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<document id="05ABB3AC2BA68F8B12B94AD01DBC72EA" ID-DOI="10.11646/zootaxa.1011.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5048990" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625105215244" checkinUser="felipe" docAuthor="Jaschhof, Mathias &amp; Hippa, Heikki" docDate="2005" docId="44454C2AFFF5CA472F6F244722650FC5" docLanguage="en" docName="zt01011p054.pdf" docOrigin="Zootaxa 1011 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1011.1.1" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Manota acutistylus Jaschhof &amp; Hippa 2005, sp. n." docType="treatment" docVersion="3" lastPageNumber="48" masterDocId="B87C3452FFDBCA682E67255B20100B0B" masterDocTitle="The genus Manota in Costa Rica (Diptera: Mycetophilidae)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="47" updateTime="1698980072965" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="85DC1404430752609EC4E0545C62D32A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A716D2E3BD0B5DB9F517BD26FDFE6DC5">
<mods:title id="E3EF73D9D9C3362C27FD467EFBB8DD45">The genus Manota in Costa Rica (Diptera: Mycetophilidae)</mods:title>
</mods:titleInfo>
<mods:name id="0B824263DB5AE849A92A94F3F6240C90" type="personal">
<mods:role id="2E71CDAD5B07AC27CA5CA8F04E05A49C">
<mods:roleTerm id="DCF48FFA32B1B36DA4C8DA451E55B73D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6035BF28B7B230222C329677FF860882">Jaschhof, Mathias</mods:namePart>
</mods:name>
<mods:name id="D894B222744B5AE025DF93ADC1385452" type="personal">
<mods:role id="11853D2EE69FBF39D6CFB857F4E9323B">
<mods:roleTerm id="6B26E56C0D585D5D7FDD19FD3942C5A3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="70D1D1FAB348405429C29FD0B3113CD3">Hippa, Heikki</mods:namePart>
</mods:name>
<mods:typeOfResource id="932DEF867CA60064D60E07DD061F3D86">text</mods:typeOfResource>
<mods:relatedItem id="0394D154B1B9496AE595480E6D5EC1F5" type="host">
<mods:titleInfo id="DDCC78EBA419E53F67ED864E472A9591">
<mods:title id="A46853FA54D2FA8212F22D7C879694D3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="93536E33B3042845D46AF70C07FE0FE3">
<mods:date id="73520B1B63E8E7844B471B75985B3400">2005</mods:date>
<mods:detail id="68869A0F8644236CF9223DD40C132BDE" type="pubDate">
<mods:number id="AD3E3D074CB049C6A7DA115F3936BB60">2005-06-24</mods:number>
</mods:detail>
<mods:detail id="4993E8D99B422D19528E29900676777A" type="volume">
<mods:number id="9FB27BF40E83F25ADC4CE02831385F0D">1011</mods:number>
</mods:detail>
<mods:detail id="13C006DDF2BAEC310224BE676A95860E" type="issue">
<mods:number id="2DB8E7F3BBF5B62D41966127C72EB84D">1</mods:number>
</mods:detail>
<mods:extent id="52C94A33805D1A76DF6A61AE431A75A4" unit="page">
<mods:start id="9AF20D03CFBE4CD8D67B676348676D7D">1</mods:start>
<mods:end id="56F719480448CE21390328C1BBDEABCD">54</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="91EDAE46D1FCDB467A69CC361112FE12">
<mods:url id="38E93F0A6B49A3C22AA9CBBC4BEDB2F8">https://biotaxa.org/Zootaxa/article/view/zootaxa.1011.1.1</mods:url>
</mods:location>
<mods:classification id="9AA20F630AC3A314E7CF3863A32DBB8F">journal article</mods:classification>
<mods:identifier id="A00BDDB113006166AD21FD1DACA26156" type="DOI">10.11646/zootaxa.1011.1.1</mods:identifier>
<mods:identifier id="762C8D003F058FB532EC8A7E99093883" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="60CCF1388ECCEC01C51D27420AD6B236" type="Zenodo-Dep">5048990</mods:identifier>
<mods:identifier id="14B2956AB4CD9A62D20AFBC78060B693" type="ZooBank">C8892524-FB03-4DA9-9894-968448E1DB47</mods:identifier>
</mods:mods>
<treatment id="44454C2AFFF5CA472F6F244722650FC5" LSID="urn:lsid:plazi:treatment:44454C2AFFF5CA472F6F244722650FC5" httpUri="http://treatment.plazi.org/id/44454C2AFFF5CA472F6F244722650FC5" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<subSubSection id="84F6AEB7FFF5CA462F6F244722B60A3E" box="[264,678,283,310]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="CC53FD3CFFF5CA462F6F244722B60A3E" blockId="46.[264,678,283,310]" box="[264,678,283,310]" pageId="46" pageNumber="47">
<heading id="971B4A50FFF5CA462F6F244722B60A3E" bold="true" box="[264,678,283,310]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
<emphasis id="FE98212EFFF5CA462F6F244722B60A3E" bold="true" box="[264,678,283,310]" pageId="46" pageNumber="47">
<taxonomicName id="0BEC86BFFFF5CA462F6F244721FE0A3D" ID-CoL="6R6JY" authority="Jaschhof &amp; Hippa, 2005" authorityName="Jaschhof &amp; Hippa" authorityYear="2005" box="[264,494,284,310]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="acutistylus" status="sp. nov.">
<emphasis id="FE98212EFFF5CA462F6F244721FE0A3D" bold="true" box="[264,494,284,310]" italics="true" pageId="46" pageNumber="47">Manota acutistylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5AB9C55FFF5CA462F92244022270A3E" box="[501,567,283,309]" pageId="46" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="54D7E1B9FFF5CA462C202440228F0A3E" box="[583,671,283,309]" captionStart="FIGURE 24" captionStartId="46.[264,367,1825,1849]" captionTargetBox="[337,1259,690,1807]" captionTargetId="figure-115@46.[310,1303,684,1822]" captionTargetPageId="46" captionText="FIGURE 24. Manota acutistylus sp. n. A: Male terminalia, ventral view (holotype). B: Male terminalia, dorsal view (paratype). Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5049064" httpUri="https://zenodo.org/record/5049064/files/figure.png" pageId="46" pageNumber="47">Fig. 24</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="84F6AEB7FFF5CA472F6F243C22650FC5" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="description">
<paragraph id="CC53FD3CFFF5CA462F6F243C22590A8A" blockId="46.[264,1324,359,625]" box="[264,585,359,385]" pageId="46" pageNumber="47">
<emphasis id="FE98212EFFF5CA462F6F243C21820A8B" box="[264,402,359,384]" italics="true" pageId="46" pageNumber="47">Description</emphasis>
. Small species.
</paragraph>
<paragraph id="CC53FD3CFFF5CA462F5F24D421BC0AF2" blockId="46.[264,1324,359,625]" pageId="46" pageNumber="47">
Male.
<emphasis id="FE98212EFFF5CA462FE224D521DF0AA3" bold="true" box="[389,463,398,424]" pageId="46" pageNumber="47">Head.</emphasis>
Antenna with fourth flagellomere 1.6 times as long as wide. Maxillary palpus with third palpomere bearing 3–4 curved sensilla; fourth palpomere with a parasegment apically.
</paragraph>
<paragraph id="CC53FD3CFFF5CA462F5F275D233D097A" blockId="46.[264,1324,359,625]" pageId="46" pageNumber="47">
<emphasis id="FE98212EFFF5CA462F5F275D218C092B" bold="true" box="[312,412,518,544]" pageId="46" pageNumber="47">Thorax.</emphasis>
Anterior basalare and laterotergite each bare. Episternum 3 with ventral portion setose.
<emphasis id="FE98212EFFF5CA462FF3277521CE0943" bold="true" box="[404,478,558,584]" pageId="46" pageNumber="47">Wing.</emphasis>
Length
<quantity id="0B1450D9FFF5CA462C272774228E0942" box="[576,670,559,585]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="46" pageNumber="47" unit="mm" value="1.5">1.5 mm</quantity>
. Membrane with a few setae along posterior margin. Sc bare. Junction of CuA2 into CuA1 incomplete.
</paragraph>
<caption id="9893ADB4FFF5CA462F6F227A22E50C50" ID-DOI="http://doi.org/10.5281/zenodo.5049064" ID-Zenodo-Dep="5049064" httpUri="https://zenodo.org/record/5049064/files/figure.png" pageId="46" pageNumber="47" startId="46.[264,367,1825,1849]" targetBox="[337,1259,690,1807]" targetPageId="46">
<paragraph id="CC53FD3CFFF5CA462F6F227A22E50C50" blockId="46.[264,1322,1825,1883]" pageId="46" pageNumber="47">
<emphasis id="FE98212EFFF5CA462F6F227A21870C32" bold="true" box="[264,407,1825,1849]" pageId="46" pageNumber="47">FIGURE 24.</emphasis>
<taxonomicName id="0BEC86BFFFF5CA462FFA227922790C32" authority="Jaschhof &amp; Hippa, 2005" authorityName="Jaschhof &amp; Hippa" authorityYear="2005" box="[413,617,1826,1849]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="acutistylus" status="sp. nov.">
<emphasis id="FE98212EFFF5CA462FFA227922790C32" box="[413,617,1826,1849]" italics="true" pageId="46" pageNumber="47">Manota acutistylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5AB9C55FFF5CA462C08227A22B80C33" box="[623,680,1825,1848]" pageId="46" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
<emphasis id="FE98212EFFF5CA462CCA227A22DA0C32" bold="true" box="[685,714,1825,1849]" pageId="46" pageNumber="47">A:</emphasis>
Male terminalia, ventral view (holotype).
<emphasis id="FE98212EFFF5CA462AEB227A24B60C32" bold="true" box="[1164,1190,1825,1849]" pageId="46" pageNumber="47">B:</emphasis>
Male terminalia, dorsal view (paratype). Scale = 0.1 mm.
</paragraph>
</caption>
<paragraph id="CC53FD3CFFF4CA472F5F244022CA081D" blockId="47.[264,1326,283,1230]" pageId="47" pageNumber="48">
<emphasis id="FE98212EFFF4CA472F5F244021D10A3E" bold="true" box="[312,449,283,309]" pageId="47" pageNumber="48">Terminalia</emphasis>
(
<figureCitation id="54D7E1B9FFF4CA472FB6244722380A3D" box="[465,552,284,310]" captionStart="FIGURE 24" captionStartId="46.[264,367,1825,1849]" captionTargetBox="[337,1259,690,1807]" captionTargetId="figure-115@46.[310,1303,684,1822]" captionTargetPageId="46" captionText="FIGURE 24. Manota acutistylus sp. n. A: Male terminalia, ventral view (holotype). B: Male terminalia, dorsal view (paratype). Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5049064" httpUri="https://zenodo.org/record/5049064/files/figure.png" pageId="47" pageNumber="48">Fig. 24</figureCitation>
). Sternite 9 with only posterior margin distinct; margin situated at mid length of gonocoxites, slightly convex medially. Ventro­lateral lobes of gonocoxites rounded, each marginally with 2–3 larger setae; interiorly with 1 seta on a separate rim. Disto­lateral lobes of gonocoxites comparatively short; with large setae directed posteriorly. Gonocoxites in position I with 1 medium­sized, blunt megaseta; in position II with 2– 3 short, blunt megasetae; in position III with 1 pointed megaseta and 1 flat, two­branched megaseta with one branch broadly rounded and the other whip­like; in position IV with 4 large setae; on dorso­posterior gonocoxal margin with an irregular comb of numerous setae directed inwards. Gonostyli small, elongate, pointed apically; each on posterior margin with 1 long, stiff seta directed inwards, 1 long seta directed dorsally, and 1 strong, recurved seta directed dorsally; on anterior margin with 1 seta subbasally. Parastylar lobes moderately sized, weak, bare (not illustrated). Tegmen evenly tapering to tip. Hypoproct tapering to tip; with long setae laterally.
</paragraph>
<paragraph id="CC53FD3CFFF4CA472F5F267F246408BD" blockId="47.[264,1326,283,1230]" pageId="47" pageNumber="48">
<emphasis id="FE98212EFFF4CA472F5F267F222D0836" box="[312,573,804,829]" italics="true" pageId="47" pageNumber="48">Diagnostic characters</emphasis>
. It is in the group of species with both anterior basalare and laterotergites bare; terminalia with sternite 9 barely discernible; the disto­lateral lobes of the gonocoxites large; and the gonostyli small and flat. The array of megasetae dorsally on the gonocoxites is species specific, as are the pointed, apically bare gonostyli.
</paragraph>
<paragraph id="CC53FD3CFFF4CA472F5F269F216D0F0D" blockId="47.[264,1326,283,1230]" pageId="47" pageNumber="48">
<emphasis id="FE98212EFFF4CA472F5F269F21A308D6" box="[312,435,964,989]" italics="true" pageId="47" pageNumber="48">Etymology</emphasis>
. From Latin,
<emphasis id="FE98212EFFF4CA472C3E269F22B508D6" box="[601,677,964,989]" italics="true" pageId="47" pageNumber="48">acutus</emphasis>
, acute, and
<emphasis id="FE98212EFFF4CA472D51269F236908D6" box="[822,889,964,989]" italics="true" pageId="47" pageNumber="48">stylus</emphasis>
, style, referring to the outline of the gonostyli.
</paragraph>
<paragraph id="CC53FD3CFFF4CA472F5F214F22650FC5" blockId="47.[264,1326,283,1230]" pageId="47" pageNumber="48">
<emphasis id="FE98212EFFF4CA472F5F214F216A0F26" box="[312,378,1044,1069]" italics="true" pageId="47" pageNumber="48">Types</emphasis>
.
<materialsCitation id="7C84F761FFF4CA472FEE214F218B0F75" collectingDate="2003-06-19" collectingDateMax="2003-07-08" collectingDateMin="2003-06-19" collectionCode="INBio" collectorName="M. Moraga &amp; A. Azofeifa &amp; K. Caballero" country="Costa Rica" elevation="200" location="Cerro Puma" municipality="Male" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<emphasis id="FE98212EFFF4CA472FEE214F21E40F26" box="[393,500,1044,1069]" italics="true" pageId="47" pageNumber="48">
<typeStatus id="1357439EFFF4CA472FEE214F21E40F26" box="[393,500,1044,1069]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingMunicipality id="2C376746FFF4CA472C64214F22520F25" box="[515,578,1044,1070]" pageId="47" pageNumber="48">Male</collectingMunicipality>
,
<collectingCountry id="B4FBBDACFFF4CA472C28214F22C20F25" box="[591,722,1044,1070]" name="Costa Rica" pageId="47" pageNumber="48">Costa Rica</collectingCountry>
,
<collectingRegion id="0E2833DEFFF4CA472CB8214F23C00F25" box="[735,976,1044,1070]" country="Costa Rica" name="Puntarenas" pageId="47" pageNumber="48">Puntarenas province</collectingRegion>
,
<location id="C933ABE7FFF4CA472DBA214F247E0F25" LSID="urn:lsid:plazi:treatment:44454C2AFFF5CA472F6F244722650FC5:C933ABE7FFF4CA472DBA214F247E0F25" box="[989,1134,1044,1070]" country="Costa Rica" municipality="Male" name="Cerro Puma" pageId="47" pageNumber="48" stateProvince="Puntarenas">Cerro Puma</location>
,
<quantity id="0B1450D9FFF4CA472A1D214F25120F25" box="[1146,1282,1044,1070]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="47" pageNumber="48" unit="m" value="200.0" valueMax="300.0" valueMin="100.0">
<elevation id="47C11A0FFFF4CA472A1D214F25120F25" box="[1146,1282,1044,1070]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="47" pageNumber="48" unit="m" value="200.0" valueMax="300.0" valueMin="100.0">100–300 m</elevation>
</quantity>
,
<date id="B852DBFCFFF4CA472B68214F21CA0F5D" pageId="47" pageNumber="48" value="2003-06-19" valueMax="2003-07-08" valueMin="2003-06-19">
<collectingDate id="A8162214FFF4CA472B68214F21CA0F5D" pageId="47" pageNumber="48" value="2003-06-19" valueMax="2003-07-08" valueMin="2003-06-19">19 June–8 July 2003</collectingDate>
</date>
,
<collectingMethod id="15AD852BFFF4CA472F80216722B30F5D" box="[487,675,1084,1110]" pageId="47" pageNumber="48">by Malaise trap</collectingMethod>
,
<collectorName id="611998EAFFF4CA472CD7216723280F5D" box="[688,824,1084,1110]" pageId="47" pageNumber="48">M. Moraga</collectorName>
,
<collectorName id="611998EAFFF4CA472D22216723C00F5D" box="[837,976,1084,1110]" pageId="47" pageNumber="48">A. Azofeifa</collectorName>
&amp;
<collectorName id="611998EAFFF4CA472D9E216724800F5D" box="[1017,1168,1084,1110]" pageId="47" pageNumber="48">K. Caballero</collectorName>
(
<collectionCode id="AAFD65F9FFF4CA472AC7216724FB0F5D" box="[1184,1259,1084,1110]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="47" pageNumber="48">INBio</collectionCode>
sample #74508)
</materialsCitation>
.
<materialsCitation id="7C84F761FFF4CA472FCD213F22610FC5" collectingDate="2003-03-11" collectingDateMax="2003-04-01" collectingDateMin="2003-03-11" collectionCode="INBio" collectorName="E. Rojas &amp; B. Gamboa &amp; W. Arana" country="Costa Rica" elevation="560" location="Sendero Espavel" municipality="Hitoy Cerere Biological Reserve" pageId="47" pageNumber="48" specimenCount="1" specimenCount-male="1" stateProvince="Limon" typeStatus="paratype">
<emphasis id="FE98212EFFF4CA472FCD213F22050F76" box="[426,533,1124,1149]" italics="true" pageId="47" pageNumber="48">
<typeStatus id="1357439EFFF4CA472FCD213F22050F76" box="[426,533,1124,1149]" pageId="47" pageNumber="48" type="paratype">Paratype</typeStatus>
</emphasis>
.
<specimenCount id="DAEA36B5FFF4CA472C40213F226C0F75" box="[551,636,1124,1150]" pageId="47" pageNumber="48" type="male">1 male</specimenCount>
,
<collectingRegion id="0E2833DEFFF4CA472CEC213F235D0F75" box="[651,845,1124,1150]" country="Costa Rica" name="Limon" pageId="47" pageNumber="48">Limón province</collectingRegion>
,
<collectingMunicipality id="2C376746FFF4CA472D3B213F24F80F75" box="[860,1256,1124,1150]" pageId="47" pageNumber="48">Hitoy Cerere Biological Reserve</collectingMunicipality>
,
<location id="C933ABE7FFF4CA472A90213F21B20FAD" LSID="urn:lsid:plazi:treatment:44454C2AFFF5CA472F6F244722650FC5:C933ABE7FFF4CA472A90213F21B20FAD" country="Costa Rica" municipality="Hitoy Cerere Biological Reserve" name="Sendero Espavel" pageId="47" pageNumber="48" stateProvince="Limon">Sendero Espavel</location>
,
<quantity id="0B1450D9FFF4CA472FC821D721EB0FAD" box="[431,507,1164,1190]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="47" pageNumber="48" unit="m" value="560.0">
<elevation id="47C11A0FFFF4CA472FC821D721EB0FAD" box="[431,507,1164,1190]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="47" pageNumber="48" unit="m" value="560.0">560 m</elevation>
</quantity>
,
<date id="B852DBFCFFF4CA472C6021D7230F0FAD" box="[519,799,1164,1190]" pageId="47" pageNumber="48" value="2003-03-11" valueMax="2003-04-01" valueMin="2003-03-11">
<collectingDate id="A8162214FFF4CA472C6021D7230F0FAD" box="[519,799,1164,1190]" pageId="47" pageNumber="48" value="2003-03-11" valueMax="2003-04-01" valueMin="2003-03-11">11 March–1 April 2003</collectingDate>
</date>
,
<collectingMethod id="15AD852BFFF4CA472D4C21D723F50FAD" box="[811,997,1164,1190]" pageId="47" pageNumber="48">by Malaise trap</collectingMethod>
,
<collectorName id="611998EAFFF4CA472D9721D724460FAD" box="[1008,1110,1164,1190]" pageId="47" pageNumber="48">E. Rojas</collectorName>
,
<collectorName id="611998EAFFF4CA472A0521D724F70FAD" box="[1122,1255,1164,1190]" pageId="47" pageNumber="48">B. Gamboa</collectorName>
&amp;
<collectorName id="611998EAFFF4CA472B6A21D7215F0FC5" pageId="47" pageNumber="48">W. Arana</collectorName>
(
<collectionCode id="AAFD65F9FFF4CA472F3821EF21BA0FC5" box="[351,426,1204,1230]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="47" pageNumber="48">INBio</collectionCode>
sample #73628)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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