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<document id="C316DE58C51CE6399FB9758F35290E62" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFFA0730532FAFF2DE7E6F8DE" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Zoosphaerium ambrense Wesener 2009, new species" docType="treatment" docUuid="890A366B-7D99-4878-B327-25174400B9C8" docUuidSource="ZooBank" docVersion="5" lastPageNumber="86" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="84" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="9E273F86EE75A6D85419B7DAE19899A0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B25A795BBDD9A3F9E1B8AB1CE43E5A5F">
<mods:title id="782BDC85817B2146E20899CC505693A0">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
</mods:titleInfo>
<mods:name id="31D092BCE016A91B26F591069C839416" type="personal">
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</mods:role>
<mods:namePart id="C0AE37009DD7C4B1A03DD24B41D1B1F7">Wesener, Thomas</mods:namePart>
</mods:name>
<mods:typeOfResource id="08CA38E16F8D89C7D9522394D626BC1E">text</mods:typeOfResource>
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<mods:title id="12D9D497BB939AA54B440652A1D32631">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4C825D505F65BE0C26363EDDC1417903">
<mods:date id="9785A03C5EAA6D03E16111BDF300E880">2009</mods:date>
<mods:detail id="9C5A290C254F0D136A57D742EDFBEC49" type="pubDate">
<mods:number id="31B9D5C919CE3E2256AACB921841CDD9">2009-05-11</mods:number>
</mods:detail>
<mods:detail id="9026F911719A75E0A3AE9F13E65CE36E" type="volume">
<mods:number id="9C0D58E65D51CCEC4025CFF0C5A40175">2097</mods:number>
</mods:detail>
<mods:detail id="65DD73D0E052F248ADE9D0BB44AE8E1E" type="issue">
<mods:number id="A7082C5EC08979F4E518DA319A5EA719">1</mods:number>
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<mods:location id="953A05A70ABE71C770F999293C8438F0">
<mods:url id="440B2F38DE8F15C9902BDEBA1059A7EA">https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1</mods:url>
</mods:location>
<mods:classification id="AC8D84CF836F1A4D159231F3D34E2DFC">journal article</mods:classification>
<mods:identifier id="102B03FD7BA3FD588D804BFE56AF60E3" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
<mods:identifier id="56827DC5457FBC39122583AC68E3FC64" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="03A8EABF25DBC0756BA8D31DE7DD4E08" type="Zenodo-Dep">5316536</mods:identifier>
<mods:identifier id="3DF06D5E180EE5C58A96F9BE4F9EDC25" type="ZooBank">0E18167A-78EB-4079-8BF4-972A4AC88C69</mods:identifier>
</mods:mods>
<treatment id="03FFBE1DFFA0730532FAFF2DE7E6F8DE" ID-DOI="http://doi.org/10.5281/zenodo.5325503" ID-Zenodo-Dep="5325503" LSID="urn:lsid:zoobank.org:act:890A366B-7D99-4878-B327-25174400B9C8" httpUri="http://treatment.plazi.org/id/03FFBE1DFFA0730532FAFF2DE7E6F8DE" lastPageId="85" lastPageNumber="86" pageId="83" pageNumber="84">
<subSubSection id="C34C5C80FFA0730332FAFF2DE166FF68" box="[151,609,151,177]" pageId="83" pageNumber="84" type="nomenclature">
<paragraph id="8BE90F0BFFA0730332FAFF2DE166FF68" blockId="83.[151,609,151,177]" box="[151,609,151,177]" pageId="83" pageNumber="84">
<heading id="D0A1B867FFA0730332FAFF2DE166FF68" bold="true" box="[151,609,151,177]" fontSize="11" level="1" pageId="83" pageNumber="84" reason="1">
<emphasis id="B922D319FFA0730332FAFF2DE166FF68" bold="true" box="[151,609,151,177]" pageId="83" pageNumber="84">
<emphasis id="B922D319FFA0730332FAFF2DE2CDFF68" bold="true" box="[151,458,151,177]" italics="true" pageId="83" pageNumber="84">
<taxonomicName id="4C567488FFA0730332FAFF2DE2C3FF68" ID-CoL="5D7GC" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,452,151,177]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="ambrense" status="sp. nov.">Zoosphaerium ambrense</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A2116E62FFA0730333BFFF2DE166FF68" box="[466,609,151,177]" pageId="83" pageNumber="84" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFA0730332FAFF5AE116FEFB" pageId="83" pageNumber="84" type="etymology">
<paragraph id="8BE90F0BFFA0730332FAFF5AE71BFF22" blockId="83.[151,1436,224,908]" box="[151,1052,224,251]" pageId="83" pageNumber="84">
<emphasis id="B922D319FFA0730332FAFF5AE287FF23" bold="true" box="[151,384,224,250]" pageId="83" pageNumber="84">Derivatio nominis:</emphasis>
<taxonomicName id="4C567488FFA0730333E5FF5BE2FEFF23" authorityName="Wesener" authorityYear="2009" box="[392,505,225,250]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="ambrense">
<emphasis id="B922D319FFA0730333E5FF5BE2FEFF23" box="[392,505,225,250]" italics="true" pageId="83" pageNumber="84">ambrense</emphasis>
</taxonomicName>
refers to the
<typeStatus id="54EDB1A9FFA0730330FEFF5BE1C2FF22" box="[659,709,225,251]" pageId="83" pageNumber="84">type</typeStatus>
locality, Montagne d'Ambre.
</paragraph>
<paragraph id="8BE90F0BFFA0730332ABFEBDE116FEFB" blockId="83.[151,1436,224,908]" box="[198,529,263,290]" pageId="83" pageNumber="84">
<emphasis id="B922D319FFA0730332ABFEBDE298FEF8" bold="true" box="[198,415,263,289]" pageId="83" pageNumber="84">Studied material:</emphasis>
2 M,
<specimenCount id="9D50C482FFA07303338BFEB2E116FEFB" box="[486,529,264,290]" pageId="83" pageNumber="84" type="generic">2 F.</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFA0730332ABFE94E0C1FE65" pageId="83" pageNumber="84" type="materials_examined">
<paragraph id="8BE90F0BFFA0730332ABFE94E293FE4F" blockId="83.[151,1436,224,908]" pageId="83" pageNumber="84">
<materialsCitation id="3B3E0556FFA0730332ABFE94E293FE4F" collectingDate="1989-11-24" collectorName="C. Lienhard" country="Madagascar" county="Geneve Mad" elevation="1050" location="Province" municipality="Locus" pageId="83" pageNumber="84" specimenCount="1" stateProvince="Antsiranana" typeStatus="holotype">
<emphasis id="B922D319FFA0730332ABFE94E23FFE91" bold="true" box="[198,312,302,328]" pageId="83" pageNumber="84">
<typeStatus id="54EDB1A9FFA0730332ABFE94E23FFE91" box="[198,312,302,328]" pageId="83" pageNumber="84" type="holotype">Holotype</typeStatus>
</emphasis>
: 1 M (
<quantity id="4CAEA2EEFFA0730333FDFE94E2E1FE91" box="[400,486,302,328]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="83" pageNumber="84" unit="mm" value="29.0">29 mm</quantity>
long),
<emphasis id="B922D319FFA07303302CFE94E028FE91" bold="true" box="[577,815,302,328]" pageId="83" pageNumber="84">
<collectingCounty id="62887787FFA07303302CFE94E1E4FE91" box="[577,739,302,328]" pageId="83" pageNumber="84">Geneve Mad</collectingCounty>
89/29
</emphasis>
.
<emphasis id="B922D319FFA07303312CFE95E0E3FE91" box="[833,996,303,328]" italics="true" pageId="83" pageNumber="84">
<collectingMunicipality id="6B8D9571FFA07303312CFE95E080FE91" box="[833,903,303,328]" pageId="83" pageNumber="84">Locus</collectingMunicipality>
typicus
</emphasis>
:
<collectingCountry id="F3414F9BFFA07303319BFE94E781FE91" box="[1014,1158,302,328]" name="Madagascar" pageId="83" pageNumber="84">Madagascar</collectingCountry>
,
<location id="8E8959D0FFA0730336F9FE94E7FBFE91" LSID="urn:lsid:plazi:treatment:03FFBE1DFFA0730532FAFF2DE7E6F8DE:8E8959D0FFA0730336F9FE94E7FBFE91" box="[1172,1276,302,328]" country="Madagascar" county="Geneve Mad" municipality="Locus" name="Province" pageId="83" pageNumber="84" stateProvince="Antsiranana">Province</location>
<collectingRegion id="4992C1E9FFA07303376AFE94E690FE91" box="[1287,1431,302,328]" country="Madagascar" name="Antsiranana" pageId="83" pageNumber="84">Antsiranana</collectingRegion>
,
<location id="8E8959D0FFA0730332FAFEEFE133FEB6" LSID="urn:lsid:plazi:treatment:03FFBE1DFFA0730532FAFF2DE7E6F8DE:8E8959D0FFA0730332FAFEEFE133FEB6" box="[151,564,341,367]" country="Madagascar" county="Geneve Mad" municipality="Locus" name="Parc National Montagne d'Ambre" pageId="83" pageNumber="84" stateProvince="Antsiranana">Parc National Montagne d'Ambre</location>
(=Ambohitra), vers le petit lac, foret primaire,
<quantity id="4CAEA2EEFFA0730336FDFEEFE631FEB6" box="[1168,1334,341,367]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.09" metricValueMin="1.01" pageId="83" pageNumber="84" unit="m" value="1050.0" valueMax="1090.0" valueMin="1010.0">
<elevation id="007BE838FFA0730336FDFEEFE631FEB6" box="[1168,1334,341,367]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.09" metricValueMin="1.01" pageId="83" pageNumber="84" unit="m" value="1050.0" valueMax="1090.0" valueMin="1010.0">1010-1090 m</elevation>
</quantity>
, leg.
<collectorName id="26A36ADDFFA0730337ECFEEFE203FE4F" pageId="83" pageNumber="84">C. Lienhard</collectorName>
,
<date id="FFE829CBFFA073033362FEC6E297FE4F" box="[271,400,380,406]" pageId="83" pageNumber="84" value="1989-11-24">
<collectingDate id="EFACD023FFA073033362FEC6E297FE4F" box="[271,400,380,406]" pageId="83" pageNumber="84" value="1989-11-24">24.xi.1989</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE90F0BFFA0730332ABFE18E0C1FE65" blockId="83.[151,1436,224,908]" box="[198,966,418,444]" pageId="83" pageNumber="84">
<materialsCitation id="3B3E0556FFA0730332ABFE18E0C5FE65" box="[198,962,418,444]" collectingDate="1989-11-24" collectorName="C. Lienhard" country="Madagascar" county="Geneve Mad" elevation="1050" location="Mad" municipality="Locus" pageId="83" pageNumber="84" specimenCount="1" stateProvince="Geneve" typeStatus="paratype">
<emphasis id="B922D319FFA0730332ABFE18E244FE65" bold="true" box="[198,323,418,444]" pageId="83" pageNumber="84">
<typeStatus id="54EDB1A9FFA0730332ABFE18E244FE65" box="[198,323,418,444]" pageId="83" pageNumber="84" type="paratype">Paratypes</typeStatus>
</emphasis>
: 1 M, 2 F,
<emphasis id="B922D319FFA0730333A9FE18E1AAFE65" bold="true" box="[452,685,418,444]" pageId="83" pageNumber="84">
<collectingRegion id="4992C1E9FFA0730333A9FE18E126FE65" box="[452,545,418,444]" country="Switzerland" name="Geneve" pageId="83" pageNumber="84">Geneve</collectingRegion>
<location id="8E8959D0FFA073033045FE18E164FE65" LSID="urn:lsid:plazi:treatment:03FFBE1DFFA0730532FAFF2DE7E6F8DE:8E8959D0FFA073033045FE18E164FE65" box="[552,611,418,444]" name="Mad" pageId="83" pageNumber="84" stateProvince="Geneve">Mad</location>
89/29
</emphasis>
: same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFA0730332ABFE72E75AFE3A" box="[198,1117,456,483]" pageId="83" pageNumber="84" type="distribution">
<paragraph id="8BE90F0BFFA0730332ABFE72E75AFE3A" blockId="83.[151,1436,224,908]" box="[198,1117,456,483]" pageId="83" pageNumber="84">
<emphasis id="B922D319FFA0730332ABFE72E26EFE3B" bold="true" box="[198,361,456,482]" pageId="83" pageNumber="84">Distribution:</emphasis>
Known only from the rainforest of Montagne d'Ambre (Fig. 50)
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFA0730432ABFE55E3E6FF01" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84" type="diagnosis">
<paragraph id="8BE90F0BFFA0730332ABFE55E608FC55" blockId="83.[151,1436,224,908]" pageId="83" pageNumber="84">
<emphasis id="B922D319FFA0730332ABFE55E24FFDD0" bold="true" box="[198,328,495,521]" pageId="83" pageNumber="84">Diagnosis:</emphasis>
Medium sized, known specimens up to
<quantity id="4CAEA2EEFFA07303314FFE4AE074FDD3" box="[802,883,496,522]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="83" pageNumber="84" unit="mm" value="34.0">34 mm</quantity>
long. Color (alternated by alcohol) completely light brown, tergites and antennae with traces of dark olive green. Collum and posterior margins of tergites with some short hair. First antennomere with groove, antennomeres 1–5 with sclerotized teeth (Fig. 57B), disc with four apical cones (Fig. 57A). First stigmatic plate with small, triangular lobe. Movable finger of posterior telopods weakly curved, length three times greater than width (Fig. 56D), with large, triangular lobe, and four thin spines (Fig. 56C), posterior side with 15–17 small sclerotized teeth (Fig. 56D). Fixed finger shorter than movable finger, reaching up to 3/4 of its length. Third podomere of anterior telopods on posterior side laterally with 1 or 2 small, isolated sclerotized teeth (Fig. 56B). One short stridulation rib on male harp (Fig. 56A) and three on each side of female washboard. Two locking carinae on anal shield, anterior one small, posterior carina three times larger than first. Operculum of vulva reaching up to apical coxal margin, medially notched, elongated into two small, well-rounded lobes. Lateral lobe slightly larger than inner one.
</paragraph>
<paragraph id="8BE90F0BFFA0730332FAF973E782F890" blockId="83.[151,1436,1737,1865]" pageId="83" pageNumber="84">
<emphasis id="B922D319FFA0730332FAF973E228F938" bold="true" box="[151,303,1737,1761]" pageId="83" pageNumber="84">FIGURE 056</emphasis>
.
<taxonomicName id="4C567488FFA073033352F970E14FF938" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[319,584,1738,1761]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="ambrense" status="sp. nov.">
<emphasis id="B922D319FFA073033352F970E14FF938" box="[319,584,1738,1761]" italics="true" pageId="83" pageNumber="84">Zoosphaerium ambrense</emphasis>
</taxonomicName>
<emphasis id="B922D319FFA07303303FF973E197F938" bold="true" box="[594,656,1737,1761]" pageId="83" pageNumber="84">
<taxonomicNameLabel id="A2116E62FFA07303303FF973E197F938" box="[594,656,1737,1761]" pageId="83" pageNumber="84" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FFA0730330CDF970E007F938" box="[672,768,1738,1761]" pageId="83" pageNumber="84" type="holotype">holotype</typeStatus>
,
<emphasis id="B922D319FFA073033160F973E027F938" bold="true" box="[781,800,1737,1761]" pageId="83" pageNumber="84">A</emphasis>
: right half of anterior telopods, anterior view;
<emphasis id="B922D319FFA073033749F973E631F938" bold="true" box="[1316,1334,1737,1761]" pageId="83" pageNumber="84">B</emphasis>
: last two podomeres of anterior telopod, lateral view;
<emphasis id="B922D319FFA07303301BF956E18EF8DD" bold="true" box="[630,649,1772,1796]" pageId="83" pageNumber="84">C</emphasis>
: posterior telopod, anterior view;
<emphasis id="B922D319FFA073033194F956E70BF8DD" bold="true" box="[1017,1036,1772,1796]" pageId="83" pageNumber="84">D</emphasis>
: chela of posterior telopod, posterior view. Abbreviation:
<emphasis id="B922D319FFA07303331DF8B5E288F8FE" bold="true" box="[368,399,1807,1831]" pageId="83" pageNumber="84">IH</emphasis>
= position of inner horns. Abbreviations:
<emphasis id="B922D319FFA073033120F8B5E074F8FE" bold="true" box="[845,883,1807,1831]" pageId="83" pageNumber="84">cav</emphasis>
= cavity;
<emphasis id="B922D319FFA0730331B3F8B5E700F8FE" bold="true" box="[990,1031,1807,1831]" pageId="83" pageNumber="84">cr-t</emphasis>
= crenulated teeth;
<emphasis id="B922D319FFA0730336B4F8B5E7F0F8FE" bold="true" box="[1241,1271,1807,1831]" pageId="83" pageNumber="84">ml</emphasis>
= membranous lobe;
<emphasis id="B922D319FFA0730332BEF88BE3F2F890" bold="true" box="[211,245,1841,1865]" pageId="83" pageNumber="84">s-p</emphasis>
= sclerotized spot;
<emphasis id="B922D319FFA0730333ABF88BE2F2F890" bold="true" box="[454,501,1841,1865]" pageId="83" pageNumber="84">scl-t</emphasis>
= sclerotized teeth;
<emphasis id="B922D319FFA0730330A0F88BE1E8F890" bold="true" box="[717,751,1841,1865]" pageId="83" pageNumber="84">SR</emphasis>
= stridulation rib. Scale bars =
<quantity id="4CAEA2EEFFA07303362DF888E786F890" box="[1088,1153,1842,1865]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="83" pageNumber="84" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="8BE90F0BFFA0730432ABF8CDE3E6FF01" blockId="83.[151,1436,1911,2014]" lastBlockId="84.[151,1436,151,371]" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84">
<emphasis id="B922D319FFA0730332ABF8CDE297F848" bold="true" box="[198,400,1911,1937]" pageId="83" pageNumber="84">Similar species:</emphasis>
<emphasis id="B922D319FFA0730333F3F8CDE1CBF849" box="[414,716,1911,1936]" italics="true" pageId="83" pageNumber="84">
<taxonomicName id="4C567488FFA0730333F3F8CDE1CFF849" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[414,712,1911,1936]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="ambrense" status="sp. nov.">Zoosphaerium ambrense</taxonomicName>
,
</emphasis>
<emphasis id="B922D319FFA0730330B7F8CDE025F848" bold="true" box="[730,802,1911,1937]" pageId="83" pageNumber="84">
<taxonomicNameLabel id="A2116E62FFA0730330B7F8CDE025F848" box="[730,802,1911,1937]" pageId="83" pageNumber="84" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
anterior and posterior telopods are of a shape not displayed among other known giant-pill millipede species (Figs 56A–D). The triangular lobe on the third podomere of the posterior telopods is quite similar to those of
<taxonomicName id="4C567488FFA0730431C7F87FE252FF6B" authority="(de Saussure &amp; Zehntner, 1897)" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" lastPageId="84" lastPageNumber="85" order="Sphaerotheriida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="piligerum">
<emphasis id="B922D319FFA0730331C7F87FE7D5F807" box="[938,1234,1989,2014]" italics="true" pageId="83" pageNumber="84">Zoosphaerium piligerum</emphasis>
(
<bibRefCitation id="EFC772FAFFA07304368AF87EE24BFF6B" author="de Saussure, H. &amp; Zehntner, L." lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84" pagination="1 - 12" refId="ref60128" refString="de Saussure, H. &amp; Zehntner, L. (1897) Atlas de l'histoire naturelle des Myriapodes, In: Grandidier (Ed.) Histoire physique, naturelle et politique de Madagascar. 27 (53), Alfred Grandidier, Paris, France, plates 1 - 12." type="book chapter" year="1897">de Saussure &amp; Zehntner, 1897</bibRefCitation>
)
</taxonomicName>
, but
<emphasis id="B922D319FFA7730433E0FF22E123FF68" box="[397,548,152,177]" italics="true" pageId="84" pageNumber="85">
<taxonomicName id="4C567488FFA7730433E0FF22E127FF68" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[397,544,152,177]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="ambrense" status="sp. nov.">Z. ambrense</taxonomicName>
,
</emphasis>
<emphasis id="B922D319FFA773043041FF2DE169FF68" bold="true" box="[556,622,151,177]" pageId="84" pageNumber="85">
<taxonomicNameLabel id="A2116E62FFA773043041FF2DE169FF68" box="[556,622,151,177]" pageId="84" pageNumber="85" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from the latter in, among other characters, the general telopod shape.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFA7730532ABFF5FE7E6F8DE" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85" type="description">
<paragraph id="8BE90F0BFFA7730432ABFF5FE28BFEAA" blockId="84.[151,1436,151,371]" pageId="84" pageNumber="85">
<emphasis id="B922D319FFA7730432ABFF5FE267FF26" bold="true" box="[198,352,229,255]" pageId="84" pageNumber="85">Description:</emphasis>
<emphasis id="B922D319FFA773043304FF5FE2F1FF27" box="[361,502,229,254]" italics="true" pageId="84" pageNumber="85">Body length</emphasis>
: Males (
<specimenCount id="9D50C482FFA77304300DFF5FE1F0FF26" box="[608,759,229,255]" pageId="84" pageNumber="85" type="generic">2 specimens</specimenCount>
): length up to
<quantity id="4CAEA2EEFFA7730431CCFF5FE0FFFF26" box="[929,1016,229,255]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="84" pageNumber="85" unit="mm" value="29.0">29 mm</quantity>
, width of thoracic shield up to
<quantity id="4CAEA2EEFFA773043704FF5FE3CFFEFF" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="84" pageNumber="85" unit="mm" value="14.1">14.1 mm</quantity>
, height of thoracic shield up to
<quantity id="4CAEA2EEFFA773043025FEB6E1A0FEFF" box="[584,679,268,294]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="84" pageNumber="85" unit="mm" value="7.8">7.8 mm</quantity>
. Females (
<specimenCount id="9D50C482FFA773043145FEB6E0C6FEFF" box="[808,961,268,294]" pageId="84" pageNumber="85" type="generic">2 specimens</specimenCount>
): length up to
<quantity id="4CAEA2EEFFA77304361DFEB6E7C0FEFF" box="[1136,1223,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="84" pageNumber="85" unit="mm" value="34.0">34 mm</quantity>
, width up to
<quantity id="4CAEA2EEFFA773043704FEB6E3CFFE95" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="84" pageNumber="85" unit="mm" value="16.8">16.8 mm</quantity>
, height up to
<quantity id="4CAEA2EEFFA77304330AFE88E2C3FE95" box="[359,452,306,332]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="84" pageNumber="85" unit="mm" value="9.4">9.4 mm</quantity>
. Too few specimens are known to evaluate if mature males are significantly smaller than mature females.
</paragraph>
<paragraph id="8BE90F0BFFA7730432FAF88FE297F84B" blockId="84.[151,1436,1845,1939]" pageId="84" pageNumber="85">
<emphasis id="B922D319FFA7730432FAF88FE22AF894" bold="true" box="[151,301,1845,1869]" pageId="84" pageNumber="85">FIGURE 057</emphasis>
.
<taxonomicName id="4C567488FFA773043357F88CE146F894" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[314,577,1846,1869]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="ambrense" status="sp. nov.">
<emphasis id="B922D319FFA773043357F88CE146F894" box="[314,577,1846,1869]" italics="true" pageId="84" pageNumber="85">Zoosphaerium ambrense</emphasis>
</taxonomicName>
<emphasis id="B922D319FFA77304302AF88FE184F894" bold="true" box="[583,643,1845,1869]" pageId="84" pageNumber="85">
<taxonomicNameLabel id="A2116E62FFA77304302AF88FE184F894" box="[583,643,1845,1869]" pageId="84" pageNumber="85" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FFA7730430FDF88FE1E8F895" box="[656,751,1845,1868]" pageId="84" pageNumber="85" type="paratype">paratype</typeStatus>
, SEM,
<emphasis id="B922D319FFA773043150F88FE057F894" bold="true" box="[829,848,1845,1869]" pageId="84" pageNumber="85">A</emphasis>
: antenna, apical view on disc;
<emphasis id="B922D319FFA7730436F5F88FE7ADF894" bold="true" box="[1176,1194,1845,1869]" pageId="84" pageNumber="85">B</emphasis>
: antenna, lateral view;
<emphasis id="B922D319FFA7730432FAF8E2E3ADF8A9" bold="true" box="[151,170,1880,1904]" pageId="84" pageNumber="85">C</emphasis>
: right mandible, mesal view. Abbreviations:
<emphasis id="B922D319FFA7730430EAF8E2E1A9F8A9" bold="true" box="[647,686,1880,1904]" pageId="84" pageNumber="85">3iT</emphasis>
= 3 inner teeth;
<emphasis id="B922D319FFA773043137F8E2E07DF8A9" bold="true" box="[858,890,1880,1904]" pageId="84" pageNumber="85">Co</emphasis>
= condylus;
<emphasis id="B922D319FFA773043669F8E2E725F8A9" bold="true" box="[1028,1058,1880,1904]" pageId="84" pageNumber="85">eT</emphasis>
= external tooth;
<emphasis id="B922D319FFA7730436B1F8E2E606F8A9" bold="true" box="[1244,1281,1880,1904]" pageId="84" pageNumber="85">mp</emphasis>
= molar plate;
<emphasis id="B922D319FFA7730432FAF8C1E3BFF84A" bold="true" box="[151,184,1915,1939]" pageId="84" pageNumber="85">pL</emphasis>
= pectinate lamella.
</paragraph>
<paragraph id="8BE90F0BFFA6730532AAFF22E61DFF01" blockId="85.[151,1436,152,1799]" pageId="85" pageNumber="86">
<emphasis id="B922D319FFA6730532AAFF22E256FF68" box="[199,337,152,177]" italics="true" pageId="85" pageNumber="86">Coloration:</emphasis>
In most specimens faded by light and alcohol. In best preserved specimens, head, collum, thoracic shield, tergites, anal shield and legs light brownish, eyes and antenna with traces of green.
</paragraph>
<paragraph id="8BE90F0BFFA6730532FAFF5FE2E6FF26" blockId="85.[151,1436,152,1799]" box="[151,481,229,255]" pageId="85" pageNumber="86">
<emphasis id="B922D319FFA6730532FAFF5FE3E7FF27" box="[151,224,229,254]" italics="true" pageId="85" pageNumber="86">Head:</emphasis>
Eyes with&gt;60 ocelli.
</paragraph>
<paragraph id="8BE90F0BFFA6730532AAFEB0E03FFEAB" blockId="85.[151,1436,152,1799]" pageId="85" pageNumber="86">
<emphasis id="B922D319FFA6730532AAFEB0E238FEFA" box="[199,319,266,291]" italics="true" pageId="85" pageNumber="86">Antennae:</emphasis>
Length of antennomeres: 1=2=3&gt;4&lt;5&lt;6 (Fig. 57B), sixth antennomere bearing disc with four apical cones (Fig. 57A). First antennomere of greatest width, with groove but without
<emphasis id="B922D319FFA6730536DFFE8BE692FE93" box="[1202,1429,305,330]" italics="true" pageId="85" pageNumber="86">sensilla basiconica</emphasis>
. Sclerotized teeth at base of antennomeres 1–5 (Fig. 57B).
</paragraph>
<paragraph id="8BE90F0BFFA6730532AAFEC5E06DFE66" blockId="85.[151,1436,152,1799]" pageId="85" pageNumber="86">
<emphasis id="B922D319FFA6730532AAFEC5E247FE41" box="[199,320,383,408]" italics="true" pageId="85" pageNumber="86">Mandible:</emphasis>
Six rows of pectinate lamellae, number of teeth declining apically to proximally (Fig. 57C). Molar plate with deep groove (Fig. 57C). Condylus damaged.
</paragraph>
<paragraph id="8BE90F0BFFA6730532AAFE76E123FDD5" blockId="85.[151,1436,152,1799]" pageId="85" pageNumber="86">
<emphasis id="B922D319FFA6730532AAFE76E296FE3C" box="[199,401,460,485]" italics="true" pageId="85" pageNumber="86">Gnathochilarium</emphasis>
: Lateral of palpi
<emphasis id="B922D319FFA673053031FE76E190FE3C" box="[604,663,460,485]" italics="true" pageId="85" pageNumber="86">circa</emphasis>
five sensory cones, located together. Depression on posterior side of palpi with some sensory cones.
</paragraph>
<paragraph id="8BE90F0BFFA6730532FAFDA3E0E0FDEA" blockId="85.[151,1436,152,1799]" box="[151,999,537,563]" pageId="85" pageNumber="86">
<emphasis id="B922D319FFA6730532FAFDA3E3F0FDEB" box="[151,247,537,562]" italics="true" pageId="85" pageNumber="86">Collum:</emphasis>
Whole surface of collum covered with few short, isolated hairs.
</paragraph>
<paragraph id="8BE90F0BFFA6730532AAFD85E1C5FD81" blockId="85.[151,1436,152,1799]" box="[199,706,574,600]" pageId="85" pageNumber="86">
<emphasis id="B922D319FFA6730532AAFD85E279FD81" box="[199,382,575,600]" italics="true" pageId="85" pageNumber="86">Thoracic shield</emphasis>
: Texture similar to tergites.
</paragraph>
<paragraph id="8BE90F0BFFA6730532AAFDDFE12CFD7F" blockId="85.[151,1436,152,1799]" pageId="85" pageNumber="86">
<emphasis id="B922D319FFA6730532AAFDDFE224FDA7" box="[199,291,613,638]" italics="true" pageId="85" pageNumber="86">Tergites</emphasis>
: Shiny, at high magnification (90x) orange-peel like texture structure becomes visible. Paratergites projecting posteriorly.
</paragraph>
<paragraph id="8BE90F0BFFA6730532AAFD08E191FCC3" blockId="85.[151,1436,152,1799]" pageId="85" pageNumber="86">
<emphasis id="B922D319FFA6730532AAFD08E267FD12" box="[199,352,690,715]" italics="true" pageId="85" pageNumber="86">Endotergum:</emphasis>
Inner section with numerous short spines and long bristles. Between marginal ridge and inner area without cuticular impressions (Fig. 63D). Externally single row of marginal bristles. Bristles short, ending far before tergite margin (Fig. 63D).
</paragraph>
<paragraph id="8BE90F0BFFA6730532AAFC9DE691FC57" blockId="85.[151,1436,152,1799]" pageId="85" pageNumber="86">
<emphasis id="B922D319FFA6730532AAFC9DE25EFC99" box="[199,345,807,832]" italics="true" pageId="85" pageNumber="86">Anal shield:</emphasis>
Sexually dimorphic, in females well-rounded, neither bell-shaped nor tapered. In females glabrous. Ventral side of anal shield with two black locking carinae, located at ventral margin. Anterior carina short, running in 90 degrees angle upwards to anal shield margin. Posterior carina three times larger than first.
</paragraph>
<paragraph id="8BE90F0BFFA6730532AAFC20E667FBDB" blockId="85.[151,1436,152,1799]" pageId="85" pageNumber="86">
<emphasis id="B922D319FFA6730532AAFC20E20FFC6A" box="[199,264,922,947]" italics="true" pageId="85" pageNumber="86">Legs:</emphasis>
Tarsi of first pair of legs with six to eight, second with
<emphasis id="B922D319FFA6730531C3FC20E0EEFC6A" box="[942,1001,922,947]" italics="true" pageId="85" pageNumber="86">circa</emphasis>
nine and third with
<emphasis id="B922D319FFA67305368DFC20E61CFC6A" box="[1248,1307,922,947]" italics="true" pageId="85" pageNumber="86">circa</emphasis>
11 ventral spines. First two leg pairs with only weakly curved claws and without apical spine. Tarsi of legs 4–21 with curved claws, 10–12 ventral spines and one apical spine. Femur 2.0 and tarsi 3.6 times longer than wide.
</paragraph>
<paragraph id="8BE90F0BFFA6730532AAFBB5E0C1FBF1" blockId="85.[151,1436,152,1799]" box="[199,966,1038,1064]" pageId="85" pageNumber="86">
<emphasis id="B922D319FFA6730532AAFBB5E28BFBF1" box="[199,396,1039,1064]" italics="true" pageId="85" pageNumber="86">Stigmatic plates:</emphasis>
First lobe short and triangular, with rounded tip.
</paragraph>
<paragraph id="8BE90F0BFFA6730532AAFB8FE27AFB33" blockId="85.[151,1436,152,1799]" pageId="85" pageNumber="86">
<emphasis id="B922D319FFA6730532AAFB8FE2FFFB97" box="[199,504,1077,1102]" italics="true" pageId="85" pageNumber="86">Female sexual characters</emphasis>
: Subanal plate with washboard, consisting of large, broad and well developed stridulation ribs, with 2 or 3 ribs on each half. Vulva large, covering more than 3/4 of coxal surface. Operculum large, reaching up to apical margin. Apical margin of operculum medially notched, forming two, well-rounded processes, lateral part slightly larger than inner one. Mesal plate long, reaching up to 2/3 of operculum's height.
</paragraph>
<paragraph id="8BE90F0BFFA6730532AAFB4DE16BFAEE" blockId="85.[151,1436,152,1799]" pageId="85" pageNumber="86">
<emphasis id="B922D319FFA6730532AAFB4DE2E7FAC9" box="[199,480,1271,1296]" italics="true" pageId="85" pageNumber="86">Male sexual characters</emphasis>
: Anal shield almost rectangular with steep slope, covered by dense field short hairs, especially around hindmost point.
</paragraph>
<paragraph id="8BE90F0BFFA6730532AAFAFEE68CF9C6" blockId="85.[151,1436,152,1799]" pageId="85" pageNumber="86">
<emphasis id="B922D319FFA6730532AAFAFEE290FA84" box="[199,407,1348,1373]" italics="true" pageId="85" pageNumber="86">Anterior telopods</emphasis>
(Figs 56A, B): Syncoxite glabrous, first podomere only with few isolated hairs. First podomere of rectangular shape, width 1.3 times larger than height, harp with one short rib. Second podomere process lobe-like and little curved with rounded edges, reaching up to 2/3 of third podomeres height, apically with sclerotized nubs. Third podomere cavity mesally in basal half with numerous small sclerotized spots, more apically with up to four thin, sclerotized spines; laterally with 1 or 2 minute sclerotized teeth. Third podomere apically with one round sclerotized spot. Podomeres 2 and 3 on both sides with isolated long hair.
</paragraph>
<paragraph id="8BE90F0BFFA6730532AAF996E7E6F8DE" blockId="85.[151,1436,152,1799]" pageId="85" pageNumber="86">
<emphasis id="B922D319FFA6730532AAF996E298F99C" box="[199,415,1580,1605]" italics="true" pageId="85" pageNumber="86">Posterior telopods</emphasis>
(Figs 56C, D): Third podomere elongated with well-rounded tip, length 3 times greater than width, only slightly tapering. Hollowed-out inner margin with one large, triangular lobe and four sclerotized spines, posterior aspect with 15–17 small, sclerotized teeth. Fixed finger straight, reaching only up to 3/4 of movable fingers length. Fixed finger not tapering, basally with single, short, sclerotized spine, apically covered with sclerotized spots. First podomere glabrous, second and third podomere covered on both sides with hair, especially at outer margins. Inner horns of syncoxite with pointed, slender tip.
</paragraph>
</subSubSection>
</treatment>
</document>
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