171 lines
20 KiB
XML
171 lines
20 KiB
XML
<document id="241BB711BF035994CEE4650B2D98500E" ID-DOI="10.5281/zenodo.188636" ID-GBIF-Dataset="7b8ff718-7c17-4741-8f1e-614858fae142" ID-ISSN="1175-5326" ID-Zenodo-Dep="188636" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461095838806" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2009" docId="03AC87BBFF9CFFF761DEC761FA877467" docLanguage="en" docName="zt02145p035.pdf" docOrigin="Zootaxa 2145" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cephennomicrus taiwanensis" docType="treatment" docVersion="9" lastPageNumber="9" masterDocId="FF95FFC3FF9BFFFF6149C134FFD37074" masterDocTitle="Cephennomicrus Reitter (Coleoptera, Staphylinidae, Scydmaeninae) of Japan and Taiwan: taxonomic notes, ten new species and comparative morphology of nomurai and taiwanensis species groups" masterLastPageNumber="35" masterPageNumber="1" pageNumber="8" updateTime="1698589186247" updateUser="plazi">
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<mods:title id="75499A4D4F89DC2EADB4540138D9C857">Cephennomicrus Reitter (Coleoptera, Staphylinidae, Scydmaeninae) of Japan and Taiwan: taxonomic notes, ten new species and comparative morphology of nomurai and taiwanensis species groups</mods:title>
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<mods:namePart id="091CFEEA7437067A48D9BE93BB5B6B75">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="7320A3C03353295221D189BAA64612AD">Zootaxa</mods:title>
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<mods:date id="448587B815C6A9C9C0CED81FFC4B7CB4">2009</mods:date>
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<mods:number id="5436DD1E1F831587C15AE8472B4D2D40">2145</mods:number>
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<mods:identifier id="7E892B86AE5967F1A72FC03C034B8C11" type="DOI">10.5281/zenodo.188636</mods:identifier>
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<treatment id="03AC87BBFF9CFFF761DEC761FA877467" ID-DOI="http://doi.org/10.5281/zenodo.6217182" ID-GBIF-Taxon="119620359" ID-Zenodo-Dep="6217182" LSID="urn:lsid:plazi:treatment:03AC87BBFF9CFFF761DEC761FA877467" httpUri="http://treatment.plazi.org/id/03AC87BBFF9CFFF761DEC761FA877467" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<subSubSection id="C31F6526FF9CFFF761DEC761FC0770AC" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="8BBA36ADFF9CFFF861DEC761FE00761B" blockId="7.[151,467,1621,1647]" box="[151,467,1621,1647]" pageId="7" pageNumber="8">
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<heading id="D0F281C1FF9CFFF861DEC761FE00761B" bold="true" box="[151,467,1621,1647]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
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<emphasis id="B971EABFFF9CFFF861DEC761FE00761B" bold="true" box="[151,467,1621,1647]" pageId="7" pageNumber="8">
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<taxonomicName id="4C054D2EFF9CFFF861DEC761FEF7761B" ID-CoL="SFPX" box="[151,292,1621,1647]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="taiwanensis">
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<emphasis id="B971EABFFF9CFFF861DEC761FEF7761B" bold="true" box="[151,292,1621,1647]" italics="true" pageId="7" pageNumber="8">taiwanensis</emphasis>
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</taxonomicName>
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species group
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBA36ADFF9CFFF761DEC7AAFC0770AC" blockId="7.[151,1437,1694,2030]" lastBlockId="8.[151,1437,152,294]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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This group is established for species with very elongate aedeagus, which in ventral view is about
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<date id="FFBB106DFF9CFFF8644CC7AAFAF076CC" box="[1285,1315,1694,1720]" pageId="7" pageNumber="8">3x</date>
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as long as broad, with asymmetrical internal armature composed of a tubular median duct with its median part surrounded by slightly or strongly asymmetrical structures forming a central complex. These characters are unique within the genus. Additionally, all species in this group share the following combination of characters: the antennal club composed of small antennomere IX and very large antennomeres X–XI; mandibles in both sexes with a long apical tooth; the head in both sexes with frontal glands, non-modified in males; the base of pronotum with four pits forming two lateral pairs, without grooves; setae covering the dorsum very short and nearly recumbent, additionally the pronotum with three pairs of long setae located near the broadest place, in hind angles and in median part of the posterior margin, opposite anterior corners of the mesoscutellum; the elytra with similar, several (minimum 3) pairs of long setae along lateral margins; male with very small, hardly noticeable median tubercle in posterior part of the metaventrite.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31F6526FF93FFF7618FC1D1FEB371C8" pageId="8" pageNumber="9" type="description">
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<paragraph id="8BBA36ADFF93FFF7618FC1D1FB4A7151" blockId="8.[151,1437,152,294]" pageId="8" pageNumber="9">
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This lineage comprises six species known from the Ryukyus and
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<collectingCountry id="F312763DFF93FFF7628EC1D1FBF2708B" box="[967,1057,229,255]" name="Taiwan" pageId="8" pageNumber="9">Taiwan</collectingCountry>
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:
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<taxonomicName id="4C054D2EFF93FFF76565C1D1FB0A708A" box="[1068,1241,229,254]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="taiwanensis">
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<emphasis id="B971EABFFF93FFF76565C1D1FB0A708A" box="[1068,1241,229,254]" italics="true" pageId="8" pageNumber="9">C. taiwanensis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C054D2EFF93FFF765A1C1D1FA4F708A" box="[1256,1436,229,254]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="iriomotensis" status="sp. nov.">
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<emphasis id="B971EABFFF93FFF765A1C1D1FA4F708A" box="[1256,1436,229,254]" italics="true" pageId="8" pageNumber="9">C. iriomotensis</emphasis>
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</taxonomicName>
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<emphasis id="B971EABFFF93FFF761DEC03FFF0A7151" bold="true" box="[151,217,267,293]" pageId="8" pageNumber="9">
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<taxonomicNameLabel id="A24257C4FF93FFF761DEC03FFF0A7151" box="[151,217,267,293]" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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,
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<taxonomicName id="4C054D2EFF93FFF761AEC038FEAF7151" box="[231,380,268,293]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nagoanus" status="sp. nov.">
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<emphasis id="B971EABFFF93FFF761AEC038FEAF7151" box="[231,380,268,293]" italics="true" pageId="8" pageNumber="9">C. nagoanus</emphasis>
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</taxonomicName>
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<emphasis id="B971EABFFF93FFF760CAC03FFE167151" bold="true" box="[387,453,267,293]" pageId="8" pageNumber="9">
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<taxonomicNameLabel id="A24257C4FF93FFF760CAC03FFE167151" box="[387,453,267,293]" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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,
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<taxonomicName id="4C054D2EFF93FFF7609DC038FDE97151" box="[468,570,268,293]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tsurui" status="sp. nov.">
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<emphasis id="B971EABFFF93FFF7609DC038FDE97151" box="[468,570,268,293]" italics="true" pageId="8" pageNumber="9">C. tsurui</emphasis>
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</taxonomicName>
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<emphasis id="B971EABFFF93FFF76308C03FFD507151" bold="true" box="[577,643,267,293]" pageId="8" pageNumber="9">
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<taxonomicNameLabel id="A24257C4FF93FFF76308C03FFD507151" box="[577,643,267,293]" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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,
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<taxonomicName id="4C054D2EFF93FFF763DBC038FC897151" box="[658,858,268,293]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="delicatissimus" status="sp. nov.">
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<emphasis id="B971EABFFF93FFF763DBC038FC897151" box="[658,858,268,293]" italics="true" pageId="8" pageNumber="9">C. delicatissimus</emphasis>
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</taxonomicName>
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<emphasis id="B971EABFFF93FFF76228C03FFC707151" bold="true" box="[865,931,267,293]" pageId="8" pageNumber="9">
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<taxonomicNameLabel id="A24257C4FF93FFF76228C03FFC707151" box="[865,931,267,293]" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, and
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<taxonomicName id="4C054D2EFF93FFF762ADC038FB9C7151" box="[996,1103,268,293]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="imago" status="sp. nov.">
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<emphasis id="B971EABFFF93FFF762ADC038FB9C7151" box="[996,1103,268,293]" italics="true" pageId="8" pageNumber="9">C. imago</emphasis>
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</taxonomicName>
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<emphasis id="B971EABFFF93FFF7651FC03FFB4A7151" bold="true" box="[1110,1177,267,293]" pageId="8" pageNumber="9">
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<taxonomicNameLabel id="A24257C4FF93FFF7651FC03FFB4A7151" box="[1110,1177,267,293]" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="8BBA36ADFF93FFF761DEC04BFEB371C8" blockId="8.[151,749,383,444]" pageId="8" pageNumber="9">
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<emphasis id="B971EABFFF93FFF761DEC04BFDB571ED" bold="true" box="[151,614,383,409]" pageId="8" pageNumber="9">
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<taxonomicName id="4C054D2EFF93FFF761DEC04BFE2971ED" box="[151,506,383,409]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="taiwanensis">
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<emphasis id="B971EABFFF93FFF761DEC04BFE2971ED" bold="true" box="[151,506,383,409]" italics="true" pageId="8" pageNumber="9">Cephennomicrus taiwanensis</emphasis>
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</taxonomicName>
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(Jałoszy
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</emphasis>
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ń
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<emphasis id="B971EABFFF93FFF7633FC04BFD3E71ED" bold="true" box="[630,749,383,409]" pageId="8" pageNumber="9">ski, 2004)</emphasis>
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(
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<figureCitation id="133E2A28FF93FFF761D6C096FED371C8" box="[159,256,418,444]" captionStart="FIGURES 15 – 20" captionStartId="9.[151,275,1790,1814]" captionTargetBox="[327,1262,437,1753]" captionTargetId="figure@9.[316,1270,426,1766]" captionTargetPageId="9" captionText="FIGURES 15 – 20. Dorsal view of Cephennomicrus (holotypes, males). 15, C. taiwanensis (Jałoszyński). 16, C. iriomotensis Jałoszyński. 17, C. nagoanus Jałoszyński. 18, C. tsurui Jałoszyński. 19, C. delicatissimus Jałoszyński. 20, C. imago Jałoszyński. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/188640/files/figure.png" pageId="8" pageNumber="9">Figs. 15</figureCitation>
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, 21, 22)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31F6526FF93FFF761DEC0C4FD187253" pageId="8" pageNumber="9" type="reference_group">
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<paragraph id="8BBA36ADFF93FFF761DEC0C4FD187253" blockId="8.[151,753,496,551]" pageId="8" pageNumber="9">
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<taxonomicName id="4C054D2EFF93FFF761DEC0C4FD517273" authority="Jaloszynski, 2004: 158" authorityName="Jaloszynski" authorityPageNumber="158" authorityYear="2004" box="[151,642,496,519]" class="Insecta" family="Scydmaenidae" genus="Neseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="taiwanensis">
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<emphasis id="B971EABFFF93FFF761DEC0C4FE557273" box="[151,390,496,519]" italics="true" pageId="8" pageNumber="9">Neseuthia taiwanensis</emphasis>
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Jałoszyński, 2004: 158
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</taxonomicName>
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,
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<figureCitation id="133E2A28FF93FFF763C4C0C4FD3F7273" box="[653,748,496,519]" captionStart-0="FIGURES 1 – 5" captionStart-1="FIGURES 6 – 9" captionStartId-0="3.[151,274,1789,1813]" captionStartId-1="5.[151,269,791,815]" captionTargetBox-0="[346,1237,466,1738]" captionTargetBox-1="[157,1423,207,748]" captionTargetId-0="figure@3.[340,1248,453,1763]" captionTargetId-1="figure@5.[151,1436,197,768]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="FIGURES 1 – 5. Dorsal view of Cephennomicrus (male holotypes). 1, C. nomurai (Jałoszyński & Hoshina). 2, C. disjunctus (Jałoszyński, S. Arai & K. Arai). 3, C. inflatus Jałoszyński. 4, C. crucifer Jałoszyński. 5, C. hobbiti (Jałoszyński & Hoshina). Scale bars: 0.2 mm." captionText-1="FIGURES 6 – 9. Aedeagus of Cephennomicrus in ventral (6, 8) and lateral (7, 9) views. 6, 7, C. inflatus Jałoszyński. 8, 9, C. crucifer Jałoszyński. Scale bars: 0.1 mm." httpUri-0="https://zenodo.org/record/188637/files/figure.png" httpUri-1="https://zenodo.org/record/188638/files/figure.png" pageId="8" pageNumber="9">Figs 1–6</figureCitation>
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.
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<taxonomicName id="4C054D2EFF93FFF761DEC324FE037253" ID-CoL="SFPX" box="[151,464,528,551]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="taiwanensis">
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<emphasis id="B971EABFFF93FFF761DEC324FE037253" box="[151,464,528,551]" italics="true" pageId="8" pageNumber="9">Cephennomicrus taiwanensis</emphasis>
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</taxonomicName>
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; Jałoszyński, 2008: 34.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31F6526FF93FFF761DEC37BFBC5721E" box="[151,1046,591,618]" pageId="8" pageNumber="9" type="materials_examined">
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||
<paragraph id="8BBA36ADFF93FFF761DEC37BFBC5721E" blockId="8.[151,1437,591,1043]" box="[151,1046,591,618]" pageId="8" pageNumber="9">
|
||
<emphasis id="B971EABFFF93FFF761DEC37BFD84721D" bold="true" box="[151,599,591,617]" pageId="8" pageNumber="9">
|
||
<typeStatus id="54BE880FFF93FFF761DEC37BFF01721D" box="[151,210,591,617]" pageId="8" pageNumber="9">Type</typeStatus>
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||
material.
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||
<typeStatus id="54BE880FFF93FFF7601DC37BFE19721D" box="[340,458,591,617]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
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||
:
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<collectingCountry id="F312763DFF93FFF7609EC37BFD81721D" box="[471,594,591,617]" name="Taiwan" pageId="8" pageNumber="9">TAIWAN</collectingCountry>
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||
:
|
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</emphasis>
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3, Taipei County, Fu-Shan (
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<collectionCode id="ED14AE68FF93FFF762E0C364FBD9721E" LSID="urn:lsid:biocol.org:col:34991" box="[937,1034,592,618]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="8" pageNumber="9">HNHM</collectionCode>
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).
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C31F6526FF93FFF7618CC342FB4B72E4" box="[197,1176,630,656]" pageId="8" pageNumber="9" type="distribution">
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||
<paragraph id="8BBA36ADFF93FFF7618CC342FB4B72E4" blockId="8.[151,1437,591,1043]" box="[197,1176,630,656]" pageId="8" pageNumber="9">
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||
<emphasis id="B971EABFFF93FFF7618CC342FEB672E4" bold="true" box="[197,357,630,656]" pageId="8" pageNumber="9">Distribution.</emphasis>
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||
Known only from the
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<typeStatus id="54BE880FFF93FFF76338C342FD7072E4" box="[625,675,630,656]" pageId="8" pageNumber="9">type</typeStatus>
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||
locality, Taipei Hsien in Northern
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<collectingCountry id="F312763DFF93FFF76573C342FB4772E4" box="[1082,1172,630,656]" name="Taiwan" pageId="8" pageNumber="9">Taiwan</collectingCountry>
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.
|
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C31F6526FF93FFF7618FC3A9FA877467" pageId="8" pageNumber="9" type="discussion">
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||
<paragraph id="8BBA36ADFF93FFF7618FC3A9FA877467" blockId="8.[151,1437,591,1043]" pageId="8" pageNumber="9">
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||
<emphasis id="B971EABFFF93FFF7618FC3A9FE9372C3" bold="true" box="[198,320,669,695]" pageId="8" pageNumber="9">Remarks.</emphasis>
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This species is difficult to distinguish from other members of the
|
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<taxonomicName id="4C054D2EFF93FFF76574C3A9FB1B72C2" box="[1085,1224,669,694]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="taiwanensis">
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||
<emphasis id="B971EABFFF93FFF76574C3A9FB1B72C2" box="[1085,1224,669,694]" italics="true" pageId="8" pageNumber="9">taiwanensis</emphasis>
|
||
</taxonomicName>
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group by external characters; subtle differences in the general body shape (
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<figureCitation id="133E2A28FF93FFF76279C3F0FC5672AA" box="[816,901,708,734]" captionStart="FIGURES 15 – 20" captionStartId="9.[151,275,1790,1814]" captionTargetBox="[327,1262,437,1753]" captionTargetId="figure@9.[316,1270,426,1766]" captionTargetPageId="9" captionText="FIGURES 15 – 20. Dorsal view of Cephennomicrus (holotypes, males). 15, C. taiwanensis (Jałoszyński). 16, C. iriomotensis Jałoszyński. 17, C. nagoanus Jałoszyński. 18, C. tsurui Jałoszyński. 19, C. delicatissimus Jałoszyński. 20, C. imago Jałoszyński. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/188640/files/figure.png" pageId="8" pageNumber="9">Fig. 15</figureCitation>
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) exist, but they are possible to notice only by direct comparison. The combination of characters unique for
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<taxonomicName id="4C054D2EFF93FFF76223C3DEFBC47377" box="[874,1047,746,771]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="taiwanensis">
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<emphasis id="B971EABFFF93FFF76223C3DEFBC47377" box="[874,1047,746,771]" italics="true" pageId="8" pageNumber="9">C. taiwanensis</emphasis>
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</taxonomicName>
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is as follows: large punctures on the pronotum; three lateral setae on each elytron; and the aedeagus (Figs. 21, 22) with long and narrow central complex, and a complicated system of ducts. The median duct is bent twice at sharp angle, the basal loop is present, and the median loop is small and nearly closed. Moreover, this species has very slender parameres, with particularly thin apical parts, and each bears one apical and one subapical seta; the latter one is distant from the apex. The base of the parameres in the only known specimen is barely visible; reexamination with a phase-contrast microscope revealed that it is similar to that in all other species of this lineage. Despite moderate body length within the group,
|
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<taxonomicName id="4C054D2EFF93FFF76327C2CDFCC97466" box="[622,794,1017,1042]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="taiwanensis">
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||
<emphasis id="B971EABFFF93FFF76327C2CDFCC97466" box="[622,794,1017,1042]" italics="true" pageId="8" pageNumber="9">C. taiwanensis</emphasis>
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</taxonomicName>
|
||
has the longest aedeagus in this distinct lineage.
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||
</paragraph>
|
||
</subSubSection>
|
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</treatment>
|
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</document> |