243 lines
30 KiB
XML
243 lines
30 KiB
XML
<document id="97870AB1636A373A2D5BB8F35EE4B142" ID-CLB-Dataset="39310" ID-DOI="10.11646/zootaxa.4055.1.1" ID-GBIF-Dataset="7109f202-9561-476f-8f75-8c8f747656c9" ID-ISSN="1175-5326" ID-Zenodo-Dep="240153" ID-ZooBank="0604E6DE-9DF8-4EEE-A225-9584143D8312" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461353560128" checkinUser="plazi" docAuthor="Have, Ten" docDate="2015" docId="03A087E6FFD08319D788FBEAD74EFAE4" docLanguage="en" docName="zt04055p073.pdf" docOrigin="Zootaxa 4055 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Phascogale tapoatafa subsp. kimberleyensis Aplin & Rhind, subsp. nov." docType="treatment" docVersion="11" lastPageNumber="40" masterDocId="FF99FF9EFFF5833ED71FFFDDD666FF80" masterDocTitle="Taxonomic revision of Phascogale tapoatafa (Meyer, 1793) (Dasyuridae; Marsupialia), including descriptions of two new subspecies and confirmation of P. pirata Thomas, 1904 as a ‘ Top End’ endemic" masterLastPageNumber="73" masterPageNumber="1" pageNumber="38" updateTime="1698661393127" updateUser="ExternalLinkService">
|
||
<mods:mods id="4BF211CFC2DE967B438DE247F2AF80EF" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="AA14BBBF11832EA9643CF7594308DD64">
|
||
<mods:title id="722A579EB4B6B4C47B0CE9A56F0F1778">Taxonomic revision of Phascogale tapoatafa (Meyer, 1793) (Dasyuridae; Marsupialia), including descriptions of two new subspecies and confirmation of P. pirata Thomas, 1904 as a ‘ Top End’ endemic</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="5EC44052AE655F90BE50D63E53637423" type="personal">
|
||
<mods:role id="DE43781D5B15A1A561E3A00F4CF903C6">
|
||
<mods:roleTerm id="B461D4A41157BB540F608250A3F1BCD0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A6399E92E42AA15EC2BD4E893D8BF3AB">Have, Ten</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="76750A30BEF8A19BC1A66F8D3D94BA62">text</mods:typeOfResource>
|
||
<mods:relatedItem id="C28678976F39631A2309B93D30E4EA82" type="host">
|
||
<mods:titleInfo id="C51F0367F4E05B5AC3AD831533C29C0E">
|
||
<mods:title id="9D4C6F18A025969B93DA89BF4857948F">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="7CDB4249184FBD6DE15B9A29BA7BC9A7">
|
||
<mods:date id="0BFD6ABF648A95F8A06716FAFF776A9A">2015</mods:date>
|
||
<mods:detail id="63438ADC74F7DBB0BD35FF40F9A404C2" type="volume">
|
||
<mods:number id="3870C4C577712C572F4C3BBEE8F03EDA">4055</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="3DBA9A8F78D0F59BDED59E499AAC579B" type="issue">
|
||
<mods:number id="3E6D451A708DC2A86B792CF19C3C0C17">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="3718B579DA0D05688B9509104D83CAC0" unit="page">
|
||
<mods:start id="82AD4F046C177B6F09D58064B123201B">1</mods:start>
|
||
<mods:end id="39404A8454179AB35E6E9CC8C1EC1A7F">73</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="084B9D0F983D068A8098D54EE618FA63">journal article</mods:classification>
|
||
<mods:identifier id="1FAB2F77E8A7E9A6D40559B2D222E51B" type="CLB-Dataset">39310</mods:identifier>
|
||
<mods:identifier id="49AF2D73351935F70558CF6DCE65780F" type="DOI">10.11646/zootaxa.4055.1.1</mods:identifier>
|
||
<mods:identifier id="E7F035E4D1EF37A83FFB3DF0DBC90ABE" type="GBIF-Dataset">7109f202-9561-476f-8f75-8c8f747656c9</mods:identifier>
|
||
<mods:identifier id="59E33FCED44E9CAC66C755C2A528261F" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="453319F992D40E8DE05D347C843D3838" type="Zenodo-Dep">240153</mods:identifier>
|
||
<mods:identifier id="536C854AE52AC066172745FD222FD2EF" type="ZooBank">0604E6DE-9DF8-4EEE-A225-9584143D8312</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03A087E6FFD08319D788FBEAD74EFAE4" ID-DOI="http://doi.org/10.5281/zenodo.6097135" ID-GBIF-Taxon="119647157" ID-Zenodo-Dep="6097135" LSID="urn:lsid:plazi:treatment:03A087E6FFD08319D788FBEAD74EFAE4" httpUri="http://treatment.plazi.org/id/03A087E6FFD08319D788FBEAD74EFAE4" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">
|
||
<subSubSection id="C313657BFFD0831BD788FBEAD5C1FBD2" box="[151,935,1079,1106]" pageId="37" pageNumber="38" type="nomenclature">
|
||
<paragraph id="8BB636F0FFD0831BD788FBEAD5C1FBD2" blockId="37.[151,935,1079,1106]" box="[151,935,1079,1106]" pageId="37" pageNumber="38">
|
||
<heading id="D0FE819CFFD0831BD788FBEAD5C1FBD2" bold="true" box="[151,935,1079,1106]" fontSize="11" level="1" pageId="37" pageNumber="38" reason="1">
|
||
<emphasis id="B97DEAE2FFD0831BD788FBEAD5C1FBD2" bold="true" box="[151,935,1079,1106]" pageId="37" pageNumber="38">
|
||
<taxonomicName id="4C094D73FFD0831BD788FBEAD570FBD2" authority="Aplin & Rhind" authorityName="Aplin & Rhind" box="[151,790,1079,1106]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="subSpecies" species="tapoatafa" status="subsp. nov." subSpecies="kimberleyensis">
|
||
<emphasis id="B97DEAE2FFD0831BD788FBEAD432FBD1" bold="true" box="[151,596,1079,1105]" italics="true" pageId="37" pageNumber="38">Phascogale tapoatafa kimberleyensis</emphasis>
|
||
Aplin & Rhind
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E5799FFD0831BD401FBE5D5C1FBD2" box="[798,935,1080,1106]" pageId="37" pageNumber="38" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C313657BFFD0831BD788FB5CD7E9FA8C" pageId="37" pageNumber="38" type="materials_examined">
|
||
<paragraph id="8BB636F0FFD0831BD788FB5CD5AAFB41" blockId="37.[151,1437,1150,2012]" pageId="37" pageNumber="38">
|
||
<materialsCitation id="3B613CADFFD0831BD788FB5CD5AEFB41" ID-GBIF-Occurrence="2465041881" collectingDate="1910-02-19" collectionCode="WAM" collectorName="Gerald F. Hill" country="Australia" latitude="-14.8" location="Pago Mission, Napier Bay, near Broome" longitude="126.43" pageId="37" pageNumber="38" specimenCode="16028" specimenCount="2" specimenCount-male="1" stateProvince="Western Australia" typeStatus="holotype">
|
||
<typeStatus id="54B28852FFD0831BD788FB5CD762FB1A" box="[151,260,1153,1178]" pageId="37" pageNumber="38" type="holotype">
|
||
<emphasis id="B97DEAE2FFD0831BD788FB5CD762FB1A" bold="true" box="[151,260,1153,1178]" pageId="37" pageNumber="38">Holotype</emphasis>
|
||
</typeStatus>
|
||
:
|
||
<collectionCode id="ED18AE35FFD0831BD60BFB5CD738FB1A" LSID="urn:lsid:biocol.org:col:34981" box="[276,350,1153,1178]" collectionName="Australia, Western Australia, Perth, Western Australian Museum" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="37" pageNumber="38">WAM</collectionCode>
|
||
16028. Adult male. Pago Mission, Napier Bay, near Broome,
|
||
<collectingRegion id="49CDF812FFD0831BD307FB5FD280FB1A" box="[1048,1254,1153,1178]" pageId="37" pageNumber="38">Western Australia</collectingRegion>
|
||
, 14
|
||
<geoCoordinate id="EE3D5037FFD0831BD211FBA3D35BFB1A" box="[1294,1341,1150,1178]" degrees="0" direction="south" minutes="8" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-0.13333334">
|
||
0 8’
|
||
<geoCoordinate id="EE3D5037FFD0831BD234FB5FD35BFB1A" box="[1323,1341,1154,1178]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="15" value="-126.01195">S</geoCoordinate>
|
||
</geoCoordinate>
|
||
126
|
||
<geoCoordinate id="EE3D5037FFD0831BD270FBA3D6CDFB40" degrees="0" direction="east" minutes="43" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="0.71666664">0 43’ E</geoCoordinate>
|
||
. Collected by Gerald F. Hill,
|
||
<date id="FFB71030FFD0831BD6ECFB74D4AFFB41" box="[499,713,1189,1217]" pageId="37" pageNumber="38" value="1910-02-19">19th February 1910</date>
|
||
. Puppet skin and skull
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB636F0FFD0831BD7D8FB12D7E9FA8C" blockId="37.[151,1437,1150,2012]" pageId="37" pageNumber="38">
|
||
