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<document id="B5F8C7CA1DF75C917C88593F6DA8C2AB" ID-CLB-Dataset="112023" ID-DOI="10.11646/zootaxa.5103.1.1" ID-GBIF-Dataset="1bf32ba1-8e0d-4435-ba04-6d7dbd0d7238" ID-ISSN="1175-5326" ID-Zenodo-Dep="6245709" ID-ZooBank="9E1A72E7-3862-44F7-B69F-ECE64B239FF9" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1645601651449" checkinUser="plazi" docAuthor="Kirejtshuk, Alexander G. &amp; Kovalev, Alexey V." docDate="2022" docId="03AC73267628D61875E0FA61FA8DFE11" docLanguage="en" docName="zootaxa.5103.1.1.pdf" docOrigin="Zootaxa 5103 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ithyphenes australiaensis Kirejtshuk &amp; Kovalev 2022, sp. nov." docType="treatment" docUuid="EF694239-FD8F-41E3-A63A-1992B09C292E" docUuidSource="ZooBank" docVersion="7" lastPageNumber="97" masterDocId="FF950B5E7674D6787577FFF3FFF4FFE7" masterDocTitle="Monograph on the Cillaeinae (Coleoptera: Nitidulidae) from the Australian Region with comments on the taxonomy of the subfamily" masterLastPageNumber="133" masterPageNumber="1" pageNumber="93" updateTime="1699299454408" updateUser="plazi">
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<mods:title id="D5CCA69A851838D853C65A46B99744AB">Monograph on the Cillaeinae (Coleoptera: Nitidulidae) from the Australian Region with comments on the taxonomy of the subfamily</mods:title>
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<mods:namePart id="3D2A3903DABB9D58DBC7BB2772ABAD2D">Kirejtshuk, Alexander G.</mods:namePart>
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<mods:namePart id="ECF98C6CDBE206346B8843D6003616FB">Kovalev, Alexey V.</mods:namePart>
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<mods:title id="C0C8F67523DCE46A663E831B1B0A9FFA">Zootaxa</mods:title>
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<mods:date id="41C88BD60DA30DF71171E1EDE645DDF4">2022</mods:date>
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<treatment id="03AC73267628D61875E0FA61FA8DFE11" ID-DOI="http://doi.org/10.5281/zenodo.6315517" ID-GBIF-Taxon="198574683" ID-Zenodo-Dep="6315517" LSID="urn:lsid:zoobank.org:act:EF694239-FD8F-41E3-A63A-1992B09C292E" httpUri="http://treatment.plazi.org/id/03AC73267628D61875E0FA61FA8DFE11" lastPageId="96" lastPageNumber="97" pageId="92" pageNumber="93">
<subSubSection id="C31F91BB7628D62475E0FA61FC8AFA4A" box="[151,894,1426,1453]" pageId="92" pageNumber="93" type="nomenclature">
<paragraph id="8BBAC2307628D62475E0FA61FC8AFA4A" blockId="92.[151,894,1426,1489]" box="[151,894,1426,1453]" pageId="92" pageNumber="93">
<emphasis id="B9711E227628D62475E0FA61FC8AFA4A" bold="true" box="[151,894,1426,1453]" pageId="92" pageNumber="93">
37.
<taxonomicName id="4C05B9B37628D62475B4FA61FD73FA4B" authority="Kirejtshuk" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[195,647,1426,1453]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="australiaensis">
<emphasis id="B9711E227628D62475B4FA61FE0CFA4A" bold="true" box="[195,504,1426,1453]" italics="true" pageId="92" pageNumber="93">Ithyphenes australiaensis</emphasis>
Kirejtshuk
</taxonomicName>
et Kovalev,
<taxonomicNameLabel id="A242A3597628D6247656FA60FC8AFA4A" box="[801,894,1427,1453]" pageId="92" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31F91BB7628D62475E0FA44FC44F9F4" pageId="92" pageNumber="93" type="description">
<paragraph id="8BBAC2307628D62475E0FA44FED7FA36" blockId="92.[151,894,1426,1489]" box="[151,291,1463,1489]" pageId="92" pageNumber="93">
<figureCitation id="133EDEB57628D62475E0FA44FF18FA36" box="[151,236,1463,1489]" captionStart="FIGURE 28" captionStartId="95.[151,250,1898,1922]" captionTargetBox="[233,1354,281,1868]" captionTargetId="figure-65@95.[229,1358,273,1875]" captionTargetPageId="95" captionText="FIGURE 28. Ithyphenes australiensis sp. nov., orig. AD, FH: holotype, male from NSW (“3 km N of Landsowne …”); E, I: paratype from QLD (“… Cape Tribulation…”); A: body, dorsal; B: head, dorsal; C: head, ventral; D: thorax, ventral; E: terminal abdominal segment, ventral; F: male anal sclerite, ventral; G: tegmen, ventral; H: penis trunk, dorsal; I: ovipositor, ventral. Scale bars: a = 1.0 mm to Fig. A; b = 0.5 mm to Figs BE; с = 0.25 mm to Figs FI." figureDoi="http://doi.org/10.5281/zenodo.6245855" httpUri="https://zenodo.org/record/6245855/files/figure.png" pageId="92" pageNumber="93">Figs 28</figureCitation>
, AI
</paragraph>
<paragraph id="8BBAC2307628D62475E0FA08FC44F9F4" blockId="92.[151,944,1531,1555]" box="[151,944,1531,1555]" pageId="92" pageNumber="93">
LSID
<uri id="FF94CE327628D62475A1FA08FC44F9F4" box="[214,944,1531,1555]" pageId="92" pageNumber="93">
urn:lsid:zoobank.org:act:
<uuid id="FFA3F8E57628D62474B9FA08FC44F9F4" box="[462,944,1531,1555]" pageId="92" pageNumber="93">EF694239-FD8F-41E3-A63A-1992B09C292E</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C31F91BB7628D62475E0F9B2FAF9F944" pageId="92" pageNumber="93" type="materials_examined">
<paragraph id="8BBAC2307628D62475E0F9B2FAF9F944" blockId="92.[151,1437,1601,2024]" pageId="92" pageNumber="93">
<emphasis id="B9711E227628D62475E0F9B2FE3FF9BC" bold="true" box="[151,459,1601,1627]" pageId="92" pageNumber="93">Type specimens examined</emphasis>
.
<materialsCitation id="3B6DC86D7628D62474AFF9B2FAF9F944" ID-GBIF-Occurrence="3878844397" collectingDate="1996-05-31" collectingDateMax="1996-06-28" collectingDateMin="1996-05-31" collectionCode="ANIC, ZIN" collectorName="P. Zborowski" country="Australia" county="Williams" elevation="10" latitude="-16.04" location="Emgd" longLatPrecision="769" longitude="145.28" municipality="May" pageId="92" pageNumber="93" specimenCount="1" typeStatus="holotype">
<emphasis id="B9711E227628D62474AFF9B2FD61F9BC" bold="true" box="[472,661,1601,1627]" pageId="92" pageNumber="93">
<collectingCountry id="F31282A07628D62474AFF9B2FDBEF9BC" box="[472,586,1601,1627]" name="Australia" pageId="92" pageNumber="93">Australia</collectingCountry>
, NSW
</emphasis>
:
<typeStatus id="54BE7C927628D62477D4F9B2FCFAF9BC" box="[675,782,1601,1627]" pageId="92" pageNumber="93" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="ED145AF57628D624762AF9B2FC52F9BC" box="[861,934,1601,1627]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="92" pageNumber="93">ANIC</collectionCode>
)—“
<locationDeviation id="9A08DB127628D62476A0F9B2FB3BF9BC" box="[983,1231,1601,1628]" location="Landsowne" pageId="92" pageNumber="93">
<quantity id="4CFD6FD57628D62476A0F9B2FBF9F9BC" box="[983,1037,1601,1627]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="92" pageNumber="93" unit="km" value="3.0">3 km</quantity>
N of Landsowne
</locationDeviation>
, NSW,
<location id="8EDA94EB7628D6247055F9B2FA9CF9BC" LSID="urn:lsid:plazi:treatment:03AC73267628D61875E0FA61FA8DFE11:8EDA94EB7628D6247055F9B2FA9CF9BC" box="[1314,1384,1601,1627]" country="Australia" county="Williams" latitude="-16.04" longLatPrecision="769" longitude="145.28" municipality="May" name="Emgd" pageId="92" pageNumber="93">Emgd</location>
, 15 29
<collectingMunicipality id="6BDE584A7628D62475CEF995FF19F967" box="[185,237,1638,1664]" pageId="92" pageNumber="93">May</collectingMunicipality>
, 1983, C/&amp; N/
<collectingCounty id="62DBBABC7628D62474E1F996FDF5F998" box="[406,513,1637,1663]" pageId="92" pageNumber="93">Williams</collectingCounty>
”, “dead ficus oblique branch”;
<emphasis id="B9711E227628D624762DF996FC63F998" bold="true" box="[858,919,1637,1663]" pageId="92" pageNumber="93">QLD</emphasis>
:
<specimenCount id="9D0309B97628D62476D3F996FBE8F967" box="[932,1052,1637,1664]" pageId="92" pageNumber="93" type="generic" typeStatus="paratype">1 paratype</specimenCount>
, female (
<collectionCode id="ED145AF57628D62471F7F996FB46F967" box="[1152,1202,1637,1664]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="92" pageNumber="93" type="Museum">ZIN</collectionCode>
)—“
<geoCoordinate id="EE31A4F77628D6247194F996FAC5F967" box="[1251,1329,1637,1664]" degrees="16.04" direction="south" orientation="latitude" pageId="92" pageNumber="93" precision="555" value="-16.04">16.04S</geoCoordinate>
<geoCoordinate id="EE31A4F77628D6247040F996FA6CF967" box="[1335,1432,1637,1664]" degrees="145.28" direction="east" orientation="longitude" pageId="92" pageNumber="93" precision="555" value="145.28">145.28E</geoCoordinate>
, QLD, CT1
<quantity id="4CFD6FD57628D624746FF97AFEA7F943" box="[280,339,1673,1700]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="92" pageNumber="93" unit="m" value="10.0">
<elevation id="002825037628D624746FF97AFEA7F943" box="[280,339,1673,1700]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="92" pageNumber="93" unit="m" value="10.0">10 m</elevation>
</quantity>
,
<quantity id="4CFD6FD57628D6247429F97AFE64F944" box="[350,400,1673,1699]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="92" pageNumber="93" unit="km" value="1.0">1km</quantity>
WNW, Cape Tribulation,
<date id="FFBBE4F07628D62477C1F97AFC35F944" box="[694,961,1673,1700]" pageId="92" pageNumber="93" value="1996-05-31" valueMax="1996-06-28" valueMin="1996-05-31">
<collectingDate id="EFFF1D187628D62477C1F97AFC35F944" box="[694,961,1673,1700]" pageId="92" pageNumber="93" value="1996-05-31" valueMax="1996-06-28" valueMin="1996-05-31">31 May—28 June 1996</collectingDate>
</date>
,
<collectorName id="26F0A7E67628D62476BBF979FB96F944" box="[972,1122,1673,1700]" pageId="92" pageNumber="93">P. Zborowski</collectorName>
,
<collectingMethod id="5244BA277628D6247119F979FB35F943" box="[1134,1217,1674,1700]" pageId="92" pageNumber="93">FI Trap</collectingMethod>
JCU.”
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31F91BB7628D62675B0F95EFCE0FE4B" lastPageId="94" lastPageNumber="95" pageId="92" pageNumber="93" type="diagnosis">
<paragraph id="8BBAC2307628D62575B0F95EFB2EFDDF" blockId="92.[151,1437,1601,2024]" lastBlockId="93.[151,1437,151,2029]" lastPageId="93" lastPageNumber="94" pageId="92" pageNumber="93">
<emphasis id="B9711E227628D62475B0F95EFECEF920" bold="true" box="[199,314,1709,1735]" pageId="92" pageNumber="93">Diagnosis</emphasis>
. This new species can be diagnosed from the Australian congeners due to the above key to Australian species of the genus. In addition to the above key,
<taxonomicName id="4C05B9B37628D624779BF922FBF8F90C" authority="Kirejtshuk &amp; Kovalev, 2022" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[748,1036,1745,1771]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="australiaensis" status="sp. nov.">
<emphasis id="B9711E227628D624779BF922FBF8F90C" box="[748,1036,1745,1771]" italics="true" pageId="92" pageNumber="93">Ithyphenes australiaensis</emphasis>
</taxonomicName>
<emphasis id="B9711E227628D6247161F922FB87F90C" bold="true" box="[1046,1139,1745,1771]" pageId="92" pageNumber="93">
<taxonomicNameLabel id="A242A3597628D6247161F922FB87F90C" box="[1046,1139,1745,1771]" pageId="92" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C05B9B37628D62471C7F921FAC2F90C" authority="Kirejtshuk &amp; Kovalev, 2022" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[1200,1334,1745,1771]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="rectifrons" status="sp. nov.">
<emphasis id="B9711E227628D62471C7F921FAC2F90C" box="[1200,1334,1745,1771]" italics="true" pageId="92" pageNumber="93">I. rectifrons</emphasis>
</taxonomicName>
<emphasis id="B9711E227628D6247037F922FA68F90C" bold="true" box="[1344,1436,1745,1771]" pageId="92" pageNumber="93">
<taxonomicNameLabel id="A242A3597628D6247037F922FA68F90C" box="[1344,1436,1745,1771]" pageId="92" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are different also in the antennomeres 2 and 3 and width of femora (see their descriptions). These two Australian species by some particular characters are somewhat similar to different Indo-Malayan species which were proposed in composition of “
<taxonomicName id="4C05B9B37628D6247400F8CEFE1BF8B0" box="[375,495,1853,1879]" class="Oligohymenophorea" family="Cinetochilidae" genus="Platynema" kingdom="Chromista" order="Hymenostomatida" pageId="92" pageNumber="93" phylum="Ciliophora" rank="genus">
<emphasis id="B9711E227628D6247400F8CEFE1BF8B0" box="[375,495,1853,1879]" italics="true" pageId="92" pageNumber="93">Platynema</emphasis>
</taxonomicName>
”, but the new species distinct from them in many other structural features. In the contrast to other formerly regarded as members of “
<taxonomicName id="4C05B9B37628D624778BF892FC80F89C" box="[764,884,1889,1915]" class="Oligohymenophorea" family="Cinetochilidae" genus="Platynema" kingdom="Chromista" order="Hymenostomatida" pageId="92" pageNumber="93" phylum="Ciliophora" rank="genus">
<emphasis id="B9711E227628D624778BF892FC80F89C" box="[764,884,1889,1915]" italics="true" pageId="92" pageNumber="93">Platynema</emphasis>
</taxonomicName>
”, this new species has the comparatively more subparallel-sided body, slightly modified anterior part of frons, shorter head, longer prothorax, and also the absence of long and thin hairs along thorax and abdomen. The anterior edge of frons and many other features of it are rather similar to those in
<taxonomicName id="4C05B9B37628D6247410F83EFDA5F800" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[359,593,1997,2023]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="breviceps" status="comb. nov.">
<emphasis id="B9711E227628D6247410F83EFDA5F800" box="[359,593,1997,2023]" italics="true" pageId="92" pageNumber="93">Ithyphenes breviceps</emphasis>
</taxonomicName>
<emphasis id="B9711E227628D624772FF83EFD2CF800" bold="true" box="[600,728,1997,2023]" pageId="92" pageNumber="93">
<taxonomicNameLabel id="A242A3597628D624772FF83EFD2CF800" box="[600,728,1997,2023]" pageId="92" pageNumber="93" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, but the new species is distinct from the latter in the longer and narrower prothorax, more expressed striae with punctures on elytra, longer abdominal segments, wider pregenal processes, strongly projecting anterior angles of mentum, markedly shorter and wider oviform ultimate labial palpomere (not subcylindrical as in
<taxonomicName id="4C05B9B37629D625775CFF2CFD58FF1F" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[555,684,222,248]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="breviceps" status="comb. nov.">
<emphasis id="B9711E227629D625775CFF2CFD58FF1F" box="[555,684,222,248]" italics="true" pageId="93" pageNumber="94">I. breviceps</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D62577C3FF2DFCC0FF1F" bold="true" box="[692,820,222,248]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D62577C3FF2DFCC0FF1F" box="[692,820,222,248]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
), antennal grooves extending behind eyes, subtruncate apex of prosternal process (not widely arcuate as in
<taxonomicName id="4C05B9B37629D62577ACFEF1FCA8FEFC" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[731,860,257,283]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="breviceps" status="comb. nov.">
<emphasis id="B9711E227629D62577ACFEF1FCA8FEFC" box="[731,860,257,283]" italics="true" pageId="93" pageNumber="94">I. breviceps</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D6257614FEF1FC17FEFB" bold="true" box="[867,995,258,284]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D6257614FEF1FC17FEFB" box="[867,995,258,284]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
), male pygidium about 1.5 × as long as wide (in
<taxonomicName id="4C05B9B37629D625758DFED5FE8FFED8" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[250,379,293,319]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="breviceps" status="comb. nov.">
<emphasis id="B9711E227629D625758DFED5FE8FFED8" box="[250,379,293,319]" italics="true" pageId="93" pageNumber="94">I. breviceps</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D62574F6FED6FDF5FED8" bold="true" box="[385,513,293,319]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D62574F6FED6FDF5FED8" box="[385,513,293,319]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
the male pygidium slightly longer than wide), weaker serration along the posterior edge of male anal sclerite. Another congener with similar anterior edge of slightly modified frons compared with
<taxonomicName id="4C05B9B37629D62575E0FE9FFE4CFE61" authority="Kirejtshuk &amp; Kovalev, 2022" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[151,440,364,390]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="australiaensis" status="sp. nov.">
<emphasis id="B9711E227629D62575E0FE9FFE4CFE61" box="[151,440,364,390]" italics="true" pageId="93" pageNumber="94">Ithyphenes australiaensis</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D62574B5FE9FFDEAFE61" bold="true" box="[450,542,364,390]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D62574B5FE9FFDEAFE61" box="[450,542,364,390]" pageId="93" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is
<taxonomicName id="4C05B9B37629D6257732FE9EFD3FFE61" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[581,715,364,390]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="planiceps" status="comb. nov.">
<emphasis id="B9711E227629D6257732FE9EFD3FFE61" box="[581,715,364,390]" italics="true" pageId="93" pageNumber="94">I. planiceps</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D62577A2FE9FFCACFE61" bold="true" box="[725,856,364,390]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D62577A2FE9FFCACFE61" box="[725,856,364,390]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, but the new species differs from the latter in the markedly weakly projecting lateral angles of frons, and also in the more convex and more subparallel-sided body, scape longer and less swollen anteriorly (in
<taxonomicName id="4C05B9B37629D6257708FE47FCF4FE2A" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[639,768,435,461]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="planiceps" status="comb. nov.">
<emphasis id="B9711E227629D6257708FE47FCF4FE2A" box="[639,768,435,461]" italics="true" pageId="93" pageNumber="94">I. planiceps</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D6257670FE47FC73FE29" bold="true" box="[775,903,436,462]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D6257670FE47FC73FE29" box="[775,903,436,462]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
the scape less than 2.0 × as long as thick), more consolidated antennal club (in
<taxonomicName id="4C05B9B37629D6257481FE2BFD8EFE16" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[502,634,471,497]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="planiceps" status="comb. nov.">
<emphasis id="B9711E227629D6257481FE2BFD8EFE16" box="[502,634,471,497]" italics="true" pageId="93" pageNumber="94">I. planiceps</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D62577F4FE24FCF1FE16" bold="true" box="[643,773,471,497]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D62577F4FE24FCF1FE16" box="[643,773,471,497]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
antennomeres 9 and 10 more separated) markedly shorter head, prothorax much longer and subrectiliner at sides (not rounded as in
<taxonomicName id="4C05B9B37629D62576BAFE08FBBBFDF2" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[973,1103,507,533]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="planiceps" status="comb. nov.">
<emphasis id="B9711E227629D62576BAFE08FBBBFDF2" box="[973,1103,507,533]" italics="true" pageId="93" pageNumber="94">I. planiceps</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D6257121FE08FB22FDF2" bold="true" box="[1110,1238,507,533]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D6257121FE08FB22FDF2" box="[1110,1238,507,533]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
), absence of very long and fine conspicuous hairs along sides of abdominal segments, markedly weaker mandibles.
</paragraph>
<paragraph id="8BBAC2307629D62575B0FDB1FF04FD98" blockId="93.[151,1437,151,2029]" pageId="93" pageNumber="94">
It can be compared also with the species formerly included into “
<taxonomicName id="4C05B9B37629D62576DCFDB1FBD7FDBB" box="[939,1059,578,604]" class="Oligohymenophorea" family="Cinetochilidae" genus="Platynema" kingdom="Chromista" order="Hymenostomatida" pageId="93" pageNumber="94" phylum="Ciliophora" rank="genus">
<emphasis id="B9711E227629D62576DCFDB1FBD7FDBB" box="[939,1059,578,604]" italics="true" pageId="93" pageNumber="94">Platynema</emphasis>
</taxonomicName>
”, particularly, with the following species:
</paragraph>
<paragraph id="8BBAC2307629D62575B0FD79FB24FC04" blockId="93.[151,1437,151,2029]" pageId="93" pageNumber="94">
with
<taxonomicName id="4C05B9B37629D625746CFD7AFDF6FD44" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Grouvelle" baseAuthorityYear="1917" box="[283,514,649,675]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="angustus" status="comb. nov.">
<emphasis id="B9711E227629D625746CFD7AFDF6FD44" box="[283,514,649,675]" italics="true" pageId="93" pageNumber="94">Ithyphenes angustus</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D625777CFD7AFD7BFD44" bold="true" box="[523,655,649,675]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D625777CFD7AFD7BFD44" box="[523,655,649,675]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
possessing many characters similar to those in
<taxonomicName id="4C05B9B37629D62571CDFD79FA85FD44" authority="Kirejtshuk &amp; Kovalev, 2022" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[1210,1393,649,675]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="australiaensis" status="sp. nov.">
<emphasis id="B9711E227629D62571CDFD79FA85FD44" box="[1210,1393,649,675]" italics="true" pageId="93" pageNumber="94">I. australiaensis</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D625700CFD7AFF3CFD20" bold="true" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D625700CFD7AFF3CFD20" pageId="93" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; however. this new species differs from the latter in the more convex, more subparallel-sided and somewhat lighter body; shorter and thicker antennomeres 2 and 3 (in
<taxonomicName id="4C05B9B37629D625764AFD22FC4EFD0D" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Grouvelle" baseAuthorityYear="1917" box="[829,954,721,746]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="angustus" status="comb. nov.">
<emphasis id="B9711E227629D625764AFD22FC4EFD0D" box="[829,954,721,746]" italics="true" pageId="93" pageNumber="94">I. angustus</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D62576B5FD23FBB1FD0D" bold="true" box="[962,1093,720,746]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D62576B5FD23FBB1FD0D" box="[962,1093,720,746]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
antennomeres 2 and 3 almost 2.0 × as long as thick); more compact antennal club (antennomeres 9 and 10 only slightly separated); weaker mandibles; slightly projecting three processes of frons (frons of
<taxonomicName id="4C05B9B37629D625761EFCEBFC11FCD6" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Grouvelle" baseAuthorityYear="1917" box="[873,997,792,817]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="angustus" status="comb. nov.">
<emphasis id="B9711E227629D625761EFCEBFC11FCD6" box="[873,997,792,817]" italics="true" pageId="93" pageNumber="94">I. angustus</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D625769BFCEBFB99FCD5" bold="true" box="[1004,1133,792,818]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D625769BFCEBFB99FCD5" box="[1004,1133,792,818]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
has only a pair of strongly projecting lateral processes (anterior angles) and deep arcuate excision between them); markedly shorter head, prothorax much longer and subrectiliner at sides (not rounded as in
<taxonomicName id="4C05B9B37629D62576E2FCACFBE5FC9F" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Grouvelle" baseAuthorityYear="1917" box="[917,1041,863,888]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="angustus" status="comb. nov.">
<emphasis id="B9711E227629D62576E2FCACFBE5FC9F" box="[917,1041,863,888]" italics="true" pageId="93" pageNumber="94">I. angustus</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D625716FFCACFB6DFC9E" bold="true" box="[1048,1177,863,889]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D625716FFCACFB6DFC9E" box="[1048,1177,863,889]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
); absence of very long and fine conspicuous hairs along sides of metaventrite and abdominal segments; wider pregenal processes; strongly projecting anterior angles of mentum; much wider antennal grooves extending behind eyes; shape of sclerites of ultimate abdominal segment in both sexes; finer serration of posterior edge of male anal sclerite;
</paragraph>
<paragraph id="8BBAC2307629D62575B0FC1DFD1FFAA0" blockId="93.[151,1437,151,2029]" pageId="93" pageNumber="94">
with
<taxonomicName id="4C05B9B37629D6257461FC1DFE58FBE0" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[278,428,1005,1031]" class="Insecta" family="Nitidulidae" genus="Orthogramma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="fuscipennis" status="comb. nov.">
<emphasis id="B9711E227629D6257461FC1DFE58FBE0" box="[278,428,1005,1031]" italics="true" pageId="93" pageNumber="94">I. fuscipennis</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D62574C3FC1EFDC1FBE0" bold="true" box="[436,565,1005,1031]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D62574C3FC1EFDC1FBE0" box="[436,565,1005,1031]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
) demonstrating a considerable similarity to
<taxonomicName id="4C05B9B37629D625715FFC1DFB29FBE0" authority="Kirejtshuk &amp; Kovalev, 2022" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[1064,1245,1005,1031]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="australiaensis" status="sp. nov.">
<emphasis id="B9711E227629D625715FFC1DFB29FBE0" box="[1064,1245,1005,1031]" italics="true" pageId="93" pageNumber="94">I. australiaensis</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D6257192FC1EFACBFBE0" bold="true" box="[1253,1343,1005,1031]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D6257192FC1EFACBFBE0" box="[1253,1343,1005,1031]" pageId="93" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; but the new species differs from the latter in the more convex, more subparallel-sided and somewhat lighter subunicolorous body; slightly projecting three processes of frons (frons of
<taxonomicName id="4C05B9B37629D6257648FBC6FC22FBA9" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[831,982,1076,1102]" class="Insecta" family="Nitidulidae" genus="Orthogramma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="fuscipennis" status="comb. nov.">
<emphasis id="B9711E227629D6257648FBC6FC22FBA9" box="[831,982,1076,1102]" italics="true" pageId="93" pageNumber="94">I. fuscipennis</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D6257697FBC7FB96FBA9" bold="true" box="[992,1122,1076,1102]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D6257697FBC7FB96FBA9" box="[992,1122,1076,1102]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
has only a pair of strongly projecting lateral processes (anterior angles) and deep arcuate excision between them); shorter antennomeres 2 and 3 (in
<taxonomicName id="4C05B9B37629D62575A1FB8FFE99FB71" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[214,365,1148,1174]" class="Insecta" family="Nitidulidae" genus="Orthogramma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="fuscipennis" status="comb. nov.">
<emphasis id="B9711E227629D62575A1FB8FFE99FB71" box="[214,365,1148,1174]" italics="true" pageId="93" pageNumber="94">I. fuscipennis</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D6257401FB8FFE0CFB71" bold="true" box="[374,504,1148,1174]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D6257401FB8FFE0CFB71" box="[374,504,1148,1174]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
at least antennomere 2 more than 2.0 × as long as thick); markedly shorter head; prothorax much longer and subrectiliner at sides (not rounded as in
<taxonomicName id="4C05B9B37629D62576DEFB53FBB5FB5E" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[937,1089,1183,1209]" class="Insecta" family="Nitidulidae" genus="Orthogramma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="fuscipennis" status="comb. nov.">
<emphasis id="B9711E227629D62576DEFB53FBB5FB5E" box="[937,1089,1183,1209]" italics="true" pageId="93" pageNumber="94">I. fuscipennis</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D625713DFB6CFB39FB5E" bold="true" box="[1098,1229,1183,1209]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D625713DFB6CFB39FB5E" box="[1098,1229,1183,1209]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
); absence of very long and fine conspicuous hairs along sides of thorax and abdominal segments; markedly weaker mandibles; wider pregenal processes; strongly projecting anterior angles of mentum; much wider antennal grooves; shape of sclerites of ultimate abdominal segment in both sexes (particularly, due to wider base and arcuate (not suparallesided) sides); finer serration of posterior edge of male anal sclerite;
</paragraph>
<paragraph id="8BBAC2307629D62575B0FAA1FB0DF9A6" blockId="93.[151,1437,151,2029]" pageId="93" pageNumber="94">
with
<taxonomicName id="4C05B9B37629D625746DFAA1FE54FA8C" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Hisamatsu" baseAuthorityYear="1985" box="[282,416,1362,1387]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="japonicus" status="comb. nov.">
<emphasis id="B9711E227629D625746DFAA1FE54FA8C" box="[282,416,1362,1387]" italics="true" pageId="93" pageNumber="94">I. japonicus</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D62574DDFAA2FDD8FA8C" bold="true" box="[426,556,1361,1387]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D62574DDFAA2FDD8FA8C" box="[426,556,1361,1387]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
demonstrating a considerable similarity to
<taxonomicName id="4C05B9B37629D6257157FAA1FB22FA8C" authority="Kirejtshuk &amp; Kovalev, 2022" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[1056,1238,1361,1387]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="australiaensis" status="sp. nov.">
<emphasis id="B9711E227629D6257157FAA1FB22FA8C" box="[1056,1238,1361,1387]" italics="true" pageId="93" pageNumber="94">I. australiaensis</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D6257197FAA2FAC8FA8C" bold="true" box="[1248,1340,1361,1387]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D6257197FAA2FAC8FA8C" box="[1248,1340,1361,1387]" pageId="93" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; but the new species differs from the latter in the more convex, more subparallel-sided and somewhat darker subunicolorous body; slightly projecting three processes of frons (frons of
<taxonomicName id="4C05B9B37629D6257630FA6AFC3BFA55" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Hisamatsu" baseAuthorityYear="1985" box="[839,975,1433,1458]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="japonicus" status="comb. nov.">
<emphasis id="B9711E227629D6257630FA6AFC3BFA55" box="[839,975,1433,1458]" italics="true" pageId="93" pageNumber="94">I. japonicus</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D62576AEFA6BFBA9FA55" bold="true" box="[985,1117,1432,1458]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D62576AEFA6BFBA9FA55" box="[985,1117,1432,1458]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
has only a pair of strongly projecting lateral processes (anterior angles) and deep arcuate excision between them); markedly shorter head; prothorax much longer and subrectiliner at sides (not rounded as in
<taxonomicName id="4C05B9B37629D62576F2FA13FBEDFA1D" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[901,1049,1504,1530]" class="Insecta" family="Nitidulidae" genus="Orthogramma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="fuscipennis" status="comb. nov.">
<emphasis id="B9711E227629D62576F2FA13FBEDFA1D" box="[901,1049,1504,1530]" italics="true" pageId="93" pageNumber="94">I. fuscipennis</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D6257168FA13FB6BFA1D" bold="true" box="[1055,1183,1504,1530]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D6257168FA13FB6BFA1D" box="[1055,1183,1504,1530]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
); absence of very long and fine conspicuous hairs along sides of abdominal segments; somewhat weaker mandibles; shape of sclerites of ultimate abdominal segment in both sexes; finer serration of the posterior edge of male anal sclerite;
</paragraph>
<paragraph id="8BBAC2307629D62575B0F9B8FC90F842" blockId="93.[151,1437,151,2029]" pageId="93" pageNumber="94">
with
<taxonomicName id="4C05B9B37629D625746AF9B8FE60F983" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Grouvelle" baseAuthorityYear="1897" box="[285,404,1611,1636]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="ritsemai" status="comb. nov.">
<emphasis id="B9711E227629D625746AF9B8FE60F983" box="[285,404,1611,1636]" italics="true" pageId="93" pageNumber="94">I. ritsemai</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D62574E9F9B9FDD7F983" bold="true" box="[414,547,1610,1636]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D62574E9F9B9FDD7F983" box="[414,547,1610,1636]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
demostrating also some similarity to
<taxonomicName id="4C05B9B37629D62576A8F9B8FB63F983" authority="Kirejtshuk &amp; Kovalev, 2022" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[991,1175,1610,1636]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="australiaensis" status="sp. nov.">
<emphasis id="B9711E227629D62576A8F9B8FB63F983" box="[991,1175,1610,1636]" italics="true" pageId="93" pageNumber="94">I. australiaensis</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D62571D5F9B9FB0BF983" bold="true" box="[1186,1279,1610,1636]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D62571D5F9B9FB0BF983" box="[1186,1279,1610,1636]" pageId="93" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; but the new species differs from the latter in the more convex, more subparallel-sided and somewhat lighter subunicolorous body; slightly projecting three processes of frons (frons of
<taxonomicName id="4C05B9B37629D6257648F961FC22F94B" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[831,982,1682,1708]" class="Insecta" family="Nitidulidae" genus="Orthogramma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="fuscipennis" status="comb. nov.">
<emphasis id="B9711E227629D6257648F961FC22F94B" box="[831,982,1682,1708]" italics="true" pageId="93" pageNumber="94">I. fuscipennis</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D6257697F961FB96F94B" bold="true" box="[992,1122,1682,1708]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D6257697F961FB96F94B" box="[992,1122,1682,1708]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
has only a pair of strongly projecting lateral processes (anterior angles) and deep arcuate excision between them); shorter antennomeres 2 and 3 (in
<taxonomicName id="4C05B9B37629D62575A1F92AFE99F914" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[214,365,1753,1779]" class="Insecta" family="Nitidulidae" genus="Orthogramma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="fuscipennis" status="comb. nov.">
<emphasis id="B9711E227629D62575A1F92AFE99F914" box="[214,365,1753,1779]" italics="true" pageId="93" pageNumber="94">I. fuscipennis</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D6257401F92AFE0CF914" bold="true" box="[374,504,1753,1779]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D6257401F92AFE0CF914" box="[374,504,1753,1779]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
at least antennomere 2 more than 2.0 × as long as thick); markedly shorter head; prothorax much longer and subrectiliner at sides (not rounded as in
<taxonomicName id="4C05B9B37629D62576DEF90EFBB5F8F1" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[937,1089,1788,1814]" class="Insecta" family="Nitidulidae" genus="Orthogramma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="fuscipennis" status="comb. nov.">
<emphasis id="B9711E227629D62576DEF90EFBB5F8F1" box="[937,1089,1788,1814]" italics="true" pageId="93" pageNumber="94">I. fuscipennis</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D625713DF90FFB39F8F1" bold="true" box="[1098,1229,1788,1814]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D625713DF90FFB39F8F1" box="[1098,1229,1788,1814]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
); absence of very long and fine conspicuous hairs along sides of thorax and abdominal segments; markedly weaker mandibles; wider pregenal processes; strongly projecting anterior angles of mentum; much wider antennal grooves; shape of sclerites of ultimate abdominal segment in both sexes (particularly, due to the wider base and arcuate (not suparallesided) sides); finer serration of the posterior edge of male anal sclerite;
</paragraph>
<paragraph id="8BBAC2307629D62675B0F85CFCE0FE4B" blockId="93.[151,1437,151,2029]" lastBlockId="94.[151,1437,151,2043]" lastPageId="94" lastPageNumber="95" pageId="93" pageNumber="94">
with
<taxonomicName id="4C05B9B37629D6257461F85CFE6FF82E" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[278,411,1967,1993]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="saundersi" status="comb. nov.">
<emphasis id="B9711E227629D6257461F85CFE6FF82E" box="[278,411,1967,1993]" italics="true" pageId="93" pageNumber="94">I. saundersi</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D62574D4F85CFDD0F82E" bold="true" box="[419,548,1967,1993]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D62574D4F85CFDD0F82E" box="[419,548,1967,1993]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
demostrating some similarity to
<taxonomicName id="4C05B9B37629D62576EEF85CFBBAF82E" authority="Kirejtshuk &amp; Kovalev, 2022" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[921,1102,1967,1993]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="australiaensis" status="sp. nov.">
<emphasis id="B9711E227629D62576EEF85CFBBAF82E" box="[921,1102,1967,1993]" italics="true" pageId="93" pageNumber="94">I. australiaensis</emphasis>
</taxonomicName>
<emphasis id="B9711E227629D6257121F85CFB44F82E" bold="true" box="[1110,1200,1967,1993]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A242A3597629D6257121F85CFB44F82E" box="[1110,1200,1967,1993]" pageId="93" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; but the new species differs from the latter in the more convex, more subparallel-sided and somewhat lighter subunicolorous body; slightly projecting three processes of frons (frons of
<taxonomicName id="4C05B9B3762AD62677A8FF64FC80FF56" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[735,884,151,177]" class="Insecta" family="Nitidulidae" genus="Orthogramma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="fuscipennis" status="comb. nov.">
<emphasis id="B9711E22762AD62677A8FF64FC80FF56" box="[735,884,151,177]" italics="true" pageId="94" pageNumber="95">I. fuscipennis</emphasis>
</taxonomicName>
<emphasis id="B9711E22762AD626760CFF64FC0FFF56" bold="true" box="[891,1019,151,177]" pageId="94" pageNumber="95">
<taxonomicNameLabel id="A242A359762AD626760CFF64FC0FFF56" box="[891,1019,151,177]" pageId="94" pageNumber="95" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
has only a pair of strongly projecting lateral processes (anterior angles) and deep arcuate excision between them); shorter antennomeres 2 and 3 (in
<taxonomicName id="4C05B9B3762AD62670FBFF48FEE1FF1E" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" class="Insecta" family="Nitidulidae" genus="Orthogramma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="fuscipennis" status="comb. nov.">
<emphasis id="B9711E22762AD62670FBFF48FEE1FF1E" italics="true" pageId="94" pageNumber="95">I. fuscipennis</emphasis>
</taxonomicName>
<emphasis id="B9711E22762AD626746BFF2CFE69FF1E" bold="true" box="[284,413,223,249]" pageId="94" pageNumber="95">
<taxonomicNameLabel id="A242A359762AD626746BFF2CFE69FF1E" box="[284,413,223,249]" pageId="94" pageNumber="95" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
at least antennomere 2 more than 2.0 × as long as thick); markedly shorter head; prothorax much longer and subrectiliner at sides (not rounded as in
<taxonomicName id="4C05B9B3762AD626766CFEF0FC45FEFA" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" baseAuthorityName="Murray" baseAuthorityYear="1864" box="[795,945,259,285]" class="Insecta" family="Nitidulidae" genus="Orthogramma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="fuscipennis" status="comb. nov.">
<emphasis id="B9711E22762AD626766CFEF0FC45FEFA" box="[795,945,259,285]" italics="true" pageId="94" pageNumber="95">I. fuscipennis</emphasis>
</taxonomicName>
<emphasis id="B9711E22762AD62676CFFEF0FBCDFEFA" bold="true" box="[952,1081,259,285]" pageId="94" pageNumber="95">
<taxonomicNameLabel id="A242A359762AD62676CFFEF0FBCDFEFA" box="[952,1081,259,285]" pageId="94" pageNumber="95" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
); absence of very long and fine conspicuous hairs along sides of thorax and abdominal segments; markedly weaker mandibles; wider pregenal processes; strongly projecting anterior angles of mentum; much wider antennal grooves; shape of sclerites of ultimate abdominal segment in both sexes (particularly, due to wider base and arcuate (not suparallesided) sides); finer serration of the posterior edge of male anal sclerite.
</paragraph>
</subSubSection>
<subSubSection id="C31F91BB762AD62675B0FE45FC59FE37" box="[199,941,438,464]" pageId="94" pageNumber="95" type="etymology">
<paragraph id="8BBAC230762AD62675B0FE45FC59FE37" blockId="94.[151,1437,151,2043]" box="[199,941,438,464]" pageId="94" pageNumber="95">
<emphasis id="B9711E22762AD62675B0FE45FEBCFE37" bold="true" box="[199,328,438,464]" pageId="94" pageNumber="95">Etymology</emphasis>
. The epithet new species is formed from the
<collectingCountry id="F31282A0762AD6267637FE45FC53FE37" box="[832,935,438,464]" name="Australia" pageId="94" pageNumber="95">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31F91BB762AD61875B0FE2AFCEFFE6E" lastPageId="96" lastPageNumber="97" pageId="94" pageNumber="95" type="description">
<paragraph id="8BBAC230762AD62675B0FE2AFE5DFD64" blockId="94.[151,1437,151,2043]" pageId="94" pageNumber="95">
<emphasis id="B9711E22762AD62675B0FE2AFE15FE13" bold="true" box="[199,481,473,500]" pageId="94" pageNumber="95">
Description of
<typeStatus id="54BE7C92762AD626740DFE29FE15FE13" box="[378,481,474,500]" pageId="94" pageNumber="95" type="holotype">holotype</typeStatus>
</emphasis>
(
<emphasis id="B9711E22762AD6267484FE29FDD9FE13" bold="true" box="[499,557,474,500]" pageId="94" pageNumber="95">male</emphasis>
). Length of body with abdomen 6.8, length without abdomen 4.0, width 1.2, height
<quantity id="4CFD6FD5762AD6267593FE0DFEC8FDFF" box="[228,316,510,536]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="94" pageNumber="95" unit="mm" value="0.7">0.7 mm</quantity>
. Elongate, subflattened dorsally and rather convex ventrally; subunicolorous reddish with somewhat darkened elytra; moderately shining; almost without pubescence, although posterior edge of pronotum with dense microsetae, abdominal sclerites with short and thin inconspicuous yellowish setae, posterior edge of anal sclerite with thin and long hairs.
</paragraph>
<paragraph id="8BBAC230762AD62675B0FD7EFC2AFCBD" blockId="94.[151,1437,151,2043]" pageId="94" pageNumber="95">Head and pronotum with punctures, slightly coarser than eye facets, separated by 3.04.0 × greater than one puncture diameter, interspaces between them smooth to slightly alutaceous; elytra with weak and slightly oblique striae coordinated with shallow punctures in bottom of these striae (punctures about as coarse as those on head and pronotum); exposed abdominal dorsal sclerites with shallower and somewhat finer punctures than those on head and pronotum, and obliterately microreticulate interspaces between them. Prosternum and metaventrite with rather fine and sparse punctures along middle, interspaces between punctures smooth.</paragraph>
<paragraph id="8BBAC230762AD62675B0FC97FB6FFA7E" blockId="94.[151,1437,151,2043]" pageId="94" pageNumber="95">Head widest behind eyes and slightly arcuately narrowing posteriorly, temples about 1.5 × as long as eye.Anterior part of frons subflattened, its anterior edge with slight median projection and one couple of slight paramedian lateral projections. Labrum slightly exposed from under frons and visible from above, with widely rounded and sparsely serrate anterior edge of lobes. Mandibles moderately stout and with bidentate apex strongly curved downwards, subtransverse anterior edge and gently rounded at side.Antennae slightly longer than head wide; scape comparatively narrow and slightly curved, about 2.0 × as long as each of antennomeres 2 and 3; antennomere 2 (pedicel) slightly thicker than antennomere 3; oblong club about 0.3× total antennal length, somewhat more than 2.0 × as long as wide, with antennomeres 9 and 10 comparable in width and length and somewhat separated from each other, and slightly wider and longer than antennomere 11, apex of terminal antennomere rounded to subacute. Pronotum about 1.3 × as long as wide and somewhat rectilinearly widening posteriorly, anterior edge almost straight and posterior one evenly convex, posterior angles widely rounded. Scutellar shield subtriangular, about 2.0 × as wide as long. Elytra slightly widening posteriorly, nearly 1.5 × as long as their combined width, moderately steeply sloping at narrowly explanate sides. Abdomen about as wide as pronotum with only distal part of segment 4 exposed dorsally and with laterosternites clearly widened posteriorly, segment 5 almost 2.0 × as wide as long, tergite VI somewhat longer than tergite V; postbasal lines of exposed tergites behind elytral apices well expressed and their lateral ends reaching middle of tergites. Pygidium nearly 1.2 × as long as wide, widely rounded at apex.</paragraph>
<paragraph id="8BBAC230762AD62675B0FA51FD9AF8A0" blockId="94.[151,1437,151,2043]" pageId="94" pageNumber="95">Pregenal processes comparatively narrow and at wide hypostomal sinuses subarcuately curved. Terminal maxillary palpomere more than 2.5 × as long as thick and subcylindrical. Terminal labial palpomere slightly longer than thick, subovoid and with oblique apex. Mentum rather widened anteriorly and with sides of mentum about 2.0 × as long as its median part. S-shaped antennal grooves somewhat divergent, their inner side quite distinct behind posterior level of eye. Median plate of prosternum almost flatly widened anteriorly from prosternal process. Prosternal process strongly widened before subtruncate apex, its apex nearly 2.0 × as wide as distance between procoxae. Distance between procoxae about 0.5 × as great as length of antennal club, and somewhat smaller than that between mesocoxae and somewhat greater than that between metacoxae. Mesothoracic median plate distinctly isolated by longer lateral ridges. Metaventrite subflattened and with clear discrimen in distal third. Metepisterna gradually narrowing posteriorly and at base about as wide as mesotarsomere 1. Abdominal ventrite 1 somewhat longer than ventrite 2 and somewhat shorter than each of ventrites 3 and 4. Hypopygidium somewhat transverse and with shallowly emarginate posterior edge.</paragraph>
<paragraph id="8BBAC230762AD62675B0F8A2FA78F81C" blockId="94.[151,1437,151,2043]" pageId="94" pageNumber="95">Pro- and mesofemora about 3.0 × as wide as corresponding tibiae and metafemora almost 3.5 × as wide as corresponding tibia. Protibia somewhat narrower than antennal club, comparatively coarsely serrate along outer edge. Meso- and metatibiae somewhat narrower than protibia, with two unclear rows of short and stout spines along outer edge. Protibia with very long and stout spur, meso-and metatibiae with moderately long spurs. Tarsi with thickest tarsomere 1 (slightly wider than antennomere 2), following tarsomeres thinner, claws simple and narrow.</paragraph>
<paragraph id="8BBAC230762BD62775B0FF64FBF7FF1E" blockId="95.[151,1437,151,249]" pageId="95" pageNumber="96">Anal sclerite with subtruncate and serrate apex bearing ten teeth (including one very small tooth at each side). Aedeagus moderately sclerotized and comparatively small, tegmen widely rounded at apex; penis trunk about as long as tegmen, somewhat arcuate at sides and nearly widely rounded at apex.</paragraph>
<caption id="DF7A92B8762BD62775E0F899FC1AF809" ID-DOI="http://doi.org/10.5281/zenodo.6245855" ID-Zenodo-Dep="6245855" httpUri="https://zenodo.org/record/6245855/files/figure.png" pageId="95" pageNumber="96" startId="95.[151,250,1898,1922]" targetBox="[233,1354,281,1868]" targetPageId="95">
<paragraph id="8BBAC230762BD62775E0F899FC1AF809" blockId="95.[151,1437,1898,2031]" pageId="95" pageNumber="96">
<emphasis id="B9711E22762BD62775E0F899FED4F864" bold="true" box="[151,288,1898,1923]" pageId="95" pageNumber="96">FIGURE 28.</emphasis>
<taxonomicName id="4C05B9B3762BD6277451F898FDEFF864" authority="Kirejtshuk &amp; Kovalev, 2022" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[294,539,1899,1923]" class="Insecta" family="Nitidulidae" genus="Ithyphenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="australiensis" status="sp. nov.">
<emphasis id="B9711E22762BD6277451F898FDEFF864" box="[294,539,1899,1923]" italics="true" pageId="95" pageNumber="96">Ithyphenes australiensis</emphasis>
</taxonomicName>
<emphasis id="B9711E22762BD6277756F898FD85F864" bold="true" box="[545,625,1899,1923]" pageId="95" pageNumber="96">
<taxonomicNameLabel id="A242A359762BD6277756F898FD85F864" box="[545,625,1899,1923]" pageId="95" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, orig. AD, FH: holotype, male from NSW (“3 km N of Landsowne …”); E, I: paratype from QLD (“… Cape Tribulation…”);
<emphasis id="B9711E22762BD627770EF87DFD7FF841" bold="true" box="[633,651,1934,1958]" pageId="95" pageNumber="96">A</emphasis>
: body, dorsal;
<emphasis id="B9711E22762BD6277669F87CFCDBF840" bold="true" box="[798,815,1935,1959]" pageId="95" pageNumber="96">B</emphasis>
: head, dorsal;
<emphasis id="B9711E22762BD62776B6F87DFC27F841" bold="true" box="[961,979,1934,1958]" pageId="95" pageNumber="96">C</emphasis>
: head, ventral;
<emphasis id="B9711E22762BD627711AF87CFB8BF840" bold="true" box="[1133,1151,1935,1959]" pageId="95" pageNumber="96">D</emphasis>
: thorax, ventral;
<emphasis id="B9711E22762BD627705CF87CFAC8F840" bold="true" box="[1323,1340,1935,1959]" pageId="95" pageNumber="96">E</emphasis>
: terminal abdominal segment, ventral;
<emphasis id="B9711E22762BD62774B3F840FE27F82C" bold="true" box="[452,467,1971,1995]" pageId="95" pageNumber="96">F</emphasis>
: male anal sclerite, ventral;
<emphasis id="B9711E22762BD627778AF841FCE5F82D" bold="true" box="[765,785,1970,1994]" pageId="95" pageNumber="96">G</emphasis>
: tegmen, ventral;
<emphasis id="B9711E22762BD62776B8F840FC17F82C" bold="true" box="[975,995,1971,1995]" pageId="95" pageNumber="96">H</emphasis>
: penis trunk, dorsal;
<emphasis id="B9711E22762BD62771B5F840FB38F82C" bold="true" box="[1218,1228,1971,1995]" pageId="95" pageNumber="96">I</emphasis>
: ovipositor, ventral. Scale bars:
<emphasis id="B9711E22762BD627747BF824FEEDF808" bold="true" box="[268,281,2007,2031]" pageId="95" pageNumber="96">a</emphasis>
= 1.0 mm to Fig. A;
<emphasis id="B9711E22762BD6277499F824FE08F808" bold="true" box="[494,508,2007,2031]" pageId="95" pageNumber="96">b</emphasis>
= 0.5 mm to Figs BE;
<emphasis id="B9711E22762BD6277786F824FD08F808" bold="true" box="[753,764,2007,2031]" pageId="95" pageNumber="96">с</emphasis>
= 0.25 mm to Figs FI.
</paragraph>
</caption>
<paragraph id="8BBAC2307614D61875B0FF64FCEFFE6E" blockId="96.[151,1437,150,502]" pageId="96" pageNumber="97">
<emphasis id="B9711E227614D61875B0FF64FEEAFF56" bold="true" box="[199,286,151,177]" pageId="96" pageNumber="97">Female</emphasis>
. Length of body with abdomen
<quantity id="4CFD6FD57614D61877F4FF65FD2FFF56" box="[643,731,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="96" pageNumber="97" unit="mm" value="5.6">5.6 mm</quantity>
. Differs from male in structure of terminal abdominal segment: more coarsely and widely rounded apex of pygidium (about 1.3 × as long as wide) bearing ten small teeth and widely rounded apex hypopygidium. Mandibles with arcuate anterior edge and roundly angular at side, their apex slightly turning downwards. Ovipositor moderately sclerotized and rather wide; gonocoxites comprising almost 0.4 of entire length, lateral lobes slightly sclerotized and about 0.5 × as long as gonocoxites in general, outer outline of gonocoxites slightly concave and sinuated posteriorly from apices of lateral lobes, moderately narrow apex with moderately long styli located at great distance from apex.
</paragraph>
</subSubSection>
<subSubSection id="C31F91BB7614D61875B0FE61FA8DFE11" pageId="96" pageNumber="97" type="distribution">
<paragraph id="8BBAC2307614D61875B0FE61FED1FE35" blockId="96.[151,1437,150,502]" pageId="96" pageNumber="97">
<emphasis id="B9711E227614D61875B0FE61FEADFE4B" bold="true" box="[199,345,402,428]" pageId="96" pageNumber="97">Distribution</emphasis>
. This species is known only from
<typeStatus id="54BE7C927614D618766CFE60FCBFFE4A" box="[795,843,403,429]" pageId="96" pageNumber="97">type</typeStatus>
locality in NSW (Landsowne) and QLD (Cape Tribulation).
</paragraph>
<paragraph id="8BBAC2307614D61875B0FE28FA8DFE11" blockId="96.[151,1437,150,502]" box="[199,1401,474,502]" pageId="96" pageNumber="97">
<emphasis id="B9711E227614D61875B0FE28FE6AFE13" bold="true" box="[199,414,474,501]" pageId="96" pageNumber="97">Notes on bionomy</emphasis>
. This species can be associated with dying and dead trees (dead Ficus obliqua branch”).
</paragraph>
</subSubSection>
</treatment>
</document>