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<document id="075E9D898EEFAFD53C9DEFECAB707C17" ID="10.11646/zootaxa.4268.2.2SLASH11053" ID-CLB-Dataset="33044" ID-DOI="10.11646/zootaxa.4268.2.2" ID-GBIF-Dataset="00fa23ba-e0e6-4a1a-959c-7192b134caf2" ID-ISSN="1175-5326" ID-Zenodo-Dep="580190" ID-ZooBank="61A285A2-F1E8-43F1-AEA8-9E64D3450108" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1494920846434" checkinUser="plazi" docAuthor="Bilton, David T." docDate="2017" docId="03E72E67E10FFF88FF6BB48FFDBA4E93" docLanguage="en" docName="zootaxa.4268.2.2SLASH11053.pdf" docOrigin="Zootaxa 4268 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mesoceration chasmum Bilton, 2017, sp. nov." docType="treatment" docVersion="8" lastPageNumber="206" masterDocId="FFDE561FE104FF87FFFCB10CFFD84E46" masterDocTitle="Water beetles from the Bokkeveld Plateau: a semi-arid hotspot of freshwater biodiversity in the Northern Cape of South Africa" masterLastPageNumber="214" masterPageNumber="191" pageNumber="202" updateTime="1698439582737" updateUser="ExternalLinkService">
<mods:mods id="9395D4F8AF2A23F88FB4808018924C66" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4C675F7C1FAD47CF5EB781D980606E77">
<mods:title id="52686EA4480D19FFED92F0E0240BB7A8">Water beetles from the Bokkeveld Plateau: a semi-arid hotspot of freshwater biodiversity in the Northern Cape of South Africa</mods:title>
</mods:titleInfo>
<mods:name id="BCDB196617BABAE45D38F8E1A6B339E1" type="personal">
<mods:role id="B194CB94086EF0CE64420280B05FEC0B">
<mods:roleTerm id="30DCC1C087085E69F471C60C6F1A348A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="231AD231D4CBE24043D5A48BE6412022">Bilton, David T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="7A30531CF146D752F8CCCE421E872504">text</mods:typeOfResource>
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<mods:title id="BEAFF07215E058227EC9109AC8BE7612">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="01421C3551CAF61623D42BEDB7207604">
<mods:date id="03EED19FBA50B0A1A53A1A1C51ADA661">2017</mods:date>
<mods:detail id="70F242F8911EE2989A6CAA1606ED57FE" type="volume">
<mods:number id="DCE294EB841666C1F28289D2BCDA76B1">4268</mods:number>
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<mods:detail id="D99FA59CA3C66B40C2CFDE31F9AD29E0" type="issue">
<mods:number id="E5B8B7994E189F5B8B16FE71BFFAA2F7">2</mods:number>
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<mods:start id="6924E742770C81DA14814EA8F28EE67C">191</mods:start>
<mods:end id="DCB31CE5FA865551678DA9BC4EF7450D">214</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="40B1F240E8E9D15322F9379FA2495D96">journal article</mods:classification>
<mods:identifier id="C33EE40FFE2BEA077B89C3616313935E" type="CLB-Dataset">33044</mods:identifier>
<mods:identifier id="A38FDE451C7638EA79BDE0E2C1D1FBE9" type="DOI">10.11646/zootaxa.4268.2.2</mods:identifier>
<mods:identifier id="C2F5B698EFA4B9953F867083F30D1B74" type="GBIF-Dataset">00fa23ba-e0e6-4a1a-959c-7192b134caf2</mods:identifier>
<mods:identifier id="8CF01A2C6A09C89B572F7AC852E76447" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="ECB508DB6351AB2D770A5D2434DB4EF5" type="Zenodo-Dep">580190</mods:identifier>
<mods:identifier id="A2D9FFF8385E5F4345BA7C48ECF9AF20" type="ZooBank">61A285A2-F1E8-43F1-AEA8-9E64D3450108</mods:identifier>
</mods:mods>
<treatment id="03E72E67E10FFF88FF6BB48FFDBA4E93" ID-DOI="http://doi.org/10.5281/zenodo.6030437" ID-GBIF-Taxon="130809871" ID-Zenodo-Dep="6030437" LSID="urn:lsid:plazi:treatment:03E72E67E10FFF88FF6BB48FFDBA4E93" httpUri="http://treatment.plazi.org/id/03E72E67E10FFF88FF6BB48FFDBA4E93" lastPageId="15" lastPageNumber="206" pageId="11" pageNumber="202">
<subSubSection id="C354CCFAE10FFF8CFF6BB48FFE814BF9" pageId="11" pageNumber="202" type="nomenclature">
<paragraph id="8BF19F71E10FFF8CFF6BB48FFE814BF9" blockId="11.[151,537,1411,1471]" pageId="11" pageNumber="202">
<taxonomicName id="4C4EE4F2E10FFF8CFF6BB48FFE614BDB" ID-CoL="4222R" box="[151,441,1411,1437]" class="Insecta" family="Hydraenidae" genus="Mesoceration" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="202" phylum="Arthropoda" rank="species" species="chasmum" status="sp. nov.">
<emphasis id="B93A4363E10FFF8CFF6BB48FFE614BDB" bold="true" box="[151,441,1411,1437]" italics="true" pageId="11" pageNumber="202">Mesoceration chasmum</emphasis>
</taxonomicName>
<emphasis id="B93A4363E10FFF8CFE43B488FDC14BDB" bold="true" box="[447,537,1412,1437]" pageId="11" pageNumber="202">
<taxonomicNameLabel id="A209FE18E10FFF8CFE43B488FDC14BDB" box="[447,537,1412,1437]" pageId="11" pageNumber="202" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="137583F4E10FFF8CFF63B4AAFF3F4BF9" box="[159,231,1446,1471]" captionStart="FIGURE 2" captionStartId="5.[151,250,1952,1974]" captionTargetBox="[183,1385,208,1893]" captionTargetId="figure@5.[174,1405,193,1918]" captionTargetPageId="5" captionText="FIGURE 2. Habitus of hydraenid holotypes. A) Parasthetops porcellus sp. nov.; B) Mesoceration castaneum sp. nov.; C) M. chasmum sp. nov.; D) M. sabulosum sp. nov." httpUri="https://zenodo.org/record/580192/files/figure.png" pageId="11" pageNumber="202">Figs 2</figureCitation>
,
<figureCitation id="137583F4E10FFF8CFF0FB4ABFEDB4BF9" box="[243,259,1447,1471]" captionStart="FIGURE 3" captionStartId="6.[151,250,1969,1991]" captionTargetBox="[205,1345,208,1904]" captionTargetId="figure@6.[188,1390,193,1930]" captionTargetPageId="6" captionText="FIGURE 3. Aedeagi of hydraenid holotypes, ventral and lateral views. A) Parasthetops porcellus sp. nov.; B) Mesoceration castaneum sp. nov.; C) M. chasmum sp. nov.; D) M. sabulosum sp. nov." httpUri="https://zenodo.org/record/580193/files/figure.png" pageId="11" pageNumber="202">3</figureCitation>
,
<figureCitation id="137583F4E10FFF8CFEF3B4AAFEC54BF9" box="[271,285,1446,1471]" captionStart="FIGURE 5" captionStartId="8.[151,250,1907,1929]" captionTargetBox="[228,1360,193,1876]" captionTargetId="figure@8.[228,1360,193,1876]" captionTargetPageId="8" captionText="FIGURE 5. Habitats on the Bokkeveld Plateau. A) Avontuur Reserve, margins of Kromvlei (Site 8); B) Avontuur Reserve, pools in drying stream (Site 7); C) Oorlogskloof Reserve, wet rocks &amp; stream below Driefontein Waterfall (Sites 9 &amp; 10; stream below large boulders in photo); D) Oorlogskloof Reserve, wet rocks below Driefontein Waterfall (Site 9 — with Andy Foggo). Photos D. T. Bilton." httpUri="https://zenodo.org/record/580195/files/figure.png" pageId="11" pageNumber="202">5</figureCitation>
&amp;
<figureCitation id="137583F4E10FFF8CFEBDB4AAFE884BF9" box="[321,336,1446,1471]" captionStart="FIGURE 6" captionStartId="9.[151,250,1953,1975]" captionTargetBox="[177,1410,197,1932]" captionTargetId="figure@9.[177,1410,193,1932]" captionTargetPageId="9" captionText="FIGURE 6. Habitats on the Bokkeveld Plateau. A – B) Avontuur Reserve, stream below Fynbos Cottage (Site 5); B) C) Oorlogskloof Reserve, Oorlogskloof River below Driefontein Waterfall (Site 11) Photos D. T. Bilton." httpUri="https://zenodo.org/record/580196/files/figure.png" pageId="11" pageNumber="202">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C354CCFAE10FFF8CFF6BB4E2FDFD4945" pageId="11" pageNumber="202" type="materials_examined">
<paragraph id="8BF19F71E10FFF8CFF6BB4E2FC3A486D" blockId="11.[151,1437,1518,2011]" pageId="11" pageNumber="202">
<materialsCitation id="3B26952CE10FFF8CFF6BB4E2FC3A486D" ID-GBIF-Occurrence="1503125145" country="South Africa" county="Bokkeveld Plateau" elevation="640" latitude="-31.511625" location="Driefontein Waterfall" longLatPrecision="1" longitude="19.001667" municipality="Oorlogskloof Canyon" pageId="11" pageNumber="202" specimenCount="1" stateProvince="Northern Cape" typeStatus="holotype">
<emphasis id="B93A4363E10FFF8CFF6BB4E2FEF64841" bold="true" box="[151,302,1518,1543]" pageId="11" pageNumber="202">
<typeStatus id="54F521D3E10FFF8CFF6BB4E2FF0A4841" box="[151,210,1518,1543]" pageId="11" pageNumber="202">Type</typeStatus>
locality
</emphasis>
.
<emphasis id="B93A4363E10FFF8CFEC1B4E2FE0F4841" bold="true" box="[317,471,1518,1543]" pageId="11" pageNumber="202">
<collectingCountry id="F359DFE1E10FFF8CFEC1B4E2FE0F4841" box="[317,471,1518,1543]" name="South Africa" pageId="11" pageNumber="202">South Africa</collectingCountry>
</emphasis>
,
<collectingRegion id="498A5193E10FFF8CFE1BB4E2FD484841" box="[487,656,1518,1543]" country="South Africa" name="Northern Cape" pageId="11" pageNumber="202">Northern Cape</collectingRegion>
,
<collectingCounty id="6290E7FDE10FFF8CFD60B4E2FCAB4841" box="[668,883,1518,1543]" pageId="11" pageNumber="202">Bokkeveld Plateau</collectingCounty>
,
<collectingMunicipality id="6B95050BE10FFF8CFC83B4E2FBAE4841" box="[895,1142,1518,1543]" pageId="11" pageNumber="202">Oorlogskloof Canyon</collectingMunicipality>
, stream amongst boulders below
<location id="8E91C9AAE10FFF8CFF1EB71EFE0C486D" LSID="urn:lsid:plazi:treatment:03E72E67E10FFF88FF6BB48FFDBA4E93:8E91C9AAE10FFF8CFF1EB71EFE0C486D" box="[226,468,1554,1579]" country="South Africa" county="Bokkeveld Plateau" latitude="-31.511625" longLatPrecision="1" longitude="19.001667" municipality="Oorlogskloof Canyon" name="Driefontein Waterfall" pageId="11" pageNumber="202" stateProvince="Northern Cape">Driefontein Waterfall</location>
,
<quantity id="4CB63294E10FFF8CFE1CB71EFDF1486C" box="[480,553,1554,1579]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="11" pageNumber="202" unit="m" value="640.0">
<elevation id="00637842E10FFF8CFE1CB71EFDF1486C" box="[480,553,1554,1579]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="11" pageNumber="202" unit="m" value="640.0">640 m</elevation>
</quantity>
,
<geoCoordinate id="EE7AF9B6E10FFF8CFDC9B71FFD11486D" box="[565,713,1554,1579]" direction="south" orientation="latitude" pageId="11" pageNumber="202" precision="1" value="-31.511625">31 30 41.85S</geoCoordinate>
<geoCoordinate id="EE7AF9B6E10FFF8CFD2CB71FFCBE486D" box="[720,870,1554,1579]" direction="east" orientation="longitude" pageId="11" pageNumber="202" precision="1" value="19.001667">19 0 6 58.98E</geoCoordinate>
(
<figureCitation id="137583F4E10FFF8CFC81B71EFC67486D" box="[893,959,1554,1579]" captionStart="FIGURE 5" captionStartId="8.[151,250,1907,1929]" captionTargetBox="[228,1360,193,1876]" captionTargetId="figure@8.[228,1360,193,1876]" captionTargetPageId="8" captionText="FIGURE 5. Habitats on the Bokkeveld Plateau. A) Avontuur Reserve, margins of Kromvlei (Site 8); B) Avontuur Reserve, pools in drying stream (Site 7); C) Oorlogskloof Reserve, wet rocks &amp; stream below Driefontein Waterfall (Sites 9 &amp; 10; stream below large boulders in photo); D) Oorlogskloof Reserve, wet rocks below Driefontein Waterfall (Site 9 — with Andy Foggo). Photos D. T. Bilton." httpUri="https://zenodo.org/record/580195/files/figure.png" pageId="11" pageNumber="202">Fig. 5</figureCitation>
C).
</materialsCitation>
</paragraph>
<paragraph id="8BF19F71E10FFF8CFF3BB73AFDAC48D1" blockId="11.[151,1437,1518,2011]" pageId="11" pageNumber="202">
<materialsCitation id="3B26952CE10FFF8CFF3BB73AFDA848D1" ID-GBIF-Occurrence="1503125132" collectingDate="2015-10-07" collectionCode="AMG" collectorName="D T Bilton" country="South Africa" elevation="640" latitude="-31.511625" location="Oorlogskloof Canyon-stream" longLatPrecision="1" longitude="19.001667" municipality="Driefontein Waterfall" pageId="11" pageNumber="202" specimenCount="3" typeStatus="holotype">
<emphasis id="B93A4363E10FFF8CFF3BB73AFEAE4809" bold="true" box="[199,374,1590,1615]" pageId="11" pageNumber="202">
<typeStatus id="54F521D3E10FFF8CFF3BB73AFEDA4809" box="[199,258,1590,1615]" pageId="11" pageNumber="202">Type</typeStatus>
material
</emphasis>
.
<emphasis id="B93A4363E10FFF8CFE70B73AFE214809" bold="true" box="[396,505,1590,1615]" pageId="11" pageNumber="202">
<typeStatus id="54F521D3E10FFF8CFE70B73AFE214809" box="[396,505,1590,1615]" pageId="11" pageNumber="202" type="holotype">Holotype</typeStatus>
</emphasis>
(male): “
<date id="FFF0B9B1E10FFF8CFD8FB73AFD014809" box="[627,729,1590,1615]" pageId="11" pageNumber="202" value="2015-10-07">
<collectingDate id="EFB44059E10FFF8CFD8FB73AFD014809" box="[627,729,1590,1615]" pageId="11" pageNumber="202" value="2015-10-07">7/x/2015</collectingDate>
</date>
<collectingCountry id="F359DFE1E10FFF8CFD14B73AFC584809" box="[744,896,1590,1615]" name="South Africa" pageId="11" pageNumber="202">South Africa</collectingCountry>
NC//
<location id="8E91C9AAE10FFF8CFC2AB73AFAEF4809" LSID="urn:lsid:plazi:treatment:03E72E67E10FFF88FF6BB48FFDBA4E93:8E91C9AAE10FFF8CFC2AB73AFAEF4809" box="[982,1335,1590,1615]" country="South Africa" latitude="-31.511625" longLatPrecision="1" longitude="19.001667" municipality="Driefontein Waterfall" name="Oorlogskloof Canyon-stream" pageId="11" pageNumber="202">Oorlogskloof Canyon—stream</location>
// below
<collectingMunicipality id="6B95050BE10FFF8CFF6BB756FE5F4835" box="[151,391,1626,1651]" pageId="11" pageNumber="202">Driefontein Waterfall</collectingMunicipality>
//
<quantity id="4CB63294E10FFF8CFE5CB756FE304834" box="[416,488,1626,1651]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="11" pageNumber="202" unit="m" value="640.0">
<elevation id="00637842E10FFF8CFE5CB756FE304834" box="[416,488,1626,1651]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="11" pageNumber="202" unit="m" value="640.0">640 m</elevation>
</quantity>
<geoCoordinate id="EE7AF9B6E10FFF8CFE0CB757FD504835" box="[496,648,1626,1651]" direction="south" orientation="latitude" pageId="11" pageNumber="202" precision="1" value="-31.511625">31 30 41.85S</geoCoordinate>
<geoCoordinate id="EE7AF9B6E10FFF8CFD6CB757FCF04835" box="[656,808,1626,1651]" direction="east" orientation="longitude" pageId="11" pageNumber="202" precision="1" value="19.001667">19 0 6 58.98E</geoCoordinate>
//
<collectorName id="26BBFAA7E10FFF8CFCBDB757FC654835" box="[833,957,1626,1651]" pageId="11" pageNumber="202">D T Bilton</collectorName>
leg.” (genitalia extracted and mounted on same card) and red
<typeStatus id="54F521D3E10FFF8CFE8EB772FE0C48D1" box="[370,468,1662,1687]" pageId="11" pageNumber="202" type="holotype">holotype</typeStatus>
label (
<collectionCode id="ED5F07B4E10FFF8CFDDCB773FDB048D1" box="[544,616,1663,1687]" country="South Africa" httpUri="http://grbio.org/cool/b9qz-7eka" name="Albany Museum" pageId="11" pageNumber="202">AMG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF19F71E10FFF8CFF3BB7AEFDFD4945" blockId="11.[151,1437,1518,2011]" pageId="11" pageNumber="202">
<materialsCitation id="3B26952CE10FFF8CFF3BB7AEFD1748FD" ID-GBIF-Occurrence="1503125142" box="[199,719,1698,1723]" collectingDate="2015-10-07" collectionCode="AMG" collectorName="D T Bilton" country="South Africa" elevation="640" latitude="-31.511625" location="Oorlogskloof Canyon-stream" longLatPrecision="1" longitude="19.001667" municipality="Driefontein Waterfall" pageId="11" pageNumber="202" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B93A4363E10FFF8CFF3BB7AEFEE748FD" bold="true" box="[199,319,1698,1723]" pageId="11" pageNumber="202">
<typeStatus id="54F521D3E10FFF8CFF3BB7AEFEE748FD" box="[199,319,1698,1723]" pageId="11" pageNumber="202" type="paratype">Paratypes</typeStatus>
</emphasis>
(2):
<specimenCount id="9D4854F8E10FFF8CFE7FB7AFFE6B48FC" box="[387,435,1699,1723]" pageId="11" pageNumber="202" type="male">1 ♂</specimenCount>
same data as holotype
</materialsCitation>
;
<materialsCitation id="3B26952CE10FFF8CFD1DB7AFFDFA4945" ID-GBIF-Occurrence="1503125140" collectingDate="2015-10-07" collectionCode="CDTB, AMG" collectorName="D T Bilton &amp; All" country="South Africa" county="Driefontein Waterfall" elevation="492" latitude="-31.514156" location="Oorlogskloof" longLatPrecision="1" longitude="19.001667" municipality="River" pageId="11" pageNumber="202" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D4854F8E10FFF8CFD1DB7AFFCCA48FC" box="[737,786,1699,1723]" pageId="11" pageNumber="202" type="male">1 ♂</specimenCount>
“
<date id="FFF0B9B1E10FFF8CFCD6B7AEFC4848FD" box="[810,912,1698,1723]" pageId="11" pageNumber="202" value="2015-10-07">
<collectingDate id="EFB44059E10FFF8CFCD6B7AEFC4848FD" box="[810,912,1698,1723]" pageId="11" pageNumber="202" value="2015-10-07">7/x/2015</collectingDate>
</date>
<collectingCountry id="F359DFE1E10FFF8CFC61B7AEFBEC48FD" box="[925,1076,1698,1723]" name="South Africa" pageId="11" pageNumber="202">South Africa</collectingCountry>
NC//
<location id="8E91C9AAE10FFF8CFB7AB7AEFAC348FD" LSID="urn:lsid:plazi:treatment:03E72E67E10FFF88FF6BB48FFDBA4E93:8E91C9AAE10FFF8CFB7AB7AEFAC348FD" box="[1158,1307,1698,1723]" country="South Africa" county="Driefontein Waterfall" latitude="-31.514156" longLatPrecision="1" longitude="19.001667" municipality="River" name="Oorlogskloof" pageId="11" pageNumber="202">Oorlogskloof</location>
Canyon—
<location id="8E91C9AAE10FFF8CFF6BB7CAFEF34899" LSID="urn:lsid:plazi:treatment:03E72E67E10FFF88FF6BB48FFDBA4E93:8E91C9AAE10FFF8CFF6BB7CAFEF34899" box="[151,299,1734,1759]" country="South Africa" county="Driefontein Waterfall" latitude="-31.514156" longLatPrecision="1" longitude="19.001667" municipality="River" name="Oorlogskloof" pageId="11" pageNumber="202">Oorlogskloof</location>
//
<collectingMunicipality id="6B95050BE10FFF8CFEB8B7CAFE5A4899" box="[324,386,1734,1759]" pageId="11" pageNumber="202">River</collectingMunicipality>
below
<collectingCounty id="6290E7FDE10FFF8CFE2AB7CAFD1F4899" box="[470,711,1734,1759]" pageId="11" pageNumber="202">Driefontein Waterfall</collectingCounty>
//
<quantity id="4CB63294E10FFF8CFD23B7CBFCFF4898" box="[735,807,1735,1759]" metricMagnitude="2" metricUnit="m" metricValue="4.92" pageId="11" pageNumber="202" unit="m" value="492.0">
<elevation id="00637842E10FFF8CFD23B7CBFCFF4898" box="[735,807,1735,1759]" metricMagnitude="2" metricUnit="m" metricValue="4.92" pageId="11" pageNumber="202" unit="m" value="492.0">492 m</elevation>
</quantity>
<geoCoordinate id="EE7AF9B6E10FFF8CFCD3B7CBFC1D4899" box="[815,965,1734,1759]" direction="south" orientation="latitude" pageId="11" pageNumber="202" precision="1" value="-31.514156">31 30 50.96S</geoCoordinate>
<geoCoordinate id="EE7AF9B6E10FFF8CFC31B7CBFBBD4899" box="[973,1125,1734,1759]" direction="east" orientation="longitude" pageId="11" pageNumber="202" precision="1" value="19.001667">19 0 6 47.86E</geoCoordinate>
//
<collectorName id="26BBFAA7E10FFF8CFB81B7CBFB214899" box="[1149,1273,1734,1759]" pageId="11" pageNumber="202">D T Bilton</collectorName>
leg.”
<collectorName id="26BBFAA7E10FFF8CFAC3B7CAFABB4899" box="[1343,1379,1734,1759]" pageId="11" pageNumber="202">All</collectorName>
with red
<typeStatus id="54F521D3E10FFF8CFF3EB7E7FEFA4945" box="[194,290,1771,1795]" pageId="11" pageNumber="202" type="paratype">paratype</typeStatus>
labels (
<collectionCode id="ED5F07B4E10FFF8CFE85B7E7FE104945" box="[377,456,1771,1795]" pageId="11" pageNumber="202">CDTB</collectionCode>
,
<collectionCode id="ED5F07B4E10FFF8CFE2FB7E7FDC14945" box="[467,537,1771,1795]" country="South Africa" httpUri="http://grbio.org/cool/b9qz-7eka" name="Albany Museum" pageId="11" pageNumber="202">AMG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C354CCFAE10FFF8BFF3BB602FE46486B" lastPageId="12" lastPageNumber="203" pageId="11" pageNumber="202" type="description">
<paragraph id="8BF19F71E10FFF8CFF3BB602FDFD490C" blockId="11.[151,1437,1518,2011]" pageId="11" pageNumber="202">
<emphasis id="B93A4363E10FFF8CFF3BB602FE894961" bold="true" box="[199,337,1806,1831]" pageId="11" pageNumber="202">Description</emphasis>
. Size:
<typeStatus id="54F521D3E10FFF8CFE63B602FDD44961" box="[415,524,1806,1831]" pageId="11" pageNumber="202" type="holotype">Holotype</typeStatus>
: BL
<quantity id="4CB63294E10FFF8CFDBEB602FD724960" box="[578,682,1806,1831]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="11" pageNumber="202" unit="mm" value="1.75">1.75 mm</quantity>
; EL 1.00 mm; EW
<quantity id="4CB63294E10FFF8CFC7AB602FC364960" box="[902,1006,1806,1831]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="11" pageNumber="202" unit="mm" value="0.65">0.65 mm</quantity>
.
<typeStatus id="54F521D3E10FFF8CFC04B603FBB24961" box="[1016,1130,1807,1831]" pageId="11" pageNumber="202" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D4854F8E10FFF8CFB89B603FB534960" box="[1141,1163,1807,1830]" pageId="11" pageNumber="202" type="male">♂</specimenCount>
s BL
<quantity id="4CB63294E10FFF8CFB34B603FAB64960" box="[1224,1390,1806,1831]" metricMagnitude="-3" metricUnit="m" metricValue="1.725" metricValueMax="1.75" metricValueMin="1.7" pageId="11" pageNumber="202" unit="mm" value="1.725" valueMax="1.75" valueMin="1.7">1.70–1.75 mm</quantity>
; EL 1.00–
<quantity id="4CB63294E10FFF8CFF2BB63EFEE5490C" box="[215,317,1842,1867]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="11" pageNumber="202" unit="mm" value="1.05">1.05 mm</quantity>
; EW
<quantity id="4CB63294E10FFF8CFE87B63EFDF9490C" box="[379,545,1842,1867]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="11" pageNumber="202" unit="mm" value="0.625" valueMax="0.65" valueMin="0.6">0.60–0.65 mm</quantity>
.
</paragraph>
<paragraph id="8BF19F71E10FFF8CFF3BB65AFADE499D" blockId="11.[151,1437,1518,2011]" pageId="11" pageNumber="202">
Colour: Dorsum (
<figureCitation id="137583F4E10FFF8CFE68B65AFE024929" box="[404,474,1878,1903]" captionStart="FIGURE 2" captionStartId="5.[151,250,1952,1974]" captionTargetBox="[183,1385,208,1893]" captionTargetId="figure@5.[174,1405,193,1918]" captionTargetPageId="5" captionText="FIGURE 2. Habitus of hydraenid holotypes. A) Parasthetops porcellus sp. nov.; B) Mesoceration castaneum sp. nov.; C) M. chasmum sp. nov.; D) M. sabulosum sp. nov." httpUri="https://zenodo.org/record/580192/files/figure.png" pageId="11" pageNumber="202">Fig. 2</figureCitation>
C) with head pitchy brown to black; ocelli paler. Pronotum and elytra pale brown, infuscated on pronotal disc and around elytral suture. Maxillary palpi pale yellow, apical 0.75 of terminal segment infuscated. Legs pale yellow, apex of terminal tarsal segments narrowly infuscated. Venter testaceous brown, meso and metaventrite and mentum pitchy black. Much of venter silvery due to dense hydrofuge vestiture.
</paragraph>
<paragraph id="8BF19F71E108FF8BFF3BB19BFA5D4FEB" blockId="12.[151,1437,151,2018]" pageId="12" pageNumber="203">Head: Labrum quadrate, widening anteriorly. Rounded apicolaterally, with v-shaped apicomedian emargination extending 0.5 of length. Sides of apicomedian emargination narrowly raised. Surface weakly shining, with strong isodiametric microreticulation and fine punctures. Long, yellow recumbent setae along lateral and anterior margins. Clypeus weakly shining, with isodiametric microreticulation and sparse, fine punctures bearing fine decumbent setae. Frontoclypeal suture weakly arcuate. Frons weakly shining on centre, with isodiametric microreticulation and sparse, medium punctures bearing white decumbent setae. Anteocellar sulci broad, opening anteriorly, rugosely microreticulate and punctate, both in sulic and between them and compound eyes. Ocelli shining. Compound eyes moderately large, occupying 0.6 lateral margin of head, 15 ommatidia in longest series.</paragraph>
<paragraph id="8BF19F71E108FF8BFF3BB0B4FB8D4CEE" blockId="12.[151,1437,151,2018]" pageId="12" pageNumber="203">Pronotum: Cordate, transverse, broadest just behind middle. Anterior margin arcuate over central 0.5, with narrow hyaline border; posterior margin weakly bisinuate around centre. Anterior angles broadly rounded; posterior angles obtusely rounded. Lateral margins sinuated from widest point to base; broadly marginated and moderately crenulated. Upper surface weakly shining, with shallow, isodiametric microreticulation and sparse, medium punctures bearing white decumbent setae. With 10 distinct fovea as follows: Anterior median fovea elongate; posterior median fovea circular. Anterior admedian foveae rounded; posterior admedian foveae elongate oval, orientated anterolaterally. Anterior and adlateral foveae small, rounded, pit-like.</paragraph>
<paragraph id="8BF19F71E108FF8BFF3BB3B8FD384DC6" blockId="12.[151,1437,151,2018]" pageId="12" pageNumber="203">Elytra: Elongate oval, broadest at middle. Subparallel over median 0.4, then attenuated to broadly rounded apices, emarginated around suture. narrowly explanate at sides. Punctures of elytral series evident to apex; larger in anterior 0.5, smaller behind. Punctures bearing decumbent setae, each seta reaching next puncture in series. Coarsely granulated between punctures. Discal series striate impressed. Series 1 and 2 confluent 0.5 from apex; series 5 and 6 separate throughout. Intervals weakly shining, finely granulated, granules with fine, white decumbent setae. Interval 8 without evident carinae.</paragraph>
<paragraph id="8BF19F71E108FF8BFF3BB281FAE94DE2" blockId="12.[151,1437,151,2018]" box="[199,1329,908,933]" pageId="12" pageNumber="203">Legs: Apical protarsomere with two stout ventral setae; basal three protarsomeres with suction setae.</paragraph>
<paragraph id="8BF19F71E108FF8BFF3BB2BCFC9E4BDA" blockId="12.[151,1437,151,2018]" pageId="12" pageNumber="203">Venter: Mentum shining, with isodiametric to elongate microreticulation; more strongly impressed towards lateral margins and sparse, fine punctures bearing decumbent setae. Submentum shining, with transverse microreticulation and sparse, fine punctures bearing decumbent setae. Genae shining, with transverse microreticulation. Gula shining, with transverse microreticulation; meshes smaller than on genae. Prosternum with weak central ridge; surface rugulose, with dense hydrofuge vestiture. Pronotal hypomeron broad, weakly shining, with strong, slightly elongate microreticulation. Meso and metaventrite with dense hydrofuge vestiture. Mesoventral plaques weak, low, forming inverted Y, stem 0.5 length of arms. Weak admedian and adlateral ridges also present. Metaventrite with broad, shallow median fovea over posterior 0.4; fovea with hydrofuge vestiture. Elytral pseudepipleurs broad, shining, without evident microreticulation; extending to apex of abdominal ventrite 5. Elytral epipleurs shining, narrow, ridge-like. Abdominal ventrites 1–4 with dense hydrofuge vestiture throughout. Abdominal ventrite 5 with semicircular glabrous patch occupying apical 0.3; glabrous area with shallow, transverse microreticulation and irregular transverse row of fine punctures bearing long, golden decumbent setae. Abdominal ventrite 6 glabrous, shining, with medium transverse microreticulation and scattered fine punctures bearing golden decumbent setae close to apex.</paragraph>
<paragraph id="8BF19F71E108FF8BFF3BB4ABFD6C484E" blockId="12.[151,1437,151,2018]" pageId="12" pageNumber="203">
Aedeagus: Elongate, with parameres attaching close to base. Main piece somwhat arcuate in ventral view, curved ventrally in lateral view, with strong, stout tooth on left hand side in ventral view. Distal lobe short, strongly curved over apical tooth of distal lobe (
<figureCitation id="137583F4E108FF8BFDB3B4E3FD49484E" box="[591,657,1519,1544]" captionStart="FIGURE 3" captionStartId="6.[151,250,1969,1991]" captionTargetBox="[205,1345,208,1904]" captionTargetId="figure@6.[188,1390,193,1930]" captionTargetPageId="6" captionText="FIGURE 3. Aedeagi of hydraenid holotypes, ventral and lateral views. A) Parasthetops porcellus sp. nov.; B) Mesoceration castaneum sp. nov.; C) M. chasmum sp. nov.; D) M. sabulosum sp. nov." httpUri="https://zenodo.org/record/580193/files/figure.png" pageId="12" pageNumber="203">Fig. 3</figureCitation>
C).
</paragraph>
<paragraph id="8BF19F71E108FF8BFF3BB718FE46486B" blockId="12.[151,1437,151,2018]" box="[199,414,1556,1581]" pageId="12" pageNumber="203">Female: Unknown.</paragraph>
</subSubSection>
<subSubSection id="C354CCFAE108FF8BFF3BB736FCC8494C" pageId="12" pageNumber="203" type="diagnosis">
<paragraph id="8BF19F71E108FF8BFF3BB736FCC8494C" blockId="12.[151,1437,151,2018]" pageId="12" pageNumber="203">
<emphasis id="B93A4363E108FF8BFF3BB736FE1E4815" bold="true" box="[199,454,1594,1619]" pageId="12" pageNumber="203">Differential diagnosis</emphasis>
. The entirely separated 5th and 6th elytral series and the structure of the aedeagus would place this species within the
<emphasis id="B93A4363E108FF8BFE23B753FD894831" box="[479,593,1631,1655]" italics="true" pageId="12" pageNumber="203">barriotum</emphasis>
group
<emphasis id="B93A4363E108FF8BFD5FB76CFD394831" box="[675,737,1632,1655]" italics="true" pageId="12" pageNumber="203">sensu</emphasis>
<bibRefCitation id="EFDFE280E108FF8BFD15B752FC4B4831" author="Perkins" box="[745,915,1630,1655]" pageId="12" pageNumber="203" refString="Perkins, P. D. (2008) New species and new collection records of Prosthetopine water beetles from southern Africa (Coleoptera: Hydraenidae). Zootaxa, 1864, 1 - 124." type="journal article" year="2008">Perkins (2008)</bibRefCitation>
. It shares with
<emphasis id="B93A4363E108FF8BFBBEB76CFB154830" box="[1090,1229,1631,1655]" italics="true" pageId="12" pageNumber="203">M. integrum</emphasis>
<bibRefCitation id="EFDFE280E108FF8BFB2AB752FAA94831" author="Perkins" box="[1238,1393,1630,1655]" pageId="12" pageNumber="203" refString="Perkins, P. D. (2008) New species and new collection records of Prosthetopine water beetles from southern Africa (Coleoptera: Hydraenidae). Zootaxa, 1864, 1 - 124." type="journal article" year="2008">Perkins, 2008</bibRefCitation>
the complete absence of a carina on the 8th elytral interval, something which is always present, to some extent, in other members of the group. It differs from
<emphasis id="B93A4363E108FF8BFDB5B7A6FD0C4887" box="[585,724,1706,1729]" italics="true" pageId="12" pageNumber="203">M. integrum</emphasis>
in the more elongate habitus, paler dorsal colouration, and the presence of distinct microreticulation on the pronotal reliefs. The aedeagi of the two species are very similar in basic structure, despite these strong external differences.
</paragraph>
</subSubSection>
<subSubSection id="C354CCFAE108FF8BFF3BB619FEE049A4" pageId="12" pageNumber="203" type="distribution">
<paragraph id="8BF19F71E108FF8BFF3BB619FEE049A4" blockId="12.[151,1437,151,2018]" pageId="12" pageNumber="203">
<emphasis id="B93A4363E108FF8BFF3BB619FE814968" bold="true" box="[199,345,1813,1838]" pageId="12" pageNumber="203">Distribution</emphasis>
. To date known only from the two adjacent localities (Sites 10 &amp; 11,
<figureCitation id="137583F4E108FF8BFB3CB619FAD24968" box="[1216,1290,1813,1838]" captionStart="FIGURE 5" captionStartId="8.[151,250,1907,1929]" captionTargetBox="[228,1360,193,1876]" captionTargetId="figure@8.[228,1360,193,1876]" captionTargetPageId="8" captionText="FIGURE 5. Habitats on the Bokkeveld Plateau. A) Avontuur Reserve, margins of Kromvlei (Site 8); B) Avontuur Reserve, pools in drying stream (Site 7); C) Oorlogskloof Reserve, wet rocks &amp; stream below Driefontein Waterfall (Sites 9 &amp; 10; stream below large boulders in photo); D) Oorlogskloof Reserve, wet rocks below Driefontein Waterfall (Site 9 — with Andy Foggo). Photos D. T. Bilton." httpUri="https://zenodo.org/record/580195/files/figure.png" pageId="12" pageNumber="203">Figs 5</figureCitation>
C &amp; 6C) in Oorlogskloof Canyon on the Bokkeveld Plateau. In the
<typeStatus id="54F521D3E108FF8BFCF6B636FCE24914" box="[778,826,1850,1874]" pageId="12" pageNumber="203">type</typeStatus>
locality specimens were netted from gravel and debris between large sandstone boulders in a heavily shaded stream below a waterfall, where they were the only water beetle recorded. A single male was also taken in the Oorlogskloof River, upstream of where the
<typeStatus id="54F521D3E108FF8BFB20B68EFAD449DC" box="[1244,1292,1922,1946]" pageId="12" pageNumber="203">type</typeStatus>
stream joins the main river. This specimen was netted from gravel below a boulder, together with abundant
<emphasis id="B93A4363E108FF8BFB0DB6AAFAC849FB" box="[1265,1296,1958,1981]" italics="true" pageId="12" pageNumber="203">M.</emphasis>
<taxonomicName id="4C4EE4F2E108FF8BFAE5B6A9FEED49A4" authority="Perkins, 2008" authorityName="Perkins" authorityYear="2008" class="Insecta" family="Hydraenidae" genus="Mesoceration" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="203" phylum="Arthropoda" rank="species" species="longipennis">
<emphasis id="B93A4363E108FF8BFAE5B6A9FA4449FB" box="[1305,1436,1957,1981]" italics="true" pageId="12" pageNumber="203">longipennis</emphasis>
<bibRefCitation id="EFDFE280E108FF8BFF6BB6C5FEED49A4" author="Perkins" box="[151,309,1993,2018]" pageId="12" pageNumber="203" refString="Perkins, P. D. (2008) New species and new collection records of Prosthetopine water beetles from southern Africa (Coleoptera: Hydraenidae). Zootaxa, 1864, 1 - 124." type="journal article" year="2008">Perkins, 2008</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF31CFF9E109FF8AFF6BB675FB4449E8" httpUri="https://zenodo.org/record/580197/files/figure.png" pageId="13" pageNumber="204" targetBox="[151,1436,196,1892]" targetPageId="13">
<paragraph id="8BF19F71E109FF8AFF6BB675FB4449E8" blockId="13.[151,1436,1913,1966]" pageId="13" pageNumber="204">
<emphasis id="B93A4363E109FF8AFF6BB675FECB49C8" bold="true" box="[151,275,1913,1935]" pageId="13" pageNumber="204">FIGURE 7.</emphasis>
Habitats on the Bokkeveld Plateau. A) Oorlogskloof Canyon, approach to Papkuilsfontein Waterfall (Site 12); B) Oorlogskloof Canyon, pools in stream above Papkuilsfontein Waterfall (Site 12). Photos D. T. Bilton.
</paragraph>
</caption>
<caption id="DF31CFF9E10AFF89FF6BB6ADFCD149B2" httpUri="https://zenodo.org/record/580198/files/figure.png" pageId="14" pageNumber="205" targetBox="[177,1410,197,1932]" targetPageId="14">
<paragraph id="8BF19F71E10AFF89FF6BB6ADFCD149B2" blockId="14.[151,1435,1953,2036]" pageId="14" pageNumber="205">
<emphasis id="B93A4363E10AFF89FF6BB6ADFECD49F0" bold="true" box="[151,277,1953,1975]" pageId="14" pageNumber="205">FIGURE 8.</emphasis>
Habitats on the Bokkeveld Plateau. A) Oorlogskloof Canyon, pools in stream above Papkuilsfontein Waterfall (Site 12); B) Oorlogskloof Reserve, stream pools close to Groot Tuin entrance (Site 13); C) Oorlogskloof Reserve, spring pool at head of stream at Groot Tuin (Site 14). Photos D. T. Bilton.
</paragraph>
</caption>
<subSubSection id="C354CCFAE10BFF88FF3BB19BFDBA4E93" pageId="15" pageNumber="206" type="etymology">
<paragraph id="8BF19F71E10BFF88FF3BB19BFDBA4E93" blockId="15.[151,1436,151,213]" pageId="15" pageNumber="206">
<emphasis id="B93A4363E10BFF88FF3BB19BFE9E4EF6" bold="true" box="[199,326,151,176]" pageId="15" pageNumber="206">Etymology</emphasis>
. From the latin
<emphasis id="B93A4363E10BFF88FDFBB194FD854EF6" box="[519,605,152,176]" italics="true" pageId="15" pageNumber="206">chasma</emphasis>
(= gorge, steep-sided depression), in reference to Oologskloof Canyon, where both known localities are situated.
</paragraph>
</subSubSection>
</treatment>
</document>
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