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<document id="9C711DB8450C99B6CE503BAB49CE6CE5" ID-DOI="10.5281/zenodo.296570" ID-GBIF-Dataset="51d302cc-a68c-4c73-a2a1-37d9f48af87c" ID-ISSN="1175-5326" ID-Zenodo-Dep="296570" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460292317526" checkinUser="plazi" docAuthor="Massa, Bruno &amp; Fontana, Paolo" docDate="2011" docId="03DA1148FFCF180FFF46F98FFCD2AFE9" docLanguage="en" docName="zt02837p047.pdf" docOrigin="Zootaxa 2837" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Sphagniana Zeuner 1941" docType="treatment" docVersion="11" lastPageNumber="15" masterDocId="FFE36930FFC21801FFD1FFA2FFE7AE6E" masterDocTitle="Supraspecific taxonomy of Palaearctic Platycleidini with unarmed prosternum: a morphological approach (Orthoptera: Tettigoniidae, Tettigoniinae)" masterLastPageNumber="47" masterPageNumber="1" pageNumber="14" updateTime="1698251656597" updateUser="plazi">
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<mods:title id="DFD40DB342A9D8FD866E09D5FE82A0CD">Supraspecific taxonomy of Palaearctic Platycleidini with unarmed prosternum: a morphological approach (Orthoptera: Tettigoniidae, Tettigoniinae)</mods:title>
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<mods:namePart id="32A9251CDFBFA8BFDE07BEB91BF5DBDD">Massa, Bruno</mods:namePart>
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<paragraph id="8BCCA05EFFCF180CFF46F98FFCECA829" blockId="13.[151,779,1581,1607]" box="[151,779,1581,1607]" pageId="13" pageNumber="14">
<heading id="D0841732FFCF180CFF46F98FFCECA829" bold="true" box="[151,779,1581,1607]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="B9077C4CFFCF180CFF46F98FFCECA829" bold="true" box="[151,779,1581,1607]" pageId="13" pageNumber="14">
<keyLead id="3082036BFFCF180CFF46F98FFCECA829" box="[151,779,1581,1607]" pageId="13" pageNumber="14">
<keyStep id="3087B8FBFFCF180CFF46F98FFCECA829" box="[151,779,1581,1607]" pageId="13" pageNumber="14">
<key id="FFE2F76FFFCF180CFF46F98FFCECA829" box="[151,779,1581,1607]" pageId="13" pageNumber="14">
13)
<taxonomicName id="4C73DBDDFFCF180CFF14F98FFDE7A829" ID-CoL="8NZD2" authority="Zeuner, 1941" authorityName="Zeuner" authorityYear="1941" box="[197,512,1581,1607]" class="Insecta" family="Tettigoniidae" genus="Sphagniana" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B9077C4CFFCF180CFF14F98FFEBFA829" bold="true" box="[197,344,1581,1607]" italics="true" pageId="13" pageNumber="14">Sphagniana</emphasis>
<bibRefCitation id="EFE2DDAFFFCF180CFE8EF98FFDE7A829" author="Zeuner" box="[351,512,1581,1607]" pageId="13" pageNumber="14" refString="Zeuner, F. E. (1941) The classification of the Decticinae hitherto included in Platycleis Fieb. or Metrioptera Wesm. (Orthoptera, Saltatoria). Transactions of Royal Entomological Society of London, 91, 1 - 50." type="journal article" year="1941">Zeuner, 1941</bibRefCitation>
</taxonomicName>
(
<figureCitation id="1348BCDBFFCF180CFDC1F98FFD94A829" box="[528,627,1581,1607]" captionStart="FIGURES 25 32" captionStartId="32.[151,264,1909,1932]" captionTargetBox="[158,1430,193,1882]" captionTargetId="figure@32.[151,1436,193,1888]" captionTargetPageId="32" captionText="FIGURES 25 32. Lateral view of the pronotum of other 8 species of Platycleidini (Sphagniana, Eobiana, Vichetia, Metrioptera, Bicolorana, Roeseliana, Zeuneriana, Broughtonia)." httpUri="https://zenodo.org/record/296588/files/figure.png" pageId="13" pageNumber="14">Figs. 25</figureCitation>
,
<figureCitation id="1348BCDBFFCF180CFD51F98FFD78A829" box="[640,671,1581,1607]" captionStart="FIGURES 48 63" captionStartId="36.[151,264,1919,1942]" captionTargetBox="[286,1301,193,1898]" captionTargetId="figure@36.[286,1301,193,1898]" captionTargetPageId="36" captionText="FIGURES 48 63. Lateral view of the ovipositor of 16 species of Platycleidini (Platycleis, Sardoplatycleis, Sepiana, Metrioptera, Roeseliana, Bicolorana, Chizuella, Eobiana, Modestana, Sphagniana, Amedegnatiana, Alticolana, Parnassiana, Squamiana, Eumetrioptera)." httpUri="https://zenodo.org/record/296600/files/figure.png" pageId="13" pageNumber="14">57</figureCitation>
,
<figureCitation id="1348BCDBFFCF180CFD7DF98FFD2CA829" box="[684,715,1581,1607]" captionStart="FIGURES 94 101" captionStartId="40.[151,264,1929,1952]" captionTargetBox="[166,1417,193,1899]" captionTargetId="figure@40.[163,1424,193,1908]" captionTargetPageId="40" captionText="FIGURES 94 101. Subgenital plate of the female of other 8 species of Platycleidini (Sepiana, Parnassiana, Amedegnatiana, Modestana, Sphagniana, Squamiana, Uludaghia)." httpUri="https://zenodo.org/record/298466/files/figure.png" pageId="13" pageNumber="14">98</figureCitation>
, 111)
</key>
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</paragraph>
<paragraph id="8BCCA05EFFCF180CFF46F9CFFB73A90E" blockId="13.[151,1436,1645,2033]" pageId="13" pageNumber="14">
<emphasis id="B9077C4CFFCF180CFF46F9CFFEF0A8E8" bold="true" box="[151,279,1645,1670]" italics="true" pageId="13" pageNumber="14">Characters</emphasis>
. Tegmina are generally abbreviated, but rarely macropterous individuals are found. Head just larger than long, ratio maximum width/length of head (from vertex to clipeus): 1.2-1.3. Pronotum depressed, widened and enlarged in the metazona, with lateral lobes bent down at a sharp angle (mainly in males), median keel obliterated; hind femora comparatively short and weak; Ƥ subgenital plate very small, wider than long; ovipositor slender, long and regularly curved near the base; hind femora/pronotum length 3: 2.4-2.5, Ƥ: 2.6-3.0;
<date id="FFCD869EFFCF180CFB4DF8A3FB28A976" box="[1180,1231,1793,1817]" pageId="13" pageNumber="14">3 X</date>
th tergite enlarged, incised in the middle and with two very expanded lobes completely covering cerci; cerci just flattened in the inner spine portion. Colour of lateral lobes of the pronotum is black bordered with a white stripe.
</paragraph>
</subSubSection>
<subSubSection id="C369F3D5FFCF180FFF16F8CEFBBCAF73" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BCCA05EFFCF180FFF16F8CEFBBCAF73" blockId="13.[151,1436,1645,2033]" lastBlockId="14.[151,1437,151,392]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="B9077C4CFFCF180CFF16F8CEFEC9A9EB" bold="true" box="[199,302,1900,1925]" italics="true" pageId="13" pageNumber="14">Remarks</emphasis>
. It is evidently separated from all here considered taxa by its characteristic male processes covering cerci, its pronotum shape and relative short femora. These characters consents definitely to separate it from
<taxonomicName id="4C73DBDDFFCF180CFA85F830FF07A9A3" class="Insecta" family="Tettigoniidae" genus="Metrioptera" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B9077C4CFFCF180CFA85F830FF07A9A3" italics="true" pageId="13" pageNumber="14">Metrioptera</emphasis>
</taxonomicName>
, by some authors considered its synonym. It has been listed by
<bibRefCitation id="EFE2DDAFFFCF180CFC74F817FB4AA9A2" author="Rentz" box="[933,1197,1972,1996]" pageId="13" pageNumber="14" refString="Rentz, D. C. F. &amp; Colless, D. H. (1990) A classification of the Shield-backed Katydids (Tettigoniinae) of the world. In: Bailey W. J. &amp; Rentz D. C. F. (eds.). The Tettigoniidae: Biology, Systematics and Evolution. Springer Verlag, Berlin, 352 - 377." type="book chapter" year="1990">Rentz &amp; Colless (1990)</bibRefCitation>
within the subgenera of
<taxonomicName id="4C73DBDDFFCF180CFF65F878FEDCA99F" box="[180,315,2010,2033]" class="Insecta" family="Tettigoniidae" genus="Metrioptera" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B9077C4CFFCF180CFF65F878FEDCA99F" box="[180,315,2010,2033]" italics="true" pageId="13" pageNumber="14">Metrioptera</emphasis>
</taxonomicName>
, but
<bibRefCitation id="EFE2DDAFFFCF180CFEA2F87AFDA1A99E" box="[371,582,2008,2033]" pageId="13" pageNumber="14" refString="Eades, D. C., Otte, D., Cigliano, M. M. &amp; Braun, H. (2010) Orthoptera Species File Online. Version 2.0 / 4.0 (accessed: December 2010). http: // Orthoptera. SpeciesFile. org." type="book chapter">
Eades
<emphasis id="B9077C4CFFCF180CFE6CF878FE12A99F" box="[445,501,2009,2033]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
(2010)
</bibRefCitation>
consider it a valid genus. We examined only
<taxonomicName id="4C73DBDDFFCF180CFBEFF87BFB01A99F" box="[1086,1254,2009,2033]" class="Insecta" family="Tettigoniidae" genus="Sphagniana" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sphagnorum">
<emphasis id="B9077C4CFFCF180CFBEFF87BFB01A99F" box="[1086,1254,2009,2033]" italics="true" pageId="13" pageNumber="14">S. sphagnorum</emphasis>
</taxonomicName>
, thus we are not able to establish if the other two species reported within this genus have the same characteristics or they should be classified in another way. We only observe that
<taxonomicName id="4C73DBDDFFCC180FFD11FF1FFCABAEBA" box="[704,844,189,212]" class="Insecta" family="Tettigoniidae" genus="Sphagniana" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ussuriana">
<emphasis id="B9077C4CFFCC180FFD11FF1FFCABAEBA" box="[704,844,189,212]" italics="true" pageId="14" pageNumber="15">S. ussuriana</emphasis>
</taxonomicName>
shows a different pronotum and ovipositor pattern. Previously,
<bibRefCitation id="EFE2DDAFFFCC180FFECAFF42FE22AE99" author="Ramme" box="[283,453,223,247]" pageId="14" pageNumber="15" refString="Ramme, W. (1951) Zur Systematik Faunistik und Biologie der Orthopteren von sudost-Europa und Vorderasien. Mitteilungen Zoologischen Museum Berlin, 27, 1 - 431, 29 pl." type="journal article" year="1951">Ramme (1951)</bibRefCitation>
wrote that the classification of
<taxonomicName id="4C73DBDDFFCC180FFCFAFF42FC56AE96" box="[811,945,224,248]" class="Insecta" family="Tettigoniidae" genus="Sphagniana" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B9077C4CFFCC180FFCFAFF42FC56AE96" box="[811,945,224,248]" italics="true" pageId="14" pageNumber="15">Sphagniana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C73DBDDFFCC180FFC39FF42FBBAAE96" box="[1000,1117,224,248]" class="Insecta" family="Tettigoniidae" genus="Alticolana" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B9077C4CFFCC180FFC39FF42FBBAAE96" box="[1000,1117,224,248]" italics="true" pageId="14" pageNumber="15">Alticolana</emphasis>
</taxonomicName>
was not clear and their species appeared much heterogeneous, some of them probably belonging to other genera.
</paragraph>
</subSubSection>
<subSubSection id="C369F3D5FFCC180FFF16FE85FCD2AFE9" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8BCCA05EFFCC180FFF16FE85FCD2AFE9" blockId="14.[151,1437,151,392]" pageId="14" pageNumber="15">
<emphasis id="B9077C4CFFCC180FFF16FE85FEB4AF2E" bold="true" box="[199,339,295,320]" italics="true" pageId="14" pageNumber="15">Distribution</emphasis>
. It is the only taxon, among those here considered, covering E Asia and N
<collectingCountry id="F364E0CEFFCC180FFB4AFE85FB18AF51" box="[1179,1279,295,319]" name="United States of America" pageId="14" pageNumber="15">America</collectingCountry>
.
<emphasis id="B9077C4CFFCC180FFF16FEEEFDE4AF0A" bold="true" box="[199,515,331,356]" pageId="14" pageNumber="15">Number of species known</emphasis>
(after
<bibRefCitation id="EFE2DDAFFFCC180FFD85FEEEFCC4AF0A" author="Eades" box="[596,803,332,356]" pageId="14" pageNumber="15" refString="Eades, D. C., Otte, D., Cigliano, M. M. &amp; Braun, H. (2010) Orthoptera Species File Online. Version 2.0 / 4.0 (accessed: December 2010). http: // Orthoptera. SpeciesFile. org." type="book chapter" year="2010">
Eades
<emphasis id="B9077C4CFFCC180FFD70FEEFFD3AAF0A" box="[673,733,332,356]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2010
</bibRefCitation>
): 3: 1)
<taxonomicName id="4C73DBDDFFCC180FFCAAFEEEFB38AF0A" authority="Kim et Kim, 2001" authorityName="Kim et Kim" authorityYear="2001" box="[891,1247,332,356]" class="Insecta" family="Tettigoniidae" genus="Metrioptera" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B9077C4CFFCC180FFCAAFEEEFC0CAF0A" box="[891,1003,332,356]" italics="true" pageId="14" pageNumber="15">monticola</emphasis>
(Kim et Kim, 2001)
</taxonomicName>
; 2)
<taxonomicName id="4C73DBDDFFCC180FFAC1FEEEFED9AFE9" authority="Walker, 1869" authorityName="Walker" authorityYear="1869" class="Insecta" family="Tettigoniidae" genus="Sphagniana" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="sphagnorum">
<emphasis id="B9077C4CFFCC180FFAC1FEEEFA7BAF0A" box="[1296,1436,332,356]" italics="true" pageId="14" pageNumber="15">sphagnorum</emphasis>
(Walker, 1869)
</taxonomicName>
(
<typeStatus id="54C81EFCFFCC180FFE9CFECDFE98AFE9" box="[333,383,367,391]" pageId="14" pageNumber="15">type</typeStatus>
species); 3)
<taxonomicName id="4C73DBDDFFCC180FFDDCFED3FCD6AFE9" authority="Uvarov, 1926" authorityName="Uvarov" authorityYear="1926" box="[525,817,367,392]" class="Insecta" family="Tettigoniidae" genus="Sphagniana" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ussuriana">
<emphasis id="B9077C4CFFCC180FFDDCFED3FD9BAFE6" box="[525,636,369,392]" italics="true" pageId="14" pageNumber="15">ussuriana</emphasis>
(Uvarov, 1926)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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