427 lines
60 KiB
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427 lines
60 KiB
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<document id="BDF7FEEB15939156CF7E552AD7FABE43" ID-CLB-Dataset="4210" ID-DOI="10.5852/ejt.2021.767.1495" ID-GBIF-Dataset="67bc6a73-4137-4de3-9dc4-ce87972f742d" ID-ISSN="2118-9773" ID-Zenodo-Dep="5514839" ID-ZooBank="9F019E02-95E7-4B6A-B5BC-01F8E26F3466" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1631927261920" checkinUser="felipe" docAuthor="Nahok, Benchawan, Tumpeesuwan, Sakboworn & Tumpeesuwan, Chanidaporn" docDate="2021" docId="03AC87E7C320FFDF9F41FD0890C2BA73" docLanguage="en" docName="ejt-767_142-166.pdf" docOrigin="European Journal of Taxonomy 767 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.767.1495" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Landouria chloritoides Nahok & Tumpeesuwan & Tumpeesuwan 2021, sp. nov." docType="treatment" docUuid="740CA614-D17B-47A3-A719-7DFC7D2CE2D5" docUuidSource="ZooBank" docVersion="11" lastPageNumber="158" masterDocId="FF95FF9FC32EFFCF9D1FFF9B931BBE42" masterDocTitle="Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902)" masterLastPageNumber="166" masterPageNumber="142" pageNumber="156" updateTime="1699449512379" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="777927C40D8D68D7D66CD8995A6CB93E">Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902)</mods:title>
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<mods:namePart id="73BBC5BECF93A94AF9E5C564FE7CC63D">Nahok, Benchawan</mods:namePart>
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<mods:affiliation id="CC812C814125E7DA224842C60453B607">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand. & Educational Research Development and Demonstration Institute, Srinakharinwirot University, Ongkharak District, Nakhon Nayok, 26120 Thailand.</mods:affiliation>
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<mods:nameIdentifier id="8EB81BB1B0B96BD0015C4894B88489F5" type="email">nahok.b@gmail.com</mods:nameIdentifier>
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<mods:namePart id="19759F54D6BF6FBC5D0819768CC8B77F">Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:affiliation id="48F50C06C293B34243C2823E1C215BA5">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand. & Palaeontological Research and Education Centre, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand.</mods:affiliation>
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<mods:nameIdentifier id="1DE7C4AB29FC4F0865D10B2ECB8C2962" type="email">stumpeesuwan@yahoo.com</mods:nameIdentifier>
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<mods:affiliation id="A0E7A416F2E0A6EDC104FD834FA78896">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand.</mods:affiliation>
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<mods:nameIdentifier id="E2B3EF94A1A00FBC4653310A1F725490" type="email">ctumpeesuwan@yahoo.com</mods:nameIdentifier>
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<taxonomicName id="4C054D72C320FFC19F41FD08906FBCEC" ID-CoL="8QFT4" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[606,884,659,686]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="156" phylum="Mollusca" rank="species" species="chloritoides" status="sp. nov.">
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<emphasis id="B971EAE3C320FFC19F41FD08906FBCEC" bold="true" box="[606,884,659,686]" italics="true" pageId="14" pageNumber="156">Landouria chloritoides</emphasis>
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<paragraph id="8BBA36F1C320FFC19F53FD4190FCBCB6" blockId="14.[358,1230,659,757]" box="[588,999,730,757]" pageId="14" pageNumber="156">
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<figureCitation id="133E2A74C320FFC19F53FD4191B7BCB6" box="[588,684,730,756]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="14" pageNumber="156">Figs 3D</figureCitation>
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,
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<figureCitation id="133E2A74C320FFC19FA6FD4191C4BCB6" box="[697,735,730,756]" captionStart="Fig" captionStartId="12.[189,232,1921,1947]" captionTargetBox="[193,1395,278,1870]" captionTargetId="figure-12@12.[193,1395,265,1883]" captionTargetPageId="12" captionText="Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514847" httpUri="https://zenodo.org/record/5514847/files/figure.png" pageId="14" pageNumber="156">4D</figureCitation>
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,
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<figureCitation id="133E2A74C320FFC19FF4FD419009BCB7" box="[747,786,730,757]" captionStart="Fig" captionStartId="15.[189,232,1816,1842]" captionTargetBox="[283,1352,278,1767]" captionTargetId="figure-15@15.[232,1356,265,1778]" captionTargetPageId="15" captionText="Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514849" httpUri="https://zenodo.org/record/5514849/files/figure.png" pageId="14" pageNumber="156">5D</figureCitation>
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,
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<figureCitation id="133E2A74C320FFC19E01FD419047BCB6" box="[798,860,730,756]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="14" pageNumber="156">6J–L</figureCitation>
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;
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<tableCitation id="C687034AC320FFC19E78FD4190FCBCB6" box="[871,999,730,756]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStart-3="Table 4" captionStartId-0="2.[185,255,857,885]" captionStartId-1="4.[186,256,600,628]" captionStartId-2="6.[185,255,695,723]" captionStartId-3="10.[185,255,544,572]" captionTargetBox-0="[195,1921,987,1444]" captionTargetBox-1="[197,1907,827,1685]" captionTargetBox-2="[212,1909,868,1619]" captionTargetBox-3="[196,1908,758,1739]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="10" captionText-0="Table 1. Shell measurements (mm) of six new species of the genus Landouria Godwin-Austen, 1918 from northeastern Thailand." captionText-1="Table 2. Locality data and identification of specimens of Landouria Godwin-Austen, 1918 used for mtDNA sequencing, including collection catalogue numbers and GenBank accession numbers. Abbreviations: NHMSU = Natural History Museum, Mahasarakham University; HNHM = Hungarian Natural History Museum; AM = Australian Museum. References: 1 = Tumpeesuwan & Tumpeesuwan (2019); 2 = Hirano et al. (2014); 3 = Köhler et al. (2018)." captionText-2="Table 3. Pairwise p-distance values of Landouria Godwin-Austen, 1918 based on mitochondrial COI sequences." captionText-3="Table 4. Comparison of shells, radulae, and genital systems of Landouria spp. from northeastern Thailand. References: 1 = Tumpeesuwan & Tumpeesuwan (2019); 2 = Nahok (2020); 3 = this study. *In Tumpeesuwan & Tumpeesuwan (2019) indicated the number of teeth per row in Landouria strobiloides varies from 3 to 53. This lowest row has only 3 teeth. It is the uppermost row, probably lacerated and peel off. Unfortunately, we could not re-checked the complete specimens in this time being, so, this point need further investigation and proof." httpUri-0="http://table.plazi.org/id/DF7A6679C32CFFCD9DA6FCC295B0BD37" httpUri-1="http://table.plazi.org/id/DF7A6679C32AFFCB9DA5FDC39249BC9F" httpUri-2="http://table.plazi.org/id/DF7A6679C328FFC99DA6FD2C96FFBC91" httpUri-3="http://table.plazi.org/id/DF7A6679C324FFC59DA6FDBB969CBCE7" pageId="14" pageNumber="156" tableUuid-0="DF7A6679C32CFFCD9DA6FCC295B0BD37" tableUuid-1="DF7A6679C32AFFCB9DA5FDC39249BC9F" tableUuid-2="DF7A6679C328FFC99DA6FD2C96FFBC91" tableUuid-3="DF7A6679C324FFC59DA6FDBB969CBCE7">Tables 1–4</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31F657AC320FFC19DA2FC87923FBDE2" pageId="14" pageNumber="156" type="reference_group">
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<paragraph id="8BBA36F1C320FFC19DA2FC87967DBD75" blockId="14.[189,1399,796,928]" box="[189,1382,796,823]" pageId="14" pageNumber="156">
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<taxonomicName id="4C054D72C320FFC19DA2FC87926BBD74" box="[189,368,796,822]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="156" phylum="Mollusca" rank="species" species="undefined-1">
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<emphasis id="B971EAE3C320FFC19DA2FC879222BD74" box="[189,313,796,822]" italics="true" pageId="14" pageNumber="156">Landouria</emphasis>
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sp.1
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</taxonomicName>
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–
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<treatmentCitation id="0AA410E0C320FFC19C91FC8791A0BD74" author="Ounchareon J." box="[398,699,796,823]" page="29" pageId="14" pageNumber="156" year="2015">
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<bibRefCitation id="EF944B00C320FFC19C91FC8791A0BD74" author="Ounchareon J." box="[398,699,796,823]" pageId="14" pageNumber="156" refId="ref10673" refString="Ounchareon J. 2015. Taxonomy of Land Snail Genus Landouria Godwin-Austen, 1918 in Northeastern Thailand. Unpublished Master thesis. Department Faculty of Science, Mahasakham University, Maha Sarakham. [In Thai with English abstract.]" type="book" year="2015">Ounchareon 2015: 29–32</bibRefCitation>
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, 44–50, 54–56, 59, 61, figs 4.2c, 4.3c, 4.4. 4.5i–l, 5.5, 5.7.
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</paragraph>
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<paragraph id="8BBA36F1C320FFC19DA2FCA49725BD18" blockId="14.[189,1399,796,928]" box="[189,1086,831,858]" pageId="14" pageNumber="156">
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<treatmentCitationGroup id="AB1511DFC320FFC19DA2FCA49725BD18" box="[189,1086,831,858]" pageId="14" pageNumber="156">
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<taxonomicName id="4C054D72C320FFC19DA2FCA4926ABD1B" box="[189,369,831,857]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="156" phylum="Mollusca" rank="species" species="undefined-2">
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<emphasis id="B971EAE3C320FFC19DA2FCA49222BD1B" box="[189,313,831,857]" italics="true" pageId="14" pageNumber="156">Landouria</emphasis>
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sp.2
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</taxonomicName>
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–
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<treatmentCitation id="0AA410E0C320FFC19C91FCA491DEBD1B" author="Buddharaksa W." box="[398,709,831,857]" page="37" pageId="14" pageNumber="156" year="2016">
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<bibRefCitation id="EF944B00C320FFC19C91FCA491DEBD1B" author="Buddharaksa W." box="[398,709,831,857]" pageId="14" pageNumber="156" refId="ref9870" refString="Buddharaksa W. 2016. Reproductive Anatomy of Some Land Pulmonate Snails in Northeastern Thailand. Unpublished Bachelor's Degree Science Project. Department Faculty of Science, Mahasarakham University, Maha Sarakham. [In Thai with English Abstract.]" type="book" year="2016">Buddharaksa 2016: 37–38</bibRefCitation>
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, 41–42, figs 4.19, 4.20, 5.1, 5.2.
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|
||
<treatmentCitationGroup id="AB1511DFC320FFC19DA2FCF9923FBDE2" pageId="14" pageNumber="156">
|
||
<taxonomicName id="4C054D72C320FFC19DA2FCF99269BD3E" box="[189,370,866,892]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="156" phylum="Mollusca" rank="species" species="undefined-4">
|
||
<emphasis id="B971EAE3C320FFC19DA2FCF99222BD3E" box="[189,313,866,892]" italics="true" pageId="14" pageNumber="156">Landouria</emphasis>
|
||
sp.4
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AA410E0C320FFC19C8FFCF991FFBD3F" author="Yingkhamhang A." box="[400,740,866,893]" page="25" pageId="14" pageNumber="156" year="2016">
|
||
<bibRefCitation id="EF944B00C320FFC19C8FFCF991FFBD3F" author="Yingkhamhang A." box="[400,740,866,893]" pageId="14" pageNumber="156" refId="ref11408" refString="Yingkhamhang A. 2016. Shell Morphology and Reproductive Anatomy of Land Snails Genus Landouria Godwin-Austen, 1918 in Northeastern Thailand. Unpublished Bachelor's Degree Science Project. Department Faculty of Science, Mahasakham University, Maha Sarakham. [In Thai with English abstract.]" type="book" year="2016">Yingkhamhang 2016: 25–26</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 39–41, 43–45, 49–52, figs 4.7, 4.8, 4.21d, 4.22d, 5.3a, 5.4a.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC320FFC19DA2FC5C92A2BA3B" pageId="14" pageNumber="156" type="diagnosis">
|
||
<paragraph id="8BBA36F1C320FFC19DA2FC5C922DBDA3" blockId="14.[189,310,967,993]" box="[189,310,967,993]" pageId="14" pageNumber="156">
|
||
<heading id="D0F2819DC320FFC19DA2FC5C922DBDA3" bold="true" box="[189,310,967,993]" fontSize="11" level="3" pageId="14" pageNumber="156" reason="3">
|
||
<emphasis id="B971EAE3C320FFC19DA2FC5C922DBDA3" bold="true" box="[189,310,967,993]" pageId="14" pageNumber="156">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C320FFC19DA2FC6E92A2BA3B" blockId="14.[188,1399,1013,1145]" pageId="14" pageNumber="156">
|
||
Shell shape similar to that of the genus
|
||
<taxonomicName id="4C054D72C320FFC19F9EFC6E9738BA52" authority="Beck, 1837" authorityName="Beck" authorityYear="1837" box="[641,1059,1013,1040]" class="Gastropoda" family="Camaenidae" genus="Chloritis" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="156" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B971EAE3C320FFC19F9EFC6E91F2BA4D" box="[641,745,1013,1039]" italics="true" pageId="14" pageNumber="156">Chloritis</emphasis>
|
||
Beck, 1837 (Camaenidae)
|
||
</taxonomicName>
|
||
, but with last whorl stout and usually slightly angular, peripheral keel blunt, suture deep, aperture oval and oblique. Flagellum small ovate, penis very large and stout compared to vagina; vagina and free oviduct short (
|
||
<figureCitation id="133E2A74C320FFC199B0FBA79615BA15" box="[1199,1294,1084,1111]" captionStart="Fig" captionStartId="15.[189,232,1816,1842]" captionTargetBox="[283,1352,278,1767]" captionTargetId="figure-15@15.[232,1356,265,1778]" captionTargetPageId="15" captionText="Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514849" httpUri="https://zenodo.org/record/5514849/files/figure.png" pageId="14" pageNumber="156">Fig. 5D</figureCitation>
|
||
). Radula with lanceolate teeth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC320FFC19DA2FB3A91E7BB4F" pageId="14" pageNumber="156" type="etymology">
|
||
<paragraph id="8BBA36F1C320FFC19DA2FB3A925FBAF9" blockId="14.[189,324,1185,1211]" box="[189,324,1185,1211]" pageId="14" pageNumber="156">
|
||
<heading id="D0F2819DC320FFC19DA2FB3A925FBAF9" bold="true" box="[189,324,1185,1211]" fontSize="11" level="3" pageId="14" pageNumber="156" reason="3">
|
||
<emphasis id="B971EAE3C320FFC19DA2FB3A925FBAF9" bold="true" box="[189,324,1185,1211]" pageId="14" pageNumber="156">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C320FFC19DA2FB5491E7BB4F" blockId="14.[189,1399,1231,1293]" pageId="14" pageNumber="156">
|
||
The specific epithet ‘
|
||
<taxonomicName id="4C054D72C320FFC19CA4FB54915CBAAB" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[443,583,1231,1257]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="156" phylum="Mollusca" rank="species" species="chloritoides">
|
||
<emphasis id="B971EAE3C320FFC19CA4FB54915CBAAB" box="[443,583,1231,1257]" italics="true" pageId="14" pageNumber="156">chloritoides</emphasis>
|
||
</taxonomicName>
|
||
’ refers to the conchological similarity between the shell of the new species and that of the camaenid genus
|
||
<taxonomicName id="4C054D72C320FFC19F93FB6991EFBB4E" authorityName="Beck" authorityYear="1837" box="[652,756,1266,1292]" class="Gastropoda" family="Camaenidae" genus="Chloritis" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="156" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B971EAE3C320FFC19F93FB6991EFBB4E" box="[652,756,1266,1292]" italics="true" pageId="14" pageNumber="156">Chloritis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC320FFC19DA2FAAF90C6B943" pageId="14" pageNumber="156" type="materials_examined">
|
||
<paragraph id="8BBA36F1C320FFC19DA2FAAF9277BB0F" blockId="14.[189,364,1331,1358]" box="[189,364,1331,1358]" pageId="14" pageNumber="156">
|
||
<heading id="D0F2819DC320FFC19DA2FAAF9277BB0F" bold="true" box="[189,364,1331,1358]" fontSize="11" level="3" pageId="14" pageNumber="156" reason="3">
|
||
<emphasis id="B971EAE3C320FFC19DA2FAAF9277BB0F" bold="true" box="[189,364,1331,1358]" pageId="14" pageNumber="156">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C320FFC19DF5FAF9966CBBA4" blockId="14.[189,1399,1378,1510]" pageId="14" pageNumber="156">
|
||
<materialsCitation id="3B6D3CACC320FFC19DF5FAF9966CBBA4" ID-GBIF-Occurrence="3357769306" collectingDate="2017-06-09" collectionCode="UW" collectorName="B. Nahok & S. Tumpeesuwan & C. Tumpeesuwan" country="Thailand" elevation="406" latitude="16.758055" location="Wat Thep Udom Wanaram" longLatPrecision="15" longitude="101.96529" municipality="Chum Phae District" pageId="14" pageNumber="156" specimenCode="NHMSU-00029" specimenCount="1" stateProvince="Khon Kaen Province" typeStatus="holotype">
|
||
<emphasis id="B971EAE3C320FFC19DF5FAF99247BB3E" bold="true" box="[234,348,1378,1404]" pageId="14" pageNumber="156">
|
||
<typeStatus id="54BE8853C320FFC19DF5FAF99247BB3E" box="[234,348,1378,1404]" pageId="14" pageNumber="156" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F3127661C320FFC19DA2FA1D9248BBE2" box="[189,339,1414,1440]" name="Thailand" pageId="14" pageNumber="156">THAILAND</collectingCountry>
|
||
•
|
||
<specimenCount id="9D03FD78C320FFC19C6FFA1E92BCBBDD" box="[368,423,1413,1439]" pageId="14" pageNumber="156">shell</specimenCount>
|
||
(SH = 12.0 mm, SW = 19.0 mm, AH =
|
||
<quantity id="4CFD9B14C320FFC19E8FFA1D90F5BBE2" box="[912,1006,1414,1440]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="14" pageNumber="156" unit="mm" value="7.7">7.7 mm</quantity>
|
||
, AW =
|
||
<quantity id="4CFD9B14C320FFC19954FA1E97B2BBE2" box="[1099,1193,1413,1440]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="14" pageNumber="156" unit="mm" value="7.3">7.3 mm</quantity>
|
||
, UW =
|
||
<quantity id="4CFD9B14C320FFC19815FA1E9672BBE2" box="[1290,1385,1413,1440]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="14" pageNumber="156" unit="mm" value="3.9">3.9 mm</quantity>
|
||
);
|
||
<collectingRegion id="49C1F813C320FFC19DA2FA3392ACBB80" box="[189,439,1448,1475]" country="Thailand" name="Khon Kaen" pageId="14" pageNumber="156">Khon Kaen Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BDEAC8BC320FFC19CDBFA3391ABBB80" box="[452,688,1448,1474]" pageId="14" pageNumber="156">Chum Phae District</collectingMunicipality>
|
||
,
|
||
<location id="8EDA602AC320FFC19FA3FA3290E7BB81" LSID="urn:lsid:plazi:treatment:03AC87E7C320FFDF9F41FD0890C2BA73:8EDA602AC320FFC19FA3FA3290E7BB81" box="[700,1020,1448,1475]" country="Thailand" latitude="16.758055" longLatPrecision="15" longitude="101.96529" municipality="Chum Phae District" name="Wat Thep Udom Wanaram" pageId="14" pageNumber="156" stateProvince="Khon Kaen Province">Wat Thep Udom Wanaram</location>
|
||
;
|
||
<geoCoordinate id="EE315036C320FFC19917FA3C9785BB81" box="[1032,1182,1447,1475]" degrees="16" direction="north" minutes="45" orientation="latitude" pageId="14" pageNumber="156" precision="15" seconds="29" value="16.758055">16°45′29″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE315036C320FFC199B3FA3C966ABB81" box="[1196,1393,1447,1475]" degrees="101" direction="east" minutes="57" orientation="longitude" pageId="14" pageNumber="156" precision="1" seconds="55.03" value="101.96529">101°57′55.03″ E</geoCoordinate>
|
||
; alt.
|
||
<quantity id="4CFD9B14C320FFC19DF9FA57922ABBA4" box="[230,305,1484,1510]" metricMagnitude="2" metricUnit="m" metricValue="4.06" pageId="14" pageNumber="156" unit="m" value="406.0">
|
||
<elevation id="0028D1C2C320FFC19DF9FA57922ABBA4" box="[230,305,1484,1510]" metricMagnitude="2" metricUnit="m" metricValue="4.06" pageId="14" pageNumber="156" unit="m" value="406.0">406 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="FFBB1031C320FFC19C24FA5792D9BBA4" box="[315,450,1484,1510]" pageId="14" pageNumber="156" value="2017-06-09">
|
||
<collectingDate id="EFFFE9D9C320FFC19C24FA5792D9BBA4" box="[315,450,1484,1510]" pageId="14" pageNumber="156" value="2017-06-09">9 Jun. 2017</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F05327C320FFC19CD3FA579126BBA4" box="[460,573,1484,1510]" pageId="14" pageNumber="156">B. Nahok</collectorName>
|
||
,
|
||
<collectorName id="26F05327C320FFC19F59FA579010BBA4" box="[582,779,1484,1510]" pageId="14" pageNumber="156">S. Tumpeesuwan</collectorName>
|
||
and
|
||
<collectorName id="26F05327C320FFC19E20FA57971CBBA4" box="[831,1031,1484,1510]" pageId="14" pageNumber="156">C. Tumpeesuwan</collectorName>
|
||
leg.;
|
||
<specimenCode id="DBA39E8AC320FFC1995BFA579618BBA4" box="[1092,1283,1484,1510]" pageId="14" pageNumber="156">NHMSU-00029</specimenCode>
|
||
. (
|
||
<figureCitation id="133E2A74C320FFC1980AFA579675BBA4" box="[1301,1390,1484,1510]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="14" pageNumber="156">Fig. 3D</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C320FFC19DF5F996915CB82C" blockId="14.[189,1398,1549,1646]" pageId="14" pageNumber="156">
|
||
<materialsCitation id="3B6D3CACC320FFC19DF5F9969158B82C" ID-GBIF-Occurrence="3357769301" accessionNumber="MN449404 and MZ435749" collectingDate="2017-06-09" collectionCode="MN, MZ" collectorName="GenBank No." country="Thailand" elevation="406" latitude="16.758055" location="THAILAND" longLatPrecision="15" longitude="101.96529" municipality="Chum Phae District" pageId="14" pageNumber="156" specimenCode="NHMSU-00030" specimenCount="22" stateProvince="Khon Kaen" typeStatus="paratype">
|
||
<emphasis id="B971EAE3C320FFC19DF5F996927CB865" bold="true" box="[234,359,1549,1575]" pageId="14" pageNumber="156">
|
||
<typeStatus id="54BE8853C320FFC19DF5F996927CB865" box="[234,359,1549,1575]" pageId="14" pageNumber="156" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F3127661C320FFC19DA2F9AA924EB809" box="[189,341,1585,1611]" name="Thailand" pageId="14" pageNumber="156">THAILAND</collectingCountry>
|
||
•
|
||
<specimenCount id="9D03FD78C320FFC19C72F9AB92D2B808" box="[365,457,1584,1610]" pageId="14" pageNumber="156">8 shells</specimenCount>
|
||
,
|
||
<specimenCount id="9D03FD78C320FFC19CCBF9AB919CB809" box="[468,647,1584,1611]" pageId="14" pageNumber="156">14 living specs</specimenCount>
|
||
; same collection data as for holotype; GenBank No.
|
||
<accessionNumber id="9456AB12C320FFC199F0F9AB966DB808" box="[1263,1398,1584,1610]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN449404" pageId="14" pageNumber="156">MN449404</accessionNumber>
|
||
and
|
||
<accessionNumber id="9456AB12C320FFC19DF0F9C8926CB82C" box="[239,375,1619,1646]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/MZ435749" pageId="14" pageNumber="156" type="GenBank">MZ435749</accessionNumber>
|
||
;
|
||
<specimenCode id="DBA39E8AC320FFC19C9DF9CF9158B82C" box="[386,579,1620,1646]" pageId="14" pageNumber="156">NHMSU-00030</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C320FFC19DA2F90E92E2B8ED" blockId="14.[189,505,1685,1711]" box="[189,505,1685,1711]" pageId="14" pageNumber="156">
|
||
<heading id="D0F2819DC320FFC19DA2F90E92E2B8ED" bold="true" box="[189,505,1685,1711]" fontSize="11" level="3" pageId="14" pageNumber="156" reason="3">
|
||
<emphasis id="B971EAE3C320FFC19DA2F90E92E2B8ED" bold="true" box="[189,505,1685,1711]" pageId="14" pageNumber="156">Other material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C320FFC19DA2F95F90C6B943" blockId="14.[189,1398,1731,1793]" pageId="14" pageNumber="156">
|
||
<materialsCitation id="3B6D3CACC320FFC19DA2F95F90C1B943" ID-GBIF-Occurrence="3357769311" collectionCode="NHMSU" country="Thailand" county="North East" elevation="336" latitude="16.830496" location="Pu Lup Cave" longLatPrecision="1" longitude="101.9861" municipality="Chum Phae District" pageId="14" pageNumber="156" specimenCode="NHMSU-00031" specimenCount="12" stateProvince="Khon Kaen Province">
|
||
<collectingCountry id="F3127661C320FFC19DA2F95F924EB89C" box="[189,341,1732,1758]" name="Thailand" pageId="14" pageNumber="156">THAILAND</collectingCountry>
|
||
•
|
||
<specimenCount id="9D03FD78C320FFC19C6FF95F92D6B89F" box="[368,461,1731,1758]" pageId="14" pageNumber="156">4 shells</specimenCount>
|
||
,
|
||
<specimenCount id="9D03FD78C320FFC19CC5F95F919AB89C" box="[474,641,1731,1758]" pageId="14" pageNumber="156">8 living specs</specimenCount>
|
||
;
|
||
<collectingCounty id="62DB4E7DC320FFC19F91F958900BB89C" box="[654,784,1731,1758]" pageId="14" pageNumber="156">North East</collectingCounty>
|
||
,
|
||
<collectingRegion id="49C1F813C320FFC19E01F9589702B89F" box="[798,1049,1731,1758]" country="Thailand" name="Khon Kaen" pageId="14" pageNumber="156">Khon Kaen Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BDEAC8BC320FFC19939F9589608B89F" box="[1062,1299,1731,1757]" pageId="14" pageNumber="156">Chum Phae District</collectingMunicipality>
|
||
,
|
||
<location id="8EDA602AC320FFC1983FF95F93E6B943" LSID="urn:lsid:plazi:treatment:03AC87E7C320FFDF9F41FD0890C2BA73:8EDA602AC320FFC1983FF95F93E6B943" country="Thailand" county="North East" latitude="16.830496" longLatPrecision="1" longitude="101.9861" municipality="Chum Phae District" name="Pu Lup Cave" pageId="14" pageNumber="156" stateProvince="Khon Kaen Province">Pu Lup Cave</location>
|
||
;
|
||
<geoCoordinate id="EE315036C320FFC19C16F97E92D8B943" box="[265,451,1765,1793]" degrees="16" direction="north" minutes="49" orientation="latitude" pageId="14" pageNumber="156" precision="1" seconds="49.78" value="16.830496">16°49′49.78″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE315036C320FFC19CD0F97E919DB943" box="[463,646,1765,1793]" degrees="101" direction="east" minutes="59" orientation="longitude" pageId="14" pageNumber="156" precision="1" seconds="9.97" value="101.9861">101°59′9.97″ E</geoCoordinate>
|
||
; alt.
|
||
<quantity id="4CFD9B14C320FFC19FA0F97C9017B943" box="[703,780,1767,1793]" metricMagnitude="2" metricUnit="m" metricValue="3.36" pageId="14" pageNumber="156" unit="m" value="336.0">
|
||
<elevation id="0028D1C2C320FFC19FA0F97C9017B943" box="[703,780,1767,1793]" metricMagnitude="2" metricUnit="m" metricValue="3.36" pageId="14" pageNumber="156" unit="m" value="336.0">336 m</elevation>
|
||
</quantity>
|
||
;
|
||
<specimenCode id="DBA39E8AC320FFC19E07F97C90C1B943" box="[792,986,1767,1793]" pageId="14" pageNumber="156">NHMSU-00031</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC320FFDF9DA2F8B391BCBC87" lastPageId="16" lastPageNumber="158" pageId="14" pageNumber="156" type="description">
|
||
<paragraph id="8BBA36F1C320FFC19DA2F8B39255B900" blockId="14.[189,334,1832,1858]" box="[189,334,1832,1858]" pageId="14" pageNumber="156">
|
||
<heading id="D0F2819DC320FFC19DA2F8B39255B900" bold="true" box="[189,334,1832,1858]" fontSize="11" level="3" pageId="14" pageNumber="156" reason="3">
|
||
<emphasis id="B971EAE3C320FFC19DA2F8B39255B900" bold="true" box="[189,334,1832,1858]" pageId="14" pageNumber="156">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C320FFC19DA2F8CC9774B998" blockId="14.[189,1400,1877,2010]" pageId="14" pageNumber="156">
|
||
SHELL (
|
||
<figureCitation id="133E2A74C320FFC19C0BF8CC926BB933" box="[276,368,1879,1905]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="14" pageNumber="156">Fig. 3D</figureCitation>
|
||
,
|
||
<tableCitation id="C687034AC320FFC19C63F8CC92C8B933" box="[380,467,1879,1905]" captionStart="Table 1" captionStartId="2.[185,255,857,885]" captionTargetBox="[195,1921,987,1444]" captionTargetPageId="2" captionText="Table 1. Shell measurements (mm) of six new species of the genus Landouria Godwin-Austen, 1918 from northeastern Thailand." httpUri="http://table.plazi.org/id/DF7A6679C32CFFCD9DA6FCC295B0BD37" pageId="14" pageNumber="156" tableUuid="DF7A6679C32CFFCD9DA6FCC295B0BD37">Table 1</tableCitation>
|
||
). Dextral, somewhat depressed, relatively large-sized. Whorls 6½, suture deep, apex obtuse, and moderately depressed, with low conical spire. Body whorl rounded, slightly keeled, red-brown in living specimens (
|
||
<figureCitation id="133E2A74C320FFC19F20F8069184B9F5" box="[575,671,1949,1975]" captionStart="Fig" captionStartId="12.[189,232,1921,1947]" captionTargetBox="[193,1395,278,1870]" captionTargetId="figure-12@12.[193,1395,265,1883]" captionTargetPageId="12" captionText="Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514847" httpUri="https://zenodo.org/record/5514847/files/figure.png" pageId="14" pageNumber="156">Fig. 4D</figureCitation>
|
||
), empty shells are paler (
|
||
<figureCitation id="133E2A74C320FFC19ECDF8069728B9F5" box="[978,1075,1949,1975]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="14" pageNumber="156">Fig. 3D</figureCitation>
|
||
). Umbilicus very deep and wide. Aperture almost circular, with slightly thickened and weakly expanded lip.
|
||
</paragraph>
|
||
<caption id="DF7A6679C321FFC09DA2F883927CB9A1" ID-DOI="http://doi.org/10.5281/zenodo.5514849" ID-Zenodo-Dep="5514849" httpUri="https://zenodo.org/record/5514849/files/figure.png" pageId="15" pageNumber="157" startId="15.[189,232,1816,1842]" targetBox="[283,1352,278,1767]" targetPageId="15">
|
||
<paragraph id="8BBA36F1C321FFC09DA2F883927CB9A1" blockId="15.[189,1399,1816,2019]" pageId="15" pageNumber="157">
|
||
<emphasis id="B971EAE3C321FFC09DA2F8839217B970" bold="true" box="[189,268,1816,1842]" pageId="15" pageNumber="157">Fig. 5.</emphasis>
|
||
Illustrations of the genital system of
|
||
<taxonomicName id="4C054D72C321FFC09FD6F8839067B971" box="[713,892,1816,1843]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="157" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B971EAE3C321FFC09FD6F883905EB970" box="[713,837,1816,1842]" italics="true" pageId="15" pageNumber="157">Landouria</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
from northeastern Thailand.
|
||
<emphasis id="B971EAE3C321FFC099C8F88397F0B970" bold="true" box="[1239,1259,1816,1842]" pageId="15" pageNumber="157">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C321FFC099E4F883923DB917" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="157" phylum="Mollusca" rank="species" species="circinata" status="sp. nov.">
|
||
<emphasis id="B971EAE3C321FFC099E4F883923DB917" italics="true" pageId="15" pageNumber="157">Landouria circinata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C321FFC09C30F8A79294B914" box="[303,399,1852,1878]" pageId="15" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype (NHMSU-00024).
|
||
<emphasis id="B971EAE3C321FFC09FF0F8A0901BB917" bold="true" box="[751,768,1851,1877]" pageId="15" pageNumber="157">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C321FFC09E0EF8A790A2B917" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[785,953,1851,1877]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="157" phylum="Mollusca" rank="species" species="tuberculata" status="sp. nov.">
|
||
<emphasis id="B971EAE3C321FFC09E0EF8A790A2B917" box="[785,953,1851,1877]" italics="true" pageId="15" pageNumber="157">L. tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C321FFC09EDCF8A79739B914" box="[963,1058,1852,1878]" pageId="15" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype (NHMSU-00026).
|
||
<emphasis id="B971EAE3C321FFC09DA0F8C593C9B93A" bold="true" box="[191,210,1886,1912]" pageId="15" pageNumber="157">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C321FFC09DF9F8C492FAB93A" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[230,481,1886,1912]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="157" phylum="Mollusca" rank="species" species="trochomorphoides" status="sp. nov.">
|
||
<emphasis id="B971EAE3C321FFC09DF9F8C492FAB93A" box="[230,481,1886,1912]" italics="true" pageId="15" pageNumber="157">L. trochomorphoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C321FFC09CF2F8C4914BB93B" box="[493,592,1887,1913]" pageId="15" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype (NHMSU-00028).
|
||
<emphasis id="B971EAE3C321FFC09EA8F8C490D7B93B" bold="true" box="[951,972,1887,1913]" pageId="15" pageNumber="157">D</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C321FFC09EFEF8C4978AB93A" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[993,1169,1886,1912]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="157" phylum="Mollusca" rank="species" species="chloritoides" status="sp. nov.">
|
||
<emphasis id="B971EAE3C321FFC09EFEF8C4978AB93A" box="[993,1169,1886,1912]" italics="true" pageId="15" pageNumber="157">L. chloritoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C321FFC09981F8C4961BB93B" box="[1182,1280,1887,1913]" pageId="15" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype (NHMSU-00030).
|
||
<emphasis id="B971EAE3C321FFC09C83F81992ABB9DE" bold="true" box="[412,432,1922,1948]" pageId="15" pageNumber="157">E</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C321FFC09CA1F819912CB9DE" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[446,567,1922,1948]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="157" phylum="Mollusca" rank="species" species="elegans" status="sp. nov.">
|
||
<emphasis id="B971EAE3C321FFC09CA1F819912CB9DE" box="[446,567,1922,1948]" italics="true" pageId="15" pageNumber="157">L. elegans</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C321FFC09F21F8199180B9DE" box="[574,667,1922,1948]" pageId="15" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype (NHMSU-00033).
|
||
<emphasis id="B971EAE3C321FFC09EEDF819971FB9DE" bold="true" box="[1010,1028,1922,1948]" pageId="15" pageNumber="157">F</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C321FFC0990DF819921AB9FD" authority="(Mollendorff, 1902)" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" baseAuthorityName="Mollendorff" baseAuthorityYear="1902" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="157" phylum="Mollusca" rank="species" species="diplogramma" status="comb. nov.">
|
||
<emphasis id="B971EAE3C321FFC0990DF81997D4B9DE" box="[1042,1231,1922,1948]" italics="true" pageId="15" pageNumber="157">L. diplogramma</emphasis>
|
||
(
|
||
<bibRefCitation id="EF944B00C321FFC099C1F81993E2B9FD" author="Mollendorff O. F." pageId="15" pageNumber="157" pagination="153 - 160" refId="ref10489" refString="Mollendorff O. F. 1902. Neue Arten und Unterarten von Fruhstorfer in Siam gesammelt. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 34 (9 - 10): 153 - 160." type="journal article" year="1902">Möllendorff, 1902</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C321FFC09C14F83E9294B9FD" box="[267,399,1957,1983]" pageId="15" pageNumber="157" rank="species">comb. nov.</taxonomicNameLabel>
|
||
(NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBA36F1C33EFFDF9DA2FE92967DBFB4" blockId="16.[188,1399,265,503]" pageId="16" pageNumber="158">
|
||
GENITAL SYSTEM (n = 3) (
|
||
<figureCitation id="133E2A74C33EFFDF9CFAFE929158BF66" box="[485,579,265,292]" captionStart="Fig" captionStartId="15.[189,232,1816,1842]" captionTargetBox="[283,1352,278,1767]" captionTargetId="figure-15@15.[232,1356,265,1778]" captionTargetPageId="15" captionText="Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514849" httpUri="https://zenodo.org/record/5514849/files/figure.png" pageId="16" pageNumber="158">Fig. 5D</figureCitation>
|
||
,
|
||
<tableCitation id="C687034AC33EFFDF9F4FFE9291B0BF61" box="[592,683,265,291]" captionStart="Table 4" captionStartId="10.[185,255,544,572]" captionTargetBox="[196,1908,758,1739]" captionTargetPageId="10" captionText="Table 4. Comparison of shells, radulae, and genital systems of Landouria spp. from northeastern Thailand. References: 1 = Tumpeesuwan & Tumpeesuwan (2019); 2 = Nahok (2020); 3 = this study. *In Tumpeesuwan & Tumpeesuwan (2019) indicated the number of teeth per row in Landouria strobiloides varies from 3 to 53. This lowest row has only 3 teeth. It is the uppermost row, probably lacerated and peel off. Unfortunately, we could not re-checked the complete specimens in this time being, so, this point need further investigation and proof." httpUri="http://table.plazi.org/id/DF7A6679C324FFC59DA6FDBB969CBCE7" pageId="16" pageNumber="158" tableUuid="DF7A6679C324FFC59DA6FDBB969CBCE7">Table 4</tableCitation>
|
||
). Penis very large and stout, longer than vagina, cylindrical, gradually tapering towards epiphallus, internally with six rather thick, corrugated longitudinal pilasters; opening of the short, grooved verge surrounded by 2–3 lobes. Epiphallus short and thick. Flagellum short, elongate ovate, without nodes, internally with three large longitudinal pilasters. Vas deferens long and slender, entering in the lower half of the flagellum. Vagina short, as long as free oviduct, internally with seven, corrugated longitudinal pilasteres varying in size. Gametolytic sac thickened at base, with a long narrow cylindrical tube, and a small oval sac at distal end. Uterus swollen, prostate gland long.
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C33EFFDF9DA2FD8591BCBC87" blockId="16.[188,1399,541,709]" pageId="16" pageNumber="158">
|
||
RADULA (n = 3). Comprises 125–129 transverse rows with 75–83 teeth per row; radular formula: (26–28)- (11–13)-1-(11–13)-(26–28). Central tooth short lanceolate, symmetric, unicuspid (
|
||
<figureCitation id="133E2A74C33EFFDF9963FDDA97D4BC1E" box="[1148,1231,577,604]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="16" pageNumber="158">Fig. 6J</figureCitation>
|
||
). Lateral teeth large and elongated lanceolate (
|
||
<figureCitation id="133E2A74C33EFFDF9F2CFDFF919CBC3D" box="[563,647,612,639]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="16" pageNumber="158">Fig. 6J</figureCitation>
|
||
). Marginal teeth lanceolate, gradually changing from unicuspid to bicuspid and finally tricuspid; endocone small; ectocone slightly oblique, with wide triangular notch and two to four lateral cusps (
|
||
<figureCitation id="133E2A74C33EFFDF9F05FD319181BC87" box="[538,666,682,709]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="16" pageNumber="158">Fig. 6K–L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC33EFFDF9DA2FD7790CCBDDD" pageId="16" pageNumber="158" type="discussion">
|
||
<paragraph id="8BBA36F1C33EFFDF9DA2FD779234BD44" blockId="16.[189,303,748,774]" box="[189,303,748,774]" pageId="16" pageNumber="158">
|
||
<heading id="D0F2819DC33EFFDF9DA2FD779234BD44" bold="true" box="[189,303,748,774]" fontSize="11" level="3" pageId="16" pageNumber="158" reason="3">
|
||
<emphasis id="B971EAE3C33EFFDF9DA2FD779234BD44" bold="true" box="[189,303,748,774]" pageId="16" pageNumber="158">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C33EFFDF9DA2FC8190CCBDDD" blockId="16.[188,1399,794,927]" pageId="16" pageNumber="158">
|
||
<taxonomicName id="4C054D72C33EFFDF9DA2FC8192D7BD76" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[189,460,794,820]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="158" phylum="Mollusca" rank="species" species="chloritoides" status="sp. nov.">
|
||
<emphasis id="B971EAE3C33EFFDF9DA2FC8192D7BD76" box="[189,460,794,820]" italics="true" pageId="16" pageNumber="158">Landouria chloritoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C33EFFDF9CCCFC809136BD77" box="[467,557,795,821]" pageId="16" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be distinguished from other species of
|
||
<taxonomicName id="4C054D72C33EFFDF9938FC8197B8BD76" authorityName="Godwin-Austen" authorityYear="1918" box="[1063,1187,794,820]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="158" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B971EAE3C33EFFDF9938FC8197B8BD76" box="[1063,1187,794,820]" italics="true" pageId="16" pageNumber="158">Landouria</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F3127661C33EFFDF99D8FC819634BD76" box="[1223,1327,794,820]" name="Thailand" pageId="16" pageNumber="158">Thailand</collectingCountry>
|
||
by its conchological similarity to the genus
|
||
<taxonomicName id="4C054D72C33EFFDF9F6CFCA691C0BD15" authorityName="Beck" authorityYear="1837" box="[627,731,829,855]" class="Gastropoda" family="Camaenidae" genus="Chloritis" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="158" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B971EAE3C33EFFDF9F6CFCA691C0BD15" box="[627,731,829,855]" italics="true" pageId="16" pageNumber="158">Chloritis</emphasis>
|
||
</taxonomicName>
|
||
and its much larger and stout penis (
|
||
<figureCitation id="133E2A74C33EFFDF9998FCA697FEBD1A" box="[1159,1253,829,856]" captionStart="Fig" captionStartId="15.[189,232,1816,1842]" captionTargetBox="[283,1352,278,1767]" captionTargetId="figure-15@15.[232,1356,265,1778]" captionTargetPageId="15" captionText="Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514849" httpUri="https://zenodo.org/record/5514849/files/figure.png" pageId="16" pageNumber="158">Fig. 5D</figureCitation>
|
||
). The radula of
|
||
<taxonomicName id="4C054D72C33EFFDF9DC2FCF99292BD39" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[221,393,865,891]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="158" phylum="Mollusca" rank="species" species="chloritoides" status="sp. nov.">
|
||
<emphasis id="B971EAE3C33EFFDF9DC2FCF99292BD39" box="[221,393,865,891]" italics="true" pageId="16" pageNumber="158">L. chloritoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C33EFFDF9C8EFCFA92F7BD39" box="[401,492,865,891]" pageId="16" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is similar to that of
|
||
<taxonomicName id="4C054D72C33EFFDF9FC5FCF99064BD39" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[730,895,865,891]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="158" phylum="Mollusca" rank="species" species="tuberculata" status="sp. nov.">
|
||
<emphasis id="B971EAE3C33EFFDF9FC5FCF99064BD39" box="[730,895,865,891]" italics="true" pageId="16" pageNumber="158">L. tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C33EFFDF9E98FCFA90F9BD39" box="[903,994,865,891]" pageId="16" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
but differs by its unicuspid lateral teeth (
|
||
<figureCitation id="133E2A74C33EFFDF9C19FC1F9242BDDD" box="[262,345,900,927]" captionStart="Fig" captionStartId="15.[189,232,1816,1842]" captionTargetBox="[283,1352,278,1767]" captionTargetId="figure-15@15.[232,1356,265,1778]" captionTargetPageId="15" captionText="Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514849" httpUri="https://zenodo.org/record/5514849/files/figure.png" pageId="16" pageNumber="158">Fig. 5J</figureCitation>
|
||
) vs the bicuspid lateral teeth of
|
||
<taxonomicName id="4C054D72C33EFFDF9FCFFC1E906EBDDC" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[720,885,900,926]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="158" phylum="Mollusca" rank="species" species="tuberculata" status="sp. nov.">
|
||
<emphasis id="B971EAE3C33EFFDF9FCFFC1E906EBDDC" box="[720,885,900,926]" italics="true" pageId="16" pageNumber="158">L. tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C33EFFDF9E62FC1E90CCBDDD" box="[893,983,901,927]" pageId="16" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC33EFFDF9DA2FC5E90C2BA73" pageId="16" pageNumber="158" type="distribution">
|
||
<paragraph id="8BBA36F1C33EFFDF9DA2FC5E924DBD9D" blockId="16.[189,342,965,991]" box="[189,342,965,991]" pageId="16" pageNumber="158">
|
||
<heading id="D0F2819DC33EFFDF9DA2FC5E924DBD9D" bold="true" box="[189,342,965,991]" fontSize="11" level="3" pageId="16" pageNumber="158" reason="3">
|
||
<emphasis id="B971EAE3C33EFFDF9DA2FC5E924DBD9D" bold="true" box="[189,342,965,991]" pageId="16" pageNumber="158">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C33EFFDF9DA2FC6F90C2BA73" blockId="16.[189,1399,1012,1074]" pageId="16" pageNumber="158">
|
||
<taxonomicName id="4C054D72C33EFFDF9DA2FC6F92D6BA4C" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[189,461,1012,1038]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="158" phylum="Mollusca" rank="species" species="chloritoides" status="sp. nov.">
|
||
<emphasis id="B971EAE3C33EFFDF9DA2FC6F92D6BA4C" box="[189,461,1012,1038]" italics="true" pageId="16" pageNumber="158">Landouria chloritoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C33EFFDF9CC9FC6F912ABA4C" box="[470,561,1012,1038]" pageId="16" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is currently known only from the limestone area around Phu Pha Man National Park, Chum Phae District,
|
||
<collectingRegion id="49C1F813C33EFFDF9F7AFB8C9045BA73" box="[613,862,1047,1074]" country="Thailand" name="Khon Kaen" pageId="16" pageNumber="158">Khon Kaen Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3127661C33EFFDF9E76FB8C90CEBA73" box="[873,981,1047,1073]" name="Thailand" pageId="16" pageNumber="158">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |