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<mods:title id="EC8A2066E8F38E1637670642C6701E2F">A new brachypterous Filchneria and two new generic records for China, Arcynopteryx and Pictetiella (Plecoptera: Perlodidae)</mods:title>
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<treatment id="03E8C610FF80FFDF88FDFF45FB9D0D8E" ID-DOI="http://doi.org/10.5281/zenodo.4759689" ID-GBIF-Taxon="159628615" ID-Zenodo-Dep="4759689" LSID="urn:lsid:plazi:treatment:03E8C610FF80FFDF88FDFF45FB9D0D8E" httpUri="http://treatment.plazi.org/id/03E8C610FF80FFDF88FDFF45FB9D0D8E" lastPageNumber="523" pageId="12" pageNumber="523">
<subSubSection id="C35B248DFF80FFDF88FDFF45FD610FD9" box="[151,717,153,180]" pageId="12" pageNumber="523" type="nomenclature">
<paragraph id="8BFE7706FF80FFDF88FDFF45FD610FD9" blockId="12.[151,717,153,216]" box="[151,717,153,180]" pageId="12" pageNumber="523">
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<taxonomicName id="4C410C85FF80FFDF88FDFF45FD610FD9" ID-CoL="4HQX7" authority="Zwick &amp; Levanidova, 1971" authorityName="Zwick &amp; Levanidova" authorityYear="1971" box="[151,717,153,180]" class="Insecta" family="Perlodidae" genus="Pictetiella" kingdom="Animalia" order="Plecoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="species" species="asiatica">
<emphasis id="B935AB14FF80FFDF88FDFF45FD610FD9" bold="true" box="[151,717,153,180]" pageId="12" pageNumber="523">
<emphasis id="B935AB14FF80FFDF88FDFF45FED80FD9" bold="true" box="[151,372,153,180]" italics="true" pageId="12" pageNumber="523">Pictetiella asiatica</emphasis>
<bibRefCitation id="EFD00AF7FF80FFDF8911FF45FD610FD9" author="Zwick, P. &amp; Levanidova, I. M." box="[379,717,153,179]" pageId="12" pageNumber="523" pagination="849 - 869" refId="ref6910" refString="Zwick, P. &amp; Levanidova, I. M. (1971) On the stonefly fauna (Plecoptera) of the Soviet Far East. Entomologicheskoe Obozrenie, 50 (4), 849 - 869." type="journal article" year="1971">Zwick &amp; Levanidova, 1971</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="C35B248DFF80FFDF88FDFF61FEE10FBD" box="[151,333,189,216]" pageId="12" pageNumber="523" type="description">
<paragraph id="8BFE7706FF80FFDF88FDFF61FEE10FBD" blockId="12.[151,717,153,216]" box="[151,333,189,216]" pageId="12" pageNumber="523">
<figureCitation id="137A6B83FF80FFDF88FDFF61FF5F0FB2" box="[151,243,189,216]" captionStart="FIGURE 15" captionStartId="10.[151,250,1970,1995]" captionTargetBox="[167,1421,1323,1950]" captionTargetId="figure@10.[167,1421,1323,1950]" captionTargetPageId="10" captionText="FIGURE 15. Locality of Arcynopteryx polaris Klapálek, 1912 and Pictetiella asiatica Zwick &amp; Levanidova, 1971 in Jilin Province of northeastern China." figureDoi="http://doi.org/10.5281/zenodo.3471232" httpUri="https://zenodo.org/record/3471232/files/figure.png" pageId="12" pageNumber="523">Figs. 15</figureCitation>
,
<figureCitation id="137A6B83FF80FFDF8895FF61FEE40FBD" box="[255,328,189,215]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStartId-0="12.[151,250,1355,1380]" captionStartId-1="12.[151,250,1972,1997]" captionTargetBox-0="[157,1436,781,1286]" captionTargetBox-1="[151,1435,1481,1907]" captionTargetId-0="figure@12.[151,1436,781,1286]" captionTargetId-1="figure@12.[151,1435,1481,1907]" captionTargetPageId-0="12" captionTargetPageId-1="12" captionText-0="FIGURE 18. Pictetiella asiatica Zwick &amp; Levanidova, 1971, male. A. habitus, dorsal view; B. habitus, ventral view; C. head and pronotum, dorsal view; D. head and thorax, ventral view; E. left forewing, dorsal view; F. left hindwing, dorsal view. Scale bar = 1 mm." captionText-1="FIGURE 19. Pictetiella asiatica Zwick &amp; Levanidova, 1971, male. A. abdomen, dorsal view; B. abdomen, ventral view; C. terminalia, dorsal view; D, terminalia, ventral view. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3471238" figureDoi-1="http://doi.org/10.5281/zenodo.3471240" httpUri-0="https://zenodo.org/record/3471238/files/figure.png" httpUri-1="https://zenodo.org/record/3471240/files/figure.png" pageId="12" pageNumber="523">1819</figureCitation>
.
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<taxonomicName id="4C410C85FF80FFDF88FDFEDEFD030E70" ID-CoL="4HQX7" authority="Zwick &amp; Levanidova, 1971: 853" authorityName="Zwick &amp; Levanidova" authorityPageNumber="853" authorityYear="1971" box="[151,687,257,282]" class="Insecta" family="Perlodidae" genus="Pictetiella" kingdom="Animalia" order="Plecoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="species" species="asiatica">
<emphasis id="B935AB14FF80FFDF88FDFEDEFEFA0E73" box="[151,342,258,282]" italics="true" pageId="12" pageNumber="523">Pictetiella asiatica</emphasis>
<treatmentCitation id="0AE05117FF80FFDF8934FEDDFD030E70" author="Zwick, P. &amp; Levanidova, I. M." box="[350,687,257,282]" page="853" pageId="12" pageNumber="523" year="1971">
<bibRefCitation id="EFD00AF7FF80FFDF8934FEDDFD030E70" author="Zwick, P. &amp; Levanidova, I. M." box="[350,687,257,282]" pageId="12" pageNumber="523" pagination="849 - 869" refId="ref6910" refString="Zwick, P. &amp; Levanidova, I. M. (1971) On the stonefly fauna (Plecoptera) of the Soviet Far East. Entomologicheskoe Obozrenie, 50 (4), 849 - 869." type="journal article" year="1971">Zwick &amp; Levanidova, 1971: 853</bibRefCitation>
</treatmentCitation>
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;
<treatmentCitation id="0AE05117FF80FFDF8AD0FEDDFCDE0E70" author="Zwick, P." box="[698,882,257,282]" page="238" pageId="12" pageNumber="523" year="1973">
<bibRefCitation id="EFD00AF7FF80FFDF8AD0FEDDFCDE0E70" author="Zwick, P." box="[698,882,257,282]" pageId="12" pageNumber="523" pagination="1 - 465" refId="ref6686" refString="Zwick, P. (1973) Insecta Plecoptera. Phylogenetisches System und Katalog. Das Tierreich, 94, 1 - 465." type="journal article" year="1973">Zwick, 1973: 238</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AE05117FF80FFDF8B17FEDDFA860E70" author="Zhiltzova, L. A. &amp; Zapekina-Dulkeit, J. I." box="[893,1322,257,282]" page="183" pageId="12" pageNumber="523" year="1986">
<bibRefCitation id="EFD00AF7FF80FFDF8B17FEDDFA860E70" author="Zhiltzova, L. A. &amp; Zapekina-Dulkeit, J. I." box="[893,1322,257,282]" pageId="12" pageNumber="523" pagination="172 - 234" refId="ref6649" refString="Zhiltzova, L. A. &amp; Zapekina-Dulkeit, J. I. (1986) 10. Otrjad Plecoptera. Vesnyanki. Opredelitej nasekomih Dalnego Vostoka SSSR, Leningrad, 1, 172 - 234." type="journal article" year="1986">Zhiltzova &amp; Zapekina-Dulkeit, 1986: 183</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AE05117FF80FFDF8D5FFEDDFEBA0E52" author="Zhiltzova, L. A." page="7" pageId="12" pageNumber="523" year="1995">
<bibRefCitation id="EFD00AF7FF80FFDF8D5FFEDDFEBA0E52" author="Zhiltzova, L. A." pageId="12" pageNumber="523" refId="ref6611" refString="Zhiltzova, L. A. (1995) Catalogue of type specimens in the collection of the Zoological Institute, Russian Academy of Sciences. Plecoptera. Zoologicheskii Institut, St. Petersburg, 37 pp." type="book" year="1995">Zhiltzova, 1995: 7</bibRefCitation>
</treatmentCitation>
; Teslenko, 2009: 698;
<treatmentCitation id="0AE05117FF80FFDF8990FEFCFC920E52" author="Teslenko, V. A. &amp; Zhiltzova, L. A." box="[506,830,288,313]" page="32" pageId="12" pageNumber="523" year="2009">
<bibRefCitation id="EFD00AF7FF80FFDF8990FEFCFC920E52" author="Teslenko, V. A. &amp; Zhiltzova, L. A." box="[506,830,288,313]" pageId="12" pageNumber="523" refId="ref6357" refString="Teslenko, V. A. &amp; Zhiltzova, L. A. (2009) Keys to the stoneflies (Insecta, Plecoptera) of Russia and adjacent countries. Imagines and larvae. Dalnauka, Vladivostok, 382 pp." type="book" year="2009">Teslenko &amp; Zhiltzova, 2009: 32</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AE05117FF80FFDF8B22FEFCFBF20E52" author="Judson, S. W. &amp; Nelson, C. R." box="[840,1118,288,313]" page="44" pageId="12" pageNumber="523" year="2012">
<bibRefCitation id="EFD00AF7FF80FFDF8B22FEFCFBF20E52" author="Judson, S. W. &amp; Nelson, C. R." box="[840,1118,288,313]" pageId="12" pageNumber="523" pagination="1 - 118" refId="ref5582" refString="Judson, S. W. &amp; Nelson, C. R. (2012) A guide to Mongolian stoneflies (Insecta: Plecoptera). Zootaxa, 3541 (1), 1 - 118. https: // doi. org / 10.11646 / zootaxa. 3541.1.1" type="journal article" year="2012">Judson &amp; Nelson, 2012: 44</bibRefCitation>
</treatmentCitation>
; Murányi, Hwang, Jeon &amp; Seo, 2016: 113.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35B248DFF80FFDF88FDFE59FDBA0D45" pageId="12" pageNumber="523" type="materials_examined">
<paragraph id="8BFE7706FF80FFDF88FDFE59FDBA0D45" blockId="12.[151,1437,389,740]" pageId="12" pageNumber="523">
<emphasis id="B935AB14FF80FFDF88FDFE59FE2C0ECA" bold="true" box="[151,384,389,416]" pageId="12" pageNumber="523">Material examined.</emphasis>
One male,
<collectingCountry id="F3563796FF80FFDF8A6AFE59FDEB0EF5" box="[512,583,389,415]" name="China" pageId="12" pageNumber="523">China</collectingCountry>
:
<collectingRegion id="4985B9E4FF80FFDF8A38FE59FD430EF5" box="[594,751,389,415]" country="China" name="Jilin" pageId="12" pageNumber="523">Jilin Province</collectingRegion>
, Changbaishan National Natural Reserve, Pond of the Visitor Center (
<figureCitation id="137A6B83FF80FFDF8898FE75FEE90EAE" box="[242,325,425,452]" captionStart="FIGURE 15" captionStartId="10.[151,250,1970,1995]" captionTargetBox="[167,1421,1323,1950]" captionTargetId="figure@10.[167,1421,1323,1950]" captionTargetPageId="10" captionText="FIGURE 15. Locality of Arcynopteryx polaris Klapálek, 1912 and Pictetiella asiatica Zwick &amp; Levanidova, 1971 in Jilin Province of northeastern China." figureDoi="http://doi.org/10.5281/zenodo.3471232" httpUri="https://zenodo.org/record/3471232/files/figure.png" pageId="12" pageNumber="523">Fig. 15</figureCitation>
),
<geoCoordinate id="EE7511C1FF80FFDF8931FE75FE4C0EAE" box="[347,480,425,452]" degrees="42.07111" direction="north" orientation="latitude" pageId="12" pageNumber="523" precision="1" value="42.07111">42.07111 N</geoCoordinate>
,
<geoCoordinate id="EE7511C1FF80FFDF8986FE75FDD20EAE" box="[492,638,425,452]" degrees="128.06358" direction="east" orientation="longitude" pageId="12" pageNumber="523" precision="1" value="128.06358">128.06358 E</geoCoordinate>
,
<quantity id="4CB9DAE3FF80FFDF8AE1FE75FD4E0EAE" box="[651,738,425,452]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="523" unit="m" value="1700.0">1700 m</quantity>
,
<date id="FFFF51C6FF80FFDF8A85FE76FC220EA9" box="[751,910,425,452]" pageId="12" pageNumber="523" value="2011-06-29">June 29, 2011</date>
, J. Morse, J. Murphy, Z. Diao (donated by Shuang Qiu in 2019, ICJUST). One male,
<collectingRegion id="4985B9E4FF80FFDF8A3CFE11FD5F0E8D" box="[598,755,461,487]" country="China" name="Jilin" pageId="12" pageNumber="523">Jilin Province</collectingRegion>
, Antu County, Erdaobaihe Town, Tributary of Jianshui River (
<figureCitation id="137A6B83FF80FFDF88CAFE2DFF5D0D66" box="[160,241,497,524]" captionStart="FIGURE 15" captionStartId="10.[151,250,1970,1995]" captionTargetBox="[167,1421,1323,1950]" captionTargetId="figure@10.[167,1421,1323,1950]" captionTargetPageId="10" captionText="FIGURE 15. Locality of Arcynopteryx polaris Klapálek, 1912 and Pictetiella asiatica Zwick &amp; Levanidova, 1971 in Jilin Province of northeastern China." figureDoi="http://doi.org/10.5281/zenodo.3471232" httpUri="https://zenodo.org/record/3471232/files/figure.png" pageId="12" pageNumber="523">Fig. 15</figureCitation>
),
<geoCoordinate id="EE7511C1FF80FFDF896FFE2DFE260D66" box="[261,394,497,524]" degrees="42.38248" direction="north" orientation="latitude" pageId="12" pageNumber="523" precision="1" value="42.38248">42.38248 N</geoCoordinate>
,
<geoCoordinate id="EE7511C1FF80FFDF89FEFE2DFD860D66" box="[404,554,497,524]" degrees="128. 13431" direction="east" orientation="longitude" pageId="12" pageNumber="523" precision="1" value="128.13431">128. 13431 E</geoCoordinate>
,
<quantity id="4CB9DAE3FF80FFDF8A5FFE2DFDD10D66" box="[565,637,497,524]" metricMagnitude="2" metricUnit="m" metricValue="7.57" pageId="12" pageNumber="523" unit="m" value="757.0">757 m</quantity>
,
<date id="FFFF51C6FF80FFDF8AEDFE2EFC8F0D61" box="[647,803,497,524]" pageId="12" pageNumber="523" value="2011-06-28">June 28, 2011</date>
, J. Morse, J. Murphy, Yun-Jun Yan, Zhi-Cai Xie (donated by Shuang Qiu in 2019, ICJUST).
</paragraph>
</subSubSection>
<subSubSection id="C35B248DFF80FFDF88ADFDE6FB9D0D8E" pageId="12" pageNumber="523" type="discussion">
<paragraph id="8BFE7706FF80FFDF88ADFDE6FB9D0D8E" blockId="12.[151,1437,389,740]" pageId="12" pageNumber="523">
<emphasis id="B935AB14FF80FFDF88ADFDE6FE970D3E" bold="true" box="[199,315,570,596]" pageId="12" pageNumber="523">Remarks.</emphasis>
Body (
<figureCitation id="137A6B83FF80FFDF89FEFDE5FE590D39" box="[404,501,569,595]" captionStart="FIGURE 18" captionStartId="12.[151,250,1355,1380]" captionTargetBox="[157,1436,781,1286]" captionTargetId="figure@12.[151,1436,781,1286]" captionTargetPageId="12" captionText="FIGURE 18. Pictetiella asiatica Zwick &amp; Levanidova, 1971, male. A. habitus, dorsal view; B. habitus, ventral view; C. head and pronotum, dorsal view; D. head and thorax, ventral view; E. left forewing, dorsal view; F. left hindwing, dorsal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3471238" httpUri="https://zenodo.org/record/3471238/files/figure.png" pageId="12" pageNumber="523">Figs. 18</figureCitation>
,
<figureCitation id="137A6B83FF80FFDF8A6EFDE5FD8D0D39" box="[516,545,569,595]" captionStart="FIGURE 19" captionStartId="12.[151,250,1972,1997]" captionTargetBox="[151,1435,1481,1907]" captionTargetId="figure@12.[151,1435,1481,1907]" captionTargetPageId="12" captionText="FIGURE 19. Pictetiella asiatica Zwick &amp; Levanidova, 1971, male. A. abdomen, dorsal view; B. abdomen, ventral view; C. terminalia, dorsal view; D, terminalia, ventral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3471240" httpUri="https://zenodo.org/record/3471240/files/figure.png" pageId="12" pageNumber="523">19</figureCitation>
) length ca.
<quantity id="4CB9DAE3FF80FFDF8AC7FDE5FCAC0D3E" box="[685,768,569,596]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="523" unit="mm" value="15.0">15 mm</quantity>
(excluding antennae and cerci). This species is originally described from
<collectingCountry id="F3563796FF80FFDF8927FD81FE360D1D" box="[333,410,605,631]" name="Russia" pageId="12" pageNumber="523">Russia</collectingCountry>
, and is now only known from
<collectingCountry id="F3563796FF80FFDF8B69FD81FCDF0D1D" box="[771,883,605,631]" name="Mongolia" pageId="12" pageNumber="523">Mongolia</collectingCountry>
,
<collectingCountry id="F3563796FF80FFDF8BEAFD81FBBE0D12" box="[896,1042,605,632]" name="North Korea" pageId="12" pageNumber="523">North Korea</collectingCountry>
, and
<collectingCountry id="F3563796FF80FFDF8C39FD81FB320D1D" box="[1107,1182,605,631]" name="Russia" pageId="12" pageNumber="523">Russia</collectingCountry>
(
<bibRefCitation id="EFD00AF7FF80FFDF8CC5FD81FA3D0D1D" author="DeWalt, R. E. &amp; Maehr, M. D. &amp; Neu-Becker, U. &amp; Stueber, G." box="[1199,1425,605,632]" pageId="12" pageNumber="523" refId="ref5460" refString="DeWalt, R. E., Maehr, M. D., Neu-Becker, U. &amp; Stueber, G. (2019) Plecoptera Species File online. Available from: http: // Plecoptera. SpeciesFile. org / (accessed 10 August 2019) https: // doi. org / 10.3390 / insects 10040099" type="book" year="2019">
DeWalt
<emphasis id="B935AB14FF80FFDF8D65FD82FAEF0D1D" box="[1295,1347,605,631]" italics="true" pageId="12" pageNumber="523">et al</emphasis>
. 2019
</bibRefCitation>
). Morphology of the specimens studied herein is almost identical with that of
<bibRefCitation id="EFD00AF7FF80FFDF8B8DFD5DFB750DF6" author="Muranyi, D. &amp; Hwang, J. M. &amp; Jeon, M. J." box="[999,1241,641,668]" pageId="12" pageNumber="523" pagination="111 - 121" refId="ref5932" refString="Muranyi, D., Hwang, J. M., Jeon, M. J. &amp; Seo-gu, G. D. (2016) New species records of systellognathan stoneflies (PLecoptera: Systellognatha) for the fauna of the Korean Peninsula. Opuscula Zoologica, Budapest, 47 (2), 111 - 121. https: // doi. org / 10.18348 / opzool. 2016.2.111" type="journal article" year="2016">
Murányi
<emphasis id="B935AB14FF80FFDF8C24FD5EFBD30DF1" box="[1102,1151,641,667]" italics="true" pageId="12" pageNumber="523">et al</emphasis>
. (2016)
</bibRefCitation>
, who redescribed the North Korean specimens also from the Changbai (Baekdu) ranges. Discovery of
<taxonomicName id="4C410C85FF80FFDF8C32FD79FB670DD5" authorityName="Illies" authorityYear="1966" box="[1112,1227,677,703]" class="Insecta" family="Perlodidae" genus="Pictetiella" kingdom="Animalia" order="Plecoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B935AB14FF80FFDF8C32FD79FB670DD5" box="[1112,1227,677,703]" italics="true" pageId="12" pageNumber="523">Pictetiella</emphasis>
</taxonomicName>
from northeastern
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fills the gap of the generic distribution between
<collectingCountry id="F3563796FF80FFDF8A90FD15FCCB0D89" box="[762,871,713,739]" name="Mongolia" pageId="12" pageNumber="523">Mongolia</collectingCountry>
and
<collectingCountry id="F3563796FF80FFDF8BF7FD15FB810D8E" box="[925,1069,713,740]" name="North Korea" pageId="12" pageNumber="523">North Korea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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