treatments-xml/data/03/AC/87/03AC87B63A0DFFE5FF52BF48FC484EF4.xml

<document id="D0E188F06E15E76BF7165E77C83B9F04" ID-CLB-Dataset="3563" ID-DOI="10.11646/zootaxa.5061.3.8" ID-GBIF-Dataset="09ca7af6-f232-4272-ba7c-0f2686d11b78" ID-ISSN="1175-5326" ID-Zenodo-Dep="5649998" ID-ZooBank="25C44022-4BE9-453B-9D96-359A00A18EC1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1636147046350" checkinUser="plazi" docAuthor="Polak, Slavko &amp; Mulaomerović, Jasminko" docDate="2021" docId="03AC87B63A0DFFE5FF52BF48FC484EF4" docLanguage="en" docName="zootaxa.5061.3.8.pdf" docOrigin="Zootaxa 5061 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rudogorites simonei Polak &amp; Mulaomerović 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="554" masterDocId="FF95FFCE3A0EFFECFFC5BE26FFE74F59" masterDocTitle="Rudogorites simonei gen. nov. and sp. nov. from Central Bosnia (Coleoptera: Leiodidae: Cholevinae: Leptodirini)" masterLastPageNumber="558" masterPageNumber="545" pageNumber="548" updateTime="1699467954132" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B9A00FE4AF3A706DDDA18EA6651945EC">Rudogorites simonei gen. nov. and sp. nov. from Central Bosnia (Coleoptera: Leiodidae: Cholevinae: Leptodirini)</mods:title>
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<mods:namePart id="E5548D638B263E853CA1244C67A45DE8">Polak, Slavko</mods:namePart>
<mods:affiliation id="B81D389451DAE78E7E94295B2D7D7382">Notranjska museum Postojna, Kolodvorska cesta 3, SI- 6230 Postojna, Slovenia.</mods:affiliation>
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<mods:namePart id="6F91FC6794AAC585E1AE447346F3539F">Mulaomerović, Jasminko</mods:namePart>
<mods:nameIdentifier id="72D8AE8E1BF1BDCC8666259D6A453D1A" type="ORCID">0000-0003-4121-5238</mods:nameIdentifier>
<mods:affiliation id="A1187658EAD4A6EC627C862459E12A3E">Center for Karst and Speleology, Branilaca Sarajeva 30, BIH- 71000 Sarajevo, Bosnia and Herzegovina. jasminko @ centarzakrs. ba; https: // orcid. org / 0000 - 0003 - 4121 - 5238</mods:affiliation>
<mods:nameIdentifier id="B3BBA626587ECBB7610918BD28862E2C" type="email">jasminko@centarzakrs.ba</mods:nameIdentifier>
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<mods:date id="2E95DABDB4A3CB1F48B06A359023CF80">2021</mods:date>
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<subSubSection id="C31F652B3A0DFFEFFF52BF48FE124ED0" box="[151,501,366,393]" pageId="3" pageNumber="548" type="nomenclature">
<paragraph id="8BBA36A03A0DFFEFFF52BF48FE124ED0" blockId="3.[151,835,366,429]" box="[151,501,366,393]" pageId="3" pageNumber="548">
<heading id="D0F281CC3A0DFFEFFF52BF48FE124ED0" bold="true" box="[151,501,366,393]" fontSize="11" level="1" pageId="3" pageNumber="548" reason="1">
<emphasis id="B971EAB23A0DFFEFFF52BF48FE124ED0" bold="true" box="[151,501,366,393]" pageId="3" pageNumber="548">
<taxonomicName id="4C054D233A0DFFEFFF52BF48FE774ED0" ID-CoL="9GCRR" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[151,400,366,393]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="548" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A0DFFEFFF52BF48FE774ED0" bold="true" box="[151,400,366,393]" italics="true" pageId="3" pageNumber="548">Rudogorites simonei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24257C93A0DFFEFFE52BF49FE124ED0" box="[407,501,367,393]" pageId="3" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31F652B3A0DFFEFFF52BFB5FCA44EF4" box="[151,835,402,429]" pageId="3" pageNumber="548" type="description">
<paragraph id="8BBA36A03A0DFFEFFF52BFB5FCA44EF4" blockId="3.[151,835,366,429]" box="[151,835,402,429]" pageId="3" pageNumber="548">
<figureCitation id="133E2A253A0DFFEFFF52BFB5FF044EF4" box="[151,227,403,429]" captionStart="FIGURE 1" captionStartId="3.[151,250,1841,1865]" captionTargetBox="[235,1356,861,1807]" captionTargetId="figure-266@3.[215,1371,849,1817]" captionTargetPageId="3" captionText="FIGURE 1. Rudogorites simonei sp. nov. male and female habitus." figureDoi="http://doi.org/10.5281/zenodo.5650000" httpUri="https://zenodo.org/record/5650000/files/figure.png" pageId="3" pageNumber="548">Figs. 1</figureCitation>
, 
<figureCitation id="133E2A253A0DFFEFFF34BFB5FE964EF4" box="[241,369,403,429]" captionStart="FIGURES 2–10" captionStartId="4.[151,264,1813,1837]" captionTargetBox="[159,1421,820,1785]" captionTargetId="figure-257@4.[159,1427,813,1790]" captionTargetPageId="4" captionText="FIGURES 2–10. Figures 2–5. pronotum, ventral view. 2, Rudogorites simonei sp. nov., male. 3, Rudogorites simonei sp. nov., female. 4, Charonites scheibeli, female. 5, Charonites matzenaueri, male. Figure 6, elytra surface detail Rudogorites simonei sp. nov., male. Figures 7–8, thorax ventral view, mesothoracic apophysis are indicated with white arrow, sutures between mesosetrnum and mesepisternum are indicated with grey arrow. 7, Rudogorites simonei sp. nov., male. 8, Charonites matzenaueri, male. Figures 9–10, thorax lateral view, ventral edge of mesosternal keel is indicated with black arrow. 9, Rudogorites simonei sp. nov., male. 10, Charonites matzenaueri, male." figureDoi="http://doi.org/10.5281/zenodo.5650002" httpUri="https://zenodo.org/record/5650002/files/figure.png" pageId="3" pageNumber="548">2, 3, 6, 7, 9</figureCitation>
, 
<figureCitation id="133E2A253A0DFFEFFEB8BFB5FDFF4EF4" box="[381,536,402,429]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1106,1130]" captionTargetBox="[172,1417,188,1074]" captionTargetId="figure-19@5.[164,1424,181,1082]" captionTargetPageId="5" captionText="FIGURES 11–16. Figures 11–14, left first leg, comb of protibial spines is indicated with black arrow. 11, Rudogorites simonei sp. nov., female. 12, Rudogorites simonei sp. nov., male. 13, Charonites scheibeli, female. 14, Charonites matzenaueri, male. Figure 15, second left leg, Rudogorites simonei sp. nov., male. Figure 16, third leg, left Rudogorites simonei sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.5650004" httpUri="https://zenodo.org/record/5650004/files/figure.png" pageId="3" pageNumber="548">11, 12, 15, 16</figureCitation>
, 
<figureCitation id="133E2A253A0DFFEFFDE1BFB5FD274EF4" box="[548,704,403,429]" captionStart="FIGURES 17–24" captionStartId="6.[151,264,1257,1281]" captionTargetBox="[173,1414,274,1225]" captionTargetId="figure-80@6.[161,1426,273,1234]" captionTargetPageId="6" captionText="FIGURES 17–24. Figures 17–20. Antenna. 17, Rudogorites simonei sp. nov., male. 18, Rudogorites simonei sp. nov., female. 19, Charonites scheibeli, female. 20, Charonites matzenaueri, male. Figures 21–23, metatergal apparatus, posterior process is indicated with black arrow. 21, Rudogorites simonei sp. nov., male. 22, Charonites matzenaueri, male. 23, Charonites scheibeli, female. Figure 24, metendosternite, Rudogorites simonei sp. nov., male." figureDoi="http://doi.org/10.5281/zenodo.5650006" httpUri="https://zenodo.org/record/5650006/files/figure.png" pageId="3" pageNumber="548">17, 18, 21, 24</figureCitation>
, 
<figureCitation id="133E2A253A0DFFEFFD09BFB4FCF24EF4" box="[716,789,402,429]" captionStart="FIGURES 25–30" captionStartId="7.[151,264,1301,1325]" captionTargetBox="[159,1428,532,1276]" captionTargetId="figure-168@7.[153,1435,525,1278]" captionTargetPageId="7" captionText="FIGURES 25–30. Rudogorites simonei sp. nov., male and female genital structures. 25, aedeagus dorsal view. 26, aedeagus lateral view, paramere apex detail is separately slightly enlarged. 27, male genital segment (IX sternite). 28, female ventrite VIIIth. 29, female genital segment, urite IXth with gonostyles. 30, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5650008" httpUri="https://zenodo.org/record/5650008/files/figure.png" pageId="3" pageNumber="548">25–30</figureCitation>
, 
<figureCitation id="133E2A253A0DFFEFFCE5BFB5FCD84EF4" box="[800,831,403,429]" captionStart="FIGURE 31" captionStartId="8.[151,250,1366,1390]" captionTargetBox="[151,1435,166,1342]" captionTargetId="figure-33@8.[151,1435,166,1342]" captionTargetPageId="8" captionText="FIGURE 31. Map of taxa distributions. White star Rudogorites simonei sp. nov., black star Protobracharhron dusinae, dark grey area Charonites spp., pale grey area distribution of genera of Apholeuonus phyletic group." figureDoi="http://doi.org/10.5281/zenodo.5650012" httpUri="https://zenodo.org/record/5650012/files/figure.png" pageId="3" pageNumber="548">31</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31F652B3A0DFFEFFF52BFFDFDC44D38" pageId="3" pageNumber="548" type="materials_examined">
<paragraph id="8BBA36A03A0DFFEFFF52BFFDFDC44D38" blockId="3.[151,1437,474,825]" pageId="3" pageNumber="548">
<emphasis id="B971EAB23A0DFFEFFF52BFFDFEC94EAD" bold="true" box="[151,302,474,501]" pageId="3" pageNumber="548">Type locality</emphasis>
: 
<materialsCitation id="3B6D3CFD3A0DFFEFFEF9BFFDFDF84D38" ID-GBIF-Occurrence="3400807310" collectorName="Hercegovine, Ev. Br." country="Bosnia and Herzegovina" county="Sarajevo in Central" elevation="700" latitude="43.831" location="Lijina Pecina" longLatPrecision="68" longitude="18.075" municipality="The cave" pageId="3" pageNumber="548" specimenCount="1">
<location id="8EDA607B3A0DFFEFFEF9BFFDFE284EAC" LSID="urn:lsid:plazi:treatment:03AC87B63A0DFFE5FF52BF48FC484EF4:8EDA607B3A0DFFEFFEF9BFFDFE284EAC" box="[316,463,475,501]" country="Bosnia and Herzegovina" county="Sarajevo in Central" latitude="43.831" longLatPrecision="68" longitude="18.075" municipality="The cave" name="Lijina Pecina" pageId="3" pageNumber="548">Lijina Pećina</location>
(
<location id="8EDA607B3A0DFFEFFE19BFFDFDBC4EAC" LSID="urn:lsid:plazi:treatment:03AC87B63A0DFFE5FF52BF48FC484EF4:8EDA607B3A0DFFEFFE19BFFDFDBC4EAC" box="[476,603,475,501]" country="Bosnia and Herzegovina" county="Sarajevo in Central" latitude="43.831" longLatPrecision="68" longitude="18.075" municipality="The cave" name="Lijina cave" pageId="3" pageNumber="548">Lijina cave</location>
) (
<location id="8EDA607B3A0DFFEFFDB5BFFDFD284EAC" LSID="urn:lsid:plazi:treatment:03AC87B63A0DFFE5FF52BF48FC484EF4:8EDA607B3A0DFFEFFDB5BFFDFD284EAC" box="[624,719,475,501]" country="Bosnia and Herzegovina" county="Sarajevo in Central" latitude="43.831" longLatPrecision="68" longitude="18.075" municipality="The cave" name="Katastar" pageId="3" pageNumber="548">Katastar</location>
speleoloških objekata 
<location id="8EDA607B3A0DFFEFFC0EBFFDFBF64EAC" LSID="urn:lsid:plazi:treatment:03AC87B63A0DFFE5FF52BF48FC484EF4:8EDA607B3A0DFFEFFC0EBFFDFBF64EAC" box="[971,1041,475,501]" country="Bosnia and Herzegovina" county="Sarajevo in Central" latitude="43.831" longLatPrecision="68" longitude="18.075" municipality="The cave" name="Bosne" pageId="3" pageNumber="548">Bosne</location>
i 
<collectorName id="26F053763A0DFFEFFBE1BFFDFAEB4EAC" box="[1060,1292,475,501]" pageId="3" pageNumber="548">Hercegovine, Ev. Br.</collectorName>
5391) which is located 
<location id="8EDA607B3A0DFFEFFEC2BFD9FE994D40" LSID="urn:lsid:plazi:treatment:03AC87B63A0DFFE5FF52BF48FC484EF4:8EDA607B3A0DFFEFFEC2BFD9FE994D40" box="[263,382,511,537]" country="Bosnia and Herzegovina" county="Sarajevo in Central" latitude="43.831" longLatPrecision="68" longitude="18.075" municipality="The cave" name="North East" pageId="3" pageNumber="548">North East</location>
of the village of 
<location id="8EDA607B3A0DFFEFFDFFBFD9FD734D40" LSID="urn:lsid:plazi:treatment:03AC87B63A0DFFE5FF52BF48FC484EF4:8EDA607B3A0DFFEFFDFFBFD9FD734D40" box="[570,660,511,537]" country="Bosnia and Herzegovina" county="Sarajevo in Central" latitude="43.831" longLatPrecision="68" longitude="18.075" municipality="The cave" name="Gunjani" pageId="3" pageNumber="548">Gunjani</location>
and 
<locationDeviation id="9A082F823A0DFFEFFD0CBFD9FC734D40" box="[713,916,511,537]" location="South East" pageId="3" pageNumber="548">
<quantity id="4CFD9B453A0DFFEFFD0CBFD9FCF14D40" box="[713,790,511,537]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="3" pageNumber="548" unit="km" value="4.4">4.4 km</quantity>
South East
</locationDeviation>
of town 
<location id="8EDA607B3A0DFFEFFC33BFD9FB354D40" LSID="urn:lsid:plazi:treatment:03AC87B63A0DFFE5FF52BF48FC484EF4:8EDA607B3A0DFFEFFC33BFD9FB354D40" box="[1014,1234,511,537]" country="Bosnia and Herzegovina" county="Sarajevo in Central" latitude="43.831" longLatPrecision="68" longitude="18.075" municipality="The cave" name="Kresevo in Kresevo" pageId="3" pageNumber="548">Kreševo in Kreševo</location>
municipality. 
<collectingMunicipality id="6BDEACDA3A0DFFEFFAB4BFD9FF2D4D64" pageId="3" pageNumber="548">The cave</collectingMunicipality>
is approximately 
<locationDeviation id="9A082F823A0DFFEFFE54BC05FD7E4D64" box="[401,665,547,573]" location="West of Sarajevo" pageId="3" pageNumber="548">
<quantity id="4CFD9B453A0DFFEFFE54BC05FE304D64" box="[401,471,547,573]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="548" unit="km" value="20.0">20 km</quantity>
West of 
<collectingCounty id="62DB4E2C3A0DFFEFFDF2BC05FD7E4D64" box="[567,665,547,573]" pageId="3" pageNumber="548">Sarajevo</collectingCounty>
</locationDeviation>
in Central 
<collectingCountry id="F31276303A0DFFEFFCD6BC05FBC34D64" box="[787,1060,547,573]" name="Bosnia and Herzegovina" pageId="3" pageNumber="548">Bosnia and Herzegovina</collectingCountry>
at a proximal altitude 
<elevation id="0028D1933A0DFFEFFAD9BC05FA7A4D64" box="[1308,1437,547,573]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="548" unit="m" value="700.0">
<quantity id="4CFD9B453A0DFFEFFAD9BC05FA854D64" box="[1308,1378,547,573]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="548" unit="m" value="700.0">700 m</quantity>
a.s.l.
