324 lines
35 KiB
XML
324 lines
35 KiB
XML
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<mods:title id="4B686D40BB954F5B31B0AC25687FC501">Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography</mods:title>
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<mods:namePart id="6D48165119C7F14FBA362D4B61A1EBFE">Merle, Didier</mods:namePart>
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<mods:namePart id="66BECA62F48A4FF475890BA970C9FDA9">Bartolini, Annachiara</mods:namePart>
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<mods:namePart id="985F2E4737DC7618AA04AD0A14101FEC">Lashari, Rafiq A.</mods:namePart>
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<mods:namePart id="531B82527D8FE9EA16FD35DB29D0B6AD">Brohi, Imdad A.</mods:namePart>
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<mods:namePart id="569FF450B7C43DAA09D7DF3C085AA5DD">Solangi, Sarfraz H.</mods:namePart>
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<mods:namePart id="8362C13091B00FFA138B10C1142BE29D">Marivaux, Laurent</mods:namePart>
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<mods:title id="FDA9265A9D22194DA95DE7E0F16C6FB8">Zootaxa</mods:title>
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<mods:date id="83D399E682417F96654075AA2C788DEF">2014</mods:date>
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<mods:number id="E66E1EB88B4291F794EABE0D25A1567F">2014-06-27</mods:number>
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<mods:number id="F9EB094EDB61E2503CEF0F1D7ACC0D77">3826</mods:number>
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<mods:start id="389D5A0E5DEC9FCC40D5DD0D55748C81">101</mods:start>
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<treatment id="03A087E00E42FFBFFF7FF919FA43FB43" ID-DOI="http://doi.org/10.5281/zenodo.6137822" ID-GBIF-Taxon="119509282" ID-Zenodo-Dep="6137822" LSID="urn:lsid:plazi:treatment:03A087E00E42FFBFFF7FF919FA43FB43" httpUri="http://treatment.plazi.org/id/03A087E00E42FFBFFF7FF919FA43FB43" lastPageId="13" lastPageNumber="114" pageId="11" pageNumber="112">
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<subSubSection id="C313657D0E42FFB9FF7FF919FED4F898" pageId="11" pageNumber="112" type="nomenclature">
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<paragraph id="8BB636F60E42FFB9FF7FF919FDC1F8B9" blockId="11.[151,539,1783,1843]" box="[151,539,1783,1810]" pageId="11" pageNumber="112">
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<heading id="D0FE819A0E42FFB9FF7FF919FDC1F8B9" bold="true" box="[151,539,1783,1810]" fontSize="11" level="1" pageId="11" pageNumber="112" reason="1">
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<emphasis id="B97DEAE40E42FFB9FF7FF919FDC1F8B9" bold="true" box="[151,539,1783,1810]" pageId="11" pageNumber="112">
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<taxonomicName id="4C094D750E42FFB9FF7FF919FE6CF8BA" authority="Merle et al., 2014" box="[151,438,1783,1809]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="vredenburgi" status="sp. nov.">
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<emphasis id="B97DEAE40E42FFB9FF7FF919FE6CF8BA" bold="true" box="[151,438,1783,1809]" italics="true" pageId="11" pageNumber="112">Lyrischapa vredenburgi</emphasis>
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</taxonomicName>
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sp. nov.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB636F60E42FFB9FF7FF8F4FED4F898" blockId="11.[151,539,1783,1843]" box="[151,270,1818,1843]" pageId="11" pageNumber="112">
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<figureCitation id="13322A730E42FFB9FF7FF8F4FED4F898" box="[151,270,1818,1843]" captionStart="FIGURE 5. A – H" captionStartId="12.[151,250,1914,1936]" captionTargetBox="[224,1371,193,1879]" captionTargetId="figure@12.[224,1372,193,1881]" captionTargetPageId="12" captionText="FIGURE 5. A – H. Lyrischapa vredenburgi sp. nov., A – B (paratype 1, CPAG. RAN. I. 12, protoconch and first teleoconch whorl in ventral (A) and dorsal (B) views, stn 4), C – E (holotype, CPAG. RAN. I. 11, ventral (C), dorsal (D) and apical (E) views, stn 4), F (paratype 2, CPAG. RAN. I. 13, ventral view, stn 4), G (paratype 3, CPAG. RAN. I. 14, ventral view, stn 4), H (paratype MNHN. F. J 12751, Cossmann coll., Jhirak). Scale bars: A – H = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.228542" httpUri="https://zenodo.org/record/228542/files/figure.png" pageId="11" pageNumber="112">Fig. 5A–H</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C313657D0E42FFB9FF7FF8B3FB45F864" pageId="11" pageNumber="112" type="reference_group">