<materialsCitation id="3B613CADFFD0831BD7D8FB12D7EDFA8C" ID-GBIF-Occurrence="2465041883" collectingDate="1909-10-20" collectionCode="WAM" collectorName="Gerald F. Hill" country="Australia" latitude="-14.8" location="Pago Mission, Napier Bay, near Broome" longitude="126.43" pageId="37" pageNumber="38" specimenCode="16029" specimenCount="2" specimenCount-male="1" stateProvince="Western Australia" typeStatus="paratype">
|
||
<typeStatus id="54B28852FFD0831BD7D8FB12D752FB68" box="[199,308,1231,1256]" pageId="37" pageNumber="38" type="paratype">
|
||
<emphasis id="B97DEAE2FFD0831BD7D8FB12D752FB68" bold="true" box="[199,308,1231,1256]" pageId="37" pageNumber="38">Paratype</emphasis>
|
||
</typeStatus>
|
||
:
|
||
<collectionCode id="ED18AE35FFD0831BD65DFB12D7EAFB68" LSID="urn:lsid:biocol.org:col:34981" box="[322,396,1231,1256]" collectionName="Australia, Western Australia, Perth, Western Australian Museum" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="37" pageNumber="38">WAM</collectionCode>
|
||
16029. Subadult male. Locality and collector as for
|
||
<typeStatus id="54B28852FFD0831BD4D6FB12D249FB68" box="[969,1071,1231,1256]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
. Collected on
|
||
<date id="FFB71030FFD0831BD3D1FB12D3FEFB67" box="[1230,1432,1228,1256]" pageId="37" pageNumber="38" value="1909-10-20">20th October 1909</date>
|
||
. Puppet skin and skull
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C313657BFFD0831BD7D8FACAD274FAAF" box="[199,1042,1303,1328]" pageId="37" pageNumber="38" type="etymology">
|
||
<paragraph id="8BB636F0FFD0831BD7D8FACAD274FAAF" blockId="37.[151,1437,1150,2012]" box="[199,1042,1303,1328]" pageId="37" pageNumber="38">
|
||
<emphasis id="B97DEAE2FFD0831BD7D8FACAD720FAB0" bold="true" box="[199,326,1303,1328]" pageId="37" pageNumber="38">Etymology</emphasis>
|
||
. Referring to the geographic region of occurrence of this taxon.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C313657BFFD0831BD7D8FAE6D352F9CF" pageId="37" pageNumber="38" type="diagnosis">
|
||
<paragraph id="8BB636F0FFD0831BD7D8FAE6D5C3F9AC" blockId="37.[151,1437,1150,2012]" pageId="37" pageNumber="38">
|
||
<emphasis id="B97DEAE2FFD0831BD7D8FAE6D727FAD4" bold="true" box="[199,321,1339,1364]" pageId="37" pageNumber="38">Diagnosis.</emphasis>
|
||
A small-bodied race of
|
||
<taxonomicName id="4C094D73FFD0831BD542FAE1D480FAD3" box="[605,742,1339,1363]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="tapoatafa">
|
||
<emphasis id="B97DEAE2FFD0831BD542FAE1D480FAD3" box="[605,742,1339,1363]" italics="true" pageId="37" pageNumber="38">P. tapoatafa</emphasis>
|
||
</taxonomicName>
|
||
that differs from the typical race of southeastern Australia in the following ways: externally, with shorter, crisper and generally paler dorsal fur, more extensive cream fur on venter, relatively shorter tail, more boldly patterned face with contrasting cream muzzle and cheek, inner metatarsal pad contacting but clearly distinct from first interdigital pad, probable lower number of teats; cranially, very similar to the other subspecies of
|
||
<taxonomicName id="4C094D73FFD0831BD54AFA11D4B8FA63" box="[597,734,1483,1507]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="tapoatafa">
|
||
<emphasis id="B97DEAE2FFD0831BD54AFA11D4B8FA63" box="[597,734,1483,1507]" italics="true" pageId="37" pageNumber="38">P. tapoatafa</emphasis>
|
||
</taxonomicName>
|
||
but smaller in all cranial dimensions for both sexes and with proportionally narrower foramen magnum, relatively deeper rostrum and proportionally larger molars; genetically, with 3.0–5.8 % sequence divergence for
|
||
<emphasis id="B97DEAE2FFD0831BD540F9C9D4F3F9AB" box="[607,661,1555,1579]" italics="true" pageId="37" pageNumber="38">cyt b</emphasis>
|
||
from other populations.
|
||
</paragraph>
|
||
<paragraph id="8BB636F0FFD0831BD7D8F9EAD352F9CF" blockId="37.[151,1437,1150,2012]" box="[199,1332,1591,1616]" pageId="37" pageNumber="38">
|
||
Additional differences from
|
||
<taxonomicName id="4C094D73FFD0831BD519F9E5D4DCF9CF" box="[518,698,1591,1615]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="subSpecies" species="tapoatafa" subSpecies="wambenger">
|
||
<emphasis id="B97DEAE2FFD0831BD519F9E5D4DCF9CF" box="[518,698,1591,1615]" italics="true" pageId="37" pageNumber="38">P. t. wambenger</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B97DEAE2FFD0831BD5EFF9E5D52BF9CF" box="[752,845,1592,1615]" italics="true" pageId="37" pageNumber="38">
|
||
<taxonomicName id="4C094D73FFD0831BD5EFF9E5D559F9CF" box="[752,831,1592,1615]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="pirat">P. pirat</taxonomicName>
|
||
a
|
||
</emphasis>
|
||
are listed under the accounts of those taxa.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C313657BFFD08319D7D8F986D74EFAE4" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" type="description">
|
||
<paragraph id="8BB636F0FFD0831BD7D8F986D4E9F918" blockId="37.[151,1437,1150,2012]" pageId="37" pageNumber="38">
|
||
<emphasis id="B97DEAE2FFD0831BD7D8F986D780F9F4" bold="true" box="[199,486,1627,1652]" pageId="37" pageNumber="38">
|
||
Description of
|
||
<typeStatus id="54B28852FFD0831BD667F986D787F9F4" box="[376,481,1627,1652]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
External measurements, taken from label: Total length 350, Ear 30; as measured on puppet skin: HB c. 200, TL c. 205, Pes c. 35.
|
||
</paragraph>
|
||
<paragraph id="8BB636F0FFD0831BD7D8F97ED412F814" blockId="37.[151,1437,1150,2012]" pageId="37" pageNumber="38">
|
||
The dorsal fur is relatively short and crisp, the contour hairs measuring
|
||
<quantity id="4CF19B15FFD0831BD311F97ED208F93B" box="[1038,1134,1699,1724]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="37" pageNumber="38" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
|
||
in the centre of the back. Overall colour of the back and flanks is pale grey with a silvery frosting. Abundant and evenly distributed blacktipped guard hairs project up to
|
||
<quantity id="4CF19B15FFD0831BD6E5F931D45DF883" box="[506,571,1772,1795]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="37" pageNumber="38" unit="mm" value="7.0">7 mm</quantity>
|
||
beyond the contour hairs. The ventral fur is a uniform cream from the vent to the chin. A diffuse, yellowish patch on the throat overlies a pigmented gular gland that measures
|
||
<quantity id="4CF19B15FFD0831BD3C3F8D2D34DF8A7" box="[1244,1323,1807,1832]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="37" pageNumber="38" unit="mm" value="25.0">25 mm</quantity>
|
||
long by
|
||
<quantity id="4CF19B15FFD0831BD291F8D2D6A5F8CB" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="37" pageNumber="38" unit="mm" value="6.0">6 mm</quantity>
|
||
wide. The boundary between flank and ventral colouration is relatively sharp. The fore- and hind-limbs are grey externally and cream internally, with a sharp demarcation, especially on the hindlimb. The scrotum is lightly pigmented and thinly clothed in cream fur.
|
||
</paragraph>
|
||
<paragraph id="8BB636F0FFD08318D7D8F842D4BCFEC0" blockId="37.[151,1437,1150,2012]" lastBlockId="38.[151,1436,151,429]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">The face is boldly patterned with a russet mid-rostral stripe that merges posteriorly into a dark grey patch on the crown, a cream preauricular patch, a dark loreal stripe, a dark postorbital spot, and distinct, black-rimmed orbital crescents. The cream fur of the chin and throat merges with a cream supralabial bar that extends partway along the upper lip, while the cheek patch is cream peppered with black. The vibrissal pads are thinly clothed in short cream fur, peppered with black. The mystacial vibrissae are thick and black, some with short gingery tipping, but the papillae are unpigmented. The pinnae are moderately large and thinly furred except along the anterior margin which supports a fringe of short black hairs.</paragraph>
|
||
<paragraph id="8BB636F0FFD38318D7D8FE91D2CAFE2D" blockId="38.[151,1436,151,429]" pageId="38" pageNumber="39">
|
||
The tail supports a black ‘brush’ measuring
|
||
<quantity id="4CF19B15FFD38318D5DFFE91D547FEE4" box="[704,801,332,357]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" pageId="38" pageNumber="39" unit="mm" value="135.0">135 mm</quantity>
|
||
. The basal portion of the tail is only moderately furred, producing the effect of an elongate ‘stem’ (c.
|
||
<quantity id="4CF19B15FFD38318D5BBFEB2D49EFE08" box="[676,760,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" pageId="38" pageNumber="39" unit="mm" value="67.0">67 mm</quantity>
|
||
). The dorsal fur of the stem is grey near the base, blending distally into off-white. The under-surface is grey blending into russet at the base of the brush.
|
||
</paragraph>
|
||
<caption id="DF766678FFD38318D788F84AD3E7F84C" httpUri="https://zenodo.org/record/240174/files/figure.png" pageId="38" pageNumber="39" targetBox="[151,1436,481,1920]" targetPageId="38">
|
||
<paragraph id="8BB636F0FFD38318D788F84AD3E7F84C" blockId="38.[151,1436,1943,1996]" pageId="38" pageNumber="39">
|
||
<emphasis id="B97DEAE2FFD38318D788F84AD77EF82C" bold="true" box="[151,280,1943,1965]" pageId="38" pageNumber="39">FIGURE 21</emphasis>
|
||
. Cranium and dentary of WAM16028, the holotype of
|
||
<taxonomicName id="4C094D73FFD38318D45BF845D23FF82D" box="[836,1113,1943,1965]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="38" pageNumber="39" phylum="Chordata" rank="subSpecies" species="tapoatafa" status="subsp. nov." subSpecies="kimberleyensis">
|
||
<emphasis id="B97DEAE2FFD38318D45BF845D23FF82D" box="[836,1113,1943,1965]" italics="true" pageId="38" pageNumber="39">P. tapoatafa kimberleyensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E5799FFD38318D37FF84AD2B5F82D" box="[1120,1235,1943,1965]" pageId="38" pageNumber="39" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
|
||
, an adult male from Pago Mission, Western Australia: a) dorsal view of cranium, b) ventral view of cranium, and c) lateral view of right dentary.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB636F0FFD28319D7D8FF4AD2D0FEBE" blockId="39.[151,1437,151,1380]" pageId="39" pageNumber="40">
|
||
The upper surface of the pes is grey with a silvery frosting, changing abruptly to white on the sides and on the digits. The plantar surface of the pes is granular and supports six conspicuously striated, raised pads. The inner metatarsal and first interdigital pads are in contact but with an obvious line of demarcation; their combined length is
|
||
<quantity id="4CF19B15FFD28319D7ABFEDFD779FE9A" box="[180,287,258,283]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="39" pageNumber="40" unit="mm" value="12.6">12.6 mm</quantity>
|
||
. The outer metatarsal pad measures
|
||
<quantity id="4CF19B15FFD28319D5D4FEDFD542FE9A" box="[715,804,258,283]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="39" pageNumber="40" unit="mm" value="6.7">6.7 mm</quantity>
|
||
and is separated by a gap of
|
||
<quantity id="4CF19B15FFD28319D399FEDED286FE9A" box="[1158,1248,259,283]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="39" pageNumber="40" unit="mm" value="2.7">2.7 mm</quantity>
|
||
from the fourth interdigital pad. The second to fourth interdigital pads are elongate and of nearly equal length.
|
||
</paragraph>
|
||
<paragraph id="8BB636F0FFD28319D7D8FE94D770FE70" blockId="39.[151,1437,151,1380]" pageId="39" pageNumber="40">
|
||
The cranium of WAM16028 is illustrated in
|
||
<figureCitation id="13322A75FFD28319D5C2FE94D537FEE1" box="[733,849,329,354]" captionStart="FIGURE 21" captionStartId="38.[151,250,1943,1965]" captionTargetBox="[151,1436,481,1920]" captionTargetId="figure@38.[151,1436,481,1922]" captionTargetPageId="38" captionText="FIGURE 21. Cranium and dentary of WAM 16028, the holotype of P. tapoatafa kimberleyensis subsp. nov., an adult male from Pago Mission, Western Australia: a) dorsal view of cranium, b) ventral view of cranium, and c) lateral view of right dentary." httpUri="https://zenodo.org/record/240174/files/figure.png" pageId="39" pageNumber="40">Figure 21</figureCitation>
|
||
. Measurements are provided in
|
||
<tableCitation id="C68B034BFFD28319D3CFFE94D34DFEE2" box="[1232,1323,329,354]" captionStart="TABLE 5" captionStartId="25.[125,203,525,545]" captionText="TABLE 5. Cranial and dental measurements of each taxon and major regional population within the P. tapoatafa group, presented separately for males (5 a) and females (5 b). Data are presented separately for three regional populations within nominotypical P. tapoatafa: 1) South Australia; 2) Victoria; 3) New South Wales and southern Queensland. Measurements of holotypes are also presented." pageId="39" pageNumber="40">Table 5</tableCitation>
|
||
. Too few specimens are available to demonstrate sexual dimorphism. However, the two adult males differ from the one adult female in ways that suggest male-biased dimorphism of the kind noted in the nomino-typical population—the male skulls are broader and more robust for their length, and have canines enlarged relative to near equal-sized cheekteeth.
|
||
</paragraph>
|
||
<paragraph id="8BB636F0FFD28319D7D8FE27D4CEFB84" blockId="39.[151,1437,151,1380]" pageId="39" pageNumber="40">
|
||
<emphasis id="B97DEAE2FFD28319D7D8FE27D7C9FD93" bold="true" box="[199,431,506,531]" pageId="39" pageNumber="40">Observed variation</emphasis>
|
||
. External measurements taken from museum specimen records are summarised in
|
||
<tableCitation id="C68B034BFFD28319D241FE27D6CEFDB6" captionStart="TABLE 4 a" captionStartId="23.[151,239,440,463]" captionTargetBox="[159,1330,564,1991]" captionTargetPageId="23" captionText="TABLE 4 a. Summary external measures of male Phascogale tapoatafa group specimens used in this study (HV headvent, TV tail-vent, Wgt weight). Obvious subadults excluded." httpUri="http://table.plazi.org/id/DF766678FFE28329D788FE65D54CFE6F" pageId="39" pageNumber="40" tableUuid="DF766678FFE28329D788FE65D54CFE6F">Table 4</tableCitation>
|
||
. The
|
||
<typeStatus id="54B28852FFD28319D7FAFDC2D72EFDB7" box="[229,328,543,567]" pageId="39" pageNumber="40" type="paratype">paratype</typeStatus>
|
||
, a subadult male, has softer, lusher fur of a more even grey colour. The tail ‘stem’ is proportionally shorter (
|
||
<quantity id="4CF19B15FFD28319D7E8FD9CD72FFDD9" box="[247,329,577,602]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="39" pageNumber="40" unit="mm" value="35.0">35 mm</quantity>
|
||
<emphasis id="B97DEAE2FFD28319D64CFD9FD70CFDD9" box="[339,362,578,601]" italics="true" pageId="39" pageNumber="40">vs</emphasis>
|
||
