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<document id="23D9775AD86F658A85BC535DEC533CE4" ID-DOI="10.11646/phytotaxa.218.3.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13636012" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725333359211" checkinUser="felipe" docAuthor="Egan, Ashley N. &amp; Pan, Bo" docDate="2015" docId="039787AE392A8462FF6A5CC60CF2F8B2" docLanguage="en" docName="phytotaxa.218.3.1.pdf" docOrigin="Phytotaxa 218 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.218.3.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Neustanthus Bentham, Pl. Jungh." docType="treatment" docVersion="2" lastPageNumber="204" masterDocId="FFAEFFD639288461FFE25A300D60FFBF" masterDocTitle="Resolution of polyphyly in Pueraria (Leguminosae, Papilionoideae): The creation of two new genera, Haymondia and Toxicopueraria, the resurrection of Neustanthus, and a new combination in Teyleria" masterLastPageNumber="226" masterPageNumber="201" pageNumber="203" updateTime="1725334893390" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B4145CDAAD9A64077A829822A0BFA7AA">Resolution of polyphyly in Pueraria (Leguminosae, Papilionoideae): The creation of two new genera, Haymondia and Toxicopueraria, the resurrection of Neustanthus, and a new combination in Teyleria</mods:title>
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<mods:namePart id="C6DDC7265D23A4580DD6405ABAFBEE54">Egan, Ashley N.</mods:namePart>
<mods:affiliation id="456BA97852684A78046B58E97C490648">US National Herbarium (US), Department of Botany, Smithsonian Institution-NMNH, MRC 166, 10 and Constitution Ave, Washington DC, 20560, USA</mods:affiliation>
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<mods:namePart id="62953A7C18FFE965CC90A4FB9F187E61">Pan, Bo</mods:namePart>
<mods:affiliation id="667E363A5897DA3C0800E6745186477C">Centre for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Menglun, Mengla, Yunnan 666303, China</mods:affiliation>
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<mods:title id="33D7E979D007E1A6678EC8F54E14132E">Phytotaxa</mods:title>
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<mods:identifier id="3DDA9C5C548CA59852C5FD541FC262D1" type="ISSN">1179-3163</mods:identifier>
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<treatment id="039787AE392A8462FF6A5CC60CF2F8B2" ID-DOI="http://doi.org/10.5281/zenodo.13636006" ID-Zenodo-Dep="13636006" LSID="urn:lsid:plazi:treatment:039787AE392A8462FF6A5CC60CF2F8B2" httpUri="http://treatment.plazi.org/id/039787AE392A8462FF6A5CC60CF2F8B2" lastPageId="3" lastPageNumber="204" pageId="2" pageNumber="203">
<subSubSection id="C3246533392A8463FF6A5CC60FB7F8B0" box="[136,727,1782,1807]" pageId="2" pageNumber="203" type="nomenclature">
<paragraph id="8B8136B8392A8463FF6A5CC60FB7F8B0" blockId="2.[136,727,1782,1807]" box="[136,727,1782,1807]" pageId="2" pageNumber="203">
<heading id="D0C981D4392A8463FF6A5CC60FB7F8B0" box="[136,727,1782,1807]" fontSize="11" level="2" pageId="2" pageNumber="203" reason="3">
<taxonomicName id="4C3E4D3B392A8463FF6A5CC60F6CF8B0" authority="Bentham, Pl. Jungh." authorityName="Bentham, Pl. Jungh." box="[136,524,1782,1807]" class="Magnoliopsida" family="Fabaceae" genus="Neustanthus" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="203" phylum="Magnoliopsida" rank="genus">
<emphasis id="B94AEAAA392A8463FF6A5CC60CF5F8B0" bold="true" box="[136,405,1782,1807]" pageId="2" pageNumber="203">
<emphasis id="B94AEAAA392A8463FF6A5CC60C7AF8B0" bold="true" box="[136,282,1782,1807]" italics="true" pageId="2" pageNumber="203">Neustanthus</emphasis>
Bentham,
</emphasis>
Pl. Jungh.
</taxonomicName>
2: 234235. 1852
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3246533392A8463FF6A5D700E84F8EA" box="[136,996,1856,1877]" pageId="2" pageNumber="203" type="type_taxon">
<paragraph id="8B8136B8392A8463FF6A5D700E84F8EA" blockId="2.[136,996,1856,1877]" box="[136,996,1856,1877]" pageId="2" pageNumber="203">
<emphasis id="B94AEAAA392A8463FF6A5D700C6FF8EA" bold="true" box="[136,271,1856,1877]" pageId="2" pageNumber="203">
<typeStatus id="5485881A392A8463FF6A5D700DDAF8EA" box="[136,186,1856,1877]" pageId="2" pageNumber="203">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C3E4D3B392A8463FEC55D700E00F8EA" authority="(Roxburgh) Bentham, Pl. Jungh." authorityName="Bentham, Pl. Jungh." baseAuthorityName="Roxburgh" box="[295,864,1856,1877]" class="Magnoliopsida" family="Fabaceae" genus="Neustanthus" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="203" phylum="Magnoliopsida" rank="species" species="phaseoloides">
<emphasis id="B94AEAAA392A8463FEC55D700F41F8EA" box="[295,545,1856,1877]" italics="true" pageId="2" pageNumber="203">Neustanthus phaseoloides</emphasis>
(Roxburgh) Bentham, Pl. Jungh.
</taxonomicName>
2: 235. 1852.
</paragraph>
</subSubSection>
<subSubSection id="C3246533392A8462FF6A5DB60C55FEBE" lastPageId="3" lastPageNumber="204" pageId="2" pageNumber="203" type="diagnosis">
<paragraph id="8B8136B8392A8462FF6A5DB60C55FEBE" blockId="2.[136,1452,1926,2059]" lastBlockId="3.[136,1452,160,1805]" lastPageId="3" lastPageNumber="204" pageId="2" pageNumber="203">
<emphasis id="B94AEAAA392A8463FF6A5DB60C65F820" bold="true" box="[136,261,1926,1951]" pageId="2" pageNumber="203">Diagnosis:</emphasis>
—Perennial herbaceous climber or creeper. Stems hirsute with spreading hairs. Leaves pinnately trifoliolate. Leaflets entire or lobed. Stipules basifixed, open and reflexed. Nodose (hereinafter refers to a highly compacted inflorescence branch or brachyblast; e.g.
<figureCitation id="13052A3D392A8463FDBE5DFE0FABF858" box="[604,715,1998,2023]" captionStart="FIGURE 1" captionStartId="4.[136,229,1922,1943]" captionTargetBox="[190,1394,195,1891]" captionTargetId="figure-15@4.[190,1398,190,1897]" captionTargetPageId="4" captionText="FIGURE 1. Neustanthus phaseoloides. A. Habit, Inflorescence. B. Upper leaf surface (detail). C. Lower leaf surface (detail). D. Stipules.E. Stipels.F.Calyx, inner view.G. Flower, lateral view. H. Flower, face view.I. Vexillum. J. Wing petal.K. Keel.L. Androecium, lateral view, surrounding stigma and apex of style. M. Fused stamens and free stamen. N. Gynoecium. O. Fruit. P.Nodes (brachyblast) of pseudoraceme. Q. Seed, lateral view. R. Seed, dorsal view. Voucher used for illustrations: AE: A.N. Egan &amp; Xu Bo 13-0817 (US); FN: A.N. Egan et al. 13-0715 (US); OR: Shiu Ying Hu 5764 (US). Illustrations by Alice R. Tangerini." figureDoi="http://doi.org/10.5281/zenodo.13636014" httpUri="https://zenodo.org/record/13636014/files/figure.png" pageId="2" pageNumber="203">Figure 1P</figureCitation>
) pseudoracemes axillary, several flowers clustered at each node. Calyx 5-lobed, the lower lobe longer, the upper two connate from the middle. Corolla blue or purple or white with purple to pink stripe. Vexillum without callosities, apex slightly emarginate, base with incurved auricles. Vexillary stamen connate to the tube. Ovules many. Fruits subterete, valves twisting upon dehiscence. Seeds barrel-shaped to oblong-elliptic.
</paragraph>
</subSubSection>
<subSubSection id="C3246533392B8462FF5F5B3C0C1DFB16" pageId="3" pageNumber="204" type="description">
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<emphasis id="B94AEAAA392B8462FF5F5B3C0C31FE9A" bold="true" box="[189,337,268,293]" pageId="3" pageNumber="204">Description:</emphasis>
—Perennial herbaceous climber or creeper. Stems slender but strong, fibrous, up to
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long, rooting at the nodes, pubescent with spreading hairs. Stipules basifixed, triangular to ovate,
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×
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, acuminate, deciduous or persistent. Leaves pinnately trifoliolate; petiole
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long, somewhat pubescent with adpressed and spreading hairs; leaflets broadly ovate, rhomboid, or ovate-rhomboid, the terminal one broader and symmetrical, entire or lobed,
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, margin entire or sinuate, lateral leaflets sometimes oblique,
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, mucronate, leaflets adpressed pubescent above, more densely below, veins conspicuous below, strongly pubescent, 3 from the base, 5 pairs of primary lateral veins, opposite or not; petiolules
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long with spreading hairs; stipels
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long, linear to lanceolate, setaceous. Inflorescences solitary, axillary pseudoracemes, unbranched, (4)10 35(45) cm long, nodose (with brachyblasts), flowering above the middle with 4 or more flowers per node; bracts subtending the nodes,
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long, caducous; pedicels
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, pubescent; bracteoles 2 per flower, lanceolate,
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. Calyx
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long, hirsute on the outside, the tube
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long, 5-lobed, the upper 2 lobes fused along half of their length or more but not entirely fused,
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long, lateral lobes obtuse to acute,
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long, the lower lobe acute to lanceolate-acuminate or subulate, longer than the others,
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long. Corolla bluish-purple or white suffused with purple, pink, or bluish-purple, vexillum obovate, often with green or yellow patch at the base,
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, base with 2 incurved auricles, without callosities; wing petals obovate-oblong, bluish-purple or white suffused with bluish-purple, slightly longer than the keel,
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, one side of base with rounded auricle, claw slender,
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long, basal margin lobed; keel petals falcate, bluish-purple or white with blue, green or purple tip,
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long, base truncate, clawed, the claw
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long. Ovary linear, hirsute,
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long with ca. 20 ovules; style glabrous,
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long, curving upward; stigma terminal, globose. Stamens diadelphous, the vexillary stamen adherent near the middle to the staminal column in bud but becoming free with age,
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long; anthers basifixed, alternating on long and short filaments. Fruits leguminous pods, subterete, grey to black, first adpressed hirsute, glabrescent,
<quantity id="4CC69B5D392B8462FE055E300F29FBA6" box="[487,585,1024,1049]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="3" pageNumber="204" unit="cm" value="8.5" valueMax="12.0" valueMin="5.0">512 cm</quantity>
×
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, valves twisting upon dehiscence, with papery partitions between the seeds. Seeds 1520 per fruit, barrel-shaped to oblong-elliptic,
<quantity id="4CC69B5D392B8462FC865E140EB5FB82" box="[868,981,1060,1085]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="3" pageNumber="204" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.54 mm</quantity>
×
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,
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thick; funicle short, deltoid; arils elongate.
</paragraph>
<paragraph id="8B8136B8392B8462FF5F5E5C0C1DFB16" blockId="3.[136,1452,160,1805]" pageId="3" pageNumber="204">
<emphasis id="B94AEAAA392B8462FF5F5E5C0C23FB3A" bold="true" box="[189,323,1132,1157]" pageId="3" pageNumber="204">Phenology:</emphasis>
—Flowering and fruiting various, dependent upon locality (see
<tableCitation id="C6BC0303392B8462FBF65E5C0908FB3A" box="[1044,1128,1132,1157]" captionStart="TABLE 2" captionStartId="2.[136,216,486,507]" captionText="TABLE 2. Character state comparisons between Pueraria s.s., the resurrected Neustanthus, Teyleria, Toxicopueraria gen. nov. and Haymondia gen. nov." pageId="3" pageNumber="204">Table 2</tableCitation>
of
<bibRefCitation id="EFAF4B49392B8462FB6D5E5C08E3FB3A" author="van der Maesen, L. J. G." box="[1167,1411,1132,1157]" pageId="3" pageNumber="204" pagination="1 - 130" refId="ref12839" refString="van der Maesen, L. J. G. (1985) Revision of the genus Pueraria DC with some notes on Teyleria Backer (Leguminosae). Wageningen Agricultural University Papers 85: 1 - 130." type="journal article" year="1985">van der Maesen 1985</bibRefCitation>
for detailed information).
</paragraph>
</subSubSection>
<subSubSection id="C3246533392B8462FF5F5E84081AFA86" pageId="3" pageNumber="204" type="distribution">
<paragraph id="8B8136B8392B8462FF5F5E84090CFAAA" blockId="3.[136,1452,160,1805]" pageId="3" pageNumber="204">
<emphasis id="B94AEAAA392B8462FF5F5E840C39FB72" bold="true" box="[189,345,1204,1229]" pageId="3" pageNumber="204">Distribution:</emphasis>
—Native to southern, eastern, and Southeast Asia:
<collectingCountry id="F3297628392B8462FC9C5E840E88FB72" box="[894,1000,1204,1229]" name="Australia" pageId="3" pageNumber="204">Australia</collectingCountry>
,
<collectingCountry id="F3297628392B8462FC125E840916FB72" box="[1008,1142,1204,1229]" name="Bangladesh" pageId="3" pageNumber="204">Bangladesh</collectingCountry>
,
<collectingCountry id="F3297628392B8462FB9C5E8409B2FB72" box="[1150,1234,1204,1229]" name="Bhutan" pageId="3" pageNumber="204">Bhutan</collectingCountry>
,
<collectingCountry id="F3297628392B8462FB385E840849FB72" box="[1242,1321,1204,1229]" name="Brunei" pageId="3" pageNumber="204">Brunei</collectingCountry>
,
<collectingCountry id="F3297628392B8462FAD35E8408C7FB72" box="[1329,1447,1204,1229]" name="Cambodia" pageId="3" pageNumber="204">Cambodia</collectingCountry>
,
<collectingCountry id="F3297628392B8462FF6A5EE80DAEFB4E" box="[136,206,1240,1265]" name="China" pageId="3" pageNumber="204">China</collectingCountry>
,
<collectingCountry id="F3297628392B8462FF3B5EE80C75FB4E" box="[217,277,1240,1265]" name="India" pageId="3" pageNumber="204">India</collectingCountry>
,
<collectingCountry id="F3297628392B8462FEC25EE80CF0FB4E" box="[288,400,1240,1265]" name="Indonesia" pageId="3" pageNumber="204">Indonesia</collectingCountry>
,
<collectingCountry id="F3297628392B8462FE795EE80CB4FB4E" box="[411,468,1240,1265]" name="Laos" pageId="3" pageNumber="204">Laos</collectingCountry>
,
<collectingCountry id="F3297628392B8462FE3D5EE80F28FB4E" box="[479,584,1240,1265]" name="Malaysia" pageId="3" pageNumber="204">Malaysia</collectingCountry>
,
<collectingCountry id="F3297628392B8462FDB65EE80FA3FB4E" box="[596,707,1240,1265]" name="Myanmar" pageId="3" pageNumber="204">Myanmar</collectingCountry>
, N. Borneo,
<collectingCountry id="F3297628392B8462FCB25EE80EF6FB4E" box="[848,918,1240,1265]" name="Nepal" pageId="3" pageNumber="204">Nepal</collectingCountry>
, New
<collectingCountry id="F3297628392B8462FC3E5EE80950FB4E" box="[988,1072,1240,1265]" name="Guinea-Bissau" pageId="3" pageNumber="204">Guinea</collectingCountry>
,
<collectingCountry id="F3297628392B8462FBD95EE809DCFB4E" box="[1083,1212,1240,1265]" name="Philippines" pageId="3" pageNumber="204">Philippines</collectingCountry>
,
<collectingCountry id="F3297628392B8462FB255EE8085BFB4E" box="[1223,1339,1240,1265]" name="Singapore" pageId="3" pageNumber="204">Singapore</collectingCountry>
,
<collectingCountry id="F3297628392B8462FAA45EE80DBAFAAA" name="Solomon Islands" pageId="3" pageNumber="204">Solomon Islands</collectingCountry>
,
<collectingCountry id="F3297628392B8462FF075ECC0C36FAAA" box="[229,342,1276,1301]" name="Sri Lanka" pageId="3" pageNumber="204">Sri Lanka</collectingCountry>
,
<collectingCountry id="F3297628392B8462FE835ECC0CD6FAAA" box="[353,438,1276,1301]" name="Taiwan" pageId="3" pageNumber="204">Taiwan</collectingCountry>
,
<collectingCountry id="F3297628392B8462FE225ECC0F46FAAA" box="[448,550,1276,1301]" name="Thailand" pageId="3" pageNumber="204">Thailand</collectingCountry>
,
<collectingCountry id="F3297628392B8462FDD35ECC0FF4FAAA" box="[561,660,1276,1301]" name="Vietnam" pageId="3" pageNumber="204">Vietnam</collectingCountry>
. Introduced widely to other tropical areas.
</paragraph>
<paragraph id="8B8136B8392B8462FF5F5F10081AFA86" blockId="3.[136,1452,160,1805]" box="[189,1402,1310,1338]" pageId="3" pageNumber="204">
<emphasis id="B94AEAAA392B8462FF5F5F100CB9FA85" bold="true" box="[189,473,1310,1338]" pageId="3" pageNumber="204">Vernacular:—ΞṘḍƀ</emphasis>
san lie ye ge (Chinese), thua sian pa (Thai), phak phit (Thai).
</paragraph>
</subSubSection>
<subSubSection id="C3246533392B8462FF5F5F740CF2F8B2" pageId="3" pageNumber="204" type="discussion">
<paragraph id="8B8136B8392B8462FF5F5F740CF2F8B2" blockId="3.[136,1452,160,1805]" pageId="3" pageNumber="204">
<emphasis id="B94AEAAA392B8462FF5F5F740C25FAE2" bold="true" box="[189,325,1348,1373]" pageId="3" pageNumber="204">Discussion:</emphasis>
<taxonomicName id="4C3E4D3B392B8462FE835F740C8BFAE2" box="[353,491,1348,1373]" class="Magnoliopsida" family="Fabaceae" genus="Neustanthus" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="204" phylum="Magnoliopsida" rank="genus">
<emphasis id="B94AEAAA392B8462FE835F740C8BFAE2" box="[353,491,1348,1373]" italics="true" pageId="3" pageNumber="204">Neustanthus</emphasis>
</taxonomicName>
is phylogenetically close to
<taxonomicName id="4C3E4D3B392B8462FCCC5F740EDFFAE2" box="[814,959,1348,1373]" class="Magnoliopsida" family="Fabaceae" genus="Sinodolichos" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEAAA392B8462FCCC5F740EDFFAE2" box="[814,959,1348,1373]" italics="true" pageId="3" pageNumber="204">Sinodolichos</emphasis>
</taxonomicName>
(Egan
<emphasis id="B94AEAAA392B8462FBF25F740929FAE2" box="[1040,1097,1348,1373]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
, in prep.;
<bibRefCitation id="EFAF4B49392B8462FB5F5F740822FAE2" author="Cagle, W." box="[1213,1346,1348,1373]" pageId="3" pageNumber="204" refId="ref11857" refString="Cagle, W. (2013) Parsing Polyphyletic Pueraria: Delimiting distinct evolutionary lineages through phylogeny. Greenville, North Carolina, U. S. A., 293 pp." type="book" year="2013">Cagle 2013</bibRefCitation>
), a genus of similar geographic distribution. Both genera are perennial, herbaceous climbers or creepers, with spreading hirsute hairs and basifixed stipules. Both have long, acuminate calyx lobes and linear, somewhat compressed, subterete fruit. However, they differ in their inflorescence structure, with
<taxonomicName id="4C3E4D3B392B8462FCF55F800EC1FA76" box="[791,929,1456,1481]" class="Magnoliopsida" family="Fabaceae" genus="Neustanthus" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="204" phylum="Magnoliopsida" rank="genus">
<emphasis id="B94AEAAA392B8462FCF55F800EC1FA76" box="[791,929,1456,1481]" italics="true" pageId="3" pageNumber="204">Neustanthus</emphasis>
</taxonomicName>
having a pseudoraceme whereas
<taxonomicName id="4C3E4D3B392B8462FAF95F8008CCFA76" box="[1307,1452,1456,1481]" class="Magnoliopsida" family="Fabaceae" genus="Sinodolichos" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEAAA392B8462FAF95F8008CCFA76" box="[1307,1452,1456,1481]" italics="true" pageId="3" pageNumber="204">Sinodolichos</emphasis>
</taxonomicName>
has a true raceme.
<taxonomicName id="4C3E4D3B392B8462FE855FE40C91FA52" box="[359,497,1492,1517]" class="Magnoliopsida" family="Fabaceae" genus="Neustanthus" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="204" phylum="Magnoliopsida" rank="genus">
<emphasis id="B94AEAAA392B8462FE855FE40C91FA52" box="[359,497,1492,1517]" italics="true" pageId="3" pageNumber="204">Neustanthus</emphasis>
</taxonomicName>
has ca. 20 seeds per pod that are usually barrel-shaped whereas
<taxonomicName id="4C3E4D3B392B8462FB095FE4081CFA52" box="[1259,1404,1492,1517]" class="Magnoliopsida" family="Fabaceae" genus="Sinodolichos" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEAAA392B8462FB095FE4081CFA52" box="[1259,1404,1492,1517]" italics="true" pageId="3" pageNumber="204">Sinodolichos</emphasis>
</taxonomicName>
has 310 seeds that are oblong. The most recent treatment of
<taxonomicName id="4C3E4D3B392B8462FCEC5FC80EDBF9AE" box="[782,955,1528,1553]" class="Magnoliopsida" family="Fabaceae" genus="Neustanthus" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="204" phylum="Magnoliopsida" rank="species" species="phaseoloides">
<emphasis id="B94AEAAA392B8462FCEC5FC80EDBF9AE" box="[782,955,1528,1553]" italics="true" pageId="3" pageNumber="204">P. phaseoloides</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFAF4B49392B8462FC045FC8098AF9AE" author="van der Maesen, L. J. G." box="[998,1258,1528,1553]" pageId="3" pageNumber="204" pagination="1 - 28" refId="ref12946" refString="van der Maesen, L. J. G. (2002) Pueraria: botanical characteristics. In: Keung, W. M. (Ed.) Pueraria: The genus Pueraria. Taylor &amp; Francis, New York City, NY, pp. 1 - 28." type="book chapter" year="2002">van der Maesen (2002)</bibRefCitation>
recognized three varieties: var.
<taxonomicName id="4C3E4D3B392B8462FEC65C2C0F32F98A" box="[292,594,1564,1589]" class="Magnoliopsida" family="Fabaceae" genus="Neustanthus" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="204" phylum="Magnoliopsida" rank="variety" species="phaseoloides" variety="javanica">
<emphasis id="B94AEAAA392B8462FEC65C2C0CD6F98A" box="[292,438,1564,1589]" italics="true" pageId="3" pageNumber="204">phaseoloides</emphasis>
, var.
<emphasis id="B94AEAAA392B8462FE105C2C0F32F98A" box="[498,594,1564,1589]" italics="true" pageId="3" pageNumber="204">javanica</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C3E4D3B392B8462FD6C5C2C0E56F98A" box="[654,822,1564,1589]" class="Magnoliopsida" family="Fabaceae" genus="Neustanthus" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="204" phylum="Magnoliopsida" rank="variety" species="phaseoloides" variety="subspicata">
var.
<emphasis id="B94AEAAA392B8462FD5C5C2C0E56F98A" box="[702,822,1564,1589]" italics="true" pageId="3" pageNumber="204">subspicata</emphasis>
</taxonomicName>
. These taxa have been variably recognized at the species level (e.g. as
<taxonomicName id="4C3E4D3B392B8462FEFD5C700F7CF9E6" box="[287,540,1600,1625]" class="Magnoliopsida" family="Fabaceae" genus="Neustanthus" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="204" phylum="Magnoliopsida" rank="species" species="javanicus">
<emphasis id="B94AEAAA392B8462FEFD5C700F7CF9E6" box="[287,540,1600,1625]" italics="true" pageId="3" pageNumber="204">Neustanthus javanicus</emphasis>
</taxonomicName>
by Bentham (1832)), varietal levels (e.g.
<bibRefCitation id="EFAF4B49392B8462FC175C700992F9E6" author="van der Maesen, L. J. G." box="[1013,1266,1600,1625]" pageId="3" pageNumber="204" pagination="1 - 130" refId="ref12839" refString="van der Maesen, L. J. G. (1985) Revision of the genus Pueraria DC with some notes on Teyleria Backer (Leguminosae). Wageningen Agricultural University Papers 85: 1 - 130." type="journal article" year="1985">van der Maesen, 1985</bibRefCitation>
,
<bibRefCitation id="EFAF4B49392B8462FB1F5C700858F9E6" author="van der Maesen, L. J. G." box="[1277,1336,1600,1625]" pageId="3" pageNumber="204" pagination="55 - 86" refId="ref12878" refString="van der Maesen, L. J. G. (1994) Pueraria, the Kudzu and its relatives, an update of the taxonomy. In: Sorensen, M. (Ed.) Proceedings of the First International Symposium on Tuberous Legumes, Gualdeloupe, F. W. I. DSR Boghandel, Fredericksberg, Denmark, pp. 55 - 86." type="book chapter" year="1994">1994</bibRefCitation>
,
<bibRefCitation id="EFAF4B49392B8462FAA65C7008E0F9E6" author="van der Maesen, L. J. G." box="[1348,1408,1600,1625]" pageId="3" pageNumber="204" pagination="1 - 28" refId="ref12946" refString="van der Maesen, L. J. G. (2002) Pueraria: botanical characteristics. In: Keung, W. M. (Ed.) Pueraria: The genus Pueraria. Taylor &amp; Francis, New York City, NY, pp. 1 - 28." type="book chapter" year="2002">2002</bibRefCitation>
), or in synonymy under
<taxonomicName id="4C3E4D3B392B8462FE855C540F74F9C2" box="[359,532,1636,1661]" class="Magnoliopsida" family="Fabaceae" genus="Neustanthus" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="204" phylum="Magnoliopsida" rank="species" species="phaseoloides">
<emphasis id="B94AEAAA392B8462FE855C540F74F9C2" box="[359,532,1636,1661]" italics="true" pageId="3" pageNumber="204">P. phaseoloides</emphasis>
</taxonomicName>
. A comprehensive sampling coupled with morphometric and/or molecular genetic work will be necessary to determine the best level at which to recognize these taxa. Pending further investigation, we choose to continue with the current recognition of these taxa at the varietal level. Resurrected names and new combinations are presented below. For a full list of synonyms and a key to the varieties and descriptions thereof, see
<bibRefCitation id="EFAF4B49392B8462FF6A5CC40CEFF8B2" author="van der Maesen, L. J. G." box="[136,399,1780,1805]" pageId="3" pageNumber="204" pagination="1 - 130" refId="ref12839" refString="van der Maesen, L. J. G. (1985) Revision of the genus Pueraria DC with some notes on Teyleria Backer (Leguminosae). Wageningen Agricultural University Papers 85: 1 - 130." type="journal article" year="1985">van der Maesen (1985)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>