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<paragraph id="8BB636F60E42FFB9FF7FF8B3FDD6F81B" blockId="11.[151,1436,1885,1999]" pageId="11" pageNumber="112">
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||
<taxonomicName id="4C094D750E42FFB9FF7FF8B3FEE2F8D8" box="[151,312,1885,1907]" class="Gastropoda" family="Volutidae" genus="Aulicina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="haimei">
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<emphasis id="B97DEAE40E42FFB9FF7FF8B3FEE2F8D8" box="[151,312,1885,1907]" italics="true" pageId="11" pageNumber="112">Aulicina haimei</emphasis>
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</taxonomicName>
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(non
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<taxonomicName id="4C094D750E42FFB9FE9BF8B3FCCBF8D8" authority="d'Archiac & Haime, 1853" authorityName="d'Archiac & Haime" authorityYear="1853" box="[371,785,1885,1907]" class="Gastropoda" family="Volutidae" genus="Voluta" kingdom="Animalia" order="Neogastropoda" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="haimei">
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<emphasis id="B97DEAE40E42FFB9FE9BF8B3FE25F8D8" box="[371,511,1885,1907]" italics="true" pageId="11" pageNumber="112">Voluta haimei</emphasis>
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d’Archiac & Haime, 1853
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</taxonomicName>
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).
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<bibRefCitation id="EF984B070E42FFB9FCCAF8B3FB85F8D8" author="Cossmann" box="[802,1119,1885,1907]" firstAuthor="Cossmann" page="25" pageId="11" pageNumber="112" refString="Cossmann, M. & Pissarro, G. (1909) The Mollusca of the Ranikot Series. Part 1, Cephalopoda and Gastropoda. Memoirs of the Geological Survey of India, Palaeontologia Indica, New Series, 3 (1), 1 - 83." type="journal article" year="1909">Cossmann & Pissarro 1909: 25</bibRefCitation>
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, pl. 2, fig. 30, pl. 8, fig. 2.
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<taxonomicName id="4C094D750E42FFB9FF7FF892FDD3F839" authority="Vredenburg, 1923" authorityName="Vredenburg" authorityYear="1923" box="[151,521,1916,1938]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="blanfordi">
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<emphasis id="B97DEAE40E42FFB9FF7FF892FE9CF839" box="[151,326,1916,1938]" italics="true" pageId="11" pageNumber="112">Athleta blanfordi</emphasis>
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<bibRefCitation id="EF984B070E42FFB9FEA7F892FDD3F839" author="Vredenburg" box="[335,521,1916,1938]" firstAuthor="Vredenburg" pageId="11" pageNumber="112" refString="Vredenburg, E. W. (1923) Indian Tertiary Gastropoda, IV. Olividae, Harpidae, Marginellidae, Volutidae and Mitridae, with comparative diagnosis of new species. Records of the Geological Survey of India, 54, 243 - 276." type="journal article" year="1923">Vredenburg, 1923</bibRefCitation>
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</taxonomicName>
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pars: 255, proposed for
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<taxonomicName id="4C094D750E42FFB9FCE6F892FE80F81B" authority="d'Archiac & Haime, 1853 sensu Cossmann & Pissarro, 1909" authorityName="d'Archiac & Haime, 1853 sensu Cossmann & Pissarro" authorityYear="1909" class="Gastropoda" family="Volutidae" genus="Aulicina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="haimei">
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<emphasis id="B97DEAE40E42FFB9FCE6F892FC6BF839" box="[782,945,1916,1938]" italics="true" pageId="11" pageNumber="112">Aulicina haimei</emphasis>
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d’Archiac & Haime, 1853 sensu
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<bibRefCitation id="EF984B070E42FFB9FAFDF892FE80F81B" author="Cossmann & Pissarro" firstAuthor="Cossmann" pageId="11" pageNumber="112" refString="Cossmann, M. & Pissarro, G. (1909) The Mollusca of the Ranikot Series. Part 1, Cephalopoda and Gastropoda. Memoirs of the Geological Survey of India, Palaeontologia Indica, New Series, 3 (1), 1 - 83." type="journal article" year="1909">Cossmann & Pissarro, 1909</bibRefCitation>
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</taxonomicName>
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, pl. 8, fig. 2 only.