<quantity id="4CF19B15FFD28319D66BFD9CD7B4FDD9" box="[372,466,577,602]" metricMagnitude="-1" metricUnit="m" metricValue="1.16" pageId="39" pageNumber="40" unit="mm" value="116.0">116 mm</quantity>
|
||
for brush). WAM1309, an adult female from Kunmunya Hill, has a well developed pouch with sparse, pale gingery hairs but the number of teats cannot be determined. The dorsal fur of this specimen is overall more gingery than the
|
||
<typeStatus id="54B28852FFD28319D51CFD55D403FD21" box="[515,613,648,673]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
|
||
or
|
||
<typeStatus id="54B28852FFD28319D595FD54D48CFD21" box="[650,746,649,673]" pageId="39" pageNumber="40" type="paratype">paratype</typeStatus>
|
||
but this is probably due to fading. The ventral fur is cream to the roots along a mid-ventral zone, from the chin to around the pouch. This is flanked on each side by a band of grey-based and cream tipped fur that extends onto the inside of the fore- and hind-limbs. The preauricular patch is more distinct in this specimen. The tail ‘stem’ is proportionally shorter again, measuring
|
||
<quantity id="4CF19B15FFD28319D386FD2ED28CFC8B" box="[1177,1258,755,780]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="39" pageNumber="40" unit="mm" value="35.0">35 mm</quantity>
|
||
<emphasis id="B97DEAE2FFD28319D3ECFD29D36CFC8B" box="[1267,1290,756,779]" italics="true" pageId="39" pageNumber="40">vs</emphasis>
|
||
<quantity id="4CF19B15FFD28319D20CFD2ED314FC8B" box="[1299,1394,755,780]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" pageId="39" pageNumber="40" unit="mm" value="125.0">125 mm</quantity>
|
||
for brush). WAM53763, a damaged roadkill specimen, is an adult female with a damaged pouch; the number of teats cannot be determined. The pes measures
|
||
<quantity id="4CF19B15FFD28319D565FCE7D4AAFCD1" box="[634,716,826,850]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="39" pageNumber="40" unit="mm" value="34.0">34 mm</quantity>
|
||
and the tail
|
||
<quantity id="4CF19B15FFD28319D47AFCE7D5A0FCD1" box="[869,966,826,850]" metricMagnitude="-1" metricUnit="m" metricValue="2.07" pageId="39" pageNumber="40" unit="mm" value="207.0">207 mm</quantity>
|
||
with a basal stem of
|
||
<quantity id="4CF19B15FFD28319D3D6FCE4D378FCD1" box="[1225,1310,825,850]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="39" pageNumber="40" unit="mm" value="55.0">55 mm</quantity>
|
||
. The outer metatarsal tubercle in this specimen is very short (
|
||
<quantity id="4CF19B15FFD28319D5C9FC83D555FCF5" box="[726,819,862,886]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="39" pageNumber="40" unit="mm" value="4.7">4.7 mm</quantity>
|
||
; gap to fourth interdigital pad
|
||
<quantity id="4CF19B15FFD28319D38EFC80D284FCF6" box="[1169,1250,861,886]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="39" pageNumber="40" unit="mm" value="3.5">3.5mm</quantity>
|
||
) and the second interdigital pad is slightly longer than the third and fourth. The combined inner metatarsal tubercle and first interdigital pad are smoothly united on the left pes but distinguished by a clear kink on the left pes. Rhind
|
||
<emphasis id="B97DEAE2FFD28319D27DFC78D3FAFC3C" box="[1378,1436,932,956]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
|
||
(2001) reported a female specimen from Mitchell Plateau (WAM27208; as KIM5) with six teats; it is not known whether this represents the typical teat number.
|
||
</paragraph>
|
||
<paragraph id="8BB636F0FFD28319D7D8FBD3D235FB34" blockId="39.[151,1437,151,1380]" pageId="39" pageNumber="40">
|
||
<emphasis id="B97DEAE2FFD28319D7D8FBD3D7DEFBA7" bold="true" box="[199,440,1038,1063]" pageId="39" pageNumber="40">Taxonomic remarks</emphasis>
|
||
. This population has attracted little previous attention, although
|
||
<bibRefCitation id="EF984B01FFD28319D3BAFBD3D3FAFBA7" author="Troughton" box="[1189,1436,1038,1063]" pageId="39" pageNumber="40" refString="Troughton, E. (1967) Furred Animals of Australia. 9 th Edition. Angus and Roberston, Sydney, 376 pp." type="book" year="1967">Troughton (1967: 32)</bibRefCitation>
|
||
remarked that “the Northern Australian race (
|
||
<taxonomicName id="4C094D73FFD28319D5B1FBEFD591FBC9" box="[686,1015,1074,1098]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="39" pageNumber="40" phylum="Chordata" rank="subSpecies" species="tapoatafa" subSpecies="pirata">
|
||
<emphasis id="B97DEAE2FFD28319D5B1FBEFD591FBC9" box="[686,1015,1074,1098]" italics="true" pageId="39" pageNumber="40">Phascogale tapoatafa pirata</emphasis>
|
||
</taxonomicName>
|
||
) … is also found in the Kimberley division, Western Australia”. Rhind
|
||
<emphasis id="B97DEAE2FFD28319D527FB8BD412FBED" box="[568,628,1109,1133]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
|
||
(2001) also allied this population with
|
||
<taxonomicName id="4C094D73FFD28319D320FB8BD2DDFBED" box="[1087,1211,1110,1133]" class="Mammalia" family="Dasyuridae" genus="Phascogale" kingdom="Animalia" order="Dasyuromorphia" pageId="39" pageNumber="40" phylum="Chordata" rank="subSpecies" species="tapoatafa" subSpecies="pirata">
|
||
<emphasis id="B97DEAE2FFD28319D320FB8BD2DDFBED" box="[1087,1211,1110,1133]" italics="true" pageId="39" pageNumber="40">P. t. pirata</emphasis>
|
||
</taxonomicName>
|
||
based on the small body size, slight brownish tinge to the dorsal fur and extensive cream venters. A total of five museum specimens are known from the entire Kimberley region (see Supplementary Table II for details).
|
||
</paragraph>
|
||
<paragraph id="8BB636F0FFD28319D7D8FB63D74EFAE4" blockId="39.[151,1437,151,1380]" pageId="39" pageNumber="40">
|
||
<emphasis id="B97DEAE2FFD28319D7D8FB63D431FB57" bold="true" box="[199,599,1214,1239]" pageId="39" pageNumber="40">Former and present distribution</emphasis>
|
||
. The few records of this taxon are scattered throughout the Kimberley region, from Yampi Sound in the west to near Kalumburu (Pago Mission) in the east (
|
||
<figureCitation id="13322A75FFD28319D369FB3CD2ABFB7A" box="[1142,1229,1249,1274]" captionStart="FIGURE 22" captionStartId="39.[151,250,1952,1974]" captionTargetBox="[313,1262,1426,1925]" captionTargetId="figure@39.[312,1263,1426,1935]" captionTargetPageId="39" captionText="FIGURE 22. Map of northern Australia, showing the distributions of P. tapoatafa kimberleyensis subsp. nov (circles) and P. pirata (triangles). The square symbol represents the location of a sight record that might pertain to either P. pirata or P. tapoatafa." httpUri="https://zenodo.org/record/240175/files/figure.png" pageId="39" pageNumber="40">Fig. 22</figureCitation>
|
||
). There are recent records from Kimbolton Station near Derby (2003; WAM53763) and from the King Leopold Range (2001; sighting by Mr Murray Ellis, pers. comm.). Undated subfossil occurrences in the southwest Kimberley are reported by
|
||
<bibRefCitation id="EF984B01FFD28319D277FAF5D742FAE4" pageId="39" pageNumber="40" refString="Start, A. N., Burbidge, A. A., McDowell, M. C. & McKenzie, N. L. (2012) The status of non-volant mammals along a rainfall gradient in the south-west Kimberley, Western Australia. Australian Mammalogy, 34, 36 - 48. http: // dx. doi. org / 10.1071 / AM 10026" type="journal article">
|
||
Start
|
||
<emphasis id="B97DEAE2FFD28319D788FA91D6A9FAE3" box="[151,207,1355,1379]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
|
||
(2012)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |