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<document id="5594873D422AA584F28972B0F0166922" ID-CLB-Dataset="6176" ID-DOI="10.11646/phytotaxa.30.1.1" ID-GBIF-Dataset="e617ba0d-6329-4481-8c82-15d70cee421c" ID-ISSN="1179-3163" ID-Zenodo-Dep="4894396" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622649201366" checkinUser="felipe" docAuthor="Levkov, Zlatko &amp; Williams, David M." docDate="2011" docId="03AC87D2B840FFEB00FDF9E6249FDD8A" docLanguage="en" docName="Phytotaxa.30.1-41.pdf.imf" docOrigin="Phytotaxa 30" docSource="http://biotaxa.org/Phytotaxa/article/view/phytotaxa.30.1.1" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Gomphonema olivaceum Brebisson 1838" docType="key" docVersion="7" lastPageNumber="33" masterDocId="FF95FFAAB85FFFCB006AFF802103DC05" masterDocTitle="Fifteen new diatom (Bacillariophyta) species from Lake Ohrid, Macedonia" masterLastPageNumber="41" masterPageNumber="1" pageNumber="32" updateTime="1698998376749" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3583CE8FE09655EBCAB8AC9AE94084E6">Fifteen new diatom (Bacillariophyta) species from Lake Ohrid, Macedonia</mods:title>
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<mods:namePart id="0F4FE142322C5E3CA1E5BEE0475556E4">Levkov, Zlatko</mods:namePart>
<mods:affiliation id="95D57430EECCFC6D9EF6A9AA7C5A80C6">Institute of Biology, Faculty of Natural Sciences, Gazi Baba bb, Skopje, R. Macedonia. E-mail: zlevkov @ iunona. pmf. ukim. edu. mk Department of Botany, the Natural History Museum, Cromwell Road, London, United Kingdom. Email: d. m. williams @ nhm. ac. uk</mods:affiliation>
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<mods:namePart id="CE7100EEDAA4BC37793FAD0999D69C1A">Williams, David M.</mods:namePart>
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<mods:title id="02F051718EE29C92238A677332A7C42A">Phytotaxa</mods:title>
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<mods:date id="7103AEC017BB4D1F164F641ACBF6F644">2011</mods:date>
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<mods:number id="6F86F9E6140169BF862804039EA7A2A6">30</mods:number>
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<mods:identifier id="0B9837290A3242E72E8127A09F97B098" type="DOI">10.11646/phytotaxa.30.1.1</mods:identifier>
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<treatment id="03AC87D2B840FFEB00FDF9E6249FDD8A" ID-DOI="http://doi.org/10.5281/zenodo.4894553" ID-GBIF-Taxon="182645413" ID-Zenodo-Dep="4894553" LSID="urn:lsid:plazi:treatment:03AC87D2B840FFEB00FDF9E6249FDD8A" httpUri="http://treatment.plazi.org/id/03AC87D2B840FFEB00FDF9E6249FDD8A" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<subSubSection id="C31F654FB840FFD400FDF9E62081DAAD" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8BBA36C4B840FFD400FDF9E62081DAAD" blockId="31.[151,1387,1638,1704]" pageId="31" pageNumber="32">
<heading id="D0F281A8B840FFD400FDF9E62081DAAD" bold="true" fontSize="12" level="1" pageId="31" pageNumber="32" reason="1">
<emphasis id="B971EAD6B840FFD400FDF9E62081DAAD" bold="true" pageId="31" pageNumber="32">
Key for identification of members of the
<taxonomicName id="4C054D47B840FFD402A8F9E62508DA87" authorityName="Brebisson" authorityYear="1838" baseAuthorityName="Hornemann" box="[706,1035,1638,1666]" class="Bacillariophyceae" family="Gomphonemataceae" genus="Gomphonema" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="31" pageNumber="32" phylum="Bacillariophyta" rank="species" species="olivaceum">
<emphasis id="B971EAD6B840FFD402A8F9E62508DA87" bold="true" box="[706,1035,1638,1666]" italics="true" pageId="31" pageNumber="32">Gomphonema olivaceum</emphasis>
</taxonomicName>
complex and related taxa from Lake
<collectingRegion id="49C1F826B840FFD40145F90B2081DAAD" box="[303,386,1675,1704]" country="Macedonia" name="Ohrid" pageId="31" pageNumber="32">Ohrid</collectingRegion>
</emphasis>
</heading>
</paragraph>
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<keyStep id="30F12E61B840FFD400FDF9582498DB0A" pageId="31" pageNumber="32">
<paragraph id="8BBA36C4B840FFD400FDF9582498DAEA" blockId="31.[151,1436,1752,2031]" box="[151,1435,1752,1775]" pageId="31" pageNumber="32">
<keyLead id="30F495F1B840FFD400FDF9582498DAEA" box="[151,1435,1752,1775]" pageId="31" pageNumber="32">
1. Valves broader than
<quantity id="4CFD9B21B840FFD401C8F95820E9DAEA" box="[418,490,1752,1775]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="31" pageNumber="32" unit="mm" value="12.0">12 µm</quantity>
, marginal lamina present (visible in girdle view) ........................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BBA36C4B840FFD400FDF9782498DB0A" blockId="31.[151,1436,1752,2031]" box="[151,1435,1784,1807]" pageId="31" pageNumber="32">
<keyLead id="30F495F1B840FFD400FDF9782498DB0A" box="[151,1435,1784,1807]" pageId="31" pageNumber="32">
-. Valves narrower than
<quantity id="4CFD9B21B840FFD401DBF97820FADB0A" box="[433,505,1784,1807]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="31" pageNumber="32" unit="mm" value="12.0">12 µm</quantity>
, marginal lamina absent ................................................................................................ 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F12E61B840FFD400FDF8982498DB6A" pageId="31" pageNumber="32">
<paragraph id="8BBA36C4B840FFD400FDF8982498DB2A" blockId="31.[151,1436,1752,2031]" box="[151,1435,1816,1839]" pageId="31" pageNumber="32">
<keyLead id="30F495F1B840FFD400FDF8982498DB2A" box="[151,1435,1816,1839]" pageId="31" pageNumber="32">
2. Central area panduriform, striae
<quantity id="4CFD9B21B840FFD40249F8982378DB2A" box="[547,635,1816,1839]" metricMagnitude="-1" metricUnit="m" metricValue="2.0955" metricValueMax="2.286" metricValueMin="1.905" pageId="31" pageNumber="32" unit="in" value="8.25" valueMax="9.0" valueMin="7.5">7.5–9 in</quantity>
<quantity id="4CFD9B21B840FFD402EBF89823C4DB2A" box="[641,711,1816,1839]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="32" unit="mm" value="10.0">10 µm</quantity>
<emphasis id="B971EAD6B840FFD402A5F898224EDB2A" box="[719,845,1816,1839]" italics="true" pageId="31" pageNumber="32">...................</emphasis>
<taxonomicName id="4C054D47B840FFD4033DF898259CDB2A" authority="Levkov" authorityName="Levkov" box="[855,1183,1816,1839]" class="Bacillariophyceae" family="Gomphonemataceae" genus="Gomphoneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="31" pageNumber="32" phylum="Bacillariophyta" rank="species" species="ohridana">
<emphasis id="B971EAD6B840FFD4033DF8982544DB2A" box="[855,1095,1816,1839]" italics="true" pageId="31" pageNumber="32">Gomphoneis ohridana</emphasis>
Levkov
</taxonomicName>
(in
<bibRefCitation id="EF944B35B840FFD404A3F8982496DB2A" author="Levkov, Z. &amp; Krstic, S. &amp; Metzeltin, D. &amp; Nakov, T." box="[1225,1429,1816,1839]" pageId="31" pageNumber="32" refId="ref20659" refString="Levkov, Z., Krstic, S., Metzeltin, D. &amp; Nakov, T. (2007) Diatoms of Lakes Prespa and Ohrid (Macedonia). Iconographia Diatomologica 16. Rugell, A. R. G. Gantner Verlag, 603 pp." type="book" year="2007">
Levkov
<emphasis id="B971EAD6B840FFD40548F8982454DB2A" box="[1314,1367,1816,1839]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
2007
</bibRefCitation>
)
</keyLead>
</paragraph>
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<keyLead id="30F495F1B840FFD400FDF8B82498DB6A" pageId="31" pageNumber="32">
-. Central area round, striae
<quantity id="4CFD9B21B840FFD401B6F8B8232EDB4A" box="[476,557,1848,1871]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="3.048" metricValueMin="2.286" pageId="31" pageNumber="32" unit="in" value="10.5" valueMax="12.0" valueMin="9.0">9–12 in</quantity>
<quantity id="4CFD9B21B840FFD40259F8B8237ADB4A" box="[563,633,1848,1871]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="32" unit="mm" value="10.0">10 µm</quantity>
...................................................................................................................... ...........................................................
<taxonomicName id="4C054D47B840FFD4020EF8D8259CDB6A" authority="Levkov, Metzeltin &amp; Krstic" authorityName="Levkov, Metzeltin &amp; Krstic" box="[612,1183,1880,1903]" class="Bacillariophyceae" family="Gomphonemataceae" genus="Gomphoneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="31" pageNumber="32" phylum="Bacillariophyta" rank="species" species="prespanenis">
<emphasis id="B971EAD6B840FFD4020EF8D82271DB6A" box="[612,882,1880,1903]" italics="true" pageId="31" pageNumber="32">Gomphoneis prespanenis</emphasis>
Levkov, Metzeltin &amp; Krstic
</taxonomicName>
(in
<bibRefCitation id="EF944B35B840FFD404A0F8D82496DB6A" author="Levkov, Z. &amp; Krstic, S. &amp; Metzeltin, D. &amp; Nakov, T." box="[1226,1429,1880,1903]" pageId="31" pageNumber="32" refId="ref20659" refString="Levkov, Z., Krstic, S., Metzeltin, D. &amp; Nakov, T. (2007) Diatoms of Lakes Prespa and Ohrid (Macedonia). Iconographia Diatomologica 16. Rugell, A. R. G. Gantner Verlag, 603 pp." type="book" year="2007">
Levkov
<emphasis id="B971EAD6B840FFD40548F8D82454DB6A" box="[1314,1367,1880,1903]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
2007
</bibRefCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F12E61B840FFD400FDF8F82498DBAA" pageId="31" pageNumber="32">
<paragraph id="8BBA36C4B840FFD400FDF8F82498DB8A" blockId="31.[151,1436,1752,2031]" box="[151,1435,1912,1935]" pageId="31" pageNumber="32">
<keyLead id="30F495F1B840FFD400FDF8F82498DB8A" box="[151,1435,1912,1935]" pageId="31" pageNumber="32">
3. Four stigmoids present in central area ......................
<taxonomicName id="4C054D47B840FFD40369F8F8259CDB8A" authority="Levkov" authorityName="Levkov" authorityYear="2007" box="[771,1183,1912,1935]" class="Bacillariophyceae" family="Gomphonemataceae" genus="Gomphonema" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="31" pageNumber="32" phylum="Bacillariophyta" rank="species" species="perolivaceoides">
<emphasis id="B971EAD6B840FFD40369F8F82544DB8A" box="[771,1095,1912,1935]" italics="true" pageId="31" pageNumber="32">Gomphonema perolivaceoides</emphasis>
Levkov
</taxonomicName>
(in
<bibRefCitation id="EF944B35B840FFD404A0F8F82496DB8A" author="Levkov, Z. &amp; Krstic, S. &amp; Metzeltin, D. &amp; Nakov, T." box="[1226,1429,1912,1935]" pageId="31" pageNumber="32" refId="ref20659" refString="Levkov, Z., Krstic, S., Metzeltin, D. &amp; Nakov, T. (2007) Diatoms of Lakes Prespa and Ohrid (Macedonia). Iconographia Diatomologica 16. Rugell, A. R. G. Gantner Verlag, 603 pp." type="book" year="2007">
Levkov
<emphasis id="B971EAD6B840FFD40548F8F82454DB8A" box="[1314,1367,1912,1935]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
2007
</bibRefCitation>
)
</keyLead>
</paragraph>
<paragraph id="8BBA36C4B840FFD400FDF8182498DBAA" blockId="31.[151,1436,1752,2031]" box="[151,1435,1944,1967]" pageId="31" pageNumber="32">
<keyLead id="30F495F1B840FFD400FDF8182498DBAA" box="[151,1435,1944,1967]" pageId="31" pageNumber="32">-. Stigmoids absent in central area ................................................................................................................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F12E61B840FFD400FDF8382498DBEA" pageId="31" pageNumber="32">
<paragraph id="8BBA36C4B840FFD400FDF838249FDBCA" blockId="31.[151,1436,1752,2031]" box="[151,1436,1976,1999]" pageId="31" pageNumber="32">
<keyLead id="30F495F1B840FFD400FDF838249FDBCA" box="[151,1436,1976,1999]" pageId="31" pageNumber="32">
4. Valves linear, narrower than
<quantity id="4CFD9B21B840FFD40190F8382336DBCA" box="[506,565,1976,1999]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="31" pageNumber="32" unit="mm" value="7.0">7 µm</quantity>
, length/breadth ration 5–7 ................................
<taxonomicName id="4C054D47B840FFD4045BF838249FDBCA" authority="Levkov" authorityName="Levkov" authorityYear="2011" box="[1073,1436,1976,1999]" class="Bacillariophyceae" family="Gomphonemataceae" genus="Gomphonema" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="31" pageNumber="32" phylum="Bacillariophyta" rank="species" species="linearoides">
<emphasis id="B971EAD6B840FFD4045BF8382447DBCA" box="[1073,1348,1976,1999]" italics="true" pageId="31" pageNumber="32">Gomphonema linearoides</emphasis>
Levkov
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBA36C4B840FFD400FDF8582498DBEA" blockId="31.[151,1436,1752,2031]" box="[151,1435,2008,2031]" pageId="31" pageNumber="32">
<keyLead id="30F495F1B840FFD400FDF8582498DBEA" box="[151,1435,2008,2031]" pageId="31" pageNumber="32">
-. Valves always broader than
<quantity id="4CFD9B21B840FFD40199F858232FDBEA" box="[499,556,2008,2031]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="31" pageNumber="32" unit="mm" value="7.0">7 µm</quantity>
............................................................................................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F12E61B87FFFEB00FDFF182498DCEA" pageId="32" pageNumber="33">
<paragraph id="8BBA36C4B87FFFEB00FDFF182498DCCA" blockId="32.[151,1436,152,399]" pageId="32" pageNumber="33">
<keyLead id="30F495F1B87FFFEB00FDFF182498DCCA" pageId="32" pageNumber="33">
5. Valves shorter than
<quantity id="4CFD9B21B87FFFEB01F1FF1820E0DCAA" box="[411,483,152,175]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="32" pageNumber="33" unit="mm" value="25.0">25 µm</quantity>
, length/bread ratio around 2 ................................................................................................ .......................................................................
<taxonomicName id="4C054D47B87FFFEB02C5FF38259CDCCA" authority="Levkov &amp; Nakov" authorityName="Levkov &amp; Nakov" authorityYear="2007" box="[687,1183,184,207]" class="Bacillariophyceae" family="Gomphonemataceae" genus="Gomphonema" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="32" pageNumber="33" phylum="Bacillariophyta" rank="species" species="subolivaceum">
<emphasis id="B971EAD6B87FFFEB02C5FF3822DDDCCA" box="[687,990,184,207]" italics="true" pageId="32" pageNumber="33">Gomphonema subolivaceum</emphasis>
Levkov &amp; Nakov
</taxonomicName>
(in
<bibRefCitation id="EF944B35B87FFFEB04A0FF382496DCCA" author="Levkov, Z. &amp; Krstic, S. &amp; Metzeltin, D. &amp; Nakov, T." box="[1226,1429,184,207]" pageId="32" pageNumber="33" refId="ref20659" refString="Levkov, Z., Krstic, S., Metzeltin, D. &amp; Nakov, T. (2007) Diatoms of Lakes Prespa and Ohrid (Macedonia). Iconographia Diatomologica 16. Rugell, A. R. G. Gantner Verlag, 603 pp." type="book" year="2007">
Levkov
<emphasis id="B971EAD6B87FFFEB0548FF382454DCCA" box="[1314,1367,184,207]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
2007
</bibRefCitation>
)
</keyLead>
</paragraph>
<paragraph id="8BBA36C4B87FFFEB00FDFF582498DCEA" blockId="32.[151,1436,152,399]" box="[151,1435,216,239]" pageId="32" pageNumber="33">
<keyLead id="30F495F1B87FFFEB00FDFF582498DCEA" box="[151,1435,216,239]" pageId="32" pageNumber="33">
-. Valves always longer than
<quantity id="4CFD9B21B87FFFEB018CFF58232DDCEA" box="[486,558,216,239]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="32" pageNumber="33" unit="mm" value="25.0">25 µm</quantity>
, length/bread ratio higher than 4 ........................................................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F12E61B87FFFEB00FDFF782498DD4A" pageId="32" pageNumber="33">
<paragraph id="8BBA36C4B87FFFEB00FDFF782498DD2A" blockId="32.[151,1436,152,399]" pageId="32" pageNumber="33">
<keyLead id="30F495F1B87FFFEB00FDFF782498DD2A" pageId="32" pageNumber="33">
6. Valves with truncate head pole, maximal width at head pole ........................................................................................ ............................................................
<taxonomicName id="4C054D47B87FFFEB0200FE98259DDD2A" authority="(Hustedt) Levkov &amp; Krstic" authorityName="Levkov &amp; Krstic" baseAuthorityName="Hustedt" box="[618,1182,280,303]" class="Bacillariophyceae" family="Gomphonemataceae" genus="Gomphonema" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="32" pageNumber="33" phylum="Bacillariophyta" rank="species" species="fonticolum">
<emphasis id="B971EAD6B87FFFEB0200FE98227BDD2A" box="[618,888,280,303]" italics="true" pageId="32" pageNumber="33">Gomphonema fonticolum</emphasis>
(Hustedt) Levkov &amp; Krstic
</taxonomicName>
(in
<bibRefCitation id="EF944B35B87FFFEB04A0FE982496DD2A" author="Levkov, Z. &amp; Krstic, S. &amp; Metzeltin, D. &amp; Nakov, T." box="[1226,1429,280,303]" pageId="32" pageNumber="33" refId="ref20659" refString="Levkov, Z., Krstic, S., Metzeltin, D. &amp; Nakov, T. (2007) Diatoms of Lakes Prespa and Ohrid (Macedonia). Iconographia Diatomologica 16. Rugell, A. R. G. Gantner Verlag, 603 pp." type="book" year="2007">
Levkov
<emphasis id="B971EAD6B87FFFEB0548FE982454DD2A" box="[1314,1367,280,303]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
2007
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)
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<keyLead id="30F495F1B87FFFEB00FDFEB82498DD4A" box="[151,1435,312,335]" pageId="32" pageNumber="33">-. Valves with broadly rounded head pole, maximal width near mid–valve .................................................................. 7</keyLead>
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7. Valves wider than
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<taxonomicName id="4C054D47B87FFFEB03B6FED82498DD6A" authority="Levkov" authorityName="Levkov" authorityYear="2011" box="[988,1435,344,367]" class="Bacillariophyceae" family="Gomphonemataceae" genus="Gomphonema" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="32" pageNumber="33" phylum="Bacillariophyta" rank="species" species="perolivaceolacuum">
<emphasis id="B971EAD6B87FFFEB03B6FED82447DD6A" box="[988,1348,344,367]" italics="true" pageId="32" pageNumber="33">Gomphonema perolivaceolacuum</emphasis>
Levkov
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<keyLead id="30F495F1B87FFFEB00FDFEF8249FDD8A" box="[151,1436,376,399]" pageId="32" pageNumber="33">
-. Valves narrower than
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<taxonomicName id="4C054D47B87FFFEB047CFEF8249FDD8A" authority="Levkov" authorityName="Levkov" authorityYear="2011" box="[1046,1436,376,399]" class="Bacillariophyceae" family="Gomphonemataceae" genus="Gomphonema" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cymbellales" pageId="32" pageNumber="33" phylum="Bacillariophyta" rank="species" species="densistriatum">
<emphasis id="B971EAD6B87FFFEB047CFEF82440DD8A" box="[1046,1347,376,399]" italics="true" pageId="32" pageNumber="33">Gomphonema densistriatum</emphasis>
Levkov
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