</elevation>
(Coordinates: 
<geoCoordinate id="EE3150673A0DFFEFFEF2BC61FE7F4D38" box="[311,408,583,609]" degrees="43.831" direction="north" orientation="latitude" pageId="3" pageNumber="548" precision="55" value="43.831">43.831N</geoCoordinate>
/ 
<geoCoordinate id="EE3150673A0DFFEFFE6BBC60FDF04D38" box="[430,535,582,609]" degrees="18.075" direction="east" orientation="longitude" pageId="3" pageNumber="548" precision="55" value="18.075">18.075 E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31F652B3A0DFFEBFF02BC4CFE484EAC" lastPageId="7" lastPageNumber="552" pageId="3" pageNumber="548" type="description">
<paragraph id="8BBA36A03A0DFFEFFF02BC4CFB254DF0" blockId="3.[151,1437,474,825]" pageId="3" pageNumber="548">
<emphasis id="B971EAB23A0DFFEFFF02BC4CFE694DDC" bold="true" box="[199,398,618,645]" pageId="3" pageNumber="548">
Description. 
<emphasis id="B971EAB23A0DFFEFFE9ABC4DFE694DDC" bold="true" box="[351,398,619,645]" italics="true" pageId="3" pageNumber="548">Size</emphasis>
</emphasis>
: Total body length (BL) 
<quantity id="4CFD9B453A0DFFEFFD67BC4CFCA34DDC" box="[674,836,618,645]" metricMagnitude="-3" metricUnit="m" metricValue="2.445" metricValueMax="2.52" metricValueMin="2.37" pageId="3" pageNumber="548" unit="mm" value="2.445" valueMax="2.52" valueMin="2.37">2.37–2.52 mm</quantity>
(M= 2.43, N= 4) in 
<specimenCount id="9D03FD293A0DFFEFFBE0BC4DFBA84DDC" box="[1061,1103,619,645]" pageId="3" pageNumber="548" type="male">♂♂</specimenCount>
and 
<quantity id="4CFD9B453A0DFFEFFB41BC4CFAC14DDC" box="[1156,1318,618,645]" metricMagnitude="-3" metricUnit="m" metricValue="2.9349999999999996" metricValueMax="3.53" metricValueMin="2.34" pageId="3" pageNumber="548" unit="mm" value="2.9349999999999996" valueMax="3.53" valueMin="2.34">2.34–3.53 mm</quantity>
(M= 2.44, N= 4) in 
<specimenCount id="9D03FD293A0DFFEFFF38BCA9FECE4DF0" box="[253,297,655,681]" pageId="3" pageNumber="548" type="female">♀♀</specimenCount>
. 
<emphasis id="B971EAB23A0DFFEFFEF0BCA8FE914DF0" bold="true" box="[309,374,654,681]" italics="true" pageId="3" pageNumber="548">Color</emphasis>
: Reddish brown, moderately shiny, antenna and legs slightly paler (
<figureCitation id="133E2A253A0DFFEFFBAABCA9FB564DF0" box="[1135,1201,655,681]" captionStart="FIGURE 1" captionStartId="3.[151,250,1841,1865]" captionTargetBox="[235,1356,861,1807]" captionTargetId="figure-266@3.[215,1371,849,1817]" captionTargetPageId="3" captionText="FIGURE 1. Rudogorites simonei sp. nov. male and female habitus." figureDoi="http://doi.org/10.5281/zenodo.5650000" httpUri="https://zenodo.org/record/5650000/files/figure.png" pageId="3" pageNumber="548">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BBA36A03A0DFFEFFF02BC94FB584C60" blockId="3.[151,1437,474,825]" pageId="3" pageNumber="548">
<emphasis id="B971EAB23A0DFFEFFF02BC94FEE84D94" bold="true" box="[199,271,690,717]" italics="true" pageId="3" pageNumber="548">Head:</emphasis>
Eyes atrophied, head dorsally cowered by sparse erected setae, occipital carina atrophied. Maxillary apical palpomere (4 
<superScript id="7C709BE83A0DFFEFFE97BCF0FEB84DBD" attach="none" box="[338,351,726,740]" fontSize="6" pageId="3" pageNumber="548">th</superScript>
) prolonged conical with slightly concave lateral edges, long of 2/3 of preapical palpomere length, preapical palpomere sub-conical, swollen with convex lateral edges. Mandible regularly curved short and armed with variable number of 1 to 3 short teeth. Antenna inserted on the middle third of head length.
</paragraph>
<caption id="DF7A66283A0DFFEFFF52B917FCA14810" ID-DOI="http://doi.org/10.5281/zenodo.5650000" ID-Zenodo-Dep="5650000" box="[151,838,1841,1866]" httpUri="https://zenodo.org/record/5650000/files/figure.png" pageId="3" pageNumber="548" startId="3.[151,250,1841,1865]" targetBox="[235,1356,861,1807]" targetPageId="3">
<paragraph id="8BBA36A03A0DFFEFFF52B917FCA14810" blockId="3.[151,838,1841,1866]" box="[151,838,1841,1866]" pageId="3" pageNumber="548">
<emphasis id="B971EAB23A0DFFEFFF52B917FEF44810" bold="true" box="[151,275,1841,1865]" pageId="3" pageNumber="548">FIGURE 1.</emphasis>
<taxonomicName id="4C054D233A0DFFEFFEDCB917FE0C4810" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[281,491,1841,1865]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="548" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A0DFFEFFEDCB917FE0C4810" box="[281,491,1841,1865]" italics="true" pageId="3" pageNumber="548">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A0DFFEFFE34B914FDA64813" bold="true" box="[497,577,1842,1866]" pageId="3" pageNumber="548">
<taxonomicNameLabel id="A24257C93A0DFFEFFE34B914FDA64813" box="[497,577,1842,1866]" pageId="3" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male and female habitus.
</paragraph>
</caption>
<paragraph id="8BBA36A03A0DFFE8FF02B95EFCF84FA0" blockId="3.[151,1437,1911,2010]" lastBlockId="4.[151,1437,150,789]" lastPageId="4" lastPageNumber="549" pageId="3" pageNumber="548">
<emphasis id="B971EAB23A0DFFEFFF02B95EFECC48CB" bold="true" box="[199,299,1912,1938]" italics="true" pageId="3" pageNumber="548">Antenna</emphasis>
: (
<figureCitation id="133E2A253A0DFFEFFE84B95EFE6A48CB" box="[321,397,1912,1938]" captionStart="FIGURE 1" captionStartId="3.[151,250,1841,1865]" captionTargetBox="[235,1356,861,1807]" captionTargetId="figure-266@3.[215,1371,849,1817]" captionTargetPageId="3" captionText="FIGURE 1. Rudogorites simonei sp. nov. male and female habitus." figureDoi="http://doi.org/10.5281/zenodo.5650000" httpUri="https://zenodo.org/record/5650000/files/figure.png" pageId="3" pageNumber="548">Figs. 1</figureCitation>
, 
<figureCitation id="133E2A253A0DFFEFFE5FB95EFE0748CB" box="[410,480,1912,1938]" captionStart="FIGURES 17–24" captionStartId="6.[151,264,1257,1281]" captionTargetBox="[173,1414,274,1225]" captionTargetId="figure-80@6.[161,1426,273,1234]" captionTargetPageId="6" captionText="FIGURES 17–24. Figures 17–20. Antenna. 17, Rudogorites simonei sp. nov., male. 18, Rudogorites simonei sp. nov., female. 19, Charonites scheibeli, female. 20, Charonites matzenaueri, male. Figures 21–23, metatergal apparatus, posterior process is indicated with black arrow. 21, Rudogorites simonei sp. nov., male. 22, Charonites matzenaueri, male. 23, Charonites scheibeli, female. Figure 24, metendosternite, Rudogorites simonei sp. nov., male." figureDoi="http://doi.org/10.5281/zenodo.5650006" httpUri="https://zenodo.org/record/5650006/files/figure.png" pageId="3" pageNumber="548">17, 18</figureCitation>
) 
<materialsCitation id="3B6D3CFD3A0DFFE8FE28B95EFCFC4FA0" ID-GBIF-Occurrence="3400807306" collectionCode="PL" lastPageId="4" lastPageNumber="549" pageId="3" pageNumber="548" specimenCount="6" specimenCount-female="3" specimenCount-male="3">
Antenna total length (AL): 
<quantity id="4CFD9B453A0DFFEFFCDEB951FC5A48CB" box="[795,957,1911,1938]" metricMagnitude="-3" metricUnit="m" metricValue="1.87" metricValueMax="1.99" metricValueMin="1.75" pageId="3" pageNumber="548" unit="mm" value="1.87" valueMax="1.99" valueMin="1.75">1.75–1.99 mm</quantity>
(M= 1.86, N= 4) in 
<specimenCount id="9D03FD293A0DFFEFFB59B95EFB2148CB" box="[1180,1222,1912,1938]" pageId="3" pageNumber="548" type="male">♂♂</specimenCount>
and 
<quantity id="4CFD9B453A0DFFEFFB3EB95EFA7A48CB" box="[1275,1437,1912,1938]" metricMagnitude="-3" metricUnit="m" metricValue="1.395" metricValueMax="1.49" metricValueMin="1.3" pageId="3" pageNumber="548" unit="mm" value="1.395" valueMax="1.49" valueMin="1.3">1.30–1.49 mm</quantity>
(M= 1.40, N= 4) in 
<specimenCount id="9D03FD293A0DFFEFFEBFB9BAFE4148EF" box="[378,422,1948,1974]" pageId="3" pageNumber="548" type="female">♀♀</specimenCount>
. AL/BL: 
<quantity id="4CFD9B453A0DFFEFFDD7B9BAFD4748EF" box="[530,672,1948,1974]" metricMagnitude="-2" metricUnit="m" metricValue="1.9431" metricValueMax="2.0066" metricValueMin="1.8796" pageId="3" pageNumber="548" unit="in" value="0.765" valueMax="0.79" valueMin="0.74">0.74–0.79 in</quantity>
<specimenCount id="9D03FD293A0DFFEFFD6DB9BAFD3548EF" box="[680,722,1948,1974]" pageId="3" pageNumber="548" type="male">♂♂</specimenCount>
and 
<quantity id="4CFD9B453A0DFFEFFCCFB9BDFC7F48EF" box="[778,920,1947,1974]" metricMagnitude="-2" metricUnit="m" metricValue="1.4605" metricValueMax="1.4986" metricValueMin="1.4224" pageId="3" pageNumber="548" unit="in" value="0.575" valueMax="0.59" valueMin="0.56">0.56–0.59 in</quantity>
<specimenCount id="9D03FD293A0DFFEFFC5AB9BAFC2D48EF" box="[927,970,1948,1974]" pageId="3" pageNumber="548" type="female">♀♀</specimenCount>
, AL/ 
<collectionCode id="ED14AE653A0DFFEFFBC2B9BAFBCD48EF" box="[1031,1066,1948,1974]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="3" pageNumber="548" type="Museum">PL</collectionCode>
: 
<quantity id="4CFD9B453A0DFFEFFBF1B9BDFB2548EF" box="[1076,1218,1947,1974]" metricMagnitude="-2" metricUnit="m" metricValue="6.985" metricValueMax="7.3152" metricValueMin="6.6548" pageId="3" pageNumber="548" unit="in" value="2.75" valueMax="2.88" valueMin="2.62">2.62–2.88 in</quantity>
<specimenCount id="9D03FD293A0DFFEFFB0FB9BAFB1348EF" box="[1226,1268,1948,1974]" pageId="3" pageNumber="548" type="male">♂♂</specimenCount>
and 2.24– 
<quantity id="4CFD9B453A0DFFEFFAAEB9BAFF4A4883" metricMagnitude="-2" metricUnit="m" metricValue="5.588" pageId="3" pageNumber="548" unit="in" value="2.2">2.20 in</quantity>
<specimenCount id="9D03FD293A0DFFEFFF72B9E6FF044883" box="[183,227,1984,2010]" pageId="3" pageNumber="548" type="female">♀♀</specimenCount>
. First antennomere distinctly shorter but not twice as short as second antennomere. Second antennomere distinctly longer than third, which is slightly longer than following 4 
<superScript id="7C709BE83A0AFFE8FC62BEB0FC534FFD" attach="left" box="[935,948,150,164]" fontSize="6" pageId="4" pageNumber="549">th</superScript>
to 6 
<superScript id="7C709BE83A0AFFE8FC2DBEB0FC124FFD" attach="left" box="[1000,1013,150,164]" fontSize="6" pageId="4" pageNumber="549">th</superScript>
antennomeres which are sub-equally long, 7 
<superScript id="7C709BE83A0AFFE8FF20BE9CFF154F91" attach="none" box="[229,242,186,200]" fontSize="6" pageId="4" pageNumber="549">th</superScript>
, 9 
<superScript id="7C709BE83A0AFFE8FECBBE9CFEFC4F91" attach="left" box="[270,283,186,200]" fontSize="6" pageId="4" pageNumber="549">th</superScript>
and 10 
<superScript id="7C709BE83A0AFFE8FEA8BE9CFE9D4F91" attach="left" box="[365,378,186,200]" fontSize="6" pageId="4" pageNumber="549">th</superScript>
antennomeres prolonged and sub-equally long, 8 
<superScript id="7C709BE83A0AFFE8FC66BE9CFC574F91" attach="left" box="[931,944,186,200]" fontSize="6" pageId="4" pageNumber="549">th</superScript>
antennomere the shortest, 11 
<superScript id="7C709BE83A0AFFE8FB3DBE9CFAE24F91" attach="left" box="[1272,1285,186,200]" fontSize="6" pageId="4" pageNumber="549">th</superScript>
antennomere the longest, elliptical. All antennomeres longer than wide
</materialsCitation>
.
</paragraph>
<paragraph id="8BBA36A03A0AFFE8FF02BF25FE1C4E18" blockId="4.[151,1437,150,789]" pageId="4" pageNumber="549">
AmL 
<specimenCount id="9D03FD293A0AFFE8FEC2BF25FED24E44" box="[263,309,259,285]" pageId="4" pageNumber="549" type="male">♂♂</specimenCount>
: 0.13–0.14; 0.19–0.20; 0.14–0.15; 0.11–0.12; 0.12–0.13; 0.12–0.13; 0.19–0.20; 0.10–0.11; 0.19–0.20; 0.20–0.020; 0.27–0.227 (N= 3).
</paragraph>
<paragraph id="8BBA36A03A0AFFE8FF02BF6DFE384ED0" blockId="4.[151,1437,150,789]" pageId="4" pageNumber="549">
AmL 
<specimenCount id="9D03FD293A0AFFE8FEC2BF6DFED44E3C" box="[263,307,331,357]" pageId="4" pageNumber="549" type="female">♀♀</specimenCount>
: 0.12–0.13; 0.18–0.19; 0.12–0.13; 0.10–0.11; 0.11–0.12; 0.11–0.12; 0.16–0.17; 0.09–0.10; 0.14–0.15; 0.14–0.15; 0.22–0.23 (N= 3).
</paragraph>
<paragraph id="8BBA36A03A0AFFE8FF02BFB5FB514EF4" blockId="4.[151,1437,150,789]" box="[199,1206,402,429]" pageId="4" pageNumber="549">
AL/AmL (in %) 
<specimenCount id="9D03FD293A0AFFE8FE46BFB5FE564EF4" box="[387,433,403,429]" pageId="4" pageNumber="549" type="male">♂♂</specimenCount>
: 7.26; 10.61; 7.82; 6.70; 7.26; 6.70; 11.17; 5.59; 10.61; 11.17; 15.08.
</paragraph>
<paragraph id="8BBA36A03A0AFFE8FF02BF91FB7C4E88" blockId="4.[151,1437,150,789]" box="[199,1179,438,465]" pageId="4" pageNumber="549">
AL/AmL (in %) 
<specimenCount id="9D03FD293A0AFFE8FE46BF91FE484E88" box="[387,431,439,465]" pageId="4" pageNumber="549" type="female">♀♀</specimenCount>
: 7.69; 11.54; 8.33; 7.05; 7.69; 7.05; 10.90; 5.77; 9.62; 9.62; 14.74.
</paragraph>
<paragraph id="8BBA36A03A0AFFE8FF02BFFDFBA34EAC" blockId="4.[151,1437,150,789]" box="[199,1092,475,501]" pageId="4" pageNumber="549">
AmL/AmW 
<specimenCount id="9D03FD293A0AFFE8FE90BFFDFE644EAC" box="[341,387,475,501]" pageId="4" pageNumber="549" type="male">♂♂</specimenCount>
: 1.63; 3.17; 2.80; 2.40; 2.60; 2.40; 2.86; 1.67; 2.71; 2.86; 3.38.
</paragraph>
<paragraph id="8BBA36A03A0AFFE8FF02BFD9FBA34D40" blockId="4.[151,1437,150,789]" box="[199,1092,510,537]" pageId="4" pageNumber="549">
AmL/AmW 
<specimenCount id="9D03FD293A0AFFE8FE90BFD9FE664D40" box="[341,385,511,537]" pageId="4" pageNumber="549" type="female">♀♀</specimenCount>
: 1.50; 3.00; 2.60; 2.20; 2.40; 2.20; 2.13; 1.80; 1.88; 1.67; 2.56.
</paragraph>
<paragraph id="8BBA36A03A0AFFE8FF02BC05FE244DDC" blockId="4.[151,1437,150,789]" pageId="4" pageNumber="549">
Vesicular gland organ (Hamman’s organ) in 7 
<superScript id="7C709BE83A0AFFE8FD02BC04FD334D69" attach="left" box="[711,724,546,560]" fontSize="6" pageId="4" pageNumber="549">th</superScript>
antennomere voluminous, spherical with short but distinct vestibulum (duct) opened to atrium. Sensory glands on distal edge of 9 
<superScript id="7C709BE83A0AFFE8FCB8BC60FC6D4D0D" attach="left" box="[893,906,582,596]" fontSize="6" pageId="4" pageNumber="549">th</superScript>
and 10 
<superScript id="7C709BE83A0AFFE8FC1FBC60FC004D0D" attach="left" box="[986,999,582,596]" fontSize="6" pageId="4" pageNumber="549">th</superScript>
antennomere present as hemi-spherical anteriorly opened vesicles.
</paragraph>
<paragraph id="8BBA36A03A0AFFE8FF02BCA9FF304C4C" blockId="4.[151,1437,150,789]" pageId="4" pageNumber="549">
<emphasis id="B971EAB23A0AFFE8FF02BCA9FEDB4DF0" bold="true" box="[199,316,655,681]" italics="true" pageId="4" pageNumber="549">Pronotum</emphasis>
: Transverse, wider than long, distinctly narrower than elytra width, making the habitus of sub-pholeuonoid aspect (
<figureCitation id="133E2A253A0AFFE8FE98BC95FE7D4D97" box="[349,410,691,718]" captionStart="FIGURE 1" captionStartId="3.[151,250,1841,1865]" captionTargetBox="[235,1356,861,1807]" captionTargetId="figure-266@3.[215,1371,849,1817]" captionTargetPageId="3" captionText="FIGURE 1. Rudogorites simonei sp. nov. male and female habitus." figureDoi="http://doi.org/10.5281/zenodo.5650000" httpUri="https://zenodo.org/record/5650000/files/figure.png" pageId="4" pageNumber="549">Fig.1</figureCitation>
). Lateral edge in dorsal observation (
<figureCitation id="133E2A253A0AFFE8FC8ABC95FC584D94" box="[847,959,691,717]" captionStart="FIGURES 2–10" captionStartId="4.[151,264,1813,1837]" captionTargetBox="[159,1421,820,1785]" captionTargetId="figure-257@4.[159,1427,813,1790]" captionTargetPageId="4" captionText="FIGURES 2–10. Figures 2–5. pronotum, ventral view. 2, Rudogorites simonei sp. nov., male. 3, Rudogorites simonei sp. nov., female. 4, Charonites scheibeli, female. 5, Charonites matzenaueri, male. Figure 6, elytra surface detail Rudogorites simonei sp. nov., male. Figures 7–8, thorax ventral view, mesothoracic apophysis are indicated with white arrow, sutures between mesosetrnum and mesepisternum are indicated with grey arrow. 7, Rudogorites simonei sp. nov., male. 8, Charonites matzenaueri, male. Figures 9–10, thorax lateral view, ventral edge of mesosternal keel is indicated with black arrow. 9, Rudogorites simonei sp. nov., male. 10, Charonites matzenaueri, male." figureDoi="http://doi.org/10.5281/zenodo.5650002" httpUri="https://zenodo.org/record/5650002/files/figure.png" pageId="4" pageNumber="549">Figs. 2, 3</figureCitation>
) slightly sinuate concave, bell-like form, maximum width on posterior edge. Dorsal face strongly pubescent, with uniform distribution of short semierect setae.
</paragraph>
<caption id="DF7A66283A0AFFE8FF52B933FD7648B8" ID-DOI="http://doi.org/10.5281/zenodo.5650002" ID-Zenodo-Dep="5650002" httpUri="https://zenodo.org/record/5650002/files/figure.png" pageId="4" pageNumber="549" startId="4.[151,264,1813,1837]" targetBox="[159,1421,820,1785]" targetPageId="4">
<paragraph id="8BBA36A03A0AFFE8FF52B933FD7648B8" blockId="4.[151,1436,1813,2017]" pageId="4" pageNumber="549">
<emphasis id="B971EAB23A0AFFE8FF52B933FEA04874" bold="true" box="[151,327,1813,1837]" pageId="4" pageNumber="549">FIGURES 2–10.</emphasis>
Figures 2–5. pronotum, ventral view. 2, 
<taxonomicName id="4C054D233A0AFFE8FD26B933FC534874" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[739,948,1813,1837]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A0AFFE8FD26B933FC534874" box="[739,948,1813,1837]" italics="true" pageId="4" pageNumber="549">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A0AFFE8FC7FB933FBED4874" bold="true" box="[954,1034,1813,1837]" pageId="4" pageNumber="549">
<taxonomicNameLabel id="A24257C93A0AFFE8FC7FB933FBED4874" box="[954,1034,1813,1837]" pageId="4" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. 3, 
<taxonomicName id="4C054D233A0AFFE8FBABB933FAD84874" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[1134,1343,1813,1837]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A0AFFE8FBABB933FAD84874" box="[1134,1343,1813,1837]" italics="true" pageId="4" pageNumber="549">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A0AFFE8FA80B933FA724874" bold="true" box="[1349,1429,1813,1837]" pageId="4" pageNumber="549">
<taxonomicNameLabel id="A24257C93A0AFFE8FA80B933FA724874" box="[1349,1429,1813,1837]" pageId="4" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. 4, 
<taxonomicName id="4C054D233A0AFFE8FEC0B91FFE314808" box="[261,470,1849,1873]" class="Insecta" family="Leiodidae" genus="Charonites" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="species" species="scheibeli">
<emphasis id="B971EAB23A0AFFE8FEC0B91FFE314808" box="[261,470,1849,1873]" italics="true" pageId="4" pageNumber="549">Charonites scheibeli</emphasis>
</taxonomicName>
, female. 5, 
<taxonomicName id="4C054D233A0AFFE8FD97B91FFCA14808" box="[594,838,1849,1873]" class="Insecta" family="Leiodidae" genus="Charonites" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="species" species="matzenaueri">
<emphasis id="B971EAB23A0AFFE8FD97B91FFCA14808" box="[594,838,1849,1873]" italics="true" pageId="4" pageNumber="549">Charonites matzenaueri</emphasis>
</taxonomicName>
, male. Figure 6, elytra surface detail 
<taxonomicName id="4C054D233A0AFFE8FB0FB91FFA7B4808" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[1226,1436,1849,1873]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A0AFFE8FB0FB91FFA7B4808" box="[1226,1436,1849,1873]" italics="true" pageId="4" pageNumber="549">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A0AFFE8FF52B97BFF00482C" bold="true" box="[151,231,1885,1909]" pageId="4" pageNumber="549">
<taxonomicNameLabel id="A24257C93A0AFFE8FF52B97BFF00482C" box="[151,231,1885,1909]" pageId="4" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. Figures 7–8, thorax ventral view, mesothoracic apophysis are indicated with white arrow, sutures between mesosetrnum and mesepisternum are indicated with grey arrow. 7, 
<taxonomicName id="4C054D233A0AFFE8FCD5B9A7FC0548C0" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[784,994,1921,1945]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A0AFFE8FCD5B9A7FC0548C0" box="[784,994,1921,1945]" italics="true" pageId="4" pageNumber="549">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A0AFFE8FC2CB9A7FBDC48C0" bold="true" box="[1001,1083,1921,1945]" pageId="4" pageNumber="549">
<taxonomicNameLabel id="A24257C93A0AFFE8FC2CB9A7FBDC48C0" box="[1001,1083,1921,1945]" pageId="4" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. 8, 
<taxonomicName id="4C054D233A0AFFE8FB67B9A7FA7148C0" box="[1186,1430,1921,1945]" class="Insecta" family="Leiodidae" genus="Charonites" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="species" species="matzenaueri">
<emphasis id="B971EAB23A0AFFE8FB67B9A7FA7148C0" box="[1186,1430,1921,1945]" italics="true" pageId="4" pageNumber="549">Charonites matzenaueri</emphasis>
</taxonomicName>
, male. Figures 9–10, thorax lateral view, ventral edge of mesosternal keel is indicated with black arrow. 9, 
<taxonomicName id="4C054D233A0AFFE8FB0EB983FA7B48E4" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[1227,1436,1957,1981]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A0AFFE8FB0EB983FA7B48E4" box="[1227,1436,1957,1981]" italics="true" pageId="4" pageNumber="549">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A0AFFE8FF52B9EFFF0F48B8" bold="true" box="[151,232,1993,2017]" pageId="4" pageNumber="549">
<taxonomicNameLabel id="A24257C93A0AFFE8FF52B9EFFF0F48B8" box="[151,232,1993,2017]" pageId="4" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. 10, 
<taxonomicName id="4C054D233A0AFFE8FE9CB9EFFDAB48B8" box="[345,588,1993,2017]" class="Insecta" family="Leiodidae" genus="Charonites" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="species" species="matzenaueri">
<emphasis id="B971EAB23A0AFFE8FE9CB9EFFDAB48B8" box="[345,588,1993,2017]" italics="true" pageId="4" pageNumber="549">Charonites matzenaueri</emphasis>
</taxonomicName>
, male.
</paragraph>
</caption>
<caption id="DF7A66283A0BFFE9FF52BA74FA9F4BEB" ID-DOI="http://doi.org/10.5281/zenodo.5650004" ID-Zenodo-Dep="5650004" httpUri="https://zenodo.org/record/5650004/files/figure.png" pageId="5" pageNumber="550" startId="5.[151,264,1106,1130]" targetBox="[172,1417,188,1074]" targetPageId="5">
<paragraph id="8BBA36A03A0BFFE9FF52BA74FA9F4BEB" blockId="5.[151,1437,1106,1203]" pageId="5" pageNumber="550">
<emphasis id="B971EAB23A0BFFE9FF52BA74FEB44B33" bold="true" box="[151,339,1106,1130]" pageId="5" pageNumber="550">FIGURES 11–16.</emphasis>
Figures 11–14, left first leg, comb of protibial spines is indicated with black arrow. 11, 
<taxonomicName id="4C054D233A0BFFE9FB0EBA74FA7B4B33" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[1227,1436,1106,1130]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="550" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A0BFFE9FB0EBA74FA7B4B33" box="[1227,1436,1106,1130]" italics="true" pageId="5" pageNumber="550">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A0BFFE9FF52BA51FF0F4BD6" bold="true" box="[151,232,1143,1167]" pageId="5" pageNumber="550">
<taxonomicNameLabel id="A24257C93A0BFFE9FF52BA51FF0F4BD6" box="[151,232,1143,1167]" pageId="5" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. 12, 
<taxonomicName id="4C054D233A0BFFE9FEAABA50FDA64BD7" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[367,577,1142,1166]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="550" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A0BFFE9FEAABA50FDA64BD7" box="[367,577,1142,1166]" italics="true" pageId="5" pageNumber="550">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A0BFFE9FD8DBA51FD7E4BD6" bold="true" box="[584,665,1143,1167]" pageId="5" pageNumber="550">
<taxonomicNameLabel id="A24257C93A0BFFE9FD8DBA51FD7E4BD6" box="[584,665,1143,1167]" pageId="5" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. 13, 
<taxonomicName id="4C054D233A0BFFE9FCC9BA50FC3A4BD7" box="[780,989,1142,1166]" class="Insecta" family="Leiodidae" genus="Charonites" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="550" phylum="Arthropoda" rank="species" species="scheibeli">
<emphasis id="B971EAB23A0BFFE9FCC9BA50FC3A4BD7" box="[780,989,1142,1166]" italics="true" pageId="5" pageNumber="550">Charonites scheibeli</emphasis>
</taxonomicName>
, female. 14, 
<taxonomicName id="4C054D233A0BFFE9FBA6BA50FAB04BD7" box="[1123,1367,1142,1166]" class="Insecta" family="Leiodidae" genus="Charonites" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="550" phylum="Arthropoda" rank="species" species="matzenaueri">
<emphasis id="B971EAB23A0BFFE9FBA6BA50FAB04BD7" box="[1123,1367,1142,1166]" italics="true" pageId="5" pageNumber="550">Charonites matzenaueri</emphasis>
</taxonomicName>
, male. Figure 15, second left leg, 
<taxonomicName id="4C054D233A0BFFE9FE63BABCFD904BEB" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[422,631,1178,1202]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="550" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A0BFFE9FE63BABCFD904BEB" box="[422,631,1178,1202]" italics="true" pageId="5" pageNumber="550">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A0BFFE9FDB8BABDFD294BEA" bold="true" box="[637,718,1179,1203]" pageId="5" pageNumber="550">
<taxonomicNameLabel id="A24257C93A0BFFE9FDB8BABDFD294BEA" box="[637,718,1179,1203]" pageId="5" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. Figure 16, third leg, left 
<taxonomicName id="4C054D233A0BFFE9FBD4BABCFB054BEB" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[1041,1250,1178,1202]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="550" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A0BFFE9FBD4BABCFB054BEB" box="[1041,1250,1178,1202]" italics="true" pageId="5" pageNumber="550">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A0BFFE9FB2CBABDFADE4BEA" bold="true" box="[1257,1337,1179,1203]" pageId="5" pageNumber="550">
<taxonomicNameLabel id="A24257C93A0BFFE9FB2CBABDFADE4BEA" box="[1257,1337,1179,1203]" pageId="5" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male.
