162 lines
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162 lines
20 KiB
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<document ID-CLB-Dataset="256667" ID-DOI="http://dx.doi.org/10.3897/phytokeys.225.99356" ID-GBIF-Dataset="9edaba47-012e-4c83-a606-8a760dc24de6" ID-PMC="PMC10209509" ID-Pensoft-Pub="1314-2003-225-115" ID-Pensoft-UUID="AB75E2E9609C5A7583F34B48EA61B976" ID-PubMed="37252326" ModsDocID="1314-2003-225-115" checkinTime="1682624511249" checkinUser="pensoft" docAuthor="Khamar, Hamid, Oualidi, Jalal El, Touhami, Amina Ouazzani & Civeyrel, Laure" docDate="2023" docId="03E9E636E25A5138948B09EAB972EE14" docLanguage="en" docName="PhytoKeys 225: 115-152" docOrigin="PhytoKeys 225" docPubDate="2023-04-27" docSource="http://dx.doi.org/10.3897/phytokeys.225.99356" docTitle="Verbascum faurei subsp. acanthifolium (Pau) Benedí & J. M. Monts., Lagascalia 20: 169. 1997." docType="treatment" docVersion="5" id="AB75E2E9609C5A7583F34B48EA61B976" lastPageNumber="115" masterDocId="AB75E2E9609C5A7583F34B48EA61B976" masterDocTitle="Nomenclature and typification in Verbascum (Scrophulariaceae) from North Africa" masterLastPageNumber="152" masterPageNumber="115" pageNumber="115" updateTime="1732936468263" updateUser="ExternalLinkService">
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<mods:titleInfo id="17587EB9A2E6DD45EDF47624D40FA069">
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<mods:title id="FFCA4537035569AB0266627661D40F1A">Nomenclature and typification in Verbascum (Scrophulariaceae) from North Africa</mods:title>
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<mods:namePart id="59148B8C307DE28D47A4C6B1E7DE2D75">Khamar, Hamid</mods:namePart>
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<mods:nameIdentifier id="B1D3E3C8E086F275AACA89E4BD633042" type="ORCID">https://orcid.org/0000-0001-6008-3264</mods:nameIdentifier>
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<mods:affiliation id="E6524CF0AF2B53D2F2C4DD598883F39C">Department of Botany and Plant Ecology, Scientific Institute, University Mohammed V in Rabat, B. P. 703, Rabat 10106, Morocco</mods:affiliation>
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<mods:nameIdentifier id="F78C6509ADF3C32DFFA8AE3587D39052" type="email">khamarhamid90@gmail.com</mods:nameIdentifier>
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<mods:namePart id="34252605E9974D5334B8DC1D088E0040">Oualidi, Jalal El</mods:namePart>
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<mods:affiliation id="7C399B1716DDA27F1C487AC75364D8E8">Department of Botany and Plant Ecology, Scientific Institute, University Mohammed V in Rabat, B. P. 703, Rabat 10106, Morocco</mods:affiliation>
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<mods:namePart id="42682822DEDA4619BE4899BCC30FC3F7">Touhami, Amina Ouazzani</mods:namePart>
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<mods:affiliation id="24856868E9871E47BB32810B3C1AF69F">Plant, Animal Productions and Agro-industry Laboratory, Department of Biology, Faculty of Sciences, Ibn Tofail University, B. P. 133 14000, Kenitra, Morocco</mods:affiliation>
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<mods:namePart id="2099F06A9B527FD12749B4F011D61F8F">Civeyrel, Laure</mods:namePart>
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<mods:affiliation id="FAF14676725BA398CB7553B3257760A1">Laboratoire Evolution & Diversite Biologique (EDB UMR 5174), Universite de Toulouse, CNRS, IRD, UPS. 118 route de Narbonne, Bat 4 R 1, 31062 Toulouse cedex 9, France</mods:affiliation>
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<mods:title id="40B2FF4A4A1CA00044346AAE256AA930">PhytoKeys</mods:title>
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<mods:date id="4663EAAF914E24FE6DE668BF7A98BDE6">2023</mods:date>
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<mods:identifier id="1E94901F988742414D87B0882634C74B" type="DOI">http://dx.doi.org/10.3897/phytokeys.225.99356</mods:identifier>
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<treatment id="03E9E636E25A5138948B09EAB972EE14" ID-GBIF-Taxon="209579280" LSID="urn:lsid:plazi:treatment:03E9E636E25A5138948B09EAB972EE14" httpUri="http://treatment.plazi.org/id/03E9E636E25A5138948B09EAB972EE14" lastPageNumber="115" pageId="0" pageNumber="115" scope_family="Scrophulariaceae">
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<subSubSection id="8D918356545876D5D7C1E5AA5FC71FAD" pageId="0" pageNumber="115" type="nomenclature">
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<paragraph id="C83C2DEDA2E5C6B45765ECEDF693E782" pageId="0" pageNumber="115">
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<taxonomicName id="0E0CA39266F34E1680F25D3882BF0FC9" LSID="03E9E636-E25A-5138-948B-09EAB972EE14" authority="(Pau) Benedi & J. M. Monts., Lagascalia 20: 169. 1997." authorityName="(Pau) Benedí & J. M. Monts., Lagascalia 20: 169. 1997." class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Verbascum faurei subsp. acanthifolium" order="Lamiales" pageId="0" pageNumber="115" rank="subSpecies" species="faurei" subSpecies="acanthifolium">
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Verbascum faurei subsp. acanthifolium (Pau)
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<normalizedToken id="D1B12A59AA97B56A839F4E5420DCBD32" originalValue="Benedí">Benedi</normalizedToken>
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& J.M.Monts., Lagascalia 20: 169. 1997.
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="7B2B82798AEFBD415193B9A05704B3FB" pageId="0" pageNumber="115" type="reference_group">
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<paragraph id="D8EBB5D535D7DD06271049DEAAE56E99" pageId="0" pageNumber="115">
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<taxonomicName id="E3F76A53C617D7CD282A3E741BC677FC" class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia acanthifolia" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="acanthifolia">Celsia acanthifolia</taxonomicName>
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≡
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<taxonomicName id="48E23590E555D091728740E7E444D01C" class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia acanthifolia" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="acanthifolia">Celsia acanthifolia</taxonomicName>
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Pau in Font Quer. Iter Maroc. n° 566. 1927. Type: [Morocco]. pr.
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<normalizedToken id="4E0E36F05010642F56FE3BD2BBAA8123" originalValue="Badú">Badu</normalizedToken>
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(Atlante rhiphaeo); Hab. in saxosis, solo schistose, 6 July 1927,
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<emphasis id="C4B2978FAB34577BC195B319EEB18167" italics="true" pageId="0" pageNumber="115">P. Font Quer, 566</emphasis>
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(lectotype, selected by
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<bibRefCitation id="00248E77EF140037A26BFC9629E9DE5F" author="Benedi, C" journalOrPublisher="Lagascalia" pageId="0" pageNumber="115" pagination="167 - 169" refId="B17" refString="Benedi, C, Montserrat, JM, 1997. Verbascum faurei (Murb.) Hub.-Mor. (Scrophulariaceae) y especies afines en el norte de Africa. Lagascalia 20 (1): 167 - 169" title="Verbascum faurei (Murb.) Hub. - Mor. (Scrophulariaceae) y especies afines en el norte de Africa." volume="20" year="1997">
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<normalizedToken id="CFB18DBD0C063529262E298650C8D89C" originalValue="Benedí">Benedi</normalizedToken>
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and Montserrat 1997
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</bibRefCitation>
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, pg. 169, first step
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<normalizedToken id="56E7DC6F0419EBE4AC3DC5097FD6B0F7" originalValue="“type”">"type"</normalizedToken>
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, as "
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<emphasis id="B4EF567F11E2BE8CA4FBA31376352F31" italics="true" pageId="0" pageNumber="115">Font Quer 566-1927</emphasis>
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"; second step, designated here: BC [BC43694]!; isolectotypes: LD [LD1222710]!, GDA [GDA39155]!, G [G00015117]!, MPU [MPU009627, MPU009628]!, BCN [BCN18029]!, BM [BM000930561]!). [image available at http://psimg.jstor.org/fsi/img/pdf/i0/10.5555/al.ap.specimen.bc43694_normal.pdf]
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</paragraph>
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<paragraph id="5543289BD2D1F4A27C82E551BF32CC93" pageId="0" pageNumber="115">
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<taxonomicName id="DFB39365A1EECDF5A09698B45F8D46B3" class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia acanthifolia" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="acanthifolia">Celsia acanthifolia</taxonomicName>
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≡
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<taxonomicName id="CB472D100203C023222AA8CA2793770C" class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia acanthifolia" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="acanthifolia">Celsia acanthifolia</taxonomicName>
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Pau, in Cavanillesia 1: 47. 1928, nom. nud.
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</paragraph>
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<paragraph id="5C73B4AFD723999D068CC3C3CFCD0727" pageId="0" pageNumber="115">
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<taxonomicName id="13ACA09DEDA80327969071D9C4B55D16" class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia faurei" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="faurei">Celsia faurei</taxonomicName>
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≡
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<taxonomicName id="F9D77693FC04E51F863F24EC197482C7" class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia faurei subsp. var. var. acanthifolia" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="variety" species="faurei" subSpecies="var." variety="acanthifolia">Celsia faurei var. acanthifolia</taxonomicName>
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(Pau) Maire in Jahand. & Maire. Cat. PI. Maroc.3: 669 (1934).
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</paragraph>
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</subSubSection>
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<subSubSection id="DD09C03FEA1D5020C30344B181F854FB" pageId="0" pageNumber="115" type="notes">
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<paragraph id="626D12A90562570A6A3AA455870ED872" pageId="0" pageNumber="115">Notes.</paragraph>
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<paragraph id="637DC61DC461BD2FB9BB34DD8727686C" pageId="0" pageNumber="115">
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<bibRefCitation id="40EA9D3655B2D0BFBA1420024E4EB7B7" author="Font Quer, P" journalOrPublisher="Feddes Repertorium" pageId="0" pageNumber="115" refId="B38" refString="Font Quer, P, 1927. Iter Maroccanum. Exsiccata schedula. Natural Science Museum, Barcelona, N° 566." title="Iter Maroccanum. Exsiccata schedula. Natural Science Museum, Barcelona, N ° 566." year="1927">Font Quer (1927)</bibRefCitation>
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based his species on a collection of plants from the Rif region that he has gathered during his 1927 campaign in Morocco. Font Quer gave the following diagnosis "Folia Acanthi mollis L., superiora oblonga, sinuato-dentata, floribus longe pedunculatis, pedicellis glanduliferis, racemosis; bractae lanceolatae brevissimae, calycis laciniae lanceolatae, margine integro, capsula globosa, obtusa calyce triplo longior" with the collection locality "hab. in saxosis, pr.
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<normalizedToken id="2368D8FABE4E29CD494A2584E948841A" originalValue="Badù">Badu</normalizedToken>
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(Atlante rhiphaeo), 1500 m. alt., solo schistoso, 6 julii". This protologue has been validly published by
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<bibRefCitation id="3AC87A943A5EA35539287BE2CA18EFD4" author="Gonzalez-Bueno, A" journalOrPublisher="Treballs de l'Institut Botanic de Barcelona" pageId="0" pageNumber="115" pagination="5 - 173" refId="B40" refString="Gonzalez-Bueno, A, 1988. Les campanyes botaniques de Pius Font i Quer al Nord d'Àfrica. Treballs de l'Institut Botanic de Barcelona 12: 5 - 173" title="Les campanyes botaniques de Pius Font i Quer al Nord d'Àfrica." volume="12" year="1988">
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<normalizedToken id="1B404AC651E059838E8ACF1415101FF3" originalValue="Gonzâlez-Bueno">Gonzalez-Bueno</normalizedToken>
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(1988)
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</bibRefCitation>
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. According to
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<bibRefCitation id="DCDB32FF13EB39B6BF263D593678D962" author="Nualart, N" journalOrPublisher="17), Oxford University Press, New York" pageId="0" pageNumber="115" publicationUrl="http://hdl.handle.net/2445/117964" refId="B84" refString="Nualart, N, 2017. Els herbaris, fonts per al coneixement de la flora. Aplicacions en conservacio i taxonomia. PhD Thesis, University of Barcelona, Spain. http://hdl.handle.net/2445/117964" title="Els herbaris, fonts per al coneixement de la flora. Aplicacions en conservacio i taxonomia. PhD Thesis, University of Barcelona, Spain." url="http://hdl.handle.net/2445/117964" year="2017">Nualart (2017)</bibRefCitation>
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,
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<bibRefCitation id="87455C5BB20C47437F12686152900790" author="Benedi, C" journalOrPublisher="Lagascalia" pageId="0" pageNumber="115" pagination="167 - 169" refId="B17" refString="Benedi, C, Montserrat, JM, 1997. Verbascum faurei (Murb.) Hub.-Mor. (Scrophulariaceae) y especies afines en el norte de Africa. Lagascalia 20 (1): 167 - 169" title="Verbascum faurei (Murb.) Hub. - Mor. (Scrophulariaceae) y especies afines en el norte de Africa." volume="20" year="1997">
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<normalizedToken id="DB6DAD56C257B4F86340A2FB6E240BD5" originalValue="Benedí">Benedi</normalizedToken>
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and Montserrat (1997)
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</bibRefCitation>
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wrongly cited the collection number of the lectotype as
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<normalizedToken id="091F134F5A1768F763659AD502A5B2EE" originalValue="“BC436934”">"BC436934"</normalizedToken>
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instead of
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<normalizedToken id="8B91D7AC3E972078EE787DF18286285A" originalValue="“BC43694”">"BC43694"</normalizedToken>
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. So, in addition to the specimen kept at Herb. BC (BC43694), seven other specimens are in agreement with the description in the protologue. They are housed at Herbs LD (LD1222710), GDA (GDA39155), G (G00015117), MPU (MPU009627, MPU009628), BCN (BCN18029), and BM (BM000930561). According to Art. 9.6 of the ICN (
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<bibRefCitation id="EC984B3B75B16B7E6A02B3FEFE6BB98C" DOI="https://doi.org/10.12705/Code.2018" author="Turland, NJ" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="115" refId="B108" refString="Turland, NJ, Wiersema, JH, Barrie, FR, Greuter, W, Hawksworth, DL, Herendeen, PS, Knapp, S, Kusber, WH, Li, DZ, Marhold, K, May, TW, McNeill, J, Monro, AM, Prado, J, Price, MJ, Smith, GF [Eds], 2018. International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashuetten. https://doi.org/10.12705/Code.2018" title="International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashuetten." url="https://doi.org/10.12705/Code.2018" year="2018">Turland et al. 2018</bibRefCitation>
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), all of these specimens and those persevered at Herb. BC must be treated as syntypes. The designation by
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<bibRefCitation id="BC7D123647968B628E7C4C6F0ED7BC50" author="Benedi, C" journalOrPublisher="Lagascalia" pageId="0" pageNumber="115" pagination="167 - 169" refId="B17" refString="Benedi, C, Montserrat, JM, 1997. Verbascum faurei (Murb.) Hub.-Mor. (Scrophulariaceae) y especies afines en el norte de Africa. Lagascalia 20 (1): 167 - 169" title="Verbascum faurei (Murb.) Hub. - Mor. (Scrophulariaceae) y especies afines en el norte de Africa." volume="20" year="1997">
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<normalizedToken id="1A9D0D534A579A7F56919326C1736625" originalValue="Benedí">Benedi</normalizedToken>
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and Montserrat (1997)
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</bibRefCitation>
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can be considered here as a first-step typification (see Art. 9.17 of the ICN,
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<bibRefCitation id="9888DF4B509EAB62F78B3B95D8C6D645" DOI="https://doi.org/10.12705/Code.2018" author="Turland, NJ" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="115" refId="B108" refString="Turland, NJ, Wiersema, JH, Barrie, FR, Greuter, W, Hawksworth, DL, Herendeen, PS, Knapp, S, Kusber, WH, Li, DZ, Marhold, K, May, TW, McNeill, J, Monro, AM, Prado, J, Price, MJ, Smith, GF [Eds], 2018. International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashuetten. https://doi.org/10.12705/Code.2018" title="International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashuetten." url="https://doi.org/10.12705/Code.2018" year="2018">Turland et al. 2018</bibRefCitation>
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). As the herbarium specimen BC43694 [mounted on three sheets]) kept at Herb. BC is the one showing the best quality of preservation of morphological features described in the protologue, we select it here as second-step lectotype of the name.
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</paragraph>
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<paragraph id="DEEB84FF5A90A244517344D5F1F22D91" pageId="0" pageNumber="115">
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The taxonomic status of
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<taxonomicName id="FA63BB781A74891315C38B5F4152834B" class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia acanthifolia" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="acanthifolia">
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<emphasis id="CD1156023FFD1842B420AEEAE8B9A90F" italics="true" pageId="0" pageNumber="115">Celsia acanthifolia</emphasis>
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</taxonomicName>
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has been assessed by some authors. Maire (in
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<bibRefCitation id="56468501FB628D53B4E046C9A3239597" author="Jahandiez, E" journalOrPublisher="Archives des Sciences" pageId="0" pageNumber="115" refId="B56" refString="Jahandiez, E, Maire, R, 1934. Catalogue des plantes du Maroc (Vol. 3). Imprimerie Minerva, Algiers, 663-669." title="Catalogue des plantes du Maroc (Vol. 3). Imprimerie Minerva, Algiers, 663 - 669." year="1934">Jahandiez and Maire 1934</bibRefCitation>
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) regarded it as a variety under
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<taxonomicName id="E752566838B6CFBA50EB8136A887DE66" genus="Ceslia" lsidName="Ceslia faurei" pageId="0" pageNumber="115" rank="species" species="faurei">
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<emphasis id="6F60BFCFBC396E1D7CF9DE9B7DB016F2" italics="true" pageId="0" pageNumber="115">Ceslia faurei</emphasis>
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</taxonomicName>
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Murb. [
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<taxonomicName id="C4C2A77B4966D8E2FD1667DCD7F55B23" class="Magnoliopsida" family="Scrophulariaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="C. faurei subsp. var. var. acanthifolia" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="variety" species="faurei" subSpecies="var." variety="acanthifolia">
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<emphasis id="748260247304B80ABEF52AE97A9A324F" italics="true" pageId="0" pageNumber="115">C. faurei var. acanthifolia</emphasis>
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</taxonomicName>
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(Pau) Maire]. Later, Benedi and Montserrat (1997) raised it to the rank of subspecies and they published it as a new combination,
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<taxonomicName id="C036D6DD99B2383A8B87B955FEE49F2C" class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" kingdom="Plantae" lsidName="Verbascum faurei subsp. acanthifolium" order="Lamiales" pageId="0" pageNumber="115" rank="subSpecies" species="faurei" subSpecies="acanthifolium">Verbascum faurei (Murb.) Hub.-Morath subsp. acanthifolium</taxonomicName>
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(Pau)
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<normalizedToken id="23B6728D284769BFBCAEA8A99664562E" originalValue="Benedí">Benedi</normalizedToken>
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& J.M.Monts.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |