561 lines
78 KiB
XML
561 lines
78 KiB
XML
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<mods:title id="A9E8CFF62AAC90428383510EAC1BC55B">Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species</mods:title>
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<mods:namePart id="51F19CEE2C4BE16E3D27BC69CDDA986D">Inkhavilay, Khamla</mods:namePart>
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<mods:affiliation id="5D80C8DFC5193FF0B7E1061EC9415168">Email: kinkhavilay @ gmail. com & urn: lsid: zoobank. org: author: 2 E 620 B 9 C- 48 BA- 48 E 6 - A 017 - FE 8 ED 2 AB 0 E 59</mods:affiliation>
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<mods:nameIdentifier id="93587071C28E4965DE8C111B3C2052A9" type="email">kinkhavilay@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5E9004872C785A2CE9AD242F10227324">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="657AAE8D969056A576159010FF7C5FC6">urn: lsid: zoobank. org: author: C 2 E 2 FA 6 B-A 3 F 9 - 4 F 33 - B 447 - B 59 B 1 BD 322 D 4 & Corresponding authors: jirasak 4 @ yahoo. com; somsak. pan @ chula. ac. th</mods:affiliation>
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<mods:nameIdentifier id="28EB3FCD7B66F4A6FFD33EFCFA2E59A6" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="B05E69EF32286DFF35C81DE4498CE3D2">Panha, Somsak</mods:namePart>
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<mods:affiliation id="6D1353301D9267DAB4DE4CBE5AB10542">urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79 & Corresponding authors: jirasak 4 @ yahoo. com; somsak. pan @ chula. ac. th</mods:affiliation>
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<mods:nameIdentifier id="A770197D7C04F0D17B3F54499B2D7581" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:title id="550922D7D551FEF38075E04780001D02">European Journal of Taxonomy</mods:title>
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<mods:date id="B407A9EC4ECF9117DFEF434CA8F1EAB0">2017</mods:date>
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<treatment id="03DAD35F6931EF11FE67AC14FB772288" ID-DOI="http://doi.org/10.5281/zenodo.3848205" ID-GBIF-Taxon="164161061" ID-Zenodo-Dep="3848205" LSID="urn:lsid:plazi:treatment:03DAD35F6931EF11FE67AC14FB772288" httpUri="http://treatment.plazi.org/id/03DAD35F6931EF11FE67AC14FB772288" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
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<subSubSection id="C36931C26931EF1FFE67AC14FB802626" box="[492,1096,810,837]" pageId="23" pageNumber="24" type="nomenclature">
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<paragraph id="8BCC62496931EF1FFE67AC14FB802626" blockId="23.[492,1096,810,872]" box="[492,1096,810,837]" pageId="23" pageNumber="24">
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<heading id="D084D5256931EF1FFE67AC14FB802626" box="[492,1096,810,837]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="24" reason="2">
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<taxonomicName id="4C7319CA6931EF1FFE67AC14FB802626" ID-CoL="66FPZ" authority="(Pfeiffer, 1861)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[492,1096,810,837]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="flavus" subGenus="Syndromus">
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<emphasis id="B907BE5B6931EF1FFE67AC14FC442626" bold="true" box="[492,908,810,837]" pageId="23" pageNumber="24">
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<emphasis id="B907BE5B6931EF1FFE67AC14FD512626" bold="true" box="[492,665,810,837]" italics="true" pageId="23" pageNumber="24">Amphidromus</emphasis>
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(
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<emphasis id="B907BE5B6931EF1FFD21AC14FCFB2626" bold="true" box="[682,819,810,837]" italics="true" pageId="23" pageNumber="24">Syndromus</emphasis>
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)
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<emphasis id="B907BE5B6931EF1FFCCFAC14FC442626" bold="true" box="[836,908,810,837]" italics="true" pageId="23" pageNumber="24">flavus</emphasis>
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</emphasis>
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(Pfeiffer, 1861)
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36931C26931EF1FFD9EAC70FBD7260B" box="[533,1055,846,872]" pageId="23" pageNumber="24" type="description">
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<paragraph id="8BCC62496931EF1FFD9EAC70FBD7260B" blockId="23.[492,1096,810,872]" box="[533,1055,846,872]" pageId="23" pageNumber="24">
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<figureCitation id="13487ECC6931EF1FFD9EAC70FD95260B" box="[533,605,846,872]" captionStart="Fig" captionStartId="4.[189,232,1991,2017]" captionTargetBox="[416,1172,981,1949]" captionTargetId="figure@4.[416,1172,981,1949]" captionTargetPageId="4" captionText="Fig. 1. Distribution map of Amphidromus spp. in Laos recognized in this study." figureDoi="http://doi.org/10.5281/zenodo.3829746" httpUri="https://zenodo.org/record/3829746/files/figure.png" pageId="23" pageNumber="24">Figs 1</figureCitation>
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,
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<figureCitation id="13487ECC6931EF1FFDE7AC70FD58260B" box="[620,656,846,872]" captionStart="Fig" captionStartId="20.[189,232,1823,1849]" captionTargetBox="[189,1399,656,1782]" captionTargetId="figure@20.[189,1399,656,1782]" captionTargetPageId="20" captionText="Fig. 9. Living snails of Amphidromus (Syndromus) spp. A. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7023). B. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7027). C–D. Amphidromus xiengensis Morlet, 1891 from Thad Kacham, Luang Phrabang, Laos (CUMZ 7035), the typical form, and var. “multifasciata” Fulton, 1896, respectively. E–F. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos (CUMZ 7041–7042). E. Typical color form. F. Monochrome yellowish color form." figureDoi="http://doi.org/10.5281/zenodo.3829762" httpUri="https://zenodo.org/record/3829762/files/figure.png" pageId="23" pageNumber="24">9B</figureCitation>
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,
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<figureCitation id="13487ECC6931EF1FFD16AC70FD71260B" box="[669,697,846,872]" captionStart="Fig" captionStartId="21.[189,232,1752,1778]" captionTargetBox="[189,1399,539,1707]" captionTargetId="figure@21.[189,1399,539,1707]" captionTargetPageId="21" captionText="Fig. 10. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7022). D. Amphidromus begini (Morlet, 1886), syntype from Strung- Trang, Cambodia (MNHN-IM-2000-1832). E-G. Amphidromus flavus (Pfeiffer, 1861) from Tam Pou Kham, Vientiane, Laos (CUMZ 7029). H. Amphidromus flavus var. “indistinctus” Pilsbry, 1900, holotype (ANSP 31486). I–K. Amphidromus flavus var. “tryoni” Pilsbry, 1900. I. Lectotype (ANSP 31488). J. Paralectotype (ANSP 252745). K. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7026). L–M. Amphidromus semitessellatus (Morlet, 1884). L. Lectotype (MNHN-IM-2000-1985). M. Specimen from Cambodia (NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3829764" httpUri="https://zenodo.org/record/3829764/files/figure.png" pageId="23" pageNumber="24">10</figureCitation>
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||
E−K, 11C–D, 12A−C;
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||
<tableCitation id="C6F157F26931EF1FFC43AC70FBD7260B" box="[968,1055,846,872]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionTargetId="graphics@3.[189,1398,1154,1193]" captionTargetPageId="3" captionText="Table 1 (continued on next page). Shell measurements of nine species of Amphidromus from Laos. Specimen catalogue numbers are indicated in parentheses." httpUri="http://table.plazi.org/id/DF0C32C16925EF0BFF36AE32FCA1242B" pageId="23" pageNumber="24" tableUuid="DF0C32C16925EF0BFF36AE32FCA1242B">Table 1</tableCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36931C26931EF1FFF36ACAEFBE22030" pageId="23" pageNumber="24" type="reference_group">
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<paragraph id="8BCC62496931EF1FFF36ACAEFC1026C8" blockId="23.[189,1398,912,1153]" box="[189,984,912,939]" pageId="23" pageNumber="24">
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<treatmentCitationGroup id="AB6345676931EF1FFF36ACAEFC1026C8" box="[189,984,912,939]" pageId="23" pageNumber="24">
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<treatmentCitation id="0AD244586931EF1FFF36ACAEFD9526C8" author="Pfeiffer L." box="[189,605,912,939]" page="194" pageId="23" pageNumber="24" year="1861">
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<taxonomicName id="4C7319CA6931EF1FFF36ACAEFD9526C8" authority="Pfeiffer, 1861 a: 194" authorityName="Pfeiffer" authorityPageNumber="194" authorityYear="1861" box="[189,605,912,939]" class="Gastropoda" family="Bulimulidae" genus="Bulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="flavus">
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<emphasis id="B907BE5B6931EF1FFF36ACAEFEA026C9" box="[189,360,912,938]" italics="true" pageId="23" pageNumber="24">Bulimus flavus</emphasis>
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<bibRefCitation id="EFE21FB86931EF1FFEFBACAEFD9526C8" author="Pfeiffer L." box="[368,605,912,939]" pageId="23" pageNumber="24" pagination="190 - 196" refId="ref19251" refString="Pfeiffer L. 1861 a. Descriptions of new land-shells, in the collection of H. Cuming, Esq. Proceedings of the Zoological Society of London 29: 190 - 196. (Published in parts, dates follow Duncan 1937.) Available from http: // biodiversitylibrary. org / page / 28672714 [accessed 9 Sept. 2013]." type="journal article" year="1861">Pfeiffer, 1861a: 194</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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||
<typeStatus id="54C8DCEB6931EF1FFDE3ACAFFD6A26C8" box="[616,674,913,939]" pageId="23" pageNumber="24">Type</typeStatus>
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locality:
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<collectingCountry id="F36422D96931EF1FFC9AACAEFC8526C9" box="[785,845,912,938]" name="Thailand" pageId="23" pageNumber="24">Siam</collectingCountry>
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[
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<collectingCountry id="F36422D96931EF1FFCD7ACAEFC0626C8" box="[860,974,912,939]" name="Thailand" pageId="23" pageNumber="24">Thailand</collectingCountry>
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].
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BCC62496931EF1FFF36AC8AFE612691" blockId="23.[189,1398,912,1153]" pageId="23" pageNumber="24">
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<treatmentCitationGroup id="AB6345676931EF1FFF36AC8AFE612691" pageId="23" pageNumber="24">
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<treatmentCitation id="0AD244586931EF1FFF36AC8AFCC526AD" author="Fulton H. C." box="[189,781,948,975]" page="81" pageId="23" pageNumber="24" year="1896">
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<taxonomicName id="4C7319CA6931EF1FFF36AC8AFCC526AD" ID-CoL="5M72P" authority="Fulton, 1896: 81" authorityName="Fulton" authorityPageNumber="81" authorityYear="1896" box="[189,781,948,975]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="variety" species="flavus" variety="proxima">
|
||
<emphasis id="B907BE5B6931EF1FFF36AC8AFE6326AD" box="[189,427,948,974]" italics="true" pageId="23" pageNumber="24">Amphidromus flavus</emphasis>
|
||
var.
|
||
<emphasis id="B907BE5B6931EF1FFE6FAC8BFD8D26AD" box="[484,581,949,974]" italics="true" pageId="23" pageNumber="24">proxima</emphasis>
|
||
<bibRefCitation id="EFE21FB86931EF1FFDC0AC8AFCC526AD" author="Fulton H. C." box="[587,781,948,974]" pageId="23" pageNumber="24" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896: 81</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
, pl. 6, fig. 4.
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<typeStatus id="54C8DCEB6931EF1FFC28AC8BFC1526AC" box="[931,989,949,975]" pageId="23" pageNumber="24">Type</typeStatus>
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locality: Unknown.
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||
<treatmentCitation id="0AD244586931EF1FFB47AC8AFEF62690" author="Sutcharit C. & Ablett J. & Tongkerd P. & Naggs F. & Panha S." page="85" pageId="23" pageNumber="24" year="2015">
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<bibRefCitation id="EFE21FB86931EF1FFB47AC8AFEF62690" author="Sutcharit C. & Ablett J. & Tongkerd P. & Naggs F. & Panha S." pageId="23" pageNumber="24" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. & Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">
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||
Sutcharit
|
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<emphasis id="B907BE5B6931EF1FFAB6AC8BFAA726AD" box="[1341,1391,948,974]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
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||
. 2015: 85
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||
</bibRefCitation>
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||
</treatmentCitation>
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||
, fig. 13c.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496931EF1FFF36ACC2FE282159" blockId="23.[189,1398,912,1153]" pageId="23" pageNumber="24">
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<treatmentCitationGroup id="AB6345676931EF1FFF36ACC2FE282159" pageId="23" pageNumber="24">
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<treatmentCitation id="0AD244586931EF1FFF36ACC2FC612175" author="Pilsbry H. A." box="[189,937,1020,1046]" page="197" pageId="23" pageNumber="24" year="1900">
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||
<taxonomicName id="4C7319CA6931EF1FFF36ACC2FC612175" ID-CoL="5M72X" authority="Pilsbry, 1900: 196 - 197" authorityName="Pilsbry" authorityPageNumber="196 - 197" authorityYear="1900" box="[189,937,1020,1046]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="variety" species="xiengensis" variety="tryoni">
|
||
<emphasis id="B907BE5B6931EF1FFF36ACC2FE212175" box="[189,489,1020,1046]" italics="true" pageId="23" pageNumber="24">Amphidromus xiengensis</emphasis>
|
||
var.
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||
<emphasis id="B907BE5B6931EF1FFDB9ACC3FDB02175" box="[562,632,1021,1046]" italics="true" pageId="23" pageNumber="24">tryoni</emphasis>
|
||
<bibRefCitation id="EFE21FB86931EF1FFD0CACC2FC612175" author="Pilsbry H. A." box="[647,937,1020,1046]" pageId="23" pageNumber="24" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry, 1900: 196–197</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
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||
, pl. 63, fig. 78.
|
||
<typeStatus id="54C8DCEB6931EF1FFB0EACC2FB772175" box="[1157,1215,1020,1046]" pageId="23" pageNumber="24">Type</typeStatus>
|
||
locality:
|
||
<collectingCountry id="F36422D96931EF1FFAB6ACC2FABE2175" box="[1341,1398,1020,1046]" name="Laos" pageId="23" pageNumber="24">Laos</collectingCountry>
|
||
Mountains,
|
||
<collectingCountry id="F36422D96931EF1FFEEBAB1EFE152159" box="[352,477,1056,1082]" name="Cambodia" pageId="23" pageNumber="24">Cambodia</collectingCountry>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496931EF1FFF36AB7AFE9C21E2" blockId="23.[189,1398,912,1153]" pageId="23" pageNumber="24">
|
||
<treatmentCitationGroup id="AB6345676931EF1FFF36AB7AFE9C21E2" pageId="23" pageNumber="24">
|
||
<treatmentCitation id="0AD244586931EF1FFF36AB7AFC96213D" author="Pilsbry H. A." box="[189,862,1091,1118]" page="192" pageId="23" pageNumber="24" year="1900">
|
||
<taxonomicName id="4C7319CA6931EF1FFF36AB7AFC96213D" ID-CoL="5M72V" authority="Pilsbry, 1900: 192" authorityName="Pilsbry" authorityPageNumber="192" authorityYear="1900" box="[189,862,1091,1118]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="variety" species="sinensis" variety="indistinctus">
|
||
<emphasis id="B907BE5B6931EF1FFF36AB7AFE0B213E" box="[189,451,1092,1118]" italics="true" pageId="23" pageNumber="24">Amphidromus sinensis</emphasis>
|
||
var.
|
||
<emphasis id="B907BE5B6931EF1FFE71AB7AFD49213D" box="[506,641,1092,1118]" italics="true" pageId="23" pageNumber="24">indistinctus</emphasis>
|
||
<bibRefCitation id="EFE21FB86931EF1FFD0CAB7DFC96213D" author="Pilsbry H. A." box="[647,862,1091,1118]" pageId="23" pageNumber="24" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry, 1900: 192</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, pl. 62, fig. 70.
|
||
<typeStatus id="54C8DCEB6931EF1FFB86AB7AFB8F213D" box="[1037,1095,1092,1118]" pageId="23" pageNumber="24">Type</typeStatus>
|
||
locality:
|
||
<collectingCountry id="F36422D96931EF1FFB38AB7AFB24213D" box="[1203,1260,1092,1118]" name="Laos" pageId="23" pageNumber="24">Laos</collectingCountry>
|
||
Mountains,
|
||
<collectingCountry id="F36422D96931EF1FFF5FAB59FE9921E2" box="[212,337,1127,1153]" name="Cambodia" pageId="23" pageNumber="24">Cambodia</collectingCountry>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496931EF1FFF36AB97FBF721A7" blockId="23.[189,1398,1193,1363]" box="[189,1087,1193,1220]" pageId="23" pageNumber="24">
|
||
<treatmentCitationGroup id="AB6345676931EF1FFF36AB97FBF721A7" box="[189,1087,1193,1220]" pageId="23" pageNumber="24">
|
||
<taxonomicName id="4C7319CA6931EF1FFF36AB97FEA021A0" authorityName="Pfeiffer" authorityYear="1861" box="[189,360,1193,1219]" class="Gastropoda" family="Bulimulidae" genus="Bulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="flavus">
|
||
<emphasis id="B907BE5B6931EF1FFF36AB97FEA021A0" box="[189,360,1193,1219]" italics="true" pageId="23" pageNumber="24">Bulimus flavus</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AD244586931EF1FFE0DAB97FDA521A7" author="Pfeiffer L." box="[390,621,1193,1220]" page="171" pageId="23" pageNumber="24" year="1861">
|
||
<bibRefCitation id="EFE21FB86931EF1FFE0DAB97FDA521A7" author="Pfeiffer L." box="[390,621,1193,1220]" pageId="23" pageNumber="24" pagination="161 - 178" refId="ref19316" refString="Pfeiffer L. 1861 b (1860 - 1866). Novitates Conchologicae. Series Prima Mollusca Extramarina, Vol. 2: 139 - 303, pls 37 - 72 (1861: 161 - 178, pls 43 - 48). Theodor Fischer Verlag, Kassel. (Published in parts, dates follow Johnson 1969.) Available from http: // biodiversitylibrary. org / item / 122926 [accessed 8 Jun. 2016]." type="journal article" year="1861">Pfeiffer 1861b: 171</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 46, figs 7–8. —
|
||
<treatmentCitation id="0AD244586931EF1FFCD6AB97FBF321A7" author="Morelet A." box="[861,1083,1193,1220]" page="265" pageId="23" pageNumber="24" year="1875">
|
||
<bibRefCitation id="EFE21FB86931EF1FFCD6AB97FBF321A7" author="Morelet A." box="[861,1083,1193,1220]" pageId="23" pageNumber="24" refId="ref18862" refString="Morelet A. 1875. Series conchyliologiques comprenant l'Enumeration de Mollusques terrestres et fluviatiles recueillies pendant le Cours de differents Voyages ainsi que la Description de plusieurs Especes nouvelles. Fourth issue: 227 - 377. Klincksieck, Paris. https: // doi. org / 10.5962 / bhl. title. 11458" type="book" year="1875">
|
||
Morelet
|
||
<date id="FFCD44896931EF1FFC4AAB97FBC821A7" box="[961,1024,1193,1220]" pageId="23" pageNumber="24" value="1875">1875</date>
|
||
: 265
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496931EF1FFF36ABF3FCD32068" blockId="23.[189,1398,1193,1363]" pageId="23" pageNumber="24">
|
||
<treatmentCitationGroup id="AB6345676931EF1FFF36ABF3FCD32068" pageId="23" pageNumber="24">
|
||
<taxonomicName id="4C7319CA6931EF1FFF36ABF3FE672184" ID-CoL="66FPZ" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[189,431,1229,1255]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="flavus">
|
||
<emphasis id="B907BE5B6931EF1FFF36ABF3FE672184" box="[189,431,1229,1255]" italics="true" pageId="23" pageNumber="24">Amphidromus flavus</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AD244586931EF1FFE59ABF3FD5E2184" author="Fulton H. C." box="[466,662,1229,1255]" page="81" pageId="23" pageNumber="24" year="1896">
|
||
<bibRefCitation id="EFE21FB86931EF1FFE59ABF3FD5E2184" author="Fulton H. C." box="[466,662,1229,1255]" pageId="23" pageNumber="24" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton 1896: 81</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AD244586931EF1FFD41ABF3FC552184" author="Ancey C. F." box="[714,925,1229,1255]" page="134" pageId="23" pageNumber="24" year="1898">
|
||
<bibRefCitation id="EFE21FB86931EF1FFD41ABF3FC552184" author="Ancey C. F." box="[714,925,1229,1255]" pageId="23" pageNumber="24" pagination="129 - 137" refId="ref17973" refString="Ancey C. F. 1898. Observations sur les mollusques terrestres et fluviatiles recueillis dans lIndo-chine et particulierement au Laos par M. H. Counillon. Bulletin du Museum d'Histoire naturelle de Marseille 1: 129 - 137." type="journal article" year="1898">Ancey 1898: 134</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AD244586931EF1FFC58ABF3FB2E2184" author="Pilsbry H. A." box="[979,1254,1229,1255]" page="197" pageId="23" pageNumber="24" year="1900">
|
||
<bibRefCitation id="EFE21FB86931EF1FFC58ABF3FB2E2184" author="Pilsbry H. A." box="[979,1254,1229,1255]" pageId="23" pageNumber="24" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry 1900: 197–198</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 63, figs 92–93. —
|
||
<treatmentCitation id="0AD244586931EF1FFEDBABCFFD14206F" author="Laidlaw F. F. & Solem A." box="[336,732,1265,1292]" page="563" pageId="23" pageNumber="24" year="1961">
|
||
<bibRefCitation id="EFE21FB86931EF1FFEDBABCFFD14206F" author="Laidlaw F. F. & Solem A." box="[336,732,1265,1292]" pageId="23" pageNumber="24" pagination="507 - 677" refId="ref18651" refString="Laidlaw F. F. & Solem A. 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana: Zoology 41: 507 - 677. Available from http: // biodiversitylibrary. org / item / 20789 [accessed 18 Jul. 2013]." type="journal article" year="1961">
|
||
Laidlaw & Solem
|
||
<date id="FFCD44896931EF1FFDA3ABCFFDAF2068" box="[552,615,1265,1291]" pageId="23" pageNumber="24" value="1961">1961</date>
|
||
: 563–564
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 619.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496931EF1FFF36AA2BFC30204C" blockId="23.[189,1398,1193,1363]" box="[189,1016,1301,1327]" pageId="23" pageNumber="24">
|
||
<treatmentCitationGroup id="AB6345676931EF1FFF36AA2BFC30204C" box="[189,1016,1301,1327]" pageId="23" pageNumber="24">
|
||
<taxonomicName id="4C7319CA6931EF1FFF36AA2BFD9A204C" box="[189,594,1301,1327]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="variety" species="flavus" variety="proximus">
|
||
<emphasis id="B907BE5B6931EF1FFF36AA2BFE64204C" box="[189,428,1301,1327]" italics="true" pageId="23" pageNumber="24">Amphidromus flavus</emphasis>
|
||
var.
|
||
<emphasis id="B907BE5B6931EF1FFE6DAA28FD9A204C" box="[486,594,1302,1327]" italics="true" pageId="23" pageNumber="24">proximus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B907BE5B6931EF1FFDD1AA28FDA1204C" box="[602,617,1302,1327]" italics="true" pageId="23" pageNumber="24">–</emphasis>
|
||
<treatmentCitation id="0AD244586931EF1FFDFBAA2BFC80204C" author="Pilsbry H. A." box="[624,840,1301,1327]" page="198" pageId="23" pageNumber="24" year="1900">
|
||
<bibRefCitation id="EFE21FB86931EF1FFDFBAA2BFC80204C" author="Pilsbry H. A." box="[624,840,1301,1327]" pageId="23" pageNumber="24" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry, 1900: 198</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 63, fig. 94.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496931EF1FFF36AA07FBE22030" blockId="23.[189,1398,1193,1363]" box="[189,1066,1336,1363]" pageId="23" pageNumber="24">
|
||
<treatmentCitationGroup id="AB6345676931EF1FFF36AA07FBE22030" box="[189,1066,1336,1363]" pageId="23" pageNumber="24">
|
||
<taxonomicName id="4C7319CA6931EF1FFF36AA07FDB52030" ID-CoL="5M72P" box="[189,637,1337,1363]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="variety" species="xiengensis" variety="proxima">
|
||
<emphasis id="B907BE5B6931EF1FFF36AA07FE2A2030" box="[189,482,1337,1363]" italics="true" pageId="23" pageNumber="24">Amphidromus xiengensis</emphasis>
|
||
var.
|
||
<emphasis id="B907BE5B6931EF1FFD97AA04FDB52030" box="[540,637,1338,1363]" italics="true" pageId="23" pageNumber="24">proxima</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AD244586931EF1FFD11AA07FC232030" author="Laidlaw F. F. & Solem A." box="[666,1003,1336,1363]" page="565" pageId="23" pageNumber="24" year="1961">
|
||
<bibRefCitation id="EFE21FB86931EF1FFD11AA07FC232030" author="Laidlaw F. F. & Solem A." box="[666,1003,1336,1363]" pageId="23" pageNumber="24" pagination="507 - 677" refId="ref18651" refString="Laidlaw F. F. & Solem A. 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana: Zoology 41: 507 - 677. Available from http: // biodiversitylibrary. org / item / 20789 [accessed 18 Jul. 2013]." type="journal article" year="1961">Laidlaw & Solem 1961: 565</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 652.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36931C26931EF10FF36AA45FEA7240F" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="materials_examined">
|
||
<paragraph id="8BCC62496931EF1FFF36AA45FE6220F6" blockId="23.[189,426,1403,1429]" box="[189,426,1403,1429]" pageId="23" pageNumber="24">
|
||
<heading id="D084D5256931EF1FFF36AA45FE6220F6" bold="true" box="[189,426,1403,1429]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
|
||
<emphasis id="B907BE5B6931EF1FFF36AA45FE6220F6" bold="true" box="[189,426,1403,1429]" pageId="23" pageNumber="24">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496931EF1FFF61AA94FE5120A7" blockId="23.[189,1399,1450,1548]" box="[234,409,1450,1476]" pageId="23" pageNumber="24">
|
||
<emphasis id="B907BE5B6931EF1FFF61AA94FE5120A7" bold="true" box="[234,409,1450,1476]" pageId="23" pageNumber="24">
|
||
<typeStatus id="54C8DCEB6931EF1FFF61AA94FEEF20A7" box="[234,295,1450,1476]" pageId="23" pageNumber="24">Type</typeStatus>
|
||
material
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496931EF1FFF36AAF0FE6A236F" blockId="23.[189,1399,1450,1548]" pageId="23" pageNumber="24">
|
||
<collectingCountry id="F36422D96931EF1FFF36AAF0FE91208B" box="[189,345,1486,1512]" name="Thailand" pageId="23" pageNumber="24">THAILAND</collectingCountry>
|
||
:
|
||
<typeStatus id="54C8DCEB6931EF1FFEEDAAF0FE16208B" box="[358,478,1486,1512]" pageId="23" pageNumber="24" type="lectotype">Lectotype</typeStatus>
|
||
,
|
||
<collectionCode id="ED62FA8C6931EF1FFE66AAF0FD94208B" box="[493,604,1486,1512]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="23" pageNumber="24" type="Museum">NHMUK</collectionCode>
|
||
19601436 (
|
||
<bibRefCitation id="EFE21FB86931EF1FFD67AAF0FC20208A" author="Sutcharit C. & Ablett J. & Tongkerd P. & Naggs F. & Panha S." box="[748,1000,1486,1513]" pageId="23" pageNumber="24" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. & Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">
|
||
Sutcharit
|
||
<emphasis id="B907BE5B6931EF1FFCE9AAF1FC50208B" box="[866,920,1486,1512]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
: fig. 7i);
|
||
<typeStatus id="54C8DCEB6931EF1FFBD3AAF0FB31208B" box="[1112,1273,1486,1512]" pageId="23" pageNumber="24" type="paralectotype">paralectotype</typeStatus>
|
||
,
|
||
<collectionCode id="ED62FA8C6931EF1FFA83AAF0FABF208B" box="[1288,1399,1486,1512]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="23" pageNumber="24" type="Museum">NHMUK</collectionCode>
|
||
19601437 (1 shell).
|
||
</paragraph>
|
||
<paragraph id="8BCC62496931EF1FFF36A90AFE7A2311" blockId="23.[189,1399,1587,1650]" pageId="23" pageNumber="24">
|
||
UNKNOWN LOCALITY:
|
||
<typeStatus id="54C8DCEB6931EF1FFD8FA90AFDB9232D" box="[516,625,1588,1614]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
|
||
of var. “
|
||
<emphasis id="B907BE5B6931EF1FFD5EA90AFCFE232E" box="[725,822,1588,1613]" italics="true" pageId="23" pageNumber="24">proxima</emphasis>
|
||
”
|
||
<bibRefCitation id="EFE21FB86931EF1FFCC1A90AFC29232E" author="Fulton H. C." box="[842,993,1587,1614]" pageId="23" pageNumber="24" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896</bibRefCitation>
|
||
,
|
||
<collectionCode id="ED62FA8C6931EF1FFC66A90AFB94232D" box="[1005,1116,1588,1614]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="23" pageNumber="24" type="Museum">NHMUK</collectionCode>
|
||
1896.6.13.48 (
|
||
<bibRefCitation id="EFE21FB86931EF1FFA80A90AFEF62311" author="Sutcharit C. & Ablett J. & Tongkerd P. & Naggs F. & Panha S." pageId="23" pageNumber="24" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. & Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">
|
||
Sutcharit
|
||
<emphasis id="B907BE5B6931EF1FFF36A966FF382312" box="[189,240,1623,1649]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
: fig. 13c).
|
||
</paragraph>
|
||
<paragraph id="8BCC62496931EF1FFF36A9A4FB7823D7" blockId="23.[189,1200,1689,1716]" box="[189,1200,1689,1716]" pageId="23" pageNumber="24">
|
||
<collectingCountry id="F36422D96931EF1FFF36A9A4FEA923D7" box="[189,353,1690,1716]" name="Cambodia" pageId="23" pageNumber="24">CAMBODIA</collectingCountry>
|
||
:
|
||
<typeStatus id="54C8DCEB6931EF1FFEFFA9A7FE2923D0" box="[372,481,1689,1715]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
|
||
of var. “
|
||
<emphasis id="B907BE5B6931EF1FFDCCA9A7FD0623D0" box="[583,718,1689,1715]" italics="true" pageId="23" pageNumber="24">indistinctus</emphasis>
|
||
”
|
||
<bibRefCitation id="EFE21FB86931EF1FFD68A9A7FC4823D7" author="Pilsbry H. A." box="[739,896,1689,1716]" pageId="23" pageNumber="24" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry, 1900</bibRefCitation>
|
||
,
|
||
<specimenCode id="DBD5CA326931EF1FFC00A9A4FBED23D0" box="[907,1061,1689,1716]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="23" pageNumber="24">ANSP 31486</specimenCode>
|
||
(
|
||
<figureCitation id="13487ECC6931EF1FFBBEA9A7FB6B23D7" box="[1077,1187,1689,1716]" captionStart="Fig" captionStartId="21.[189,232,1752,1778]" captionTargetBox="[189,1399,539,1707]" captionTargetId="figure@21.[189,1399,539,1707]" captionTargetPageId="21" captionText="Fig. 10. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7022). D. Amphidromus begini (Morlet, 1886), syntype from Strung- Trang, Cambodia (MNHN-IM-2000-1832). E-G. Amphidromus flavus (Pfeiffer, 1861) from Tam Pou Kham, Vientiane, Laos (CUMZ 7029). H. Amphidromus flavus var. “indistinctus” Pilsbry, 1900, holotype (ANSP 31486). I–K. Amphidromus flavus var. “tryoni” Pilsbry, 1900. I. Lectotype (ANSP 31488). J. Paralectotype (ANSP 252745). K. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7026). L–M. Amphidromus semitessellatus (Morlet, 1884). L. Lectotype (MNHN-IM-2000-1985). M. Specimen from Cambodia (NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3829764" httpUri="https://zenodo.org/record/3829764/files/figure.png" pageId="23" pageNumber="24">
|
||
Fig. 10
|
||
<collectionCode id="ED62FA8C6931EF1FFB00A9A4FB6B23D7" box="[1163,1187,1690,1716]" country="Finland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15618" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="23" pageNumber="24" type="Herbarium">H</collectionCode>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC62496931EF1FFF36A9E2FE35227A" blockId="23.[189,1399,1755,1818]" pageId="23" pageNumber="24">
|
||
<collectingCountry id="F36422D96931EF1FFF36A9E2FEA92395" box="[189,353,1756,1782]" name="Cambodia" pageId="23" pageNumber="24">CAMBODIA</collectingCountry>
|
||
:
|
||
<typeStatus id="54C8DCEB6931EF1FFEFEA9E2FE222395" box="[373,490,1756,1782]" pageId="23" pageNumber="24" type="lectotype">Lectotype</typeStatus>
|
||
of var. “
|
||
<emphasis id="B907BE5B6931EF1FFDDFA9E2FD522396" box="[596,666,1756,1781]" italics="true" pageId="23" pageNumber="24">tryoni</emphasis>
|
||
”
|
||
<bibRefCitation id="EFE21FB86931EF1FFD24A9E5FC852395" author="Pilsbry H. A." box="[687,845,1755,1782]" pageId="23" pageNumber="24" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry, 1900</bibRefCitation>
|
||
,
|
||
<specimenCode id="DBD5CA326931EF1FFCD3A9E2FC3B2395" box="[856,1011,1756,1782]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="23" pageNumber="24">ANSP 31488</specimenCode>
|
||
(
|
||
<figureCitation id="13487ECC6931EF1FFB8EA9E5FBAF2395" box="[1029,1127,1755,1782]" captionStart="Fig" captionStartId="21.[189,232,1752,1778]" captionTargetBox="[189,1399,539,1707]" captionTargetId="figure@21.[189,1399,539,1707]" captionTargetPageId="21" captionText="Fig. 10. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7022). D. Amphidromus begini (Morlet, 1886), syntype from Strung- Trang, Cambodia (MNHN-IM-2000-1832). E-G. Amphidromus flavus (Pfeiffer, 1861) from Tam Pou Kham, Vientiane, Laos (CUMZ 7029). H. Amphidromus flavus var. “indistinctus” Pilsbry, 1900, holotype (ANSP 31486). I–K. Amphidromus flavus var. “tryoni” Pilsbry, 1900. I. Lectotype (ANSP 31488). J. Paralectotype (ANSP 252745). K. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7026). L–M. Amphidromus semitessellatus (Morlet, 1884). L. Lectotype (MNHN-IM-2000-1985). M. Specimen from Cambodia (NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3829764" httpUri="https://zenodo.org/record/3829764/files/figure.png" pageId="23" pageNumber="24">
|
||
Fig. 10
|
||
<collectionCode id="ED62FA8C6931EF1FFBD7A9E2FBAF2395" box="[1116,1127,1756,1782]" country="Romania" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14415" lsid="urn:lsid:biocol.org:col:14415" name="&quot;Alexandru Ioan Cuza&quot; University" pageId="23" pageNumber="24" type="Herbarium">I</collectionCode>
|
||
</figureCitation>
|
||
); paralectoypes,
|
||
<specimenCode id="DBD5CA326931EF1FFAA6A9E2FEDD2279" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="23" pageNumber="24">ANSP 252745</specimenCode>
|
||
(2 shells,
|
||
<figureCitation id="13487ECC6931EF1FFE05A9C1FE38227A" box="[398,496,1791,1817]" captionStart="Fig" captionStartId="21.[189,232,1752,1778]" captionTargetBox="[189,1399,539,1707]" captionTargetId="figure@21.[189,1399,539,1707]" captionTargetPageId="21" captionText="Fig. 10. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7022). D. Amphidromus begini (Morlet, 1886), syntype from Strung- Trang, Cambodia (MNHN-IM-2000-1832). E-G. Amphidromus flavus (Pfeiffer, 1861) from Tam Pou Kham, Vientiane, Laos (CUMZ 7029). H. Amphidromus flavus var. “indistinctus” Pilsbry, 1900, holotype (ANSP 31486). I–K. Amphidromus flavus var. “tryoni” Pilsbry, 1900. I. Lectotype (ANSP 31488). J. Paralectotype (ANSP 252745). K. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7026). L–M. Amphidromus semitessellatus (Morlet, 1884). L. Lectotype (MNHN-IM-2000-1985). M. Specimen from Cambodia (NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3829764" httpUri="https://zenodo.org/record/3829764/files/figure.png" pageId="23" pageNumber="24">
|
||
Fig. 10
|
||
<collectionCode id="ED62FA8C6931EF1FFE68A9C1FE38227A" box="[483,496,1791,1817]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14647" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="23" pageNumber="24" type="Herbarium">J</collectionCode>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC62496931EF1FFF61A87FFE6E2238" blockId="23.[189,1399,1857,2027]" box="[234,422,1857,1883]" pageId="23" pageNumber="24">
|
||
<emphasis id="B907BE5B6931EF1FFF61A87FFE6E2238" bold="true" box="[234,422,1857,1883]" pageId="23" pageNumber="24">Other material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496931EF10FF36A85BFEA7240F" blockId="23.[189,1399,1857,2027]" lastBlockId="24.[189,1398,267,364]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
|
||
<collectingCountry id="F36422D96931EF1FFF36A85BFEC6221C" box="[189,270,1893,1919]" name="Laos" pageId="23" pageNumber="24">LAOS</collectingCountry>
|
||
: 5 shells, park at the temple mountain, Luang Phrabang (
|
||
<collectionCode id="ED62FA8C6931EF1FFC42A85BFBF2221C" box="[969,1082,1893,1919]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="23" pageNumber="24" type="Museum">NHMUK</collectionCode>
|
||
ex. Brandt collection No. 17320); 20 shells, Tempelberg (Temple Mount), Luang Phrabang (
|
||
<collectionCode id="ED62FA8C6931EF1FFC7DA8B7FBFF22C0" box="[1014,1079,1929,1955]" country="Germany" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="23" pageNumber="24" type="Museum">ZMB</collectionCode>
|
||
ex. Lehmann collection); 33 shells, Wat Phousy (temple), Luang Phrabang (
|
||
<collectionCode id="ED62FA8C6931EF1FFCA9A893FCB122A4" box="[802,889,1965,1991]" country="0" name="Cameroon University, Museum of Zoology" pageId="23" pageNumber="24">CUMZ</collectionCode>
|
||
7025); 3 shells (
|
||
<figureCitation id="13487ECC6931EF1FFBC7A893FB7422A4" box="[1100,1212,1965,1991]" captionStart="Fig" captionStartId="21.[189,232,1752,1778]" captionTargetBox="[189,1399,539,1707]" captionTargetId="figure@21.[189,1399,539,1707]" captionTargetPageId="21" captionText="Fig. 10. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7022). D. Amphidromus begini (Morlet, 1886), syntype from Strung- Trang, Cambodia (MNHN-IM-2000-1832). E-G. Amphidromus flavus (Pfeiffer, 1861) from Tam Pou Kham, Vientiane, Laos (CUMZ 7029). H. Amphidromus flavus var. “indistinctus” Pilsbry, 1900, holotype (ANSP 31486). I–K. Amphidromus flavus var. “tryoni” Pilsbry, 1900. I. Lectotype (ANSP 31488). J. Paralectotype (ANSP 252745). K. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7026). L–M. Amphidromus semitessellatus (Morlet, 1884). L. Lectotype (MNHN-IM-2000-1985). M. Specimen from Cambodia (NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3829764" httpUri="https://zenodo.org/record/3829764/files/figure.png" pageId="23" pageNumber="24">
|
||
Fig. 10
|
||
<collectionCode id="ED62FA8C6931EF1FFB2FA893FB7422A4" box="[1188,1212,1965,1991]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15867" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="23" pageNumber="24" type="Herbarium">K</collectionCode>
|
||
</figureCitation>
|
||
,
|
||
<collectionCode id="ED62FA8C6931EF1FFB40A893FAE822A4" box="[1227,1312,1965,1991]" country="0" name="Cameroon University, Museum of Zoology" pageId="23" pageNumber="24">CUMZ</collectionCode>
|
||
7026),
|
||
<specimenCount id="9D75A9C06931EF1FFF36A8EFFE9C2288" box="[189,340,2001,2027]" pageId="23" pageNumber="24" type="generic">7 specimens</specimenCount>
|
||
, in ethanol (
|
||
<figureCitation id="13487ECC6931EF1FFE7BA8EFFD9C2288" box="[496,596,2001,2027]" captionStart="Fig" captionStartId="20.[189,232,1823,1849]" captionTargetBox="[189,1399,656,1782]" captionTargetId="figure@20.[189,1399,656,1782]" captionTargetPageId="20" captionText="Fig. 9. Living snails of Amphidromus (Syndromus) spp. A. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7023). B. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7027). C–D. Amphidromus xiengensis Morlet, 1891 from Thad Kacham, Luang Phrabang, Laos (CUMZ 7035), the typical form, and var. “multifasciata” Fulton, 1896, respectively. E–F. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos (CUMZ 7041–7042). E. Typical color form. F. Monochrome yellowish color form." figureDoi="http://doi.org/10.5281/zenodo.3829762" httpUri="https://zenodo.org/record/3829762/files/figure.png" pageId="23" pageNumber="24">
|
||
Figs 9
|
||
<collectionCode id="ED62FA8C6931EF1FFDB4A8EFFD9C2288" box="[575,596,2001,2027]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15534" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="23" pageNumber="24" type="Herbarium">B</collectionCode>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="13487ECC6931EF1FFDEEA8EFFDB72288" box="[613,639,2001,2027]" captionStart="Fig" captionStartId="22.[189,232,1844,1870]" captionTargetBox="[189,1399,298,1799]" captionTargetId="figure@22.[189,1399,298,1799]" captionTargetPageId="22" captionText="Fig. 11. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7023). C–D. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vagina chamber (CUMZ 7027)." figureDoi="http://doi.org/10.5281/zenodo.3829766" httpUri="https://zenodo.org/record/3829766/files/figure.png" pageId="23" pageNumber="24">11</figureCitation>
|
||
C–D, 12A−C,
|
||
<collectionCode id="ED62FA8C6931EF1FFCBFA8EFFC412288" box="[820,905,2001,2027]" country="0" name="Cameroon University, Museum of Zoology" pageId="23" pageNumber="24">CUMZ</collectionCode>
|
||
7027), Ban Na Deauy, Luang Phrabang District, Luang Phrabang; 5 shells, Mouhot’s Tomb,
|
||
<collectionCode id="ED62FA8C693EEF10FCB1AE32FC842445" box="[826,844,268,294]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="24" pageNumber="25" type="Herbarium">E</collectionCode>
|
||
bank of Kan River, Luang Phrabang District, Luang Phrabang (
|
||
<collectionCode id="ED62FA8C693EEF10FE00AE11FE2A242A" box="[395,482,303,329]" country="0" name="Cameroon University, Museum of Zoology" pageId="24" pageNumber="25">CUMZ</collectionCode>
|
||
7028); 16 shells (
|
||
<figureCitation id="13487ECC693EEF10FD3BAE10FCC9242A" box="[688,769,302,329]" captionStart="Fig" captionStartId="21.[189,232,1752,1778]" captionTargetBox="[189,1399,539,1707]" captionTargetId="figure@21.[189,1399,539,1707]" captionTargetPageId="21" captionText="Fig. 10. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7022). D. Amphidromus begini (Morlet, 1886), syntype from Strung- Trang, Cambodia (MNHN-IM-2000-1832). E-G. Amphidromus flavus (Pfeiffer, 1861) from Tam Pou Kham, Vientiane, Laos (CUMZ 7029). H. Amphidromus flavus var. “indistinctus” Pilsbry, 1900, holotype (ANSP 31486). I–K. Amphidromus flavus var. “tryoni” Pilsbry, 1900. I. Lectotype (ANSP 31488). J. Paralectotype (ANSP 252745). K. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7026). L–M. Amphidromus semitessellatus (Morlet, 1884). L. Lectotype (MNHN-IM-2000-1985). M. Specimen from Cambodia (NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3829764" httpUri="https://zenodo.org/record/3829764/files/figure.png" pageId="24" pageNumber="25">Fig. 10</figureCitation>
|
||
E−G), Tam Pou Kham, Vang Vieng District,
|
||
<collectingRegion id="49B7ACAB693EEF10FA8DAE10FABE242B" box="[1286,1398,302,328]" country="Laos" name="Viangchan" pageId="24" pageNumber="25">Vientiane</collectingRegion>
|
||
(
|
||
<collectionCode id="ED62FA8C693EEF10FF4EAE6CFED4240F" box="[197,284,338,364]" country="0" name="Cameroon University, Museum of Zoology" pageId="24" pageNumber="25">CUMZ</collectionCode>
|
||
7029).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36931C2693EEF11FF36AEADFEF72139" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
|
||
<paragraph id="8BCC6249693EEF10FF36AEADFE8624CE" blockId="24.[189,334,403,429]" box="[189,334,403,429]" pageId="24" pageNumber="25">
|
||
<heading id="D084D525693EEF10FF36AEADFE8624CE" bold="true" box="[189,334,403,429]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
|
||
<emphasis id="B907BE5B693EEF10FF36AEADFE8624CE" bold="true" box="[189,334,403,429]" pageId="24" pageNumber="25">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCC6249693EEF11FF36AEFCFE8D2446" blockId="24.[189,1399,450,652]" lastBlockId="25.[189,1399,267,540]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
|
||
Shell sinistral, small, ovate conical, small, thin and glossy. Spire conical; apex acute yellowish, with black spot on tip. Whorls 5 and 6 convex; suture depressed; last whorl rounded to well rounded. Periostracum transparent to thin corneous. Last whorl processes uniform yellowish; Bands 1 to 3 usually absent; Bands 4 and 5 absent or present with indistinct bands; Band 6 usually absent or sometimes present as scantly reddish band. Parietal callus thin and transparent.Aperture ovate to sub-ovate; peristome weakly thickened and shortly expanded; lip whitish. Columella straight and white. Umbilicus opened to narrowly opened.
|
||
<emphasis id="B907BE5B693FEF11FF61AE35FE8D2446" bold="true" box="[234,325,267,293]" pageId="25" pageNumber="26">Radula</emphasis>
|
||
</paragraph>
|
||
<caption id="DF0C32C1693EEF10FF36A81EFB382289" ID-DOI="http://doi.org/10.5281/zenodo.3829768" ID-Zenodo-Dep="3829768" httpUri="https://zenodo.org/record/3829768/files/figure.png" pageId="24" pageNumber="25" startId="24.[189,232,1824,1850]" targetBox="[194,1391,713,1777]" targetPageId="24">
|
||
<paragraph id="8BCC6249693EEF10FF36A81EFB382289" blockId="24.[189,1399,1824,2026]" pageId="24" pageNumber="25">
|
||
<emphasis id="B907BE5B693EEF10FF36A81EFED42259" bold="true" box="[189,284,1824,1850]" pageId="24" pageNumber="25">Fig. 12.</emphasis>
|
||
SEM images of the radula.
|
||
<emphasis id="B907BE5B693EEF10FDFFA81EFD652259" bold="true" box="[628,685,1824,1850]" pageId="24" pageNumber="25">A–C</emphasis>
|
||
.
|
||
<taxonomicName id="4C7319CA693EEF10FD34A81EFBBA2258" authority="(Pfeiffer, 1861)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[703,1138,1824,1851]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="flavus">
|
||
<emphasis id="B907BE5B693EEF10FD34A81EFC782259" box="[703,944,1824,1850]" italics="true" pageId="24" pageNumber="25">Amphidromus flavus</emphasis>
|
||
(Pfeiffer, 1861)
|
||
</taxonomicName>
|
||
from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7027).
|
||
<emphasis id="B907BE5B693EEF10FDF4A87DFD7D223E" bold="true" box="[639,693,1859,1885]" pageId="24" pageNumber="25">D–F</emphasis>
|
||
.
|
||
<taxonomicName id="4C7319CA693EEF10FD49A87DFB4B223E" authority="Morlet, 1891" authorityName="Morlet" authorityYear="1891" box="[706,1155,1859,1885]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="xiengensis">
|
||
<emphasis id="B907BE5B693EEF10FD49A87DFC2D223E" box="[706,997,1859,1885]" italics="true" pageId="24" pageNumber="25">Amphidromus xiengensis</emphasis>
|
||
Morlet, 1891
|
||
</taxonomicName>
|
||
from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7037).
|
||
<emphasis id="B907BE5B693EEF10FD10A858FD0422E3" bold="true" box="[667,716,1894,1920]" pageId="24" pageNumber="25">G–I</emphasis>
|
||
.
|
||
<taxonomicName id="4C7319CA693EEF10FD55A858FB3022E3" authority="Mollendorff, 1898" authorityName="Mollendorff" authorityYear="1898" box="[734,1272,1894,1920]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="fuscolabris">
|
||
<emphasis id="B907BE5B693EEF10FD55A858FBC722E3" box="[734,1039,1894,1920]" italics="true" pageId="24" pageNumber="25">Amphidromus fuscolabris</emphasis>
|
||
<bibRefCitation id="EFE21FB8693EEF10FB90A858FB3022E3" author="Mollendorff O. F. von" box="[1051,1272,1894,1920]" pageId="24" pageNumber="25" pagination="65 - 85" refId="ref18807" refString="Mollendorff O. F. von. 1898. Die Binnenmollusken Annams. Nachrichtsblatt der deutschen Malakozoologischen Gesellschaft 30: 65 - 85. Available from http: // www. biodiversitylibrary. org / item / 89671 # page / 261 / mode / 1 up [accessed 21 Apr. 2017]." type="journal article" year="1898">Möllendorff, 1898</bibRefCitation>
|
||
</taxonomicName>
|
||
from Ban Phone, Sekong, Laos (CUMZ 7041). A, D, G = central tooth with the first to fifth or sixth lateral teeth; B, E, H = lateral teeth with the tricuspid marginal teeth transition; C, F, I = outermost marginal teeth. Numbers indicate the order of the lateral and marginal teeth. Central tooth indicated by ‘C’.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCC6249693FEF11FF36AE10FC53277F" blockId="25.[189,1399,267,540]" pageId="25" pageNumber="26">
|
||
Teeth arranged in V-shaped rows, each row contains about 166 (84-(7-10)-1-(8-12)-81) teeth. Central tooth tricuspid; mesocone of triangular shape; ectocones small, with pointed cusp. Lateral teeth tricuspid; endocone small and rounded cusps; mesocone large and truncated cusp; ectocone very small, pointed cusp and located at base of teeth. Lateral teeth gradually transformed to elongated marginal teeth. Marginal teeth tricuspid, start around tooth number 8 to 12; endocone elongate with dull cusps and separated from mesocome by wide notch; mesocone large, elongate and curved cusps; ectocone small, with pointed cusps or sometimes serrated shape (
|
||
<figureCitation id="13487ECC693FEF11FD77AD3CFC98277F" box="[764,848,514,540]" captionStart="Fig" captionStartId="24.[189,232,1824,1850]" captionTargetBox="[194,1391,713,1777]" captionTargetId="figure@24.[189,1398,706,1783]" captionTargetPageId="24" captionText="Fig. 12. SEM images of the radula. A–C. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7027). D–F. Amphidromus xiengensis Morlet, 1891 from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7037). G–I. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos (CUMZ 7041). A, D, G = central tooth with the first to fifth or sixth lateral teeth; B, E, H = lateral teeth with the tricuspid marginal teeth transition; C, F, I = outermost marginal teeth. Numbers indicate the order of the lateral and marginal teeth. Central tooth indicated by ‘C’." figureDoi="http://doi.org/10.5281/zenodo.3829768" httpUri="https://zenodo.org/record/3829768/files/figure.png" pageId="25" pageNumber="26">Fig. 12</figureCitation>
|
||
A−C).
|
||
</paragraph>
|
||
<paragraph id="8BCC6249693FEF11FF61AD7CFE6C273F" blockId="25.[189,1399,578,745]" box="[234,420,578,604]" pageId="25" pageNumber="26">
|
||
<emphasis id="B907BE5B693FEF11FF61AD7CFE6C273F" bold="true" box="[234,420,578,604]" pageId="25" pageNumber="26">Genital organs</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC6249693FEF11FF36AD5BFC84278A" blockId="25.[189,1399,578,745]" pageId="25" pageNumber="26">
|
||
Atrium (at) rather long (n = 5). Penis (p) long, cylindrical and enlarged from middle to end. Epiphallus (e) cylindrical, length longer than that of penis; flagellum (fl) short and one fourth as long as epiphallus; appendix absent. Penial retractor muscle (pr) thickened and relatively short. Vas deferens (vd) small tube, connecting epiphallus and free oviduct (
|
||
<figureCitation id="13487ECC693FEF11FD5EADF1FCF7278A" box="[725,831,719,745]" captionStart="Fig" captionStartId="22.[189,232,1844,1870]" captionTargetBox="[189,1399,298,1799]" captionTargetId="figure@22.[189,1399,298,1799]" captionTargetPageId="22" captionText="Fig. 11. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7023). C–D. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vagina chamber (CUMZ 7027)." figureDoi="http://doi.org/10.5281/zenodo.3829766" httpUri="https://zenodo.org/record/3829766/files/figure.png" pageId="25" pageNumber="26">Fig. 11C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC6249693FEF11FF36AC31FBCC262F" blockId="25.[189,1398,783,845]" pageId="25" pageNumber="26">
|
||
Internal penial wall corrugated into thin penial pilasters, which form fringe around penial verge. Penial verge conical, with smooth surface and orifice open near tip (
|
||
<figureCitation id="13487ECC693FEF11FC00AC0CFC3E262F" box="[907,1014,818,844]" captionStart="Fig" captionStartId="22.[189,232,1844,1870]" captionTargetBox="[189,1399,298,1799]" captionTargetId="figure@22.[189,1399,298,1799]" captionTargetPageId="22" captionText="Fig. 11. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7023). C–D. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vagina chamber (CUMZ 7027)." figureDoi="http://doi.org/10.5281/zenodo.3829766" httpUri="https://zenodo.org/record/3829766/files/figure.png" pageId="25" pageNumber="26">Fig. 11D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC6249693FEF11FF36AC4CFE052695" blockId="25.[189,1399,882,1014]" pageId="25" pageNumber="26">
|
||
Vagina cylindrical, long, slender about two times as long as penis length. Gametolytic duct long, proximal to genital orifice almost same diameter as vagina, distally tapering; distal to genital orifice small tube of about same length as proximal part and connected to gametolytic sac. Oviduct and albumin gland small (
|
||
<figureCitation id="13487ECC693FEF11FEDEACE2FE772695" box="[341,447,988,1014]" captionStart="Fig" captionStartId="22.[189,232,1844,1870]" captionTargetBox="[189,1399,298,1799]" captionTargetId="figure@22.[189,1399,298,1799]" captionTargetPageId="22" captionText="Fig. 11. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7023). C–D. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vagina chamber (CUMZ 7027)." figureDoi="http://doi.org/10.5281/zenodo.3829766" httpUri="https://zenodo.org/record/3829766/files/figure.png" pageId="25" pageNumber="26">Fig. 11C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC6249693FEF11FF36AB22FEF72139" blockId="25.[189,1399,1052,1114]" pageId="25" pageNumber="26">
|
||
Internally, vaginal wall shows longitudinal vaginal pilasters (vp); pilasters very narrow and thin (
|
||
<figureCitation id="13487ECC693FEF11FF4DAB01FEF92139" box="[198,305,1087,1114]" captionStart="Fig" captionStartId="22.[189,232,1844,1870]" captionTargetBox="[189,1399,298,1799]" captionTargetId="figure@22.[189,1399,298,1799]" captionTargetPageId="22" captionText="Fig. 11. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7023). C–D. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vagina chamber (CUMZ 7027)." figureDoi="http://doi.org/10.5281/zenodo.3829766" httpUri="https://zenodo.org/record/3829766/files/figure.png" pageId="25" pageNumber="26">Fig. 11D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36931C2693FEF11FF36AB41FB6C21AB" box="[189,1188,1151,1225]" pageId="25" pageNumber="26" type="distribution">
|
||
<paragraph id="8BCC6249693FEF11FF36AB41FE9E21FA" blockId="25.[189,342,1151,1177]" box="[189,342,1151,1177]" pageId="25" pageNumber="26">
|
||
<heading id="D084D525693FEF11FF36AB41FE9E21FA" bold="true" box="[189,342,1151,1177]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
|
||
<emphasis id="B907BE5B693FEF11FF36AB41FE9E21FA" bold="true" box="[189,342,1151,1177]" pageId="25" pageNumber="26">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCC6249693FEF11FF36AB90FB6C21AB" blockId="25.[189,1188,1198,1225]" box="[189,1188,1198,1225]" pageId="25" pageNumber="26">
|
||
The distribution range of the species is from
|
||
<collectingRegion id="49B7ACAB693FEF11FD40AB90FCF321AB" box="[715,827,1198,1224]" country="Laos" name="Viangchan" pageId="25" pageNumber="26">Vientiane</collectingRegion>
|
||
to Luang Phrabang Provinces.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36931C2693FEF11FF36ABCEFB772288" pageId="25" pageNumber="26" type="discussion">
|
||
<paragraph id="8BCC6249693FEF11FF36ABCEFEE72069" blockId="25.[189,303,1264,1290]" box="[189,303,1264,1290]" pageId="25" pageNumber="26">
|
||
<heading id="D084D525693FEF11FF36ABCEFEE72069" bold="true" box="[189,303,1264,1290]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
|
||
<emphasis id="B907BE5B693FEF11FF36ABCEFEE72069" bold="true" box="[189,303,1264,1290]" pageId="25" pageNumber="26">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCC6249693FEF11FF36AA20FAF5234C" blockId="25.[189,1399,1310,1584]" pageId="25" pageNumber="26">
|
||
<taxonomicName id="4C7319CA693FEF11FF36AA20FE65205B" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[189,429,1310,1336]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="flavus">
|
||
<emphasis id="B907BE5B693FEF11FF36AA20FE65205B" box="[189,429,1310,1336]" italics="true" pageId="25" pageNumber="26">Amphidromus flavus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C7319CA693FEF11FDC5AA21FC45205A" authority="(Benson, 1851)" baseAuthorityName="Benson" baseAuthorityYear="1851" box="[590,909,1310,1337]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="sinensis">
|
||
<emphasis id="B907BE5B693FEF11FDC5AA21FD05205B" box="[590,717,1311,1337]" italics="true" pageId="25" pageNumber="26">A. sinensis</emphasis>
|
||
(Benson, 1851)
|
||
</taxonomicName>
|
||
in having a smaller shell, the spire more ovate and conical, with a single reddish band on the penultimate whorl. It can be distinguished from
|
||
<taxonomicName id="4C7319CA693FEF11FAD6AA7CFEFF201C" authorityName="Morlet" authorityYear="1891" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="xiengensis">
|
||
<emphasis id="B907BE5B693FEF11FAD6AA7CFEFF201C" italics="true" pageId="25" pageNumber="26">A. xiengensis</emphasis>
|
||
</taxonomicName>
|
||
in having a smaller shell, ovate conical, without any band on yellowish ground color or on shell sculpture. It differs from
|
||
<taxonomicName id="4C7319CA693FEF11FDABAAB6FC3A20C0" authority="Sutcharit & Panha, 2015" authorityName="Sutcharit & Panha" authorityYear="2015" box="[544,1010,1416,1443]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="globonevilli">
|
||
<emphasis id="B907BE5B693FEF11FDABAAB6FD0420C1" box="[544,716,1416,1442]" italics="true" pageId="25" pageNumber="26">A. globonevilli</emphasis>
|
||
<bibRefCitation id="EFE21FB8693FEF11FD58AAB6FC3A20C0" author="Sutcharit C. & Ablett J. & Tongkerd P. & Naggs F. & Panha S." box="[723,1010,1416,1443]" pageId="25" pageNumber="26" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. & Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">Sutcharit & Panha, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
by having a larger shell size, and it is smaller than
|
||
<taxonomicName id="4C7319CA693FEF11FE0EAA92FC8620A5" authority="Sutcharit & Panha, 2015" authorityName="Sutcharit & Panha" authorityYear="2015" box="[389,846,1451,1478]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="principalis">
|
||
<emphasis id="B907BE5B693FEF11FE0EAA92FDEC20A6" box="[389,548,1451,1478]" italics="true" pageId="25" pageNumber="26">A. principalis</emphasis>
|
||
<bibRefCitation id="EFE21FB8693FEF11FDA0AA95FC8620A5" author="Sutcharit C. & Ablett J. & Tongkerd P. & Naggs F. & Panha S." box="[555,846,1451,1478]" pageId="25" pageNumber="26" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. & Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">Sutcharit & Panha, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
. In comparison,
|
||
<taxonomicName id="4C7319CA693FEF11FB84AA92FB7320A6" authorityName="Sutcharit & Panha" authorityYear="2015" box="[1039,1211,1451,1478]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="globonevilli">
|
||
<emphasis id="B907BE5B693FEF11FB84AA92FB7320A6" box="[1039,1211,1451,1478]" italics="true" pageId="25" pageNumber="26">A. globonevilli</emphasis>
|
||
</taxonomicName>
|
||
has an elongate conical shell and a faint yellow spiral band below the periphery, while
|
||
<taxonomicName id="4C7319CA693FEF11FC67AAF1FB42208A" authorityName="Sutcharit & Panha" authorityYear="2015" box="[1004,1162,1487,1513]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="principalis">
|
||
<emphasis id="B907BE5B693FEF11FC67AAF1FB42208A" box="[1004,1162,1487,1513]" italics="true" pageId="25" pageNumber="26">A. principalis</emphasis>
|
||
</taxonomicName>
|
||
has a more elongate conical shell, last whorl without any band, aperture ovate. In our collection we found two
|
||
<typeStatus id="54C8DCEB693FEF11FB4FAACCFACA236F" box="[1220,1282,1522,1548]" pageId="25" pageNumber="26">types</typeStatus>
|
||
identified as
|
||
<taxonomicName id="4C7319CA693FEF11FF56A928FE89234C" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[221,321,1557,1584]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="flavus">
|
||
<emphasis id="B907BE5B693FEF11FF56A928FE89234C" box="[221,321,1557,1584]" italics="true" pageId="25" pageNumber="26">A. flavus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7319CA693FEF11FEF0A928FE17234C" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[379,479,1557,1584]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="flavus">
|
||
<emphasis id="B907BE5B693FEF11FEF0A928FE17234C" box="[379,479,1557,1584]" italics="true" pageId="25" pageNumber="26">A. flavus</emphasis>
|
||
</taxonomicName>
|
||
var. “
|
||
<emphasis id="B907BE5B693FEF11FDACA928FD40234C" box="[551,648,1558,1583]" italics="true" pageId="25" pageNumber="26">proxima</emphasis>
|
||
”, living sympatrically at Luang Phrabang and Vang Vieng.
|
||
</paragraph>
|
||
<paragraph id="8BCC6249693FEF11FF36A969FE4423F7" blockId="25.[189,1398,1623,1684]" pageId="25" pageNumber="26">
|
||
Three subspecific entities have been proposed; however, we recognize them as a single biological species,
|
||
<taxonomicName id="4C7319CA693FEF11FEABA944FE4D23F7" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[288,389,1658,1684]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="flavus">
|
||
<emphasis id="B907BE5B693FEF11FEABA944FE4D23F7" box="[288,389,1658,1684]" italics="true" pageId="25" pageNumber="26">A. flavus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCC6249693FEF11FF36A992FDAE2388" blockId="25.[189,1399,1708,1771]" pageId="25" pageNumber="26">
|
||
1. Typical form: monochrome yellowish shell and Bands 1 to 6 absent (
|
||
<bibRefCitation id="EFE21FB8693FEF11FB84A992FACE23A4" author="Sutcharit C. & Ablett J. & Tongkerd P. & Naggs F. & Panha S." box="[1039,1286,1708,1735]" pageId="25" pageNumber="26" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. & Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">
|
||
Sutcharit
|
||
<emphasis id="B907BE5B693FEF11FB08A993FB7023A5" box="[1155,1208,1708,1734]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
: fig. 7i–j, for the
|
||
<typeStatus id="54C8DCEB693FEF11FE99A9EEFEB62389" box="[274,382,1744,1770]" pageId="25" pageNumber="26" type="lectotype">lectotype</typeStatus>
|
||
and
|
||
<typeStatus id="54C8DCEB693FEF11FE3DA9EEFD922388" box="[438,602,1744,1771]" pageId="25" pageNumber="26" type="paralectotype">paralectotype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC6249693FEF11FF36A83CFC8C2223" blockId="25.[189,1398,1794,1856]" pageId="25" pageNumber="26">
|
||
2. var. “
|
||
<emphasis id="B907BE5B693FEF11FE9CA83DFEB0227F" box="[279,376,1795,1820]" italics="true" pageId="25" pageNumber="26">proxima</emphasis>
|
||
”
|
||
<bibRefCitation id="EFE21FB8693FEF11FE00A83CFDEC227F" author="Fulton H. C." box="[395,548,1794,1820]" pageId="25" pageNumber="26" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896</bibRefCitation>
|
||
: monochrome yellowish shell, Bands 4 and 5 are indistinct pale brownish bands (
|
||
<bibRefCitation id="EFE21FB8693FEF11FE9AA81BFDCD2223" author="Sutcharit C. & Ablett J. & Tongkerd P. & Naggs F. & Panha S." box="[273,517,1829,1856]" pageId="25" pageNumber="26" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. & Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">
|
||
Sutcharit
|
||
<emphasis id="B907BE5B693FEF11FE0FA818FE7F225C" box="[388,439,1829,1855]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
: fig. 13c, for the
|
||
<typeStatus id="54C8DCEB693FEF11FD40A81BFCFF2223" box="[715,823,1829,1856]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC6249693FEF11FF36A866FC4A22DB" blockId="25.[189,1398,1879,1976]" pageId="25" pageNumber="26">
|
||
3. var. “
|
||
<emphasis id="B907BE5B693FEF11FE95A867FEAC2211" box="[286,356,1881,1906]" italics="true" pageId="25" pageNumber="26">tryoni</emphasis>
|
||
”
|
||
<bibRefCitation id="EFE21FB8693FEF11FEF0A866FDD42211" author="Pilsbry H. A." box="[379,540,1880,1906]" pageId="25" pageNumber="26" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry, 1900</bibRefCitation>
|
||
: Band 1 yellowish, Bands 2 and 3 absent, Bands 4 and 5 brownish and Band 6 absent. In some specimens, brownish blotches of Bands 2 and 3 present in the earlier whorls and disappeared in the penultimate and last whorls (
|
||
<figureCitation id="13487ECC693FEF11FD66A8A0FC8922DB" box="[749,833,1950,1976]" captionStart="Fig" captionStartId="21.[189,232,1752,1778]" captionTargetBox="[189,1399,539,1707]" captionTargetId="figure@21.[189,1399,539,1707]" captionTargetPageId="21" captionText="Fig. 10. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7022). D. Amphidromus begini (Morlet, 1886), syntype from Strung- Trang, Cambodia (MNHN-IM-2000-1832). E-G. Amphidromus flavus (Pfeiffer, 1861) from Tam Pou Kham, Vientiane, Laos (CUMZ 7029). H. Amphidromus flavus var. “indistinctus” Pilsbry, 1900, holotype (ANSP 31486). I–K. Amphidromus flavus var. “tryoni” Pilsbry, 1900. I. Lectotype (ANSP 31488). J. Paralectotype (ANSP 252745). K. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7026). L–M. Amphidromus semitessellatus (Morlet, 1884). L. Lectotype (MNHN-IM-2000-1985). M. Specimen from Cambodia (NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3829764" httpUri="https://zenodo.org/record/3829764/files/figure.png" pageId="25" pageNumber="26">Fig. 10</figureCitation>
|
||
I−K).
|
||
</paragraph>
|
||
<paragraph id="8BCC6249693FEF11FF36A8EFFB772288" blockId="25.[189,1215,2001,2027]" box="[189,1215,2001,2027]" pageId="25" pageNumber="26">
|
||
4. var. “
|
||
<emphasis id="B907BE5B693FEF11FE91A8EFFE692288" box="[282,417,2001,2027]" italics="true" pageId="25" pageNumber="26">indistinctus</emphasis>
|
||
”
|
||
<bibRefCitation id="EFE21FB8693FEF11FE3EA8EFFD9C2288" author="Pilsbry H. A." box="[437,596,2001,2027]" pageId="25" pageNumber="26" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry, 1900</bibRefCitation>
|
||
: identical to var. “
|
||
<emphasis id="B907BE5B693FEF11FCA2A8ECFC422288" box="[809,906,2002,2027]" italics="true" pageId="25" pageNumber="26">proxima</emphasis>
|
||
<bibRefCitation id="EFE21FB8693FEF11FC19A8EFFBE22288" author="Fulton H. C." box="[914,1066,2001,2027]" pageId="25" pageNumber="26" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896</bibRefCitation>
|
||
” (
|
||
<figureCitation id="13487ECC693FEF11FBCEA8EFFB7A2288" box="[1093,1202,2001,2027]" captionStart="Fig" captionStartId="21.[189,232,1752,1778]" captionTargetBox="[189,1399,539,1707]" captionTargetId="figure@21.[189,1399,539,1707]" captionTargetPageId="21" captionText="Fig. 10. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7022). D. Amphidromus begini (Morlet, 1886), syntype from Strung- Trang, Cambodia (MNHN-IM-2000-1832). E-G. Amphidromus flavus (Pfeiffer, 1861) from Tam Pou Kham, Vientiane, Laos (CUMZ 7029). H. Amphidromus flavus var. “indistinctus” Pilsbry, 1900, holotype (ANSP 31486). I–K. Amphidromus flavus var. “tryoni” Pilsbry, 1900. I. Lectotype (ANSP 31488). J. Paralectotype (ANSP 252745). K. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7026). L–M. Amphidromus semitessellatus (Morlet, 1884). L. Lectotype (MNHN-IM-2000-1985). M. Specimen from Cambodia (NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3829764" httpUri="https://zenodo.org/record/3829764/files/figure.png" pageId="25" pageNumber="26">Fig. 10H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |