361 lines
36 KiB
XML
361 lines
36 KiB
XML
<document id="8BBCE151302BEEBC31A01A225170AA8B" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFFFD735F32FAFBD2E69CFA6B" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Zoosphaerium libidinosum" docType="key" docVersion="4" lastPageNumber="16" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="15" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="5E3C309E1799C66D65135AEF6DE3A11C">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
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<mods:namePart id="F571CD7E19A04B78F8EA5F1775130F7F">Wesener, Thomas</mods:namePart>
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<mods:title id="DCD57CEEE3E07C5CF9413B6B874330E3">Zootaxa</mods:title>
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<mods:part id="32D0EA803474B6A9293B5C89F7348592">
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<mods:date id="45BC46BE61233F3AA5A532E71FB2A672">2009</mods:date>
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<mods:number id="9ACD8B8FDD22D96B2ACA53267D3DFECB">2009-05-11</mods:number>
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<mods:number id="BF671222A2084ECCC3508C10AB069705">2097</mods:number>
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<mods:url id="66C17249EF03C30E858A3D1CC19E7E40">https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1</mods:url>
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<mods:classification id="5CE94A969034771F00F14FDED276AA78">journal article</mods:classification>
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<mods:identifier id="FC8BDFFBF86EA3A6F522DE2DBF2E78FB" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
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<mods:identifier id="C12D38FC65D6C83066A791076141E9D6" type="ISSN">11755334</mods:identifier>
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<treatment id="03FFBE1DFFFD735F32FAFBD2E69CFA6B" ID-DOI="http://doi.org/10.5281/zenodo.5325439" ID-Zenodo-Dep="5325439" LSID="urn:lsid:plazi:treatment:03FFBE1DFFFD735F32FAFBD2E69CFA6B" httpUri="http://treatment.plazi.org/id/03FFBE1DFFFD735F32FAFBD2E69CFA6B" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
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<subSubSection id="C34C5C80FFFD735E32FAFBD2E1FAFB5B" box="[151,765,1128,1155]" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="8BE90F0BFFFD735E32FAFBD2E1FAFB5B" blockId="14.[151,765,1128,1155]" box="[151,765,1128,1155]" pageId="14" pageNumber="15">
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<heading id="D0A1B867FFFD735E32FAFBD2E1FAFB5B" bold="true" box="[151,765,1128,1155]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
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<emphasis id="B922D319FFFD735E32FAFBD2E1FAFB5B" bold="true" box="[151,765,1128,1155]" pageId="14" pageNumber="15">
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Key to species of the
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<taxonomicName id="4C567488FFFD735E33F1FBD3E14BFB5A" ID-CoL="5D7HG" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[412,588,1129,1155]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="libidinosum">
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<emphasis id="B922D319FFFD735E33F1FBD3E14BFB5A" bold="true" box="[412,588,1129,1155]" italics="true" pageId="14" pageNumber="15">Z. libidinosum</emphasis>
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</taxonomicName>
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species-group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34C5C80FFFD735F32FAFB08E69CFA6B" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="key">
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<keyStep id="30A217AEFFFD735E32FAFB08E692FB35" pageId="14" pageNumber="15">
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||
<paragraph id="8BE90F0BFFFD735E32FAFB08E69CFB13" blockId="14.[151,1436,1202,2023]" box="[151,1435,1202,1226]" pageId="14" pageNumber="15">
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<keyLead id="30A7AC3EFFFD735E32FAFB08E69CFB13" box="[151,1435,1202,1226]" pageId="14" pageNumber="15">
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<emphasis id="B922D319FFFD735E32FAFB08E3A3FB13" bold="true" box="[151,164,1202,1226]" pageId="14" pageNumber="15">1</emphasis>
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. Anal shield without locking carinae (in small, immature individuals sometimes traces of carinae are visible). .......
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<emphasis id="B922D319FFFD735E37E3FB08E69CFB13" bold="true" box="[1422,1435,1202,1226]" pageId="14" pageNumber="15">2</emphasis>
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</keyLead>
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</paragraph>
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<paragraph id="8BE90F0BFFFD735E32FAFB6FE692FB35" blockId="14.[151,1436,1202,2023]" box="[151,1429,1236,1260]" pageId="14" pageNumber="15">
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<keyLead id="30A7AC3EFFFD735E32FAFB6FE692FB35" box="[151,1429,1236,1260]" pageId="14" pageNumber="15">
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- Anal shield with two obvious locking carinae. ...........................................................................................................
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<emphasis id="B922D319FFFD735E37E5FB6EE692FB35" bold="true" box="[1416,1429,1236,1260]" pageId="14" pageNumber="15">6</emphasis>
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</keyLead>
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||
</paragraph>
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||
</keyStep>
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||
<keyStep id="30A217AEFFFD735E32FAFB4DE69CFAAE" pageId="14" pageNumber="15">
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||
<paragraph id="8BE90F0BFFFD735E32FAFB4DE69CFAEB" blockId="14.[151,1436,1202,2023]" pageId="14" pageNumber="15">
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||
<keyLead id="30A7AC3EFFFD735E32FAFB4DE69CFAEB" pageId="14" pageNumber="15">
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<emphasis id="B922D319FFFD735E32FAFB4DE3A3FAD6" bold="true" box="[151,164,1271,1295]" pageId="14" pageNumber="15">2</emphasis>
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. Operculum of vulva at least weakly notched apically (Fig. 7C). Fifth antennomere with field of
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<emphasis id="B922D319FFFD735E36A2FB42E69CFAD6" box="[1231,1435,1272,1295]" italics="true" pageId="14" pageNumber="15">sensilla basiconica</emphasis>
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apically (Fig. 8C) .........................................................................................................................................................
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<emphasis id="B922D319FFFD735E37E3FAA0E69CFAEB" bold="true" box="[1422,1435,1306,1330]" pageId="14" pageNumber="15">3</emphasis>
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</keyLead>
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||
</paragraph>
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<paragraph id="8BE90F0BFFFD735E32FAFA87E69CFAAE" blockId="14.[151,1436,1202,2023]" pageId="14" pageNumber="15">
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<keyLead id="30A7AC3EFFFD735E32FAFA87E69CFAAE" pageId="14" pageNumber="15">
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- Operculum of vulva well-rounded apically (Fig. 14A). No
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<emphasis id="B922D319FFFD735E3120FA87E71CFA8D" box="[845,1051,1341,1364]" italics="true" pageId="14" pageNumber="15">sensilla basiconica</emphasis>
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present on fifth antennomere (Fig. 15A). ............................................................................................................................................................................
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<emphasis id="B922D319FFFD735E37E3FAE5E69CFAAE" bold="true" box="[1422,1435,1375,1399]" pageId="14" pageNumber="15">4</emphasis>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30A217AEFFFD735E32FAFA38E69CF9B3" pageId="14" pageNumber="15">
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<paragraph id="8BE90F0BFFFD735E32FAFA38E692F9DB" blockId="14.[151,1436,1202,2023]" pageId="14" pageNumber="15">
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<keyLead id="30A7AC3EFFFD735E32FAFA38E692F9DB" pageId="14" pageNumber="15">
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<emphasis id="B922D319FFFD735E32FAFA38E3A3FA43" bold="true" box="[151,164,1410,1434]" pageId="14" pageNumber="15">3</emphasis>
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. Tergites dark green, in alcohol almost blackish, posterior margin light brown to yellow (Fig. 6A). Tarsus of walking legs broad, only 3.0 times longer than wide (Fig. 7A). Third podomere of anterior telopods with four large, sclerotized teeth (Fig. 7F). Endotergum marginal bristles strongly extending beyond tergite (Fig. 23B). ....................
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<emphasis id="B922D319FFFD735E32AAFA50E655F9D8" box="[199,1362,1514,1537]" italics="true" pageId="14" pageNumber="15">
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...........................................................................................................................................................
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<taxonomicName id="4C567488FFFD735E36BFFA50E648F9D8" ID-CoL="5D7H5" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1234,1359,1514,1537]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="haackeri" status="sp. nov.">Z. haackeri</taxonomicName>
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,
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</emphasis>
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<emphasis id="B922D319FFFD735E3734FA50E692F9DB" bold="true" box="[1369,1429,1514,1538]" pageId="14" pageNumber="15">
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<taxonomicNameLabel id="A2116E62FFFD735E3734FA50E692F9DB" box="[1369,1429,1514,1538]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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||
</keyLead>
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||
</paragraph>
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<paragraph id="8BE90F0BFFFD735E32FAF9B7E69CF9B3" blockId="14.[151,1436,1202,2023]" pageId="14" pageNumber="15">
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<keyLead id="30A7AC3EFFFD735E32FAF9B7E69CF9B3" pageId="14" pageNumber="15">
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||
- Tergites white, only dorso-laterally on each side with light, shiny-green field (Fig. 1C). Tarsus of walking legs of usual size, 3.8 times longer than wide. Third podomere of anterior telopods with single large, sclerotized tooth (Fig. 10D). Endotergum marginal bristles slightly extending beyond tergite (Fig. 23C).
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<emphasis id="B922D319FFFD735E365DF9E8E65EF9B0" box="[1072,1369,1618,1641]" italics="true" pageId="14" pageNumber="15">
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||
.............................
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<taxonomicName id="4C567488FFFD735E3699F9E8E652F9B0" ID-CoL="5D7G9" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1268,1365,1618,1641]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="album" status="sp. nov.">Z. album</taxonomicName>
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,
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</emphasis>
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<emphasis id="B922D319FFFD735E370DF9E8E69CF9B3" bold="true" box="[1376,1435,1618,1642]" pageId="14" pageNumber="15">
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<taxonomicNameLabel id="A2116E62FFFD735E370DF9E8E69CF9B3" box="[1376,1435,1618,1642]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30A217AEFFFD735E32FAF9CEE69CF8E3" pageId="14" pageNumber="15">
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<paragraph id="8BE90F0BFFFD735E32FAF9CEE69CF92D" blockId="14.[151,1436,1202,2023]" pageId="14" pageNumber="15">
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<keyLead id="30A7AC3EFFFD735E32FAF9CEE69CF92D" pageId="14" pageNumber="15">
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<emphasis id="B922D319FFFD735E32FAF9CEE3A3F955" bold="true" box="[151,164,1652,1676]" pageId="14" pageNumber="15">4</emphasis>
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. Male harp with two, weakly-developed stridulation ribs (Fig. 14B). Anterior telopod second podomere process slightly extending beyond third podomere (Fig. 14B). Third podomere laterally with plateau and mesally with sharp, prominent process (Figs 14B–D). Tergites light green with distinct, light-brown posterior margin (Fig. 6B). .............
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<emphasis id="B922D319FFFD735E32AAF967E65EF92D" box="[199,1369,1757,1780]" italics="true" pageId="14" pageNumber="15">
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.............................................................................................................................................................
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<taxonomicName id="4C567488FFFD735E368FF967E652F92D" ID-CoL="5D7GW" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1250,1365,1757,1780]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="discolor" status="sp. nov.">Z. discolor</taxonomicName>
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,
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</emphasis>
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<emphasis id="B922D319FFFD735E370DF966E69CF92D" bold="true" box="[1376,1435,1756,1780]" pageId="14" pageNumber="15">
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||
<taxonomicNameLabel id="A2116E62FFFD735E370DF966E69CF92D" box="[1376,1435,1756,1780]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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||
</keyLead>
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||
</paragraph>
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<paragraph id="8BE90F0BFFFD735E32FAF8BAE69CF8E3" blockId="14.[151,1436,1202,2023]" pageId="14" pageNumber="15">
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||
<keyLead id="30A7AC3EFFFD735E32FAF8BAE69CF8E3" pageId="14" pageNumber="15">
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- Male harp with two stridulation ribs. Anterior telopod second podomere process shorter than third podomere. Third podomere of anterior telopods only laterally with plateau. ........................................................................................
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<emphasis id="B922D319FFFD735E37E3F898E69CF8E3" bold="true" box="[1422,1435,1826,1850]" pageId="14" pageNumber="15">5</emphasis>
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||
</keyLead>
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||
</paragraph>
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||
</keyStep>
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||
<keyStep id="30A217AEFFFD735F32FAF8FEE69CFF76" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
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||
<paragraph id="8BE90F0BFFFD735E32FAF8FEE69BF878" blockId="14.[151,1436,1202,2023]" pageId="14" pageNumber="15">
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||
<keyLead id="30A7AC3EFFFD735E32FAF8FEE69BF878" pageId="14" pageNumber="15">
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||
<emphasis id="B922D319FFFD735E32FAF8FEE3A3F885" bold="true" box="[151,164,1860,1884]" pageId="14" pageNumber="15">5</emphasis>
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||
. Tergites with dark-green, light green pattern, posteriorly with distinct, thin light-brown margin. First antennomere with
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||
<emphasis id="B922D319FFFD735E3292F8D2E2CBF8A6" box="[255,460,1896,1919]" italics="true" pageId="14" pageNumber="15">sensilla basiconica</emphasis>
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||
. Anal shield glabrous. Endotergum with>3 rows of marginal bristles, strongly extending beyond tergite. ....................................................................................
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||
<taxonomicName id="4C567488FFFD735E31CAF831E69BF878" ID-CoL="5D7HG" authority="(de Saussure & Zehntner, 1897)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[935,1436,1930,1954]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="libidinosum">
|
||
<emphasis id="B922D319FFFD735E31CAF831E746F87B" box="[935,1089,1931,1954]" italics="true" pageId="14" pageNumber="15">Z. libidinosum</emphasis>
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||
(
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||
<bibRefCitation id="EFC772FAFFFD735E3622F830E693F878" author="de Saussure, H. & Zehntner, L." box="[1103,1428,1930,1953]" pageId="14" pageNumber="15" pagination="1 - 12" refId="ref60128" refString="de Saussure, H. & Zehntner, L. (1897) Atlas de l'histoire naturelle des Myriapodes, In: Grandidier (Ed.) Histoire physique, naturelle et politique de Madagascar. 27 (53), Alfred Grandidier, Paris, France, plates 1 - 12." type="book chapter" year="1897">de Saussure & Zehntner, 1897</bibRefCitation>
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)
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||
</taxonomicName>
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||
</keyLead>
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||
</paragraph>
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<paragraph id="8BE90F0BFFFD735F32FAF817E69CFF76" blockId="14.[151,1436,1202,2023]" lastBlockId="15.[151,1436,151,1458]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
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||
<keyLead id="30A7AC3EFFFD735F32FAF817E69CFF76" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
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||
- Tergites dark, almost blackish-green, posteriorly with distinct and wide light brown margin (Fig. 17A). First antennomere without
|
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<emphasis id="B922D319FFFD735E33D8F86AE184F83E" box="[437,643,2000,2023]" italics="true" pageId="14" pageNumber="15">sensilla basiconica</emphasis>
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||
(Fig. 19A). Anal shield covered with numerous short hair. Endotergum with single row of marginal bristles, slightly extending beyond tergite (Fig. 23E).
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<emphasis id="B922D319FFFC735F365DFF22E65EFF76" box="[1072,1369,152,175]" italics="true" pageId="15" pageNumber="16">
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||
............................
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<taxonomicName id="4C567488FFFC735F3680FF22E652FF76" ID-CoL="5D7HJ" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1261,1365,152,175]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="mitoho" status="sp. nov.">Z. mitoho</taxonomicName>
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||
,
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||
</emphasis>
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<emphasis id="B922D319FFFC735F370DFF2DE69CFF76" bold="true" box="[1376,1435,151,175]" pageId="15" pageNumber="16">
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||
<taxonomicNameLabel id="A2116E62FFFC735F370DFF2DE69CFF76" box="[1376,1435,151,175]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
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||
</keyStep>
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||
<keyStep id="30A217AEFFFC735F32FAFF00E69CFF2D" pageId="15" pageNumber="16">
|
||
<paragraph id="8BE90F0BFFFC735F32FAFF00E69CFF0B" blockId="15.[151,1436,151,1458]" box="[151,1435,186,210]" pageId="15" pageNumber="16">
|
||
<keyLead id="30A7AC3EFFFC735F32FAFF00E69CFF0B" box="[151,1435,186,210]" pageId="15" pageNumber="16">
|
||
<emphasis id="B922D319FFFC735F32FAFF00E3A3FF0B" bold="true" box="[151,164,186,210]" pageId="15" pageNumber="16">6</emphasis>
|
||
. Second and third podomeres of posterior telopods covered with numerous minute hairs (Fig. 21E). .......................
|
||
<emphasis id="B922D319FFFC735F37E3FF00E69CFF0B" bold="true" box="[1422,1435,186,210]" pageId="15" pageNumber="16">7</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFFC735F32FAFF67E69CFF2D" blockId="15.[151,1436,151,1458]" box="[151,1435,220,244]" pageId="15" pageNumber="16">
|
||
<keyLead id="30A7AC3EFFFC735F32FAFF67E69CFF2D" box="[151,1435,220,244]" pageId="15" pageNumber="16">
|
||
- Second and third podomere of posterior telopods glabrous or covered with long setae. ...........................................
|
||
<emphasis id="B922D319FFFC735F37E3FF66E69CFF2D" bold="true" box="[1422,1435,220,244]" pageId="15" pageNumber="16">8</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A217AEFFFC735F32FAFF45E69CFE3E" pageId="15" pageNumber="16">
|
||
<paragraph id="8BE90F0BFFFC735F32FAFF45E69CFEA6" blockId="15.[151,1436,151,1458]" pageId="15" pageNumber="16">
|
||
<keyLead id="30A7AC3EFFFC735F32FAFF45E69CFEA6" pageId="15" pageNumber="16">
|
||
<emphasis id="B922D319FFFC735F32FAFF45E3A3FECE" bold="true" box="[151,164,255,279]" pageId="15" pageNumber="16">7</emphasis>
|
||
. Tergites glossy, completely jade-like green. Male harp with two basally fused stridulation ribs. Third podomere of anterior telopods mesally with sharp, prominent process. Endotergum with one row of rounded cuticular impressions. Posterior telopod fixed finger basally without membranous lobe. ............................................................
|
||
<emphasis id="B922D319FFFC735F32AAFED2E0B6FEA6" box="[199,945,360,383]" italics="true" pageId="15" pageNumber="16">................................................................................................................</emphasis>
|
||
<taxonomicName id="4C567488FFFC735F31D9FED2E69CFEA6" ID-CoL="5D7GG" authority="(de Saussure & Zehntner, 1902)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1902" box="[948,1435,360,383]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="anomalum">
|
||
<emphasis id="B922D319FFFC735F31D9FED2E747FEA6" box="[948,1088,360,383]" italics="true" pageId="15" pageNumber="16">Z. anomalum</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC772FAFFFC735F3622FED2E693FEA6" author="de Saussure, H. & Zehntner, L." box="[1103,1428,360,383]" pageId="15" pageNumber="16" pagination="1 - 356" refId="ref60181" refString="de Saussure, H. & Zehntner, L. (1902) Myriapodes de Madagascar, In: Grandidier (Ed.), Histoire physique, naturelle et politique de Madagascar. 27 (53), Alfred Grandidier, Paris, France, i - viii, pp. 1 - 356, plates 13 - 15." type="journal article" year="1902">de Saussure & Zehntner, 1902</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFFC735F32FAFE30E69CFE3E" blockId="15.[151,1436,151,1458]" pageId="15" pageNumber="16">
|
||
<keyLead id="30A7AC3EFFFC735F32FAFE30E69CFE3E" pageId="15" pageNumber="16">
|
||
- Tergites glossy, jade-like green, with shiny dark-green pattern (Fig. 17B). Male harp with two stridulation ribs (Fig. 21B). Third podomere of anterior telopods without process (Fig. 21C). Endotergum without cuticular impressions (Fig. 23F). Posterior telopod fixed finger with membranous lobe (Fig. 21E).
|
||
<emphasis id="B922D319FFFC735F3199FE6AE65EFE3E" box="[1012,1369,464,487]" italics="true" pageId="15" pageNumber="16">
|
||
..............................
|
||
<taxonomicName id="4C567488FFFC735F36ADFE6AE652FE3E" ID-CoL="7GH2R" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1216,1365,464,487]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pulchellum" status="sp. nov.">Z. pulchellum</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B922D319FFFC735F370DFE75E69CFE3E" bold="true" box="[1376,1435,463,487]" pageId="15" pageNumber="16">
|
||
<taxonomicNameLabel id="A2116E62FFFC735F370DFE75E69CFE3E" box="[1376,1435,463,487]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A217AEFFFC735F32FAFE48E691FDF5" pageId="15" pageNumber="16">
|
||
<paragraph id="8BE90F0BFFFC735F32FAFE48E69CFDD3" blockId="15.[151,1436,151,1458]" box="[151,1435,498,522]" pageId="15" pageNumber="16">
|
||
<keyLead id="30A7AC3EFFFC735F32FAFE48E69CFDD3" box="[151,1435,498,522]" pageId="15" pageNumber="16">
|
||
<emphasis id="B922D319FFFC735F32FAFE48E3A3FDD3" bold="true" box="[151,164,498,522]" pageId="15" pageNumber="16">8</emphasis>
|
||
. Posterior end of anal shield in both sexes excavated. ..................................................................................................
|
||
<emphasis id="B922D319FFFC735F37E3FE48E69CFDD3" bold="true" box="[1422,1435,498,522]" pageId="15" pageNumber="16">9</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFFC735F32FAFDAFE691FDF5" blockId="15.[151,1436,151,1458]" box="[151,1430,532,556]" pageId="15" pageNumber="16">
|
||
<keyLead id="30A7AC3EFFFC735F32FAFDAFE691FDF5" box="[151,1430,532,556]" pageId="15" pageNumber="16">
|
||
- Posterior end of anal shield well-rounded.................................................................................................................
|
||
<emphasis id="B922D319FFFC735F3716FDAEE691FDF5" bold="true" box="[1403,1430,532,556]" pageId="15" pageNumber="16">12</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A217AEFFFC735F32FAFD8DE69CFD4D" pageId="15" pageNumber="16">
|
||
<paragraph id="8BE90F0BFFFC735F32FAFD8DE69CFDAB" blockId="15.[151,1436,151,1458]" pageId="15" pageNumber="16">
|
||
<keyLead id="30A7AC3EFFFC735F32FAFD8DE69CFDAB" pageId="15" pageNumber="16">
|
||
<emphasis id="B922D319FFFC735F32FAFD8DE3A3FD96" bold="true" box="[151,164,567,591]" pageId="15" pageNumber="16">9</emphasis>
|
||
. Tergites glossy, green. Male harp with two weakly-developed stridulation ribs (Fig. 24A). Anterior telopod third podomere with single, large, sclerotized tooth (Fig. 24B).
|
||
<emphasis id="B922D319FFFC735F315EFDE0E65EFDA8" box="[819,1369,602,625]" italics="true" pageId="15" pageNumber="16">
|
||
...........................................................
|
||
<taxonomicName id="4C567488FFFC735F36D2FDE0E652FDA8" ID-CoL="5D7GY" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1215,1365,602,625]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="endemicum" status="sp. nov.">Z. endemicum</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B922D319FFFC735F370DFDE0E69CFDAB" bold="true" box="[1376,1435,602,626]" pageId="15" pageNumber="16">
|
||
<taxonomicNameLabel id="A2116E62FFFC735F370DFDE0E69CFDAB" box="[1376,1435,602,626]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFFC735F32FAFDC7E69CFD4D" blockId="15.[151,1436,151,1458]" box="[151,1435,636,660]" pageId="15" pageNumber="16">
|
||
<keyLead id="30A7AC3EFFFC735F32FAFDC7E69CFD4D" box="[151,1435,636,660]" pageId="15" pageNumber="16">
|
||
- Tergites matte, brownish. Male harp with two well-developed stridulation ribs (Fig. 26D). ...................................
|
||
<emphasis id="B922D319FFFC735F37ECFDC6E69CFD4D" bold="true" box="[1409,1435,636,660]" pageId="15" pageNumber="16">10</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A217AEFFFC735F32FAFD25E69BFCC6" pageId="15" pageNumber="16">
|
||
<paragraph id="8BE90F0BFFFC735F32FAFD25E690FD03" blockId="15.[151,1436,151,1458]" pageId="15" pageNumber="16">
|
||
<keyLead id="30A7AC3EFFFC735F32FAFD25E690FD03" pageId="15" pageNumber="16">
|
||
<emphasis id="B922D319FFFC735F32FAFD25E3B5FD6E" bold="true" box="[151,178,671,695]" pageId="15" pageNumber="16">10</emphasis>
|
||
. Locking carinae on anal shield of similar length. Third podomere of posterior telopods with
|
||
<emphasis id="B922D319FFFC735F369BFD1AE62BFD6E" box="[1270,1324,672,695]" italics="true" pageId="15" pageNumber="16">circa</emphasis>
|
||
28 large, sclerotized teeth (Fig. 26H). Antenna with more than 40 apical cones (Fig. 27C).
|
||
<emphasis id="B922D319FFFC735F3647FD78E65FFD00" box="[1066,1368,706,729]" italics="true" pageId="15" pageNumber="16">
|
||
......................
|
||
<taxonomicName id="4C567488FFFC735F36D2FD78E654FD00" ID-CoL="5D7JG" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1215,1363,706,729]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="xerophilum" status="sp. nov.">Z. xerophilum</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B922D319FFFC735F3731FD78E690FD03" bold="true" box="[1372,1431,706,730]" pageId="15" pageNumber="16">
|
||
<taxonomicNameLabel id="A2116E62FFFC735F3731FD78E690FD03" box="[1372,1431,706,730]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFFC735F32FAFD5FE69BFCC6" blockId="15.[151,1436,151,1458]" pageId="15" pageNumber="16">
|
||
<keyLead id="30A7AC3EFFFC735F32FAFD5FE69BFCC6" pageId="15" pageNumber="16">
|
||
- Posterior locking carina at least two times as long as anterior carina. Third podomere of posterior telopods with less than 25 large, sclerotized teeth. Antenna with less than 30 apical cones. ................................................................
|
||
<emphasis id="B922D319FFFC735F37EEFCBDE69BFCC6" bold="true" box="[1411,1436,775,799]" pageId="15" pageNumber="16">11</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A217AEFFFC735F32FAFC90E69CFC5E" pageId="15" pageNumber="16">
|
||
<paragraph id="8BE90F0BFFFC735F32FAFC90E69BFCBD" blockId="15.[151,1436,151,1458]" pageId="15" pageNumber="16">
|
||
<keyLead id="30A7AC3EFFFC735F32FAFC90E69BFCBD" pageId="15" pageNumber="16">
|
||
<emphasis id="B922D319FFFC735F32FAFC90E3B7FC9B" bold="true" box="[151,176,810,834]" pageId="15" pageNumber="16">11</emphasis>
|
||
. Posterior locking carina of anal shield more than three times longer than anterior carina.............................................
|
||
<emphasis id="B922D319FFFC735F32AAFCF7E0D4FCBD" box="[199,979,845,868]" italics="true" pageId="15" pageNumber="16">.....................................................................................................................</emphasis>
|
||
<taxonomicName id="4C567488FFFC735F31B4FCF7E68FFCBD" ID-CoL="5D7GH" authority="Wesener & Sierwald, 2005" authorityName="Wesener & Sierwald" authorityYear="2005" box="[985,1416,845,868]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="arborealis">
|
||
<emphasis id="B922D319FFFC735F31B4FCF7E763FCBD" box="[985,1124,845,868]" italics="true" pageId="15" pageNumber="16">Z. arborealis</emphasis>
|
||
Wesener & Sierwald, 2005
|
||
</taxonomicName>
|
||
A
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFFC735F32FAFCCAE69CFC5E" blockId="15.[151,1436,151,1458]" box="[151,1435,880,903]" pageId="15" pageNumber="16">
|
||
<keyLead id="30A7AC3EFFFC735F32FAFCCAE69CFC5E" box="[151,1435,880,903]" pageId="15" pageNumber="16">
|
||
- Posterior locking carina
|
||
<emphasis id="B922D319FFFC735F33BFFCCAE10FFC5E" box="[466,520,880,903]" italics="true" pageId="15" pageNumber="16">circa</emphasis>
|
||
twice as long as anterior one
|
||
<emphasis id="B922D319FFFC735F312AFCCAE730FC5E" box="[839,1079,880,903]" italics="true" pageId="15" pageNumber="16">....................................</emphasis>
|
||
<taxonomicName id="4C567488FFFC735F3654FCCAE69CFC5E" ID-CoL="5D7HD" authority="(Brolemann, 1922)" baseAuthorityName="Brolemann" baseAuthorityYear="1922" box="[1081,1435,880,903]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="lambertoni">
|
||
<emphasis id="B922D319FFFC735F3654FCCAE7CDFC5E" box="[1081,1226,880,903]" italics="true" pageId="15" pageNumber="16">Z. lambertoni</emphasis>
|
||
(Brolemann, 1922)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A217AEFFFC735F32FAFC28E691FB45" pageId="15" pageNumber="16">
|
||
<paragraph id="8BE90F0BFFFC735F32FAFC28E69CFBCB" blockId="15.[151,1436,151,1458]" pageId="15" pageNumber="16">
|
||
<keyLead id="30A7AC3EFFFC735F32FAFC28E69CFBCB" pageId="15" pageNumber="16">
|
||
<emphasis id="B922D319FFFC735F32FAFC28E3B5FC73" bold="true" box="[151,178,914,938]" pageId="15" pageNumber="16">12</emphasis>
|
||
. Anal shield glabrous, golden (Fig. 17C). Tergites with unique pitchblack/reddish-brown pattern (Fig. 17C). Anterior telopod third podomere without large, sclerotized teeth (Fig. 28E). First antennomere without
|
||
<emphasis id="B922D319FFFC735F36A3FC0FE69CFC15" box="[1230,1435,949,972]" italics="true" pageId="15" pageNumber="16">sensilla basiconica</emphasis>
|
||
(Fig. 30A). Sclerotized teeth present on antennomeres 1–4 (Fig. 30A). Endotergum marginal bristles not extending beyond tergite margin (Fig. 35C).
|
||
<emphasis id="B922D319FFFC735F3042FC41E65EFBCB" box="[559,1369,1019,1042]" italics="true" pageId="15" pageNumber="16">
|
||
.........................................................................................................
|
||
<taxonomicName id="4C567488FFFC735F3682FC41E65EFBCB" ID-CoL="5D7GJ" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1263,1369,1019,1042]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="aureum" status="sp. nov.">Z. aureum</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="B922D319FFFC735F370DFC40E69CFBCB" bold="true" box="[1376,1435,1018,1042]" pageId="15" pageNumber="16">
|
||
<taxonomicNameLabel id="A2116E62FFFC735F370DFC40E69CFBCB" box="[1376,1435,1018,1042]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFFC735F32FAFBA7E691FB45" blockId="15.[151,1436,151,1458]" pageId="15" pageNumber="16">
|
||
<keyLead id="30A7AC3EFFFC735F32FAFBA7E691FB45" pageId="15" pageNumber="16">
|
||
- Anal shield covered with few, minute hair, of dirty brownish color. Tergites with light brown, blackish coloration pattern. Anterior telopod third podomere laterally with at least single large, sclerotized tooth (Fig. 32B). First antennomere with
|
||
<emphasis id="B922D319FFFC735F33F4FBD9E16DFBA3" box="[409,618,1123,1146]" italics="true" pageId="15" pageNumber="16">sensilla basiconica</emphasis>
|
||
(Fig. 33A). Sclerotized teeth present on antennomeres 1–5 (Fig. 33A). Endotergum marginal bristles slightly extending beyond tergite margin (Fig. 35D). ..............................................
|
||
<emphasis id="B922D319FFFC735F3716FB3EE691FB45" bold="true" box="[1403,1430,1156,1180]" pageId="15" pageNumber="16">13</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A217AEFFFC735F32FAFB1DE69CFA6B" pageId="15" pageNumber="16">
|
||
<paragraph id="8BE90F0BFFFC735F32FAFB1DE69BFAFE" blockId="15.[151,1436,151,1458]" pageId="15" pageNumber="16">
|
||
<keyLead id="30A7AC3EFFFC735F32FAFB1DE69BFAFE" pageId="15" pageNumber="16">
|
||
<emphasis id="B922D319FFFC735F32FAFB1DE3B5FB66" bold="true" box="[151,178,1191,1215]" pageId="15" pageNumber="16">13</emphasis>
|
||
. Legs short, femur 1.45 times longer than wide, tarsus 3.2 times longer than wide. Third pair of legs without apical spine. Third podomere of posterior telopods with
|
||
<emphasis id="B922D319FFFC735F3086FB71E026FB3B" box="[747,801,1227,1250]" italics="true" pageId="15" pageNumber="16">circa</emphasis>
|
||
18 large, sclerotized teeth. Endotergum with two rows of marginal bristles. Posterior locking carina four time longer than anterior carina. Antenna with 25 or less apical cones.
|
||
<emphasis id="B922D319FFFC735F337CFAAAE0D1FAFE" box="[273,982,1296,1319]" italics="true" pageId="15" pageNumber="16">..........................................................................................................</emphasis>
|
||
<taxonomicName id="4C567488FFFC735F31B8FAAAE69BFAFE" ID-CoL="5D7HP" authority="(de Saussure & Zehntner, 1897)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[981,1436,1296,1319]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="priapus">
|
||
<emphasis id="B922D319FFFC735F31B8FAAAE746FAFE" box="[981,1089,1296,1319]" italics="true" pageId="15" pageNumber="16">Z. priapus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC772FAFFFC735F3622FAAAE693FAFE" author="de Saussure, H. & Zehntner, L." box="[1103,1428,1296,1319]" pageId="15" pageNumber="16" pagination="1 - 12" refId="ref60128" refString="de Saussure, H. & Zehntner, L. (1897) Atlas de l'histoire naturelle des Myriapodes, In: Grandidier (Ed.) Histoire physique, naturelle et politique de Madagascar. 27 (53), Alfred Grandidier, Paris, France, plates 1 - 12." type="book chapter" year="1897">de Saussure & Zehntner, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFFC735F32FAFA88E69CFA6B" blockId="15.[151,1436,151,1458]" pageId="15" pageNumber="16">
|
||
<keyLead id="30A7AC3EFFFC735F32FAFA88E69CFA6B" pageId="15" pageNumber="16">
|
||
- Legs of usual size, femur 1.9 times longer than wide, tarsus 4.2 times longer than wide. Third pair of legs with apical spine. Third podomere of posterior telopods with
|
||
<emphasis id="B922D319FFFC735F30BAFAEFE00AFAB5" box="[727,781,1365,1388]" italics="true" pageId="15" pageNumber="16">circa</emphasis>
|
||
22 large, sclerotized teeth (Fig. 32D). Endotergum with three rows of marginal bristles (Fig. 35D). Locking carinae of equal size. Antenna with 35–86 apical cones (Fig. 33B). ... ..................................................................................................................................................
|
||
<taxonomicName id="4C567488FFFC735F36CDFA20E65EFA68" ID-CoL="5D7HT" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1184,1369,1434,1457]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pseudopriapus" status="sp. nov.">
|
||
<emphasis id="B922D319FFFC735F36CDFA20E65EFA68" box="[1184,1369,1434,1457]" italics="true" pageId="15" pageNumber="16">Z. pseudopriapus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B922D319FFFC735F370DFA20E69CFA6B" bold="true" box="[1376,1435,1434,1458]" pageId="15" pageNumber="16">
|
||
<taxonomicNameLabel id="A2116E62FFFC735F370DFA20E69CFA6B" box="[1376,1435,1434,1458]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |