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<document id="1444E7AEAB6EAA138370CDCE078BBCB4" ID-DOI="10.11646/zootaxa.3920.1.1" ID-GBIF-Dataset="7289c86d-0ed1-4e93-b7b3-bc596042d854" ID-ISSN="1175-5326" ID-Zenodo-Dep="287886" ID-ZooBank="025F9918-F9F6-4049-B7CD-5AC632094F5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460994190417" checkinUser="plazi" docAuthor="Kataev, Boris M. &amp; Liang, Hongbin" docDate="2015" docId="039787812E5AFFD994FD4D1CEAF7E95D" docLanguage="en" docName="zt03920p039.pdf" docOrigin="Zootaxa 3920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpalus (Pseudoophonus) sericatus Tschitscherine 1906" docType="treatment" docVersion="7" lastPageNumber="25" masterDocId="FFAEFFF92E4DFFC1946A4A2BE927EF1A" masterDocTitle="Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)" masterLastPageNumber="39" masterPageNumber="1" pageNumber="24" updateTime="1698562432667" updateUser="plazi">
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<mods:title id="4ECC053B79B937B8E11028E91CE86B3D">Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="3F4C4937A34EACB1BE1F9ABC3B8F6113">Kataev, Boris M.</mods:namePart>
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<mods:namePart id="958CDF82639778DD00A5864B5A001F1B">Liang, Hongbin</mods:namePart>
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<mods:title id="18054EF3AB8068D9933551B01A07F927">Zootaxa</mods:title>
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<mods:date id="37E186DD5DD53F5C24432F1B69C97BF0">2015</mods:date>
<mods:detail id="87764C0286CDD8E045A135338EEE647B" type="volume">
<mods:number id="6E4BE4754AEEEAF12BF22166CFD3780D">3920</mods:number>
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<mods:identifier id="A40197FD4E4AF18365309D23FB5F0482" type="DOI">10.11646/zootaxa.3920.1.1</mods:identifier>
<mods:identifier id="438F25A086C34BD387947A07A713742B" type="GBIF-Dataset">7289c86d-0ed1-4e93-b7b3-bc596042d854</mods:identifier>
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<treatment id="039787812E5AFFD994FD4D1CEAF7E95D" ID-DOI="http://doi.org/10.5281/zenodo.6114280" ID-GBIF-Taxon="127671643" ID-Zenodo-Dep="6114280" LSID="urn:lsid:plazi:treatment:039787812E5AFFD994FD4D1CEAF7E95D" httpUri="http://treatment.plazi.org/id/039787812E5AFFD994FD4D1CEAF7E95D" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<subSubSection id="C324651C2E5AFFD694FD4D1CE800E869" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8B8136972E5AFFD694FD4D1CEA7CE848" blockId="23.[151,859,1847,1907]" box="[151,859,1847,1874]" pageId="23" pageNumber="24">
<heading id="D0C981FB2E5AFFD694FD4D1CEA7CE848" bold="true" box="[151,859,1847,1874]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<taxonomicName id="4C3E4D142E5AFFD694FD4D1CEA7CE848" ID-CoL="6LDTH" authority="Tschitscherine, 1906" authorityName="Tschitscherine" authorityYear="1906" box="[151,859,1847,1874]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="sericatus" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E5AFFD694FD4D1CEA7CE848" bold="true" box="[151,859,1847,1874]" pageId="23" pageNumber="24">
<emphasis id="B94AEA852E5AFFD694FD4D1CE82EE84B" bold="true" box="[151,265,1847,1873]" italics="true" pageId="23" pageNumber="24">Harpalus</emphasis>
(
<emphasis id="B94AEA852E5AFFD695704D1CE8FDE84B" bold="true" box="[282,474,1847,1873]" italics="true" pageId="23" pageNumber="24">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E5AFFD695814D1CEB70E84B" bold="true" box="[491,599,1847,1873]" italics="true" pageId="23" pageNumber="24">sericatus</emphasis>
<bibRefCitation id="EFAF4B662E5AFFD696344D13EA7CE848" author="Tschitscherine" box="[606,859,1848,1874]" pageId="23" pageNumber="24" refString="Tschitscherine, T. (1906) Notes detachees sur les Harpalini de l'Asie orientale. Horae Societatis Entomologicae Rossicae, 37 (3 / 4), 247 - 292." type="journal article" year="1906">Tschitschérine, 1906</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8136972E5AFFD694FD4D71E800E869" blockId="23.[151,859,1847,1907]" box="[151,295,1882,1907]" pageId="23" pageNumber="24">
(
<figureCitation id="13052A122E5AFFD694F54D71E838E869" box="[159,287,1882,1907]" captionStart="FIGURES 48 – 51" captionStartId="24.[151,263,1124,1147]" captionTargetBox="[204,1384,201,1096]" captionTargetId="figure@24.[196,1392,195,1103]" captionTargetPageId="24" captionText="FIGURES 48 – 51. Harpalus, median lobe of aedeagus. 48 – 49, H. davidi (Gansu); 50 – 51, H. sericatus (Sichuan, lectotype). 48, 50, view from the left side; 49, 51, dorsal view. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287896/files/figure.png" pageId="23" pageNumber="24">Figs 50–51</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E5AFFD694FD4D88ECA5E8A2" box="[151,1410,1954,1976]" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="8B8136972E5AFFD694FD4D88ECA5E8A2" blockId="23.[151,1410,1954,1976]" box="[151,1410,1954,1976]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="AB2E11B92E5AFFD694FD4D88ECA5E8A2" box="[151,1410,1954,1976]" pageId="23" pageNumber="24">
<taxonomicName id="4C3E4D142E5AFFD694FD4D88EBFFE8A2" ID-CoL="6LDTH" authority="Tschitscherine, 1906: 255" authorityName="Tschitscherine" authorityPageNumber="255" authorityYear="1906" box="[151,728,1954,1976]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="sericatus" subGenus="Ophonus">
<emphasis id="B94AEA852E5AFFD694FD4D88E9D0E8A2" box="[151,247,1955,1976]" italics="true" pageId="23" pageNumber="24">Harpalus</emphasis>
(
<emphasis id="B94AEA852E5AFFD6956F4D88E846E8A2" box="[261,353,1955,1976]" italics="true" pageId="23" pageNumber="24">Ophonus</emphasis>
)
<emphasis id="B94AEA852E5AFFD695054D88E8ECE8A2" box="[367,459,1955,1976]" italics="true" pageId="23" pageNumber="24">sericatus</emphasis>
<treatmentCitation id="0A9F10862E5AFFD695BB4D89EBFFE8A2" author="Tschitscherine" box="[465,728,1954,1976]" page="255" pageId="23" pageNumber="24" year="1906">
<bibRefCitation id="EFAF4B662E5AFFD695BB4D89EB81E8A2" author="Tschitscherine" box="[465,678,1954,1976]" pageId="23" pageNumber="24" refString="Tschitscherine, T. (1906) Notes detachees sur les Harpalini de l'Asie orientale. Horae Societatis Entomologicae Rossicae, 37 (3 / 4), 247 - 292." type="journal article" year="1906">Tschitschérine, 1906</bibRefCitation>
: 255
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="548588352E5AFFD696884D88EA33E8A2" box="[738,788,1955,1976]" pageId="23" pageNumber="24">Type</typeStatus>
locality: &quot;Betw. Sin-pu-guan and Tun-khia&quot;, Sichuan,
<collectingCountry id="F32976072E5AFFD691544D89EC59E8A2" box="[1342,1406,1954,1976]" name="China" pageId="23" pageNumber="24">China</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF41661F2E55FFD994FD4E4FEA05EB83" httpUri="https://zenodo.org/record/287896/files/figure.png" pageId="24" pageNumber="25" targetBox="[204,1384,201,1096]" targetPageId="24">
<paragraph id="8B8136972E55FFD994FD4E4FEA05EB83" blockId="24.[151,1436,1124,1177]" pageId="24" pageNumber="25">
<emphasis id="B94AEA852E55FFD994FD4E4FE874EB60" bold="true" box="[151,339,1124,1147]" pageId="24" pageNumber="25">FIGURES 48–51.</emphasis>
<taxonomicName id="4C3E4D142E55FFD995324E4EE89FEB60" box="[344,440,1125,1146]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA852E55FFD995324E4EE89FEB60" box="[344,440,1125,1146]" italics="true" pageId="24" pageNumber="25">Harpalus</emphasis>
</taxonomicName>
, median lobe of aedeagus. 48–49,
<taxonomicName id="4C3E4D142E55FFD9977C4E4EEAE8EB60" authority="(Gansu)" baseAuthorityName="Gansu" box="[790,975,1124,1146]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="B94AEA852E55FFD9977C4E4EEA53EB60" box="[790,884,1125,1146]" italics="true" pageId="24" pageNumber="25">H. davidi</emphasis>
(Gansu)
</taxonomicName>
; 50–51,
<taxonomicName id="4C3E4D142E55FFD9904E4E4EEDB8EB60" box="[1060,1183,1125,1146]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sericatus">
<emphasis id="B94AEA852E55FFD9904E4E4EEDB8EB60" box="[1060,1183,1125,1146]" italics="true" pageId="24" pageNumber="25">H. sericatus</emphasis>
</taxonomicName>
(Sichuan, lectotype). 48, 50, view from the left side; 49, 51, dorsal view. Scale = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C324651C2E55FFD994FD4EEDE9CDEA75" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="8B8136972E55FFD994FD4EEDEA2BEA3D" blockId="24.[151,1437,1222,1607]" pageId="24" pageNumber="25">
<emphasis id="B94AEA852E55FFD994FD4EEDE819EBC5" bold="true" box="[151,318,1222,1247]" pageId="24" pageNumber="25">
<typeStatus id="548588352E55FFD994FD4EEDE9F5EBC5" box="[151,210,1222,1247]" pageId="24" pageNumber="25">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="548588352E55FFD995264EECE8E6EBC5" box="[332,449,1223,1247]" pageId="24" pageNumber="25" type="lectotype">Lectotype</typeStatus>
: ♂, labeled &quot;Mzhd. Sin-pu-guan i Tun-khia
<date id="FF8010572E55FFD997C54EECED02EBC5" box="[943,1061,1223,1247]" pageId="24" pageNumber="25" value="1993-08-12">12.VIII.93</date>
Potanin&quot; [in Cyrillic], &quot;
<taxonomicName id="4C3E4D142E55FFD9915E4EECECBBEBC4" box="[1332,1436,1223,1246]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="36" phylum="Arthropoda" rank="species" species="sericatus">
<emphasis id="B94AEA852E55FFD9915E4EECECBBEBC4" box="[1332,1436,1223,1246]" italics="true" pageId="24" pageNumber="25">Sericatus</emphasis>
</taxonomicName>
m., typ., Tschitscherin det.&quot; (
<collectionCode id="ED2FAE522E55FFD995874EC1EB06EA18" box="[493,545,1258,1282]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="24" pageNumber="25">ZIN</collectionCode>
).
<typeStatus id="548588352E55FFD996534EC1EBC6EA19" box="[569,737,1258,1283]" pageId="24" pageNumber="25" type="paralectotype">Paralectotypes</typeStatus>
: 2 ♀, labeled &quot;Mzhd. Ku-chang i Sing-pu-guan
<date id="FF8010572E55FFD991754EC1ECB0EA18" box="[1311,1431,1258,1282]" pageId="24" pageNumber="25" value="1993-08-24">24.VIII.93</date>
, Potanin&quot;, &quot;
<taxonomicName id="4C3E4D142E55FFD9957B4F24E85EEA3C" box="[273,377,1295,1318]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="36" phylum="Arthropoda" rank="species" species="sericatus">
<emphasis id="B94AEA852E55FFD9957B4F24E85EEA3C" box="[273,377,1295,1318]" italics="true" pageId="24" pageNumber="25">Sericatus</emphasis>
</taxonomicName>
m., Typ., Tschitscherin det.&quot; (
<collectionCode id="ED2FAE522E55FFD996A14F24EBD8EA3D" box="[715,767,1295,1319]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="24" pageNumber="25">ZIN</collectionCode>
).
</paragraph>
<paragraph id="8B8136972E55FFD994AD4F19E9CDEA75" blockId="24.[151,1437,1222,1607]" pageId="24" pageNumber="25">
<emphasis id="B94AEA852E55FFD994AD4F19EB14EA51" bold="true" box="[199,563,1330,1355]" pageId="24" pageNumber="25">Additional material examined</emphasis>
.
<collectingCountry id="F32976072E55FFD9962E4F19EBAAEA51" box="[580,653,1330,1355]" name="China" pageId="24" pageNumber="25">
<emphasis id="B94AEA852E55FFD9962E4F19EBAAEA51" bold="true" box="[580,653,1330,1355]" pageId="24" pageNumber="25">China</emphasis>
</collectingCountry>
.
<specimenCount id="9D38FD1E2E55FFD996F44F19EBEAEA50" box="[670,717,1330,1354]" pageId="24" pageNumber="25" type="generic">2 ♂</specimenCount>
, SICHUAN, Venchuan env.,
<date id="FF8010572E55FFD9904B4F19EDCFEA51" box="[1057,1256,1330,1355]" pageId="24" pageNumber="25" value="1995-04-27" valueMax="1995-05-19" valueMin="1995-04-27">27.IV.–19.V.1995</date>
, Vl. Beneš leg. (cWR).
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E55FFD994AD4F51EBBFE938" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="8B8136972E55FFD994AD4F51EBBFE938" blockId="24.[151,1437,1222,1607]" pageId="24" pageNumber="25">
<emphasis id="B94AEA852E55FFD994AD4F51E81DEA89" bold="true" box="[199,314,1402,1427]" pageId="24" pageNumber="25">Diagnosis</emphasis>
. This species is very similar in external characters and male genitalia to
<taxonomicName id="4C3E4D142E55FFD990184F50EDFCEA88" box="[1138,1243,1402,1426]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="B94AEA852E55FFD990184F50EDFCEA88" box="[1138,1243,1402,1426]" italics="true" pageId="24" pageNumber="25">H. davidi</emphasis>
</taxonomicName>
, but differs from the latter in having elytra mat, pubescent on two lateral intervals and with impunctate, superficial striae, protibia with three ventroapical spines, median lobe of aedeagus with smaller basal bulb, smaller apical capitulum and internal sac with medial patch of smaller spines (
<figureCitation id="13052A122E55FFD996AA4FCDEA63EAE5" box="[704,836,1510,1535]" captionStart="FIGURES 48 – 51" captionStartId="24.[151,263,1124,1147]" captionTargetBox="[204,1384,201,1096]" captionTargetId="figure@24.[196,1392,195,1103]" captionTargetPageId="24" captionText="FIGURES 48 – 51. Harpalus, median lobe of aedeagus. 48 – 49, H. davidi (Gansu); 50 – 51, H. sericatus (Sichuan, lectotype). 48, 50, view from the left side; 49, 51, dorsal view. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287896/files/figure.png" pageId="24" pageNumber="25">Figs 50–51</figureCitation>
). Metafemur has five to seven setigerous pores along posterior margin. Body length
<quantity id="4CC69B722E55FFD995854C21EBB3E938" box="[495,660,1546,1571]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" metricValueMax="1.28" metricValueMin="1.16" pageId="24" pageNumber="25" unit="mm" value="12.2" valueMax="12.8" valueMin="11.6">11.6–12.8 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E55FFD994AD4C05EAF7E95D" box="[199,976,1582,1607]" pageId="24" pageNumber="25" type="distribution">
<paragraph id="8B8136972E55FFD994AD4C05EAF7E95D" blockId="24.[151,1437,1222,1607]" box="[199,976,1582,1607]" pageId="24" pageNumber="25">
<emphasis id="B94AEA852E55FFD994AD4C05E87EE95D" bold="true" box="[199,345,1582,1607]" pageId="24" pageNumber="25">Distribution</emphasis>
. Known only from Sichuan,
<collectingCountry id="F32976072E55FFD996F54C05EBC5E95D" box="[671,738,1582,1607]" name="China" pageId="24" pageNumber="25">China</collectingCountry>
(
<bibRefCitation id="EFAF4B662E55FFD9969A4C04EAE3E95D" author="Kataev" box="[752,964,1582,1607]" pageId="24" pageNumber="25" refString="Kataev, B. M., Wrase, D. W. &amp; Ito, N. (2003) Harpalina. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
Kataev
<emphasis id="B94AEA852E55FFD997224C04EAA6E95C" box="[840,897,1582,1606]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
2003
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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