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<document id="02DBC625195387245F53C9A0BAA667E8" ID="10.11646/zootaxa.4267.1.1SLASH11021" ID-CLB-Dataset="33061" ID-DOI="10.5281/zenodo.575641" ID-GBIF-Dataset="810b55e4-b8ba-4368-bf9c-8517ee33e7d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="575641" ID-ZooBank="8920893E-E5F8-482A-A60D-7A248D2B0DCB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1494577389872" checkinUser="plazi" docAuthor="Skelley, Paul, Xu, Guang, Tang, William, Lindström, Anders J., Marler, Thomas, Khuraijam, Jibankumar Singh, Singh, Rita &amp; Rich, Stephen" docDate="2017" docId="03A38781FFC9FFBEFF70FE37FBC5C084" docLanguage="en" docName="zootaxa.4267.1.1SLASH11021.pdf" docOrigin="Zootaxa 4267 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cycadophila (Strobilophila) assamensis Skelley, Xu &amp; Tang, new species" docType="treatment" docVersion="8" lastPageNumber="50" masterDocId="FF9AFFF9FFE6FF8FFFE7FFEBFFD5C639" masterDocTitle="Review of Cycadophila Xu, Tang &amp; Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species" masterLastPageNumber="63" masterPageNumber="1" pageNumber="48" updateTime="1698439510239" updateUser="ExternalLinkService">
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<mods:title id="060DDAA7D1478CD9ACCE1FE6CDA485AA">Review of Cycadophila Xu, Tang &amp; Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species</mods:title>
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<mods:namePart id="49D9EB9F187F47412105BDE45E684749">Tang, William</mods:namePart>
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<mods:namePart id="7A0AC9CC39DF7C170D8B2AECC70136E9">Lindström, Anders J.</mods:namePart>
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<mods:namePart id="603192081E76E2EE649198CB3127CBDD">Khuraijam, Jibankumar Singh</mods:namePart>
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<treatment id="03A38781FFC9FFBEFF70FE37FBC5C084" ID-DOI="http://doi.org/10.5281/zenodo.5999227" ID-GBIF-Taxon="130809507" ID-Zenodo-Dep="5999227" LSID="urn:lsid:plazi:treatment:03A38781FFC9FFBEFF70FE37FBC5C084" httpUri="http://treatment.plazi.org/id/03A38781FFC9FFBEFF70FE37FBC5C084" lastPageId="49" lastPageNumber="50" pageId="47" pageNumber="48">
<subSubSection id="C310651CFFC9FFA0FF70FE37FEF2C42E" pageId="47" pageNumber="48" type="nomenclature">
<paragraph id="8BB53697FFC9FFA0FF70FE37FBD9C7CF" blockId="47.[151,1036,476,535]" box="[151,1036,476,502]" pageId="47" pageNumber="48">
<heading id="D0FD81FBFFC9FFA0FF70FE37FBD9C7CF" bold="true" box="[151,1036,476,502]" fontSize="11" level="1" pageId="47" pageNumber="48" reason="1">
<emphasis id="B97EEA85FFC9FFA0FF70FE37FBD9C7CF" bold="true" box="[151,1036,476,502]" pageId="47" pageNumber="48">
<taxonomicName id="4C0A4D14FFC9FFA0FF70FE37FCBBC7CF" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[151,878,476,502]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="assamensis" status="sp. nov." subGenus="Strobilophila">
<emphasis id="B97EEA85FFC9FFA0FF70FE37FEFBC7CF" bold="true" box="[151,302,476,502]" italics="true" pageId="47" pageNumber="48">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFC9FFA0FED8FE37FE09C7CF" bold="true" box="[319,476,476,502]" italics="true" pageId="47" pageNumber="48">Strobilophila</emphasis>
)
<emphasis id="B97EEA85FFC9FFA0FE0AFE37FDA1C7CF" bold="true" box="[493,628,476,502]" italics="true" pageId="47" pageNumber="48">assamensis</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
,
<taxonomicNameLabel id="A24D57FEFFC9FFA0FC9BFE37FBD9C7CF" box="[892,1036,476,502]" pageId="47" pageNumber="48" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB53697FFC9FFA0FF70FE15FEF2C42E" blockId="47.[151,1036,476,535]" box="[151,295,510,535]" pageId="47" pageNumber="48">
<figureCitation id="13312A12FFC9FFA0FF70FE15FF25C42E" box="[151,240,510,535]" captionStart="FIGURE 16" captionStartId="48.[151,250,1886,1908]" captionTargetBox="[151,1435,339,1864]" captionTargetId="figure@48.[151,1435,339,1864]" captionTargetPageId="48" captionText="FIGURE 16. Photographs of Cycadophila (Strobilophila) assamensis, female holotype: A) dorsal habitus; B) ventral habitus; C) lateral habitus; D) head and prosternum, ventral; E) male genitalia; F) male paratype, abdominal ventrite V; G) female holotype, abdominal ventrite V." httpUri="https://zenodo.org/record/575657/files/figure.png" pageId="47" pageNumber="48">Figs. 16</figureCitation>
AG
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFC9FFA0FF70FDADFAEDC237" pageId="47" pageNumber="48" type="diagnosis">
<paragraph id="8BB53697FFC9FFA0FF70FDADFD67C562" blockId="47.[151,1437,582,2011]" pageId="47" pageNumber="48">
<emphasis id="B97EEA85FFC9FFA0FF70FDADFE8FC466" bold="true" box="[151,346,582,607]" pageId="47" pageNumber="48">Adult diagnosis.</emphasis>
A member of
<taxonomicName id="4C0A4D14FFC9FFA0FDE2FDADFC97C466" authority="Strobilophila" authorityName="Strobilophila" box="[517,834,582,607]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFC9FFA0FDE2FDADFD41C466" box="[517,660,582,607]" italics="true" pageId="47" pageNumber="48">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFC9FFA0FD40FDADFCECC466" box="[679,825,582,607]" italics="true" pageId="47" pageNumber="48">Strobilophila</emphasis>
)
</taxonomicName>
as discussed above, more readily distinguished by a combination of characters than any single unique character, and by analysis of the 16S rRNA mitochondrial gene.
<taxonomicName id="4C0A4D14FFC9FFA0FF70FD65FE79C49F" box="[151,428,654,679]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="B97EEA85FFC9FFA0FF70FD65FE79C49F" box="[151,428,654,679]" italics="true" pageId="47" pageNumber="48">Cycadophila assamensis</emphasis>
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can be distinguished by the distally dilated protibia bearing narrowly spaced teeth along the apical third of the lateral margin, the distinct furrow on each side of pronotal base, elytral disc being mostly glabrous, the nearly straight basal pronotal margin, the lateral pronotal carina being abruptly curved inward at posterior angle pore which has a small acute projection behind it, and the terminal abdominal ventrite being sexually dimorphic (males with surface evenly coarsely punctate and setose; females with surface entirely setose and bearing a glossy area near the medial apex).
</paragraph>
<paragraph id="8BB53697FFC9FFA0FF20FC8DFAEDC237" blockId="47.[151,1437,582,2011]" pageId="47" pageNumber="48">
<emphasis id="B97EEA85FFC9FFA0FF20FC8DFE4FC546" bold="true" box="[199,410,870,895]" pageId="47" pageNumber="48">Adult description</emphasis>
. Length
<quantity id="4CF29B72FFC9FFA0FDE5FC8DFD7FC547" box="[514,682,870,895]" metricMagnitude="-3" metricUnit="m" metricValue="4.94" metricValueMax="5.19" metricValueMin="4.69" pageId="47" pageNumber="48" unit="mm" value="4.94" valueMax="5.19" valueMin="4.69">4.695.19 mm</quantity>
; width 1.8 82.00 mm. Body in dorsal view elongate, sides nearly straight, greatest width at middle of elytra; in lateral view convex dorsally. General body and appendage color solid reddish-brown [available specimens poorly preserved, darker; image of fresh specimen has color identical to
<taxonomicName id="4C0A4D14FFC9FFA0FA65FC44FED1C5D2" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="tansachai">
<emphasis id="B97EEA85FFC9FFA0FA65FC44FED1C5D2" italics="true" pageId="47" pageNumber="48">C. tansachai</emphasis>
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]; dorsal surface distinctly punctate, surface appearing slightly dulled, dorsally appearing glabrous, with short procumbent hairs associated with punctation, ventrally mostly covered with short procumbent setae.
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFC9FFBEFF20FBF1FCD9C4F4" lastPageId="49" lastPageNumber="50" pageId="47" pageNumber="48" type="description">
<paragraph id="8BB53697FFC9FFA0FF20FBF1FDA4C076" blockId="47.[151,1437,582,2011]" pageId="47" pageNumber="48">
<emphasis id="B97EEA85FFC9FFA0FF20FBF1FED0C20A" bold="true" box="[199,261,1050,1075]" italics="true" pageId="47" pageNumber="48">Head</emphasis>
in dorsal view conical, gradually narrowed apically, surface flat to slightly convex, distinctly punctured; width 1.00
<quantity id="4CF29B72FFC9FFA0FEFAFBD4FE51C26F" box="[285,388,1087,1111]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="47" pageNumber="48" unit="mm" value="1.1">1.10 mm</quantity>
; dorsal interocular distance
<quantity id="4CF29B72FFC9FFA0FD58FBD5FCB6C26F" box="[703,867,1086,1111]" metricMagnitude="-4" metricUnit="m" metricValue="6.95" metricValueMax="7.3" metricValueMin="6.6" pageId="47" pageNumber="48" unit="mm" value="0.695" valueMax="0.73" valueMin="0.66">0.660.73 mm</quantity>
, head width/dorsal interocular distance ratio 1.50 1.52; ventral interocular distance
<quantity id="4CF29B72FFC9FFA0FDC4FB89FD18C243" box="[547,717,1122,1147]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="4.9" metricValueMin="4.6" pageId="47" pageNumber="48" unit="mm" value="0.475" valueMax="0.49" valueMin="0.46">0.460.49 mm</quantity>
, head width/ventral interocular distance ratio 2.162.25. Eye globular; bordered dorsally with supraocular stria; projected laterally, with large black facets and short, inconspicuous interfacetal setae. Frons with short, dark, subcuticular suture on each side, approximately 1/4 width of frons in length, extending obliquely to lateral margin above and anterior to antennal insertion. Antennal length about 2/3 pronotal width, and slightly longer than head width, antennomere relative lengths from base to end approximately 19:13:13:10:10:10:10:10:15:15:19; antennomere I (scape) fairly large, slightly elongate, antennomere II slightly smaller than III; IVVIII small, width equals length; club fairly large, IXXI similar in length, XI globular. Clypeus truncate anteriorly, anterior margin with long setae; densely punctate; somewhat emarginate, with narrow margin. Maxillary palpomeres II, III slightly longer than wide; IV elongate oval, apex densely papillate; relative lengths of IIIV approximately 3:3:5. Labium with mentum pentagonal with carina forming triangular plate projecting medially, lacking distinct lateral pockets; labial palpomere III elongate-oval.
<collectingRegion id="49CEF875FFC9FFA0FF70FA04FF23C03F" box="[151,246,1519,1542]" country="Myanmar" name="Chin" pageId="47" pageNumber="48">Mentum</collectingRegion>
and submentum with moderate punctation and setation, setae short. Gular area smooth, without setae or punctation, except near suture with submentum where a shallow transversely elongate fovea is present bearing dense setose punctures, fovea dark brown.
</paragraph>
<paragraph id="8BB53697FFC9FFA0FF20F9B1FA48C1E2" blockId="47.[151,1437,582,2011]" pageId="47" pageNumber="48">
<emphasis id="B97EEA85FFC9FFA0FF20F9B1FEC8C04A" bold="true" box="[199,285,1626,1651]" italics="true" pageId="47" pageNumber="48">Thorax</emphasis>
with pronotum transverse in dorsal view, with marginal beads on all sides; length/width ratio 0.65 0.68, convex, evenly weakly arcuate most of length; anterior angles rounded; lateral carina with thickened bead, bead bearing scattered setose punctures; posterior angle exteriorly angulate, indented at pore; posterior margin projecting medially; with narrow, short longitudinal furrow posterolaterally, beginning at pore on posterior margin, extending forward and ending abruptly, length of furrows relative to pronotal length 0.220.24. Prosternum with anterior margin slightly emarginate, finely denticulate with fringe of long, anteriorly directed setae; posterior prosternal process round and convex. Hypomeron with coarse punctures laterally, medially with longitudinal striations. Scutellar shield transverse, posterior margin projected, pentagonal. Elytra elongate, nearly parallel sided, convex; length/width 3.283.65, greatest width near midlength; with marginal bead basally, striae not impressed, scutellary striole with 1012 punctures; intervals of striae with distinct punctures. All elytral punctures bearing a single short seta; seta visible but barely emerging out of puncture. Punctation on meso- and metaventrite distinct.
</paragraph>
<paragraph id="8BB53697FFD6FFBFFF70FF7CFB7AC724" blockId="48.[151,1436,151,285]" pageId="48" pageNumber="49">Metaventrite long, convex laterally, slightly impressed medially, metathoracic discrimen extending about ¾ metaventrite length. Legs stout, similar in length; femora compressed laterally; tibiae shorter than femora, dilated to wide apex. Protibia triangularly dilated at apex, apex and apical 1/3 of lateral margin with stout spinules, marginal spinules separated by less than length. Meso- and metatibia less triangularly dilated.</paragraph>
<caption id="DF75661FFFD6FFBFFF70F8B5FE02C180" httpUri="https://zenodo.org/record/575657/files/figure.png" pageId="48" pageNumber="49" targetBox="[151,1435,339,1864]" targetPageId="48">
<paragraph id="8BB53697FFD6FFBFFF70F8B5FE02C180" blockId="48.[151,1436,1886,1977]" pageId="48" pageNumber="49">
<emphasis id="B97EEA85FFD6FFBFFF70F8B5FEF5C14D" bold="true" box="[151,288,1886,1908]" pageId="48" pageNumber="49">FIGURE 16.</emphasis>
Photographs of
<taxonomicName id="4C0A4D14FFD6FFBFFE2EF8B5FC8AC14D" box="[457,863,1886,1908]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="assamensis" subGenus="Strobilophila">
<emphasis id="B97EEA85FFD6FFBFFE2EF8B5FD9EC14D" box="[457,587,1886,1908]" italics="true" pageId="48" pageNumber="49">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFD6FFBFFDBDF8B5FD0BC14D" box="[602,734,1886,1908]" italics="true" pageId="48" pageNumber="49">Strobilophila</emphasis>
)
<emphasis id="B97EEA85FFD6FFBFFD0AF8B4FC8AC14D" box="[749,863,1887,1908]" italics="true" pageId="48" pageNumber="49">assamensis</emphasis>
</taxonomicName>
, female holotype: A) dorsal habitus; B) ventral habitus; C) lateral habitus; D) head and prosternum, ventral; E) male genitalia; F) male paratype, abdominal ventrite V; G) female holotype, abdominal ventrite V.
</paragraph>
</caption>
<paragraph id="8BB53697FFD7FFBEFF20FF7CFF0DC7E9" blockId="49.[151,1437,151,1725]" pageId="49" pageNumber="50">
<emphasis id="B97EEA85FFD7FFBEFF20FF7CFEE0C689" bold="true" box="[199,309,151,176]" italics="true" pageId="49" pageNumber="50">Abdomen</emphasis>
with 5 ventrites bearing fine punctures; anterior margin with intercoxal process narrow, with triangular point anteromedially, lateral edges slightly projected, lateral and posterior margins arcuate, converging posteriorly; anterior and posterior margins of ventrites more or less straight or weakly bisinuate (
<figureCitation id="13312A12FFD7FFBEFB1CFF34FA8DC6C1" box="[1275,1368,223,248]" captionStart="FIGURE 16" captionStartId="48.[151,250,1886,1908]" captionTargetBox="[151,1435,339,1864]" captionTargetId="figure@48.[151,1435,339,1864]" captionTargetPageId="48" captionText="FIGURE 16. Photographs of Cycadophila (Strobilophila) assamensis, female holotype: A) dorsal habitus; B) ventral habitus; C) lateral habitus; D) head and prosternum, ventral; E) male genitalia; F) male paratype, abdominal ventrite V; G) female holotype, abdominal ventrite V." httpUri="https://zenodo.org/record/575657/files/figure.png" pageId="49" pageNumber="50">Figs. 16</figureCitation>
FG); ventrite I lacking subcoxal line, slightly longer medially than II; IIV each with pair short hairs located adjacent to midline (often abraded); IIIV subequal in length; V slightly longer than IV with lateral margins converging posteriorly to a rounded apex bearing row of short appressed, densely arranged setae; ventrite V surface of male evenly coarsely and rugosely punctate, and entirely setose (similar to
<taxonomicName id="4C0A4D14FFD7FFBEFC24FE9BFB81C7B1" box="[963,1108,367,392]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="tansachai">
<emphasis id="B97EEA85FFD7FFBEFC24FE9BFB81C7B1" box="[963,1108,367,392]" italics="true" pageId="49" pageNumber="50">C. tansachai</emphasis>
</taxonomicName>
in
<figureCitation id="13312A12FFD7FFBEFB99FE84FB07C7B1" box="[1150,1234,367,392]" captionStart="FIGURE 19" captionStartId="55.[151,250,1953,1975]" captionTargetBox="[207,1380,193,1924]" captionTargetId="figure@55.[207,1380,193,1932]" captionTargetPageId="55" captionText="FIGURE 19. Photographs of Cycadophila (Strobilophila) tansachai, paratypes: A) dorsal habitus; B) ventral habitus; C) lateral habitus; D) head and prosternum, ventral; E) male genitalia; F) male abdominal ventrite V; G) female abdominal ventrite V; H I) inflated endophallus, lateral and ventral views." httpUri="https://zenodo.org/record/575660/files/figure.png" pageId="49" pageNumber="50">Fig. 19</figureCitation>
F), female evenly setose, basally with coarse punctures, with small area of fine punctures and glossy surface near medial apex (
<figureCitation id="13312A12FFD7FFBEFA88FE7FFF61C7E9" captionStart="FIGURE 16" captionStartId="48.[151,250,1886,1908]" captionTargetBox="[151,1435,339,1864]" captionTargetId="figure@48.[151,1435,339,1864]" captionTargetPageId="48" captionText="FIGURE 16. Photographs of Cycadophila (Strobilophila) assamensis, female holotype: A) dorsal habitus; B) ventral habitus; C) lateral habitus; D) head and prosternum, ventral; E) male genitalia; F) male paratype, abdominal ventrite V; G) female holotype, abdominal ventrite V." httpUri="https://zenodo.org/record/575657/files/figure.png" pageId="49" pageNumber="50">Fig. 16</figureCitation>
G).
</paragraph>
<paragraph id="8BB53697FFD7FFBEFF20FE37FCD9C4F4" blockId="49.[151,1437,151,1725]" pageId="49" pageNumber="50">
<emphasis id="B97EEA85FFD7FFBEFF20FE37FEBBC7CC" bold="true" box="[199,366,476,501]" italics="true" pageId="49" pageNumber="50">Male genitalia</emphasis>
(tegmen and median lobe of aedeagus) twisted approximately 45° toward the left side of animal and resting on side. Tegmen sclerotized, triangular, gently twisted; with anterior region ring-like, posterior region sheath-like, posterior margin slightly arcuate; lateral margins gradually converging posteriorly; dorsally with 2 large, elongate, somewhat coniform parameres. Parameres with long setae apically; length/width ratio 2.2 0. Aedeagus with median lobe short, slender, highly sclerotized, apex long and acuminate. Ratio of median lobe length to penile strut length 1:5.
<emphasis id="B97EEA85FFD7FFBEFDE5FD64FD8FC491" bold="true" box="[514,602,655,680]" italics="true" pageId="49" pageNumber="50">Female</emphasis>
similar to male; except with pygidium truncate not as well developed; and dimorphic abdominal ventrite V surface as noted above.
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFD7FFBEFF20FD3CFD72C245" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="8BB53697FFD7FFBEFF20FD3CFCE0C4C9" blockId="49.[151,1437,151,1725]" box="[199,821,727,752]" pageId="49" pageNumber="50">
<materialsCitation id="3B623CCAFFD7FFBEFF20FD3CFCE0C4C9" ID-GBIF-Occurrence="1502685004" box="[199,821,727,752]" country="India" location="North Guwahati." municipality="Kamrup" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Assam" typeStatus="holotype">
<emphasis id="B97EEA85FFD7FFBEFF20FD3CFEB0C4C9" bold="true" box="[199,357,727,752]" pageId="49" pageNumber="50">
<typeStatus id="54B18835FFD7FFBEFF20FD3CFED7C4C9" box="[199,258,727,752]" pageId="49" pageNumber="50">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F31D7607FFD7FFBEFE8BFD3CFE7FC4C9" box="[364,426,727,752]" name="India" pageId="49" pageNumber="50">India</collectingCountry>
:
<collectingRegion id="49CEF875FFD7FFBEFE53FD33FDD6C4C9" box="[436,515,728,752]" country="India" name="Assam" pageId="49" pageNumber="50">Assam</collectingRegion>
,
<collectingMunicipality id="6BD1ACEDFFD7FFBEFDE9FD33FDB8C4D6" box="[526,621,728,751]" pageId="49" pageNumber="50">Kamrup</collectingMunicipality>
,
<location id="8ED5604CFFD7FFBEFD9FFD3CFCE0C4C9" LSID="urn:lsid:plazi:treatment:03A38781FFC9FFBEFF70FE37FBC5C084:8ED5604CFFD7FFBEFD9FFD3CFCE0C4C9" box="[632,821,727,752]" country="India" municipality="Kamrup" name="North Guwahati." pageId="49" pageNumber="50" stateProvince="Assam">North Guwahati.</location>
</materialsCitation>
</paragraph>
<paragraph id="8BB53697FFD7FFBEFF20FD17FDC1C52C" blockId="49.[151,1437,151,1725]" box="[199,532,764,789]" pageId="49" pageNumber="50">
<materialsCitation id="3B623CCAFFD7FFBEFF20FD17FDC1C52C" ID-GBIF-Occurrence="1502685000" box="[199,532,764,789]" country="India" location="Assam" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Assam">
<emphasis id="B97EEA85FFD7FFBEFF20FD17FEC6C52C" bold="true" box="[199,275,764,789]" pageId="49" pageNumber="50">Range</emphasis>
.
<collectingCountry id="F31D7607FFD7FFBEFEC6FD17FE8EC52C" box="[289,347,764,789]" name="India" pageId="49" pageNumber="50">India</collectingCountry>
(
<collectingRegion id="49CEF875FFD7FFBEFE8EFD17FE6CC52D" box="[361,441,764,788]" country="India" name="Assam" pageId="49" pageNumber="50">Assam</collectingRegion>
,
<collectingRegion id="49CEF875FFD7FFBEFE22FD17FDDDC52C" box="[453,520,764,789]" country="India" name="Bihar" pageId="49" pageNumber="50">Bihar</collectingRegion>
).
</materialsCitation>
</paragraph>
<paragraph id="8BB53697FFD7FFBEFF20FCF4FD72C245" blockId="49.[151,1437,151,1725]" pageId="49" pageNumber="50">
<materialsCitation id="3B623CCAFFD7FFBEFF20FCF4FCCCC5B9" ID-GBIF-Occurrence="1502684997" collectionCode="PHOTO" collectorName="K. J. Singh" country="India" county="Material" location="North Guwahati" municipality="Kamrup" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Assam" typeStatus="holotype">
<emphasis id="B97EEA85FFD7FFBEFF20FCF4FDFAC501" bold="true" box="[199,559,799,824]" pageId="49" pageNumber="50">
<collectingCounty id="62D44E1BFFD7FFBEFF20FCF4FEFAC501" box="[199,303,799,824]" pageId="49" pageNumber="50">Material</collectingCounty>
examined.
<typeStatus id="54B18835FFD7FFBEFE25FCF4FDFAC501" box="[450,559,799,824]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
</emphasis>
(by designation) female with the following labels: 1) [rectangular; white; printed in black ink] “
<emphasis id="B97EEA85FFD7FFBEFE6BFCAFFE0AC564" bold="true" box="[396,479,836,861]" pageId="49" pageNumber="50">
<collectingCountry id="F31D7607FFD7FFBEFE6BFCAFFE0AC564" box="[396,479,836,861]" name="India" pageId="49" pageNumber="50">INDIA</collectingCountry>
</emphasis>
:
<emphasis id="B97EEA85FFD7FFBEFE0AFCAFFDE9C564" bold="true" box="[493,572,836,861]" pageId="49" pageNumber="50">
<collectingRegion id="49CEF875FFD7FFBEFE0AFCAFFDE9C564" box="[493,572,836,861]" country="India" name="Assam" pageId="49" pageNumber="50">Assam</collectingRegion>
</emphasis>
,
<collectingMunicipality id="6BD1ACEDFFD7FFBEFDADFCAEFD7CC565" box="[586,681,837,860]" pageId="49" pageNumber="50">Kamrup</collectingMunicipality>
,
<location id="8ED5604CFFD7FFBEFD54FCAFFCBEC564" LSID="urn:lsid:plazi:treatment:03A38781FFC9FFBEFF70FE37FBC5C084:8ED5604CFFD7FFBEFD54FCAFFCBEC564" box="[691,875,836,861]" country="India" county="Material" municipality="Kamrup" name="North Guwahati" pageId="49" pageNumber="50" stateProvince="Assam">North Guwahati</location>
, ex
<specimenCount id="9D0CFD1EFFD7FFBEFC70FCAEFC7FC565" box="[919,938,837,860]" pageId="49" pageNumber="50" type="male"></specimenCount>
cone
<taxonomicName id="4C0A4D14FFD7FFBEFC00FCAEFB36C565" box="[999,1251,836,860]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis id="B97EEA85FFD7FFBEFC00FCAEFB4FC565" box="[999,1178,837,860]" italics="true" pageId="49" pageNumber="50">Cycas pectinata</emphasis>
, 2008
</taxonomicName>
,
<collectorName id="26FF5341FFD7FFBEFB09FCAEFABAC564" box="[1262,1391,836,861]" pageId="49" pageNumber="50">K. J. Singh</collectorName>
”; 2) [rectangular; pink; printed in bold black ink] “
<collectionCode id="ED1BAE52FFD7FFBEFD4DFC83FCDAC5B9" box="[682,783,872,896]" pageId="49" pageNumber="50">PHOTO</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B623CCAFFD7FFBEFCC1FC83FC66C5D5" ID-GBIF-Occurrence="1502684984" collectionCode="FSCA" collectorName="Deposited" country="India" county="June-July" location="North Guwahati" municipality="Kamrup" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="ASSAM" typeStatus="holotype">
3) [rectangular; white; printed in black ink] “collection date
<collectingCounty id="62D44E1BFFD7FFBEFF34FC67FEEAC59C" box="[211,319,908,933]" pageId="49" pageNumber="50">June-July</collectingCounty>
2007”; 4) [rectangular; white; hand written in black ink] “
<collectingCountry id="F31D7607FFD7FFBEFBE8FC67FBB0C59D" box="[1039,1125,908,932]" name="India" pageId="49" pageNumber="50">INDIA</collectingCountry>
:
<collectingRegion id="49CEF875FFD7FFBEFB92FC67FB0DC59D" box="[1141,1240,908,932]" country="India" name="Assam" pageId="49" pageNumber="50">ASSAM</collectingRegion>
,
<collectingMunicipality id="6BD1ACEDFFD7FFBEFB0DFC66FA9CC59D" box="[1258,1353,909,932]" pageId="49" pageNumber="50">Kamrup</collectingMunicipality>
,
<location id="8ED5604CFFD7FFBEFABCFC67FED2C5F1" LSID="urn:lsid:plazi:treatment:03A38781FFC9FFBEFF70FE37FBC5C084:8ED5604CFFD7FFBEFABCFC67FED2C5F1" country="India" county="June-July" municipality="Kamrup" name="North Guwahati" pageId="49" pageNumber="50" stateProvince="ASSAM">North Guwahati</location>
,
<taxonomicName id="4C0A4D14FFD7FFBEFEFFFC5BFE07C5F1" box="[280,466,943,968]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="pectinata">Cycas pectinata</taxonomicName>
, NG1”; 5) [rectangular; red; printed in black ink] “
<typeStatus id="54B18835FFD7FFBEFBA1FC5BFB09C5F1" box="[1094,1244,944,968]" pageId="49" pageNumber="50" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D0CFD1EFFD7FFBEFB0DFC5BFB2AC5FE" box="[1258,1279,944,967]" pageId="49" pageNumber="50" type="male"></specimenCount>
<taxonomicName id="4C0A4D14FFD7FFBEFAEAFC44FD4DC5D5" authority="P. Skelley, G. Xu &amp; W. Tang 2017" authorityName="P. Skelley, G. Xu &amp; W. Tang" authorityYear="2017" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="B97EEA85FFD7FFBEFAEAFC44FEC0C5D5" italics="true" pageId="49" pageNumber="50">Cycadophila assamensis</emphasis>
P. Skelley, G. Xu &amp; W. Tang 2017
</taxonomicName>
”.
<collectorName id="26FF5341FFD7FFBEFD4AFC3FFCCAC5D4" box="[685,799,980,1005]" pageId="49" pageNumber="50">Deposited</collectorName>
in the
<collectionCode id="ED1BAE52FFD7FFBEFC8EFC3FFC66C5D5" box="[873,947,980,1004]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="49" pageNumber="50">FSCA</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B623CCAFFD7FFBEFC5AFC3FFD04C20D" ID-GBIF-Occurrence="1502684983" collectionCode="FSCA" collectorName="Guwahati &amp; Campus &amp; K. J. Singh &amp; Deposited" country="India" county="June-July" location="North Guwahati" municipality="Kamrup" pageId="49" pageNumber="50" specimenCount="4" specimenCount-male="1" stateProvince="Assam" typeStatus="allotype">
<emphasis id="B97EEA85FFD7FFBEFC5AFC3FFBF7C5D4" bold="true" box="[957,1058,980,1005]" pageId="49" pageNumber="50">
<typeStatus id="54B18835FFD7FFBEFC5AFC3FFBF7C5D4" box="[957,1058,980,1005]" pageId="49" pageNumber="50" type="allotype">Allotype</typeStatus>
</emphasis>
male and
<specimenCount id="9D0CFD1EFFD7FFBEFB72FC3FFACAC5D4" box="[1173,1311,980,1005]" pageId="49" pageNumber="50" type="generic" typeStatus="paratypes">
3
<emphasis id="B97EEA85FFD7FFBEFB4EFC3FFACAC5D4" bold="true" box="[1193,1311,980,1005]" pageId="49" pageNumber="50">paratypes</emphasis>
</specimenCount>
: same data as holotype (
<emphasis id="B97EEA85FFD7FFBEFECFFC1CFE5AC229" bold="true" box="[296,399,1015,1040]" pageId="49" pageNumber="50">
<typeStatus id="54B18835FFD7FFBEFECFFC1CFE5AC229" box="[296,399,1015,1040]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
</emphasis>
&amp;
<emphasis id="B97EEA85FFD7FFBEFE53FC1CFDC6C229" bold="true" box="[436,531,1015,1040]" pageId="49" pageNumber="50">
<typeStatus id="54B18835FFD7FFBEFE53FC1CFDC6C229" box="[436,531,1015,1040]" pageId="49" pageNumber="50" type="allotype">allotype</typeStatus>
</emphasis>
,
<collectionCode id="ED1BAE52FFD7FFBEFDC6FC13FDBFC229" box="[545,618,1016,1040]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="49" pageNumber="50">FSCA</collectionCode>
, 1);
<emphasis id="B97EEA85FFD7FFBEFD7BFC1CFD3BC229" bold="true" box="[668,750,1015,1040]" pageId="49" pageNumber="50">
<collectingCountry id="F31D7607FFD7FFBEFD7BFC1CFD3BC229" box="[668,750,1015,1040]" name="India" pageId="49" pageNumber="50">INDIA</collectingCountry>
</emphasis>
:
<emphasis id="B97EEA85FFD7FFBEFD19FC1CFC98C229" bold="true" box="[766,845,1015,1040]" pageId="49" pageNumber="50">
<collectingRegion id="49CEF875FFD7FFBEFD19FC1CFC98C229" box="[766,845,1015,1040]" country="India" name="Assam" pageId="49" pageNumber="50">Assam</collectingRegion>
</emphasis>
,
<collectorName id="26FF5341FFD7FFBEFCBBFC1CFC1CC229" box="[860,969,1015,1040]" pageId="49" pageNumber="50">Guwahati</collectorName>
IIT
<collectorName id="26FF5341FFD7FFBEFC1BFC13FB8EC229" box="[1020,1115,1016,1040]" pageId="49" pageNumber="50">Campus</collectorName>
, ex
<specimenCount id="9D0CFD1EFFD7FFBEFB6FFC13FB48C236" box="[1160,1181,1016,1039]" pageId="49" pageNumber="50" type="male"></specimenCount>
cone
<taxonomicName id="4C0A4D14FFD7FFBEFB06FC13FF07C20D" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis id="B97EEA85FFD7FFBEFB06FC13FA40C229" box="[1249,1429,1016,1040]" italics="true" pageId="49" pageNumber="50">Cycas pectinata</emphasis>
, 2008
</taxonomicName>
,
<collectorName id="26FF5341FFD7FFBEFF03FBF6FEBBC20C" box="[228,366,1052,1077]" pageId="49" pageNumber="50">K. J. Singh</collectorName>
(2).
<collectorName id="26FF5341FFD7FFBEFE54FBF7FDF1C20C" box="[435,548,1052,1077]" pageId="49" pageNumber="50">Deposited</collectorName>
in the
<collectionCode id="ED1BAE52FFD7FFBEFD60FBF7FD04C20D" box="[647,721,1052,1076]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="49" pageNumber="50">FSCA</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B623CCAFFD7FFBEFD05FBF7FD76C245" ID-GBIF-Occurrence="1502685019" collectingDate="2015-03" collectionCode="FSCA" collectorName="K. J. Singh &amp; Deposited" country="India" county="Others" location="West Champaran" municipality="One" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Bihar" typeStatus="allotype">
<emphasis id="B97EEA85FFD7FFBEFD05FBF7FC6AC20C" bold="true" box="[738,959,1052,1077]" pageId="49" pageNumber="50">
<collectingCounty id="62D44E1BFFD7FFBEFD05FBF7FCE0C20C" box="[738,821,1052,1077]" pageId="49" pageNumber="50">Others</collectingCounty>
examined:
</emphasis>
<collectingMunicipality id="6BD1ACEDFFD7FFBEFC2AFBF7FC29C20D" box="[973,1020,1052,1076]" pageId="49" pageNumber="50">One</collectingMunicipality>
additional adult male collected in association with larvae, currently considered this species:
<emphasis id="B97EEA85FFD7FFBEFCDFFBD4FC5FC261" bold="true" box="[824,906,1087,1112]" pageId="49" pageNumber="50">
<collectingCountry id="F31D7607FFD7FFBEFCDFFBD4FC5FC261" box="[824,906,1087,1112]" name="India" pageId="49" pageNumber="50">INDIA</collectingCountry>
</emphasis>
:
<emphasis id="B97EEA85FFD7FFBEFC7BFBD4FC35C261" bold="true" box="[924,992,1087,1112]" pageId="49" pageNumber="50">
<collectingRegion id="49CEF875FFD7FFBEFC7BFBD4FC35C261" box="[924,992,1087,1112]" country="India" name="Bihar" pageId="49" pageNumber="50">Bihar</collectingRegion>
</emphasis>
,
<location id="8ED5604CFFD7FFBEFC16FBABFB62C261" LSID="urn:lsid:plazi:treatment:03A38781FFC9FFBEFF70FE37FBC5C084:8ED5604CFFD7FFBEFC16FBABFB62C261" box="[1009,1207,1087,1112]" country="India" county="Others" municipality="One" name="West Champaran" pageId="49" pageNumber="50" stateProvince="Bihar">West Champaran</location>
, Valmiki Nat. Park,
<date id="FFB41057FFD7FFBEFF70FB8EFED2C244" box="[151,263,1124,1149]" pageId="49" pageNumber="50" value="2015-03">
<collectingDate id="EFF0E9BFFFD7FFBEFF70FB8EFED2C244" box="[151,263,1124,1149]" pageId="49" pageNumber="50" value="2015-03">Mar 2015</collectingDate>
</date>
,
<collectorName id="26FF5341FFD7FFBEFEF4FB8EFE5BC244" box="[275,398,1124,1149]" pageId="49" pageNumber="50">K. J. Singh</collectorName>
(1).
<collectorName id="26FF5341FFD7FFBEFE23FB8FFDE3C244" box="[452,566,1124,1149]" pageId="49" pageNumber="50">Deposited</collectorName>
in
<collectionCode id="ED1BAE52FFD7FFBEFDBDFB8FFD76C245" box="[602,675,1124,1148]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="49" pageNumber="50">FSCA</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFD7FFBEFF20FB6CFBD8C299" box="[199,1037,1159,1184]" pageId="49" pageNumber="50" type="etymology">
<paragraph id="8BB53697FFD7FFBEFF20FB6CFBD8C299" blockId="49.[151,1437,151,1725]" box="[199,1037,1159,1184]" pageId="49" pageNumber="50">
<emphasis id="B97EEA85FFD7FFBEFF20FB6CFE93C299" bold="true" box="[199,326,1159,1184]" pageId="49" pageNumber="50">Etymology</emphasis>
. Named for the State where the
<typeStatus id="54B18835FFD7FFBEFD57FB63FD35C299" box="[688,736,1160,1184]" pageId="49" pageNumber="50">type</typeStatus>
specimens were collected.
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFD7FFBEFF20FB47FBC5C084" pageId="49" pageNumber="50" type="discussion">
<paragraph id="8BB53697FFD7FFBEFF20FB47FC7EC334" blockId="49.[151,1437,151,1725]" pageId="49" pageNumber="50">
<emphasis id="B97EEA85FFD7FFBEFF20FB47FEE1C2FC" bold="true" box="[199,308,1196,1221]" pageId="49" pageNumber="50">Remarks</emphasis>
. A female
<typeStatus id="54B18835FFD7FFBEFE5DFB47FDC9C2FC" box="[442,540,1196,1221]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
was chosen because it was the most intact specimen available for study and because it shows the sexual dimorphism used to distinguish the species.
<taxonomicName id="4C0A4D14FFD7FFBEFC27FB24FB01C2D1" box="[960,1236,1231,1256]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="B97EEA85FFD7FFBEFC27FB24FB01C2D1" box="[960,1236,1231,1256]" italics="true" pageId="49" pageNumber="50">Cycadophila assamensis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C0A4D14FFD7FFBEFF70FB1EFEF1C334" box="[151,292,1268,1293]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="tansachai">
<emphasis id="B97EEA85FFD7FFBEFF70FB1EFEF1C334" box="[151,292,1268,1293]" italics="true" pageId="49" pageNumber="50">C. tansachai</emphasis>
</taxonomicName>
, differing most notably in abdominal sexual dimorphism.
</paragraph>
<paragraph id="8BB53697FFD7FFBEFF20FAFCFF2FC014" blockId="49.[151,1437,151,1725]" pageId="49" pageNumber="50">
Paucity of Indian materials prevents us from performing certain analyses and answering certain questions. Adult morphology allow us to distinguish the populations from
<collectingCountry id="F31D7607FFD7FFBEFC62FAD7FC6BC36C" box="[901,958,1340,1365]" name="India" pageId="49" pageNumber="50">India</collectingCountry>
and propose a name for future research. Larvae from
<collectingRegion id="49CEF875FFD7FFBEFECFFAB4FEB3C341" box="[296,358,1375,1400]" country="India" name="Bihar" pageId="49" pageNumber="50">Bihar</collectingRegion>
are also morphologically distinguishable from
<emphasis id="B97EEA85FFD7FFBEFC90FA8BFC5FC34E" box="[887,906,1376,1399]" italics="true" pageId="49" pageNumber="50">C</emphasis>
. (
<emphasis id="B97EEA85FFD7FFBEFC46FAB4FBE6C341" box="[929,1075,1375,1400]" italics="true" pageId="49" pageNumber="50">Strobilophila</emphasis>
) larvae collected in
<collectingCountry id="F31D7607FFD7FFBEFAF1FAB4FAAFC341" box="[1302,1402,1375,1400]" name="Thailand" pageId="49" pageNumber="50">Thailand</collectingCountry>
by the posteriad-tilting orientation of their t1 tubercle (see Key to larvae above) vs. the vertically oriented t1 turbercle in Thai specimens. While the
<collectingRegion id="49CEF875FFD7FFBEFE10FA43FD91C3F9" box="[503,580,1448,1472]" country="India" name="Assam" pageId="49" pageNumber="50">Assam</collectingRegion>
and
<collectingRegion id="49CEF875FFD7FFBEFD66FA4CFD6AC3F9" box="[641,703,1447,1472]" country="India" name="Bihar" pageId="49" pageNumber="50">Bihar</collectingRegion>
populations are presented here as conspecific, we have doubts requiring further analyses. Thus, the
<typeStatus id="54B18835FFD7FFBEFDA4FA26FDA6C3DC" box="[579,627,1485,1509]" pageId="49" pageNumber="50">type</typeStatus>
series is limited to the
<collectingRegion id="49CEF875FFD7FFBEFC6EFA27FC00C3DD" box="[905,981,1484,1508]" country="India" name="Assam" pageId="49" pageNumber="50">Assam</collectingRegion>
population where we are able to make morphological comparisons with all known species. As noted above, molecular analyses failed with available material.
</paragraph>
<paragraph id="8BB53697FFD7FFBEFF20F9DCFBC5C084" blockId="49.[151,1437,151,1725]" pageId="49" pageNumber="50">
The host for this species in
<collectingRegion id="49CEF875FFD7FFBEFE10F9D3FD93C069" box="[503,582,1592,1616]" country="India" name="Assam" pageId="49" pageNumber="50">Assam</collectingRegion>
,
<taxonomicName id="4C0A4D14FFD7FFBEFDB7F9D3FCD6C069" box="[592,771,1592,1616]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis id="B97EEA85FFD7FFBEFDB7F9D3FCD6C069" box="[592,771,1592,1616]" italics="true" pageId="49" pageNumber="50">Cycas pectinata</emphasis>
</taxonomicName>
, occurs at atypically low elevations for the species (
<quantity id="4CF29B72FFD7FFBEFAA0F9DCFF65C04D" metricMagnitude="2" metricUnit="m" metricValue="1.22" metricValueMax="1.92" metricValueMin="0.52" pageId="49" pageNumber="50" unit="m" value="122.0" valueMax="192.0" valueMin="52.0">52192 m</quantity>
; see
<bibRefCitation id="EF9B4B66FFD7FFBEFF0DF9B7FE0DC04D" author="Singh" box="[234,472,1628,1653]" pageId="49" pageNumber="50" refString="Singh, K. J. &amp; Singh, R. (2014) Population assessment and distribution of Cycas pectinata" type="book" year="2014">Singh &amp; Singh 2014</bibRefCitation>
) vs. over
<quantity id="4CF29B72FFD7FFBEFDABF9B7FD40C04D" box="[588,661,1628,1653]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="49" pageNumber="50" unit="m" value="500.0">500 m</quantity>
for other localities of
<taxonomicName id="4C0A4D14FFD7FFBEFC7EF9B6FB9AC04D" box="[921,1103,1629,1652]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis id="B97EEA85FFD7FFBEFC7EF9B6FB9AC04D" box="[921,1103,1629,1652]" italics="true" pageId="49" pageNumber="50">Cycas pectinata</emphasis>
</taxonomicName>
in the states of
<collectingRegion id="49CEF875FFD7FFBEFAEDF9B7FABFC04C" box="[1290,1386,1628,1653]" country="India" name="Manipur" pageId="49" pageNumber="50">Manipur</collectingRegion>
and
<collectingRegion id="49CEF875FFD7FFBEFF70F96AFEFEC0A1" box="[151,299,1663,1688]" country="India" name="West Bengal" pageId="49" pageNumber="50">West Bengal</collectingRegion>
,
<collectingCountry id="F31D7607FFD7FFBEFEDDF994FEA1C0A1" box="[314,372,1663,1688]" name="India" pageId="49" pageNumber="50">India</collectingCountry>
or in
<collectingCountry id="F31D7607FFD7FFBEFE26F994FDF0C0A1" box="[449,549,1663,1688]" name="Thailand" pageId="49" pageNumber="50">Thailand</collectingCountry>
and provides evidence that this beetle is geographically isolated from other populations of the
<emphasis id="B97EEA85FFD7FFBEFE8FF94FFE2FC084" box="[360,506,1700,1725]" italics="true" pageId="49" pageNumber="50">Strobilophila</emphasis>
complex. See also remarks under
<emphasis id="B97EEA85FFD7FFBEFC9CF94EFBC5C084" box="[891,1040,1700,1725]" italics="true" pageId="49" pageNumber="50">
<taxonomicName id="4C0A4D14FFD7FFBEFC9CF94EFBD9C084" box="[891,1036,1700,1725]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="kwaiensis">C. kwaiensis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>