin embracing the scutal area 
<collectionCode id="ED1BAE12302C873312BCF89B0805F8D0" box="[239,266,1875,1898]" pageId="44" pageNumber="45">III</collectionCode>
(
<figureCitation id="13312A52302C8733134AF89A087DF8D0" box="[281,370,1874,1898]" captionStart="Figure 24" captionStartId="44.[130,195,1534,1558]" captionTargetBox="[129,1473,144,1496]" captionTargetId="figure-270@44.[129,1473,144,1496]" captionTargetPageId="44" captionText="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." figureDoi="http://doi.org/10.5281/zenodo.14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="44" pageNumber="45">Figs 24A</figureCitation>
, 
<figureCitation id="13312A52302C8733132DF89A08A7F8D0" box="[382,424,1874,1898]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="44" pageNumber="45">25A</figureCitation>
, 
<figureCitation id="13312A52302C873313E7F89A08D1F8D0" box="[436,478,1874,1898]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="44" pageNumber="45">26A</figureCitation>
). Scutal area 
<collectionCode id="ED1BAE12302C87331035F89B0B8EF8D0" box="[614,641,1875,1898]" pageId="44" pageNumber="45">III</collectionCode>
with a pair of paramedian outstanding subconical spines (
<emphasis id="B97EEAC5302C87331016F8BA0B41F833" box="[581,590,1906,1929]" italics="true" pageId="44" pageNumber="45">c</emphasis>
. 14× the ordinary tubercles) (
<figureCitation id="13312A52302C873312A4F859087EF813" box="[247,369,1937,1961]" captionStart="Figure 24" captionStartId="44.[130,195,1534,1558]" captionTargetBox="[129,1473,144,1496]" captionTargetId="figure-270@44.[129,1473,144,1496]" captionTargetPageId="44" captionText="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." figureDoi="http://doi.org/10.5281/zenodo.14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="44" pageNumber="45">Figs 24A–C</figureCitation>
, 
<figureCitation id="13312A52302C8733132FF85908A9F813" box="[380,422,1937,1961]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="44" pageNumber="45">25A</figureCitation>
, 
<figureCitation id="13312A52302C873313E1F8590B14F813" box="[434,539,1937,1961]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="44" pageNumber="45">26A, C–D</figureCitation>
). Scutal area IV central portion with a transversal row of five–six prominent tubercles (
<emphasis id="B97EEAC5302C87331117F9B00A42F935" box="[836,845,1656,1679]" italics="true" pageId="44" pageNumber="45">c</emphasis>
. 2× the ordinary tubercles) (
<figureCitation id="13312A52302C87331627F9BF0DC0F935" box="[1140,1231,1655,1679]" captionStart="Figure 24" captionStartId="44.[130,195,1534,1558]" captionTargetBox="[129,1473,144,1496]" captionTargetId="figure-270@44.[129,1473,144,1496]" captionTargetPageId="44" captionText="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." figureDoi="http://doi.org/10.5281/zenodo.14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="44" pageNumber="45">Figs 24C</figureCitation>
, 
<figureCitation id="13312A52302C87331689F9BF0C0BF935" box="[1242,1284,1655,1679]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="44" pageNumber="45">25A</figureCitation>
, 
<figureCitation id="13312A52302C87331742F9BF0C55F934" box="[1297,1370,1655,1679]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="44" pageNumber="45">26A, D</figureCitation>
). DS posterior margin and free tergites I– 
<collectionCode id="ED1BAE12302C8733162DF95F0D97F914" box="[1150,1176,1687,1710]" pageId="44" pageNumber="45">III</collectionCode>
each with a transversal row of tubercles, growing in size towards the central portion (
<figureCitation id="13312A52302C87331731F97E0CB3F974" box="[1378,1468,1718,1742]" captionStart="Figure 24" captionStartId="44.[130,195,1534,1558]" captionTargetBox="[129,1473,144,1496]" captionTargetId="figure-270@44.[129,1473,144,1496]" captionTargetPageId="44" captionText="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." figureDoi="http://doi.org/10.5281/zenodo.14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="44" pageNumber="45">Figs 24C</figureCitation>
, 
<figureCitation id="13312A52302C8733116AF91D0A6BF957" box="[825,868,1749,1773]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="44" pageNumber="45">26A</figureCitation>
). Anal operculum tuberculate.
</paragraph>
<caption id="DF75665F302C873312D1FA3608B3F989" ID-DOI="http://doi.org/10.5281/zenodo.14802132" ID-Zenodo-Dep="14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="44" pageNumber="45" startId="44.[130,195,1534,1558]" targetBox="[129,1473,144,1496]" targetPageId="44" targetType="figure">
<paragraph id="8BB536D7302C873312D1FA3608B3F989" blockId="44.[129,1412,1534,1587]" pageId="44" pageNumber="45">
<emphasis id="B97EEAC5302C873312D1FA3609E9F9AC" bold="true" box="[130,230,1534,1558]" pageId="44" pageNumber="45">Figure 24.</emphasis>
Habitus 
<emphasis id="B97EEAC5302C87331369FA37087BF9AC" box="[314,372,1535,1558]" italics="true" pageId="44" pageNumber="45">in vivo</emphasis>
of 
<taxonomicName id="4C0A4D54302C873313C2FA370B3DF9AC" authorityName="Carvalho &amp; Kury" authorityYear="2025" box="[401,562,1535,1558]" class="Arachnida" family="Gonyleptidae" genus="Iamarinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pontesi">
<emphasis id="B97EEAC5302C873313C2FA370B3DF9AC" box="[401,562,1535,1558]" italics="true" pageId="44" pageNumber="45">Iamarinus pontesi</emphasis>
</taxonomicName>
, from Brazil, Paraná, 
<taxonomicName id="4C0A4D54302C873310A9FA360A59F9AC" authority="Arapoti, Guaraquecaba" authorityName="Arapoti, Guaraquecaba" box="[762,854,1534,1558]" class="Insecta" family="Pseudococcidae" genus="Antonina" kingdom="Animalia" order="Hemiptera" pageId="44" pageNumber="25" phylum="Arthropoda" rank="genus">Antonina</taxonomicName>
. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E).
</paragraph>
</caption>
<paragraph id="8BB536D7302C8732116AF8DD09D9FB33" blockId="44.[825,1474,1812,1962]" lastBlockId="45.[113,763,980,1161]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<emphasis id="B97EEAC5302C8733116AF8DD0A71F896" box="[825,894,1813,1836]" italics="true" pageId="44" pageNumber="45">Venter:</emphasis>
Cx I– 
<collectionCode id="ED1BAE12302C87331194F8DD0AEEF896" box="[967,993,1813,1836]" pageId="44" pageNumber="45">III</collectionCode>
sub-parallel to each other, each with ventral longitudinal rows of 8–13 setiferous tubercles (Cx I rows with higher and sharper tubercles than the others). Cx II with a retroventral distal row of six acuminated tubercles. Cx 
<collectionCode id="ED1BAE12302C87331730F8BB0C71F830" box="[1379,1406,1907,1930]" pageId="44" pageNumber="45">III</collectionCode>
with a retroventral distal row of 10 acuminated tubercles. Cx IV much larger than the others, directed obliquely (
<figureCitation id="13312A52302D87321076FC1C0B72FC56" box="[549,637,980,1004]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">Fig. 25C</figureCitation>
). Intercoxal bridges are well marked (
<figureCitation id="13312A52302D8732132EFC3C08D9FBB6" box="[381,470,1012,1036]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">Fig. 25C</figureCitation>
). Stigmatic area Y-inverted-shaped, clearly sunken concerning Cx IV’s distal part (
<figureCitation id="13312A52302D873210C0FBDB0BE5FB91" box="[659,746,1043,1067]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">Fig. 25C</figureCitation>
). Cx IV covered by ordinary tubercles (
<figureCitation id="13312A52302D873213B9FBFA0B4EFBF0" box="[490,577,1074,1098]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">Fig. 25C</figureCitation>
). Stigmata are visible (
<figureCitation id="13312A52302D873212FBFB9A080FFBD0" box="[168,256,1106,1130]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">Fig. 25C</figureCitation>
). Free sternites with a transverse row of ordinary tubercles.
</paragraph>
<caption id="DF75665F302D87321222FCA00D3BFC26" ID-DOI="http://doi.org/10.5281/zenodo.14802134" ID-Zenodo-Dep="14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46" startId="45.[113,178,872,896]" targetBox="[113,1457,144,834]" targetPageId="45" targetType="figure">
<paragraph id="8BB536D7302D87321222FCA00D3BFC26" blockId="45.[113,1394,872,925]" pageId="45" pageNumber="46">
<emphasis id="B97EEAC5302D87321222FCA009DAFC3A" bold="true" box="[113,213,872,896]" pageId="45" pageNumber="46">Figure 25.</emphasis>
<taxonomicName id="4C0A4D54302D87321289FCA10874FC3A" authorityName="Carvalho &amp; Kury" authorityYear="2025" box="[218,379,873,896]" class="Arachnida" family="Gonyleptidae" genus="Iamarinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pontesi">
<emphasis id="B97EEAC5302D87321289FCA10874FC3A" box="[218,379,873,896]" italics="true" pageId="45" pageNumber="46">Iamarinus pontesi</emphasis>
</taxonomicName>
, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view. E, same, lateral view. F, same, ventral view. Scale bars = 5 mm.
</paragraph>
</caption>
<paragraph id="8BB536D7302D87321222FB6B09C9FAA3" blockId="45.[113,762,1187,1305]" pageId="45" pageNumber="46">
<emphasis id="B97EEAC5302D87321222FB6B09DCFB01" box="[113,211,1187,1211]" italics="true" pageId="45" pageNumber="46">Chelicera:</emphasis>
Basichelicerite elongate, bulla well marked (
<figureCitation id="13312A52302D873210C7FB6B0BE6FB01" box="[660,745,1187,1211]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">Fig. 26A</figureCitation>
), with marginal setiferous tubercles—two or three lateral ectal, one or two posteriors, two lateral mesal (
<figureCitation id="13312A52302D87321074FB2A0B8FFB40" box="[551,640,1250,1274]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">Fig. 26A</figureCitation>
); hand not swollen.
</paragraph>
<paragraph id="8BB536D7302D87321222FAFB09FDFA52" blockId="45.[113,765,1331,1512]" pageId="45" pageNumber="46">
<emphasis id="B97EEAC5302D87321222FAFB09D9FAF1" box="[113,214,1331,1355]" italics="true" pageId="45" pageNumber="46">Pedipalps:</emphasis>
Tr with two geminated ventral setiferous tubercles (
<figureCitation id="13312A52302D8732122FFA9A09DAFAD0" box="[124,213,1362,1386]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">Fig. 25C</figureCitation>
). Fe with a ventral basal and a mesal distal setiferous tubercle (
<figureCitation id="13312A52302D87321284FABA083FFA30" box="[215,304,1394,1418]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">Fig. 25C</figureCitation>
). Pa unarmed (
<figureCitation id="13312A52302D87321385FABA0B3FFA30" box="[470,560,1394,1418]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">Fig. 25C</figureCitation>
). Ti with two rows of four (IiIi) spines on ventro-mesal and ventro-ectal faces. Ta with two rows of spines—three (IIi) ventro-mesal and four (IiIi) ventro-ectal.
</paragraph>
<paragraph id="8BB536D7302D872F1222F9C20842FE9D" blockId="45.[112,764,1545,1977]" lastBlockId="48.[127,777,146,295]" lastPageId="48" lastPageNumber="49" pageId="45" pageNumber="46">
<emphasis id="B97EEAC5302D87321222F9C209ABF99B" box="[113,164,1546,1569]" italics="true" pageId="45" pageNumber="46">Legs:</emphasis>
All the unmentioned podomeres are unarmed or without relevant armature. Cx I–II dorsal proximal face with anterior and posterior basal apophyses (linked with ozopores); simple ones on Cx I, prominent ones on Cx II (posterior apophysis bifurcated, with the anterior bud larger and swollen). Tr I– 
<collectionCode id="ED1BAE12302D873210FAF94F0BCCF924" box="[681,707,1671,1694]" pageId="45" pageNumber="46">III</collectionCode>
each with several ventral tubercles. Fe I–II straight (
<figureCitation id="13312A52302D87321001F96E0BA6F904" box="[594,681,1702,1726]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">Fig. 25A</figureCitation>
). Fe 
<collectionCode id="ED1BAE12302D8732108CF96E0BF5F907" box="[735,762,1702,1725]" pageId="45" pageNumber="46">III</collectionCode>
sub-straight (
<figureCitation id="13312A52302D873212AAF90D0842F967" box="[249,333,1733,1757]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">Fig.25A</figureCitation>
). Fe and Ti I–II with prodorsal, proventral, retroventral, and retrodorsal rows of small tubercles. Fe 
<collectionCode id="ED1BAE12302D873210E2F92D0BC3F946" box="[689,716,1765,1788]" pageId="45" pageNumber="46">III</collectionCode>
and Ti 
<collectionCode id="ED1BAE12302D873212C0F8CC09A1F8A1" box="[147,174,1796,1819]" pageId="45" pageNumber="46">III</collectionCode>
with prodorsal, proventral, retroventral, and retrodorsal rows of tubercles (Fe 
<collectionCode id="ED1BAE12302D8732130AF8EC087BF881" box="[345,372,1828,1851]" pageId="45" pageNumber="46">III</collectionCode>
proventral and retroventral tubercles larger and sharper than others). Fe 
<collectionCode id="ED1BAE12302D87321384F88B08FDF8E0" box="[471,498,1859,1882]" pageId="45" pageNumber="46">III</collectionCode>