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</paragraph>
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<paragraph id="8BB636F60E42FFB9FF7FF857FB45F864" blockId="11.[151,1436,1885,1999]" box="[151,1183,1977,1999]" pageId="11" pageNumber="112">
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<taxonomicName id="4C094D750E42FFB9FF7FF857FEAEF864" box="[151,372,1977,1999]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="sismondai">
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<emphasis id="B97DEAE40E42FFB9FF7FF857FEAEF864" box="[151,372,1977,1999]" italics="true" pageId="11" pageNumber="112">Lyrischapa sismondai</emphasis>
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</taxonomicName>
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(non
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<taxonomicName id="4C094D750E42FFB9FE58F857FC94F864" authority="d'Archiac & Haime, 1853" authorityName="d'Archiac & Haime" authorityYear="1853" box="[432,846,1977,1999]" class="Gastropoda" family="Volutidae" genus="Voluta" kingdom="Animalia" order="Neogastropoda" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="haimei">
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<emphasis id="B97DEAE40E42FFB9FE58F857FDE6F864" box="[432,572,1977,1999]" italics="true" pageId="11" pageNumber="112">Voluta haimei</emphasis>
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d’Archiac & Haime, 1853
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</taxonomicName>
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).
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<bibRefCitation id="EF984B070E42FFB9FC88F857FC39F864" author="Givens" box="[864,995,1977,1999]" firstAuthor="Givens" pageId="11" pageNumber="112" refString="Givens, C. R. (1991) Old world Tethyan occurrences of Lyrischapa (Gastropoda: Volutidae) and biogeographic implications. Journal of Paleontology, 65, 661 - 670." type="journal article" year="1991">Givens 1991</bibRefCitation>
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, 666, figs 4.3, 4.6.
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||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF76667E0E45FFBEFF7FF894FC87F840" ID-DOI="http://doi.org/10.5281/zenodo.228542" httpUri="https://zenodo.org/record/228542/files/figure.png" pageId="12" pageNumber="113" targetBox="[224,1371,193,1879]" targetPageId="12">
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||
<paragraph id="8BB636F60E45FFBEFF7FF894FC87F840" blockId="12.[151,1436,1914,2027]" pageId="12" pageNumber="113">
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||
<emphasis id="B97DEAE40E45FFBEFF7FF894FE89F83B" bold="true" box="[151,339,1914,1936]" pageId="12" pageNumber="113">FIGURE 5. A–H.</emphasis>
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<taxonomicName id="4C094D750E45FFBEFEB1F894FD90F83B" box="[345,586,1914,1936]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="113" phylum="Mollusca" rank="species" species="vredenburgi" status="sp. nov.">
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<emphasis id="B97DEAE40E45FFBEFEB1F894FD90F83B" box="[345,586,1914,1936]" italics="true" pageId="12" pageNumber="113">Lyrischapa vredenburgi</emphasis>
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</taxonomicName>
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<emphasis id="B97DEAE40E45FFBEFDB9F894FD7BF83B" bold="true" box="[593,673,1914,1936]" pageId="12" pageNumber="113">sp. nov.</emphasis>
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, A–B (paratype 1, CPAG.RAN.I.12, protoconch and first teleoconch whorl in ventral (A) and dorsal (B) views, stn 4), C–E (holotype, CPAG.RAN.I.11, ventral (C), dorsal (D) and apical (E) views, stn 4), F (paratype 2, CPAG.RAN.I.13, ventral view, stn 4), G (paratype 3, CPAG.RAN.I.14, ventral view, stn 4), H (paratype MNHN.F.
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<accessionNumber id="945AAB150E45FFBEFF17F83BFE92F840" box="[255,328,2005,2027]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J12751" pageId="12" pageNumber="113" type="EnaNcbi">J12751</accessionNumber>
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, Cossmann coll., Jhirak). Scale bars: A–H = 10 mm.
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</paragraph>
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</caption>
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<subSubSection id="C313657D0E44FFBFFF7FFF79FA5CFF1B" box="[151,1414,151,176]" pageId="13" pageNumber="114" type="etymology">
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<paragraph id="8BB636F60E44FFBFFF7FFF79FA5CFF1B" blockId="13.[151,1437,151,1257]" box="[151,1414,151,176]" pageId="13" pageNumber="114">
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<emphasis id="B97DEAE40E44FFBFFF7FFF79FEC4FF1B" bold="true" box="[151,286,151,176]" pageId="13" pageNumber="114">Etymology.</emphasis>
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Dedicated to E. W. Vredenburg for his work on the geology and paleontology of the Ranikot Group.
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="C313657D0E44FFBFFF2FFF52FC46FECF" pageId="13" pageNumber="114" type="materials_examined">
|
||
<paragraph id="8BB636F60E44FFBFFF2FFF52FB6DFF7E" blockId="13.[151,1437,151,1257]" box="[199,1207,188,213]" pageId="13" pageNumber="114">
|
||
<emphasis id="B97DEAE40E44FFBFFF2FFF52FEBFFF7E" bold="true" box="[199,357,188,213]" pageId="13" pageNumber="114">
|
||
<typeStatus id="54B288540E44FFBFFF2FFF52FED8FF7E" box="[199,258,188,213]" pageId="13" pageNumber="114">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Stn 4: Lakhra Dome, Lakhra village section, base of the Lakhra Formation.
|
||
</paragraph>
|
||
<paragraph id="8BB636F60E44FFBFFF2FFF31FAD2FEEB" blockId="13.[151,1437,151,1257]" pageId="13" pageNumber="114">
|
||
<emphasis id="B97DEAE40E44FFBFFF2FFF31FEAEFF53" bold="true" box="[199,372,223,248]" pageId="13" pageNumber="114">
|
||
<typeStatus id="54B288540E44FFBFFF2FFF31FED8FF53" box="[199,258,223,248]" pageId="13" pageNumber="114">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
<typeStatus id="54B288540E44FFBFFE93FF31FE39FF53" box="[379,483,223,248]" pageId="13" pageNumber="114" type="holotype">Holotype</typeStatus>
|
||
(stn 4:
|
||
<collectionCode id="ED18AE330E44FFBFFDDFFF0EFD52FF53" box="[567,648,224,248]" pageId="13" pageNumber="114">CPAG</collectionCode>
|
||
.RAN.
|
||
<date id="FFB710360E44FFBFFD26FF0EFD29FF53" box="[718,755,224,248]" pageId="13" pageNumber="114" value="2011-01">I.11</date>
|
||
, cast
|
||
<collectionCode id="ED18AE330E44FFBFFCD8FF31FC51FF53" LSID="urn:lsid:biocol.org:col:34988" box="[816,907,223,248]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="114">MNHN</collectionCode>
|
||
.F.
|
||
<accessionNumber id="945AAB150E44FFBFFC49FF31FC25FF53" box="[929,1023,223,248]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A50349" pageId="13" pageNumber="114" type="EnaNcbi">A50349</accessionNumber>
|
||
),
|
||
<typeStatus id="54B288540E44FFBFFBFAFF0EFBA8FF53" box="[1042,1138,224,248]" pageId="13" pageNumber="114" type="paratype">paratype</typeStatus>
|
||
1 (stn 4:
|
||
<collectionCode id="ED18AE330E44FFBFFB33FF0EFAF1FF53" box="[1243,1323,224,248]" pageId="13" pageNumber="114">CPAG</collectionCode>
|
||
.RAN.
|
||
<date id="FFB710360E44FFBFFA99FF0EFA42FF53" box="[1393,1432,224,248]" pageId="13" pageNumber="114" value="2012-01">I.12</date>
|
||
, cast
|
||
<collectionCode id="ED18AE330E44FFBFFF3DFEEAFEEAFEB6" LSID="urn:lsid:biocol.org:col:34988" box="[213,304,260,285]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="114">MNHN</collectionCode>
|
||
.F.A50350),
|
||
<typeStatus id="54B288540E44FFBFFE2AFEEBFDF8FEB6" box="[450,546,261,285]" pageId="13" pageNumber="114" type="paratype">paratype</typeStatus>
|
||
2 (stn 4:
|
||
<collectionCode id="ED18AE330E44FFBFFD51FEEAFCD0FEB7" box="[697,778,260,284]" pageId="13" pageNumber="114">CPAG</collectionCode>
|
||
.RAN.
|
||
<date id="FFB710360E44FFBFFCB8FEEAFCACFEB7" box="[848,886,260,284]" pageId="13" pageNumber="114" value="2013-01">I.13</date>
|
||
, cast
|
||
<collectionCode id="ED18AE330E44FFBFFC22FEEAFBFCFEB6" LSID="urn:lsid:biocol.org:col:34988" box="[970,1062,260,285]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="114">MNHN</collectionCode>
|
||
.F.
|
||
<accessionNumber id="945AAB150E44FFBFFBD4FEEAFB43FEB6" box="[1084,1177,260,285]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A50351" pageId="13" pageNumber="114" type="EnaNcbi">A50351</accessionNumber>
|
||
),
|
||
<typeStatus id="54B288540E44FFBFFB50FEEBFAC2FEB6" box="[1208,1304,261,285]" pageId="13" pageNumber="114" type="paratype">paratype</typeStatus>
|
||
3 (stn 4:
|
||
<collectionCode id="ED18AE330E44FFBFFF7FFEC6FF3DFEEB" box="[151,231,296,320]" pageId="13" pageNumber="114">CPAG</collectionCode>
|
||
.RAN.
|
||
<date id="FFB710360E44FFBFFEC5FEC6FE8EFEEB" box="[301,340,296,320]" pageId="13" pageNumber="114" value="1914-01">I.14</date>
|
||
, cast
|
||
<collectionCode id="ED18AE330E44FFBFFE7AFEC9FE37FEEB" LSID="urn:lsid:biocol.org:col:34988" box="[402,493,295,320]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="114">MNHN</collectionCode>
|
||
.F.A50352), three
|
||
<typeStatus id="54B288540E44FFBFFD5BFEC6FCC4FEEB" box="[691,798,296,320]" pageId="13" pageNumber="114" type="paratype">paratypes</typeStatus>
|
||
(Jhirak, Cossmann coll.
|
||
<collectionCode id="ED18AE330E44FFBFFBDDFEC9FB48FEEB" LSID="urn:lsid:biocol.org:col:34988" box="[1077,1170,295,320]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="114">MNHN</collectionCode>
|
||
.F.
|
||
<accessionNumber id="945AAB150E44FFBFFB42FEC9FB26FEEB" box="[1194,1276,295,320]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J12751" pageId="13" pageNumber="114" type="EnaNcbi">J12751</accessionNumber>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB636F60E44FFBFFF2FFEA2FC46FECF" blockId="13.[151,1437,151,1257]" box="[199,924,332,357]" pageId="13" pageNumber="114">
|
||
<emphasis id="B97DEAE40E44FFBFFF2FFEA2FE5BFECE" bold="true" box="[199,385,332,357]" pageId="13" pageNumber="114">Other material.</emphasis>
|
||
16 spm (stn 4:
|
||
<collectionCode id="ED18AE330E44FFBFFDC7FEA2FD50FECF" LSID="urn:lsid:biocol.org:col:34988" box="[559,650,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="114">MNHN</collectionCode>
|
||
); 3 spm (stn 5:
|
||
<collectionCode id="ED18AE330E44FFBFFCDDFEA2FC4AFECF" LSID="urn:lsid:biocol.org:col:34988" box="[821,912,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="114">MNHN</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C313657D0E44FFBFFF2FFE81FA43FB43" pageId="13" pageNumber="114" type="description">
|
||
<paragraph id="8BB636F60E44FFBFFF2FFE81FC10FCBE" blockId="13.[151,1437,151,1257]" pageId="13" pageNumber="114">
|
||
<emphasis id="B97DEAE40E44FFBFFF2FFE81FE83FE23" bold="true" box="[199,345,367,392]" pageId="13" pageNumber="114">Description.</emphasis>
|
||
Shell biconical, wide, H 77–85, D
|
||
<quantity id="4CF19B130E44FFBFFCFAFE81FC48FE2C" box="[786,914,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="5.95" metricValueMax="6.5" metricValueMin="5.4" pageId="13" pageNumber="114" unit="mm" value="59.5" valueMax="65.0" valueMin="54.0">54–65 mm</quantity>
|
||
(
|
||
<typeStatus id="54B288540E44FFBFFC4FFE81FBD1FE23" box="[935,1035,367,392]" pageId="13" pageNumber="114" type="holotype">holotype</typeStatus>
|
||
(subadult) H 38.3, D
|
||
<quantity id="4CF19B130E44FFBFFAC9FE9EFA4BFE23" box="[1313,1425,368,392]" metricMagnitude="-2" metricUnit="m" metricValue="3.11" pageId="13" pageNumber="114" unit="mm" value="31.1">31.1 mm</quantity>
|
||
). Protoconch smooth, globose, slightly deviated, of two whorls; H 16, D
|
||
<quantity id="4CF19B130E44FFBFFC05FE7AFB9AFE07" box="[1005,1088,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="114" unit="mm" value="11.0">11 mm</quantity>
|
||
when complete (
|
||
<figureCitation id="13322A730E44FFBFFAF8FE7AFA4AFE06" box="[1296,1424,404,429]" captionStart="FIGURE 5. A – H" captionStartId="12.[151,250,1914,1936]" captionTargetBox="[224,1371,193,1879]" captionTargetId="figure@12.[224,1372,193,1881]" captionTargetPageId="12" captionText="FIGURE 5. A – H. Lyrischapa vredenburgi sp. nov., A – B (paratype 1, CPAG. RAN. I. 12, protoconch and first teleoconch whorl in ventral (A) and dorsal (B) views, stn 4), C – E (holotype, CPAG. RAN. I. 11, ventral (C), dorsal (D) and apical (E) views, stn 4), F (paratype 2, CPAG. RAN. I. 13, ventral view, stn 4), G (paratype 3, CPAG. RAN. I. 14, ventral view, stn 4), H (paratype MNHN. F. J 12751, Cossmann coll., Jhirak). Scale bars: A – H = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.228542" httpUri="https://zenodo.org/record/228542/files/figure.png" pageId="13" pageNumber="114">Fig. 5A–B</figureCitation>
|
||
). Transition from protoconch to teleoconch not defined. Teleoconch of 3 to 3.5 whorls. Spire low, occupying 10% of total shell height. Spiral whorls almost flat, separated by shallow suture undulating between shoulder spines of preceding whorl. Last whorl rather triangular. Spiral sculpture on spire of obsolete threads, visible only on first teleoconch whorl. On juvenile specimens of one whorl, last whorl sculptured with very fine spiral threads covering three abapical quarters of shell, threads disappearing on succeeding whorls. Axial sculpture of spiny shoulder costae only. First whorl: 11 costae, becoming more prominent on second half of whorl. Second whorl: 9–10 costae; third whorl: 9–12 costae. No shoulder sinus. Aperture narrow, occupying 87% of total height, 28% of diameter. Inner lip straight, columella with five strong folds succeeded posteriorly by one weaker fold; inner lip callus narrow. Outer lip thin, not thickened externally. Siphonal canal short; siphonal notch weak; siphonal fasciole low, weak, but delineated by carina spreading dorsally on gerontic specimens.
|
||
</paragraph>
|
||
<paragraph id="8BB636F60E44FFBFFF2FFCF1FEE3FB6F" blockId="13.[151,1437,151,1257]" pageId="13" pageNumber="114">
|
||
<emphasis id="B97DEAE40E44FFBFFF2FFCF1FEB6FC93" bold="true" box="[199,364,799,824]" pageId="13" pageNumber="114">Comparisons.</emphasis>
|
||
From the Lakhra Formation, this species can be compared with
|
||
<taxonomicName id="4C094D750E44FFBFFB4FFCF1FEACFCF7" authority="Vredenburg, 1923" authorityName="Vredenburg" authorityYear="1923" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="blanfordi">
|
||
<emphasis id="B97DEAE40E44FFBFFB4FFCF1FA46FC93" box="[1191,1436,799,824]" italics="true" pageId="13" pageNumber="114">Lyrischapa blanfordi</emphasis>
|
||
(
|
||
<bibRefCitation id="EF984B070E44FFBFFF76FCAAFEB4FCF7" author="Vredenburg" box="[158,366,836,861]" firstAuthor="Vredenburg" pageId="13" pageNumber="114" refString="Vredenburg, E. W. (1923) Indian Tertiary Gastropoda, IV. Olividae, Harpidae, Marginellidae, Volutidae and Mitridae, with comparative diagnosis of new species. Records of the Geological Survey of India, 54, 243 - 276." type="journal article" year="1923">Vredenburg, 1923</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Both species share obsolete threads on the first teleoconch whorl and the disappearance of the spiral sculpture on the last few whorls.
|
||
<collectingCountry id="F31E76660E44FFBFFD8EFC86FD5FFC2B" box="[614,645,872,896]" name="American Samoa" pageId="13" pageNumber="114">As</collectingCountry>
|
||
in
|
||
<taxonomicName id="4C094D750E44FFBFFD47FC87FC8DFC2B" box="[687,855,871,896]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="vredenburgi">
|
||
<emphasis id="B97DEAE40E44FFBFFD47FC87FC8DFC2B" box="[687,855,871,896]" italics="true" pageId="13" pageNumber="114">L. vredenburgi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C094D750E44FFBFFC80FC87FC2AFC2B" box="[872,1008,871,896]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="blanfordi">
|
||
<emphasis id="B97DEAE40E44FFBFFC80FC87FC2AFC2B" box="[872,1008,871,896]" italics="true" pageId="13" pageNumber="114">L. blanfordi</emphasis>
|
||
</taxonomicName>
|
||
exhibits shoulder spines and lacks a shoulder sinus.
|
||
<taxonomicName id="4C094D750E44FFBFFEAEFC63FE11FC0E" box="[326,459,908,933]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="blanfordi">
|
||
<emphasis id="B97DEAE40E44FFBFFEAEFC63FE11FC0E" box="[326,459,908,933]" italics="true" pageId="13" pageNumber="114">L. blanfordi</emphasis>
|
||
</taxonomicName>
|
||
differs in its less numerous costae on the last whorl (8–9 instead of 9–12), its shoulder spines oriented adapically and its narrower shape. The number of columellar folds varies from 5 to 6, whereas those of
|
||
<taxonomicName id="4C094D750E44FFBFFF14FC3BFE59FC46" box="[252,387,980,1005]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="blanfordi">
|
||
<emphasis id="B97DEAE40E44FFBFFF14FC3BFE59FC46" box="[252,387,980,1005]" italics="true" pageId="13" pageNumber="114">L. blanfordi</emphasis>
|
||
</taxonomicName>
|
||
vary from 6 to 8.
|
||
<taxonomicName id="4C094D750E44FFBFFDB5FC3BFCDEFC46" box="[605,772,980,1005]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="vredenburgi">
|
||
<emphasis id="B97DEAE40E44FFBFFDB5FC3BFCDEFC46" box="[605,772,980,1005]" italics="true" pageId="13" pageNumber="114">L. vredenburgi</emphasis>
|
||
</taxonomicName>
|
||
also has a strong similarity to
|
||
<taxonomicName id="4C094D750E44FFBFFB84FC3BFA46FC47" authority="Douville, 1929" authorityName="Douville" authorityYear="1929" box="[1132,1436,980,1005]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="crassa">
|
||
<emphasis id="B97DEAE40E44FFBFFB84FC3BFB0FFC47" box="[1132,1237,981,1004]" italics="true" pageId="13" pageNumber="114">L. crassa</emphasis>
|
||
(
|
||
<bibRefCitation id="EF984B070E44FFBFFB0EFC3AFA4EFC47" author="Douville" box="[1254,1428,980,1005]" firstAuthor="Douville" pageId="13" pageNumber="114" refString="Douville, H. (1929) Les couches a Cardita beaumonti dans le Sind. Memoirs of the Geological Survey of India, Palaeontologia Indica, New Series, 10 (3), 27 - 73." type="journal article" year="1929">Douvillé, 1929</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
described from the Danian of the Kadhro Formation, at Jakhmari and at Ranikot. The two species share a wide biconic shape, a low spire, 12 costae on the last whorl and the presence of shoulder spines. The
|
||
<typeStatus id="54B288540E44FFBFFB29FBF3FB2BFB9E" box="[1217,1265,1053,1077]" pageId="13" pageNumber="114">type</typeStatus>
|
||
specimen of
|
||
<taxonomicName id="4C094D750E44FFBFFA6DFBF3FF3AFBF3" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="crassa">
|
||
<emphasis id="B97DEAE40E44FFBFFA6DFBF3FF3AFBF3" italics="true" pageId="13" pageNumber="114">L. crassa</emphasis>
|
||
</taxonomicName>
|
||
is poorly preserved, but it apparently differs from
|
||
<taxonomicName id="4C094D750E44FFBFFCF6FBAEFC1FFBF3" box="[798,965,1087,1112]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="vredenburgi">
|
||
<emphasis id="B97DEAE40E44FFBFFCF6FBAEFC1FFBF3" box="[798,965,1087,1112]" italics="true" pageId="13" pageNumber="114">A. vredenburgi</emphasis>
|
||
</taxonomicName>
|
||
by having only four columellar folds and more rounded shoulder spines.
|
||
<taxonomicName id="4C094D750E44FFBFFDE1FB8BFDAFFBD7" box="[521,629,1125,1148]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="crassa">
|
||
<emphasis id="B97DEAE40E44FFBFFDE1FB8BFDAFFBD7" box="[521,629,1125,1148]" italics="true" pageId="13" pageNumber="114">L. crassa</emphasis>
|
||
</taxonomicName>
|
||
was originally assigned to
|
||
<taxonomicName id="4C094D750E44FFBFFC57FB8AFBB2FBD6" box="[959,1128,1124,1149]" class="Gastropoda" genus="Diconomorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEAE40E44FFBFFC57FB8AFBB2FBD6" box="[959,1128,1124,1149]" italics="true" pageId="13" pageNumber="114">Diconomorpha</emphasis>
|
||
</taxonomicName>
|
||
, but the close similarities between
|
||
<taxonomicName id="4C094D750E44FFBFFF12FB66FEB8FB0B" box="[250,354,1160,1184]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="crassa">
|
||
<emphasis id="B97DEAE40E44FFBFFF12FB66FEB8FB0B" box="[250,354,1160,1184]" italics="true" pageId="13" pageNumber="114">A. crassa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C094D750E44FFBFFE7FFB66FDE6FB0B" box="[407,572,1159,1184]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="vredenburgi">
|
||
<emphasis id="B97DEAE40E44FFBFFE7FFB66FDE6FB0B" box="[407,572,1159,1184]" italics="true" pageId="13" pageNumber="114">A. vredenburgi</emphasis>
|
||
</taxonomicName>
|
||
constitute a further argument to consider
|
||
<taxonomicName id="4C094D750E44FFBFFBE4FB69FB6CFB0B" box="[1036,1206,1159,1184]" class="Gastropoda" genus="Diconomorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEAE40E44FFBFFBE4FB69FB6CFB0B" box="[1036,1206,1159,1184]" italics="true" pageId="13" pageNumber="114">Diconomorpha</emphasis>
|
||
</taxonomicName>
|
||
as a junior synonym of
|
||
<taxonomicName id="4C094D750E44FFBFFF5DFB42FEE8FB6E" box="[181,306,1196,1221]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEAE40E44FFBFFF5DFB42FEE8FB6E" box="[181,306,1196,1221]" italics="true" pageId="13" pageNumber="114">Lyrischapa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB636F60E44FFBFFF2FFB21FA43FB43" blockId="13.[151,1437,151,1257]" box="[199,1433,1231,1257]" pageId="13" pageNumber="114">
|
||
<emphasis id="B97DEAE40E44FFBFFF2FFB21FE62FB43" bold="true" box="[199,440,1231,1256]" pageId="13" pageNumber="114">Stratigraphic range.</emphasis>
|
||
Lakhra Formation at Lakhra and Jhirak; Nammal Formation, Salt Range (
|
||
<bibRefCitation id="EF984B070E44FFBFFB10FB21FA56FB43" author="Givens" box="[1272,1420,1231,1256]" firstAuthor="Givens" pageId="13" pageNumber="114" refString="Givens, C. R. (1991) Old world Tethyan occurrences of Lyrischapa (Gastropoda: Volutidae) and biogeographic implications. Journal of Paleontology, 65, 661 - 670." type="journal article" year="1991">Givens 1991</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |