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<document id="63B6F6867C85E85ADE2DAA4A054F8A7F" ID-DOI="10.1093/zoolinnean/zlae023" ID-ISSN="0024-4082" ID-Zenodo-Dep="14802079" ID-ZooBank="BB99D24-3973-4413-B127-BDAA83186FA3" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1738154460006" checkinUser="plazi" docAuthor="Carvalho, Rafael N. &amp; Kury, Adriano B." docDate="2025" docId="03A387C130288735110BF93E0D68F923" docLanguage="en" docName="zlae023.pdf" docOrigin="Zoological Journal of the Linnean Society (zlae 023) 203 (1)" docSource="https://doi.org/10.1093/zoolinnean/zlae023" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Iamarinus fenax, gen. et comb. nov." docType="treatment" docVersion="2" lastPageNumber="43" masterDocId="FF9AFFB93000871F1253FFC8090FFFBA" masterDocTitle="Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species" masterLastPageNumber="65" masterPageNumber="1" pageNumber="41" updateTime="1738675863454" updateUser="ExternalLinkService">
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<mods:title id="3C38D1D8167BB44C7022593737CC55D1">Further draining of Discocyrtus to expand Neopachylinae (Opiliones, Gonyleptidae): absorption of taxa and establishment of new genera and species</mods:title>
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<mods:namePart id="2EC534EEAB736CE53D290613DC564085">Carvalho, Rafael N.</mods:namePart>
<mods:affiliation id="4E1216A9CC778794FFEB88ADC2948D81">Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro (UFRJ), Quinta da Boa Vista s / n, São Cristóvão, 20.940 - 040, Rio de Janeiro, Brazil &amp; Departamento de Zoologia, Instituto de Biologia Roberto de Alcantara Gomes, Universidade do Estado do Rio de Janeiro (UERJ), Rua São Francisco Xavier 524, Maracanã, 20.550 - 900, Rio de Janeiro, Brazil</mods:affiliation>
<mods:nameIdentifier id="CE5A22F9972EF08986FD5EF7449102B2" type="email">rafaelcarvalhobio@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="C6F5CDA93DBC8712F96E31259C7F066A">Kury, Adriano B.</mods:namePart>
<mods:affiliation id="9A40A065374C6F0292E4B126993495B7">Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro (UFRJ), Quinta da Boa Vista s / n, São Cristóvão, 20.940 - 040, Rio de Janeiro, Brazil</mods:affiliation>
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<mods:date id="AF5D3CD8C9DA6D5E3BB6A2D234DFE38A">2025</mods:date>
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<subSubSection id="C310655C30288737110BF93E0DB9F8F5" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="8BB536D730288737110BF93E0DB9F8F5" blockId="40.[851,1449,1782,1871]" pageId="40" pageNumber="41">
<emphasis id="B97EEAC530288737110BF93E0DB9F8F5" bold="true" pageId="40" pageNumber="41">
<taxonomicName id="4C0A4D5430288737110BF93E0C6FF895" authority="(Kury, Pinto-da-Rocha &amp; Carvalho, 2018)" baseAuthorityName="Kury, Pinto-da-Rocha &amp; Carvalho" baseAuthorityYear="2018" class="Arachnida" family="Gonyleptidae" genus="Iamarinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="fenax" status="gen. et comb. nov.">
<emphasis id="B97EEAC530288737110BF93E0D1EF8AA" bold="true" box="[856,1041,1782,1808]" italics="true" pageId="40" pageNumber="41">Iamarinus fenax</emphasis>
(Kury, Pinto-da-Rocha &amp; Carvalho
<emphasis id="B97EEAC53028873717DEF93E0CACF8AA" bold="true" box="[1421,1443,1782,1808]" italics="true" pageId="40" pageNumber="41">in</emphasis>
<bibRefCitation id="EF9B4B26302887371100F8DE0C57F895" author="Kury AB &amp; Pinto-da-Rocha R &amp; Gruber J" box="[851,1368,1813,1840]" pageId="40" pageNumber="41" pagination="355 - 69" refId="ref53736" refString="Kury AB, Pinto-da-Rocha R, Gruber J et al. Clarification on three species of Discocyrtus Holmberg, 1878 with convoluted taxonomic stories (Opiliones, Gonyleptidae, Pachylinae). Journal of Arachnology 2018 b; 46: 355 - 69. https: // doi. org / 10.1636 / joa-s- 17 - 069.1" type="journal article" year="2018">Kury, Pinto-da-Rocha, Gruber &amp; Carvalho, 2018</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A24D57BE302887371737F8DE0DB9F8F5" pageId="40" pageNumber="41" rank="species">gen. et comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C310655C30288737166BF8AC0DCCF8C4" box="[1080,1219,1892,1918]" pageId="40" pageNumber="41" type="description">
<paragraph id="8BB536D730288737166BF8AC0DCCF8C4" blockId="40.[825,1475,1892,1986]" box="[1080,1219,1892,1918]" pageId="40" pageNumber="41">
<emphasis id="B97EEAC530288737166BF8AC0DCCF8C4" bold="true" box="[1080,1219,1892,1918]" pageId="40" pageNumber="41">
(
<figureCitation id="13312A52302887371610F8AC0DB6F8C4" box="[1091,1209,1892,1918]" captionStart-0="Figure 22" captionStart-1="15" captionStartId-0="42.[126,191,1272,1296]" captionStartId-1="5.[113,139,194,218]" captionTargetBox-0="[129,1473,144,1233]" captionTargetBox-1="[116,1453,147,816]" captionTargetId-0="figure-388@42.[129,1473,144,1233]" captionTargetId-1="figure-515@43.[113,1457,144,820]" captionTargetPageId-0="42" captionTargetPageId-1="43" captionText-0="Figure 22. Iamarinus fenax, ♀ (MHNCI 6305). A, habitus, dorsal view.B, detail of the mesotergum and free tergites I–III, dorsal view. C, habitus, ventral view.D, same, lateral view.Scale bars = 5 mm." captionText-1="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" figureDoi-0="http://doi.org/10.5281/zenodo.14802128" httpUri-0="https://zenodo.org/record/14802128/files/figure.png" pageId="40" pageNumber="41">Figs 22–23</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C310655C30288737116AF8430C2DF878" pageId="40" pageNumber="41" type="reference_group">
<paragraph id="8BB536D730288737116AF8430C2DF878" blockId="40.[825,1475,1892,1986]" pageId="40" pageNumber="41">
<treatmentCitationGroup id="AB1A11F930288737116AF8430C2DF878" pageId="40" pageNumber="41">
•
<taxonomicName id="4C0A4D5430288737111FF8440D38F819" ID-CoL="7VS97" authority="Kury" authorityName="Kury" box="[844,1079,1931,1955]" class="Arachnida" family="Gonyleptidae" genus="Discocyrtus" kingdom="Animalia" order="Opiliones" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="fenax">
<emphasis id="B97EEAC530288737111FF8440AF5F819" box="[844,1018,1931,1955]" italics="true" pageId="40" pageNumber="41">Discocyrtus fenax</emphasis>
Kury
</taxonomicName>
, Pinto-da-Rocha &amp; Carvalho
<emphasis id="B97EEAC530288737173CF8440C8DF819" box="[1391,1410,1932,1955]" italics="true" pageId="40" pageNumber="41">in</emphasis>
<treatmentCitation id="0AAB10C63028873717D9F8430C12F878" author="Kury AB &amp; Pinto-da-Rocha R &amp; Gruber J" page="367" pageId="40" pageNumber="41" year="2018">
<bibRefCitation id="EF9B4B263028873717D9F8430C12F878" author="Kury AB &amp; Pinto-da-Rocha R &amp; Gruber J" pageId="40" pageNumber="41" pagination="355 - 69" refId="ref53736" refString="Kury AB, Pinto-da-Rocha R, Gruber J et al. Clarification on three species of Discocyrtus Holmberg, 1878 with convoluted taxonomic stories (Opiliones, Gonyleptidae, Pachylinae). Journal of Arachnology 2018 b; 46: 355 - 69. https: // doi. org / 10.1636 / joa-s- 17 - 069.1" type="journal article" year="2018">Kury, Pinto-da-Rocha, Gruber &amp; Carvalho 2018: 367</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C310655C3029873613D4FF5908C0FE30" pageId="41" pageNumber="42" type="materials_examined">
<paragraph id="8BB536D73029873613D4FF5908EAFF10" blockId="41.[113,764,144,300]" box="[391,485,144,170]" pageId="41" pageNumber="42">
<emphasis id="B97EEAC53029873613D4FF5908EAFF10" box="[391,485,144,170]" italics="true" pageId="41" pageNumber="42">
<typeStatus id="54B188753029873613D4FF5908BAFF10" box="[391,437,145,170]" pageId="41" pageNumber="42">Type</typeStatus>
data
</emphasis>
</paragraph>
<paragraph id="8BB536D7302987361222FF7F083EFE96" blockId="41.[113,764,144,300]" pageId="41" pageNumber="42">
<emphasis id="B97EEAC5302987361222FF7F082AFF74" box="[113,293,182,206]" italics="true" pageId="41" pageNumber="42">
<taxonomicName id="4C0A4D54302987361222FF7F082EFF74" box="[113,289,182,206]" class="Arachnida" family="Gonyleptidae" genus="Discocyrtus" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="fenax">Discocyrtus fenax</taxonomicName>
:
</emphasis>
<materialsCitation id="3B623C8A30298736137DFF7109A7FEB7" collectionCode="MZSP, BR" country="Brazil" location="Itajai" municipality="Itajai" pageId="41" pageNumber="42" specimenCode="MZSP 18158, BR470" specimenCount="1" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="holotype">
<specimenCount id="9D0CFD5E30298736137DFF710847FF77" box="[302,328,185,205]" count="1" pageId="41" pageNumber="42" type="male">♂</specimenCount>
<typeStatus id="54B1887530298736131EFF7E08A6FF74" box="[333,425,182,206]" pageId="41" pageNumber="42" type="holotype">holotype</typeStatus>
(
<specimenCode id="DBAC9EAC3029873613EBFF7E0B46FF74" box="[440,585,182,206]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="42" type="Museum">MZSP 18158</specimenCode>
, examined) from
<collectingCountry id="F31D7647302987361220FF1E09C1FF57" box="[115,206,214,237]" name="Brazil" pageId="41" pageNumber="42">BRAZIL</collectingCountry>
, state of
<collectingRegion id="49CEF835302987361365FF1E08DCFF54" box="[310,467,214,238]" country="Brazil" name="Santa Catarina" pageId="41" pageNumber="42">Santa Catarina</collectingRegion>
, close to road
<specimenCode id="DBAC9EAC30298736102BFF1E0BCEFF54" box="[632,705,214,238]" collectionCode="BR" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="41" pageNumber="42">BR470</specimenCode>
, near
<collectingMunicipality id="6BD1ACAD302987361222FF3D09A7FEB7" box="[113,168,245,269]" pageId="41" pageNumber="42">Itajaí</collectingMunicipality>
</materialsCitation>
;
<materialsCitation id="3B623C8A3029873612E0FF300822FE96" collectionCode="SMF" country="Brazil" location="Joinville" municipality="Joinville" pageId="41" pageNumber="42" specimenCode="RI 812" specimenCount="1" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D0CFD5E3029873612E0FF3009C2FEB6" box="[179,205,248,268]" count="1" pageId="41" pageNumber="42" type="male">♂</specimenCount>
<typeStatus id="54B18875302987361287FF3D0822FEB7" box="[212,301,245,269]" pageId="41" pageNumber="42" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED1BAE12302987361368FF3D087EFEB7" box="[315,369,245,269]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="41" pageNumber="42">SMF</collectionCode>
<specimenCode id="DBAC9EAC30298736132BFF3D08CCFEB7" box="[376,451,245,269]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="41" pageNumber="42">RI 812</specimenCode>
) from
<collectingCountry id="F31D7647302987361059FF3D0B6AFEB6" box="[522,613,245,268]" name="Brazil" pageId="41" pageNumber="42">BRAZIL</collectingCountry>
, state of
<collectingRegion id="49CEF835302987361091FF3D09C2FE96" country="Brazil" name="Santa Catarina" pageId="41" pageNumber="42">Santa Catarina</collectingRegion>
,
<collectingMunicipality id="6BD1ACAD302987361285FEDC0822FE96" box="[214,301,276,300]" pageId="41" pageNumber="42">Joinville</collectingMunicipality>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB536D73029873613C2FE8408D5FEDC" blockId="41.[113,474,332,394]" box="[401,474,332,358]" pageId="41" pageNumber="42">
<emphasis id="B97EEAC53029873613C2FE8408D5FEDC" box="[401,474,332,358]" italics="true" pageId="41" pageNumber="42">Records</emphasis>
</paragraph>
<paragraph id="8BB536D7302987361222FEBA08C0FE30" blockId="41.[113,474,332,394]" box="[113,463,370,394]" pageId="41" pageNumber="42">Without further literature records.</paragraph>
</subSubSection>
<subSubSection id="C310655C3029873613D4FE630830FC98" pageId="41" pageNumber="42" type="diagnosis">
<paragraph id="8BB536D73029873613D4FE6308EBFE7E" blockId="41.[113,763,427,802]" box="[391,484,427,452]" pageId="41" pageNumber="42">
<emphasis id="B97EEAC53029873613D4FE6308EBFE7E" box="[391,484,427,452]" italics="true" pageId="41" pageNumber="42">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BB536D7302987361222FE190830FC98" blockId="41.[113,763,427,802]" pageId="41" pageNumber="42">
<taxonomicName id="4C0A4D54302987361222FE19081EFE52" ID-CoL="C8QGN" baseAuthorityName="Kury, Pinto-da-Rocha &amp; Carvalho" baseAuthorityYear="2018" box="[113,273,464,488]" class="Arachnida" family="Gonyleptidae" genus="Iamarinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="fenax">
<emphasis id="B97EEAC5302987361222FE19081EFE52" box="[113,273,464,488]" italics="true" pageId="41" pageNumber="42">Iamarinus fenax</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C0A4D54302987361079FE190B8CFE52" ID-CoL="C8QLN" authorityName="Carvalho &amp; Kury" authorityYear="2025" box="[554,643,465,488]" class="Arachnida" family="Gonyleptidae" genus="Iamarinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="pontesi">
<emphasis id="B97EEAC5302987361079FE190B8CFE52" box="[554,643,465,488]" italics="true" pageId="41" pageNumber="42">I. pontesi</emphasis>
</taxonomicName>
due to: (1) Scutal area I with two pairs of paramedian conspicuous tubercles (
<bibRefCitation id="EF9B4B263029873612F8FDC7086AFD9D" author="Kury AB &amp; Pinto-da-Rocha R &amp; Gruber J" box="[171,357,527,551]" pageId="41" pageNumber="42" pagination="355 - 69" refId="ref53736" refString="Kury AB, Pinto-da-Rocha R, Gruber J et al. Clarification on three species of Discocyrtus Holmberg, 1878 with convoluted taxonomic stories (Opiliones, Gonyleptidae, Pachylinae). Journal of Arachnology 2018 b; 46: 355 - 69. https: // doi. org / 10.1636 / joa-s- 17 - 069.1" type="journal article" year="2018">
Kury
<emphasis id="B97EEAC53029873612BBFDD80814FD9D" box="[232,283,527,551]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 10A); (2) Cx IV prodorsal distal apophysis with distal portion forming a 90° angle in relation to the longitudinal axis (
<bibRefCitation id="EF9B4B26302987361309FD860B1DFDDC" author="Kury AB &amp; Pinto-da-Rocha R &amp; Gruber J" box="[346,530,590,614]" pageId="41" pageNumber="42" pagination="355 - 69" refId="ref53736" refString="Kury AB, Pinto-da-Rocha R, Gruber J et al. Clarification on three species of Discocyrtus Holmberg, 1878 with convoluted taxonomic stories (Opiliones, Gonyleptidae, Pachylinae). Journal of Arachnology 2018 b; 46: 355 - 69. https: // doi. org / 10.1636 / joa-s- 17 - 069.1" type="journal article" year="2018">
Kury
<emphasis id="B97EEAC53029873613C4FD8608C6FDDC" box="[407,457,590,614]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 10A); (3) Cx IV prodorsal distal apophysis elongated, bent centrad and distally sinuous (
<bibRefCitation id="EF9B4B26302987361283FD450870FD1E" author="Kury AB &amp; Pinto-da-Rocha R &amp; Gruber J" box="[208,383,652,677]" pageId="41" pageNumber="42" pagination="355 - 69" refId="ref53736" refString="Kury AB, Pinto-da-Rocha R, Gruber J et al. Clarification on three species of Discocyrtus Holmberg, 1878 with convoluted taxonomic stories (Opiliones, Gonyleptidae, Pachylinae). Journal of Arachnology 2018 b; 46: 355 - 69. https: // doi. org / 10.1636 / joa-s- 17 - 069.1" type="journal article" year="2018">
Kury
<emphasis id="B97EEAC5302987361359FD450836FD1E" box="[266,313,652,676]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 10A); (4) Cx IV prodorsal distal apophysis without a secondary branch (
<bibRefCitation id="EF9B4B26302987361045FD640BC3FD7E" author="Kury AB &amp; Pinto-da-Rocha R &amp; Gruber J" box="[534,716,684,708]" pageId="41" pageNumber="42" pagination="355 - 69" refId="ref53736" refString="Kury AB, Pinto-da-Rocha R, Gruber J et al. Clarification on three species of Discocyrtus Holmberg, 1878 with convoluted taxonomic stories (Opiliones, Gonyleptidae, Pachylinae). Journal of Arachnology 2018 b; 46: 355 - 69. https: // doi. org / 10.1636 / joa-s- 17 - 069.1" type="journal article" year="2018">
Kury
<emphasis id="B97EEAC5302987361001FD640B8BFD7E" box="[594,644,684,708]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 10D); (5) Fe IV sub-straight (
<bibRefCitation id="EF9B4B263029873613FDFD030B6EFD59" author="Kury AB &amp; Pinto-da-Rocha R &amp; Gruber J" box="[430,609,715,739]" pageId="41" pageNumber="42" pagination="355 - 69" refId="ref53736" refString="Kury AB, Pinto-da-Rocha R, Gruber J et al. Clarification on three species of Discocyrtus Holmberg, 1878 with convoluted taxonomic stories (Opiliones, Gonyleptidae, Pachylinae). Journal of Arachnology 2018 b; 46: 355 - 69. https: // doi. org / 10.1636 / joa-s- 17 - 069.1" type="journal article" year="2018">
Kury
<emphasis id="B97EEAC53029873613BAFD040B15FD59" box="[489,538,715,739]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 10E); (6) Ti IV with an apical regular spine on retroventral face (
<bibRefCitation id="EF9B4B263029873610FDFD2309DFFC98" author="Kury AB &amp; Pinto-da-Rocha R &amp; Gruber J" pageId="41" pageNumber="42" pagination="355 - 69" refId="ref53736" refString="Kury AB, Pinto-da-Rocha R, Gruber J et al. Clarification on three species of Discocyrtus Holmberg, 1878 with convoluted taxonomic stories (Opiliones, Gonyleptidae, Pachylinae). Journal of Arachnology 2018 b; 46: 355 - 69. https: // doi. org / 10.1636 / joa-s- 17 - 069.1" type="journal article" year="2018">
Kury
<emphasis id="B97EEAC53029873610BAFD230985FC98" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 10H).
</paragraph>
</subSubSection>
<subSubSection id="C310655C3029873611BBFF590CBCFED1" pageId="41" pageNumber="42" type="materials_examined">
<paragraph id="8BB536D73029873611BBFF590DFAFF10" blockId="41.[810,1460,144,363]" box="[1000,1269,144,170]" pageId="41" pageNumber="42">
<emphasis id="B97EEAC53029873611BBFF590DFAFF10" box="[1000,1269,144,170]" italics="true" pageId="41" pageNumber="42">Non-type material examined</emphasis>
</paragraph>
<paragraph id="8BB536D7302987361179FF7F0CBCFED1" blockId="41.[810,1460,144,363]" pageId="41" pageNumber="42">
<materialsCitation id="3B623C8A302987361179FF7F0C02FF74" box="[810,1293,182,206]" collectorName="Blumenau" country="Brazil" location="Santa Catarina" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Santa Catarina">
<collectingCountry id="F31D7647302987361179FF7F0A87FF74" box="[810,904,183,206]" name="Brazil" pageId="41" pageNumber="42">BRAZIL</collectingCountry>
: state of
<collectingRegion id="49CEF8353029873611ABFF7E0D96FF74" box="[1016,1177,182,206]" country="Brazil" name="Santa Catarina" pageId="41" pageNumber="42">Santa Catarina</collectingRegion>
:
<emphasis id="B97EEAC53029873616F9FF7E0C02FF74" box="[1194,1293,182,206]" italics="true" pageId="41" pageNumber="42">
<collectorName id="26FF53013029873616F9FF7E0C02FF74" box="[1194,1293,182,206]" pageId="41" pageNumber="42">Blumenau</collectorName>
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B623C8A302987361771FF7E0D9EFF54" collectingDate="1987-03" collectionCode="MZSP" collectorName="Gofferje, C. N." country="Brazil" pageId="41" pageNumber="42" specimenCode="MZSP 36268" specimenCount="5" specimenCount-male="5">
<specimenCount id="9D0CFD5E302987361771FF7E0C58FF77" box="[1314,1367,182,206]" count="5" pageId="41" pageNumber="42" type="male">5 ♂</specimenCount>
(
<specimenCode id="DBAC9EAC302987361738FF7E0A7FFF54" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="42" type="Museum">MZSP 36268</specimenCode>
),
<date id="FFB410173029873611D1FF1E0ADEFF54" box="[898,977,214,238]" pageId="41" pageNumber="42" value="1987-03">
<collectingDate id="EFF0E9FF3029873611D1FF1E0ADEFF54" box="[898,977,214,238]" pageId="41" pageNumber="42" value="1987-03">iii.1987</collectingDate>
</date>
,
<collectorName id="26FF5301302987361189FF1E0D63FF54" box="[986,1132,214,238]" pageId="41" pageNumber="42">Gofferjé, C.N.</collectorName>
<emphasis id="B97EEAC5302987361621FF1E0D9EFF54" box="[1138,1169,214,238]" italics="true" pageId="41" pageNumber="42">leg.</emphasis>
</materialsCitation>
;
<materialsCitation id="3B623C8A3029873616CEFF1E0CA0FF57" box="[1181,1455,213,238]" collectionCode="MHNCI" country="Brazil" pageId="41" pageNumber="42" specimenCode="MHNCI 6305" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
<specimenCount id="9D0CFD5E3029873616CEFF1E0DC5FF57" box="[1181,1226,214,238]" count="1" pageId="41" pageNumber="42" type="male">1 ♂</specimenCount>
<specimenCount id="9D0CFD5E302987361683FF1E0DF2FF57" box="[1232,1277,214,238]" count="3" pageId="41" pageNumber="42" type="female">3 ♀</specimenCount>
(
<specimenCode id="DBAC9EAC302987361759FF1E0CA8FF57" box="[1290,1447,213,238]" collectionCode="MHNCI" country="0" httpUri="http://grbio.org/cool/ej4f-8m62" name="Museu de Historia Natural Capao de Imbuia (Brazil)" pageId="41" pageNumber="42">MHNCI 6305</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B623C8A302987361178FF3D0C4BFEB7" box="[811,1348,245,269]" collectingDate="1988-01" collectionCode="MZSP" collectorName="Gofferje, C. N." country="Brazil" pageId="41" pageNumber="42" specimenCode="MZSP 36294" specimenCount="6" specimenCount-female="4" specimenCount-male="2">
<specimenCount id="9D0CFD5E302987361178FF3D0A57FEB6" box="[811,856,245,269]" count="2" pageId="41" pageNumber="42" type="male">2 ♂</specimenCount>
<specimenCount id="9D0CFD5E30298736110DFF3D0A82FEB6" box="[862,909,245,269]" count="4" pageId="41" pageNumber="42" type="female">4 ♀</specimenCount>
(
<specimenCode id="DBAC9EAC3029873611C9FF3D0D21FEB7" box="[922,1070,245,269]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="42" type="Museum">MZSP 36294</specimenCode>
),
<date id="FFB41017302987361613FF3D0D8EFEB7" box="[1088,1153,245,269]" pageId="41" pageNumber="42" value="1988-01">
<collectingDate id="EFF0E9FF302987361613FF3D0D8EFEB7" box="[1088,1153,245,269]" pageId="41" pageNumber="42" value="1988-01">i.1988</collectingDate>
</date>
,
<collectorName id="26FF53013029873616DFFF3D0C11FEB7" box="[1164,1310,245,269]" pageId="41" pageNumber="42">Gofferjé, C.N.</collectorName>
<emphasis id="B97EEAC5302987361776FF3D0C4BFEB7" box="[1317,1348,245,269]" italics="true" pageId="41" pageNumber="42">leg.</emphasis>
</materialsCitation>
;
<materialsCitation id="3B623C8A302987361702FF3D0C97FE96" collectingDate="1987-03" collectionCode="MHNCI" collectorName="Gofferje, C. N." country="Brazil" location="Centro" pageId="41" pageNumber="42" specimenCode="MHNCI 6348" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="9D0CFD5E302987361702FF3D0C8FFEB6" box="[1361,1408,245,269]" count="2" pageId="41" pageNumber="42" type="male">2 ♂</specimenCount>
<specimenCount id="9D0CFD5E3029873617D6FF3D0CBBFEB6" box="[1413,1460,245,269]" count="1" pageId="41" pageNumber="42" type="female">1 ♀</specimenCount>
(
<specimenCode id="DBAC9EAC302987361162FEDC0ADFFE96" box="[817,976,276,300]" collectionCode="MHNCI" country="0" httpUri="http://grbio.org/cool/ej4f-8m62" name="Museu de Historia Natural Capao de Imbuia (Brazil)" pageId="41" pageNumber="42">MHNCI 6348</specimenCode>
),
<location id="8ED5600C3029873611B0FEDC0D22FE96" LSID="urn:lsid:plazi:treatment:03A387C130288735110BF93E0D68F923:8ED5600C3029873611B0FEDC0D22FE96" box="[995,1069,276,300]" country="Brazil" name="Centro" pageId="41" pageNumber="42">Centro</location>
,
<date id="FFB4101730298736166BFEDC0D88FE96" box="[1080,1159,276,300]" pageId="41" pageNumber="42" value="1987-03">
<collectingDate id="EFF0E9FF30298736166BFEDC0D88FE96" box="[1080,1159,276,300]" pageId="41" pageNumber="42" value="1987-03">iii.1987</collectingDate>
</date>
,
<collectorName id="26FF53013029873616C0FEDC0C22FE96" box="[1171,1325,276,300]" pageId="41" pageNumber="42">Gofferjé, C. N.</collectorName>
<emphasis id="B97EEAC5302987361765FEDC0C97FE96" box="[1334,1432,276,300]" italics="true" pageId="41" pageNumber="42">leg. Penha</emphasis>
</materialsCitation>
:
<materialsCitation id="3B623C8A3029873617F5FEDC0C95FEF6" collectingDate="1989-01" collectionCode="MHNCI" collectorName="Gofferje, C. N." country="Brazil" location="Armacao" pageId="41" pageNumber="42" specimenCode="MHNCI 6822" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
<specimenCount id="9D0CFD5E3029873617F5FEDC0A49FEF1" count="1" pageId="41" pageNumber="42" type="male">1 ♂</specimenCount>
<specimenCount id="9D0CFD5E302987361119FEFC0A7AFEF1" box="[842,885,308,332]" count="3" pageId="41" pageNumber="42" type="female">3 ♀</specimenCount>
(
<specimenCode id="DBAC9EAC3029873611D1FEFC0D10FEF6" box="[898,1055,308,332]" collectionCode="MHNCI" country="0" httpUri="http://grbio.org/cool/ej4f-8m62" name="Museu de Historia Natural Capao de Imbuia (Brazil)" pageId="41" pageNumber="42">MHNCI 6822</specimenCode>
),
<location id="8ED5600C30298736167CFEFC0D82FEF6" LSID="urn:lsid:plazi:treatment:03A387C130288735110BF93E0D68F923:8ED5600C30298736167CFEFC0D82FEF6" box="[1071,1165,308,332]" country="Brazil" name="Armacao" pageId="41" pageNumber="42">Armação</location>
,
<date id="FFB410173029873616C5FEFC0DD8FEF6" box="[1174,1239,308,332]" pageId="41" pageNumber="42" value="1989-01">
<collectingDate id="EFF0E9FF3029873616C5FEFC0DD8FEF6" box="[1174,1239,308,332]" pageId="41" pageNumber="42" value="1989-01">i.1989</collectingDate>
</date>
,
<collectorName id="26FF53013029873616B2FEFC0C79FEF1" box="[1249,1398,308,332]" pageId="41" pageNumber="42">Gofferjé, C. N.</collectorName>
<emphasis id="B97EEAC5302987361728FEFC0C95FEF6" box="[1403,1434,308,332]" italics="true" pageId="41" pageNumber="42">leg.</emphasis>
</materialsCitation>
;
<materialsCitation id="3B623C8A3029873617F5FEFC0CBCFED1" collectingDate="1969-12" collectionCode="MZSP" collectorName="Gofferje, C. N." country="Brazil" location="Armacao" pageId="41" pageNumber="42" specimenCode="MZSP 36288" specimenCount="22" specimenCount-female="14" specimenCount-male="8">
<specimenCount id="9D0CFD5E3029873617F5FEFC0A4BFED0" count="8" pageId="41" pageNumber="42" type="male">8 ♂</specimenCount>
<specimenCount id="9D0CFD5E30298736111AFE9B0A8EFED0" box="[841,897,339,363]" count="14" pageId="41" pageNumber="42" type="female">14 ♀</specimenCount>
2 juv (
<specimenCode id="DBAC9EAC302987361190FE9B0D5AFED1" box="[963,1109,339,363]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="42" type="Museum">MZSP 36288</specimenCode>
), idem,
<date id="FFB410173029873616F7FE9B0DF8FED1" box="[1188,1271,339,363]" pageId="41" pageNumber="42" value="1969-12">
<collectingDate id="EFF0E9FF3029873616F7FE9B0DF8FED1" box="[1188,1271,339,363]" pageId="41" pageNumber="42" value="1969-12">xii.1969</collectingDate>
</date>
,
<collectorName id="26FF5301302987361753FE9B0C9FFED1" box="[1280,1424,339,363]" pageId="41" pageNumber="42">Gofferjé, C.N.</collectorName>
<emphasis id="B97EEAC53029873617C0FE9B0CBCFED1" box="[1427,1459,339,363]" italics="true" pageId="41" pageNumber="42">leg.</emphasis>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C310655C3029873611FAFE420AD6FE52" pageId="41" pageNumber="42" type="distribution">
<paragraph id="8BB536D73029873611FAFE420C3BFE1E" blockId="41.[810,1459,394,488]" box="[937,1332,394,421]" pageId="41" pageNumber="42">
<emphasis id="B97EEAC53029873611FAFE420C3BFE1E" box="[937,1332,394,421]" italics="true" pageId="41" pageNumber="42">Distribution (new records with an asterisk)</emphasis>
</paragraph>
<paragraph id="8BB536D7302987361179FE790AD6FE52" blockId="41.[810,1459,394,488]" pageId="41" pageNumber="42">
<collectingCountry id="F31D7647302987361179FE790A87FE72" box="[810,904,433,456]" name="Brazil" pageId="41" pageNumber="42">BRAZIL</collectingCountry>
: state of
<collectingRegion id="49CEF8353029873611BFFE790D86FE73" box="[1004,1161,433,457]" country="Brazil" name="Santa Catarina" pageId="41" pageNumber="42">Santa Catarina</collectingRegion>
: Blumenau*, Itajaí, Joinville, Penha* (
<figureCitation id="13312A523029873611D0FE180AC7FE52" box="[899,968,464,488]" captionStart="Figure 21" captionStartId="41.[113,178,1926,1950]" captionTargetBox="[118,1456,908,1896]" captionTargetId="figure-344@41.[115,1459,905,1899]" captionTargetPageId="41" captionText="Figure 21. South Brazil,showing distribution of Iamarinus species. Shaded areas in the background are Morrone’s regionalization of the Neotropics (‘provinces’; Morrone et al. 2022): Araucaria Forest province (in light green) and Atlantic province (in dark green)." figureDoi="http://doi.org/10.5281/zenodo.14802126" httpUri="https://zenodo.org/record/14802126/files/figure.png" pageId="41" pageNumber="42">Fig. 21</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C310655C302987351640FDCE0D68F923" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
<paragraph id="8BB536D7302987361640FDCE0DC5FD9A" blockId="41.[810,1320,518,581]" box="[1043,1226,518,544]" pageId="41" pageNumber="42">
<emphasis id="B97EEAC5302987361640FDCE0DC5FD9A" box="[1043,1226,518,544]" italics="true" pageId="41" pageNumber="42">Male’s redescription</emphasis>
</paragraph>
<paragraph id="8BB536D7302987361179FDE40C27FDFE" blockId="41.[810,1320,518,581]" box="[810,1320,556,581]" pageId="41" pageNumber="42">
See the original description in
<bibRefCitation id="EF9B4B26302987361633FDE50C2CFDFE" author="Kury AB &amp; Pinto-da-Rocha R &amp; Gruber J" box="[1120,1315,556,581]" pageId="41" pageNumber="42" pagination="355 - 69" refId="ref53736" refString="Kury AB, Pinto-da-Rocha R, Gruber J et al. Clarification on three species of Discocyrtus Holmberg, 1878 with convoluted taxonomic stories (Opiliones, Gonyleptidae, Pachylinae). Journal of Arachnology 2018 b; 46: 355 - 69. https: // doi. org / 10.1636 / joa-s- 17 - 069.1" type="journal article" year="2018">
Kury
<emphasis id="B97EEAC53029873616CAFDE50DC7FDFE" box="[1177,1224,556,580]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
(2018b)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BB536D730298736164BFDAC0DCAFDC4" blockId="41.[809,1459,612,800]" box="[1048,1221,612,638]" pageId="41" pageNumber="42">
<emphasis id="B97EEAC530298736164BFDAC0DCAFDC4" box="[1048,1221,612,638]" italics="true" pageId="41" pageNumber="42">Female description</emphasis>
</paragraph>
<paragraph id="8BB536D7302987351179FD420D4AFA11" blockId="41.[809,1459,612,800]" lastBlockId="42.[825,1475,1364,1451]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
<emphasis id="B97EEAC5302987361179FD420AD1FD18" bold="true" box="[810,990,650,674]" pageId="41" pageNumber="42">
(
<collectionCode id="ED1BAE12302987361162FD420A9DFD18" box="[817,914,650,674]" country="0" httpUri="http://grbio.org/cool/ej4f-8m62" name="Museu de Historia Natural Capao de Imbuia (Brazil)" pageId="41" pageNumber="42">MHNCI</collectionCode>
6305)
</emphasis>
(
<figureCitation id="13312A523029873611A1FD420D67FD18" box="[1010,1128,650,674]" captionStart-0="Figure 22" captionStart-1="15" captionStartId-0="42.[126,191,1272,1296]" captionStartId-1="5.[113,139,194,218]" captionTargetBox-0="[129,1473,144,1233]" captionTargetBox-1="[116,1453,147,816]" captionTargetId-0="figure-388@42.[129,1473,144,1233]" captionTargetId-1="figure-515@43.[113,1457,144,820]" captionTargetPageId-0="42" captionTargetPageId-1="43" captionText-0="Figure 22. Iamarinus fenax, ♀ (MHNCI 6305). A, habitus, dorsal view.B, detail of the mesotergum and free tergites I–III, dorsal view. C, habitus, ventral view.D, same, lateral view.Scale bars = 5 mm." captionText-1="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" figureDoi-0="http://doi.org/10.5281/zenodo.14802128" httpUri-0="https://zenodo.org/record/14802128/files/figure.png" pageId="41" pageNumber="42">Figs 22–23</figureCitation>
). DS, measurements: CW 2.9, CL 2.2, AW 5.3, AL 3.3; Leg I–IV measurements in
<tableCitation id="C688036C302987361701FD620CA4FD78" box="[1362,1451,682,706]" captionStart="Table 13" captionStartId="43.[113,168,964,988]" captionTargetBox="[113,762,1010,1218]" captionTargetId="graphics-571@43.[113,762,998,1232]" captionTargetPageId="43" captionText="Table 13. Leg measurements of Iamarinus fenax, ♀ (MHNCI 6305)" httpUri="http://table.plazi.org/id/DF75665F302B87341222FC0C0BF8FC66" pageId="41" pageNumber="42" tableUuid="DF75665F302B87341222FC0C0BF8FC66">Table 13</tableCitation>
; Right/left tarsal (distitarsal) counts: 6(3)/6(3) - 9(3)/9(3) - 7/7 - 7/7. DS lambda (
<figureCitation id="13312A52302987361645FD200D7EFCBA" box="[1046,1137,744,768]" captionStart="Figure 22" captionStartId="42.[126,191,1272,1296]" captionTargetBox="[129,1473,144,1233]" captionTargetId="figure-388@42.[129,1473,144,1233]" captionTargetPageId="42" captionText="Figure 22. Iamarinus fenax, ♀ (MHNCI 6305). A, habitus, dorsal view.B, detail of the mesotergum and free tergites I–III, dorsal view. C, habitus, ventral view.D, same, lateral view.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802128" httpUri="https://zenodo.org/record/14802128/files/figure.png" pageId="41" pageNumber="42">Figs 22A</figureCitation>
,
<figureCitation id="13312A5230298736162EFD200DA6FCBA" box="[1149,1193,744,768]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[116,1453,147,816]" captionTargetId="figure-515@43.[113,1457,144,820]" captionTargetPageId="43" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="41" pageNumber="42">23C</figureCitation>
). Ocularium elliptical (in dorsal view), high (
<emphasis id="B97EEAC53029873611A9FCC00D0CFCA5" box="[1018,1027,776,799]" italics="true" pageId="41" pageNumber="42">c</emphasis>
. 2.5× the eye diameter), perpendicularly placed on the central portion of the carapace (
<figureCitation id="13312A52302A873510D2FA9C0A0CFAD1" box="[641,771,1364,1388]" captionStart="Figure 22" captionStartId="42.[126,191,1272,1296]" captionTargetBox="[129,1473,144,1233]" captionTargetId="figure-388@42.[129,1473,144,1233]" captionTargetPageId="42" captionText="Figure 22. Iamarinus fenax, ♀ (MHNCI 6305). A, habitus, dorsal view.B, detail of the mesotergum and free tergites I–III, dorsal view. C, habitus, ventral view.D, same, lateral view.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802128" httpUri="https://zenodo.org/record/14802128/files/figure.png" pageId="42" pageNumber="43">Figs 22A, D</figureCitation>
,
<figureCitation id="13312A52302A873512D2FABB09C3FA31" box="[129,204,1395,1419]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[116,1453,147,816]" captionTargetId="figure-515@43.[113,1457,144,820]" captionTargetPageId="43" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="42" pageNumber="43">23B–C</figureCitation>
). Ocularium with a pair of divergent spines (
<emphasis id="B97EEAC5302A873510FDFABC0BB8FA31" box="[686,695,1396,1419]" italics="true" pageId="42" pageNumber="43">c</emphasis>
. 3× the eye diameter) (
<figureCitation id="13312A52302A87351376FA5B08ADFA10" box="[293,418,1427,1451]" captionStart="Figure 22" captionStartId="42.[126,191,1272,1296]" captionTargetBox="[129,1473,144,1233]" captionTargetId="figure-388@42.[129,1473,144,1233]" captionTargetPageId="42" captionText="Figure 22. Iamarinus fenax, ♀ (MHNCI 6305). A, habitus, dorsal view.B, detail of the mesotergum and free tergites I–III, dorsal view. C, habitus, ventral view.D, same, lateral view.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802128" httpUri="https://zenodo.org/record/14802128/files/figure.png" pageId="42" pageNumber="43">Figs 22A, D</figureCitation>
,
<figureCitation id="13312A52302A873513FCFA5B08F4FA11" box="[431,507,1427,1451]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[116,1453,147,816]" captionTargetId="figure-515@43.[113,1457,144,820]" captionTargetPageId="43" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="42" pageNumber="43">23B–C</figureCitation>
).
<collectingCountry id="F31D7647302A87351040FA5B0B3DFA11" box="[531,562,1427,1451]" name="American Samoa" pageId="42" pageNumber="43">AS</collectingCountry>
lateral margins with one spiniform apophysis at scutal areas II–
<collectionCode id="ED1BAE12302A8735100BFA7A0B7CFA73" box="[600,627,1458,1481]" pageId="42" pageNumber="43">III</collectionCode>
(
<figureCitation id="13312A52302A873510D4FA7A0A0CFA70" box="[647,771,1458,1482]" captionStart="Figure 22" captionStartId="42.[126,191,1272,1296]" captionTargetBox="[129,1473,144,1233]" captionTargetId="figure-388@42.[129,1473,144,1233]" captionTargetPageId="42" captionText="Figure 22. Iamarinus fenax, ♀ (MHNCI 6305). A, habitus, dorsal view.B, detail of the mesotergum and free tergites I–III, dorsal view. C, habitus, ventral view.D, same, lateral view.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802128" httpUri="https://zenodo.org/record/14802128/files/figure.png" pageId="42" pageNumber="43">Figs 22A–B</figureCitation>
,
<figureCitation id="13312A52302A873512D2FA1909A2FA53" box="[129,173,1489,1513]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[116,1453,147,816]" captionTargetId="figure-515@43.[113,1457,144,820]" captionTargetPageId="43" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="42" pageNumber="43">23C</figureCitation>
). Scutal area
<collectionCode id="ED1BAE12302A87351319FA1A086AFA53" box="[330,357,1490,1513]" pageId="42" pageNumber="43">III</collectionCode>
with a paramedian pair of subconical spines (
<figureCitation id="13312A52302A87351284FA390878F9B2" box="[215,375,1521,1545]" captionStart="Figure 22" captionStartId="42.[126,191,1272,1296]" captionTargetBox="[129,1473,144,1233]" captionTargetId="figure-388@42.[129,1473,144,1233]" captionTargetPageId="42" captionText="Figure 22. Iamarinus fenax, ♀ (MHNCI 6305). A, habitus, dorsal view.B, detail of the mesotergum and free tergites I–III, dorsal view. C, habitus, ventral view.D, same, lateral view.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802128" httpUri="https://zenodo.org/record/14802128/files/figure.png" pageId="42" pageNumber="43">Figs 22A–B, D</figureCitation>
,
<figureCitation id="13312A52302A873513DBFA3908BAF9B3" box="[392,437,1521,1545]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[116,1453,147,816]" captionTargetId="figure-515@43.[113,1457,144,820]" captionTargetPageId="43" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="42" pageNumber="43">23C</figureCitation>
). Scutal area IV with ordinary tubercles (
<figureCitation id="13312A52302A873512A0F9D80861F992" box="[243,366,1552,1576]" captionStart="Figure 22" captionStartId="42.[126,191,1272,1296]" captionTargetBox="[129,1473,144,1233]" captionTargetId="figure-388@42.[129,1473,144,1233]" captionTargetPageId="42" captionText="Figure 22. Iamarinus fenax, ♀ (MHNCI 6305). A, habitus, dorsal view.B, detail of the mesotergum and free tergites I–III, dorsal view. C, habitus, ventral view.D, same, lateral view.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802128" httpUri="https://zenodo.org/record/14802128/files/figure.png" pageId="42" pageNumber="43">Figs 22A–B</figureCitation>
,
<figureCitation id="13312A52302A8735132FF9D808C7F992" box="[380,456,1552,1576]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[116,1453,147,816]" captionTargetId="figure-515@43.[113,1457,144,820]" captionTargetPageId="43" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="42" pageNumber="43">23A–C</figureCitation>
). Cx IV narrower than males, with the prodorsal distal apophysis reduced to a single spine and a reduced retroventral distal apophysis (
<figureCitation id="13312A52302A87351031F9870A0CF9DD" box="[610,771,1615,1639]" captionStart="Figure 22" captionStartId="42.[126,191,1272,1296]" captionTargetBox="[129,1473,144,1233]" captionTargetId="figure-388@42.[129,1473,144,1233]" captionTargetPageId="42" captionText="Figure 22. Iamarinus fenax, ♀ (MHNCI 6305). A, habitus, dorsal view.B, detail of the mesotergum and free tergites I–III, dorsal view. C, habitus, ventral view.D, same, lateral view.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802128" httpUri="https://zenodo.org/record/14802128/files/figure.png" pageId="42" pageNumber="43">Figs 22A, C–D</figureCitation>
,
<figureCitation id="13312A52302A873512D2F9A609A2F93C" box="[129,173,1646,1670]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[116,1453,147,816]" captionTargetId="figure-515@43.[113,1457,144,820]" captionTargetPageId="43" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="42" pageNumber="43">23C</figureCitation>
). Tr IV prodorsal distal half and prolateral entire length portions with prominent tubercles (
<figureCitation id="13312A52302A87351043F9450B99F91F" box="[528,662,1677,1701]" captionStart="Figure 22" captionStartId="42.[126,191,1272,1296]" captionTargetBox="[129,1473,144,1233]" captionTargetId="figure-388@42.[129,1473,144,1233]" captionTargetPageId="42" captionText="Figure 22. Iamarinus fenax, ♀ (MHNCI 6305). A, habitus, dorsal view.B, detail of the mesotergum and free tergites I–III, dorsal view. C, habitus, ventral view.D, same, lateral view.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802128" httpUri="https://zenodo.org/record/14802128/files/figure.png" pageId="42" pageNumber="43">Figs 22A, C</figureCitation>
,
<figureCitation id="13312A52302A873510F4F9450BDBF91F" box="[679,724,1677,1701]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[116,1453,147,816]" captionTargetId="figure-515@43.[113,1457,144,820]" captionTargetPageId="43" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="42" pageNumber="43">23D</figureCitation>
). Tr IV with a proximal and a distal conical apophysis (distalmost largest) on retrolateral face, with a reduced conical apophysis on the central portion (
<figureCitation id="13312A52302A873513C2F92308FAF8B9" box="[401,501,1771,1795]" captionStart="Figure 22" captionStartId="42.[126,191,1272,1296]" captionTargetBox="[129,1473,144,1233]" captionTargetId="figure-388@42.[129,1473,144,1233]" captionTargetPageId="42" captionText="Figure 22. Iamarinus fenax, ♀ (MHNCI 6305). A, habitus, dorsal view.B, detail of the mesotergum and free tergites I–III, dorsal view. C, habitus, ventral view.D, same, lateral view.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802128" httpUri="https://zenodo.org/record/14802128/files/figure.png" pageId="42" pageNumber="43">Figs 22C</figureCitation>
,
<figureCitation id="13312A52302A87351055F9230B5EF8B9" box="[518,593,1771,1795]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[116,1453,147,816]" captionTargetId="figure-515@43.[113,1457,144,820]" captionTargetPageId="43" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="42" pageNumber="43">23D–E</figureCitation>
). Fe IV straight and thinner than the males (
<figureCitation id="13312A52302A873513EBF8C30B55F899" box="[440,602,1803,1827]" captionStart="Figure 22" captionStartId="42.[126,191,1272,1296]" captionTargetBox="[129,1473,144,1233]" captionTargetId="figure-388@42.[129,1473,144,1233]" captionTargetPageId="42" captionText="Figure 22. Iamarinus fenax, ♀ (MHNCI 6305). A, habitus, dorsal view.B, detail of the mesotergum and free tergites I–III, dorsal view. C, habitus, ventral view.D, same, lateral view.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802128" httpUri="https://zenodo.org/record/14802128/files/figure.png" pageId="42" pageNumber="43">Figs 22A, C–D</figureCitation>
,
<figureCitation id="13312A52302A8735103BF8C30BBCF899" box="[616,691,1803,1827]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[116,1453,147,816]" captionTargetId="figure-515@43.[113,1457,144,820]" captionTargetPageId="43" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="42" pageNumber="43">23D–E</figureCitation>
). Fe IV dorsal face with four spines (one on proximal third, two on central third and one on distal third) (
<figureCitation id="13312A52302A8735104EF8810B93F8DB" box="[541,668,1865,1889]" captionStart="Figure 22" captionStartId="42.[126,191,1272,1296]" captionTargetBox="[129,1473,144,1233]" captionTargetId="figure-388@42.[129,1473,144,1233]" captionTargetPageId="42" captionText="Figure 22. Iamarinus fenax, ♀ (MHNCI 6305). A, habitus, dorsal view.B, detail of the mesotergum and free tergites I–III, dorsal view. C, habitus, ventral view.D, same, lateral view.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14802128" httpUri="https://zenodo.org/record/14802128/files/figure.png" pageId="42" pageNumber="43">Figs 22A, D</figureCitation>
,
<figureCitation id="13312A52302A873510F9F8810BD8F8DB" box="[682,727,1865,1889]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[116,1453,147,816]" captionTargetId="figure-515@43.[113,1457,144,820]" captionTargetPageId="43" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="42" pageNumber="43">23D</figureCitation>
). Fe IV prolateral face with a row of acuminated tubercles (
<figureCitation id="13312A52302A873510B0F8A109C4F81A" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[116,1453,147,816]" captionTargetId="figure-515@43.[113,1457,144,820]" captionTargetPageId="43" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="42" pageNumber="43">Fig. 23D–E</figureCitation>
). Fe IV proventral and retroventral faces with a row of acuminated tubercles on the proximal half and three–four conical spines on distal half (
<figureCitation id="13312A52302A8735163DFA9C0DCAFAD6" box="[1134,1221,1364,1388]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[116,1453,147,816]" captionTargetId="figure-515@43.[113,1457,144,820]" captionTargetPageId="43" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="42" pageNumber="43">Fig. 23E</figureCitation>
). Fe IV retrolateral face with a row of conical spines (iiiI on the proximal half, Iii on the distal half) (
<figureCitation id="13312A52302A873511EEFA5B0D3CFA11" box="[957,1075,1427,1451]" captionStart="15" captionStartId="5.[113,139,194,218]" captionTargetBox="[116,1453,147,816]" captionTargetId="figure-515@43.[113,1457,144,820]" captionTargetPageId="43" captionText="15. Penis, glans, ventral process, flabellum, frame2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) middle axis (Fig. 11D) (1) double V-shaped axis 16. Penis, glans, ventral process, flabellum, middle axis, type1,2,3,4,5,6 (DELTRAN, L = 3, CI = 100, RI = 100) (0) scallop-shaped (Carvalho and Kury 2023: Fig. 8D) (1) hand-shaped (Fig.15D) (2) foliaceus (Carvalho et al. 2018: fig. 5D) (3) fan-shaped 17. Penis, glans, ventral process, flabellum, middle axis, type of ornamentation2,3,4,5 (DELTRAN, L = 3, CI = 100, RI = 100) (0) continuous serrulation (Carvalho and Kury 2023: fig.8D) (1) discrete spines (Fig.19D) 18. Penis, glans, ventral process, flabellum, middle axis, arrangement of serrulation2,3 (N/O, L = 2, CI = 50, RI = 50) (0) serrulate,multi-layered (Pinto-da-Rocha and Bragagnolo 2010: fig.23) (1) incised, single-layered 19. Penis, glans, ventral process, diameter2,3,4,5,6 (DELTRAN, L = 9, CI = 44, RI = 70) (0) two-thirds of the stylus (1) half of the stylus (Fig.19C) (2) wider than the stylus (3) almost the same diameter of the stylus (Fig. 33D) (4) a third or less than the stylus (Carvalho and Kury 2020: fig.18E) 20.Penis, VP, lateral outlines of the basal half2,3,4,5,6 (N/O, L = 3, CI = 66, RI = 50) (0) straight (1) elliptical (Figs 15A–B) (2) diagonal, narrowing basally (Kury and Carvalho 2016: fig.8A) 21. Penis, VP, outline of the basal half elliptical, laterally expanded than distal part2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Fig. 15A–B) (1) present (Carvalho and Kury 2018: fig.6A–B) 22.Penis, VP, basal convex format (in lateral view)1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) absent (Carvalho et al. 2018: fig. 5B) (1) present (Fig.19C) 23. Penis, VP, distal portion without a groove, type1,2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) trapezoidal (widest apically) (Fig. 15A–B) (1) trapezoidal (narrower apically) (Carvalho and Kury 2023: fig.18A) 24.Penis, VP, distal and basal width ratio2,3,4,6 (N/O, L = 5, CI = 40, RI = 62) (0) same proportion (Fig. 15A–B) (1) basal twice wider than distal (Kury and Carvalho 2016: fig.8F) (2) basal one-third wider than distal (Fig.27A) 25. Penis, VP, MS A2, position on the VP3 (ACCTRAN, L = 8, CI = 25, RI = 50) (0) lateral (Fig. 19C) (1) basal (Fig. 33B) (2) dorsal (Fig.15C) 26.Penis, VP, MS B12,3,4,5 (N/O, L = 1, CI = 100, RI = 100) (0) absent (1) present (Fig.26C) 27.Penis, VP, MS B1, type2,3,4,5(ACCTRAN, L = 7, CI = 14, RI = 60) (0) only the apical portion exposed (Fig.33B) (1) more than the apical portion exposed (Fig. 15C) 28.Penis, VP, MS C, diameter1,2,3 (N/O, L = 1, CI = 100, RI = 100) (0) half of the stylus diameter (Fig.33A) (1) one third or less of the stylus diameter (Kury and Carvalho 2016: fig. 8D) 29.Penis, VP, MS C, length (compared to the MS A)1,2,3 (N/O, L = 2, CI = 50, RI = 83) (0) equal or less (Fig. 33A) (1) larger (Kury and Carvalho 2016: fig.8D) 30.Penis, VP, shape of the field of type 1 microsetae1,2,3,4,5,6 (N/O, L = 2, CI = 100, RI = 100) (0) field strongly reduced to a pair of latero-basal patches (1) field entire, occupying most of VP (Fig. 19B) (2) field restricted to the proximal third of VP (Bragagnolo and Pinto-da-Rocha 2009: fig.8C) 31. Ch, basal article, relation with the DS anterior margin2,3,4,5,6 (N/O, L = 1, CI = 100, RI = 100) (0) covered mainly by the DS anterior margin (1) well exposed (Fig. 18A)" pageId="42" pageNumber="43">Fig. 23D–E</figureCitation>
).
</paragraph>
<caption id="DF75665F302987361222F84E0C13F801" ID-DOI="http://doi.org/10.5281/zenodo.14802126" ID-Zenodo-Dep="14802126" httpUri="https://zenodo.org/record/14802126/files/figure.png" pageId="41" pageNumber="42" startId="41.[113,178,1926,1950]" targetBox="[118,1456,908,1896]" targetPageId="41" targetType="figure">
<paragraph id="8BB536D7302987361222F84E0C13F801" blockId="41.[113,1386,1926,1979]" pageId="41" pageNumber="42">
<emphasis id="B97EEAC5302987361222F84E09D9F825" bold="true" box="[113,214,1926,1951]" pageId="41" pageNumber="42">Figure 21.</emphasis>
South Brazil, showing distribution of
<taxonomicName id="4C0A4D54302987361068F84F0BE9F825" box="[571,742,1927,1951]" class="Arachnida" family="Gonyleptidae" genus="Iamarinus" kingdom="Animalia" order="Opiliones" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97EEAC5302987361068F84F0B97F824" box="[571,664,1927,1950]" italics="true" pageId="41" pageNumber="42">Iamarinus</emphasis>
species.
</taxonomicName>
Shaded areas in the background are Morrone’s regionalization of the Neotropics (‘provinces’;
<bibRefCitation id="EF9B4B2630298736130FF86B0B19F801" author="Morrone JJ &amp; Escalante T &amp; Rodriguez-Tapia G" box="[348,534,1955,1979]" pageId="41" pageNumber="42" refId="ref54045" refString="Morrone JJ, Escalante T, Rodriguez-Tapia G et al. Biogeographic regionalization of the Neotropical region: new map and shapefile. Anais da Academia Brasileira de Ciencias 2022; 94: e 20211167. https: // doi. org / 10.1590 / 0001 - 3765202220211167" type="journal volume" year="2022">
Morrone
<emphasis id="B97EEAC53029873613E4F86B08EDF801" box="[439,482,1955,1979]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
2022
</bibRefCitation>
): Araucaria Forest province (in light green) and Atlantic province (in dark green).
</paragraph>
</caption>
<caption id="DF75665F302A8735122DFB300BC8FA96" ID-DOI="http://doi.org/10.5281/zenodo.14802128" ID-Zenodo-Dep="14802128" httpUri="https://zenodo.org/record/14802128/files/figure.png" pageId="42" pageNumber="43" startId="42.[126,191,1272,1296]" targetBox="[129,1473,144,1233]" targetPageId="42" targetType="figure">
<paragraph id="8BB536D7302A8735122DFB300BC8FA96" blockId="42.[126,1420,1272,1324]" pageId="42" pageNumber="43">
<emphasis id="B97EEAC5302A8735122DFB3009ECFAAA" bold="true" box="[126,227,1272,1296]" pageId="42" pageNumber="43">Figure 22.</emphasis>
<taxonomicName id="4C0A4D54302A873512BAFB310873FAAA" baseAuthorityName="Kury, Pinto-da-Rocha &amp; Carvalho" baseAuthorityYear="2018" box="[233,380,1272,1296]" class="Arachnida" family="Gonyleptidae" genus="Iamarinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="fenax">
<emphasis id="B97EEAC5302A873512BAFB310873FAAA" box="[233,380,1272,1296]" italics="true" pageId="42" pageNumber="43">Iamarinus fenax</emphasis>
</taxonomicName>
, ♀ (MHNCI 6305). A, habitus, dorsal view. B, detail of the mesotergum and free tergites I–III, dorsal view. C, habitus, ventral view. D, same, lateral view. Scale bars = 5 mm.
</paragraph>
</caption>
<paragraph id="8BB536D7302A8735116AFA0D0D68F923" blockId="42.[825,1475,1477,1689]" pageId="42" pageNumber="43">
<emphasis id="B97EEAC5302A8735116AFA0D0D1EFA66" box="[825,1041,1477,1501]" italics="true" pageId="42" pageNumber="43">Intraspecific variation:</emphasis>
Some variations among
<emphasis id="B97EEAC5302A87351755FA0D0CCEFA67" box="[1286,1473,1477,1501]" italics="true" pageId="42" pageNumber="43">minor morph males</emphasis>
and
<emphasis id="B97EEAC5302A87351136FA2D0D2DFA46" box="[869,1058,1508,1532]" italics="true" pageId="42" pageNumber="43">major morph males</emphasis>
were detected: (1) DS narrower; (2) Cx IV prolateral and retrolateral apophyses reduced; and (3) Fe IV thinner, with reduced proximal spines. Variation among
<emphasis id="B97EEAC5302A873517DBF9EB0AB9F9E0" italics="true" pageId="42" pageNumber="43">major morph males</emphasis>
: the Fe IV torsion, which can be more or less prominent. No relevant intraspecific variation among females was detected in the material studied.
</paragraph>
</subSubSection>
</treatment>
</document>
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