with an apical retrodorsal spur (
<figureCitation id="13312A52302D873212E3F8AA0808F8C0" box="[176,263,1890,1914]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">Fig. 25A</figureCitation>
). Cx IV reaches scutal area IV (
<figureCitation id="13312A52302D87321005F8AA0BBEF8C0" box="[598,689,1890,1914]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">Figs 25A</figureCitation>
, 
<figureCitation id="13312A52302D873210ECF8AA0BE6F8C0" box="[703,745,1890,1914]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">26A</figureCitation>
). Cx IV tuberculate between prodorsal and ventral faces (
<figureCitation id="13312A52302D87321083F84909B3F802" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">Figs 25A–C</figureCitation>
, 
<figureCitation id="13312A52302D8732129FF86809F8F802" box="[204,247,1952,1976]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">26A</figureCitation>
). Cx IV with a prodorsal subconical apophysis, slightly bent to posterior, bearing a small accessory blunt branch on its central posterior third (
<figureCitation id="13312A52302D87321634FC3C0DEBFBB6" box="[1127,1252,1012,1036]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">Figs 25A–C</figureCitation>
, 
<figureCitation id="13312A52302D873216A2FC3C0C54FBB1" box="[1265,1371,1012,1036]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">26A, D–E</figureCitation>
). Cx IV with a short retrolateral apophysis, fused with a small secondary branch (
<figureCitation id="13312A52302D873211D3FBFA0AD5FBF1" box="[896,986,1074,1099]" captionStart="Figure 24" captionStartId="44.[130,195,1534,1558]" captionTargetBox="[129,1473,144,1496]" captionTargetId="figure-270@44.[129,1473,144,1496]" captionTargetPageId="44" captionText="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." figureDoi="http://doi.org/10.5281/zenodo.14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="45" pageNumber="46">Figs 24C</figureCitation>
, 
<figureCitation id="13312A52302D873211B7FBFA0D24FBF1" box="[996,1067,1074,1099]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">25A, C</figureCitation>
, 
<figureCitation id="13312A52302D87321666FBFA0D9BFBF0" box="[1077,1172,1074,1098]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">26A, F, H</figureCitation>
). Tr IV rectangle-shaped (in dorsal view) (
<figureCitation id="13312A52302D873211E8FB9A0D38FBD0" box="[955,1079,1106,1130]" captionStart="Figure 24" captionStartId="44.[130,195,1534,1558]" captionTargetBox="[129,1473,144,1496]" captionTargetId="figure-270@44.[129,1473,144,1496]" captionTargetPageId="44" captionText="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." figureDoi="http://doi.org/10.5281/zenodo.14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="45" pageNumber="46">Figs 24A–C</figureCitation>
, 
<figureCitation id="13312A52302D87321610FB9A0D81FBD0" box="[1091,1166,1106,1130]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">25A–C</figureCitation>
, 
<figureCitation id="13312A52302D873216C9FB9A0C0EFBD3" box="[1178,1281,1106,1130]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">26A, F–H</figureCitation>
). Tr IV proximal portion with a conical apophysis on prolateral and retrolateral faces (
<figureCitation id="13312A52302D87321138FB580AEBFB13" box="[875,996,1168,1193]" captionStart="Figure 24" captionStartId="44.[130,195,1534,1558]" captionTargetBox="[129,1473,144,1496]" captionTargetId="figure-270@44.[129,1473,144,1496]" captionTargetPageId="44" captionText="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." figureDoi="http://doi.org/10.5281/zenodo.14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="45" pageNumber="46">Figs 24A–C</figureCitation>
, 
<figureCitation id="13312A52302D873211BDFB580D5CFB12" box="[1006,1107,1168,1192]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">26A, F–H</figureCitation>
). Tr IV distal portion with a screwdriver tip-shaped transversal apophysis, dorsally covered with four tubercles, on prodorsal face (
<figureCitation id="13312A52302D873216D7FB070DD1FB5D" box="[1156,1246,1231,1255]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="45" pageNumber="46">Figs 25A</figureCitation>
, 
<figureCitation id="13312A52302D873216B8FB070C69FB5D" box="[1259,1382,1231,1255]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">26A, F–G, I</figureCitation>
). Tr IV with two proximal and one distal subconical prominent tubercles on prolateral face (
<figureCitation id="13312A52302D8732164CFAC60D7BFA9C" box="[1055,1140,1294,1318]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">Fig. 26F</figureCitation>
). Tr IV tuberculate on ventral face (
<figureCitation id="13312A52302D87321136FAE50ACEFAFF" box="[869,961,1325,1349]" captionStart="Figure 24" captionStartId="44.[130,195,1534,1558]" captionTargetBox="[129,1473,144,1496]" captionTargetId="figure-270@44.[129,1473,144,1496]" captionTargetPageId="44" captionText="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." figureDoi="http://doi.org/10.5281/zenodo.14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="45" pageNumber="46">Figs 24C</figureCitation>
, 
<figureCitation id="13312A52302D8732119DFAE50D1DFAFF" box="[974,1042,1325,1349]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">26G–I</figureCitation>
). Tr IV distal portion with a subconical apophysis on retrolateral face (
<figureCitation id="13312A52302D87321623FA840C0EFADF" box="[1136,1281,1356,1381]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">Figs 26F, G–I</figureCitation>
). Fe IV sinuous, arched (1) on the proximal portion towards the prodorsal face and (2) on the distal portion towards the retroventral face (
<figureCitation id="13312A52302D873217DAFA430A7AFA79" captionStart="Figure 24" captionStartId="44.[130,195,1534,1558]" captionTargetBox="[129,1473,144,1496]" captionTargetId="figure-270@44.[129,1473,144,1496]" captionTargetPageId="44" captionText="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." figureDoi="http://doi.org/10.5281/zenodo.14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="45" pageNumber="46">Figs 24A–C</figureCitation>
, 
<figureCitation id="13312A52302D873211D2FA620AB0FA78" box="[897,959,1450,1474]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">26F–I</figureCitation>
). Fe IV dorsal face with four spines on the proximal half and two spines on the distal third (
<figureCitation id="13312A52302D873216A6FA020C8CFA5B" box="[1269,1411,1482,1506]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">Figs 26F–G, I</figureCitation>
). Fe IV prodorsal face with a row of ordinary tubercles intercalated by two subconical spines on the central third (
<figureCitation id="13312A52302D8732175EF9C00C8CF99A" box="[1293,1411,1544,1568]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">Fig. 26F–G</figureCitation>
). Fe IV prolateral face with a row of tubercles divided in (1) ordinary tubercles on the proximal third, (2) prominent tubercles on the central third (
<emphasis id="B97EEAC5302D873211B4F9AF0AFFF9C4" box="[999,1008,1639,1662]" italics="true" pageId="45" pageNumber="46">c</emphasis>
. 2.5×–3× the ordinary ones), and (3) outstanding tubercles on the distal third (
<emphasis id="B97EEAC5302D873216EFF94F0DCAF924" box="[1212,1221,1671,1694]" italics="true" pageId="45" pageNumber="46">c</emphasis>
. 3.5×–4× the ordinary ones) (
<figureCitation id="13312A52302D8732112BF96D0AFEF907" box="[888,1009,1701,1725]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">Fig. 26F–H</figureCitation>
). Fe IV proventral face with (1) a row of ordinary tubercles on proximal and central thirds and (2) outstanding subconical tubercles interpolated by ordinary ones and a distalmost spine on the distal third (
<figureCitation id="13312A52302D873216F0F8CB0C13F8A1" box="[1187,1308,1795,1819]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">Fig. 26G–H</figureCitation>
). Fe IV ventral face with a prominent tubercle on the proximal fifth (
<figureCitation id="13312A52302D8732171AF8EA0CADF880" box="[1353,1442,1826,1850]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">Fig. 26H</figureCitation>
). Fe IV retroventral face with subconical outstanding tubercles on proximal and distal thirds and prominent tubercles on the central third (
<figureCitation id="13312A52302D873211C1F8480AF0F822" box="[914,1023,1920,1944]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="45" pageNumber="46">Fig. 26H–I</figureCitation>
). Fe IV retrolateral face with six spines (the two distalmost larger than the Fe IV diameter) on proximal third and with a pair of outstanding spines (distalmost largest) on the central third (
<figureCitation id="13312A52302F87301355F88B089CF8E1" box="[262,403,1859,1883]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="47" pageNumber="48">Fig. 26F, H–I</figureCitation>
). Fe IV retrodorsal face with two outstanding subconical tubercles and three prominent tubercles on the proximal third and three conical spines on the distal third (
<figureCitation id="13312A52302F8730122FF86909E9F803" box="[124,230,1953,1977]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="47" pageNumber="48">Fig. 26F, I</figureCitation>
). Pa IV dorsal face tuberculate (
<figureCitation id="13312A52302F8730106FF8690BC6F803" box="[572,713,1953,1977]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="47" pageNumber="48">Fig. 26F–G, I</figureCitation>
). Pa IV proventral and retroventral faces with rows of four and three spines, respectively (
<figureCitation id="13312A52302F87301656F88B0D78F8E1" box="[1029,1143,1859,1883]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="47" pageNumber="48">Fig. 26G–I</figureCitation>
). Ti IV with all faces (except ventral face) containing longitudinal rows of acuminated tubercles (
<figureCitation id="13312A52302F87301130F84A0AC1F820" box="[867,974,1922,1946]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="47" pageNumber="48">Fig. 26F–I</figureCitation>
). Ti IV retrolateral face with three-four prominent subconical tubercles on proximal half (
<figureCitation id="13312A52302F873016A9F8690C8BF803" box="[1274,1412,1953,1977]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="47" pageNumber="48">Fig. 26F, H–I</figureCitation>
). Ti IV proventral and retroventral faces with spines on the distal half (the retroventral distalmost spine is bifurcated) (
<figureCitation id="13312A523030872F103EFF790BDAFF73" box="[621,725,177,201]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="48" pageNumber="49">Fig. 26F–I</figureCitation>
). Mt IV with all faces containing longitudinal rows of small subconical tubercles (
<figureCitation id="13312A523030872F12BDFF380832FEB2" box="[238,317,240,264]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="48" pageNumber="49">Fig. 26J</figureCitation>
). Mt IV with proventral and retroventral apical spurs (
<figureCitation id="13312A523030872F12BCFEC70832FE9D" box="[239,317,271,295]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[130,1474,146,1751]" captionTargetId="figure-11@46.[127,1477,143,1754]" captionTargetPageId="46" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="48" pageNumber="49">Fig. 26J</figureCitation>
).
</paragraph>
<caption id="DF75665F302E8731122CF8C80881F8D6" ID-DOI="http://doi.org/10.5281/zenodo.14802138" ID-Zenodo-Dep="14802138" httpUri="https://zenodo.org/record/14802138/files/figure.png" pageId="46" pageNumber="47" startId="46.[127,192,1792,1816]" targetBox="[130,1474,146,1751]" targetPageId="46" targetType="figure">
<paragraph id="8BB536D7302E8731122CF8C80881F8D6" blockId="46.[127,1448,1792,1900]" pageId="46" pageNumber="47">
<emphasis id="B97EEAC5302E8731122CF8C809ECF8A2" bold="true" box="[127,227,1792,1816]" pageId="46" pageNumber="47">Figure 26.</emphasis>
<taxonomicName id="4C0A4D54302E873112B4F8C90887F8A2" authorityName="Carvalho &amp; Kury" authorityYear="2025" box="[231,392,1793,1816]" class="Arachnida" family="Gonyleptidae" genus="Iamarinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pontesi">
<emphasis id="B97EEAC5302E873112B4F8C90887F8A2" box="[231,392,1793,1816]" italics="true" pageId="46" pageNumber="47">Iamarinus pontesi</emphasis>
</taxonomicName>
, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, ocularium, frontal view. C, armature of scutal area III, posterior view. D, habitus, lateral view. E, detail of the right coxa IV prodorsal apophysis, dorsal view. F, right trochanter–tibia IV, dorsal view. G, same, prolateral view. H, same, ventral view. I, same, retrolateral view. J, right metatarsus and tarsus IV, retrolateral view. Scale bars: 1 mm (B–C, E), 5 mm (A, D, F–J).
</paragraph>
</caption>
<caption id="DF75665F302F87301221FCE40CA4FCFF" ID-DOI="http://doi.org/10.5281/zenodo.14802140" ID-Zenodo-Dep="14802140" box="[114,1451,812,837]" httpUri="https://zenodo.org/record/14802140/files/figure.png" pageId="47" pageNumber="48" startId="47.[114,177,812,836]" targetBox="[117,1455,150,771]" targetPageId="47" targetType="figure">
<paragraph id="8BB536D7302F87301221FCE40CA4FCFF" blockId="47.[114,1451,812,837]" box="[114,1451,812,837]" pageId="47" pageNumber="48">
<emphasis id="B97EEAC5302F87301221FCE409DAFCFE" bold="true" box="[114,213,812,836]" pageId="47" pageNumber="48">Figure 27.</emphasis>
<taxonomicName id="4C0A4D54302F8730128BFCE5087BFCFE" authorityName="Carvalho &amp; Kury" authorityYear="2025" box="[216,372,813,836]" class="Arachnida" family="Gonyleptidae" genus="Iamarinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pontesi">
<emphasis id="B97EEAC5302F8730128BFCE5087BFCFE" box="[216,372,813,836]" italics="true" pageId="47" pageNumber="48">Iamarinus pontesi</emphasis>
</taxonomicName>
, ♂ paratype (MNRJ 60562), genitalia, distal part. A, dorsal view.B, lateral view. C, ventral view. Scale bars = 100 μm.
</paragraph>
</caption>
<caption id="DF75665F302F87301221F96B0B37F96D" ID-DOI="http://doi.org/10.5281/zenodo.14802142" ID-Zenodo-Dep="14802142" httpUri="https://zenodo.org/record/14802142/files/figure.png" pageId="47" pageNumber="48" startId="47.[114,179,1699,1723]" targetBox="[117,1455,887,1657]" targetPageId="47" targetType="figure">
<paragraph id="8BB536D7302F87301221F96B0B37F96D" blockId="47.[112,1447,1699,1751]" pageId="47" pageNumber="48">
<emphasis id="B97EEAC5302F87301221F96B09D8F901" bold="true" box="[114,215,1699,1723]" pageId="47" pageNumber="48">Figure 28.</emphasis>
<taxonomicName id="4C0A4D54302F8730128AF96C0875F901" authorityName="Carvalho &amp; Kury" authorityYear="2025" box="[217,378,1700,1723]" class="Arachnida" family="Gonyleptidae" genus="Iamarinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pontesi">
<emphasis id="B97EEAC5302F8730128AF96C0875F901" box="[217,378,1700,1723]" italics="true" pageId="47" pageNumber="48">Iamarinus pontesi</emphasis>
</taxonomicName>
, ♀ paratype (MNRJ 60562). A, ocularium, frontal view. B, habitus, dorsal view. C, right trochanter and femur IV, dorsal view. Scale bars: 1 mm (A), 3 mm (B–C).
</paragraph>
</caption>
<paragraph id="8BB536D73030872F12D2FE810BADFC8C" blockId="48.[128,778,329,823]" pageId="48" pageNumber="49">
<emphasis id="B97EEAC53030872F12D2FE8108B9FEDA" box="[129,438,329,353]" italics="true" pageId="48" pageNumber="49">
Coloration (in vivo) (
<figureCitation id="13312A523030872F130DFE8108ABFEDB" box="[350,420,329,353]" captionStart="Figure 24" captionStartId="44.[130,195,1534,1558]" captionTargetBox="[129,1473,144,1496]" captionTargetId="figure-270@44.[129,1473,144,1496]" captionTargetPageId="44" captionText="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." figureDoi="http://doi.org/10.5281/zenodo.14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="48" pageNumber="49">Fig. 24</figureCitation>
):
</emphasis>
DS and Cx–Tr IV background 
<emphasis id="B97EEAC53030872F12D2FEA0081CFEC5" box="[129,275,360,384]" italics="true" pageId="48" pageNumber="49">Blackish Green</emphasis>
(152). Ocularium spines, scutal areas II– 
<collectionCode id="ED1BAE123030872F1091FEA00BD3FEC5" box="[706,732,360,383]" pageId="48" pageNumber="49">III</collectionCode>
and Cx IV prodorsal distal apophysis 
<emphasis id="B97EEAC53030872F1380FE4F0B65FE25" box="[467,618,391,415]" italics="true" pageId="48" pageNumber="49">Brownish Black</emphasis>
(65). Carapace and scutal areas I–IV with tubercles 
<emphasis id="B97EEAC53030872F13ADFE6F0BA6FE04" box="[510,681,423,447]" italics="true" pageId="48" pageNumber="49">Brownish Orange</emphasis>
(54). 
<collectingCountry id="F31D76473030872F10B8FE6F0A05FE05" box="[747,778,423,447]" name="American Samoa" pageId="48" pageNumber="49">AS</collectingCountry>
lateral margins with prominent tubercles 
<taxonomicName id="4C0A4D543030872F107EFE0F0BA9FE64" authority="Brown" authorityName="Brown" box="[557,678,455,478]" genus="Deep" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" rank="genus">
<emphasis id="B97EEAC53030872F107EFE0F0BA9FE64" box="[557,678,455,478]" italics="true" pageId="48" pageNumber="49">Deep Brown</emphasis>
</taxonomicName>
(56). DS posterior margin and free tergites I– 
<collectionCode id="ED1BAE123030872F1043FE2E0B25FE47" box="[528,554,486,509]" pageId="48" pageNumber="49">III</collectionCode>
with tubercles 
<emphasis id="B97EEAC53030872F1085FE2D0800FDA7" italics="true" pageId="48" pageNumber="49">Dark Orange Yellow</emphasis>
(72). Ch and Pp glossier background 
<emphasis id="B97EEAC53030872F10FFFDCD09F3FD81" italics="true" pageId="48" pageNumber="49">Moderate Yellow Green</emphasis>
(120), with honeycombed 
<emphasis id="B97EEAC53030872F105CFDEC0A07FD81" box="[527,776,548,572]" italics="true" pageId="48" pageNumber="49">Dark Grayish Olive Green</emphasis>
(128) reticle. Tr I– 
<collectionCode id="ED1BAE123030872F1317FD8C0851FDE1" box="[324,350,580,603]" pageId="48" pageNumber="49">III</collectionCode>
background 
<emphasis id="B97EEAC53030872F13BBFD8C0BC1FDE1" box="[488,718,579,603]" italics="true" pageId="48" pageNumber="49">Strong Yellowish Brown</emphasis>
(74), with a dorsal distal semicircle 
<emphasis id="B97EEAC53030872F13E2FDAB0B26FDC1" box="[433,553,611,635]" italics="true" pageId="48" pageNumber="49">Light Yellow</emphasis>
(86). Fe–Pa I– 
<collectionCode id="ED1BAE123030872F1090FDAB0BD2FDC0" box="[707,733,611,634]" pageId="48" pageNumber="49">III</collectionCode>
and Ti I– 
<collectionCode id="ED1BAE123030872F12E7FD4A09C1FD23" box="[180,206,642,665]" pageId="48" pageNumber="49">III</collectionCode>
proximal half background 
<emphasis id="B97EEAC53030872F13B7FD4A0BBEFD20" box="[484,689,642,666]" italics="true" pageId="48" pageNumber="49">Dark Grayish Brown</emphasis>
(62). Ti I–IV distal half background 
<emphasis id="B97EEAC53030872F13F5FD690B74FD03" box="[422,635,673,697]" italics="true" pageId="48" pageNumber="49">Dark Greenish Yellow</emphasis>
(103). Fe–Pa IV and Ti IV proximal half background 
<emphasis id="B97EEAC53030872F1058FD090B9BFD63" box="[523,660,705,729]" italics="true" pageId="48" pageNumber="49">Reddish Black</emphasis>
(24). Tr IV with apophyses and prominent tubercles 
<emphasis id="B97EEAC53030872F1075FD280BA0FD42" box="[550,687,736,760]" italics="true" pageId="48" pageNumber="49">Moderate Red</emphasis>
(15). Fe IV with spines and prominent tubercles 
<taxonomicName id="4C0A4D543030872F1042FCC80B80FCAD" authority="Orange" authorityName="Orange" box="[529,655,768,791]" genus="Deep" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" rank="genus">
<emphasis id="B97EEAC53030872F1042FCC80B80FCAD" box="[529,655,768,791]" italics="true" pageId="48" pageNumber="49">Deep Orange</emphasis>
</taxonomicName>
(51). Pa–Ti IV with spines and tubercles 
<taxonomicName id="4C0A4D543030872F13F6FCD70B68FC8D" authority="Orange Yellow" authorityName="Orange Yellow" box="[421,615,799,823]" genus="Deep" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" rank="genus">
<emphasis id="B97EEAC53030872F13F6FCD70B68FC8D" box="[421,615,799,823]" italics="true" pageId="48" pageNumber="49">Deep Orange Yellow</emphasis>
</taxonomicName>
(69).
</paragraph>
<paragraph id="8BB536D73030872F12D2FC900A7DFE1F" blockId="48.[129,779,856,1256]" lastBlockId="48.[825,1475,146,421]" pageId="48" pageNumber="49">
<emphasis id="B97EEAC53030872F12D2FC900817FCCA" box="[129,280,856,880]" italics="true" pageId="48" pageNumber="49">Male genitalia:</emphasis>
VP slightly divided into a distal half forming a trapezium (widest at the apex) with latero-apical flaps and a proximal half elliptical (1.5× wider than distal part) (
<figureCitation id="13312A523030872F10F8FC5F099CFC74" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[117,1455,150,771]" captionTargetId="figure-162@47.[114,1458,146,774]" captionTargetPageId="47" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="48" pageNumber="49">Fig. 27A, C</figureCitation>
). VP ventral surface entirely covered with microsetae of 
<typeStatus id="54B188753030872F1088FC7E0A06FC74" box="[731,777,950,974]" pageId="48" pageNumber="49">type</typeStatus>
1. All macrosetae cylindrical, inserted on lateral of VP. MS A1–A3 thick and acuminated, on the basal half of the VP (
<figureCitation id="13312A523030872F10B0FC3D09C4FB96" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[117,1455,150,771]" captionTargetId="figure-162@47.[114,1458,146,774]" captionTargetPageId="47" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="48" pageNumber="49">Fig. 27A–C</figureCitation>
). MS B1 short, inserted ventrally, close to A3 (
<figureCitation id="13312A523030872F10F1FBDC0BF7FB96" box="[674,760,1044,1068]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[117,1455,150,771]" captionTargetId="figure-162@47.[114,1458,146,774]" captionTargetPageId="47" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="48" pageNumber="49">Fig. 27C</figureCitation>
). MS C1–C3 thick and acuminated, forming a longitudinal row on the distal half of VP (
<figureCitation id="13312A523030872F13DBFB9B0B0FFBD1" box="[392,512,1107,1131]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[117,1455,150,771]" captionTargetId="figure-162@47.[114,1458,146,774]" captionTargetPageId="47" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="48" pageNumber="49">Fig. 27A–C</figureCitation>
). MS D1 short, closer to C3 than A1 (
<figureCitation id="13312A523030872F135AFBBA088FFB30" box="[265,384,1138,1162]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[117,1455,150,771]" captionTargetId="figure-162@47.[114,1458,146,774]" captionTargetPageId="47" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="48" pageNumber="49">Fig. 27A–C</figureCitation>
). MS E1–E2 very reduced, located on the laterodistal flange of VP—E1 between C1–C2, E2 between C2–C3 (
<figureCitation id="13312A523030872F128CFB79085AFB73" box="[223,341,1201,1225]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[117,1455,150,771]" captionTargetId="figure-162@47.[114,1458,146,774]" captionTargetPageId="47" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="48" pageNumber="49">Fig. 27B–C</figureCitation>
). Glans sac arising from the middle bulge on the podium, not extended as a dorsal process (
<figureCitation id="13312A523030872F10D5FB180BF6FB52" box="[646,761,1232,1256]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[117,1455,150,771]" captionTargetId="figure-162@47.[114,1458,146,774]" captionTargetPageId="47" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="48" pageNumber="49">Fig. 27A–B</figureCitation>
). Stylus and its ventral process axis fused basally, forming a pedestal above the glans (
<figureCitation id="13312A523030872F1671FF790D76FF73" box="[1058,1145,177,201]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[117,1455,150,771]" captionTargetId="figure-162@47.[114,1458,146,774]" captionTargetPageId="47" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="48" pageNumber="49">Fig. 27B</figureCitation>
). Stylus cylindrical, bent at the distal part (forming a plateau) and armed with a set of ventral subapical spines (
<figureCitation id="13312A523030872F11BCFF380D6EFEB2" box="[1007,1121,240,264]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[117,1455,150,771]" captionTargetId="figure-162@47.[114,1458,146,774]" captionTargetPageId="47" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="48" pageNumber="49">Fig. 27A–B</figureCitation>
). Stylus without any expansion or flattening, 
<emphasis id="B97EEAC53030872F11F6FED80AEFFE9D" box="[933,992,272,295]" italics="true" pageId="48" pageNumber="49">in situ</emphasis>
reaching the distal margin of VP (
<figureCitation id="13312A523030872F176FFEC70CBDFE9D" box="[1340,1458,271,295]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[117,1455,150,771]" captionTargetId="figure-162@47.[114,1458,146,774]" captionTargetPageId="47" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="48" pageNumber="49">Fig. 27A–C</figureCitation>
). Ventral process sinuous, as long and thinner than the stylus (
<figureCitation id="13312A523030872F17CFFEE70A70FEDC" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[117,1455,150,771]" captionTargetId="figure-162@47.[114,1458,146,774]" captionTargetPageId="47" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="48" pageNumber="49">Fig. 27A–B</figureCitation>
). Flabellum slightly bent ventrad, hand-shaped, with a comb of straight and acuminated spines on lateral faces (
<figureCitation id="13312A523030872F17CFFEA50A6DFE1F" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[117,1455,150,771]" captionTargetId="figure-162@47.[114,1458,146,774]" captionTargetPageId="47" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="48" pageNumber="49">Fig. 27B</figureCitation>
).
</paragraph>
<paragraph id="8BB536D73030872F116AFE0E0AE8FB8B" blockId="48.[825,1475,454,1073]" pageId="48" pageNumber="49">
<emphasis id="B97EEAC53030872F116AFE0E0C88FE67" box="[825,1415,454,478]" italics="true" pageId="48" pageNumber="49">
Female (
<collectionCode id="ED1BAE123030872F11C1FE0E0AD6FE67" box="[914,985,454,477]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="48" pageNumber="49">MNRJ</collectionCode>
60562) (
<figureCitation id="13312A523030872F1615FE0F0DCDFE64" box="[1094,1218,454,478]" captionStart="Figure 24" captionStartId="44.[130,195,1534,1558]" captionTargetBox="[129,1473,144,1496]" captionTargetId="figure-270@44.[129,1473,144,1496]" captionTargetPageId="44" captionText="Figure 24.Habitus in vivo of Iamarinus pontesi, from Brazil,Paraná, Antonina. ♂ (A–C) and ♀ (D–F). Photographs of Miguel Medrano (A–C) and Adriano Kury (D–E)." figureDoi="http://doi.org/10.5281/zenodo.14802132" httpUri="https://zenodo.org/record/14802132/files/figure.png" pageId="48" pageNumber="49">Figs 24D–E</figureCitation>
, 
<figureCitation id="13312A523030872F1681FE0E0C15FE64" box="[1234,1306,454,478]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="48" pageNumber="49">25D–F</figureCitation>
, 
<figureCitation id="13312A523030872F1779FE0E0C7BFE64" box="[1322,1396,454,478]" captionStart="Figure 28" captionStartId="47.[114,179,1699,1723]" captionTargetBox="[117,1455,887,1657]" captionTargetId="figure-128@47.[114,1458,884,1660]" captionTargetPageId="47" captionText="Figure 28.Iamarinus pontesi, ♀ paratype (MNRJ 60562).A, ocularium, frontal view.B, habitus, dorsal view.C, right trochanter and femur IV, dorsal view. Scale bars: 1 mm (A), 3 mm (B–C)." figureDoi="http://doi.org/10.5281/zenodo.14802142" httpUri="https://zenodo.org/record/14802142/files/figure.png" pageId="48" pageNumber="49">28A–C</figureCitation>
):
</emphasis>
DS, measurements: CW 3.0, CL 2.3, AW 5.3, AL 3.4; Leg I–IV measurements in 
<tableCitation id="C688036C3030872F1654FDCD0D63FDA6" box="[1031,1132,516,541]" captionStart="Table 15" captionStartId="48.[129,184,1679,1703]" captionTargetBox="[129,777,1753,1961]" captionTargetId="graphics-23@48.[129,778,1741,1975]" captionTargetPageId="48" captionText="Table 15. Leg measurements of Iamarinus pontesi, ♀ (MNRJ 60562)" httpUri="http://table.plazi.org/id/DF75665F3030872F12D2F94709C8F978" pageId="48" pageNumber="49" tableUuid="DF75665F3030872F12D2F94709C8F978">Table 15</tableCitation>
; Right/left tarsal (distitarsal) counts: 6(3)/6(3) - 10(3)/10(3) - 7/7 - 7/7. Ocularium longitudinally reduced than males (
<figureCitation id="13312A523030872F16DAFD8B0C0AFDE1" box="[1161,1285,579,603]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="48" pageNumber="49">Figs 25D–E</figureCitation>
, 
<figureCitation id="13312A523030872F1743FD8B0C34FDE1" box="[1296,1339,579,603]" captionStart="Figure 28" captionStartId="47.[114,179,1699,1723]" captionTargetBox="[117,1455,887,1657]" captionTargetId="figure-128@47.[114,1458,884,1660]" captionTargetPageId="47" captionText="Figure 28.Iamarinus pontesi, ♀ paratype (MNRJ 60562).A, ocularium, frontal view.B, habitus, dorsal view.C, right trochanter and femur IV, dorsal view. Scale bars: 1 mm (A), 3 mm (B–C)." figureDoi="http://doi.org/10.5281/zenodo.14802142" httpUri="https://zenodo.org/record/14802142/files/figure.png" pageId="48" pageNumber="49">28A</figureCitation>
). Scutal area 
<collectionCode id="ED1BAE123030872F116AFDAB0A5BFDC0" box="[825,852,611,634]" pageId="48" pageNumber="49">III</collectionCode>
with a paramedian pair of conical spines (larger than males) (
<figureCitation id="13312A523030872F1117FD4A0AC7FD20" box="[836,968,642,666]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="48" pageNumber="49">Figs 25D–E</figureCitation>
, 
<figureCitation id="13312A523030872F1189FD4A0D0CFD20" box="[986,1027,642,666]" captionStart="Figure 28" captionStartId="47.[114,179,1699,1723]" captionTargetBox="[117,1455,887,1657]" captionTargetId="figure-128@47.[114,1458,884,1660]" captionTargetPageId="47" captionText="Figure 28.Iamarinus pontesi, ♀ paratype (MNRJ 60562).A, ocularium, frontal view.B, habitus, dorsal view.C, right trochanter and femur IV, dorsal view. Scale bars: 1 mm (A), 3 mm (B–C)." figureDoi="http://doi.org/10.5281/zenodo.14802142" httpUri="https://zenodo.org/record/14802142/files/figure.png" pageId="48" pageNumber="49">28B</figureCitation>
). Scutal area IV covered with ordinary tubercles (
<figureCitation id="13312A523030872F11FFFD690D23FD03" box="[940,1068,673,697]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="48" pageNumber="49">Figs 25D–E</figureCitation>
, 
<figureCitation id="13312A523030872F1669FD690D6CFD03" box="[1082,1123,673,697]" captionStart="Figure 28" captionStartId="47.[114,179,1699,1723]" captionTargetBox="[117,1455,887,1657]" captionTargetId="figure-128@47.[114,1458,884,1660]" captionTargetPageId="47" captionText="Figure 28.Iamarinus pontesi, ♀ paratype (MNRJ 60562).A, ocularium, frontal view.B, habitus, dorsal view.C, right trochanter and femur IV, dorsal view. Scale bars: 1 mm (A), 3 mm (B–C)." figureDoi="http://doi.org/10.5281/zenodo.14802142" httpUri="https://zenodo.org/record/14802142/files/figure.png" pageId="48" pageNumber="49">28B</figureCitation>
). Cx IV is narrower than males, with the prodorsal distal apophysis reduced to a single spine and a reduced retroventral distal apophysis (
<figureCitation id="13312A523030872F1756FD280C72FD42" box="[1285,1405,736,760]" captionStart="Figure 25" captionStartId="45.[113,178,872,896]" captionTargetBox="[113,1457,144,834]" captionTargetId="figure-706@45.[113,1457,144,834]" captionTargetPageId="45" captionText="Figure 25. Iamarinus pontesi, ♂ holotype (MNRJ 2789). A, habitus, dorsal view. B, same, lateral view. C, same, ventral view. ♀ paratype (MNRJ 60562). D, habitus, dorsal view.E, same, lateral view.F, same, ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802134" httpUri="https://zenodo.org/record/14802134/files/figure.png" pageId="48" pageNumber="49">Figs 25D–F</figureCitation>
, 
<figureCitation id="13312A523030872F17DBFD280CBEFD42" box="[1416,1457,736,760]" captionStart="Figure 28" captionStartId="47.[114,179,1699,1723]" captionTargetBox="[117,1455,887,1657]" captionTargetId="figure-128@47.[114,1458,884,1660]" captionTargetPageId="47" captionText="Figure 28.Iamarinus pontesi, ♀ paratype (MNRJ 60562).A, ocularium, frontal view.B, habitus, dorsal view.C, right trochanter and femur IV, dorsal view. Scale bars: 1 mm (A), 3 mm (B–C)." figureDoi="http://doi.org/10.5281/zenodo.14802142" httpUri="https://zenodo.org/record/14802142/files/figure.png" pageId="48" pageNumber="49">28B</figureCitation>
). Tr IV unarmed on prodorsal and prolateral faces (
<figureCitation id="13312A523030872F1769FD370CBDFCAD" box="[1338,1458,767,791]" captionStart="Figure 28" captionStartId="47.[114,179,1699,1723]" captionTargetBox="[117,1455,887,1657]" captionTargetId="figure-128@47.[114,1458,884,1660]" captionTargetPageId="47" captionText="Figure 28.Iamarinus pontesi, ♀ paratype (MNRJ 60562).A, ocularium, frontal view.B, habitus, dorsal view.C, right trochanter and femur IV, dorsal view. Scale bars: 1 mm (A), 3 mm (B–C)." figureDoi="http://doi.org/10.5281/zenodo.14802142" httpUri="https://zenodo.org/record/14802142/files/figure.png" pageId="48" pageNumber="49">Fig. 28B–C</figureCitation>
). Tr IV retrolateral face with a conical apophysis on proximal and distal thirds (distal largest), and a prominent subconical tubercle on central third (
<figureCitation id="13312A523030872F1615FC950D90FCCF" box="[1094,1183,861,885]" captionStart="Figure 28" captionStartId="47.[114,179,1699,1723]" captionTargetBox="[117,1455,887,1657]" captionTargetId="figure-128@47.[114,1458,884,1660]" captionTargetPageId="47" captionText="Figure 28.Iamarinus pontesi, ♀ paratype (MNRJ 60562).A, ocularium, frontal view.B, habitus, dorsal view.C, right trochanter and femur IV, dorsal view. Scale bars: 1 mm (A), 3 mm (B–C)." figureDoi="http://doi.org/10.5281/zenodo.14802142" httpUri="https://zenodo.org/record/14802142/files/figure.png" pageId="48" pageNumber="49">Fig. 28C</figureCitation>
). Fe IV straight and thinner than males (
<figureCitation id="13312A523030872F1193FCB50D13FC2F" box="[960,1052,893,917]" captionStart="Figure 28" captionStartId="47.[114,179,1699,1723]" captionTargetBox="[117,1455,887,1657]" captionTargetId="figure-128@47.[114,1458,884,1660]" captionTargetPageId="47" captionText="Figure 28.Iamarinus pontesi, ♀ paratype (MNRJ 60562).A, ocularium, frontal view.B, habitus, dorsal view.C, right trochanter and femur IV, dorsal view. Scale bars: 1 mm (A), 3 mm (B–C)." figureDoi="http://doi.org/10.5281/zenodo.14802142" httpUri="https://zenodo.org/record/14802142/files/figure.png" pageId="48" pageNumber="49">Fig. 28C</figureCitation>
). Fe IV dorsal face with two spines on proximal and distal halves (
<figureCitation id="13312A523030872F1630FC540DB0FC0E" box="[1123,1215,924,948]" captionStart="Figure 28" captionStartId="47.[114,179,1699,1723]" captionTargetBox="[117,1455,887,1657]" captionTargetId="figure-128@47.[114,1458,884,1660]" captionTargetPageId="47" captionText="Figure 28.Iamarinus pontesi, ♀ paratype (MNRJ 60562).A, ocularium, frontal view.B, habitus, dorsal view.C, right trochanter and femur IV, dorsal view. Scale bars: 1 mm (A), 3 mm (B–C)." figureDoi="http://doi.org/10.5281/zenodo.14802142" httpUri="https://zenodo.org/record/14802142/files/figure.png" pageId="48" pageNumber="49">Fig. 28C</figureCitation>
). Fe IV retrolateral face with (1) a prominent tubercle on proximal third; (2) an outstanding spine (largest than Fe IV diameter) on central third; and (3) two prominent tubercles (distalmost largest) on distal third (
<figureCitation id="13312A523030872F112EFBD10AD9FB8B" box="[893,982,1049,1073]" captionStart="Figure 28" captionStartId="47.[114,179,1699,1723]" captionTargetBox="[117,1455,887,1657]" captionTargetId="figure-128@47.[114,1458,884,1660]" captionTargetPageId="47" captionText="Figure 28.Iamarinus pontesi, ♀ paratype (MNRJ 60562).A, ocularium, frontal view.B, habitus, dorsal view.C, right trochanter and femur IV, dorsal view. Scale bars: 1 mm (A), 3 mm (B–C)." figureDoi="http://doi.org/10.5281/zenodo.14802142" httpUri="https://zenodo.org/record/14802142/files/figure.png" pageId="48" pageNumber="49">Fig. 28C</figureCitation>
).
</paragraph>
<paragraph id="8BB536D73030872F116AFB9E0C45FA32" blockId="48.[824,1475,1110,1416]" pageId="48" pageNumber="49">
<emphasis id="B97EEAC53030872F116AFB9E0D1BFBD4" box="[825,1044,1110,1134]" italics="true" pageId="48" pageNumber="49">Intraspecific variation:</emphasis>
Some variations among the 
<emphasis id="B97EEAC53030872F1713FB9F0A7FFB37" italics="true" pageId="48" pageNumber="49">major morph males</emphasis>
were detected: (1) Cx IV prodorsal distal apophysis length, with the main branch larger or reduced in comparison to 
<typeStatus id="54B188753030872F17D9FB5D0A65FB76" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
; (2) Fe IV dorsal row of spines with reduced spines; and (3) Fe IV slightly thinner and more extensive than the 
<typeStatus id="54B188753030872F1733FB1B0CB1FB51" box="[1376,1470,1235,1259]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
. Among the 
<emphasis id="B97EEAC53030872F11E5FB3B0D78FAB1" box="[950,1143,1267,1291]" italics="true" pageId="48" pageNumber="49">minor morph males</emphasis>
(
<figureCitation id="13312A523030872F16DAFB3B0DEEFAB1" box="[1161,1249,1267,1291]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[113,1457,143,578]" captionTargetId="figure-830@29.[113,1457,143,578]" captionTargetPageId="29" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="48" pageNumber="49">Fig. 11A</figureCitation>
) (compared to 
<emphasis id="B97EEAC53030872F17DBFB3B0A75FA90" italics="true" pageId="48" pageNumber="49">major morph</emphasis>
): (1) DS narrower than males; (2) Cx IV distal apophysis reduced on prolateral and retrolateral faces; and (3) Fe and Ti IV thinner, with reduced spines. No relevant intraspecific variation among females was detected in the material studied.
</paragraph>
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