</paragraph>
</caption>
<paragraph id="8BBA36A03A0BFFE9FF02BAC7FC174A3E" blockId="5.[151,1437,1249,2032]" pageId="5" pageNumber="550">
<materialsCitation id="3B6D3CFD3A0BFFE9FF02BAC7FD564BA2" ID-GBIF-Occurrence="3400807302" box="[199,689,1249,1276]" collectionCode="PL" pageId="5" pageNumber="550" specimenCount="1" specimenCount-male="1">
<collectionCode id="ED14AE653A0BFFE9FF02BAC7FF0B4BA2" box="[199,236,1249,1275]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="5" pageNumber="550" type="Museum">PL</collectionCode>
: 
<quantity id="4CFD9B453A0BFFE9FF3DBAC7FE7B4BA2" box="[248,412,1249,1275]" metricMagnitude="-4" metricUnit="m" metricValue="6.45" metricValueMax="6.8999999999999995" metricValueMin="6.0" pageId="5" pageNumber="550" unit="mm" value="0.645" valueMax="0.69" valueMin="0.6">0.60–0.69 mm</quantity>
(M= 0.63, N= 5) in 
<specimenCount id="9D03FD293A0BFFE9FD42BAC7FD564BA2" box="[647,689,1249,1275]" pageId="5" pageNumber="550" type="male">♂♂</specimenCount>
</materialsCitation>
and 
<quantity id="4CFD9B453A0BFFE9FD2FBAC7FC694BA2" box="[746,910,1249,1276]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" metricValueMax="6.8" metricValueMin="5.8" pageId="5" pageNumber="550" unit="mm" value="0.63" valueMax="0.68" valueMin="0.58">0.58–0.68 mm</quantity>
(M= 0.64, N= 6) in 
<specimenCount id="9D03FD293A0BFFE9FBBCBAC7FB434BA2" box="[1145,1188,1249,1275]" pageId="5" pageNumber="550" type="female">♀♀</specimenCount>
; PaW: 
<quantity id="4CFD9B453A0BFFE9FB3CBAC7FA7A4BA2" box="[1273,1437,1249,1276]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.4" metricValueMin="4.9" pageId="5" pageNumber="550" unit="mm" value="0.515" valueMax="0.54" valueMin="0.49">0.49–0.54 mm</quantity>
(M= 0.51, N= 2) in 
<specimenCount id="9D03FD293A0BFFE9FEBCBB23FE444A46" box="[377,419,1285,1311]" pageId="5" pageNumber="550" type="male">♂♂</specimenCount>
and 
<quantity id="4CFD9B453A0BFFE9FE1EBB23FD994A46" box="[475,638,1285,1312]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" metricValueMax="5.5" metricValueMin="5.3" pageId="5" pageNumber="550" unit="mm" value="0.54" valueMax="0.55" valueMin="0.53">0.53–0.55 mm</quantity>
(M= 0.54, N= 3) in 
<specimenCount id="9D03FD293A0BFFE9FCA2BB23FC754A46" box="[871,914,1285,1311]" pageId="5" pageNumber="550" type="female">♀♀</specimenCount>
; PcW: 
<quantity id="4CFD9B453A0BFFE9FC23BB23FB6E4A46" box="[998,1161,1285,1311]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="8.299999999999999" metricValueMin="7.7" pageId="5" pageNumber="550" unit="mm" value="0.8" valueMax="0.83" valueMin="0.77">0.77–0.83 mm</quantity>
(M= 0.81, N= 5) in 
<specimenCount id="9D03FD293A0BFFE9FAB7BB23FA7B4A46" box="[1394,1436,1285,1311]" pageId="5" pageNumber="550" type="male">♂♂</specimenCount>
and 
<quantity id="4CFD9B453A0BFFE9FF02BB0FFE8C4A1A" box="[199,363,1321,1347]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" metricValueMax="8.9" metricValueMin="7.9" pageId="5" pageNumber="550" unit="mm" value="0.84" valueMax="0.89" valueMin="0.79">0.79–0.89 mm</quantity>
(M= 0.83, N= 6) in 
<specimenCount id="9D03FD293A0BFFE9FD96BB0FFD994A1A" box="[595,638,1321,1347]" pageId="5" pageNumber="550" type="female">♀♀</specimenCount>
; PpW: 
<quantity id="4CFD9B453A0BFFE9FD17BB0FFC914A1A" box="[722,886,1321,1347]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" metricValueMax="8.9" metricValueMin="8.7" pageId="5" pageNumber="550" unit="mm" value="0.88" valueMax="0.89" valueMin="0.87">0.87–0.89 mm</quantity>
(M= 0.88, N= 5) in 
<specimenCount id="9D03FD293A0BFFE9FB9BBB0FFB6F4A1A" box="[1118,1160,1321,1347]" pageId="5" pageNumber="550" type="male">♂♂</specimenCount>
and 
<quantity id="4CFD9B453A0BFFE9FB05BB0FFA844A1A" box="[1216,1379,1321,1347]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="9.7" metricValueMin="9.3" pageId="5" pageNumber="550" unit="mm" value="0.95" valueMax="0.97" valueMin="0.93">0.93–0.97 mm</quantity>
(M= 
<geoCoordinate id="EE3150673A0BFFE9FF52BB6BFF034A3E" box="[151,228,1357,1383]" degrees="0.96" direction="north" orientation="latitude" pageId="5" pageNumber="550" precision="555" value="0.96">0.96 N</geoCoordinate>
= 6) in 
<specimenCount id="9D03FD293A0BFFE9FEF0BB6BFE874A3E" box="[309,352,1357,1383]" pageId="5" pageNumber="550" type="female">♀♀</specimenCount>
; 
<materialsCitation id="3B6D3CFD3A0BFFE9FEABBB6BFCE24A3E" ID-GBIF-Occurrence="3400807307" box="[366,773,1357,1383]" collectionCode="PL" pageId="5" pageNumber="550" specimenCount="1" specimenCount-male="1">
<collectionCode id="ED14AE653A0BFFE9FEABBB6BFE694A3E" box="[366,398,1357,1383]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="5" pageNumber="550" type="Museum">PL</collectionCode>
/PpW on average M= 
<quantity id="4CFD9B453A0BFFE9FD43BB6BFD334A3E" box="[646,724,1357,1383]" metricMagnitude="-2" metricUnit="m" metricValue="1.8288" pageId="5" pageNumber="550" unit="in" value="0.72">0.72 in</quantity>
<specimenCount id="9D03FD293A0BFFE9FD1EBB6BFCE24A3E" box="[731,773,1357,1383]" pageId="5" pageNumber="550" type="male">♂♂</specimenCount>
</materialsCitation>
and M= 
<quantity id="4CFD9B453A0BFFE9FCAEBB6BFC5E4A3E" box="[875,953,1357,1383]" metricMagnitude="-2" metricUnit="m" metricValue="1.7018" pageId="5" pageNumber="550" unit="in" value="0.67">0.67 in</quantity>
<specimenCount id="9D03FD293A0BFFE9FC7ABB6BFC0C4A3E" box="[959,1003,1357,1383]" pageId="5" pageNumber="550" type="female">♀♀</specimenCount>
.
</paragraph>
<paragraph id="8BBA36A03A0BFFE9FF02BB56FE7549F2" blockId="5.[151,1437,1249,2032]" pageId="5" pageNumber="550">
<emphasis id="B971EAB23A0BFFE9FF02BB56FEC14AD2" bold="true" box="[199,294,1392,1419]" italics="true" pageId="5" pageNumber="550">Thorax:</emphasis>
Mesocoxal cavities separated by a keel, not confluent. Mesosternal keel (carena mesosternale) well developed, elevated and flattened, lanceolate, posteriorly reduced and thickened, extended to the posterior edge of coxal cavities (
<figureCitation id="133E2A253A0BFFE9FE81BB9FFE544A8A" box="[324,435,1465,1491]" captionStart="FIGURES 2–10" captionStartId="4.[151,264,1813,1837]" captionTargetBox="[159,1421,820,1785]" captionTargetId="figure-257@4.[159,1427,813,1790]" captionTargetPageId="4" captionText="FIGURES 2–10. Figures 2–5. pronotum, ventral view. 2, Rudogorites simonei sp. nov., male. 3, Rudogorites simonei sp. nov., female. 4, Charonites scheibeli, female. 5, Charonites matzenaueri, male. Figure 6, elytra surface detail Rudogorites simonei sp. nov., male. Figures 7–8, thorax ventral view, mesothoracic apophysis are indicated with white arrow, sutures between mesosetrnum and mesepisternum are indicated with grey arrow. 7, Rudogorites simonei sp. nov., male. 8, Charonites matzenaueri, male. Figures 9–10, thorax lateral view, ventral edge of mesosternal keel is indicated with black arrow. 9, Rudogorites simonei sp. nov., male. 10, Charonites matzenaueri, male." figureDoi="http://doi.org/10.5281/zenodo.5650002" httpUri="https://zenodo.org/record/5650002/files/figure.png" pageId="5" pageNumber="550">Figs. 7, 9</figureCitation>
). In lateral view keel is elevated with ventral edge straight, not dentate and entire from thorax colum to the posterior edge of mesocoxal cavities (
<figureCitation id="133E2A253A0BFFE9FCD8BBFBFC864AAE" box="[797,865,1501,1527]" captionStart="FIGURES 2–10" captionStartId="4.[151,264,1813,1837]" captionTargetBox="[159,1421,820,1785]" captionTargetId="figure-257@4.[159,1427,813,1790]" captionTargetPageId="4" captionText="FIGURES 2–10. Figures 2–5. pronotum, ventral view. 2, Rudogorites simonei sp. nov., male. 3, Rudogorites simonei sp. nov., female. 4, Charonites scheibeli, female. 5, Charonites matzenaueri, male. Figure 6, elytra surface detail Rudogorites simonei sp. nov., male. Figures 7–8, thorax ventral view, mesothoracic apophysis are indicated with white arrow, sutures between mesosetrnum and mesepisternum are indicated with grey arrow. 7, Rudogorites simonei sp. nov., male. 8, Charonites matzenaueri, male. Figures 9–10, thorax lateral view, ventral edge of mesosternal keel is indicated with black arrow. 9, Rudogorites simonei sp. nov., male. 10, Charonites matzenaueri, male." figureDoi="http://doi.org/10.5281/zenodo.5650002" httpUri="https://zenodo.org/record/5650002/files/figure.png" pageId="5" pageNumber="550">Fig. 9</figureCitation>
). Mesosternal intercoxal apophysis bifid, short and narrow, equally long as wide (
<figureCitation id="133E2A253A0BFFE9FE34B827FDD04942" box="[497,567,1537,1563]" captionStart="FIGURES 2–10" captionStartId="4.[151,264,1813,1837]" captionTargetBox="[159,1421,820,1785]" captionTargetId="figure-257@4.[159,1427,813,1790]" captionTargetPageId="4" captionText="FIGURES 2–10. Figures 2–5. pronotum, ventral view. 2, Rudogorites simonei sp. nov., male. 3, Rudogorites simonei sp. nov., female. 4, Charonites scheibeli, female. 5, Charonites matzenaueri, male. Figure 6, elytra surface detail Rudogorites simonei sp. nov., male. Figures 7–8, thorax ventral view, mesothoracic apophysis are indicated with white arrow, sutures between mesosetrnum and mesepisternum are indicated with grey arrow. 7, Rudogorites simonei sp. nov., male. 8, Charonites matzenaueri, male. Figures 9–10, thorax lateral view, ventral edge of mesosternal keel is indicated with black arrow. 9, Rudogorites simonei sp. nov., male. 10, Charonites matzenaueri, male." figureDoi="http://doi.org/10.5281/zenodo.5650002" httpUri="https://zenodo.org/record/5650002/files/figure.png" pageId="5" pageNumber="550">Fig. 7</figureCitation>
). Suture between mesosetrnum and mesepisternum is partly atrophied on its anterior side, not extending to the colum (
<figureCitation id="133E2A253A0BFFE9FDA2B803FD4D4966" box="[615,682,1573,1599]" captionStart="FIGURES 2–10" captionStartId="4.[151,264,1813,1837]" captionTargetBox="[159,1421,820,1785]" captionTargetId="figure-257@4.[159,1427,813,1790]" captionTargetPageId="4" captionText="FIGURES 2–10. Figures 2–5. pronotum, ventral view. 2, Rudogorites simonei sp. nov., male. 3, Rudogorites simonei sp. nov., female. 4, Charonites scheibeli, female. 5, Charonites matzenaueri, male. Figure 6, elytra surface detail Rudogorites simonei sp. nov., male. Figures 7–8, thorax ventral view, mesothoracic apophysis are indicated with white arrow, sutures between mesosetrnum and mesepisternum are indicated with grey arrow. 7, Rudogorites simonei sp. nov., male. 8, Charonites matzenaueri, male. Figures 9–10, thorax lateral view, ventral edge of mesosternal keel is indicated with black arrow. 9, Rudogorites simonei sp. nov., male. 10, Charonites matzenaueri, male." figureDoi="http://doi.org/10.5281/zenodo.5650002" httpUri="https://zenodo.org/record/5650002/files/figure.png" pageId="5" pageNumber="550">Fig. 7</figureCitation>
). Metendosternite (furca or Crowson’s organ) “V” shaped with dorsal arms twice as long as stalk (
<figureCitation id="133E2A253A0BFFE9FE38B86FFDB5493A" box="[509,594,1609,1635]" captionStart="FIGURES 17–24" captionStartId="6.[151,264,1257,1281]" captionTargetBox="[173,1414,274,1225]" captionTargetId="figure-80@6.[161,1426,273,1234]" captionTargetPageId="6" captionText="FIGURES 17–24. Figures 17–20. Antenna. 17, Rudogorites simonei sp. nov., male. 18, Rudogorites simonei sp. nov., female. 19, Charonites scheibeli, female. 20, Charonites matzenaueri, male. Figures 21–23, metatergal apparatus, posterior process is indicated with black arrow. 21, Rudogorites simonei sp. nov., male. 22, Charonites matzenaueri, male. 23, Charonites scheibeli, female. Figure 24, metendosternite, Rudogorites simonei sp. nov., male." figureDoi="http://doi.org/10.5281/zenodo.5650006" httpUri="https://zenodo.org/record/5650006/files/figure.png" pageId="5" pageNumber="550">Fig. 24</figureCitation>
). Metatergal apparatus with medial longitudinal groove (posterior process) strongly reduced, rudimentary and oversized by scutellum posterior beak (
<figureCitation id="133E2A253A0BFFE9FC13B84BFBCE49DE" box="[982,1065,1645,1671]" captionStart="FIGURES 17–24" captionStartId="6.[151,264,1257,1281]" captionTargetBox="[173,1414,274,1225]" captionTargetId="figure-80@6.[161,1426,273,1234]" captionTargetPageId="6" captionText="FIGURES 17–24. Figures 17–20. Antenna. 17, Rudogorites simonei sp. nov., male. 18, Rudogorites simonei sp. nov., female. 19, Charonites scheibeli, female. 20, Charonites matzenaueri, male. Figures 21–23, metatergal apparatus, posterior process is indicated with black arrow. 21, Rudogorites simonei sp. nov., male. 22, Charonites matzenaueri, male. 23, Charonites scheibeli, female. Figure 24, metendosternite, Rudogorites simonei sp. nov., male." figureDoi="http://doi.org/10.5281/zenodo.5650006" httpUri="https://zenodo.org/record/5650006/files/figure.png" pageId="5" pageNumber="550">Fig. 21</figureCitation>
). Scutellum triangular, wide than long, mostly glabrous.
</paragraph>
<paragraph id="8BBA36A03A0BFFE9FF02B892FBD748DA" blockId="5.[151,1437,1249,2032]" pageId="5" pageNumber="550">
<emphasis id="B971EAB23A0BFFE9FF02B892FEE84996" bold="true" box="[199,271,1716,1743]" italics="true" pageId="5" pageNumber="550">Elytra</emphasis>
: Elytra elongated-oval with maximum width on medial part of elytra (
<figureCitation id="133E2A253A0BFFE9FBEAB893FB964996" box="[1071,1137,1717,1743]" captionStart="FIGURE 1" captionStartId="3.[151,250,1841,1865]" captionTargetBox="[235,1356,861,1807]" captionTargetId="figure-266@3.[215,1371,849,1817]" captionTargetPageId="3" captionText="FIGURE 1. Rudogorites simonei sp. nov. male and female habitus." figureDoi="http://doi.org/10.5281/zenodo.5650000" httpUri="https://zenodo.org/record/5650000/files/figure.png" pageId="5" pageNumber="550">Fig. 1</figureCitation>
). Sparsely pubescent with uniform short and semi erected, not recumbent setae (
<figureCitation id="133E2A253A0BFFE9FCCFB8FFFCB549AA" box="[778,850,1753,1779]" captionStart="FIGURE 1" captionStartId="3.[151,250,1841,1865]" captionTargetBox="[235,1356,861,1807]" captionTargetId="figure-266@3.[215,1371,849,1817]" captionTargetPageId="3" captionText="FIGURE 1. Rudogorites simonei sp. nov. male and female habitus." figureDoi="http://doi.org/10.5281/zenodo.5650000" httpUri="https://zenodo.org/record/5650000/files/figure.png" pageId="5" pageNumber="550">Figs.1</figureCitation>
, 
<figureCitation id="133E2A253A0BFFE9FCA4B8FFFC9649AD" box="[865,881,1753,1780]" captionStart="FIGURES 2–10" captionStartId="4.[151,264,1813,1837]" captionTargetBox="[159,1421,820,1785]" captionTargetId="figure-257@4.[159,1427,813,1790]" captionTargetPageId="4" captionText="FIGURES 2–10. Figures 2–5. pronotum, ventral view. 2, Rudogorites simonei sp. nov., male. 3, Rudogorites simonei sp. nov., female. 4, Charonites scheibeli, female. 5, Charonites matzenaueri, male. Figure 6, elytra surface detail Rudogorites simonei sp. nov., male. Figures 7–8, thorax ventral view, mesothoracic apophysis are indicated with white arrow, sutures between mesosetrnum and mesepisternum are indicated with grey arrow. 7, Rudogorites simonei sp. nov., male. 8, Charonites matzenaueri, male. Figures 9–10, thorax lateral view, ventral edge of mesosternal keel is indicated with black arrow. 9, Rudogorites simonei sp. nov., male. 10, Charonites matzenaueri, male." figureDoi="http://doi.org/10.5281/zenodo.5650002" httpUri="https://zenodo.org/record/5650002/files/figure.png" pageId="5" pageNumber="550">6</figureCitation>
). Elytral punctation strong with large, deep and widely spaced foveolated punctures, not aligned in transversal strigae. Each seta growing from the bottom of a depressed puncture (
<figureCitation id="133E2A253A0BFFE9FE45B907FE214865" box="[384,454,1825,1852]" captionStart="FIGURES 2–10" captionStartId="4.[151,264,1813,1837]" captionTargetBox="[159,1421,820,1785]" captionTargetId="figure-257@4.[159,1427,813,1790]" captionTargetPageId="4" captionText="FIGURES 2–10. Figures 2–5. pronotum, ventral view. 2, Rudogorites simonei sp. nov., male. 3, Rudogorites simonei sp. nov., female. 4, Charonites scheibeli, female. 5, Charonites matzenaueri, male. Figure 6, elytra surface detail Rudogorites simonei sp. nov., male. Figures 7–8, thorax ventral view, mesothoracic apophysis are indicated with white arrow, sutures between mesosetrnum and mesepisternum are indicated with grey arrow. 7, Rudogorites simonei sp. nov., male. 8, Charonites matzenaueri, male. Figures 9–10, thorax lateral view, ventral edge of mesosternal keel is indicated with black arrow. 9, Rudogorites simonei sp. nov., male. 10, Charonites matzenaueri, male." figureDoi="http://doi.org/10.5281/zenodo.5650002" httpUri="https://zenodo.org/record/5650002/files/figure.png" pageId="5" pageNumber="550">Fig. 6</figureCitation>
). Sutural carina absent. Elytra distinctly wider in 
<specimenCount id="9D03FD293A0BFFE9FC38B907FBC04862" box="[1021,1063,1825,1851]" pageId="5" pageNumber="550" type="female">♀♀</specimenCount>
than in 
<specimenCount id="9D03FD293A0BFFE9FB43B907FB544862" box="[1158,1203,1825,1851]" pageId="5" pageNumber="550" type="male">♂♂</specimenCount>
. EL: 1.49.– 
<quantity id="4CFD9B453A0BFFE9FAF3B907FA7A4862" box="[1334,1437,1825,1851]" metricMagnitude="-3" metricUnit="m" metricValue="1.69" pageId="5" pageNumber="550" unit="mm" value="1.69">1.69 mm</quantity>
(M= 1.58, N= 3) in 
<specimenCount id="9D03FD293A0BFFE9FEB4B963FE7C4806" box="[369,411,1861,1887]" pageId="5" pageNumber="550" type="male">♂♂</specimenCount>
and 
<quantity id="4CFD9B453A0BFFE9FE15B963FD954806" box="[464,626,1861,1888]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" metricValueMax="1.74" metricValueMin="1.5" pageId="5" pageNumber="550" unit="mm" value="1.62" valueMax="1.74" valueMin="1.5">1.50–1.74 mm</quantity>
(M= 1.61, N= 3) in 
<specimenCount id="9D03FD293A0BFFE9FC97B963FC9A4806" box="[850,893,1861,1887]" pageId="5" pageNumber="550" type="female">♀♀</specimenCount>
; EW: 
<quantity id="4CFD9B453A0BFFE9FC07B963FB834806" box="[962,1124,1861,1887]" metricMagnitude="-3" metricUnit="m" metricValue="1.185" metricValueMax="1.23" metricValueMin="1.14" pageId="5" pageNumber="550" unit="mm" value="1.185" valueMax="1.23" valueMin="1.14">1.14–1.23 mm</quantity>
(M= 1.18, N= 3) in 
<specimenCount id="9D03FD293A0BFFE9FA81B963FA894806" box="[1348,1390,1861,1887]" pageId="5" pageNumber="550" type="male">♂♂</specimenCount>
and 
<quantity id="4CFD9B453A0BFFE9FF52B94FFEDD48DA" box="[151,314,1897,1924]" metricMagnitude="-3" metricUnit="m" metricValue="1.205" metricValueMax="1.25" metricValueMin="1.16" pageId="5" pageNumber="550" unit="mm" value="1.205" valueMax="1.25" valueMin="1.16">1.16–1.25 mm</quantity>
(M= 1.20, N= 3) in 
<specimenCount id="9D03FD293A0BFFE9FDDAB94FFDAD48DA" box="[543,586,1897,1923]" pageId="5" pageNumber="550" type="female">♀♀</specimenCount>
; ratio EL/EW: 
<quantity id="4CFD9B453A0BFFE9FD30B94FFCA448DA" box="[757,835,1897,1923]" metricMagnitude="-2" metricUnit="m" metricValue="3.5559999999999996" pageId="5" pageNumber="550" unit="in" value="1.4">1.40 in</quantity>
<specimenCount id="9D03FD293A0BFFE9FC8FB94FFC9348DA" box="[842,884,1897,1923]" pageId="5" pageNumber="550" type="male">♂♂</specimenCount>
and 
<quantity id="4CFD9B453A0BFFE9FC6FB94FFC1F48DA" box="[938,1016,1897,1924]" metricMagnitude="-2" metricUnit="m" metricValue="3.429" pageId="5" pageNumber="550" unit="in" value="1.35">1.35 in</quantity>
<specimenCount id="9D03FD293A0BFFE9FC3AB94FFBCC48DA" box="[1023,1067,1897,1923]" pageId="5" pageNumber="550" type="female">♀♀</specimenCount>
.
</paragraph>
<paragraph id="8BBA36A03A0BFFEAFF02B9ABFC004FA0" blockId="5.[151,1437,1249,2032]" lastBlockId="6.[151,1437,151,249]" lastPageId="6" lastPageNumber="551" pageId="5" pageNumber="550">
<emphasis id="B971EAB23A0BFFE9FF02B9ABFF1A48FE" bold="true" box="[199,253,1933,1959]" italics="true" pageId="5" pageNumber="550">Legs</emphasis>
: Moderately long, not retractile, similar to other species of the 
<taxonomicName id="4C054D233A0BFFE9FC28B9ABFB9848FE" authorityName="sensu Jeannel" authorityYear="1910" box="[1005,1151,1933,1959]" class="Insecta" family="Leiodidae" genus="Apholeuonus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="550" phylum="Arthropoda" rank="genus">
<emphasis id="B971EAB23A0BFFE9FC28B9ABFB9848FE" box="[1005,1151,1933,1959]" italics="true" pageId="5" pageNumber="550">Apholeuonus</emphasis>
</taxonomicName>
phyletic group (
<figureCitation id="133E2A253A0BFFE9FA81B9ABFA7248FE" box="[1348,1429,1933,1959]" captionStart="FIGURE 1" captionStartId="3.[151,250,1841,1865]" captionTargetBox="[235,1356,861,1807]" captionTargetId="figure-266@3.[215,1371,849,1817]" captionTargetPageId="3" captionText="FIGURE 1. Rudogorites simonei sp. nov. male and female habitus." figureDoi="http://doi.org/10.5281/zenodo.5650000" httpUri="https://zenodo.org/record/5650000/files/figure.png" pageId="5" pageNumber="550">Figs. 1</figureCitation>
, 
<figureCitation id="133E2A253A0BFFE9FF52B997FF074895" box="[151,224,1969,1996]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1106,1130]" captionTargetBox="[172,1417,188,1074]" captionTargetId="figure-19@5.[164,1424,181,1082]" captionTargetPageId="5" captionText="FIGURES 11–16. Figures 11–14, left first leg, comb of protibial spines is indicated with black arrow. 11, Rudogorites simonei sp. nov., female. 12, Rudogorites simonei sp. nov., male. 13, Charonites scheibeli, female. 14, Charonites matzenaueri, male. Figure 15, second left leg, Rudogorites simonei sp. nov., male. Figure 16, third leg, left Rudogorites simonei sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.5650004" httpUri="https://zenodo.org/record/5650004/files/figure.png" pageId="5" pageNumber="550">11–16</figureCitation>
). Claws long, regularly curved, slender and sharp in all legs. Tarsal empodium between the claws with one long bifid empodial seta (distinctly split in two at the base) in all legs (
<figureCitation id="133E2A253A0BFFE9FC09B9F3FB4848A9" box="[972,1199,2005,2032]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1106,1130]" captionTargetBox="[172,1417,188,1074]" captionTargetId="figure-19@5.[164,1424,181,1082]" captionTargetPageId="5" captionText="FIGURES 11–16. Figures 11–14, left first leg, comb of protibial spines is indicated with black arrow. 11, Rudogorites simonei sp. nov., female. 12, Rudogorites simonei sp. nov., male. 13, Charonites scheibeli, female. 14, Charonites matzenaueri, male. Figure 15, second left leg, Rudogorites simonei sp. nov., male. Figure 16, third leg, left Rudogorites simonei sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.5650004" httpUri="https://zenodo.org/record/5650004/files/figure.png" pageId="5" pageNumber="550">Figs. 11, 12, 15, 16</figureCitation>
). Male protarsi five-segmented (
<figureCitation id="133E2A253A08FFEAFEDBBEB1FE974FE8" box="[286,368,151,177]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1106,1130]" captionTargetBox="[172,1417,188,1074]" captionTargetId="figure-19@5.[164,1424,181,1082]" captionTargetPageId="5" captionText="FIGURES 11–16. Figures 11–14, left first leg, comb of protibial spines is indicated with black arrow. 11, Rudogorites simonei sp. nov., female. 12, Rudogorites simonei sp. nov., male. 13, Charonites scheibeli, female. 14, Charonites matzenaueri, male. Figure 15, second left leg, Rudogorites simonei sp. nov., male. Figure 16, third leg, left Rudogorites simonei sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.5650004" httpUri="https://zenodo.org/record/5650004/files/figure.png" pageId="6" pageNumber="551">Fig. 12</figureCitation>
) and female protarsi four-segmented (
<figureCitation id="133E2A253A08FFEAFCD8BEB1FC884FE8" box="[797,879,151,177]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1106,1130]" captionTargetBox="[172,1417,188,1074]" captionTargetId="figure-19@5.[164,1424,181,1082]" captionTargetPageId="5" captionText="FIGURES 11–16. Figures 11–14, left first leg, comb of protibial spines is indicated with black arrow. 11, Rudogorites simonei sp. nov., female. 12, Rudogorites simonei sp. nov., male. 13, Charonites scheibeli, female. 14, Charonites matzenaueri, male. Figure 15, second left leg, Rudogorites simonei sp. nov., male. Figure 16, third leg, left Rudogorites simonei sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.5650004" httpUri="https://zenodo.org/record/5650004/files/figure.png" pageId="6" pageNumber="551">Fig. 11</figureCitation>
); PtarL: 
<quantity id="4CFD9B453A08FFEAFC17BEB1FB924FE8" box="[978,1141,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="4.1499999999999995" metricValueMax="4.2" metricValueMin="4.1" pageId="6" pageNumber="551" unit="mm" value="0.415" valueMax="0.42" valueMin="0.41">0.41–0.42 mm</quantity>
in 
<specimenCount id="9D03FD293A08FFEAFB5CBEB1FB244FE8" box="[1177,1219,151,177]" pageId="6" pageNumber="551" type="male">♂♂</specimenCount>
and 
<quantity id="4CFD9B453A08FFEAFB3CBEB1FA7A4FE8" box="[1273,1437,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="3.9" metricValueMin="3.5999999999999996" pageId="6" pageNumber="551" unit="mm" value="0.375" valueMax="0.39" valueMin="0.36">0.36–0.39 mm</quantity>
in 
<specimenCount id="9D03FD293A08FFEAFF71BE9DFF384F8C" box="[180,223,187,213]" pageId="6" pageNumber="551" type="female">♀♀</specimenCount>
; PtarW: 
<quantity id="4CFD9B453A08FFEAFE86BE9CFE004F8C" box="[323,487,186,213]" metricMagnitude="-5" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="6" pageNumber="551" unit="mm" value="0.045" valueMax="0.05" valueMin="0.04">0.04–0.05 mm</quantity>
in 
<specimenCount id="9D03FD293A08FFEAFDCEBE9DFDD24F8C" box="[523,565,187,213]" pageId="6" pageNumber="551" type="male">♂♂</specimenCount>
and 
<quantity id="4CFD9B453A08FFEAFDA9BE9DFD374F8C" box="[620,720,187,213]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="551" unit="mm" value="0.04">0.04 mm</quantity>
in 
<specimenCount id="9D03FD293A08FFEAFD31BE9DFCF84F8C" box="[756,799,187,213]" pageId="6" pageNumber="551" type="female">♀♀</specimenCount>
, not distinctly dilated in both sexes. Male protarsomeres equipped with dense, terminally hook-like adhesive (tenant) setae (
<figureCitation id="133E2A253A08FFEAFC43BEF9FC3D4FA0" box="[902,986,223,249]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1106,1130]" captionTargetBox="[172,1417,188,1074]" captionTargetId="figure-19@5.[164,1424,181,1082]" captionTargetPageId="5" captionText="FIGURES 11–16. Figures 11–14, left first leg, comb of protibial spines is indicated with black arrow. 11, Rudogorites simonei sp. nov., female. 12, Rudogorites simonei sp. nov., male. 13, Charonites scheibeli, female. 14, Charonites matzenaueri, male. Figure 15, second left leg, Rudogorites simonei sp. nov., male. Figure 16, third leg, left Rudogorites simonei sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.5650004" httpUri="https://zenodo.org/record/5650004/files/figure.png" pageId="6" pageNumber="551">Fig. 12</figureCitation>
).
</paragraph>
<caption id="DF7A66283A08FFEAFF52BACFFC944A37" ID-DOI="http://doi.org/10.5281/zenodo.5650006" ID-Zenodo-Dep="5650006" httpUri="https://zenodo.org/record/5650006/files/figure.png" pageId="6" pageNumber="551" startId="6.[151,264,1257,1281]" targetBox="[173,1414,274,1225]" targetPageId="6">
<paragraph id="8BBA36A03A08FFEAFF52BACFFC944A37" blockId="6.[151,1436,1257,1390]" pageId="6" pageNumber="551">
<emphasis id="B971EAB23A08FFEAFF52BACFFEB34A5B" bold="true" box="[151,340,1257,1282]" pageId="6" pageNumber="551">FIGURES 17–24.</emphasis>
Figures 17–20. Antenna. 17, 
<taxonomicName id="4C054D233A08FFEAFD44BACCFCB54A58" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[641,850,1258,1282]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A08FFEAFD44BACCFCB54A58" box="[641,850,1258,1282]" italics="true" pageId="6" pageNumber="551">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A08FFEAFC9CBACCFC4E4A5B" bold="true" box="[857,937,1258,1282]" pageId="6" pageNumber="551">
<taxonomicNameLabel id="A24257C93A08FFEAFC9CBACCFC4E4A5B" box="[857,937,1258,1282]" pageId="6" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. 18, 
<taxonomicName id="4C054D233A08FFEAFBDEBACCFB0A4A58" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[1051,1261,1258,1282]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A08FFEAFBDEBACCFB0A4A58" box="[1051,1261,1258,1282]" italics="true" pageId="6" pageNumber="551">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A08FFEAFB36BACCFAA34A5B" bold="true" box="[1267,1348,1258,1282]" pageId="6" pageNumber="551">
<taxonomicNameLabel id="A24257C93A08FFEAFB36BACCFAA34A5B" box="[1267,1348,1258,1282]" pageId="6" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. 19, 
<taxonomicName id="4C054D233A08FFEAFF7BBB28FE684A7F" box="[190,399,1294,1318]" class="Insecta" family="Leiodidae" genus="Charonites" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="scheibeli">
<emphasis id="B971EAB23A08FFEAFF7BBB28FE684A7F" box="[190,399,1294,1318]" italics="true" pageId="6" pageNumber="551">Charonites scheibeli</emphasis>
</taxonomicName>
, female. 20, 
<taxonomicName id="4C054D233A08FFEAFDD3BB28FCED4A7C" box="[534,778,1294,1318]" class="Insecta" family="Leiodidae" genus="Charonites" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="matzenaueri">
<emphasis id="B971EAB23A08FFEAFDD3BB28FCED4A7C" box="[534,778,1294,1318]" italics="true" pageId="6" pageNumber="551">Charonites matzenaueri</emphasis>
</taxonomicName>
, male. Figures 21–23, metatergal apparatus, posterior process is indicated with black arrow. 21, 
<taxonomicName id="4C054D233A08FFEAFE16BB14FD444A10" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[467,675,1330,1354]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A08FFEAFE16BB14FD444A10" box="[467,675,1330,1354]" italics="true" pageId="6" pageNumber="551">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A08FFEAFD6DBB14FD1F4A13" bold="true" box="[680,760,1330,1354]" pageId="6" pageNumber="551">
<taxonomicNameLabel id="A24257C93A08FFEAFD6DBB14FD1F4A13" box="[680,760,1330,1354]" pageId="6" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. 22, 
<taxonomicName id="4C054D233A08FFEAFCA3BB14FBBE4A10" box="[870,1113,1330,1354]" class="Insecta" family="Leiodidae" genus="Charonites" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="matzenaueri">
<emphasis id="B971EAB23A08FFEAFCA3BB14FBBE4A10" box="[870,1113,1330,1354]" italics="true" pageId="6" pageNumber="551">Charonites matzenaueri</emphasis>
</taxonomicName>
, male. 23, 
<taxonomicName id="4C054D233A08FFEAFB02BB14FA714A13" box="[1223,1430,1330,1354]" class="Insecta" family="Leiodidae" genus="Charonites" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="scheibeli">
<emphasis id="B971EAB23A08FFEAFB02BB14FA714A13" box="[1223,1430,1330,1354]" italics="true" pageId="6" pageNumber="551">Charonites scheibeli</emphasis>
</taxonomicName>
, female. Figure 24, metendosternite, 
<taxonomicName id="4C054D233A08FFEAFDC3BB70FD304A34" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[518,727,1366,1390]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A08FFEAFDC3BB70FD304A34" box="[518,727,1366,1390]" italics="true" pageId="6" pageNumber="551">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A08FFEAFD18BB70FCC94A37" bold="true" box="[733,814,1366,1390]" pageId="6" pageNumber="551">
<taxonomicNameLabel id="A24257C93A08FFEAFD18BB70FCC94A37" box="[733,814,1366,1390]" pageId="6" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
</paragraph>
</caption>
<paragraph id="8BBA36A03A08FFEAFF02BBBAFC644933" blockId="6.[151,1437,1436,2038]" pageId="6" pageNumber="551">
Protibial internal and ventral apical spur short and tridentate or polydentate. Protibia armed with ring of apical flattened spines that continues to the externo-lateral row of strong spines forming a comb. These comb spines are flattened, short and wide, almost triangular and stretching in straight continuous line on entire two thirds of protibia length. Protibia characteristically curved and the widest at the middle (
<figureCitation id="133E2A253A08FFEAFC72B82EFBD8497B" box="[951,1087,1544,1570]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1106,1130]" captionTargetBox="[172,1417,188,1074]" captionTargetId="figure-19@5.[164,1424,181,1082]" captionTargetPageId="5" captionText="FIGURES 11–16. Figures 11–14, left first leg, comb of protibial spines is indicated with black arrow. 11, Rudogorites simonei sp. nov., female. 12, Rudogorites simonei sp. nov., male. 13, Charonites scheibeli, female. 14, Charonites matzenaueri, male. Figure 15, second left leg, Rudogorites simonei sp. nov., male. Figure 16, third leg, left Rudogorites simonei sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.5650004" httpUri="https://zenodo.org/record/5650004/files/figure.png" pageId="6" pageNumber="551">Figs. 11, 12</figureCitation>
). PtibL: 
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in 
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, 
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<specimenCount id="9D03FD293A08FFEAFEEAB80AFEBD491F" box="[303,346,1580,1606]" pageId="6" pageNumber="551" type="female">♀♀</specimenCount>
; PtibW: 
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mm
</quantity>
in 
<specimenCount id="9D03FD293A08FFEAFD6DB80AFD33491F" box="[680,724,1580,1606]" pageId="6" pageNumber="551" type="male">♂♂</specimenCount>
, 
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mm
</quantity>
in 
<specimenCount id="9D03FD293A08FFEAFC0FB80AFC12491F" box="[970,1013,1580,1606]" pageId="6" pageNumber="551" type="female">♀♀</specimenCount>
, widest at the middle of tibia length, 
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mm
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in 
<specimenCount id="9D03FD293A08FFEAFEBFB876FE414933" box="[378,422,1616,1642]" pageId="6" pageNumber="551" type="male">♂♂</specimenCount>
, 
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mm
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in 
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wide at the apex.
</paragraph>
<paragraph id="8BBA36A03A08FFEAFF02B852FB8C49A2" blockId="6.[151,1437,1436,2038]" pageId="6" pageNumber="551">
Mesotarsi and metatarsi five-segmented in both sexes (
<figureCitation id="133E2A253A08FFEAFC8FB852FC3D49D6" box="[842,986,1652,1679]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1106,1130]" captionTargetBox="[172,1417,188,1074]" captionTargetId="figure-19@5.[164,1424,181,1082]" captionTargetPageId="5" captionText="FIGURES 11–16. Figures 11–14, left first leg, comb of protibial spines is indicated with black arrow. 11, Rudogorites simonei sp. nov., female. 12, Rudogorites simonei sp. nov., male. 13, Charonites scheibeli, female. 14, Charonites matzenaueri, male. Figure 15, second left leg, Rudogorites simonei sp. nov., male. Figure 16, third leg, left Rudogorites simonei sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.5650004" httpUri="https://zenodo.org/record/5650004/files/figure.png" pageId="6" pageNumber="551">Figs. 15, 16</figureCitation>
). Mesotibia and metatibia internal and ventral apical spur prolonged and multi-toothed. Mesotibia and metatibia ring of apical spines prolonged and of approximately equal length, forming a basket. Externo-lateral edge of mesotibia and metatibia armed with weak spines scattered without order. All femora distinctly expanded at apex (
<figureCitation id="133E2A253A08FFEAFCA3B8C6FC5349A3" box="[870,948,1760,1786]" captionStart="FIGURE 1" captionStartId="3.[151,250,1841,1865]" captionTargetBox="[235,1356,861,1807]" captionTargetId="figure-266@3.[215,1371,849,1817]" captionTargetPageId="3" captionText="FIGURE 1. Rudogorites simonei sp. nov. male and female habitus." figureDoi="http://doi.org/10.5281/zenodo.5650000" httpUri="https://zenodo.org/record/5650000/files/figure.png" pageId="6" pageNumber="551">Figs. 1</figureCitation>
, 
<figureCitation id="133E2A253A08FFEAFC07B8C6FBBA49A2" box="[962,1117,1760,1787]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1106,1130]" captionTargetBox="[172,1417,188,1074]" captionTargetId="figure-19@5.[164,1424,181,1082]" captionTargetPageId="5" captionText="FIGURES 11–16. Figures 11–14, left first leg, comb of protibial spines is indicated with black arrow. 11, Rudogorites simonei sp. nov., female. 12, Rudogorites simonei sp. nov., male. 13, Charonites scheibeli, female. 14, Charonites matzenaueri, male. Figure 15, second left leg, Rudogorites simonei sp. nov., male. Figure 16, third leg, left Rudogorites simonei sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.5650004" httpUri="https://zenodo.org/record/5650004/files/figure.png" pageId="6" pageNumber="551">11, 12, 15, 16</figureCitation>
).
</paragraph>
<paragraph id="8BBA36A03A08FFEBFF02B925FE164E44" blockId="6.[151,1437,1436,2038]" lastBlockId="7.[151,1436,150,501]" lastPageId="7" lastPageNumber="552" pageId="6" pageNumber="551">
<emphasis id="B971EAB23A08FFEAFF02B925FE894847" bold="true" box="[199,366,1795,1822]" italics="true" pageId="6" pageNumber="551">Male genitalia</emphasis>
: Aedeagus 
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long, broad and truncate with median lobe more or less equally wide all of its length in dorsal view (
<figureCitation id="133E2A253A08FFEAFDC5B90EFDB1481A" box="[512,598,1832,1859]" captionStart="FIGURES 25–30" captionStartId="7.[151,264,1301,1325]" captionTargetBox="[159,1428,532,1276]" captionTargetId="figure-168@7.[153,1435,525,1278]" captionTargetPageId="7" captionText="FIGURES 25–30. Rudogorites simonei sp. nov., male and female genital structures. 25, aedeagus dorsal view. 26, aedeagus lateral view, paramere apex detail is separately slightly enlarged. 27, male genital segment (IX sternite). 28, female ventrite VIIIth. 29, female genital segment, urite IXth with gonostyles. 30, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5650008" httpUri="https://zenodo.org/record/5650008/files/figure.png" pageId="6" pageNumber="551">Fig. 25</figureCitation>
). In lateral view (
<figureCitation id="133E2A253A08FFEAFCE4B90EFC93481A" box="[801,884,1832,1859]" captionStart="FIGURES 25–30" captionStartId="7.[151,264,1301,1325]" captionTargetBox="[159,1428,532,1276]" captionTargetId="figure-168@7.[153,1435,525,1278]" captionTargetPageId="7" captionText="FIGURES 25–30. Rudogorites simonei sp. nov., male and female genital structures. 25, aedeagus dorsal view. 26, aedeagus lateral view, paramere apex detail is separately slightly enlarged. 27, male genital segment (IX sternite). 28, female ventrite VIIIth. 29, female genital segment, urite IXth with gonostyles. 30, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5650008" httpUri="https://zenodo.org/record/5650008/files/figure.png" pageId="6" pageNumber="551">Fig. 26</figureCitation>
) median lobe tubular and strongly curved in the middle and with sharply deflexed apex. Internal sac of aedeagus without strong sclerotized structures only on its medio apical side; some weakly sclerotized or partly hyaline structures visible. No basal “Y” sclerotized structure present. Basal lamina of median lobe wide and basally invaginated backwards. Ventral lamina of tegmen parallel with median lobe and extended along basal lamina. Parameres (lateral styles) straight, mostly evenly wide all its length, parallel with median lobe, shorter than median lobe, with two apical long setae and one subapical (interno-lateral) short seta (
<figureCitation id="133E2A253A09FFEBFEACBEB1FE144FEB" box="[361,499,150,178]" captionStart="FIGURES 25–30" captionStartId="7.[151,264,1301,1325]" captionTargetBox="[159,1428,532,1276]" captionTargetId="figure-168@7.[153,1435,525,1278]" captionTargetPageId="7" captionText="FIGURES 25–30. Rudogorites simonei sp. nov., male and female genital structures. 25, aedeagus dorsal view. 26, aedeagus lateral view, paramere apex detail is separately slightly enlarged. 27, male genital segment (IX sternite). 28, female ventrite VIIIth. 29, female genital segment, urite IXth with gonostyles. 30, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5650008" httpUri="https://zenodo.org/record/5650008/files/figure.png" pageId="7" pageNumber="552">Figs. 25, 26</figureCitation>
). Paramere apex with small terminal hyaline membranous appendage. Male genital segment (IX sternite, urosternite) in lateral view reduced to a chitinous ring. In dorsal view (
<figureCitation id="133E2A253A09FFEBFB4BBE9DFB394F8C" box="[1166,1246,187,213]" captionStart="FIGURES 25–30" captionStartId="7.[151,264,1301,1325]" captionTargetBox="[159,1428,532,1276]" captionTargetId="figure-168@7.[153,1435,525,1278]" captionTargetPageId="7" captionText="FIGURES 25–30. Rudogorites simonei sp. nov., male and female genital structures. 25, aedeagus dorsal view. 26, aedeagus lateral view, paramere apex detail is separately slightly enlarged. 27, male genital segment (IX sternite). 28, female ventrite VIIIth. 29, female genital segment, urite IXth with gonostyles. 30, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5650008" httpUri="https://zenodo.org/record/5650008/files/figure.png" pageId="7" pageNumber="552">Fig. 27</figureCitation>
) with prolonged, rounded ventral lobe and small prolonged process on posterior side of its sternal part and with prolonged narrow lateral stylets on its tergal part.
</paragraph>
<paragraph id="8BBA36A03A09FFEBFF02BF00FE484EAC" blockId="7.[151,1436,150,501]" pageId="7" pageNumber="552">
<emphasis id="B971EAB23A09FFEBFF02BF00FE6C4E18" bold="true" box="[199,395,294,321]" italics="true" pageId="7" pageNumber="552">Female genitalia</emphasis>
: Female ventrite 
<superScript id="7C709BE83A09FFEBFD94BF01FD694E6D" attach="left" box="[593,654,294,321]" fontSize="6" pageId="7" pageNumber="552">VIIIth</superScript>
(
<figureCitation id="133E2A253A09FFEBFD5BBF01FD174E18" box="[670,752,295,321]" captionStart="FIGURES 25–30" captionStartId="7.[151,264,1301,1325]" captionTargetBox="[159,1428,532,1276]" captionTargetId="figure-168@7.[153,1435,525,1278]" captionTargetPageId="7" captionText="FIGURES 25–30. Rudogorites simonei sp. nov., male and female genital structures. 25, aedeagus dorsal view. 26, aedeagus lateral view, paramere apex detail is separately slightly enlarged. 27, male genital segment (IX sternite). 28, female ventrite VIIIth. 29, female genital segment, urite IXth with gonostyles. 30, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5650008" httpUri="https://zenodo.org/record/5650008/files/figure.png" pageId="7" pageNumber="552">Fig. 28</figureCitation>
) strongly pubescent on proximal part with anterior stout and prolonged apophysis (spiculum ventrale). Genital segment (urite 
<superScript id="7C709BE83A09FFEBFC44BF6DFC4C4E01" attach="left" box="[897,939,330,357]" fontSize="6" pageId="7" pageNumber="552">IXth</superScript>
) well sclerotized and developed (
<figureCitation id="133E2A253A09FFEBFAFCBF6DFA774E3C" box="[1337,1424,331,357]" captionStart="FIGURES 25–30" captionStartId="7.[151,264,1301,1325]" captionTargetBox="[159,1428,532,1276]" captionTargetId="figure-168@7.[153,1435,525,1278]" captionTargetPageId="7" captionText="FIGURES 25–30. Rudogorites simonei sp. nov., male and female genital structures. 25, aedeagus dorsal view. 26, aedeagus lateral view, paramere apex detail is separately slightly enlarged. 27, male genital segment (IX sternite). 28, female ventrite VIIIth. 29, female genital segment, urite IXth with gonostyles. 30, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5650008" httpUri="https://zenodo.org/record/5650008/files/figure.png" pageId="7" pageNumber="552">Fig. 29</figureCitation>
). Sub-gonocoxite with one stylus, gonocoxite with one basal, one medial and two subapical styles, apical gonostylus with one the longest stylus. Spermatheca (
<figureCitation id="133E2A253A09FFEBFDB0BFB5FD204EF4" box="[629,711,403,429]" captionStart="FIGURES 25–30" captionStartId="7.[151,264,1301,1325]" captionTargetBox="[159,1428,532,1276]" captionTargetId="figure-168@7.[153,1435,525,1278]" captionTargetPageId="7" captionText="FIGURES 25–30. Rudogorites simonei sp. nov., male and female genital structures. 25, aedeagus dorsal view. 26, aedeagus lateral view, paramere apex detail is separately slightly enlarged. 27, male genital segment (IX sternite). 28, female ventrite VIIIth. 29, female genital segment, urite IXth with gonostyles. 30, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5650008" httpUri="https://zenodo.org/record/5650008/files/figure.png" pageId="7" pageNumber="552">Fig. 30</figureCitation>
) 
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long and 
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wide, sausage-like or short subbilobed, slightly more sclerotize on its both lobes. Hyaline gland attached at the proximal part of spermatheca in small sclerotized nodule.
</paragraph>
</subSubSection>
<caption id="DF7A66283A09FFEBFF52BB33FC554A2F" ID-DOI="http://doi.org/10.5281/zenodo.5650008" ID-Zenodo-Dep="5650008" httpUri="https://zenodo.org/record/5650008/files/figure.png" pageId="7" pageNumber="552" startId="7.[151,264,1301,1325]" targetBox="[159,1428,532,1276]" targetPageId="7">
<paragraph id="8BBA36A03A09FFEBFF52BB33FC554A2F" blockId="7.[151,1436,1301,1398]" pageId="7" pageNumber="552">
<emphasis id="B971EAB23A09FFEBFF52BB33FEB14A77" bold="true" box="[151,342,1301,1326]" pageId="7" pageNumber="552">FIGURES 25–30.</emphasis>
<taxonomicName id="4C054D233A09FFEBFE98BB30FDD74A74" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[349,560,1302,1326]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A09FFEBFE98BB30FDD74A74" box="[349,560,1302,1326]" italics="true" pageId="7" pageNumber="552">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A09FFEBFDFDBB30FD6D4A77" bold="true" box="[568,650,1302,1326]" pageId="7" pageNumber="552">
<taxonomicNameLabel id="A24257C93A09FFEBFDFDBB30FD6D4A77" box="[568,650,1302,1326]" pageId="7" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male and female genital structures. 25, aedeagus dorsal view. 26, aedeagus lateral view, paramere apex detail is separately slightly enlarged. 27, male genital segment (IX sternite). 28, female ventrite 
<superScript id="7C709BE83A09FFEBFF52BB78FF294A35" attach="left" box="[151,206,1374,1397]" fontSize="6" pageId="7" pageNumber="552">VIIIth</superScript>
. 29, female genital segment, urite 
<superScript id="7C709BE83A09FFEBFDE8BB78FDB44A35" attach="left" box="[557,595,1374,1397]" fontSize="6" pageId="7" pageNumber="552">IXth</superScript>
with gonostyles. 30, spermatheca.
</paragraph>
</caption>
<subSubSection id="C31F652B3A09FFEBFF02BB82FC2C4ABB" pageId="7" pageNumber="552" type="diagnosis">
<paragraph id="8BBA36A03A09FFEBFF02BB82FC2C4ABB" blockId="7.[151,1437,1444,2040]" pageId="7" pageNumber="552">
<emphasis id="B971EAB23A09FFEBFF02BB82FE284AE7" bold="true" box="[199,463,1444,1470]" pageId="7" pageNumber="552">Differential diagnosis.</emphasis>
So far, only 
<taxonomicName id="4C054D233A09FFEBFDA7BB82FCAD4AE7" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[610,842,1444,1470]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="simonei">
<emphasis id="B971EAB23A09FFEBFDA7BB82FCAD4AE7" box="[610,842,1444,1470]" italics="true" pageId="7" pageNumber="552">Rudogorites simonei</emphasis>
</taxonomicName>
is known from this genus. It can be easily separated from only similar species of genus 
<taxonomicName id="4C054D233A09FFEBFDE4BBE1FD7B4AB8" authorityName="Apfelbeck" authorityYear="1907" box="[545,668,1479,1505]" class="Insecta" family="Leiodidae" genus="Charonites" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="552" phylum="Arthropoda" rank="genus">
<emphasis id="B971EAB23A09FFEBFDE4BBE1FD7B4AB8" box="[545,668,1479,1505]" italics="true" pageId="7" pageNumber="552">Charonites</emphasis>
</taxonomicName>
(see description of genus).
</paragraph>
</subSubSection>
<subSubSection id="C31F652B3A09FFEBFF02BBCDFEC24915" pageId="7" pageNumber="552" type="etymology">
<paragraph id="8BBA36A03A09FFEBFF02BBCDFEC24915" blockId="7.[151,1437,1444,2040]" pageId="7" pageNumber="552">
<emphasis id="B971EAB23A09FFEBFF02BBCDFEA9495C" bold="true" box="[199,334,1515,1541]" pageId="7" pageNumber="552">Etymology.</emphasis>
The species name 
<taxonomicName id="4C054D233A09FFEBFDECBBCAFD67495C" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[553,640,1516,1541]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="simonei">
<emphasis id="B971EAB23A09FFEBFDECBBCAFD67495C" box="[553,640,1516,1541]" italics="true" pageId="7" pageNumber="552">simonei</emphasis>
</taxonomicName>
is dedicated in honour of Simone Milanolo (Sarajevo), a well-known karstologist and speleologist, who first explored the Lijina Pećina cave and collected most of the specimens of this new species.
</paragraph>
</subSubSection>
<subSubSection id="C31F652B3A09FFEBFF02B873FB5E483E" pageId="7" pageNumber="552" type="distribution">
<paragraph id="8BBA36A03A09FFEBFF02B873FB5E483E" blockId="7.[151,1437,1444,2040]" pageId="7" pageNumber="552">
<emphasis id="B971EAB23A09FFEBFF02B873FE864936" bold="true" box="[199,353,1621,1647]" pageId="7" pageNumber="552">Distribution.</emphasis>
The new species is known only from its 
<typeStatus id="54BE88023A09FFEBFCF2B870FC804929" box="[823,871,1622,1648]" pageId="7" pageNumber="552">type</typeStatus>
locality, the cave “Lijina Pećina” near the village Gunjani in area known as Bosansko Rudogorje in Central 
<collectingCountry id="F31276303A09FFEBFCEDB85FFC9E49CA" box="[808,889,1657,1683]" name="Bosnia and Herzegovina" pageId="7" pageNumber="552">Bosnia</collectingCountry>
. The cave is situated in an isolated strip of limestone rock of Devon age, 
<quantity id="4CFD9B453A09FFEBFE71B8BAFE1C49EF" box="[436,507,1692,1718]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="552" unit="km" value="20.0">20 km</quantity>
long and only a few km wide, that stretches in the direction SE-NW from Lepenica river up to Vrbas river spring. This karstic strip of limestone is superficially completely isolated from the rest of the Dinaric karstic massifs on its western side by layers of Triassic sandstones, Permian sandstones and various Silurian non-karstic rocks (
<bibRefCitation id="EF944B513A09FFEBFEACB921FD874879" author="Jovanovic, R. &amp; Mojicevic, M. &amp; Tokic, S. &amp; Rokic, Lj." box="[361,608,1798,1825]" pageId="7" pageNumber="552" refId="ref10233" refString="Jovanovic, R., Mojicevic, M., Tokic, S. &amp; Rokic, Lj. (1962 - 1967) Osnovna geoloska karta. SFRJ, List Sarajevo." type="book" year="1962">
Jovanović 
<emphasis id="B971EAB23A09FFEBFE21B921FDFA4878" box="[484,541,1799,1825]" italics="true" pageId="7" pageNumber="552">et al.</emphasis>
1962
</bibRefCitation>
-1967; 
<bibRefCitation id="EF944B513A09FFEBFD6BB921FC204878" author="Sofilj J. &amp; Zivanovic, M." box="[686,967,1798,1827]" pageId="7" pageNumber="552" refId="ref11265" refString="Sofilj J. &amp; Zivanovic, M. (1965 - 1971) Osnovna geoloska karta. SFRJ, List Prozor." type="book" year="1965">Sofilj &amp; Živanović 1965</bibRefCitation>
-1971). Two caves are known in this area, the Lijina cave that was formed at the contact of Devonian limestone rich in barite and quartz keratophyre ore and the Dusina cave, which was formed at the contact of Devonian limestone and quartz porphyry.
</paragraph>
</subSubSection>
<subSubSection id="C31F652B3A09FFE4FF02B957FD7D481B" lastPageId="8" lastPageNumber="553" pageId="7" pageNumber="552" type="biology_ecology">
<paragraph id="8BBA36A03A09FFEBFF02B957FB0C48A1" blockId="7.[151,1437,1444,2040]" pageId="7" pageNumber="552">
<emphasis id="B971EAB23A09FFEBFF02B957FECB48D2" bold="true" box="[199,300,1905,1931]" pageId="7" pageNumber="552">Ecology.</emphasis>
Specimens were collected from the bottom of the cave, among the rock debris filled with soil, about 
<quantity id="4CFD9B453A09FFEBFF52B9B3FF0748E9" box="[151,224,1941,1968]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="552" unit="m" value="200.0">200 m</quantity>
from its entrance in complete darkness and without superficial meteorological influence. No other precise meteorological data are available. Besides the leptodirine beetles, the sphodrine ground beetle 
<emphasis id="B971EAB23A09FFEBFB78B99CFF2048AE" italics="true" pageId="7" pageNumber="552">Laeomostenus cavicola</emphasis>
and troglomorphic centipede 
<taxonomicName id="4C054D233A09FFEBFDD2B9FBFC9848AE" authority="(Kos 2021)" baseAuthorityName="Kos" baseAuthorityYear="2021" box="[535,895,2013,2039]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="stygius">
<emphasis id="B971EAB23A09FFEBFDD2B9FBFD6648AE" box="[535,641,2013,2039]" italics="true" pageId="7" pageNumber="552">Lithobius</emphasis>
cf. 
<emphasis id="B971EAB23A09FFEBFD69B9F8FD1D48AE" box="[684,762,2014,2039]" italics="true" pageId="7" pageNumber="552">stygius</emphasis>
(
<bibRefCitation id="EF944B513A09FFEBFCCDB9FBFC9048AE" author="Kos, A." box="[776,887,2013,2039]" pageId="7" pageNumber="552" refId="ref10345" refString="Kos, A. (2021) Genetic variability of the Centipede Lithobius stygius species complex in the Northern Dinarides. M. SC. Thesis, University of Ljubljana, Ljubljana, 49 pp." type="book" year="2021">Kos 2021</bibRefCitation>
)
</taxonomicName>
have been collected in the cave.
</paragraph>
<caption id="DF7A66283A06FFE4FF52BB70FBBB4ACA" ID-DOI="http://doi.org/10.5281/zenodo.5650012" ID-Zenodo-Dep="5650012" httpUri="https://zenodo.org/record/5650012/files/figure.png" pageId="8" pageNumber="553" startId="8.[151,250,1366,1390]" targetBox="[151,1435,166,1342]" targetPageId="8">
<paragraph id="8BBA36A03A06FFE4FF52BB70FBBB4ACA" blockId="8.[151,1436,1366,1427]" pageId="8" pageNumber="553">
<emphasis id="B971EAB23A06FFE4FF52BB70FEC64A37" bold="true" box="[151,289,1366,1390]" pageId="8" pageNumber="553">FIGURE 31.</emphasis>
Map of taxa distributions. White star 
<taxonomicName id="4C054D233A06FFE4FD69BB70FC984A37" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[684,895,1366,1390]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A06FFE4FD69BB70FC984A37" box="[684,895,1366,1390]" italics="true" pageId="8" pageNumber="553">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A06FFE4FC43BB70FC3F4A37" bold="true" box="[902,984,1366,1390]" pageId="8" pageNumber="553">
<taxonomicNameLabel id="A24257C93A06FFE4FC43BB70FC3F4A37" box="[902,984,1366,1390]" pageId="8" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, black star 
<taxonomicName id="4C054D233A06FFE4FB97BB70FA864A37" box="[1106,1377,1366,1390]" class="Insecta" family="Leiodidae" genus="Protobracharhron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="species" species="dusinae">
<emphasis id="B971EAB23A06FFE4FB97BB70FA864A37" box="[1106,1377,1366,1390]" italics="true" pageId="8" pageNumber="553">Protobracharhron dusinae</emphasis>
</taxonomicName>
, dark grey area 
<taxonomicName id="4C054D233A06FFE4FF3EBB5CFE7A4ACB" box="[251,413,1402,1426]" class="Insecta" family="Leiodidae" genus="Charonites" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B971EAB23A06FFE4FF3EBB5CFE8C4ACB" box="[251,363,1402,1426]" italics="true" pageId="8" pageNumber="553">Charonites</emphasis>
spp.
</taxonomicName>
, pale grey area distribution of genera of 
<taxonomicName id="4C054D233A06FFE4FCF2BB5CFC5C4ACB" authorityName="sensu Jeannel" authorityYear="1910" box="[823,955,1402,1426]" class="Insecta" family="Leiodidae" genus="Apholeuonus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="genus">
<emphasis id="B971EAB23A06FFE4FCF2BB5CFC5C4ACB" box="[823,955,1402,1426]" italics="true" pageId="8" pageNumber="553">Apholeuonus</emphasis>
</taxonomicName>
phyletic group.
</paragraph>
</caption>
<paragraph id="8BBA36A03A06FFE4FF02BBE6FE3949D7" blockId="8.[151,1437,1472,2039]" pageId="8" pageNumber="553">
The cave consists of one meander developing at three different levels. Two levels are dry, while the lower one is crossed by a small stream. The spring is located just a few tens of meters below the cave entrance. The cave was visited several times by cavers from the 
<collectingRegion id="49C1F8423A06FFE4FD9BB82FFD4E497A" box="[606,681,1545,1571]" country="France" name="Centre" pageId="8" pageNumber="553">Centre</collectingRegion>
for Karst and Speleology from Sarajevo during spring of 2016 and 2017. Although the explorations are not yet completed, about 
<quantity id="4CFD9B453A06FFE4FC91B80BFC7C491E" box="[852,923,1581,1607]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="553" unit="m" value="800.0">800 m</quantity>
of the cave have been surveyed so far. Many channels have already been partially visited, however are still to be mapped, so the total length of the cave can be expected to be close to 
<quantity id="4CFD9B453A06FFE4FE58B853FE3E49D7" box="[413,473,1652,1679]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="553" unit="km" value="1.0">1 km</quantity>
.
</paragraph>
<paragraph id="8BBA36A03A06FFE4FF02B8BEFD7D481B" blockId="8.[151,1437,1472,2039]" pageId="8" pageNumber="553">The paint signs on the cave wall in the initial part of the cave testifies that the cave was already partially explored, although there is no evidence of any written record in the literature. The cave was also used by the prehistoric inhabitants in the Iron Age, since the remains of pottery and traces of fire were found deep inside the cave. In the Ottoman period the cave served as a place of solitude, probably belonging to a dervish order, as shown by the remains of a wall and carved holes for beams.</paragraph>
</subSubSection>
<subSubSection id="C31F652B3A06FFE5FF02B96BFC484EF4" lastPageId="9" lastPageNumber="554" pageId="8" pageNumber="553" type="materials_examined">
<paragraph id="8BBA36A03A06FFE4FF02B96BFE6348AE" blockId="8.[151,1437,1472,2039]" pageId="8" pageNumber="553">
<emphasis id="B971EAB23A06FFE4FF02B96BFEB2483F" bold="true" box="[199,341,1868,1895]" pageId="8" pageNumber="553">Type series.</emphasis>
<materialsCitation id="3B6D3CFD3A06FFE4FE9AB96AFB7848D3" ID-GBIF-Occurrence="3400807308" collectionCode="DMHF" pageId="8" pageNumber="553" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54BE88023A06FFE4FE9AB96AFE20483F" box="[351,455,1868,1894]" pageId="8" pageNumber="553" type="holotype">Holotype</typeStatus>
<specimenCount id="9D03FD293A06FFE4FE14B96BFE0F483E" box="[465,488,1869,1895]" pageId="8" pageNumber="553" type="male">♂</specimenCount>
, glued to a white card, pinned dry, aedeagus dissected and preserved immersed in 
<collectionCode id="ED14AE653A06FFE4FF52B957FF0F48D2" box="[151,232,1905,1931]" pageId="8" pageNumber="553">DMHF</collectionCode>
on a smaller transparent board pinned under the card-mounted specimens, labeled
</materialsCitation>
: 
<materialsCitation id="3B6D3CFD3A06FFE4FB69B957FE9848AE" ID-GBIF-Occurrence="3400807303" collectingDate="2016-05-01" collectorName="Milanolo, S. &amp; Prelic, M." country="Bosnia and Herzegovina" determinerName="Polak &amp; Mulaomerovic" location="Bosnia and Herzegovina" pageId="8" pageNumber="553" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54BE88023A06FFE4FB69B957FAA748D2" box="[1196,1344,1905,1931]" pageId="8" pageNumber="553" type="holotype">HOLOTYPE</typeStatus>
| 
<taxonomicName id="4C054D233A06FFE4FA9DB956FEA448F7" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A06FFE4FA9DB956FEA448F7" italics="true" pageId="8" pageNumber="553">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A06FFE4FE8FB9B3FE4348F6" bold="true" box="[330,420,1941,1967]" pageId="8" pageNumber="553">
<taxonomicNameLabel id="A24257C93A06FFE4FE8FB9B3FE4348F6" box="[330,420,1941,1967]" pageId="8" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<specimenCount id="9D03FD293A06FFE4FE6EB9B3FE2748F6" box="[427,448,1941,1967]" pageId="8" pageNumber="553" type="male">♂</specimenCount>
| 
<determinerName id="7E66DA643A06FFE4FE11B9B2FDF348F7" box="[468,532,1940,1966]" pageId="8" pageNumber="553">Polak</determinerName>
&amp; 
<determinerName id="7E66DA643A06FFE4FDFDB9B2FD3A48F7" box="[568,733,1940,1966]" pageId="8" pageNumber="553">Mulaomerović</determinerName>
det., [rectangular red label, printed]. Second label: 
<collectingCountry id="F31276303A06FFE4FADAB9B2FECC488B" name="Bosnia and Herzegovina" pageId="8" pageNumber="553">Bosnia and Herzegovina</collectingCountry>
: Lijina Pećina, Gunjani, Kreševo; 
<date id="FFBB10603A06FFE4FD70B99EFCFE488A" box="[693,793,1976,2003]" pageId="8" pageNumber="553" value="2016-05-01">
<collectingDate id="EFFFE9883A06FFE4FD70B99EFCFE488A" box="[693,793,1976,2003]" pageId="8" pageNumber="553" value="2016-05-01">1.5.2016</collectingDate>
</date>
, 
<collectorName id="26F053763A06FFE4FCE2B99EFC53488A" box="[807,948,1976,2003]" pageId="8" pageNumber="553">Milanolo, S.</collectorName>
&amp; 
<collectorName id="26F053763A06FFE4FC19B99EFBAA488A" box="[988,1101,1976,2003]" pageId="8" pageNumber="553">Prelić, M.</collectorName>
leg. [rectangular white label, printed]—(
<emphasis id="B971EAB23A06FFE4FED0B9FAFE9348AF" bold="true" box="[277,372,2012,2038]" pageId="8" pageNumber="553">NMBiH</emphasis>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BBA36A03A07FFE5FF02BEB1FC654FE8" blockId="9.[151,1437,151,429]" box="[199,898,151,177]" pageId="9" pageNumber="554">
<typeStatus id="54BE88023A07FFE5FF02BEB1FEDF4FE8" box="[199,312,151,177]" pageId="9" pageNumber="554" type="paratype">Paratypes</typeStatus>
: specimens glued to white card, pinned dry, labeled:
</paragraph>
<paragraph id="8BBA36A03A07FFE5FF02BE9DFC484EF4" blockId="9.[151,1437,151,429]" pageId="9" pageNumber="554">
<typeStatus id="54BE88023A07FFE5FF02BE9DFEB54F8C" box="[199,338,187,213]" pageId="9" pageNumber="554" type="paratype">PARATYPE</typeStatus>
| 
<taxonomicName id="4C054D233A07FFE5FEA7BE9DFDAA4F8D" authority="Polak &amp; Mulaomerović, 2021" authorityName="Polak &amp; Mulaomerović" authorityYear="2021" box="[354,589,187,213]" class="Insecta" family="Leiodidae" genus="Rudogorites" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="simonei" status="sp. nov.">
<emphasis id="B971EAB23A07FFE5FEA7BE9DFDAA4F8D" box="[354,589,187,213]" italics="true" pageId="9" pageNumber="554">Rudogorites simonei</emphasis>
</taxonomicName>
<emphasis id="B971EAB23A07FFE5FD92BE9DFD534F8C" bold="true" box="[599,692,187,213]" pageId="9" pageNumber="554">
<taxonomicNameLabel id="A24257C93A07FFE5FD92BE9DFD534F8C" box="[599,692,187,213]" pageId="9" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<determinerName id="7E66DA643A07FFE5FD7ABE9DFD184F8C" box="[703,767,187,213]" pageId="9" pageNumber="554">Polak</determinerName>
&amp; 
<determinerName id="7E66DA643A07FFE5FCECBE9DFC294F8C" box="[809,974,187,213]" pageId="9" pageNumber="554">Mulaomerović</determinerName>
det. [rectangular yellow label, printed]. Second label: [rectangular white label, printed], 
<materialsCitation id="3B6D3CFD3A07FFE5FD70BEF9FDA64E44" ID-GBIF-Occurrence="3400807309" collectingDate="2016-08-16" collectorName="Milanolo, S." pageId="9" pageNumber="554" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="9D03FD293A07FFE5FD70BEF9FD384FA0" box="[693,735,223,249]" pageId="9" pageNumber="554" type="female">1 ♀</specimenCount>
, 
<date id="FFBB10603A07FFE5FD2BBEF9FC874FA0" box="[750,864,223,249]" pageId="9" pageNumber="554" value="2016-08-16">
<collectingDate id="EFFFE9883A07FFE5FD2BBEF9FC874FA0" box="[750,864,223,249]" pageId="9" pageNumber="554" value="2016-08-16">16.8.2016</collectingDate>
</date>
, 
<collectorName id="26F053763A07FFE5FCA9BEF9FC104FA0" box="[876,1015,223,249]" pageId="9" pageNumber="554">Milanolo, S.</collectorName>
leg., same locality than the 
<typeStatus id="54BE88023A07FFE5FAF7BEF9FA7F4FA0" box="[1330,1432,223,249]" pageId="9" pageNumber="554" type="holotype">holotype</typeStatus>
; pinned dry, not dissected—(
<emphasis id="B971EAB23A07FFE5FE12BF24FDD14E45" bold="true" box="[471,566,258,284]" pageId="9" pageNumber="554">NMBiH</emphasis>
)
</materialsCitation>
; 
<materialsCitation id="3B6D3CFD3A07FFE5FD95BF25FDD44E18" ID-GBIF-Occurrence="3400807304" collectingDate="2017-02-05" collectionCode="PMSL" collectorName="Milanolo, S." pageId="9" pageNumber="554" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="9D03FD293A07FFE5FD95BF25FD9B4E44" box="[592,636,259,285]" pageId="9" pageNumber="554" type="male">1 ♂</specimenCount>
and 
<specimenCount id="9D03FD293A07FFE5FD70BF25FD064E44" box="[693,737,259,285]" pageId="9" pageNumber="554" type="female">1 ♀</specimenCount>
, 
<date id="FFBB10603A07FFE5FD35BF24FCB24E44" box="[752,853,258,285]" pageId="9" pageNumber="554" value="2017-02-05">
<collectingDate id="EFFFE9883A07FFE5FD35BF24FCB24E44" box="[752,853,258,285]" pageId="9" pageNumber="554" value="2017-02-05">5.2.2017</collectingDate>
</date>
, 
<collectorName id="26F053763A07FFE5FCA7BF25FC094E44" box="[866,1006,259,285]" pageId="9" pageNumber="554">Milanolo, S.</collectorName>
leg., same locality than the 
<typeStatus id="54BE88023A07FFE5FAF7BF25FA7F4E44" box="[1330,1432,259,285]" pageId="9" pageNumber="554" type="holotype">holotype</typeStatus>
; pinned dry, not dissected—(
<emphasis id="B971EAB23A07FFE5FE1EBF00FDCE4E19" bold="true" box="[475,553,294,320]" pageId="9" pageNumber="554">
<collectionCode id="ED14AE653A07FFE5FE1EBF00FDCE4E19" box="[475,553,294,320]" country="Slovenia" httpUri="http://grbio.org/cool/4mab-159w" name="Slovenian Museum of Natural History (Prirodosloveni Muzej Slovenije)" pageId="9" pageNumber="554">PMSL</collectionCode>
</emphasis>
)
</materialsCitation>
. 
<materialsCitation id="3B6D3CFD3A07FFE5FD87BF01FDDE4E3C" ID-GBIF-Occurrence="3400807301" collectingDate="2017-02-05" collectionCode="NMPO" collectorName="Milanolo, S." pageId="9" pageNumber="554" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="9D03FD293A07FFE5FD87BF01FD884E18" box="[578,623,295,321]" pageId="9" pageNumber="554" type="male">1 ♂</specimenCount>
and 
<specimenCount id="9D03FD293A07FFE5FD6EBF01FD304E18" box="[683,727,295,321]" pageId="9" pageNumber="554" type="female">1 ♀</specimenCount>
, 
<date id="FFBB10603A07FFE5FD2DBF00FCAA4E18" box="[744,845,294,321]" pageId="9" pageNumber="554" value="2017-02-05">
<collectingDate id="EFFFE9883A07FFE5FD2DBF00FCAA4E18" box="[744,845,294,321]" pageId="9" pageNumber="554" value="2017-02-05">5.2.2017</collectingDate>
</date>
, 
<collectorName id="26F053763A07FFE5FC9EBF01FC0F4E18" box="[859,1000,295,321]" pageId="9" pageNumber="554">Milanolo, S.</collectorName>
leg., same locality than the 
<typeStatus id="54BE88023A07FFE5FAF7BF01FA7F4E18" box="[1330,1432,295,321]" pageId="9" pageNumber="554" type="holotype">holotype</typeStatus>
; pinned dry, not dissected—(
<emphasis id="B971EAB23A07FFE5FE1CBF6CFDC84E3D" bold="true" box="[473,559,330,356]" pageId="9" pageNumber="554">
<collectionCode id="ED14AE653A07FFE5FE1CBF6CFDC84E3D" box="[473,559,330,356]" pageId="9" pageNumber="554">NMPO</collectionCode>
</emphasis>
)
</materialsCitation>
. 
<materialsCitation id="3B6D3CFD3A07FFE5FD8DBF6DFC4E4EF4" ID-GBIF-Occurrence="3400807305" collectingDate="2016-08-16" collectionCode="NMPO" collectorName="Milanolo, S." country="Bosnia and Herzegovina" determinerName="Polak &amp; Mulaomerovic" location="Bosnia and Herzegovina" pageId="9" pageNumber="554" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="paratype">
Specimens dissected (body parts, aedeagus, genital segments, protarsi, and antenna) preserved in in Solakryl BMX (Xylene diluting) media on glass microscope slides. 
<specimenCount id="9D03FD293A07FFE5FB16BF49FAF24ED0" box="[1235,1301,367,393]" pageId="9" pageNumber="554" type="male">2 ♂♂</specimenCount>
and 
<specimenCount id="9D03FD293A07FFE5FA96BF49FA714ED0" box="[1363,1430,367,393]" pageId="9" pageNumber="554" type="female">3 ♀♀</specimenCount>
, 
<date id="FFBB10603A07FFE5FF52BFB5FEED4EF4" box="[151,266,403,429]" pageId="9" pageNumber="554" value="2016-08-16">
<collectingDate id="EFFFE9883A07FFE5FF52BFB5FEED4EF4" box="[151,266,403,429]" pageId="9" pageNumber="554" value="2016-08-16">16.8.2016</collectingDate>
</date>
, 
<collectorName id="26F053763A07FFE5FED0BFB5FE474EF4" box="[277,416,403,429]" pageId="9" pageNumber="554">Milanolo, S.</collectorName>
leg., same locality as the holotype—(
<emphasis id="B971EAB23A07FFE5FC8CBFB4FC784EF5" bold="true" box="[841,927,402,428]" pageId="9" pageNumber="554">
<collectionCode id="ED14AE653A07FFE5FC8CBFB4FC784EF5" box="[841,927,402,428]" pageId="9" pageNumber="554">NMPO</collectionCode>
</emphasis>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>