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447 lines
107 KiB
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<document id="30600F6A9E43BB8FB2DC9E8DC5012E72" ID-DOI="10.1590/2358-2936e2023006" ID-ISSN="2358-2936" ID-Zenodo-Dep="10716764" ID-ZooBank="4970E3C2-A1DB-4AAE8961-FBAA26505ED4AB" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1709038466501" checkinUser="felipe" docAuthor="Rodríguez-Cabrera, Tomás M. & de Armas, Luis F." docDate="2023" docId="03E987EBFF04FFEFFC1DFCFCFC9FFCAA" docLanguage="en" docName="Nauplius.31.e2023006.pdf" docOrigin="Nauplius (e 2023006) 31" docSource="http://dx.doi.org/10.1590/2358-2936e2023006" docStyle="DocumentStyle:64D98C772C5FBF4D3A503C51F9604E62.1:Nauplius.2017-.journal_article" docStyleId="64D98C772C5FBF4D3A503C51F9604E62" docStyleName="Nauplius.2017-.journal_article" docStyleVersion="1" docTitle="Acanthoniscus Gosse 1851" docType="treatment" docVersion="3" lastPageNumber="6" masterDocId="FFD0FF93FF06FFEAFFA3FFD4FFBBFF89" masterDocTitle="Taxonomy of the enigmatic genus Acanthoniscus Gosse, 1851 (Isopoda: Oniscidea: Armadillidae), from Jamaica, with the description of a new species" masterLastPageNumber="25" masterPageNumber="1" pageNumber="3" updateTime="1712233860446" updateUser="jonas" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="862BDECA8E767352CC641C05658E7E14">Taxonomy of the enigmatic genus Acanthoniscus Gosse, 1851 (Isopoda: Oniscidea: Armadillidae), from Jamaica, with the description of a new species</mods:title>
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<mods:namePart id="ABD179D9F3E9D084B78381884C7D258E">Rodríguez-Cabrera, Tomás M.</mods:namePart>
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<mods:affiliation id="FDF96B5D807BB1C73B337AF25A73DE40">Instituto de Ecología y Sistemática. Boyeros, La Habana 11900, Cuba. E-mail: tomasmichel. rodriguez @ gmail. com & DB-</mods:affiliation>
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<mods:namePart id="8A14626230ABE5FFEF758A326687F8D9">de Armas, Luis F.</mods:namePart>
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<mods:affiliation id="F90E3BA818BEFBCF24DE9AD538B1842F">Sociedad Cubana de Zoología. San Antonio de los Baños, Artemisa 38100, Cuba. E-mail: luisdearmas 1945 @ gmail. com & - A</mods:affiliation>
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<mods:title id="6A70B2E55538A9DCA3AFAEE426DEC80F">Nauplius</mods:title>
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<mods:date id="9155C8CAE786EF2BD557165737BAA21E">2023</mods:date>
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<mods:title id="71F0471CAA7099DB6A6B68515E17E0CD">e 2023006</mods:title>
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Genus
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<taxonomicName id="4C404D7EFF04FFE8FBB4FCFCFAD6FCCF" ID-CoL="7NMLX" authority="Gosse, 1851" authorityName="Gosse" authorityYear="1851" box="[1047,1389,808,838]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B934EAEFFF04FFE8FBB4FCFCFB77FCCC" bold="true" box="[1047,1228,808,837]" italics="true" pageId="2" pageNumber="3">Acanthoniscus</emphasis>
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<bibRefCitation id="EFD14B0CFF04FFE8FB70FCFCFAD6FCCF" author="Gosse PH" box="[1235,1389,808,838]" pageId="2" pageNumber="3" refId="ref15567" refString="Gosse PH 1851. A naturalist's sojourn in Jamaica. London, Longman, Brown, Green and Longmans, 508 p. DOI: https: // www. doi. org / 10.5962 / bhl. title. 33264" type="book" year="1851">Gosse, 1851</bibRefCitation>
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<paragraph id="8BFF36FDFF04FFE8FBF7FC84FB6CFCE4" blockId="2.[958,1389,808,877]" box="[1108,1239,848,877]" pageId="2" pageNumber="3">
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(
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<figureCitation id="137B2A78FF04FFE8FBFCFC84FB75FCE4" box="[1119,1230,848,877]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStart-7="Figure 8" captionStartId-0="3.[181,252,1611,1634]" captionStartId-1="6.[181,252,1756,1779]" captionStartId-2="10.[843,913,586,609]" captionStartId-3="11.[181,252,1628,1651]" captionStartId-4="12.[182,251,1692,1715]" captionStartId-5="13.[181,252,1788,1811]" captionStartId-6="14.[181,252,1786,1809]" captionStartId-7="15.[181,252,1523,1546]" captionTargetBox-0="[181,1467,151,1587]" captionTargetBox-1="[206,1452,168,1723]" captionTargetBox-2="[843,1467,151,563]" captionTargetBox-3="[246,1407,152,1593]" captionTargetBox-4="[312,1351,473,1658]" captionTargetBox-5="[267,1379,821,1745]" captionTargetBox-6="[387,1266,935,1753]" captionTargetBox-7="[192,1460,173,1475]" captionTargetId-0="figure-19@3.[181,1467,151,1587]" captionTargetId-1="figure-175@6.[181,1466,148,1735]" captionTargetId-2="figure-639@10.[843,1467,148,565]" captionTargetId-3="figure-20@11.[229,1420,148,1608]" captionTargetId-4="figure-134@12.[266,1382,465,1671]" captionTargetId-5="figure-164@13.[232,1415,817,1764]" captionTargetId-6="figure-324@14.[356,1292,932,1765]" captionTargetId-7="figure-248@15.[181,1466,151,1499]" captionTargetPageId-0="3" captionTargetPageId-1="6" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="13" captionTargetPageId-6="14" captionTargetPageId-7="15" captionText-0="Figure 1. Distribution of the genus Acanthoniscus in Jamaica. Above: type locality of A. spiniger at the hills near Bluefields (yellow dot), the type locality of A.richardsonae sp.nov.in the Blue Mountains (red dot), and a second location in the Blue Mountains where a specimen presumably of A. richardsonae sp. nov. was photographed (orange dot). The main collection sites of Henry G. Hubbard in 1877 are depicted by green dots, notice the proximity of Stony Hill to the distribution of A. richardsonae sp.nov.Below: The hills known as Surinam Quarters near Bluefields, referred by Gosse (1851) as “Bluefields Mountain,” Westmoreland Parish, constitute the type locality of A. spiniger." captionText-1="Figure 2. Body parts of Acanthoniscus spiniger (female holotype BMNH 1973.478.1). A, Overall, dorsal view; B, overall, ventral view; C, right pereon-epimera 1–7 in ventral view; D, overall, lateral view; E, right pleon-epimera 3–5 in ventral view; F, overall, frontal view; G, cephalothorax in dorsal view; H, cephalothorax in frontal view; I, cephalothorax in ventral view; J, pleotelson and uropods in dorsal view; K, pleotelson and uropods inventral view; L, right uropodin ventral view; M, arrangement of dorsal spines, the diameters of the circles correspond to the relative size of the spines. Drawings of the corresponding structures modified from Kinahan (1859) are included in J and L. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–K), 0.5 mm (L). Original photographs by The Trustees of the BMNH." captionText-2="Figure 3. Right lateral view of a female paratype Acanthoniscus richardsonaesp.nov.showingcolorationinalcohol.Scale bar=1 mm." captionText-3="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." captionText-4="Figure 5.Otherappendagesof Acanthoniscus richardsonae sp.nov.A, Second antenna;B,detail offlagellum of secondantennadepicting position of aesthetasc groups (small arrows); C, detail of the distal article of the flagellum of the second antenna slightly turned over showing the aesthetasc groups; D, detail of the distal-most portion of the flagellum of the second antenna showing the apical cone; E, first antenna; F, detail of the third article of the first antenna showing the aesthetascs; G, male left seventh pereiopod; H, detail of ischium ofmale seventh pereiopod; I, detailof dactylus ofmale seventh pereiopodin anterior view; J, detailof dactylus ofmale seventh pereiopod in posterior view; K, male left first pereiopod, showing the areas on the carpus and propodus covered with the cleaning apparatus or antennal brush (dashed outlines); L, detail of carpus ofmale first pereiopod (anterior surface).Abbreviations:basis (bs), ischium(is),merus(me),carpus (ca),propodus (pr),flagellum(fl),and dactylus(da).Drawingofthe corresponding structuremodified from Richardson (1909) is included inA. Scale bars = 1 mm (A), 0.5 mm (G,K), 0.3mm (B), 0.1 mm (C–E, H–J,L),0.05 mm (F)." captionText-5="Figure 6.Mouthparts of Acanthoniscus richardsonae sp. nov. A, Distal portion of right (R) and left (L) mandibles in anterior views, showing lacinia mobilis (lm), hairy lobe (hl), penicils (pn), and tuft of setae (ts); B, first maxilla, with detailof distalportion of outer endite (ex) and inner endite (en); C, second maxilla; D, detail of the distal-most portion of the second maxilla; E, hypopharynx; F, detail of the median region of hypopharynx; G, maxilliped; H, detail of the distal-most portion of maxilliped in posterior view; I, detail of the distal-most portion of maxilliped inanterior view.Drawings of the corresponding structures modified from Richardson (1909) are included in B, C, and G. Scale bars = 0.25 mm." captionText-6="Figure 7.Pleopods of a male Acanthoniscus richardsonae sp.nov.A, First pleopod; B, detail of first pleopod endopodite tip; C, detail of first pleopod exopodite; D, second pleopod, with detail of the two setae on exopodite outer margin (arrow); E, third pleopod exopodite;F, detail ofthe outer-basal regionof the thirdpleopod exopoditeshowing the respiratory structures (thick arrow indicates the position of the spiracle); G, detail of the outer-basal region of the third pleopod exopodite showing the spiracle partially closed from above; H, fourth pleopod exopodite; I, fifth pleopod exopodite. Abbreviations: endopodite (en) and exopodite (ex). Scale bars = 0.5 mm (A–H), 0.25 mm (I)." captionText-7="Figure 8. Selected body parts of Acanthoniscus for comparison with other taxa within Armadillidae. Cubaris murina Brandt, 1833, showing the ground pattern in Armadillidae (A–E): A, Posterior region of the body in dorsal view; B, posterior region of the body in ventral view; C, pleotelson in dorsal view; D, uropod in dorsal view; E, middle region of the sternite 7 of a male showing the bilobed caudal process on the posterior margin, which is considered an autapomorphy of Armadillidae.Acanthoniscus richardsonae sp. nov.(F–H): F, pleotelson in dorsal view; G, uropod in dorsal view; H, middle region of the sternite 7 of a male.Drawings of the pleotelson and uropods in dorsal views of some morphologically divergent taxa within Armadillidae. I, Echinodillo cavaticus Green, 1963; J, Echinodillo montanus Jackson, 1933; K, Laureola vietnamensis Kwon, Ferrara and Taiti, 1992; L, Laureola dubia Schmalfuss and Ferrara,1983; M, Schismadillo rouxi Verhoeff,1926; N, Merulanella wahrbergi Verhoeff,1926; O, Nesodillo incisus Verhoeff, 1926. Drawings modified from Verhoeff (1926) (M–O), Jackson (1933) (J), Green (1963) (I), Schmalfuss and Ferrara (1983) (L), and Kwon et al. (1992) (K). Abbreviations: endopodite (en), exopodite (ex), sympodite (sy)." figureDoi-0="http://doi.org/10.5281/zenodo.10716766" figureDoi-1="http://doi.org/10.5281/zenodo.10716768" figureDoi-2="http://doi.org/10.5281/zenodo.10716770" figureDoi-3="http://doi.org/10.5281/zenodo.10716774" figureDoi-4="http://doi.org/10.5281/zenodo.10716776" figureDoi-5="http://doi.org/10.5281/zenodo.10716778" figureDoi-6="http://doi.org/10.5281/zenodo.10716780" figureDoi-7="http://doi.org/10.5281/zenodo.10716782" httpUri-0="https://zenodo.org/record/10716766/files/figure.png" httpUri-1="https://zenodo.org/record/10716768/files/figure.png" httpUri-2="https://zenodo.org/record/10716770/files/figure.png" httpUri-3="https://zenodo.org/record/10716774/files/figure.png" httpUri-4="https://zenodo.org/record/10716776/files/figure.png" httpUri-5="https://zenodo.org/record/10716778/files/figure.png" httpUri-6="https://zenodo.org/record/10716780/files/figure.png" httpUri-7="https://zenodo.org/record/10716782/files/figure.png" pageId="2" pageNumber="3">Figs. 1–8</figureCitation>
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)
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</paragraph>
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<taxonomicName id="4C404D7EFF04FFE8FCE8FC4AFB02FC32" ID-CoL="7NMLX" authority="White, 1847: 99" authorityName="White" authorityPageNumber="99" authorityYear="1847" box="[843,1209,926,955]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B934EAEFFF04FFE8FCE8FC4AFC49FC32" box="[843,1010,926,955]" italics="true" pageId="2" pageNumber="3">Acanthoniscus</emphasis>
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White, 1847: 99
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</taxonomicName>
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(Nomen nudum). —
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<treatmentCitation id="0AE110ECFF04FFE8FCD2FC10FB28FC68" author="Schomburgk RH" box="[881,1171,964,994]" page="658" pageId="2" pageNumber="3" year="1848">
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<bibRefCitation id="EFD14B0CFF04FFE8FCD2FC10FB28FC68" author="Schomburgk RH" box="[881,1171,964,994]" pageId="2" pageNumber="3" refId="ref17559" refString="Schomburgk RH 1848. The history of Barbados; comprising a geographical and statistical description of the island; a sketch of the historical events since the settlement; and an account of its geology and natural productions. London, Longman, Brown, Green and Longmans, xx + 722 p. https: // www. biodiversitylibrary. org / bibliography / 182786" type="book" year="1848">Schomburgk, 1848: 658</bibRefCitation>
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(Nomen nudum).
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<treatmentCitation id="0AE110ECFF04FFE8FCE8FC3FFB03FB81" author="Gosse PH" box="[843,1208,1003,1033]" page="65" pageId="2" pageNumber="3" year="1851">
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<taxonomicName id="4C404D7EFF04FFE8FCE8FC3FFB03FB81" ID-CoL="7NMLX" authority="Gosse, 1851: 65" authorityName="Gosse" authorityPageNumber="65" authorityYear="1851" box="[843,1208,1003,1033]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B934EAEFFF04FFE8FCE8FC3FFC49FB81" box="[843,1010,1003,1032]" italics="true" pageId="2" pageNumber="3">Acanthoniscus</emphasis>
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<bibRefCitation id="EFD14B0CFF04FFE8FC5BFC38FB03FB81" author="Gosse PH" box="[1016,1208,1003,1033]" pageId="2" pageNumber="3" refId="ref15567" refString="Gosse PH 1851. A naturalist's sojourn in Jamaica. London, Longman, Brown, Green and Longmans, 508 p. DOI: https: // www. doi. org / 10.5962 / bhl. title. 33264" type="book" year="1851">Gosse, 1851: 65</bibRefCitation>
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.
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</paragraph>
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<treatmentCitation id="0AE110ECFF04FFE8FCE8FBC6FAB2FBB9" author="Kinahan JR" box="[843,1289,1042,1072]" page="198" pageId="2" pageNumber="3" year="1859">
|
||
<taxonomicName id="4C404D7EFF04FFE8FCE8FBC6FAB2FBB9" ID-CoL="7NMLX" authority="Kinahan, 1859: 197 - 198" authorityName="Kinahan" authorityPageNumber="197 - 198" authorityYear="1859" box="[843,1289,1042,1072]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EAEFFF04FFE8FCE8FBC6FC51FBA6" box="[843,1002,1042,1071]" italics="true" pageId="2" pageNumber="3">Acanthoniscus</emphasis>
|
||
<bibRefCitation id="EFD14B0CFF04FFE8FC4DFBC6FAB2FBB9" author="Kinahan JR" box="[1006,1289,1042,1072]" pageId="2" pageNumber="3" pagination="125 - 135" refId="ref16294" refString="Kinahan JR 1859. On the genus Platyarthrus (Brandt); with notices of allied undescribed genera. Natural History Review, 6: 125 - 135, pl. 14." type="journal article" year="1859">Kinahan,1859:197–198</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
,plate200–201. —
|
||
<treatmentCitation id="0AE110ECFF04FFE8FC36FBEDFB22FBDF" author="Budde-Lund G" box="[917,1177,1081,1111]" page="5" pageId="2" pageNumber="3" year="1879">
|
||
<bibRefCitation id="EFD14B0CFF04FFE8FC36FBEDFB22FBDF" author="Budde-Lund G" box="[917,1177,1081,1111]" pageId="2" pageNumber="3" refId="ref14654" refString="Budde-Lund G 1879. Prospectus generum specierumque Crustaceorum Isopodum Terrestrium. p. 1 - 10. Copenhagen." type="book" year="1879">Budde-Lund, 1879: 5</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AE110ECFF04FFE8FB6FFBEDFC24FBF4" author="Budde-Lund G" page="241" pageId="2" pageNumber="3" year="1885">
|
||
<bibRefCitation id="EFD14B0CFF04FFE8FB6FFBEDFC24FBF4" author="Budde-Lund G" pageId="2" pageNumber="3" refId="ref14673" refString="Budde-Lund G 1885. Crustacea Isopoda terrestria per familias et genera et species descripta. Copenhagen, Havniae, 319 p. DOI: https: // www. doi. org / 10.5962 / bhl. title. 109769" type="book" year="1885">Budde-Lund, 1885: 241</bibRefCitation>
|
||
</treatmentCitation>
|
||
.—
|
||
<treatmentCitation id="0AE110ECFF04FFE8FC6AFBB4FB77FBF4" author="Richardson H" box="[969,1228,1120,1149]" page="562" pageId="2" pageNumber="3" year="1901">
|
||
<bibRefCitation id="EFD14B0CFF04FFE8FC6AFBB4FB77FBF4" author="Richardson H" box="[969,1228,1120,1149]" pageId="2" pageNumber="3" pagination="493 - 579" refId="ref16936" refString="Richardson H 1901. Key to the isopods of the Atlantic coast of North America with descriptions of new and little known species. Proceedings of the United States National Museum, 23: 493 - 579. DOI: https: // www. doi. org / 10.5479 / si. 00963801.23 - 1222.493" type="journal article" year="1901">Richardson, 1901:562</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 569.—
|
||
<treatmentCitation id="0AE110ECFF04FFE8FA8EFBB4FB8CFB2D" author="Richardson H" page="636" pageId="2" pageNumber="3" year="1905">
|
||
<bibRefCitation id="EFD14B0CFF04FFE8FA8EFBB4FB8CFB2D" author="Richardson H" pageId="2" pageNumber="3" refId="ref16992" refString="Richardson H 1905. A monograph on the isopods of North America. Bulletin of the United States National Museum, 54: xxiii, 727 p., 740 figs. DOI: https: // www. doi. org / 10.5479 / si. 03629236.54. i" type="book" year="1905">Richardson, 1905: 636–637</bibRefCitation>
|
||
</treatmentCitation>
|
||
(repeated the description of
|
||
<bibRefCitation id="EFD14B0CFF04FFE8FCD2FB7AFB96FB42" author="Kinahan JR" box="[881,1069,1198,1227]" pageId="2" pageNumber="3" pagination="125 - 135" refId="ref16294" refString="Kinahan JR 1859. On the genus Platyarthrus (Brandt); with notices of allied undescribed genera. Natural History Review, 6: 125 - 135, pl. 14." type="journal article" year="1859">Kinahan, 1859</bibRefCitation>
|
||
). —
|
||
<treatmentCitation id="0AE110ECFF04FFE8FBCFFB7AFA36FB42" author="Richardson H" box="[1132,1421,1198,1227]" page="432" pageId="2" pageNumber="3" year="1909">
|
||
<bibRefCitation id="EFD14B0CFF04FFE8FBCFFB7AFA36FB42" author="Richardson H" box="[1132,1421,1198,1227]" pageId="2" pageNumber="3" pagination="431 - 434" refId="ref17042" refString="Richardson H 1909. The isopod crustacean Acanthoniscus spiniger Kinahan redescribed. Proceedings of the United States National Museum, 36: 431 - 434. DOI: https: // www. doi. org / 10.5479 / si. 00963801.36 - 1678.431" type="journal article" year="1909">Richardson, 1909: 432</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AE110ECFF04FFE8FCD2FB01FBCEFB7B" author="Budde-Lund G" box="[881,1141,1237,1266]" page="11" pageId="2" pageNumber="3" year="1910">
|
||
<bibRefCitation id="EFD14B0CFF04FFE8FCD2FB01FBCEFB7B" author="Budde-Lund G" box="[881,1141,1237,1266]" pageId="2" pageNumber="3" pagination="1905 - 1906" refId="ref14713" refString="Budde-Lund G 1910. Crustacea, 2. Isopoda. In: Wissenschaftliche Ergebnisse der schwedischen zoologischen Expedition nach dem Kilimandjaro, dem Meru und den umgebenden Massaisteppen Deutsch-Ostafrikas, 1905 - 1906. p. 3 - 20, pls. 1 - 2. Stockholm, P. Palmquists Aktebolag. https: // www. biodiversitylibrary. org / item / 16983 # page / 651 / mode / 1 up" type="book chapter" year="1910">Budde-Lund, 1910: 11</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AE110ECFF04FFE8FB02FB01FA28FB7B" author="Arcangeli A" box="[1185,1427,1237,1266]" page="135" pageId="2" pageNumber="3" year="1927">
|
||
<bibRefCitation id="EFD14B0CFF04FFE8FB02FB01FA28FB7B" author="Arcangeli A" box="[1185,1427,1237,1266]" pageId="2" pageNumber="3" pagination="126 - 141" refId="ref14187" refString="Arcangeli A 1927. Revisione dei generi degli isopodi terrestri. I. a Nota. - Sopra alcuni generi di Africa e di America. Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 66: 126 - 141. https: // www. biodiversitylibrary. org / item / 264438 - page / 144 / mode / 1 up" type="journal article" year="1927">Arcangeli, 1927: 135</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AE110ECFF04FFE8FCCCFB29FB1DFA90" author="Van Name W" box="[879,1190,1276,1305]" page="401" pageId="2" pageNumber="3" year="1936">
|
||
<bibRefCitation id="EFD14B0CFF04FFE8FCCCFB29FB1DFA90" author="Van Name W" box="[879,1190,1276,1305]" pageId="2" pageNumber="3" pagination="1 - 535" refId="ref17876" refString="Van Name W 1936. The American land and freshwater isopod Crustacea. Bulletin of the American Museum of natural History, 71: 1 - 535. https: // hdl. handle. net / 2246 / 1185" type="journal article" year="1936">Van Name, 1936: 401–402</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AE110ECFF04FFE8FB73FB28FC5FFAC9" author="Clark PF & Presswell B" page="154" pageId="2" pageNumber="3" year="2001">
|
||
<bibRefCitation id="EFD14B0CFF04FFE8FB73FB28FC5FFAC9" author="Clark PF & Presswell B" pageId="2" pageNumber="3" pagination="149 - 166" refId="ref14827" refString="Clark PF and Presswell B 2001. Adam White: the crustacean years. The Raffles Bulletin of Zoology, 49: 149 - 166. https: // lkcnhm. nus. edu. sg / wp-content / uploads / sites / 10 / app / uploads / 2017 / 06 / 49 rbz 149 - 166. pdf" type="journal article" year="2001">Clark and Presswell, 2001:154</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AE110ECFF04FFE8FBACFAF7FB41FAC9" author="Schmalfuss H" box="[1039,1274,1315,1344]" page="4" pageId="2" pageNumber="3" year="2003">
|
||
<bibRefCitation id="EFD14B0CFF04FFE8FBACFAF7FB41FAC9" author="Schmalfuss H" box="[1039,1274,1315,1344]" pageId="2" pageNumber="3" refId="ref17197" refString="Schmalfuss H 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde, Serie A, 654: 341 p. https: // www. biodiversitylibrary. org / item / 293136 - page / 1 / mode / 1 up" type="book" year="2003">Schmalfuss, 2003: 4</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AE110ECFF04FFE8FA86FAF7FBE8FAEE" author="Schmidt C & Leistikow, A" page="4" pageId="2" pageNumber="3" year="2004">
|
||
<bibRefCitation id="EFD14B0CFF04FFE8FA86FAF7FBE8FAEE" author="Schmidt C & Leistikow, A" pageId="2" pageNumber="3" pagination="1 - 118" refId="ref17528" refString="Schmidt C and Leistikow, A 2004. Catalogue of genera of the terrestrial Isopoda (Crustacea: Isopoda: Oniscidea). Steenstrupia, 28: 1 - 118." type="journal article" year="2004">Schmidt and Leistikow, 2004: 4</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<bibRefCitation id="EFD14B0CFF04FFE8FBDFFA9EFAE1FAEE" author="Boyko CB & Bruce NL & Hadfield KA & Merrin KL & Ota Y & Poore GCB & Taiti S & Schotte M & Wilson GDF" box="[1148,1370,1354,1383]" pageId="2" pageNumber="3" refId="ref14575" refString="Boyko CB; Bruce NL; Hadfield KA; Merrin KL; Ota Y; Poore GCB; Taiti S; Schotte M and Wilson GDF (Eds.) (2008 onwards). World Marine, Freshwater and Terrestrial Isopod Crustaceans database. Acanthoniscus spiniger Kinahan, 1859. Available at https: // www. marinespecies. org / aphia. php? p = taxdetails & id = 258660. Accessed on 22 January 2022" type="journal volume" year="2008">Boyko et al., 2008</bibRefCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A6576FF04FFE8FCD2FA4DFC49FA52" pageId="2" pageNumber="3" type="type_taxon">
|
||
<paragraph id="8BFF36FDFF04FFE8FCD2FA4DFC49FA52" blockId="2.[843,1467,1432,1500]" pageId="2" pageNumber="3">
|
||
<emphasis id="B934EAEFFF04FFE8FCD2FA4DFC46FA3C" box="[881,1021,1433,1461]" italics="true" pageId="2" pageNumber="3">
|
||
<typeStatus id="54FB885FFF04FFE8FCD2FA4DFC13FA3C" box="[881,936,1433,1461]" pageId="2" pageNumber="3">Type</typeStatus>
|
||
species
|
||
</emphasis>
|
||
.
|
||
<taxonomicName id="4C404D7EFF04FFE8FBAEFA4CFA0EFA3C" authority="Gosse, 1851" authorityName="" (Gosse" authorityYear="1851" box="[1037,1461,1432,1461]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="spiniger">
|
||
<emphasis id="B934EAEFFF04FFE8FBAEFA4CFAACFA3C" box="[1037,1303,1432,1461]" italics="true" pageId="2" pageNumber="3">Acanthoniscus spiniger</emphasis>
|
||
<bibRefCitation id="EFD14B0CFF04FFE8FA83FA4CFA0EFA3C" author="Gosse PH" box="[1312,1461,1432,1461]" pageId="2" pageNumber="3" refId="ref15567" refString="Gosse PH 1851. A naturalist's sojourn in Jamaica. London, Longman, Brown, Green and Longmans, 508 p. DOI: https: // www. doi. org / 10.5962 / bhl. title. 33264" type="book" year="1851">Gosse, 1851</bibRefCitation>
|
||
</taxonomicName>
|
||
, by monotypy.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A6576FF04FFE8FCD2F9D8FC1FF9D9" pageId="2" pageNumber="3" type="taxon_list">
|
||
<paragraph id="8BFF36FDFF04FFE8FCD2F9D8FC1FF9D9" blockId="2.[843,1466,1548,1616]" pageId="2" pageNumber="3">
|
||
<emphasis id="B934EAEFFF04FFE8FCD2F9D8FBD6F9A0" box="[881,1133,1548,1577]" italics="true" pageId="2" pageNumber="3">Other species included</emphasis>
|
||
.
|
||
<taxonomicName id="4C404D7EFF04FFE8FBDAF9D8FA01F9A0" authority="Rodríguez-Cabrera & de Armas, 2023" authorityName="Rodríguez-Cabrera & de Armas" authorityYear="2023" box="[1145,1466,1548,1577]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
|
||
<emphasis id="B934EAEFFF04FFE8FBDAF9D8FA01F9A0" box="[1145,1466,1548,1577]" italics="true" pageId="2" pageNumber="3">Acanthoniscus richardsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2075794FF04FFE8FCE8F9E0FC1FF9D9" box="[843,932,1588,1616]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A6576FF04FFEEFCD2F956FBEEFE4C" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="diagnosis">
|
||
<paragraph id="8BFF36FDFF04FFEEFCD2F956FBEEFE4C" blockId="2.[842,1467,1665,1967]" lastBlockId="4.[841,1468,151,453]" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
|
||
<emphasis id="B934EAEFFF04FFE8FCD2F956FC53F917" box="[881,1000,1666,1694]" italics="true" pageId="2" pageNumber="3">Diagnosis</emphasis>
|
||
(emended). Body of fragile general appearance, with endoantennal conglobation ability involving all body segments (
|
||
<figureCitation id="137B2A78FF04FFE8FB09F91BFAA7F965" box="[1194,1308,1743,1772]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStartId-0="6.[181,252,1756,1779]" captionStartId-1="10.[843,913,586,609]" captionStartId-2="11.[181,252,1628,1651]" captionTargetBox-0="[206,1452,168,1723]" captionTargetBox-1="[843,1467,151,563]" captionTargetBox-2="[246,1407,152,1593]" captionTargetId-0="figure-175@6.[181,1466,148,1735]" captionTargetId-1="figure-639@10.[843,1467,148,565]" captionTargetId-2="figure-20@11.[229,1420,148,1608]" captionTargetPageId-0="6" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Figure 2. Body parts of Acanthoniscus spiniger (female holotype BMNH 1973.478.1). A, Overall, dorsal view; B, overall, ventral view; C, right pereon-epimera 1–7 in ventral view; D, overall, lateral view; E, right pleon-epimera 3–5 in ventral view; F, overall, frontal view; G, cephalothorax in dorsal view; H, cephalothorax in frontal view; I, cephalothorax in ventral view; J, pleotelson and uropods in dorsal view; K, pleotelson and uropods inventral view; L, right uropodin ventral view; M, arrangement of dorsal spines, the diameters of the circles correspond to the relative size of the spines. Drawings of the corresponding structures modified from Kinahan (1859) are included in J and L. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–K), 0.5 mm (L). Original photographs by The Trustees of the BMNH." captionText-1="Figure 3. Right lateral view of a female paratype Acanthoniscus richardsonaesp.nov.showingcolorationinalcohol.Scale bar=1 mm." captionText-2="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi-0="http://doi.org/10.5281/zenodo.10716768" figureDoi-1="http://doi.org/10.5281/zenodo.10716770" figureDoi-2="http://doi.org/10.5281/zenodo.10716774" httpUri-0="https://zenodo.org/record/10716768/files/figure.png" httpUri-1="https://zenodo.org/record/10716770/files/figure.png" httpUri-2="https://zenodo.org/record/10716774/files/figure.png" pageId="2" pageNumber="3">Figs. 2–4</figureCitation>
|
||
); coxal plates enlarged and fused with tergites, with more or less complex inner tooth-like processes forming cleft or schisma (except coxal plate 4, which lacks tooth-like processes) (
|
||
<figureCitation id="137B2A78FF04FFE8FC42F8BFFBF6F801" box="[993,1101,1899,1928]" captionStart="Figure 2" captionStartId="6.[181,252,1756,1779]" captionTargetBox="[206,1452,168,1723]" captionTargetId="figure-175@6.[181,1466,148,1735]" captionTargetPageId="6" captionText="Figure 2. Body parts of Acanthoniscus spiniger (female holotype BMNH 1973.478.1). A, Overall, dorsal view; B, overall, ventral view; C, right pereon-epimera 1–7 in ventral view; D, overall, lateral view; E, right pleon-epimera 3–5 in ventral view; F, overall, frontal view; G, cephalothorax in dorsal view; H, cephalothorax in frontal view; I, cephalothorax in ventral view; J, pleotelson and uropods in dorsal view; K, pleotelson and uropods inventral view; L, right uropodin ventral view; M, arrangement of dorsal spines, the diameters of the circles correspond to the relative size of the spines. Drawings of the corresponding structures modified from Kinahan (1859) are included in J and L. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–K), 0.5 mm (L). Original photographs by The Trustees of the BMNH." figureDoi="http://doi.org/10.5281/zenodo.10716768" httpUri="https://zenodo.org/record/10716768/files/figure.png" pageId="2" pageNumber="3">Figs. 2C</figureCitation>
|
||
,
|
||
<figureCitation id="137B2A78FF04FFE8FBF8F8BFFB13F801" box="[1115,1192,1899,1928]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="2" pageNumber="3">4F–G</figureCitation>
|
||
). Cephalothorax with upper margin of frontal shield irregular, forming lateral and median lobes (not antennal lobes); profrons with wide shallow depressions on each side divided by very low median ridge holding second antennae; dorsal surface with small spiniform tubercles and/or medium-sized, club-like spines (
|
||
<figureCitation id="137B2A78FF02FFEEFE6EFEE7FDA7FED9" box="[461,540,307,336]" captionStart="Figure 2" captionStartId="6.[181,252,1756,1779]" captionTargetBox="[206,1452,168,1723]" captionTargetId="figure-175@6.[181,1466,148,1735]" captionTargetPageId="6" captionText="Figure 2. Body parts of Acanthoniscus spiniger (female holotype BMNH 1973.478.1). A, Overall, dorsal view; B, overall, ventral view; C, right pereon-epimera 1–7 in ventral view; D, overall, lateral view; E, right pleon-epimera 3–5 in ventral view; F, overall, frontal view; G, cephalothorax in dorsal view; H, cephalothorax in frontal view; I, cephalothorax in ventral view; J, pleotelson and uropods in dorsal view; K, pleotelson and uropods inventral view; L, right uropodin ventral view; M, arrangement of dorsal spines, the diameters of the circles correspond to the relative size of the spines. Drawings of the corresponding structures modified from Kinahan (1859) are included in J and L. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–K), 0.5 mm (L). Original photographs by The Trustees of the BMNH." figureDoi="http://doi.org/10.5281/zenodo.10716768" httpUri="https://zenodo.org/record/10716768/files/figure.png" pageId="4" pageNumber="5">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="137B2A78FF02FFEEFD8BFEE7FD81FED9" box="[552,570,307,336]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="4" pageNumber="5">4</figureCitation>
|
||
). Eyes composed of 16 ommatidia (
|
||
<figureCitation id="137B2A78FF02FFEEFED4FE8EFE71FEFE" box="[375,458,346,375]" captionStart="Figure 2" captionStartId="6.[181,252,1756,1779]" captionTargetBox="[206,1452,168,1723]" captionTargetId="figure-175@6.[181,1466,148,1735]" captionTargetPageId="6" captionText="Figure 2. Body parts of Acanthoniscus spiniger (female holotype BMNH 1973.478.1). A, Overall, dorsal view; B, overall, ventral view; C, right pereon-epimera 1–7 in ventral view; D, overall, lateral view; E, right pleon-epimera 3–5 in ventral view; F, overall, frontal view; G, cephalothorax in dorsal view; H, cephalothorax in frontal view; I, cephalothorax in ventral view; J, pleotelson and uropods in dorsal view; K, pleotelson and uropods inventral view; L, right uropodin ventral view; M, arrangement of dorsal spines, the diameters of the circles correspond to the relative size of the spines. Drawings of the corresponding structures modified from Kinahan (1859) are included in J and L. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–K), 0.5 mm (L). Original photographs by The Trustees of the BMNH." figureDoi="http://doi.org/10.5281/zenodo.10716768" httpUri="https://zenodo.org/record/10716768/files/figure.png" pageId="4" pageNumber="5">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="137B2A78FF02FFEEFE7AFE8EFE50FEFE" box="[473,491,346,375]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="4" pageNumber="5">4</figureCitation>
|
||
). Pereon tergites covered with long, slightly sinuous dorsolateral spines and smaller posteromedian and lateral spines, sometimes with paramedian spiniform tubercles; with lateral spiniform tubercle near anterior margin of pereonites 2–7 and directed forward, as part of outer structures involved during conglobation;epimera1enlarged,with posterior angle and outer margin broadly rounded, anterior angle more acutely produced; epimera 2–7 enlarged and with acutely produced outer angle; male sternite 7 simple, without bilobed caudal process (
|
||
<figureCitation id="137B2A78FF02FFEEFF62FCD2FE88FCAA" box="[193,307,774,803]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStartId-0="6.[181,252,1756,1779]" captionStartId-1="10.[843,913,586,609]" captionStartId-2="11.[181,252,1628,1651]" captionTargetBox-0="[206,1452,168,1723]" captionTargetBox-1="[843,1467,151,563]" captionTargetBox-2="[246,1407,152,1593]" captionTargetId-0="figure-175@6.[181,1466,148,1735]" captionTargetId-1="figure-639@10.[843,1467,148,565]" captionTargetId-2="figure-20@11.[229,1420,148,1608]" captionTargetPageId-0="6" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Figure 2. Body parts of Acanthoniscus spiniger (female holotype BMNH 1973.478.1). A, Overall, dorsal view; B, overall, ventral view; C, right pereon-epimera 1–7 in ventral view; D, overall, lateral view; E, right pleon-epimera 3–5 in ventral view; F, overall, frontal view; G, cephalothorax in dorsal view; H, cephalothorax in frontal view; I, cephalothorax in ventral view; J, pleotelson and uropods in dorsal view; K, pleotelson and uropods inventral view; L, right uropodin ventral view; M, arrangement of dorsal spines, the diameters of the circles correspond to the relative size of the spines. Drawings of the corresponding structures modified from Kinahan (1859) are included in J and L. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–K), 0.5 mm (L). Original photographs by The Trustees of the BMNH." captionText-1="Figure 3. Right lateral view of a female paratype Acanthoniscus richardsonaesp.nov.showingcolorationinalcohol.Scale bar=1 mm." captionText-2="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi-0="http://doi.org/10.5281/zenodo.10716768" figureDoi-1="http://doi.org/10.5281/zenodo.10716770" figureDoi-2="http://doi.org/10.5281/zenodo.10716774" httpUri-0="https://zenodo.org/record/10716768/files/figure.png" httpUri-1="https://zenodo.org/record/10716770/files/figure.png" httpUri-2="https://zenodo.org/record/10716774/files/figure.png" pageId="4" pageNumber="5">Figs. 2–4</figureCitation>
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,
|
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<figureCitation id="137B2A78FF02FFEEFEE3FCD2FEEBFCAA" box="[320,336,774,803]" captionStart="Figure 8" captionStartId="15.[181,252,1523,1546]" captionTargetBox="[192,1460,173,1475]" captionTargetId="figure-248@15.[181,1466,151,1499]" captionTargetPageId="15" captionText="Figure 8. Selected body parts of Acanthoniscus for comparison with other taxa within Armadillidae. Cubaris murina Brandt, 1833, showing the ground pattern in Armadillidae (A–E): A, Posterior region of the body in dorsal view; B, posterior region of the body in ventral view; C, pleotelson in dorsal view; D, uropod in dorsal view; E, middle region of the sternite 7 of a male showing the bilobed caudal process on the posterior margin, which is considered an autapomorphy of Armadillidae.Acanthoniscus richardsonae sp. nov.(F–H): F, pleotelson in dorsal view; G, uropod in dorsal view; H, middle region of the sternite 7 of a male.Drawings of the pleotelson and uropods in dorsal views of some morphologically divergent taxa within Armadillidae. I, Echinodillo cavaticus Green, 1963; J, Echinodillo montanus Jackson, 1933; K, Laureola vietnamensis Kwon, Ferrara and Taiti, 1992; L, Laureola dubia Schmalfuss and Ferrara,1983; M, Schismadillo rouxi Verhoeff,1926; N, Merulanella wahrbergi Verhoeff,1926; O, Nesodillo incisus Verhoeff, 1926. Drawings modified from Verhoeff (1926) (M–O), Jackson (1933) (J), Green (1963) (I), Schmalfuss and Ferrara (1983) (L), and Kwon et al. (1992) (K). Abbreviations: endopodite (en), exopodite (ex), sympodite (sy)." figureDoi="http://doi.org/10.5281/zenodo.10716782" httpUri="https://zenodo.org/record/10716782/files/figure.png" pageId="4" pageNumber="5">8</figureCitation>
|
||
). Pleon tergites 3–4 with pair of small posteromedian spines or spiniform tubercles; epimera of pleonites 3–5 enlarged and acutely produced (
|
||
<figureCitation id="137B2A78FF02FFEEFD4EFC80FF52FC11" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStartId-0="6.[181,252,1756,1779]" captionStartId-1="10.[843,913,586,609]" captionStartId-2="11.[181,252,1628,1651]" captionTargetBox-0="[206,1452,168,1723]" captionTargetBox-1="[843,1467,151,563]" captionTargetBox-2="[246,1407,152,1593]" captionTargetId-0="figure-175@6.[181,1466,148,1735]" captionTargetId-1="figure-639@10.[843,1467,148,565]" captionTargetId-2="figure-20@11.[229,1420,148,1608]" captionTargetPageId-0="6" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Figure 2. Body parts of Acanthoniscus spiniger (female holotype BMNH 1973.478.1). A, Overall, dorsal view; B, overall, ventral view; C, right pereon-epimera 1–7 in ventral view; D, overall, lateral view; E, right pleon-epimera 3–5 in ventral view; F, overall, frontal view; G, cephalothorax in dorsal view; H, cephalothorax in frontal view; I, cephalothorax in ventral view; J, pleotelson and uropods in dorsal view; K, pleotelson and uropods inventral view; L, right uropodin ventral view; M, arrangement of dorsal spines, the diameters of the circles correspond to the relative size of the spines. Drawings of the corresponding structures modified from Kinahan (1859) are included in J and L. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–K), 0.5 mm (L). Original photographs by The Trustees of the BMNH." captionText-1="Figure 3. Right lateral view of a female paratype Acanthoniscus richardsonaesp.nov.showingcolorationinalcohol.Scale bar=1 mm." captionText-2="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi-0="http://doi.org/10.5281/zenodo.10716768" figureDoi-1="http://doi.org/10.5281/zenodo.10716770" figureDoi-2="http://doi.org/10.5281/zenodo.10716774" httpUri-0="https://zenodo.org/record/10716768/files/figure.png" httpUri-1="https://zenodo.org/record/10716770/files/figure.png" httpUri-2="https://zenodo.org/record/10716774/files/figure.png" pageId="4" pageNumber="5">Figs. 2–4</figureCitation>
|
||
). Pleotelson “hourglass-shaped,” with basal part wider than distal part, distal margin broadly convex and ending in 2 triangular projections separated by notch, basal half with 2 dorsal spines 1.3–1.5 times as long as pleotelson, widely surpassing its distal margin and uropod exopodite tip (
|
||
<figureCitation id="137B2A78FF02FFEEFDDFFBE9FD6FFBD2" box="[636,724,1085,1115]" captionStart="Figure 2" captionStartId="6.[181,252,1756,1779]" captionTargetBox="[206,1452,168,1723]" captionTargetId="figure-175@6.[181,1466,148,1735]" captionTargetPageId="6" captionText="Figure 2. Body parts of Acanthoniscus spiniger (female holotype BMNH 1973.478.1). A, Overall, dorsal view; B, overall, ventral view; C, right pereon-epimera 1–7 in ventral view; D, overall, lateral view; E, right pleon-epimera 3–5 in ventral view; F, overall, frontal view; G, cephalothorax in dorsal view; H, cephalothorax in frontal view; I, cephalothorax in ventral view; J, pleotelson and uropods in dorsal view; K, pleotelson and uropods inventral view; L, right uropodin ventral view; M, arrangement of dorsal spines, the diameters of the circles correspond to the relative size of the spines. Drawings of the corresponding structures modified from Kinahan (1859) are included in J and L. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–K), 0.5 mm (L). Original photographs by The Trustees of the BMNH." figureDoi="http://doi.org/10.5281/zenodo.10716768" httpUri="https://zenodo.org/record/10716768/files/figure.png" pageId="4" pageNumber="5">Figs. 2</figureCitation>
|
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,
|
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<figureCitation id="137B2A78FF02FFEEFD40FBEAFD4FFBD2" box="[739,756,1086,1115]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="4" pageNumber="5">4</figureCitation>
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,
|
||
<figureCitation id="137B2A78FF02FFEEFCA7FBEAFCAEFBD2" box="[772,789,1086,1115]" captionStart="Figure 8" captionStartId="15.[181,252,1523,1546]" captionTargetBox="[192,1460,173,1475]" captionTargetId="figure-248@15.[181,1466,151,1499]" captionTargetPageId="15" captionText="Figure 8. Selected body parts of Acanthoniscus for comparison with other taxa within Armadillidae. Cubaris murina Brandt, 1833, showing the ground pattern in Armadillidae (A–E): A, Posterior region of the body in dorsal view; B, posterior region of the body in ventral view; C, pleotelson in dorsal view; D, uropod in dorsal view; E, middle region of the sternite 7 of a male showing the bilobed caudal process on the posterior margin, which is considered an autapomorphy of Armadillidae.Acanthoniscus richardsonae sp. nov.(F–H): F, pleotelson in dorsal view; G, uropod in dorsal view; H, middle region of the sternite 7 of a male.Drawings of the pleotelson and uropods in dorsal views of some morphologically divergent taxa within Armadillidae. I, Echinodillo cavaticus Green, 1963; J, Echinodillo montanus Jackson, 1933; K, Laureola vietnamensis Kwon, Ferrara and Taiti, 1992; L, Laureola dubia Schmalfuss and Ferrara,1983; M, Schismadillo rouxi Verhoeff,1926; N, Merulanella wahrbergi Verhoeff,1926; O, Nesodillo incisus Verhoeff, 1926. Drawings modified from Verhoeff (1926) (M–O), Jackson (1933) (J), Green (1963) (I), Schmalfuss and Ferrara (1983) (L), and Kwon et al. (1992) (K). Abbreviations: endopodite (en), exopodite (ex), sympodite (sy)." figureDoi="http://doi.org/10.5281/zenodo.10716782" httpUri="https://zenodo.org/record/10716782/files/figure.png" pageId="4" pageNumber="5">8</figureCitation>
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). Uropod sympodite asymmetric and dorsoventrally f lattened, filling gap between pleotelson and fifth pleon epimera, broader at base, medial margin deeply concave and irregular, with inner lobe above and basal with respect to insertion point of exopodite which, together with wide transversal dorsal furrow, engaging with pleotelson, outer margin from slightly to widely convex, and medial-posterior angle acutely produced (more or less sharp-pointed); uropod endopodite laterally compressed (outer surface concave to engage with medial margin of uropod sympodite), inserted near base of sympodite and not surpassing distal margin of pleotelson, but it may surpass posterior margin of uropod sympodite (e.g.,
|
||
<taxonomicName id="4C404D7EFF02FFEEFDE7F98BFD0EF9F2" authorityName="" (Gosse" authorityYear="1851" box="[580,693,1631,1659]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="spiniger">
|
||
<emphasis id="B934EAEFFF02FFEEFDE7F98BFD0EF9F2" box="[580,693,1631,1659]" italics="true" pageId="4" pageNumber="5">A. spiniger</emphasis>
|
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</taxonomicName>
|
||
); uropod exopodite extremely long (longer than sympodite), narrow, and slightly curved (stick-shaped), inserted on medial margin of sympodite (visible dorsally and ventrally) and widely surpassing posterior margin of sympodite (by 60–80% of its length) and posterior margin of pleotelson (
|
||
<figureCitation id="137B2A78FF02FFEEFE10F89CFDBBF8EC" box="[435,512,1864,1893]" captionStart="Figure 2" captionStartId="6.[181,252,1756,1779]" captionTargetBox="[206,1452,168,1723]" captionTargetId="figure-175@6.[181,1466,148,1735]" captionTargetPageId="6" captionText="Figure 2. Body parts of Acanthoniscus spiniger (female holotype BMNH 1973.478.1). A, Overall, dorsal view; B, overall, ventral view; C, right pereon-epimera 1–7 in ventral view; D, overall, lateral view; E, right pleon-epimera 3–5 in ventral view; F, overall, frontal view; G, cephalothorax in dorsal view; H, cephalothorax in frontal view; I, cephalothorax in ventral view; J, pleotelson and uropods in dorsal view; K, pleotelson and uropods inventral view; L, right uropodin ventral view; M, arrangement of dorsal spines, the diameters of the circles correspond to the relative size of the spines. Drawings of the corresponding structures modified from Kinahan (1859) are included in J and L. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–K), 0.5 mm (L). Original photographs by The Trustees of the BMNH." figureDoi="http://doi.org/10.5281/zenodo.10716768" httpUri="https://zenodo.org/record/10716768/files/figure.png" pageId="4" pageNumber="5">Figs. 2</figureCitation>
|
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,
|
||
<figureCitation id="137B2A78FF02FFEEFDAAF89CFDA2F8EC" box="[521,537,1864,1893]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="4" pageNumber="5">4</figureCitation>
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,
|
||
<figureCitation id="137B2A78FF02FFEEFD81F89CFD89F8EC" box="[546,562,1864,1893]" captionStart="Figure 8" captionStartId="15.[181,252,1523,1546]" captionTargetBox="[192,1460,173,1475]" captionTargetId="figure-248@15.[181,1466,151,1499]" captionTargetPageId="15" captionText="Figure 8. Selected body parts of Acanthoniscus for comparison with other taxa within Armadillidae. Cubaris murina Brandt, 1833, showing the ground pattern in Armadillidae (A–E): A, Posterior region of the body in dorsal view; B, posterior region of the body in ventral view; C, pleotelson in dorsal view; D, uropod in dorsal view; E, middle region of the sternite 7 of a male showing the bilobed caudal process on the posterior margin, which is considered an autapomorphy of Armadillidae.Acanthoniscus richardsonae sp. nov.(F–H): F, pleotelson in dorsal view; G, uropod in dorsal view; H, middle region of the sternite 7 of a male.Drawings of the pleotelson and uropods in dorsal views of some morphologically divergent taxa within Armadillidae. I, Echinodillo cavaticus Green, 1963; J, Echinodillo montanus Jackson, 1933; K, Laureola vietnamensis Kwon, Ferrara and Taiti, 1992; L, Laureola dubia Schmalfuss and Ferrara,1983; M, Schismadillo rouxi Verhoeff,1926; N, Merulanella wahrbergi Verhoeff,1926; O, Nesodillo incisus Verhoeff, 1926. Drawings modified from Verhoeff (1926) (M–O), Jackson (1933) (J), Green (1963) (I), Schmalfuss and Ferrara (1983) (L), and Kwon et al. (1992) (K). Abbreviations: endopodite (en), exopodite (ex), sympodite (sy)." figureDoi="http://doi.org/10.5281/zenodo.10716782" httpUri="https://zenodo.org/record/10716782/files/figure.png" pageId="4" pageNumber="5">8</figureCitation>
|
||
). Pereiopod dactylus with simple, straight to curved dactylar seta, inner claw, sickle-shaped ungual seta (shorter than inner claw and not surpassing basal half of outer claw length), and other smaller setae similar to aesthetascs of antennae (
|
||
<figureCitation id="137B2A78FF02FFEEFC49FF31FBE9FE8B" box="[1002,1106,229,258]" captionStart="Figure 5" captionStartId="12.[182,251,1692,1715]" captionTargetBox="[312,1351,473,1658]" captionTargetId="figure-134@12.[266,1382,465,1671]" captionTargetPageId="12" captionText="Figure 5.Otherappendagesof Acanthoniscus richardsonae sp.nov.A, Second antenna;B,detail offlagellum of secondantennadepicting position of aesthetasc groups (small arrows); C, detail of the distal article of the flagellum of the second antenna slightly turned over showing the aesthetasc groups; D, detail of the distal-most portion of the flagellum of the second antenna showing the apical cone; E, first antenna; F, detail of the third article of the first antenna showing the aesthetascs; G, male left seventh pereiopod; H, detail of ischium ofmale seventh pereiopod; I, detailof dactylus ofmale seventh pereiopodin anterior view; J, detailof dactylus ofmale seventh pereiopod in posterior view; K, male left first pereiopod, showing the areas on the carpus and propodus covered with the cleaning apparatus or antennal brush (dashed outlines); L, detail of carpus ofmale first pereiopod (anterior surface).Abbreviations:basis (bs), ischium(is),merus(me),carpus (ca),propodus (pr),flagellum(fl),and dactylus(da).Drawingofthe corresponding structuremodified from Richardson (1909) is included inA. Scale bars = 1 mm (A), 0.5 mm (G,K), 0.3mm (B), 0.1 mm (C–E, H–J,L),0.05 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10716776" httpUri="https://zenodo.org/record/10716776/files/figure.png" pageId="4" pageNumber="5">Fig. 5I, J</figureCitation>
|
||
). First antenna 3-jointed (
|
||
<figureCitation id="137B2A78FF02FFEEFA2EFF31FC2FFEA0" captionStart="Figure 5" captionStartId="12.[182,251,1692,1715]" captionTargetBox="[312,1351,473,1658]" captionTargetId="figure-134@12.[266,1382,465,1671]" captionTargetPageId="12" captionText="Figure 5.Otherappendagesof Acanthoniscus richardsonae sp.nov.A, Second antenna;B,detail offlagellum of secondantennadepicting position of aesthetasc groups (small arrows); C, detail of the distal article of the flagellum of the second antenna slightly turned over showing the aesthetasc groups; D, detail of the distal-most portion of the flagellum of the second antenna showing the apical cone; E, first antenna; F, detail of the third article of the first antenna showing the aesthetascs; G, male left seventh pereiopod; H, detail of ischium ofmale seventh pereiopod; I, detailof dactylus ofmale seventh pereiopodin anterior view; J, detailof dactylus ofmale seventh pereiopod in posterior view; K, male left first pereiopod, showing the areas on the carpus and propodus covered with the cleaning apparatus or antennal brush (dashed outlines); L, detail of carpus ofmale first pereiopod (anterior surface).Abbreviations:basis (bs), ischium(is),merus(me),carpus (ca),propodus (pr),flagellum(fl),and dactylus(da).Drawingofthe corresponding structuremodified from Richardson (1909) is included inA. Scale bars = 1 mm (A), 0.5 mm (G,K), 0.3mm (B), 0.1 mm (C–E, H–J,L),0.05 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10716776" httpUri="https://zenodo.org/record/10716776/files/figure.png" pageId="4" pageNumber="5">Fig. 5E–F</figureCitation>
|
||
). Second antenna long and slender, sum of its articles being 2.5 times width of cephalothorax, flagellum 2-jointed and with 2 groups of aesthetascs on distal article (
|
||
<figureCitation id="137B2A78FF02FFEEFB82FE55FB1EFE14" box="[1057,1189,384,414]" captionStart="Figure 5" captionStartId="12.[182,251,1692,1715]" captionTargetBox="[312,1351,473,1658]" captionTargetId="figure-134@12.[266,1382,465,1671]" captionTargetPageId="12" captionText="Figure 5.Otherappendagesof Acanthoniscus richardsonae sp.nov.A, Second antenna;B,detail offlagellum of secondantennadepicting position of aesthetasc groups (small arrows); C, detail of the distal article of the flagellum of the second antenna slightly turned over showing the aesthetasc groups; D, detail of the distal-most portion of the flagellum of the second antenna showing the apical cone; E, first antenna; F, detail of the third article of the first antenna showing the aesthetascs; G, male left seventh pereiopod; H, detail of ischium ofmale seventh pereiopod; I, detailof dactylus ofmale seventh pereiopodin anterior view; J, detailof dactylus ofmale seventh pereiopod in posterior view; K, male left first pereiopod, showing the areas on the carpus and propodus covered with the cleaning apparatus or antennal brush (dashed outlines); L, detail of carpus ofmale first pereiopod (anterior surface).Abbreviations:basis (bs), ischium(is),merus(me),carpus (ca),propodus (pr),flagellum(fl),and dactylus(da).Drawingofthe corresponding structuremodified from Richardson (1909) is included inA. Scale bars = 1 mm (A), 0.5 mm (G,K), 0.3mm (B), 0.1 mm (C–E, H–J,L),0.05 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10716776" httpUri="https://zenodo.org/record/10716776/files/figure.png" pageId="4" pageNumber="5">Fig. 5A–D</figureCitation>
|
||
). Pleopods with single spiracle lungs (
|
||
<figureCitation id="137B2A78FF02FFEEFBA2FE7CFBFEFE4C" box="[1025,1093,424,453]" captionStart="Figure 7" captionStartId="14.[181,252,1786,1809]" captionTargetBox="[387,1266,935,1753]" captionTargetId="figure-324@14.[356,1292,932,1765]" captionTargetPageId="14" captionText="Figure 7.Pleopods of a male Acanthoniscus richardsonae sp.nov.A, First pleopod; B, detail of first pleopod endopodite tip; C, detail of first pleopod exopodite; D, second pleopod, with detail of the two setae on exopodite outer margin (arrow); E, third pleopod exopodite;F, detail ofthe outer-basal regionof the thirdpleopod exopoditeshowing the respiratory structures (thick arrow indicates the position of the spiracle); G, detail of the outer-basal region of the third pleopod exopodite showing the spiracle partially closed from above; H, fourth pleopod exopodite; I, fifth pleopod exopodite. Abbreviations: endopodite (en) and exopodite (ex). Scale bars = 0.5 mm (A–H), 0.25 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.10716780" httpUri="https://zenodo.org/record/10716780/files/figure.png" pageId="4" pageNumber="5">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3F6675FF05FFE9FF16F99FFE60F88B" ID-DOI="http://doi.org/10.5281/zenodo.10716766" ID-Zenodo-Dep="10716766" httpUri="https://zenodo.org/record/10716766/files/figure.png" pageId="3" pageNumber="4" startId="3.[181,252,1611,1634]" targetBox="[181,1467,151,1587]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BFF36FDFF05FFE9FF16F99FFE60F88B" blockId="3.[179,1466,1610,1794]" pageId="3" pageNumber="4">
|
||
<emphasis id="B934EAEFFF05FFE9FF16F99FFEA3F9EB" bold="true" box="[181,280,1611,1634]" pageId="3" pageNumber="4">Figure 1.</emphasis>
|
||
Distribution of the genus
|
||
<taxonomicName id="4C404D7EFF05FFE9FDB8F99EFD19F9EB" authorityName="Kinahan" authorityYear="1859" box="[539,674,1610,1634]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EAEFFF05FFE9FDB8F99EFD19F9EB" box="[539,674,1610,1634]" italics="true" pageId="3" pageNumber="4">Acanthoniscus</emphasis>
|
||
</taxonomicName>
|
||
in Jamaica. Above: type locality of
|
||
<taxonomicName id="4C404D7EFF05FFE9FC5CF99EFBD8F9EB" authorityName="" (Gosse" authorityYear="1851" box="[1023,1123,1610,1634]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="spiniger">
|
||
<emphasis id="B934EAEFFF05FFE9FC5CF99EFBD8F9EB" box="[1023,1123,1610,1634]" italics="true" pageId="3" pageNumber="4">A. spiniger</emphasis>
|
||
</taxonomicName>
|
||
at the hills near Bluefields (yellow dot), the type locality of
|
||
<taxonomicName id="4C404D7EFF05FFE9FE07F9BEFD8DF90B" authority="Rodríguez-Cabrera & de Armas, 2023" authorityName="Rodríguez-Cabrera & de Armas" authorityYear="2023" box="[420,566,1642,1666]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
|
||
<emphasis id="B934EAEFFF05FFE9FE07F9BEFD8DF90B" box="[420,566,1642,1666]" italics="true" pageId="3" pageNumber="4">A. richardsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2075794FF05FFE9FD99F9BEFD38F908" box="[570,643,1642,1665]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in the Blue Mountains (red dot), and a second location in the Blue Mountains where a specimen presumably of
|
||
<taxonomicName id="4C404D7EFF05FFE9FE1AF95EFDF6F92B" authority="Rodríguez-Cabrera & de Armas, 2023" authorityName="Rodríguez-Cabrera & de Armas" authorityYear="2023" box="[441,589,1674,1698]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
|
||
<emphasis id="B934EAEFFF05FFE9FE1AF95EFDF6F92B" box="[441,589,1674,1698]" italics="true" pageId="3" pageNumber="4">A. richardsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2075794FF05FFE9FDF1F95EFD27F928" box="[594,668,1674,1697]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was photographed (orange dot). The main collection sites of Henry G. Hubbard in 1877 are depicted by green dots, notice the proximity of Stony Hill to the distribution of
|
||
<taxonomicName id="4C404D7EFF05FFE9FB91F97EFB7DF94B" authority="Rodríguez-Cabrera & de Armas, 2023" authorityName="Rodríguez-Cabrera & de Armas" authorityYear="2023" box="[1074,1222,1706,1730]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
|
||
<emphasis id="B934EAEFFF05FFE9FB91F97EFB7DF94B" box="[1074,1222,1706,1730]" italics="true" pageId="3" pageNumber="4">A. richardsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2075794FF05FFE9FB69F97EFAAFF948" box="[1226,1300,1706,1729]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Below: The hills known as Surinam Quarters near Bluefields, referred by
|
||
<bibRefCitation id="EFD14B0CFF05FFE9FD40F91EFCD5F96B" author="Gosse PH" box="[739,878,1738,1762]" pageId="3" pageNumber="4" refId="ref15567" refString="Gosse PH 1851. A naturalist's sojourn in Jamaica. London, Longman, Brown, Green and Longmans, 508 p. DOI: https: // www. doi. org / 10.5962 / bhl. title. 33264" type="book" year="1851">Gosse (1851)</bibRefCitation>
|
||
as “Bluefields Mountain,” Westmoreland Parish, constitute the type locality of
|
||
<taxonomicName id="4C404D7EFF05FFE9FED0F93EFE6CF88B" authorityName="" (Gosse" authorityYear="1851" box="[371,471,1770,1794]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="spiniger">A. spiniger</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35A6576FF02FFEEFCD2FE21FB6AFD9A" box="[881,1233,501,531]" pageId="4" pageNumber="5" type="distribution">
|
||
<paragraph id="8BFF36FDFF02FFEEFCD2FE21FB6AFD9A" blockId="4.[881,1233,501,531]" box="[881,1233,501,531]" pageId="4" pageNumber="5">
|
||
<emphasis id="B934EAEFFF02FFEEFCD2FE21FBBBFD9B" box="[881,1024,501,530]" italics="true" pageId="4" pageNumber="5">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F357766DFF02FFEEFBAEFE21FBD6FD9B" box="[1037,1133,501,530]" name="Jamaica" pageId="4" pageNumber="5">Jamaica</collectingCountry>
|
||
(
|
||
<figureCitation id="137B2A78FF02FFEEFBDCFE21FB7BFD9A" box="[1151,1216,501,531]" captionStart="Figure 1" captionStartId="3.[181,252,1611,1634]" captionTargetBox="[181,1467,151,1587]" captionTargetId="figure-19@3.[181,1467,151,1587]" captionTargetPageId="3" captionText="Figure 1. Distribution of the genus Acanthoniscus in Jamaica. Above: type locality of A. spiniger at the hills near Bluefields (yellow dot), the type locality of A.richardsonae sp.nov.in the Blue Mountains (red dot), and a second location in the Blue Mountains where a specimen presumably of A. richardsonae sp. nov. was photographed (orange dot). The main collection sites of Henry G. Hubbard in 1877 are depicted by green dots, notice the proximity of Stony Hill to the distribution of A. richardsonae sp.nov.Below: The hills known as Surinam Quarters near Bluefields, referred by Gosse (1851) as “Bluefields Mountain,” Westmoreland Parish, constitute the type locality of A. spiniger." figureDoi="http://doi.org/10.5281/zenodo.10716766" httpUri="https://zenodo.org/record/10716766/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A6576FF02FFEFFCD2FD90FC9FFCAA" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="discussion">
|
||
<paragraph id="8BFF36FDFF02FFEEFCD2FD90FB42FB84" blockId="4.[841,1467,579,1037]" pageId="4" pageNumber="5">
|
||
<emphasis id="B934EAEFFF02FFEEFCD2FD90FBABFDE9" box="[881,1040,580,608]" italics="true" pageId="4" pageNumber="5">Comparisons.</emphasis>
|
||
The genus
|
||
<taxonomicName id="4C404D7EFF02FFEEFB37FD97FA82FDE9" authorityName="Kinahan" authorityYear="1859" box="[1172,1337,579,608]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EAEFFF02FFEEFB37FD97FA82FDE9" box="[1172,1337,579,608]" italics="true" pageId="4" pageNumber="5">Acanthoniscus</emphasis>
|
||
</taxonomicName>
|
||
very much resembles other specialized(spiny) forms of Old World
|
||
<taxonomicName id="4C404D7EFF02FFEEFCEAFD45FC53FD27" authorityName="Brandt" authorityYear="1831" box="[841,1000,657,686]" class="Malacostraca" family="Armadillidae" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Armadillidae</taxonomicName>
|
||
, such as the genera
|
||
<taxonomicName id="4C404D7EFF02FFEEFB61FD45FC30FD5C" authority="Collinge, 1922" authorityName="Collinge" authorityYear="1922" class="Malacostraca" family="Armadillidae" genus="Calmanesia" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EAEFFF02FFEEFB61FD45FAF3FD27" box="[1218,1352,657,686]" italics="true" pageId="4" pageNumber="5">Calmanesia</emphasis>
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFAEDFD45FC30FD5C" author="Collinge W" pageId="4" pageNumber="5" pagination="107 - 113" refId="ref14887" refString="Collinge W 1922. Two new terrestrial isopods from Madagascar. Journal of the Linnean Society, Zoology, 35: 107 - 113, pl. 9. DOI: https: // www. doi. org / 10.1111 / j. 1096 - 3642.1922. tb 00466. x" type="journal article" year="1922">Collinge, 1922</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C404D7EFF02FFEEFC3BFD6CFB6BFD5C" authority="Jackson, 1933" authorityName="Jackson" authorityYear="1933" box="[920,1232,696,725]" class="Malacostraca" family="Armadillidae" genus="Echinodillo" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EAEFFF02FFEEFC3BFD6CFBA0FD5C" box="[920,1051,696,725]" italics="true" pageId="4" pageNumber="5">Echinodillo</emphasis>
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFB87FD6CFB6BFD5C" author="Jackson H" box="[1060,1232,696,725]" pageId="4" pageNumber="5" pagination="145 - 162" refId="ref16063" refString="Jackson H 1933. Marquesan terrestrial Isopoda. Bulletin of the Bernice P. Bishop Museum, 114: 145 - 162." type="journal article" year="1933">Jackson, 1933</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C404D7EFF02FFEEFB7DFD6CFC30FD75" authority="Barnard, 1960" authorityName="Barnard" authorityYear="1960" class="Malacostraca" family="Armadillidae" genus="Laureola" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EAEFFF02FFEEFB7DFD6CFAFCFD5C" box="[1246,1351,696,725]" italics="true" pageId="4" pageNumber="5">Laureola</emphasis>
|
||
Barnard, 1960
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C404D7EFF02FFEEFC37FD0BFA0DFD75" authority="Kwon, Ferrara and Taiti, 1992" authorityName="Kwon, Ferrara and Taiti" authorityYear="1992" box="[916,1462,735,764]" class="Malacostraca" family="Armadillidae" genus="Pseudolaureola" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EAEFFF02FFEEFC37FD0BFBFFFD75" box="[916,1092,735,764]" italics="true" pageId="4" pageNumber="5">Pseudolaureola</emphasis>
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFBE9FD34FA0DFD75" author="Kwon D & Ferrara F & Taiti S" box="[1098,1462,735,764]" pageId="4" pageNumber="5" pagination="645 - 653" refId="ref16327" refString="Kwon D; Ferrara F and Taiti S 1992. Two new species of Laureola Barnard, 1960 from India and Vietnam (Crustacea, Oniscidea, Armadillidae). Revue suisse de Zoologie, 99: 645 - 653. DOI: https: // www. doi. org / 10.5962 / bhl. part. 79844" type="journal article" year="1992">Kwon, Ferrara and Taiti, 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C404D7EFF02FFEEFCDEFCD2FB05FCAA" authority="Jackson, 1933" authorityName="Jackson" authorityYear="1933" box="[893,1214,774,803]" class="Malacostraca" family="Armadillidae" genus="Tridentodillo" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EAEFFF02FFEEFCDEFCD2FBAFFCAA" box="[893,1044,774,803]" italics="true" pageId="4" pageNumber="5">Tridentodillo</emphasis>
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFBBAFCD2FB05FCAA" author="Jackson H" box="[1049,1214,774,803]" pageId="4" pageNumber="5" pagination="145 - 162" refId="ref16063" refString="Jackson H 1933. Marquesan terrestrial Isopoda. Bulletin of the Bernice P. Bishop Museum, 114: 145 - 162." type="journal article" year="1933">Jackson, 1933</bibRefCitation>
|
||
</taxonomicName>
|
||
(e.g.,
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFAA2FCD2FA0EFCAA" author="Collinge W" box="[1281,1461,774,803]" pageId="4" pageNumber="5" pagination="107 - 113" refId="ref14887" refString="Collinge W 1922. Two new terrestrial isopods from Madagascar. Journal of the Linnean Society, Zoology, 35: 107 - 113, pl. 9. DOI: https: // www. doi. org / 10.1111 / j. 1096 - 3642.1922. tb 00466. x" type="journal article" year="1922">Collinge, 1922</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFCEAFCF9FC49FCC3" author="Jackson H" box="[841,1010,813,842]" pageId="4" pageNumber="5" pagination="145 - 162" refId="ref16063" refString="Jackson H 1933. Marquesan terrestrial Isopoda. Bulletin of the Bernice P. Bishop Museum, 114: 145 - 162." type="journal article" year="1933">Jackson, 1933</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFC5FFCF9FB10FCC3" author="Barnard K" box="[1020,1195,813,842]" pageId="4" pageNumber="5" pagination="67 - 111" refId="ref14477" refString="Barnard K 1958. Terrestrial isopods and amphipods from Madagascar. Memoires de l'Institute scientifique de Madagascar, Serie A, 12: 67 - 111." type="journal article" year="1958">Barnard, 1958</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFB17FCF9FAB8FCC3" author="Barnard K" box="[1204,1283,813,842]" pageId="4" pageNumber="5" pagination="59 - 61" refId="ref14504" refString="Barnard K 1960 a. A new species of Calmanesia from Madagascar. Memoires de l'Institute scientifique de Madagascar, Serie A, 14: 59 - 61." type="journal article" year="1960">1960a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFAAFFCF9FAE7FCC3" author="Barnard K" box="[1292,1372,813,842]" pageId="4" pageNumber="5" pagination="45 - 55" refId="ref14533" refString="Barnard K 1960 b. Terrestrial Isopoda from the Transvaal. Annals of the Natal Museum, 15: 45 - 55. https: // journals. co. za / doi / epdf / 10.10520 / AJA 03040798 _ 794" type="journal article" year="1960">1960b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFAC5FCF9FC32FCF8" author="Green A" pageId="4" pageNumber="5" pagination="77 - 80" refId="ref15608" refString="Green A 1963. A new species of Echinodillo (Isopoda, Oniscoidea, Armadillidae) from Flinders Island, Tasmania. Papers and Proceedings of the Royal Society of Tasmania, 97: 77 - 80. https: // eprints. utas. edu. au / 14329 / 1 / 1963 _ Green _ New _ species _ Echinodillo. pdf" type="journal article" year="1963">Green, 1963</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFC31FC80FB95FCF8" author="Vandel A" box="[914,1070,852,881]" pageId="4" pageNumber="5" pagination="385 - 426" refId="ref18018" refString="Vandel A 1977. La faune terrestre de l'Ile de Sainte-Helene. 4 me partie. 1. Isopodes terrestres. Annales du Musee royal de l'Afrique centrale, Serie in 8 °, Sciences zoologiques, 220: 385 - 426, pl. II." type="journal article" year="1977">Vandel, 1977</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFB9BFC81FABEFCF8" author="Kwon D & Ferrara F & Taiti S" box="[1080,1285,852,881]" pageId="4" pageNumber="5" pagination="645 - 653" refId="ref16327" refString="Kwon D; Ferrara F and Taiti S 1992. Two new species of Laureola Barnard, 1960 from India and Vietnam (Crustacea, Oniscidea, Armadillidae). Revue suisse de Zoologie, 99: 645 - 653. DOI: https: // www. doi. org / 10.5962 / bhl. part. 79844" type="journal article" year="1992">Kwon et al., 1992</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFAADFC80FA16FCF8" author="Dalens H" box="[1294,1453,852,881]" pageId="4" pageNumber="5" pagination="93 - 100" refId="ref15028" refString="Dalens H 1998. Isopodes terrestres de Nouvelle-Caledonie. II. Sur une nouvelle espece du genre Pseudolaureola (Crustacea, Oniscidea). Zoosystema, 20: 93 - 100. https: // sciencepress. mnhn. fr / sites / default / files / articles / pdf / z 1998 n 1 a 4. pdf" type="journal article" year="1998">Dalens, 1998</bibRefCitation>
|
||
). However,
|
||
<taxonomicName id="4C404D7EFF02FFEEFC60FCAFFBD2FC11" authorityName="Kinahan" authorityYear="1859" box="[963,1129,891,920]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EAEFFF02FFEEFC60FCAFFBD2FC11" box="[963,1129,891,920]" italics="true" pageId="4" pageNumber="5">Acanthoniscus</emphasis>
|
||
</taxonomicName>
|
||
can be readily distinguished from all those other genera by the combination of its unique pleotelson-uropod pattern and its spine arrangement (see Diagnosis above).
|
||
</paragraph>
|
||
<paragraph id="8BFF36FDFF02FFEEFCD2FBE9FA01F979" blockId="4.[841,1469,1085,1971]" pageId="4" pageNumber="5">
|
||
<emphasis id="B934EAEFFF02FFEEFCD2FBE9FC67FBD3" box="[881,988,1085,1114]" italics="true" pageId="4" pageNumber="5">Remarks</emphasis>
|
||
.
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFC4FFBE9FB65FBD2" author="Schmalfuss H" box="[1004,1246,1085,1115]" pageId="4" pageNumber="5" refId="ref17197" refString="Schmalfuss H 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde, Serie A, 654: 341 p. https: // www. biodiversitylibrary. org / item / 293136 - page / 1 / mode / 1 up" type="book" year="2003">Schmalfuss (2003)</bibRefCitation>
|
||
tentatively listed
|
||
<taxonomicName id="4C404D7EFF02FFEEFCE8FBB0FC50FB08" authorityName="Kinahan" authorityYear="1859" box="[843,1003,1124,1153]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EAEFFF02FFEEFCE8FBB0FC50FB08" box="[843,1003,1124,1153]" italics="true" pageId="4" pageNumber="5">Acanthoniscus</emphasis>
|
||
</taxonomicName>
|
||
and its
|
||
<typeStatus id="54FB885FFF02FFEEFB9EFBB1FBC9FB08" box="[1085,1138,1125,1153]" pageId="4" pageNumber="5">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="4C404D7EFF02FFEEFB71FBB1FAF9FB08" authorityName="" (Gosse" authorityYear="1851" box="[1234,1346,1125,1153]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="spiniger">
|
||
<emphasis id="B934EAEFFF02FFEEFB71FBB1FAF9FB08" box="[1234,1346,1125,1153]" italics="true" pageId="4" pageNumber="5">A.spiniger</emphasis>
|
||
</taxonomicName>
|
||
, under the family
|
||
<taxonomicName id="4C404D7EFF02FFEEFC39FB5FFB84FB21" box="[922,1087,1163,1192]" family="Delatorreidae" pageId="4" pageNumber="5" rank="family">Delatorreidae</taxonomicName>
|
||
(it is still listed as such by
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFAC8FB5FFC6FFB46" author="Boyko CB & Bruce NL & Hadfield KA & Merrin KL & Ota Y & Poore GCB & Taiti S & Schotte M & Wilson GDF" pageId="4" pageNumber="5" refId="ref14575" refString="Boyko CB; Bruce NL; Hadfield KA; Merrin KL; Ota Y; Poore GCB; Taiti S; Schotte M and Wilson GDF (Eds.) (2008 onwards). World Marine, Freshwater and Terrestrial Isopod Crustaceans database. Acanthoniscus spiniger Kinahan, 1859. Available at https: // www. marinespecies. org / aphia. php? p = taxdetails & id = 258660. Accessed on 22 January 2022" type="journal volume" year="2008">Boyko et al., 2008</bibRefCitation>
|
||
, an on-line database), probably because several species in this family (genera
|
||
<taxonomicName id="4C404D7EFF02FFEEFB4EFB0DFC41FA94" authority="de Saussure, 1857" authorityName="de Saussure" authorityYear="1857" class="Malacostraca" family="Delatorreiidae" genus="Pseudarmadillo" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EAEFFF02FFEEFB4EFB0DFA27FB7F" box="[1261,1436,1241,1270]" italics="true" pageId="4" pageNumber="5">Pseudarmadillo</emphasis>
|
||
de Saussure, 1857
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C404D7EFF02FFEEFB93FAD4FBAFFACD" authority="Armas and Juarrero de Verona, 1999" authorityName="Armas and Juarrero de Verona" authorityYear="1999" class="Malacostraca" family="Delatorreiidae" genus="Cuzcodinella" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EAEFFF02FFEEFB93FAD4FB73FA94" box="[1072,1224,1280,1309]" italics="true" pageId="4" pageNumber="5">Cuzcodinella</emphasis>
|
||
Armas and Juarrero de Verona, 1999
|
||
</taxonomicName>
|
||
) have the body covered with long and sharp-pointed spines similar to
|
||
<taxonomicName id="4C404D7EFF02FFEEFAB7FA9AFA06FAE2" authorityName="Kinahan" authorityYear="1859" box="[1300,1469,1358,1387]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EAEFFF02FFEEFAB7FA9AFA06FAE2" box="[1300,1469,1358,1387]" italics="true" pageId="4" pageNumber="5">Acanthoniscus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFC20FAA1FB88FA1B" author="Vandel A" box="[899,1075,1397,1426]" pageId="4" pageNumber="5" pagination="35 - 37" refId="ref17916" refString="Vandel A 1973 a. Les isopodes terrestres et cavernicoles de l'ile de Cuba. p. 35 - 37. In: Orghidan T; Nunez Jimenez A; Botosaneanu L; Decou V; Negrea S and Vina Bayes N (Eds.), Resultats des Expeditions Biospeologiques Cubano- Roumaines a Cuba. I. Bucarest, Editura Academiei Republicii Socialiste Romania." type="book chapter" year="1973">Vandel, 1973a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFB9CFAA1FC37FA30" author="Armas LF de & Juarrero de Varona A" pageId="4" pageNumber="5" pagination="1 - 42" refId="ref14343" refString="Armas LF de and Juarrero de Varona A 1999. Sistematica de la familia Delatorreidae (Isopoda: Oniscidea) en Cuba. Avicennia, 10 / 11: 1 - 42. https: // www. biodiversitylibrary. org / item / 152480 - page / 7 / mode / 1 up" type="journal article" year="1999">Armas and Juarrero de Varona, 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFC36FA49FAE0FA30" author="Juarrero de Varona A & Armas LF de" box="[917,1371,1436,1465]" pageId="4" pageNumber="5" pagination="97 - 102" refId="ref16227" refString="Juarrero de Varona A and Armas LF de 2003 a. A new species of terrestrial isopod (Oniscidea: Delatorreidae) from Cuba. Avicennia, 16: 97 - 102." type="journal article" year="2003">Juarrero de Varona and Armas, 2003a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFAC7FA48FA0DFA30" author="Juarrero de Varona A & Armas LF de" box="[1380,1462,1436,1465]" pageId="4" pageNumber="5" pagination="21 - 26" refId="ref16260" refString="Juarrero de Varona A and Armas LF de 2003 b. Especie nueva de Pseudarmadillo (Isopoda: Oniscidea: Delatorreidae) de Cuba suroriental. Solenodon, 2: 21 - 26." type="journal article" year="2003">2003b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFCEAFA17FA91FA69" author="Armas LF de & Rodriguez-Cabrera TM" box="[841,1322,1475,1504]" pageId="4" pageNumber="5" pagination="1 - 6" refId="ref14397" refString="Armas LF de and Rodriguez-Cabrera TM 2016. Nueva especie de Cuzcodinella (Isopoda: Oniscidea: Delatorreidae) de la provincia de Holguin, Cuba. Boletin de la Sociedad Entomologica Aragonesa, 59: 1 - 6. https: // www. researchgate. net / publication / 340816201 _ una _ nueva _ especie _ de _ cuzcodinella _ isopoda _ oniscidea _ delatorreidae _ de _ la _ provincia _ de _ holguin _ cuba" type="journal article" year="2016">Armas and Rodríguez-Cabrera, 2016</bibRefCitation>
|
||
). However,
|
||
<taxonomicName id="4C404D7EFF02FFEEFCE8FA3EFC56F98E" box="[843,1005,1514,1543]" family="Delatorreidae" pageId="4" pageNumber="22" rank="family">Delatorreidae</taxonomicName>
|
||
strongly differ from
|
||
<taxonomicName id="4C404D7EFF02FFEEFB74FA3EFACFF98E" authorityName="Brandt" authorityYear="1831" box="[1239,1396,1514,1543]" class="Malacostraca" family="Armadillidae" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Armadillidae</taxonomicName>
|
||
in the cephalothorax (presence of antennal lobes and furrows in
|
||
<taxonomicName id="4C404D7EFF02FFEEFCCAF9ECFBA8F9DC" box="[873,1043,1592,1621]" family="Delatorreidae" pageId="4" pageNumber="22" rank="family">Delatorreidae</taxonomicName>
|
||
), number of eye ommatidia (six in
|
||
<taxonomicName id="4C404D7EFF02FFEEFCEEF98AFC43F9F2" box="[845,1016,1630,1659]" family="Delatorreidae" pageId="4" pageNumber="22" rank="family">Delatorreidae</taxonomicName>
|
||
), and the uropod pattern (exopodite very short and inserted on the inner corner of the posterior margin of the sympodite in
|
||
<taxonomicName id="4C404D7EFF02FFEEFAAAF978FA09F940" box="[1289,1458,1708,1737]" family="Delatorreidae" pageId="4" pageNumber="22" rank="family">Delatorreidae</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFCF1F900FB43F978" author="Armas LF de & Juarrero de Varona A" box="[850,1272,1747,1777]" pageId="4" pageNumber="5" pagination="1 - 42" refId="ref14343" refString="Armas LF de and Juarrero de Varona A 1999. Sistematica de la familia Delatorreidae (Isopoda: Oniscidea) en Cuba. Avicennia, 10 / 11: 1 - 42. https: // www. biodiversitylibrary. org / item / 152480 - page / 7 / mode / 1 up" type="journal article" year="1999">Armas and Juarrero de Varona, 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFAA2F907FA16F979" author="Schmidt C" box="[1281,1453,1747,1776]" pageId="4" pageNumber="5" pagination="3 - 179" refId="ref17365" refString="Schmidt C 2003. Contribution to the phylogenetic system of the Crinocheta (Crustacea, Isopoda). Part 2. (Oniscoidea to Armadillidiidae). Mitteilungen aus dem Museum fur Naturkunde in Berlin, Zoologische Reihe, 79 (1): 3 - 179. DOI: https: // www. doi. org / 10.1002 / mmnz. 20030790102" type="journal article" year="2003">Schmidt, 2003</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFF36FDFF02FFEFFCD2F92EFD39FD9B" blockId="4.[841,1469,1085,1971]" lastBlockId="5.[179,806,151,803]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
The general pattern of the uropod in
|
||
<taxonomicName id="4C404D7EFF02FFEEFABBF92EFA01F89E" authorityName="Kinahan" authorityYear="1859" box="[1304,1466,1786,1815]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EAEFFF02FFEEFABBF92EFA01F89E" box="[1304,1466,1786,1815]" italics="true" pageId="4" pageNumber="5">Acanthoniscus</emphasis>
|
||
</taxonomicName>
|
||
led
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFCD8F8F5FA95F8B7" author="Schmidt C & Leistikow, A" box="[891,1326,1825,1854]" pageId="4" pageNumber="5" pagination="1 - 118" refId="ref17528" refString="Schmidt C and Leistikow, A 2004. Catalogue of genera of the terrestrial Isopoda (Crustacea: Isopoda: Oniscidea). Steenstrupia, 28: 1 - 118." type="journal article" year="2004">Schmidt and Leistikow (2004: 4)</bibRefCitation>
|
||
to suggest that it might belong to the heterogeneous family
|
||
<taxonomicName id="4C404D7EFF02FFEEFCE8F8BBFBB6F805" authorityName="Verhoeff" authorityYear="1938" box="[843,1037,1903,1932]" class="Malacostraca" family="Scleropactidae" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Scleropactidae</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFB84F8BBFB40F805" author="Schmidt C" box="[1063,1275,1903,1932]" pageId="4" pageNumber="5" pagination="275 - 352" refId="ref17298" refString="Schmidt C 2002. Contribution to the phylogenetic system of the Crinocheta (Crustacea, Isopoda). Part 1. (Olibrinidae to Scyphacidae s. str.). Mitteilungen aus dem Museum fur Naturkunde Berlin, Zoologische Reihe, 78 (2): 275 - 352. DOI: https: // www. doi. org / 10.1002 / mmnz. 20020780207" type="journal article" year="2002">Schmidt, 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFAB2F8BBFAE3F805" author="Schmidt C" box="[1297,1368,1903,1932]" pageId="4" pageNumber="5" pagination="3 - 179" refId="ref17365" refString="Schmidt C 2003. Contribution to the phylogenetic system of the Crinocheta (Crustacea, Isopoda). Part 2. (Oniscoidea to Armadillidiidae). Mitteilungen aus dem Museum fur Naturkunde in Berlin, Zoologische Reihe, 79 (1): 3 - 179. DOI: https: // www. doi. org / 10.1002 / mmnz. 20030790102" type="journal article" year="2003">2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFACCF8BBFA0FF805" author="Schmidt C" box="[1391,1460,1903,1932]" pageId="4" pageNumber="5" refId="ref17429" refString="Schmidt C 2007. Revision of the Neotropical Scleropactidae (Crustacea: Oniscidea). ZoologicalJournaloftheLinneanSociety, 151 (Suppl. 1): 1 - 339. DOI: https: // www. doi. org / 10.1111 / j. 1096 - 3642.2007.00286. x" type="url" year="2007">2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF02FFEEFCE8F842FC2BF83A" author="Schmidt C" box="[843,912,1942,1971]" pageId="4" pageNumber="5" pagination="191 - 226" refId="ref17477" refString="Schmidt C 2008. Phylogeny of the Terrestrial Isopoda (Oniscidea): a Review. Arthropod Systematics & Phylogeny, 66 (2): 191 - 226. https: // www. zobodat. at / pdf / Arthropod- Systematics-Phylogeny _ 66 _ 0191 - 0226. pdf" type="journal article" year="2008">2008</bibRefCitation>
|
||
). The flagellum of the second antenna may be either 2-jointed or 3-jointed in
|
||
<taxonomicName id="4C404D7EFF03FFEFFDCDFF43FC9EFF3D" authorityName="Verhoeff" authorityYear="1938" box="[622,805,151,180]" class="Malacostraca" family="Scleropactidae" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Scleropactidae</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFD14B0CFF03FFEFFF1DFF6AFECDFF52" author="Schmidt C" box="[190,374,190,219]" pageId="5" pageNumber="6" pagination="3 - 179" refId="ref17365" refString="Schmidt C 2003. Contribution to the phylogenetic system of the Crinocheta (Crustacea, Isopoda). Part 2. (Oniscoidea to Armadillidiidae). Mitteilungen aus dem Museum fur Naturkunde in Berlin, Zoologische Reihe, 79 (1): 3 - 179. DOI: https: // www. doi. org / 10.1002 / mmnz. 20030790102" type="journal article" year="2003">Schmidt, 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF03FFEFFE23FF6AFE7AFF52" author="Schmidt C" box="[384,449,190,219]" pageId="5" pageNumber="6" refId="ref17429" refString="Schmidt C 2007. Revision of the Neotropical Scleropactidae (Crustacea: Oniscidea). ZoologicalJournaloftheLinneanSociety, 151 (Suppl. 1): 1 - 339. DOI: https: // www. doi. org / 10.1111 / j. 1096 - 3642.2007.00286. x" type="url" year="2007">2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B0CFF03FFEFFE68FF6AFDB5FF52" author="Schmidt C" box="[459,526,190,219]" pageId="5" pageNumber="6" pagination="191 - 226" refId="ref17477" refString="Schmidt C 2008. Phylogeny of the Terrestrial Isopoda (Oniscidea): a Review. Arthropod Systematics & Phylogeny, 66 (2): 191 - 226. https: // www. zobodat. at / pdf / Arthropod- Systematics-Phylogeny _ 66 _ 0191 - 0226. pdf" type="journal article" year="2008">2008</bibRefCitation>
|
||
). In
|
||
<taxonomicName id="4C404D7EFF03FFEFFDE3FF6AFD49FF52" authorityName="Verhoeff" authorityYear="1938" box="[576,754,190,219]" class="Malacostraca" family="Scleropactidae" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Scleropactidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C404D7EFF03FFEFFF16FF31FEE3FE8B" authorityName="Kinahan" authorityYear="1859" box="[181,344,229,258]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EAEFFF03FFEFFF16FF31FEE3FE8B" box="[181,344,229,258]" italics="true" pageId="5" pageNumber="6">Acanthoniscus</emphasis>
|
||
</taxonomicName>
|
||
the uropod exopodite is inserted on the medial margin of the sympodite and widely surpasses its posterior margin(see Discussion for other divergent forms in
|
||
<taxonomicName id="4C404D7EFF03FFEFFE83FE8EFE7DFEFE" authorityName="Brandt" authorityYear="1831" box="[288,454,346,375]" class="Malacostraca" family="Armadillidae" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Armadillidae</taxonomicName>
|
||
;
|
||
<figureCitation id="137B2A78FF03FFEFFE73FE8EFDADFEFE" box="[464,534,346,375]" captionStart="Figure 8" captionStartId="15.[181,252,1523,1546]" captionTargetBox="[192,1460,173,1475]" captionTargetId="figure-248@15.[181,1466,151,1499]" captionTargetPageId="15" captionText="Figure 8. Selected body parts of Acanthoniscus for comparison with other taxa within Armadillidae. Cubaris murina Brandt, 1833, showing the ground pattern in Armadillidae (A–E): A, Posterior region of the body in dorsal view; B, posterior region of the body in ventral view; C, pleotelson in dorsal view; D, uropod in dorsal view; E, middle region of the sternite 7 of a male showing the bilobed caudal process on the posterior margin, which is considered an autapomorphy of Armadillidae.Acanthoniscus richardsonae sp. nov.(F–H): F, pleotelson in dorsal view; G, uropod in dorsal view; H, middle region of the sternite 7 of a male.Drawings of the pleotelson and uropods in dorsal views of some morphologically divergent taxa within Armadillidae. I, Echinodillo cavaticus Green, 1963; J, Echinodillo montanus Jackson, 1933; K, Laureola vietnamensis Kwon, Ferrara and Taiti, 1992; L, Laureola dubia Schmalfuss and Ferrara,1983; M, Schismadillo rouxi Verhoeff,1926; N, Merulanella wahrbergi Verhoeff,1926; O, Nesodillo incisus Verhoeff, 1926. Drawings modified from Verhoeff (1926) (M–O), Jackson (1933) (J), Green (1963) (I), Schmalfuss and Ferrara (1983) (L), and Kwon et al. (1992) (K). Abbreviations: endopodite (en), exopodite (ex), sympodite (sy)." figureDoi="http://doi.org/10.5281/zenodo.10716782" httpUri="https://zenodo.org/record/10716782/files/figure.png" pageId="5" pageNumber="6">Fig. 8</figureCitation>
|
||
), but the pleotelson in
|
||
<taxonomicName id="4C404D7EFF03FFEFFF15FE55FED9FE17" authorityName="Verhoeff" authorityYear="1938" box="[182,354,385,414]" class="Malacostraca" family="Scleropactidae" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Scleropactidae</taxonomicName>
|
||
is subtriangular and much shorter than the uropod sympodite, with the uropod endopodite widely surpassing the distal margins of both the uropod sympodite and the pleotelson.
|
||
</paragraph>
|
||
<paragraph id="8BFF36FDFF03FFEFFF78FDC9FC9FFCAA" blockId="5.[179,806,151,803]" pageId="5" pageNumber="6">
|
||
In a key to the genera of the
|
||
<taxonomicName id="4C404D7EFF03FFEFFDBBFDC8FEFFFDE9" authority=", Richardson (1901: 562)" authorityName="Richardson" authorityPageNumber="562" authorityYear="1901" class="Malacostraca" family="Oniscidae" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">
|
||
Oniscidae,
|
||
<bibRefCitation id="EFD14B0CFF03FFEFFD38FDC8FEFFFDE9" author="Richardson H" pageId="5" pageNumber="6" pagination="493 - 579" refId="ref16936" refString="Richardson H 1901. Key to the isopods of the Atlantic coast of North America with descriptions of new and little known species. Proceedings of the United States National Museum, 23: 493 - 579. DOI: https: // www. doi. org / 10.5479 / si. 00963801.23 - 1222.493" type="journal article" year="1901">Richardson (1901: 562)</bibRefCitation>
|
||
</taxonomicName>
|
||
included
|
||
<taxonomicName id="4C404D7EFF03FFEFFE62FD97FDD1FDE9" authorityName="Kinahan" authorityYear="1859" box="[449,618,579,608]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EAEFFF03FFEFFE62FD97FDD1FDE9" box="[449,618,579,608]" italics="true" pageId="5" pageNumber="6">Acanthoniscus</emphasis>
|
||
</taxonomicName>
|
||
as having “the flagellum of the external antennae quadri-articulate.” However, this information must be erroneous, since the f lagellum of the second antenna in this genus was unknown until the present contribution, as stated also by
|
||
<bibRefCitation id="EFD14B0CFF03FFEFFF76FCD2FE76FCAA" author="Kinahan JR" box="[213,461,774,803]" pageId="5" pageNumber="6" pagination="125 - 135" refId="ref16294" refString="Kinahan JR 1859. On the genus Platyarthrus (Brandt); with notices of allied undescribed genera. Natural History Review, 6: 125 - 135, pl. 14." type="journal article" year="1859">Kinahan (1859: 198)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFD14B0CFF03FFEFFDA1FCD2FC9BFCAA" author="Richardson H" box="[514,800,774,803]" pageId="5" pageNumber="6" pagination="431 - 434" refId="ref17042" refString="Richardson H 1909. The isopod crustacean Acanthoniscus spiniger Kinahan redescribed. Proceedings of the United States National Museum, 36: 431 - 434. DOI: https: // www. doi. org / 10.5479 / si. 00963801.36 - 1678.431" type="journal article" year="1909">Richardson (1909: 432)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |