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<document id="C9022CB70AD346A163974961E7F01E44" ID-DOI="10.11646/zootaxa.3940.1.1" ID-GBIF-Dataset="5013eaff-bb37-4be3-a1ea-43c11d0cf5d1" ID-ISSN="1175-5326" ID-Zenodo-Dep="288204" ID-ZooBank="B5D45D1B-4B52-4F0B-8AF6-B587F8857475" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461023701082" checkinUser="plazi" docAuthor="Cowie, Robert H." docDate="2015" docId="03FFE6320A36F543FF68CCD0A9F6FA06" docLanguage="en" docName="zt03940p092.pdf" docOrigin="Zootaxa 3940 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ampullaria wernei" docType="treatment" docVersion="7" lastPageNumber="52" masterDocId="FFC69E4A0A05F570FFFFCC2EAA34FF91" masterDocTitle="The recent apple snails of Africa and Asia (Mollusca: Gastropoda: Ampullariidae: Afropomus, Forbesopomus, Lanistes, Pila, Saulea): a nomenclatural and type catalogue. The apple snails of the Americas: addenda and corrigenda" masterLastPageNumber="92" masterPageNumber="1" pageNumber="52" updateTime="1698571224122" updateUser="plazi">
<mods:mods id="4524379348EAB80AAD5D03315179A15D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2A0191F9B845B34EA2A0D490332E471C">The recent apple snails of Africa and Asia (Mollusca: Gastropoda: Ampullariidae: Afropomus, Forbesopomus, Lanistes, Pila, Saulea): a nomenclatural and type catalogue. The apple snails of the Americas: addenda and corrigenda</mods:title>
</mods:titleInfo>
<mods:name id="8DC65ABD2396AFC09BD1743C141E8968" type="personal">
<mods:role id="4191A749932D0FCFF29395ADB1A40911">
<mods:roleTerm id="E1F5BBE092878AD5BA4DBC4BBA1A67CB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8FC5107F3CFF583EE884D90A4FBB5540">Cowie, Robert H.</mods:namePart>
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<mods:title id="BC83E957B7001AA49B880F0D0493B2A0">Zootaxa</mods:title>
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<mods:part id="F3E2E0F5D40748DB89151BC47D22E88D">
<mods:date id="8264A904ABEEF44E72E59452B1407274">2015</mods:date>
<mods:detail id="61BAB6C69701EF543EFB170ED2C64D73" type="volume">
<mods:number id="31D9E8CEF2D41EE586A5BCF2C49129F8">3940</mods:number>
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<mods:classification id="6B5D7B91FEE5A8ACE55D0D1B04C16251">journal article</mods:classification>
<mods:identifier id="F39F96F760F7B79C8EE5DE148322DCBC" type="DOI">10.11646/zootaxa.3940.1.1</mods:identifier>
<mods:identifier id="043F8B42033B834CCC5D07ABEBD8F6BA" type="GBIF-Dataset">5013eaff-bb37-4be3-a1ea-43c11d0cf5d1</mods:identifier>
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<treatment id="03FFE6320A36F543FF68CCD0A9F6FA06" ID-DOI="http://doi.org/10.5281/zenodo.5173381" ID-GBIF-Taxon="127687665" ID-Zenodo-Dep="5173381" LSID="urn:lsid:plazi:treatment:03FFE6320A36F543FF68CCD0A9F6FA06" httpUri="http://treatment.plazi.org/id/03FFE6320A36F543FF68CCD0A9F6FA06" lastPageNumber="52" pageId="51" pageNumber="52">
<subSubSection id="C34C04AF0A36F543FF68CCD0AADCFE86" box="[151,232,254,279]" pageId="51" pageNumber="52" type="nomenclature">
<paragraph id="8BE957240A36F543FF68CCD0AADCFE86" blockId="51.[151,1436,254,1215]" box="[151,232,254,279]" pageId="51" pageNumber="52">
<heading id="D0A1E0480A36F543FF68CCD0AADCFE86" bold="true" box="[151,232,254,279]" fontSize="10" level="2" pageId="51" pageNumber="52" reason="2">
<taxonomicName id="4C562CA70A36F543FF68CCD0AADCFE86" ID-CoL="D86C" box="[151,232,254,279]" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="wernei">
<emphasis id="B9228B360A36F543FF68CCD0AADCFE86" bold="true" box="[151,232,254,279]" pageId="51" pageNumber="52">wernei</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C04AF0A36F543FF38CD0CAB8CFD82" pageId="51" pageNumber="52" type="reference_group">
<paragraph id="8BE957240A36F543FF38CD0CAB8CFD82" blockId="51.[151,1436,254,1215]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="AB46700A0A36F543FF38CD0CAB8CFD82" infered="true" pageId="51" pageNumber="52">
<taxonomicName id="4C562CA70A36F543FF38CD0CA8AFFEAA" ID-CoL="D86C" authority="Philippi, 1851: 19" authorityName="Philippi" authorityPageNumber="19" authorityYear="1851" box="[199,667,290,315]" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="wernei">
<emphasis id="B9228B360A36F543FF38CD0CAB91FEAA" box="[199,421,290,315]" italics="true" pageId="51" pageNumber="52">Ampullaria wernei</emphasis>
<treatmentCitation id="0AF771350A36F543FE47CD0CA8AFFEAA" author="Philippi" box="[440,667,290,315]" page="19" pageId="51" pageNumber="52" year="1851">
<bibRefCitation id="EFC72AD50A36F543FE47CD0CA853FEAA" author="Philippi" box="[440,615,290,315]" pageId="51" pageNumber="52" refString="Philippi, R. A. (1851 - [1852]) Die Gattung Ampullaria. In: Kuster, H. C. (Ed.), Systematisches Conchylien-Cabinet von Martini und Chemnitz. Neu herausgegeben und vervollstandigt. Ersten Bandes zwanzigste Abtheilung. Bauer &amp; Raspe, Nurnberg [= Nuremberg], pp. 1 - 74, pls. A, 1 - 21." type="book chapter" year="1851">Philippi, 1851</bibRefCitation>
: 19
</treatmentCitation>
</taxonomicName>
, pl. 5, fig. 4 [1852a: 54, pl. 17, fig. 2; 1852b: 22].
<emphasis id="B9228B360A36F543FA98CD0AAAC2FECE" italics="true" pageId="51" pageNumber="52">
<typeStatus id="54EDE9860A36F543FA98CD0AAFADFEAA" box="[1383,1433,292,315]" pageId="51" pageNumber="52">Type</typeStatus>
material
</emphasis>
—
<typeStatus id="54EDE9860A36F543FEEDCD68AB4CFECE" box="[274,376,326,351]" pageId="51" pageNumber="52" type="lectotype">lectotype</typeStatus>
(
<bibRefCitation id="EFC72AD50A36F543FE78CD68A969FECE" author="Kohler" box="[391,861,326,351]" pageId="51" pageNumber="52" refString="Kohler, F. &amp; Glaubrecht, M. (2006) The types of Ampullariidae Gray, 1824 (Mollusca, Gastropoda) in the Malacological Collection of the Natural History Museum, Berlin: an annotated catalogue with lectotype designations. Mitteilungen aus dem Museum fur Naturkunde Berlin, Zoologische Reihe, 82 (1), 198 - 215." type="journal article" year="2006">Köhler &amp; Glaubrecht (2006: 212, fig. 3L)</bibRefCitation>
: ZMB 1335 (the shell figured as
<taxonomicName id="4C562CA70A36F543FB32CD69AF2DFECF" ID-CoL="D86C" box="[1229,1305,327,350]" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="wernei">
<emphasis id="B9228B360A36F543FB32CD69AF2DFECF" box="[1229,1305,327,350]" italics="true" pageId="51" pageNumber="52">wernei</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC72AD50A36F543FABCCD68AB45FE12" author="Philippi" pageId="51" pageNumber="52" refString="Philippi, R. A. (1851 - [1852]) Die Gattung Ampullaria. In: Kuster, H. C. (Ed.), Systematisches Conchylien-Cabinet von Martini und Chemnitz. Neu herausgegeben und vervollstandigt. Ersten Bandes zwanzigste Abtheilung. Bauer &amp; Raspe, Nurnberg [= Nuremberg], pp. 1 - 74, pls. A, 1 - 21." type="book chapter" year="1851">Philippi (1851: pl. 5, fig. 4)</bibRefCitation>
; =
<typeStatus id="54EDE9860A36F543FE6ACD44ABC3FE12" box="[405,503,362,387]" pageId="51" pageNumber="52" type="holotype">holotype</typeStatus>
of
<emphasis id="B9228B360A36F543FDE1CD42A855FE12" box="[542,609,364,387]" italics="true" pageId="51" pageNumber="52">minor</emphasis>
Bourguignat);
<typeStatus id="54EDE9860A36F543FCF2CD44A99DFE12" box="[781,937,362,387]" pageId="51" pageNumber="52" type="paralectotype">paralectotype</typeStatus>
: the shell figured by
<bibRefCitation id="EFC72AD50A36F543FB68CD44AADBFE36" author="Philippi" pageId="51" pageNumber="52" refString="Philippi, R. A. (1852 b) Centuria quinta testaceorum novorum. (Contin.) Zeitschrift fur Malakozoologie, 9 (2), 20 - 29. [25 March; p. 32]" type="journal article" year="1852" yearSuffix="a">Philippi (1852a, pl. 17, fig. 2)</bibRefCitation>
, probably MNHNCL 15962 (1 spm.).
<emphasis id="B9228B360A36F543FCF7CDA1A99DFE36" box="[776,937,398,423]" italics="true" pageId="51" pageNumber="52">
<typeStatus id="54EDE9860A36F543FCF7CDA1A908FE37" box="[776,828,399,422]" pageId="51" pageNumber="52">Type</typeStatus>
locality
</emphasis>
—“der weisse Nil” [the White Nile].
<emphasis id="B9228B360A36F543FF68CD9CAB14FE5A" box="[151,288,434,459]" italics="true" pageId="51" pageNumber="52">Distribution</emphasis>
—Scattered reports from a large area including southern
<collectingCountry id="F34117B40A36F543FC5BCD9CAE36FE5A" box="[932,1026,434,459]" name="Somalia" pageId="51" pageNumber="52">Somalia</collectingCountry>
,
<collectingCountry id="F34117B40A36F543FBF0CD9CAE6CFE5A" box="[1039,1112,434,459]" name="Sudan" pageId="51" pageNumber="52">Sudan</collectingCountry>
, possibly
<collectingCountry id="F34117B40A36F543FB36CD9DAF21FE5A" box="[1225,1301,435,459]" name="Kenya" pageId="51" pageNumber="52">Kenya</collectingCountry>
, Lake
<collectingCountry id="F34117B40A36F543FA9ECD9CAFA8FE5A" box="[1377,1436,434,459]" name="Chad" pageId="51" pageNumber="52">Chad</collectingCountry>
and westwards to the
<collectingCountry id="F34117B40A36F543FE66CDF8ABEDFE7E" box="[409,473,470,495]" name="Niger" pageId="51" pageNumber="52">Niger</collectingCountry>
River in
<collectingCountry id="F34117B40A36F543FDB2CDF8A8B6FE7E" box="[589,642,470,495]" name="Serbia" pageId="51" pageNumber="52">Mali</collectingCountry>
and southwestern
<collectingCountry id="F34117B40A36F543FCA0CDF8A982FE7E" box="[863,950,470,495]" name="Nigeria" pageId="51" pageNumber="52">Nigeria</collectingCountry>
, and western
<collectingCountry id="F34117B40A36F543FBA5CDF8AAD6FD82" name="Democratic Republic of the Congo" pageId="51" pageNumber="52">Democratic Republic of the Congo</collectingCountry>
(
<treatmentCitation id="0AF771350A36F543FF0FCDD5AB98FD82" author="Brown" box="[240,428,506,531]" page="56" pageId="51" pageNumber="52" year="1994">
<bibRefCitation id="EFC72AD50A36F543FF0FCDD5AB98FD82" author="Brown" box="[240,428,506,531]" pageId="51" pageNumber="52" refString="Brown, D. S. (1994) Freshwater snails of Africa and their medical importance. Second edition. Taylor &amp; Francis, London, x + 608 + [1] pp." type="journal article" year="1994">Brown 1994: 56</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34C04AF0A36F543FF38CE30AAC0FA96" pageId="51" pageNumber="52" type="discussion">
<paragraph id="8BE957240A36F543FF38CE30AB4FFB2E" blockId="51.[151,1436,254,1215]" pageId="51" pageNumber="52">
<emphasis id="B9228B360A36F543FF38CE30AB1DFDA6" box="[199,297,542,567]" italics="true" pageId="51" pageNumber="52">Remarks</emphasis>
. Both figures of
<bibRefCitation id="EFC72AD50A36F543FE16CE30A8BBFDA6" author="Philippi" box="[489,655,542,567]" pageId="51" pageNumber="52" refString="Philippi, R. A. (1851 - [1852]) Die Gattung Ampullaria. In: Kuster, H. C. (Ed.), Systematisches Conchylien-Cabinet von Martini und Chemnitz. Neu herausgegeben und vervollstandigt. Ersten Bandes zwanzigste Abtheilung. Bauer &amp; Raspe, Nurnberg [= Nuremberg], pp. 1 - 74, pls. A, 1 - 21." type="book chapter" year="1851">Philippi (1851</bibRefCitation>
: pl. 5, fig. 4, 1852a: pl. 17, fig. 2) were referenced by
<bibRefCitation id="EFC72AD50A36F543FB0CCE30AA8AFDCA" author="Philippi" pageId="51" pageNumber="52" refString="Philippi, R. A. (1851 - [1852]) Die Gattung Ampullaria. In: Kuster, H. C. (Ed.), Systematisches Conchylien-Cabinet von Martini und Chemnitz. Neu herausgegeben und vervollstandigt. Ersten Bandes zwanzigste Abtheilung. Bauer &amp; Raspe, Nurnberg [= Nuremberg], pp. 1 - 74, pls. A, 1 - 21." type="book chapter" year="1851">Philippi (1851: 19)</bibRefCitation>
, although pl.
<date id="FFE871E40A36F543FEAACE6DAB9CFDCA" box="[341,424,579,603]" pageId="51" pageNumber="52" value="1917-05">17 may</date>
not have been published until 1852, so both specimens are part of the
<typeStatus id="54EDE9860A36F543FB4ACE6DAED1FDCA" box="[1205,1253,579,603]" pageId="51" pageNumber="52">type</typeStatus>
series of
<taxonomicName id="4C562CA70A36F543FAAFCE6AA830FDEE" authority="Philippi. Bourguignat (1879: 32)" authorityName="Philippi. Bourguignat" authorityPageNumber="32" authorityYear="1879" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="wernei">
<emphasis id="B9228B360A36F543FAAFCE6AAFA8FDCA" box="[1360,1436,580,603]" italics="true" pageId="51" pageNumber="52">wernei</emphasis>
Philippi.
<bibRefCitation id="EFC72AD50A36F543FF02CE48A830FDEE" author="Bourguignat" box="[253,516,614,639]" pageId="51" pageNumber="52" refString="Bourguignat, J. R. (1879) Description de divers especes terrestres et fluviatiles et de differents genres de mollusques de l'Egypte, de l'Abyssinie, de Zanzibar, du Senegal et du centre de l'Afrique. Imprimerie de Madame Veuve Bouchard- Huzard, Jules Tremblay, Gendre et Successeur, Paris, 54 pp. http: // dx. doi. org / 10.5962 / bhl. title. 50008" type="book" year="1879">Bourguignat (1879: 32)</bibRefCitation>
</taxonomicName>
referenced the shell illustrated in pl. 17, fig. 2 as the “échantillon
<typeStatus id="54EDE9860A36F543FB1ACE49AF2EFDEE" box="[1253,1306,615,639]" pageId="51" pageNumber="52">type</typeStatus>
”. However, as Bourguignat often used the term “
<typeStatus id="54EDE9860A36F543FDC1CEA5A846FD32" box="[574,626,651,675]" pageId="51" pageNumber="52">type</typeStatus>
” to mean “typical” (E. Neubert pers. comm.
<date id="FFE871E40A36F543FB83CEA4AF6AFD32" box="[1148,1374,650,675]" pageId="51" pageNumber="52" value="2014-09-25">25 September 2014</date>
), this cannot be considered definitively as a
<typeStatus id="54EDE9860A36F543FDB2CE80A880FD56" box="[589,692,686,711]" pageId="51" pageNumber="52" type="lectotype">lectotype</typeStatus>
designation, and similarly nor can the usage of
<bibRefCitation id="EFC72AD50A36F543FB25CE80AAF9FD7A" author="Germain" pageId="51" pageNumber="52" refString="Germain, L. (1905 b) Sur les mollusques recueillis par les membres de la mission Foureau-Lamy dans le centre africain (suite). Bulletin du Museum d'Histoire naturelle, 11 (5), 327 - 331. [after 27 June; date of the 84 th reunion on the first page of the issue]" type="journal article" year="1905" yearSuffix="b">Germain (1905b: 329)</bibRefCitation>
.
<bibRefCitation id="EFC72AD50A36F543FF27CEFCABEBFD7A" author="Bourguignat" box="[216,479,722,747]" pageId="51" pageNumber="52" refString="Bourguignat, J. R. (1879) Description de divers especes terrestres et fluviatiles et de differents genres de mollusques de l'Egypte, de l'Abyssinie, de Zanzibar, du Senegal et du centre de l'Afrique. Imprimerie de Madame Veuve Bouchard- Huzard, Jules Tremblay, Gendre et Successeur, Paris, 54 pp. http: // dx. doi. org / 10.5962 / bhl. title. 50008" type="book" year="1879">Bourguignat (1879: 32)</bibRefCitation>
considered the shell in the other illustration of Philippi (pl. 5, fig. 4) to be sufficiently distinct from that of pl. 17, fig. 2 to warrant its recognition as variety
<emphasis id="B9228B360A36F543FC5ACED9A9DCFC9F" box="[933,1000,759,782]" italics="true" pageId="51" pageNumber="52">minor</emphasis>
Bourguignat (see
<emphasis id="B9228B360A36F543FB49CED9AECDFC9F" box="[1206,1273,759,782]" italics="true" pageId="51" pageNumber="52">minor</emphasis>
Bourguignat), the shell of pl. 5, fig. 4, as the only representative of
<emphasis id="B9228B360A36F543FD1ECF32A917FCA2" box="[737,803,796,819]" italics="true" pageId="51" pageNumber="52">minor</emphasis>
, thereby becoming the
<typeStatus id="54EDE9860A36F543FBD6CF34AEBFFCA2" box="[1065,1163,794,819]" pageId="51" pageNumber="52" type="holotype">holotype</typeStatus>
by monotypy.
<bibRefCitation id="EFC72AD50A36F543FACDCF34ABC1FCC6" author="Kohler" pageId="51" pageNumber="52" refString="Kohler, F. &amp; Glaubrecht, M. (2006) The types of Ampullariidae Gray, 1824 (Mollusca, Gastropoda) in the Malacological Collection of the Natural History Museum, Berlin: an annotated catalogue with lectotype designations. Mitteilungen aus dem Museum fur Naturkunde Berlin, Zoologische Reihe, 82 (1), 198 - 215." type="journal article" year="2006">Köhler &amp; Glaubrecht (2006: 212, fig. 3L)</bibRefCitation>
designated this specimen (pl. 5, fig. 4) as the
<typeStatus id="54EDE9860A36F543FC0DCF10AE6CFCC6" box="[1010,1112,830,855]" pageId="51" pageNumber="52" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C562CA70A36F543FB83CF11AEFCFCC7" box="[1148,1224,831,854]" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="wernei">
<emphasis id="B9228B360A36F543FB83CF11AEFCFCC7" box="[1148,1224,831,854]" italics="true" pageId="51" pageNumber="52">wernei</emphasis>
</taxonomicName>
, thereby rendering
<taxonomicName id="4C562CA70A36F543FF68CF4AAB7DFCEA" authority="Philippi" authorityName="Philippi" box="[151,329,866,891]" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="wernei">
<emphasis id="B9228B360A36F543FF68CF4AAAD7FCEA" box="[151,227,868,891]" italics="true" pageId="51" pageNumber="52">wernei</emphasis>
Philippi
</taxonomicName>
and
<emphasis id="B9228B360A36F543FE74CF4AABFAFCEA" box="[395,462,868,891]" italics="true" pageId="51" pageNumber="52">minor</emphasis>
Bourguignat objective synonyms. If Bourguignat was correct in thinking the two illustrations to be of different species, unfortunately it is the specimen that he considered to be typical of
<taxonomicName id="4C562CA70A36F543FAAFCFA9AAC4FC52" authority="Philippi" authorityName="Philippi" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="wernei">
<emphasis id="B9228B360A36F543FAAFCFA9AFA8FC0F" box="[1360,1436,903,926]" italics="true" pageId="51" pageNumber="52">wernei</emphasis>
Philippi
</taxonomicName>
(pl. 17, fig. 2) that would require a new name.
<bibRefCitation id="EFC72AD50A36F543FCFACF84A9D0FC52" author="Billotte" box="[773,996,938,963]" pageId="51" pageNumber="52" refString="Billotte, R. (1885) Recensement des ampullaires du continent africain, precede de diagnoses d'ampullaires nouvelles. Bulletin de la Societe malacologique de France, 2, 103 - 112, pl. 6. [July; p. 112]" type="journal article" year="1885">Billotte (1885: 107)</bibRefCitation>
treated
<emphasis id="B9228B360A36F543FBC0CF82AEB6FC52" box="[1087,1154,940,963]" italics="true" pageId="51" pageNumber="52">minor</emphasis>
Bourguignat as a variety of
<taxonomicName id="4C562CA70A36F543FF49CFE0ABC8FC76" authority="Billotte. Kobelt" authorityName="Billotte. Kobelt" box="[182,508,974,999]" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="charmesiana">
<emphasis id="B9228B360A36F543FF49CFE0AB73FC76" box="[182,327,974,999]" italics="true" pageId="51" pageNumber="52">charmesiana</emphasis>
Billotte. Kobelt
</taxonomicName>
(1912b: 62–63) discussed the confusion surrounding the three names and came to essentially the same conclusion that if considered distinct, the shell illustrated in pl. 17, fig. 2 would require a new name.
<bibRefCitation id="EFC72AD50A36F543FF1DC838ABA5FBBE" author="Pain" box="[226,401,1046,1071]" pageId="51" pageNumber="52" refString="Pain, T. (1961) Revision of the African Ampullariidae species of the genus Pila Roding [sic] 1798 (Mesogastropoda, Architaenioglossa, Mollusca). Musee Royal de l'Afrique Centrale-Tervuren, Belgique, Annales - Serie in 8 ° - Sciences Zoologiques, 96, [1] - [29]. [May; p. [29 ]]" type="journal article" year="1961">
<collectingCountry id="F34117B40A36F543FF1DC838AB20FBBE" box="[226,276,1046,1071]" name="Spain" pageId="51" pageNumber="52">Pain</collectingCountry>
(1961: 19)
</bibRefCitation>
listed
<emphasis id="B9228B360A36F543FE23C839A82BFBBF" box="[476,543,1047,1070]" italics="true" pageId="51" pageNumber="52">minor</emphasis>
Bourguignat (as “Kobelt”) as a synonym of
<taxonomicName id="4C562CA70A36F543FBE8C839AF1DFBBF" authority="Philippi. Reports" authorityName="Philippi. Reports" box="[1047,1321,1046,1071]" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="wernei">
<emphasis id="B9228B360A36F543FBE8C839AE57FBBF" box="[1047,1123,1047,1070]" italics="true" pageId="51" pageNumber="52">wernei</emphasis>
Philippi. Reports
</taxonomicName>
of
<taxonomicName id="4C562CA70A36F543FAAFC839AFA8FBBF" box="[1360,1436,1047,1070]" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="wernei">
<emphasis id="B9228B360A36F543FAAFC839AFA8FBBF" box="[1360,1436,1047,1070]" italics="true" pageId="51" pageNumber="52">wernei</emphasis>
</taxonomicName>
further south than the distribution above, e.g.
<collectingCountry id="F34117B40A36F543FD4EC814A971FBC2" box="[689,837,1082,1107]" name="Mozambique" pageId="51" pageNumber="52">Mozambique</collectingCountry>
and Lake Albert (
<bibRefCitation id="EFC72AD50A36F543FBE0C814AF13FBC2" author="Alderson" box="[1055,1319,1082,1107]" pageId="51" pageNumber="52" refString="Alderson, E. G. (1925) Studies in Ampullaria. Heffer, Cambridge, xx + 102 pp., 19 pls." type="book" year="1925">Alderson 1925: 89–90</bibRefCitation>
),
<collectingCountry id="F34117B40A36F543FAC1C814AFACFBC2" box="[1342,1432,1082,1107]" name="Uganda" pageId="51" pageNumber="52">Uganda</collectingCountry>
,
<collectingCountry id="F34117B40A36F543FF68C870AAC9FBE6" box="[151,253,1118,1143]" name="Tanzania" pageId="51" pageNumber="52">Zanzibar</collectingCountry>
,
<collectingCountry id="F34117B40A36F543FEF8C870AB53FBE6" box="[263,359,1118,1143]" name="Namibia" pageId="51" pageNumber="52">Namibia</collectingCountry>
and
<collectingCountry id="F34117B40A36F543FE61C870ABC4FBE6" box="[414,496,1118,1143]" name="Angola" pageId="51" pageNumber="52">Angola</collectingCountry>
(
<bibRefCitation id="EFC72AD50A36F543FE00C870A89FFBE6" author="Connolly" box="[511,683,1118,1143]" pageId="51" pageNumber="52" refString="Connolly, M. (1939) A monographic survey of South African non-marine Mollusca. Annals of the South African Museum, 33 (1), i - iii, 1 - 660, pls. 1 - 19. [December; front wrapper of issue]" type="journal article" year="1939">Connolly 1939</bibRefCitation>
: 553;
<bibRefCitation id="EFC72AD50A36F543FD10C870A950FBE6" author="Pain" box="[751,868,1118,1143]" pageId="51" pageNumber="52" refString="Pain, T. (1961) Revision of the African Ampullariidae species of the genus Pila Roding [sic] 1798 (Mesogastropoda, Architaenioglossa, Mollusca). Musee Royal de l'Afrique Centrale-Tervuren, Belgique, Annales - Serie in 8 ° - Sciences Zoologiques, 96, [1] - [29]. [May; p. [29 ]]" type="journal article" year="1961">
<collectingCountry id="F34117B40A36F543FD10C870A915FBE6" box="[751,801,1118,1143]" name="Spain" pageId="51" pageNumber="52">Pain</collectingCountry>
1961
</bibRefCitation>
: 19), are probably incorrect identifications (
<bibRefCitation id="EFC72AD50A36F543FAB1C871AB34FB0A" author="Brown" pageId="51" pageNumber="52" refString="Brown, D. S. (1994) Freshwater snails of Africa and their medical importance. Second edition. Taylor &amp; Francis, London, x + 608 + [1] pp." type="journal article" year="1994">Brown 1994: 56</bibRefCitation>
).
<taxonomicName id="4C562CA70A36F543FEEAC8ACAB70FB0A" box="[277,324,1154,1179]" class="Gastropoda" family="Ampullariidae" genus="Pila" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="51" pageNumber="52" phylum="Mollusca" rank="genus">
<emphasis id="B9228B360A36F543FEEAC8ACAB70FB0A" box="[277,324,1154,1179]" italics="true" pageId="51" pageNumber="52">Pila</emphasis>
</taxonomicName>
,
<emphasis id="B9228B360A36F543FEABC8AAABB3FB0A" box="[340,391,1156,1179]" italics="true" pageId="51" pageNumber="52">teste</emphasis>
<bibRefCitation id="EFC72AD50A36F543FE6EC8ACA8BEFB0A" author="Connolly" box="[401,650,1154,1179]" pageId="51" pageNumber="52" refString="Connolly, M. (1939) A monographic survey of South African non-marine Mollusca. Annals of the South African Museum, 33 (1), i - iii, 1 - 660, pls. 1 - 19. [December; front wrapper of issue]" type="journal article" year="1939">Connolly (1939: 553)</bibRefCitation>
,
<bibRefCitation id="EFC72AD50A36F543FD67C8ACA97FFB0A" author="Pain" box="[664,843,1154,1179]" pageId="51" pageNumber="52" refString="Pain, T. (1961) Revision of the African Ampullariidae species of the genus Pila Roding [sic] 1798 (Mesogastropoda, Architaenioglossa, Mollusca). Musee Royal de l'Afrique Centrale-Tervuren, Belgique, Annales - Serie in 8 ° - Sciences Zoologiques, 96, [1] - [29]. [May; p. [29 ]]" type="journal article" year="1961">
<collectingCountry id="F34117B40A36F543FD67C8ACA8FEFB0A" box="[664,714,1154,1179]" name="Spain" pageId="51" pageNumber="52">Pain</collectingCountry>
(1961: 18)
</bibRefCitation>
and
<bibRefCitation id="EFC72AD50A36F543FC7AC8ADAE62FB0A" author="Brown" box="[901,1110,1154,1179]" pageId="51" pageNumber="52" refString="Brown, D. S. (1994) Freshwater snails of Africa and their medical importance. Second edition. Taylor &amp; Francis, London, x + 608 + [1] pp." type="journal article" year="1994">Brown (1994: 56)</bibRefCitation>
. Misspelled as “
<emphasis id="B9228B360A36F543FAECC8AAAF5EFB0A" box="[1299,1386,1156,1179]" italics="true" pageId="51" pageNumber="52">werneri</emphasis>
” by
<bibRefCitation id="EFC72AD50A36F543FF68C888AB43FB2E" author="Gaudion" box="[151,375,1190,1215]" pageId="51" pageNumber="52" refString="Gaudion, H. (1880) Liste alphabetique des especes du genre Ampullaria de Lamarck. Bulletin de la Societe d'Etude des Sciences naturelles de Beziers, 4 [for 1879], 20 - 43. [1880 on title page of volume]" type="journal article" year="1880">Gaudion (1880: 42)</bibRefCitation>
.
</paragraph>
<paragraph id="8BE957240A36F543FF68C8C0AAC0FA96" blockId="51.[151,1436,1262,1431]" box="[151,244,1262,1287]" pageId="51" pageNumber="52">
<taxonomicName id="4C562CA70A36F543FF68C8C0AAC0FA96" box="[151,244,1262,1287]" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="winkleyi">
<emphasis id="B9228B360A36F543FF68C8C0AAC0FA96" box="[151,244,1262,1287]" italics="true" pageId="51" pageNumber="52">winkleyi</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C34C04AF0A36F543FF38C93CA9F6FA06" pageId="51" pageNumber="52" type="reference_group">
<paragraph id="8BE957240A36F543FF38C93CA9F6FA06" blockId="51.[151,1436,1262,1431]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="AB46700A0A36F543FF38C93CA9F6FA06" infered="true" pageId="51" pageNumber="52">
<taxonomicName id="4C562CA70A36F543FF38C93CA8B3FABA" authority="Pilsbry, 1901: 189" authorityName="Pilsbry" authorityPageNumber="189" authorityYear="1901" box="[199,647,1298,1323]" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="winkleyi">
<emphasis id="B9228B360A36F543FF38C93CAB98FABA" box="[199,428,1298,1323]" italics="true" pageId="51" pageNumber="52">Ampullaria winkleyi</emphasis>
<treatmentCitation id="0AF771350A36F543FE4AC93CA8B3FABA" author="Pilsbry" box="[437,647,1298,1323]" page="189" pageId="51" pageNumber="52" year="1901">
<bibRefCitation id="EFC72AD50A36F543FE4AC93CA87AFABA" author="Pilsbry" box="[437,590,1298,1323]" pageId="51" pageNumber="52" refString="Pilsbry, H. A. (1901) New species of mollusks from South Africa and Burma. Proceedings of the Academy of Natural Sciences of Philadelphia, 53, 188 - 190, pl. 5 (part). [2 May; front matter of volume]" type="journal article" year="1901">Pilsbry, 1901</bibRefCitation>
: 189
</treatmentCitation>
</taxonomicName>
, pl. 5, figs. 2, 3.
<emphasis id="B9228B360A36F543FCB3C93AA9DCFABA" box="[844,1000,1298,1323]" italics="true" pageId="51" pageNumber="52">
<typeStatus id="54EDE9860A36F543FCB3C93AA9B4FABA" box="[844,896,1300,1323]" pageId="51" pageNumber="52">Type</typeStatus>
material
</emphasis>
—
<typeStatus id="54EDE9860A36F543FBFBC93CAE5EFABA" box="[1028,1130,1298,1323]" pageId="51" pageNumber="52" type="lectotype">lectotype</typeStatus>
(
<treatmentCitation id="0AF771350A36F543FB84C93CAF0BFABA" author="Baker" box="[1147,1343,1298,1323]" page="168" pageId="51" pageNumber="52" year="1964">
<bibRefCitation id="EFC72AD50A36F543FB84C93CAF0BFABA" author="Baker" box="[1147,1343,1298,1323]" pageId="51" pageNumber="52" refString="Baker, H. B. (1964) Type land snails in the Academy of Natural Sciences of Philadelphia. Part III. Limnophile and thalassophile Pulmonata. Part IV. Land and fresh-water Prosobranchia. Proceedings of the Academy of Natural Sciences of Philadelphia, 116 (4), 149 - 193. [20 October; cover of issue, contents table of volume]" type="journal article" year="1964">
<collectingCountry id="F34117B40A36F543FB84C93CAEF4FABA" box="[1147,1216,1298,1323]" name="United States Pacific Island Wildlife Refuges" pageId="51" pageNumber="52">Baker</collectingCountry>
1964: 168
</bibRefCitation>
</treatmentCitation>
): ANSP 76011;
<typeStatus id="54EDE9860A36F543FF14C918ABA5FADE" box="[235,401,1334,1359]" pageId="51" pageNumber="52" type="paralectotype">paralectotypes</typeStatus>
: ANSP 365369 (13 spms.), UMMZ 93221 (3 spms.).
<emphasis id="B9228B360A36F543FC15C919AE4DFADE" box="[1002,1145,1334,1359]" italics="true" pageId="51" pageNumber="52">
<typeStatus id="54EDE9860A36F543FC15C919AE2AFADF" box="[1002,1054,1335,1358]" pageId="51" pageNumber="52">Type</typeStatus>
locality
</emphasis>
—“Henzada,
<collectingCountry id="F34117B40A36F543FAEDC919AF50FADE" box="[1298,1380,1335,1359]" name="Myanmar" pageId="51" pageNumber="52">Burma</collectingCountry>
”.
<emphasis id="B9228B360A36F543FF38C974AB1DFAE2" box="[199,297,1370,1395]" italics="true" pageId="51" pageNumber="52">Remarks</emphasis>
. Valid species,
<emphasis id="B9228B360A36F543FE1DC972A821FAE2" box="[482,533,1372,1395]" italics="true" pageId="51" pageNumber="52">teste</emphasis>
G.B. Sowerby III (1910: 62) and
<treatmentCitation id="0AF771350A36F543FC5AC975AEB9FAE2" author="Preston" box="[933,1165,1370,1395]" page="103" pageId="51" pageNumber="52" year="1915">
<bibRefCitation id="EFC72AD50A36F543FC5AC975AEB9FAE2" author="Preston" box="[933,1165,1370,1395]" pageId="51" pageNumber="52" refString="Preston, H. B. (1915) The fauna of British India, including Ceylon and Burma. Mollusca. (Freshwater Gastropoda &amp; Pelecypoda). Taylor and Francis, London, xiv + 244 pp. [March; cover page]" type="book" year="1915">Preston (1915: 103)</bibRefCitation>
</treatmentCitation>
. Synonym of
<emphasis id="B9228B360A36F543FACDC974AFA8FAE2" box="[1330,1436,1370,1395]" italics="true" pageId="51" pageNumber="52">theobaldi</emphasis>
Hanley &amp; Theobald,
<emphasis id="B9228B360A36F543FE7CC951AB82FA07" box="[387,438,1407,1430]" italics="true" pageId="51" pageNumber="52">teste</emphasis>
<treatmentCitation id="0AF771350A36F543FE42C950A8AEFA06" author="Prashad" box="[445,666,1406,1431]" page="589" pageId="51" pageNumber="52" year="1923">
<bibRefCitation id="EFC72AD50A36F543FE42C950A857FA06" author="Prashad" box="[445,611,1406,1431]" pageId="51" pageNumber="52" refString="Prashad, B. (1923) Revision of Kobelt's nomenclature of the Indian Ampullariidae. Journal and Proceedings of the Asiatic Society of Bengal, New Series, 18 (10 [for 1922]), 585 - 591 [30 November; p. 591]" type="journal article" year="1923">Prashad (1923</bibRefCitation>
: 589
</treatmentCitation>
, 1925: 77), followed here.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
</treatment>
</document>
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