634 lines
74 KiB
XML
634 lines
74 KiB
XML
<document id="08200BEFD9637D882533C1E37A287365" ID-DOI="10.11646/zootaxa.3616.2.3" ID-GBIF-Dataset="7d107b23-120b-40e1-b213-78ad6bdaedbf" ID-ISSN="1175-5326" ID-Zenodo-Dep="220758" ID-ZooBank="D6547090-2354-428C-B7EA-59C8B3795394" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460575715977" checkinUser="plazi" docAuthor="Larson, Helen K., Foster, Ralph, Humphreys, William F. & Stevens, Mark I." docDate="2013" docId="03A087D5FFB2D96DF7C36A7A0871FE76" docLanguage="en" docName="zt03616p150.pdf" docOrigin="Zootaxa 3616 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Milyeringa veritas Whitley 1945" docType="treatment" docVersion="12" lastPageNumber="149" masterDocId="FF99FFADFFBBD963F7546E2B0C1CFFF1" masterDocTitle="A new species of the blind cave gudgeon Milyeringa (Pisces: Gobioidei, Eleotridae) from Barrow Island, Western Australia, with a redescription of M. veritas Whitley" masterLastPageNumber="150" masterPageNumber="135" pageNumber="144" updateTime="1698323070799" updateUser="plazi">
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<mods:titleInfo id="2993862EB6CA3BE7F9AEE59924A457A9">
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<mods:title id="E533793F60B1DE679490983AA21A748D">A new species of the blind cave gudgeon Milyeringa (Pisces: Gobioidei, Eleotridae) from Barrow Island, Western Australia, with a redescription of M. veritas Whitley</mods:title>
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</mods:titleInfo>
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<mods:name id="E32B49B08ABD925963D794E6BE6F87C5" type="personal">
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<mods:roleTerm id="67BBCB7801CE94D64E14B5D739F2F0F7">Author</mods:roleTerm>
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<mods:namePart id="D4118BF3E8FCA4DB35149D20685C0AAF">Larson, Helen K.</mods:namePart>
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</mods:name>
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<mods:roleTerm id="4F237B1E32EB0BF993CB1F7DD22B44A4">Author</mods:roleTerm>
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<mods:namePart id="29CE326E8D0D09502C692DFA595A4733">Foster, Ralph</mods:namePart>
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</mods:role>
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<mods:namePart id="0408A428F5F07C16F220FC85E0CB31DA">Humphreys, William F.</mods:namePart>
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<mods:namePart id="2F9F9F45B92AF1D13784C00208BCA648">Stevens, Mark I.</mods:namePart>
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<mods:typeOfResource id="D94FD5923E73EB84F26861E9D3A14801">text</mods:typeOfResource>
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<mods:title id="62CD81A83162DFD7695E8D63F2DF7486">Zootaxa</mods:title>
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<mods:date id="2077545320C61C949F5DDB52F9613F48">2013</mods:date>
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<mods:detail id="37A46457A1D1F2D96B01404FB9130E13" type="volume">
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<mods:number id="8C415308F40E119896A53081BED1EA80">3616</mods:number>
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<mods:classification id="C7A9E86D98EFC10DFC5B6A05CBAB9718">journal article</mods:classification>
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<mods:identifier id="683333E613776DAE62BBA2D43433324C" type="DOI">10.11646/zootaxa.3616.2.3</mods:identifier>
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<mods:identifier id="DA2281918DAFF3FBF81781E92B8BE183" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D607EC37424A6BF795366C0665D284E2" type="Zenodo-Dep">220758</mods:identifier>
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<mods:identifier id="C08A3599222B54F7102194E2E959BAA5" type="ZooBank">D6547090-2354-428C-B7EA-59C8B3795394</mods:identifier>
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<treatment id="03A087D5FFB2D96DF7C36A7A0871FE76" ID-DOI="http://doi.org/10.5281/zenodo.5613148" ID-GBIF-Taxon="119560048" ID-Zenodo-Dep="5613148" LSID="urn:lsid:plazi:treatment:03A087D5FFB2D96DF7C36A7A0871FE76" httpUri="http://treatment.plazi.org/id/03A087D5FFB2D96DF7C36A7A0871FE76" lastPageId="14" lastPageNumber="149" pageId="9" pageNumber="144">
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<subSubSection id="C3136548FFB2D96AF7C36A7A0D90FB01" pageId="9" pageNumber="144" type="nomenclature">
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<paragraph id="8BB636C3FFB2D96AF7C36A7A0E39FB9B" blockId="9.[151,549,1104,1164]" box="[151,549,1104,1131]" pageId="9" pageNumber="144">
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<heading id="D0FE81AFFFB2D96AF7C36A7A0E39FB9B" bold="true" box="[151,549,1104,1131]" fontSize="11" level="1" pageId="9" pageNumber="144" reason="1">
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<taxonomicName id="4C094D40FFB2D96AF7C36A7A0E39FB9B" ID-CoL="43DZM" authority="Whitley, 1945" authorityName="Whitley" authorityYear="1945" box="[151,549,1104,1131]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="veritas">
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<emphasis id="B97DEAD1FFB2D96AF7C36A7A0E39FB9B" bold="true" box="[151,549,1104,1131]" pageId="9" pageNumber="144">
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<emphasis id="B97DEAD1FFB2D96AF7C36A7A0D6FFB9A" bold="true" box="[151,371,1105,1131]" italics="true" pageId="9" pageNumber="144">Milyeringa veritas</emphasis>
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Whitley, 1945
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BB636C3FFB2D96AF7C36A5F0D2BFB7D" blockId="9.[151,549,1104,1164]" box="[151,311,1140,1164]" pageId="9" pageNumber="144">Cave gudgeon</paragraph>
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<paragraph id="8BB636C3FFB2D96AF7C36A970D90FB01" blockId="9.[151,1436,1211,1296]" pageId="9" pageNumber="144">
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<taxonomicName id="4C094D40FFB2D96AF7C36A970E3FFB20" authority="Whitley, 1945: 36" authorityName="Whitley" authorityPageNumber="36" authorityYear="1945" box="[151,547,1211,1234]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="veritas">
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<emphasis id="B97DEAD1FFB2D96AF7C36A970D44FB23" box="[151,344,1212,1234]" italics="true" pageId="9" pageNumber="144">Milyeringa veritas</emphasis>
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Whitley, 1945: 36
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</taxonomicName>
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–37 (Milyering, Yardie,
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<quantity id="4CF19B26FFB2D96AF47E6A970F9AFB20" box="[810,902,1211,1233]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="9" pageNumber="144" unit="mi" value="20.0">20 miles</quantity>
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south-west of Vlamingh Head, North-west Cape, Western
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<collectingCountry id="F31E7653FFB2D96AF64B6AF10D9DFB01" box="[287,385,1242,1264]" name="Australia" pageId="9" pageNumber="144">Australia</collectingCountry>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3136548FFB2D96AF7C36AD20FF2FAE1" box="[151,1006,1273,1296]" pageId="9" pageNumber="144" type="description">
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<paragraph id="8BB636C3FFB2D96AF7C36AD20FF2FAE1" blockId="9.[151,1436,1211,1296]" box="[151,1006,1273,1296]" pageId="9" pageNumber="144">
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<taxonomicName id="4C094D40FFB2D96AF7C36AD20E5CFAFE" authority="Chakrabarty, 2010: 22" authorityName="Chakrabarty" authorityPageNumber="22" authorityYear="2010" box="[151,576,1273,1295]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="brooksi">
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<emphasis id="B97DEAD1FFB2D96AF7C36AD20D44FAFE" box="[151,344,1273,1295]" italics="true" pageId="9" pageNumber="144">Milyeringa brooksi</emphasis>
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Chakrabarty, 2010: 22
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</taxonomicName>
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–25 (Pilgonaman Well, North West Cape).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3136548FFB2D96AF7C36B1008ADF9F6" pageId="9" pageNumber="144" type="diagnosis">
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<paragraph id="8BB636C3FFB2D96AF7C36B1008ADF9F6" blockId="9.[151,1437,1339,1977]" pageId="9" pageNumber="144">
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<emphasis id="B97DEAD1FFB2D96AF7C36B100D0DFAA5" bold="true" box="[151,273,1339,1364]" pageId="9" pageNumber="144">Diagnosis.</emphasis>
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A
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<taxonomicName id="4C094D40FFB2D96AF6626B170DAEFAA5" box="[310,434,1340,1364]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="144" phylum="Chordata" rank="genus">
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<emphasis id="B97DEAD1FFB2D96AF6626B170DAEFAA5" box="[310,434,1340,1364]" italics="true" pageId="9" pageNumber="144">Milyeringa</emphasis>
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</taxonomicName>
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with small first dorsal fin of II–IV spines, usually III; 6–8 segmented second dorsal fin rays; I,6–8 anal fin rays; pectoral fin rays 11–14; pelvic fin rays I,4; 16–17 segmented caudal fin rays; caudal and pectoral fin rays all unbranched; body fully scaled with scales usually extending forward onto predorsal, 22–29 lateral scales, head short with reduced numbers of rows of sensory papillae on the head and body, posteroventral margin of preopercle forming expanded bony flange; vertebrae 10–12+14–15 (
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<quantity id="4CF19B26FFB2D96AF3166BE708B0FA15" box="[1090,1196,1484,1508]" metricMagnitude="-1" metricUnit="m" metricValue="6.35" metricValueMax="6.604" metricValueMin="6.096" pageId="9" pageNumber="144" unit="in" value="25.0" valueMax="26.0" valueMin="24.0">24–26 in</quantity>
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total), two pre-anal pterygiophores, one epural. Found in caves and wells of North-West Cape, Western
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<collectingCountry id="F31E7653FFB2D96AF3176BC408B2F9F6" box="[1091,1198,1519,1543]" name="Australia" pageId="9" pageNumber="144">Australia</collectingCountry>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3136548FFB2D968F79D68380DE8FF06" lastPageId="11" lastPageNumber="146" pageId="9" pageNumber="144" type="materials_examined">
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||
<paragraph id="8BB636C3FFB2D96AF79D683808C3F849" blockId="9.[151,1437,1339,1977]" pageId="9" pageNumber="144">
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||
<emphasis id="B97DEAD1FFB2D96AF79D68380DAAF9DD" bold="true" box="[201,438,1555,1580]" pageId="9" pageNumber="144">Material examined.</emphasis>
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WESTERN
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<collectingCountry id="F31E7653FFB2D96AF51B683F0EEEF9DD" box="[591,754,1556,1580]" name="Australia" pageId="9" pageNumber="144">AUSTRALIA</collectingCountry>
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: WAM P.2913,
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<typeStatus id="54B28861FFB2D96AF4FF683F0811F9DD" box="[939,1037,1556,1580]" pageId="9" pageNumber="144" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="4C094D40FFB2D96AF36E683F0911F9DD" box="[1082,1293,1556,1580]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="veritas">
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<emphasis id="B97DEAD1FFB2D96AF36E683F0911F9DD" box="[1082,1293,1556,1580]" italics="true" pageId="9" pageNumber="144">Milyeringa veritas</emphasis>
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</taxonomicName>
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,
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<quantity id="4CF19B26FFB2D96AF24B683F096DF9DD" box="[1311,1393,1556,1580]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="9" pageNumber="144" unit="mm" value="39.0">39 mm</quantity>
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SL female, Yardie Station; WAM P.28330-001,
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<typeStatus id="54B28861FFB2D96AF5D6681C0EF8F9BE" box="[642,740,1591,1615]" pageId="9" pageNumber="144" type="holotype">holotype</typeStatus>
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||
of
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<taxonomicName id="4C094D40FFB2D96AF45C68130FC2F9A1" box="[776,990,1592,1616]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="brooksi">
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<emphasis id="B97DEAD1FFB2D96AF45C68130FC2F9A1" box="[776,990,1592,1616]" italics="true" pageId="9" pageNumber="144">Milyeringa brooksi</emphasis>
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</taxonomicName>
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,
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<quantity id="4CF19B26FFB2D96AF4BF681C0852F9BE" box="[1003,1102,1591,1615]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" pageId="9" pageNumber="144" unit="mm" value="36.5">36.5 mm</quantity>
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SL female, Pilgonaman Well, North-west Cape, M. Newton,
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<date id="FFB71003FFB2D96AF55768770E8EF985" box="[515,658,1628,1652]" pageId="9" pageNumber="144" value="1984-07-08">8 July 1984</date>
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; WAM P.29242-001,
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<typeStatus id="54B28861FFB2D96AF4C668760FE1F984" box="[914,1021,1629,1653]" pageId="9" pageNumber="144" type="paratype">paratypes</typeStatus>
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of
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<taxonomicName id="4C094D40FFB2D96AF3786877091BF985" box="[1068,1287,1628,1652]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="brooksi">
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<emphasis id="B97DEAD1FFB2D96AF3786877091BF985" box="[1068,1287,1628,1652]" italics="true" pageId="9" pageNumber="144">Milyeringa brooksi</emphasis>
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</taxonomicName>
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, 2(36–38.5), Exmouth, B. Vine and party,
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<date id="FFB71003FFB2D96AF6B168540E9CF966" box="[485,640,1663,1689]" pageId="9" pageNumber="144" value="1985-05-19">19 May 1985</date>
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; WAM P.28262-001, 8(12.5–40.5), Milyering Well, North-west Cape; WAM P.33161-001, 2(34–36), Kubura Well, Cape Range, W.F. Humphreys,
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<date id="FFB71003FFB2D96AF34D688F08B4F94D" box="[1049,1192,1700,1724]" pageId="9" pageNumber="144" value="1993-07-01">1 July 1993</date>
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; WAM P.33157-001, 2(22–24), Five Mile Well, Cape Range, B. Vine,
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<date id="FFB71003FFB2D96AF59868EC0F78F92E" box="[716,868,1735,1759]" pageId="9" pageNumber="144" value="1989-07-15">15 July 1989</date>
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; WAM P.33168-001, 1(22.5), Exmouth bore field Water Corporation bore 18, Cape Range, Kinhill,
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<date id="FFB71003FFB2D96AF59668C70F48F8F5" box="[706,852,1772,1796]" pageId="9" pageNumber="144" value="2001-04-01">1 April 2001</date>
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; WAM P.33163-001, 1(13.5), Ampolex site D, Cape Range, R.D. Brooks,
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<date id="FFB71003FFB2D96AF6F469240E90F8D6" box="[416,652,1807,1831]" pageId="9" pageNumber="144" value="1995-11-12">12 November 1995</date>
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; WAM P.33160-001, 3(31–37), Kubura Well, Cape Range, W.F. Humphreys,
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<date id="FFB71003FFB2D96AF67E691F0DFAF8BD" box="[298,486,1844,1868]" pageId="9" pageNumber="144" value="1993-08-12">12 August 1993</date>
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; WAM P.33158-001, 1(32.5), Cape Range, W.F. Humphreys,
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<date id="FFB71003FFB2D96AF3E4691F0955F8BD" box="[1200,1353,1844,1868]" pageId="9" pageNumber="144" value="1993-07-15">15 July 1993</date>
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; WAM P.33159-001, 2(25–30.5), Kubura Well, Cape Range, W.F. Humphreys,
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<date id="FFB71003FFB2D96AF4936973087EF881" box="[967,1122,1880,1905]" pageId="9" pageNumber="144" value="1993-05-26">26 May 1993</date>
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; WAM P.33164-001, 1(13), Cape Range, A. Poole and S. Eberhard,
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<date id="FFB71003FFB2D96AF50969570F27F865" box="[605,827,1916,1940]" pageId="9" pageNumber="144" value="1997-09-22">22 September 1997</date>
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; AMS I.25504-001, 1(31), cleared and stained, well
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<quantity id="4CF19B26FFB2D96AF2DA69570CA7F849" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="144" unit="km" value="3.0">3 km</quantity>
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S of Mangrove Bay, Cape Range National Park, D. Hoese and D. Rennis,
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<date id="FFB71003FFB2D96AF4AA698B08C7F849" box="[1022,1243,1952,1976]" pageId="9" pageNumber="144" value="1985-09-13">13 September 1985</date>
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.
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</paragraph>
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<caption id="DF76664BFFB1D969F7C36EB3096FFF3D" ID-Table-UUID="DF76664BFFB1D969F7C36EB3096FFF3D" httpUri="http://table.plazi.org/id/DF76664BFFB1D969F7C36EB3096FFF3D" pageId="10" pageNumber="145" targetBox="[151,1436,231,802]" targetIsTable="true" targetPageId="10">
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||
<paragraph id="8BB636C3FFB1D969F7C36EB3096FFF3D" blockId="10.[151,1436,152,205]" pageId="10" pageNumber="145">
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<emphasis id="B97DEAD1FFB1D969F7C36EB30D19FF5C" bold="true" box="[151,261,152,173]" pageId="10" pageNumber="145">TABLE 3.</emphasis>
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Measurements of specimens of
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<taxonomicName id="4C094D40FFB1D969F51B6EB30F10FF5F" box="[591,780,152,174]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="145" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB1D969F51B6EB30F10FF5F" box="[591,780,152,174]" italics="true" pageId="10" pageNumber="145">Milyeringa veritas</emphasis>
|
||
</taxonomicName>
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||
, expressed as percentage of standard length (SL) or head length (HL). Missing data is usually due to specimen being damaged or distorted. (POP = preopercular posterior margin). N = 24.
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</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB636C3FFB1D969F5006ECC093BFCD3" pageId="10" pageNumber="145">
|
||
<table id="F909C463FFB1269CF7C36ECC0980FCD3" box="[151,1436,231,802]" gridcols="6" gridrows="14" pageId="10" pageNumber="145">
|
||
<tr id="35393481FFB1269CF7C36ECC0980FEEB" box="[151,1436,231,282]" gridrow="0" pageId="10" pageNumber="145" rowspan-0="1">
|
||
<th id="76E85DFDFFB1269CF5006ECC0EA9FEEB" box="[596,693,231,282]" gridcol="1" gridrow="0" pageId="10" pageNumber="145">
|
||
Holotype
|
||
<taxonomicName id="4C094D40FFB1D969F5006F2E0EA9FEEB" box="[596,693,261,282]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="145" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB1D969F5006F2E0EA9FEEB" box="[596,693,261,282]" italics="true" pageId="10" pageNumber="145">M. veritas</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76E85DFDFFB1269CF5AA6ECC0F79FEEB" box="[766,869,231,282]" gridcol="2" gridrow="0" pageId="10" pageNumber="145">
|
||
Holotype
|
||
<taxonomicName id="4C094D40FFB1D969F5AA6F2E0F79FEEB" box="[766,869,261,282]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="145" phylum="Chordata" rank="species" species="brooksi">
|
||
<emphasis id="B97DEAD1FFB1D969F5AA6F2E0F79FEEB" box="[766,869,261,282]" italics="true" pageId="10" pageNumber="145">M. brooksi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76E85DFDFFB1269CF4FD6ECC0FC3FEEB" box="[937,991,231,282]" gridcol="3" gridrow="0" pageId="10" pageNumber="145">Mean</th>
|
||
<th id="76E85DFDFFB1269CF3076ECC08ABFEEB" box="[1107,1207,231,282]" gridcol="4" gridrow="0" pageId="10" pageNumber="145">Maximum</th>
|
||
<th id="76E85DFDFFB1269CF3A96ECC0980FEEB" box="[1277,1436,231,282]" gridcol="5" gridrow="0" pageId="10" pageNumber="145">Minimum</th>
|
||
</tr>
|
||
<tr id="35393481FFB1269CF7C36F060980FEB3" box="[151,1436,301,322]" gridrow="1" pageId="10" pageNumber="145">
|
||
<th id="76E85DFDFFB1269CF7C36F060D8EFEB3" box="[151,402,301,322]" gridcol="0" gridrow="1" pageId="10" pageNumber="145">Standard length</th>
|
||
<td id="76E85DFDFFB1269CF5006F060EA9FEB3" box="[596,693,301,322]" gridcol="1" gridrow="1" pageId="10" pageNumber="145">39.0</td>
|
||
<td id="76E85DFDFFB1269CF5AA6F060F79FEB3" box="[766,869,301,322]" gridcol="2" gridrow="1" pageId="10" pageNumber="145">36.5</td>
|
||
<td id="76E85DFDFFB1269CF4FD6F060FC3FEB3" box="[937,991,301,322]" gridcol="3" gridrow="1" pageId="10" pageNumber="145">29</td>
|
||
<td id="76E85DFDFFB1269CF3076F0608ABFEB3" box="[1107,1207,301,322]" gridcol="4" gridrow="1" pageId="10" pageNumber="145">40.5</td>
|
||
<td id="76E85DFDFFB1269CF3A96F060980FEB3" box="[1277,1436,301,322]" gridcol="5" gridrow="1" pageId="10" pageNumber="145">12.5</td>
|
||
</tr>
|
||
<tr id="35393481FFB1269CF7C36F7E0980FE9B" box="[151,1436,341,362]" gridrow="2" pageId="10" pageNumber="145">
|
||
<th id="76E85DFDFFB1269CF7C36F7E0D8EFE9B" box="[151,402,341,362]" gridcol="0" gridrow="2" pageId="10" pageNumber="145">Head length in SL</th>
|
||
<td id="76E85DFDFFB1269CF5006F7E0EA9FE9B" box="[596,693,341,362]" gridcol="1" gridrow="2" pageId="10" pageNumber="145">40.3</td>
|
||
<td id="76E85DFDFFB1269CF5AA6F7E0F79FE9B" box="[766,869,341,362]" gridcol="2" gridrow="2" pageId="10" pageNumber="145">40.0</td>
|
||
<td id="76E85DFDFFB1269CF4FD6F7E0FC3FE9B" box="[937,991,341,362]" gridcol="3" gridrow="2" pageId="10" pageNumber="145">40.0</td>
|
||
<td id="76E85DFDFFB1269CF3076F7E08ABFE9B" box="[1107,1207,341,362]" gridcol="4" gridrow="2" pageId="10" pageNumber="145">42.3</td>
|
||
<td id="76E85DFDFFB1269CF3A96F7E0980FE9B" box="[1277,1436,341,362]" gridcol="5" gridrow="2" pageId="10" pageNumber="145">36.9</td>
|
||
</tr>
|
||
<tr id="35393481FFB1269CF7C36F560980FE63" box="[151,1436,381,402]" gridrow="3" pageId="10" pageNumber="145">
|
||
<th id="76E85DFDFFB1269CF7C36F560D8EFE63" box="[151,402,381,402]" gridcol="0" gridrow="3" pageId="10" pageNumber="145">Head depth in HL</th>
|
||
<td id="76E85DFDFFB1269CF5006F560EA9FE63" box="[596,693,381,402]" gridcol="1" gridrow="3" pageId="10" pageNumber="145">42.7</td>
|
||
<td id="76E85DFDFFB1269CF5AA6F560F79FE63" box="[766,869,381,402]" gridcol="2" gridrow="3" pageId="10" pageNumber="145">46.6</td>
|
||
<td id="76E85DFDFFB1269CF4FD6F560FC3FE63" box="[937,991,381,402]" gridcol="3" gridrow="3" pageId="10" pageNumber="145">42.2</td>
|
||
<td id="76E85DFDFFB1269CF3076F5608ABFE63" box="[1107,1207,381,402]" gridcol="4" gridrow="3" pageId="10" pageNumber="145">53.5</td>
|
||
<td id="76E85DFDFFB1269CF3A96F560980FE63" box="[1277,1436,381,402]" gridcol="5" gridrow="3" pageId="10" pageNumber="145">35.4</td>
|
||
</tr>
|
||
<tr id="35393481FFB1269CF7C36F8E0980FE4B" box="[151,1436,421,442]" gridrow="4" pageId="10" pageNumber="145">
|
||
<th id="76E85DFDFFB1269CF7C36F8E0D8EFE4B" box="[151,402,421,442]" gridcol="0" gridrow="4" pageId="10" pageNumber="145">Head width in HL</th>
|
||
<td id="76E85DFDFFB1269CF5006F8E0EA9FE4B" box="[596,693,421,442]" gridcol="1" gridrow="4" pageId="10" pageNumber="145">68.2</td>
|
||
<td id="76E85DFDFFB1269CF5AA6F8E0F79FE4B" box="[766,869,421,442]" gridcol="2" gridrow="4" pageId="10" pageNumber="145">67.8</td>
|
||
<td id="76E85DFDFFB1269CF4FD6F8E0FC3FE4B" box="[937,991,421,442]" gridcol="3" gridrow="4" pageId="10" pageNumber="145">61.0</td>
|
||
<td id="76E85DFDFFB1269CF3076F8E08ABFE4B" box="[1107,1207,421,442]" gridcol="4" gridrow="4" pageId="10" pageNumber="145">73.7</td>
|
||
<td id="76E85DFDFFB1269CF3A96F8E0980FE4B" box="[1277,1436,421,442]" gridcol="5" gridrow="4" pageId="10" pageNumber="145">49.5</td>
|
||
</tr>
|
||
<tr id="35393481FFB1269CF7C36FE60980FE13" box="[151,1436,461,482]" gridrow="5" pageId="10" pageNumber="145">
|
||
<th id="76E85DFDFFB1269CF7C36FE60D8EFE13" box="[151,402,461,482]" gridcol="0" gridrow="5" pageId="10" pageNumber="145">Body depth at anus in SL</th>
|
||
<td id="76E85DFDFFB1269CF5006FE60EA9FE13" box="[596,693,461,482]" gridcol="1" gridrow="5" pageId="10" pageNumber="145">17.7</td>
|
||
<td id="76E85DFDFFB1269CF5AA6FE60F79FE13" box="[766,869,461,482]" gridcol="2" gridrow="5" pageId="10" pageNumber="145">17.5</td>
|
||
<td id="76E85DFDFFB1269CF4FD6FE60FC3FE13" box="[937,991,461,482]" gridcol="3" gridrow="5" pageId="10" pageNumber="145">16.9</td>
|
||
<td id="76E85DFDFFB1269CF3076FE608ABFE13" box="[1107,1207,461,482]" gridcol="4" gridrow="5" pageId="10" pageNumber="145">20.0</td>
|
||
<td id="76E85DFDFFB1269CF3A96FE60980FE13" box="[1277,1436,461,482]" gridcol="5" gridrow="5" pageId="10" pageNumber="145">12.3</td>
|
||
</tr>
|
||
<tr id="35393481FFB1269CF7C36FDE0980FDFB" box="[151,1436,501,522]" gridrow="6" pageId="10" pageNumber="145">
|
||
<th id="76E85DFDFFB1269CF7C36FDE0D8EFDFB" box="[151,402,501,522]" gridcol="0" gridrow="6" pageId="10" pageNumber="145">Caud. ped. length in SL</th>
|
||
<td id="76E85DFDFFB1269CF5006FDE0EA9FDFB" box="[596,693,501,522]" gridcol="1" gridrow="6" pageId="10" pageNumber="145">22.8</td>
|
||
<td id="76E85DFDFFB1269CF5AA6FDE0F79FDFB" box="[766,869,501,522]" gridcol="2" gridrow="6" pageId="10" pageNumber="145">21.1</td>
|
||
<td id="76E85DFDFFB1269CF4FD6FDE0FC3FDFB" box="[937,991,501,522]" gridcol="3" gridrow="6" pageId="10" pageNumber="145">24.3</td>
|
||
<td id="76E85DFDFFB1269CF3076FDE08ABFDFB" box="[1107,1207,501,522]" gridcol="4" gridrow="6" pageId="10" pageNumber="145">27.7</td>
|
||
<td id="76E85DFDFFB1269CF3A96FDE0980FDFB" box="[1277,1436,501,522]" gridcol="5" gridrow="6" pageId="10" pageNumber="145">20.0</td>
|
||
</tr>
|
||
<tr id="35393481FFB1269CF7C36C360980FDC3" box="[151,1436,541,562]" gridrow="7" pageId="10" pageNumber="145">
|
||
<th id="76E85DFDFFB1269CF7C36C360D8EFDC3" box="[151,402,541,562]" gridcol="0" gridrow="7" pageId="10" pageNumber="145">Caud. ped. depth in SL</th>
|
||
<td id="76E85DFDFFB1269CF5006C360EA9FDC3" box="[596,693,541,562]" gridcol="1" gridrow="7" pageId="10" pageNumber="145">10.0</td>
|
||
<td id="76E85DFDFFB1269CF5AA6C360F79FDC3" box="[766,869,541,562]" gridcol="2" gridrow="7" pageId="10" pageNumber="145">10.4</td>
|
||
<td id="76E85DFDFFB1269CF4FD6C360FC3FDC3" box="[937,991,541,562]" gridcol="3" gridrow="7" pageId="10" pageNumber="145">10.2</td>
|
||
<td id="76E85DFDFFB1269CF3076C3608ABFDC3" box="[1107,1207,541,562]" gridcol="4" gridrow="7" pageId="10" pageNumber="145">11.8</td>
|
||
<td id="76E85DFDFFB1269CF3A96C360980FDC3" box="[1277,1436,541,562]" gridcol="5" gridrow="7" pageId="10" pageNumber="145">7.7</td>
|
||
</tr>
|
||
<tr id="35393481FFB1269CF7C36C6E0980FDAB" box="[151,1436,581,602]" gridrow="8" pageId="10" pageNumber="145">
|
||
<th id="76E85DFDFFB1269CF7C36C6E0D8EFDAB" box="[151,402,581,602]" gridcol="0" gridrow="8" pageId="10" pageNumber="145">Snout to POP in HL</th>
|
||
<td id="76E85DFDFFB1269CF5006C6E0EA9FDAB" box="[596,693,581,602]" gridcol="1" gridrow="8" pageId="10" pageNumber="145">-</td>
|
||
<td id="76E85DFDFFB1269CF5AA6C6E0F79FDAB" box="[766,869,581,602]" gridcol="2" gridrow="8" pageId="10" pageNumber="145">71.9</td>
|
||
<td id="76E85DFDFFB1269CF4FD6C6E0FC3FDAB" box="[937,991,581,602]" gridcol="3" gridrow="8" pageId="10" pageNumber="145">69.1</td>
|
||
<td id="76E85DFDFFB1269CF3076C6E08ABFDAB" box="[1107,1207,581,602]" gridcol="4" gridrow="8" pageId="10" pageNumber="145">72.9</td>
|
||
<td id="76E85DFDFFB1269CF3A96C6E0980FDAB" box="[1277,1436,581,602]" gridcol="5" gridrow="8" pageId="10" pageNumber="145">65.5</td>
|
||
</tr>
|
||
<tr id="35393481FFB1269CF7C36C460980FD73" box="[151,1436,621,642]" gridrow="9" pageId="10" pageNumber="145">
|
||
<th id="76E85DFDFFB1269CF7C36C460D8EFD73" box="[151,402,621,642]" gridcol="0" gridrow="9" pageId="10" pageNumber="145">Jaw length in HL</th>
|
||
<td id="76E85DFDFFB1269CF5006C460EA9FD73" box="[596,693,621,642]" gridcol="1" gridrow="9" pageId="10" pageNumber="145">40.1</td>
|
||
<td id="76E85DFDFFB1269CF5AA6C460F79FD73" box="[766,869,621,642]" gridcol="2" gridrow="9" pageId="10" pageNumber="145">43.8</td>
|
||
<td id="76E85DFDFFB1269CF4FD6C460FC3FD73" box="[937,991,621,642]" gridcol="3" gridrow="9" pageId="10" pageNumber="145">37.9</td>
|
||
<td id="76E85DFDFFB1269CF3076C4608ABFD73" box="[1107,1207,621,642]" gridcol="4" gridrow="9" pageId="10" pageNumber="145">43.8</td>
|
||
<td id="76E85DFDFFB1269CF3A96C460980FD73" box="[1277,1436,621,642]" gridcol="5" gridrow="9" pageId="10" pageNumber="145">27.1</td>
|
||
</tr>
|
||
<tr id="35393481FFB1269CF7C36CBE0980FD5B" box="[151,1436,661,682]" gridrow="10" pageId="10" pageNumber="145">
|
||
<th id="76E85DFDFFB1269CF7C36CBE0D8EFD5B" box="[151,402,661,682]" gridcol="0" gridrow="10" pageId="10" pageNumber="145">Opercle length in HL</th>
|
||
<td id="76E85DFDFFB1269CF5006CBE0EA9FD5B" box="[596,693,661,682]" gridcol="1" gridrow="10" pageId="10" pageNumber="145">-</td>
|
||
<td id="76E85DFDFFB1269CF5AA6CBE0F79FD5B" box="[766,869,661,682]" gridcol="2" gridrow="10" pageId="10" pageNumber="145">33.6</td>
|
||
<td id="76E85DFDFFB1269CF4FD6CBE0FC3FD5B" box="[937,991,661,682]" gridcol="3" gridrow="10" pageId="10" pageNumber="145">34.6</td>
|
||
<td id="76E85DFDFFB1269CF3076CBE08ABFD5B" box="[1107,1207,661,682]" gridcol="4" gridrow="10" pageId="10" pageNumber="145">39.8</td>
|
||
<td id="76E85DFDFFB1269CF3A96CBE0980FD5B" box="[1277,1436,661,682]" gridcol="5" gridrow="10" pageId="10" pageNumber="145">27.8</td>
|
||
</tr>
|
||
<tr id="35393481FFB1269CF7C36C960980FD23" box="[151,1436,701,722]" gridrow="11" pageId="10" pageNumber="145">
|
||
<th id="76E85DFDFFB1269CF7C36C960D8EFD23" box="[151,402,701,722]" gridcol="0" gridrow="11" pageId="10" pageNumber="145">Pectoral fin in SL</th>
|
||
<td id="76E85DFDFFB1269CF5006C960EA9FD23" box="[596,693,701,722]" gridcol="1" gridrow="11" pageId="10" pageNumber="145">-</td>
|
||
<td id="76E85DFDFFB1269CF5AA6C960F79FD23" box="[766,869,701,722]" gridcol="2" gridrow="11" pageId="10" pageNumber="145">18.6</td>
|
||
<td id="76E85DFDFFB1269CF4FD6C960FC3FD23" box="[937,991,701,722]" gridcol="3" gridrow="11" pageId="10" pageNumber="145">19.3</td>
|
||
<td id="76E85DFDFFB1269CF3076C9608ABFD23" box="[1107,1207,701,722]" gridcol="4" gridrow="11" pageId="10" pageNumber="145">28.9</td>
|
||
<td id="76E85DFDFFB1269CF3A96C960980FD23" box="[1277,1436,701,722]" gridcol="5" gridrow="11" pageId="10" pageNumber="145">12.3</td>
|
||
</tr>
|
||
<tr id="35393481FFB1269CF7C36CCE0980FD0B" box="[151,1436,741,762]" gridrow="12" pageId="10" pageNumber="145">
|
||
<th id="76E85DFDFFB1269CF7C36CCE0D8EFD0B" box="[151,402,741,762]" gridcol="0" gridrow="12" pageId="10" pageNumber="145">Pelvic fin in SL</th>
|
||
<td id="76E85DFDFFB1269CF5006CCE0EA9FD0B" box="[596,693,741,762]" gridcol="1" gridrow="12" pageId="10" pageNumber="145">-</td>
|
||
<td id="76E85DFDFFB1269CF5AA6CCE0F79FD0B" box="[766,869,741,762]" gridcol="2" gridrow="12" pageId="10" pageNumber="145">19.7</td>
|
||
<td id="76E85DFDFFB1269CF4FD6CCE0FC3FD0B" box="[937,991,741,762]" gridcol="3" gridrow="12" pageId="10" pageNumber="145">17.0</td>
|
||
<td id="76E85DFDFFB1269CF3076CCE08ABFD0B" box="[1107,1207,741,762]" gridcol="4" gridrow="12" pageId="10" pageNumber="145">26.9</td>
|
||
<td id="76E85DFDFFB1269CF3A96CCE0980FD0B" box="[1277,1436,741,762]" gridcol="5" gridrow="12" pageId="10" pageNumber="145">7.7</td>
|
||
</tr>
|
||
<tr id="35393481FFB1269CF7C36D260980FCD3" box="[151,1436,781,802]" gridrow="13" pageId="10" pageNumber="145">
|
||
<th id="76E85DFDFFB1269CF7C36D260D8EFCD3" box="[151,402,781,802]" gridcol="0" gridrow="13" pageId="10" pageNumber="145">Caudal fin in SL</th>
|
||
<td id="76E85DFDFFB1269CF5006D260EA9FCD3" box="[596,693,781,802]" gridcol="1" gridrow="13" pageId="10" pageNumber="145">-</td>
|
||
<td id="76E85DFDFFB1269CF5AA6D260F79FCD3" box="[766,869,781,802]" gridcol="2" gridrow="13" pageId="10" pageNumber="145">-</td>
|
||
<td id="76E85DFDFFB1269CF4FD6D260FC3FCD3" box="[937,991,781,802]" gridcol="3" gridrow="13" pageId="10" pageNumber="145">28.2</td>
|
||
<td id="76E85DFDFFB1269CF3076D2608ABFCD3" box="[1107,1207,781,802]" gridcol="4" gridrow="13" pageId="10" pageNumber="145">39.2</td>
|
||
<td id="76E85DFDFFB1269CF3A96D260980FCD3" box="[1277,1436,781,802]" gridcol="5" gridrow="13" pageId="10" pageNumber="145">18.8</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF76664BFFB1D969F7C369840E23F815" httpUri="https://zenodo.org/record/220766/files/figure.png" pageId="10" pageNumber="145" targetBox="[286,1291,870,1949]" targetPageId="10">
|
||
<paragraph id="8BB636C3FFB1D969F7C369840E23F815" blockId="10.[151,1436,1967,2020]" pageId="10" pageNumber="145">
|
||
<emphasis id="B97DEAD1FFB1D969F7C369840D11F834" bold="true" box="[151,269,1967,1990]" pageId="10" pageNumber="145">FIGURE 9</emphasis>
|
||
. Lateral (A) and dorsal (B) views of head of
|
||
<taxonomicName id="4C094D40FFB1D969F5B669840F83F837" box="[738,927,1967,1990]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="145" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB1D969F5B669840F83F837" box="[738,927,1967,1990]" italics="true" pageId="10" pageNumber="145">Milyeringa veritas</emphasis>
|
||
</taxonomicName>
|
||
, WAM P.28262-001, 37 mm SL female, showing sensory papillae pattern. Scales indicated.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB636C3FFB0D968F79D6EB30DE8FF06" blockId="11.[151,1436,151,1358]" pageId="11" pageNumber="146">
|
||
<emphasis id="B97DEAD1FFB0D968F79D6EB30D83FF41" box="[201,415,152,176]" italics="true" pageId="11" pageNumber="146">X-rayed specimens</emphasis>
|
||
: F03762, 1, North West Cape, West
|
||
<collectingCountry id="F31E7653FFB0D968F4106EBC0FB2FF5E" box="[836,942,151,175]" name="Australia" pageId="11" pageNumber="146">Australia</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE3D5004FFB0D968F4EE6EBC083EFF5E" box="[954,1058,151,175]" direction="south" orientation="latitude" pageId="11" pageNumber="146" precision="925" value="-21.783333">21° 47' S</geoCoordinate>
|
||
<geoCoordinate id="EE3D5004FFB0D968F37F6EBC08BAFF5E" box="[1067,1190,151,175]" direction="east" orientation="longitude" pageId="11" pageNumber="146" precision="925" value="114.166664">114° 10' E</geoCoordinate>
|
||
; F05491, 3, township rockhole, Exmouth, West
|
||
<collectingCountry id="F31E7653FFB0D968F6EA6E970E34FF25" box="[446,552,188,212]" name="Australia" pageId="11" pageNumber="146">Australia</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE3D5004FFB0D968F5676E970E87FF25" box="[563,667,188,212]" direction="south" orientation="latitude" pageId="11" pageNumber="146" precision="925" value="-21.933332">21° 56' S</geoCoordinate>
|
||
<geoCoordinate id="EE3D5004FFB0D968F5F06E970F05FF25" box="[676,793,188,212]" direction="east" orientation="longitude" pageId="11" pageNumber="146" precision="925" value="114.11667">114° 7' E’</geoCoordinate>
|
||
); F05492, 3, rockhole
|
||
<quantity id="4CF19B26FFB0D968F34F6E97084AFF25" box="[1051,1110,188,212]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="146" unit="km" value="6.0">6 km</quantity>
|
||
north of Yardie Creek, West
|
||
<collectingCountry id="F31E7653FFB0D968F7C36EF40D1DFF06" box="[151,257,223,247]" name="Australia" pageId="11" pageNumber="146">Australia</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE3D5004FFB0D968F6586EF40D6EFF06" box="[268,370,223,247]" direction="south" orientation="latitude" pageId="11" pageNumber="146" precision="925" value="-22.333334">22° 20' S</geoCoordinate>
|
||
<geoCoordinate id="EE3D5004FFB0D968F62C6EF40DF3FF06" box="[376,495,223,247]" direction="east" orientation="longitude" pageId="11" pageNumber="146" precision="925" value="113.85">113° 51' E</geoCoordinate>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136548FFB0D96FF7C36F280E36F937" lastPageId="12" lastPageNumber="147" pageId="11" pageNumber="146" type="description">
|
||
<paragraph id="8BB636C3FFB0D968F7C36F280DC4FECC" blockId="11.[151,1436,151,1358]" pageId="11" pageNumber="146">
|
||
<emphasis id="B97DEAD1FFB0D968F7C36F280D35FEED" bold="true" box="[151,297,259,284]" pageId="11" pageNumber="146">Description.</emphasis>
|
||
Based on
|
||
<specimenCount id="9D0FFD4AFFB0D968F6C96F2F0E24FEED" box="[413,568,260,284]" pageId="11" pageNumber="146" type="generic">23 specimens</specimenCount>
|
||
,
|
||
<quantity id="4CF19B26FFB0D968F5176F2F0EFAFEED" box="[579,742,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="4.05" metricValueMin="1.25" pageId="11" pageNumber="146" unit="mm" value="26.5" valueMax="40.5" valueMin="12.5">12.5–40.5 mm</quantity>
|
||
SL. Counts and morphometrics of the
|
||
<typeStatus id="54B28861FFB0D968F3CC6F2F08E6FEED" box="[1176,1274,260,284]" pageId="11" pageNumber="146" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C094D40FFB0D968F24B6F2F0CFFFECC" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB0D968F24B6F2F0CFFFECC" italics="true" pageId="11" pageNumber="146">Milyeringa veritas</emphasis>
|
||
</taxonomicName>
|
||
indicated by asterisk.
|
||
</paragraph>
|
||
<paragraph id="8BB636C3FFB0D968F79D6F6D0836FDE3" blockId="11.[151,1436,151,1358]" pageId="11" pageNumber="146">
|
||
First dorsal spines II–IV*, modally III; second dorsal rays 6–8*, modally 8; anal rays I,6–8, modally I,7* pectoral rays 11–14, modally 13; pelvic rays I,4; segmented caudal rays 16–17*, in 8/8 (in 4) or 9/8* pattern (16), all fin rays unbranched; lateral scale count 22–29*, modally 26; transverse scales backward 9–12, modally 10 (
|
||
<quantity id="4CF19B26FFB0D968F2D56FA50CB1FE3B" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="11" pageNumber="146" unit="in" value="11.0">11 in</quantity>
|
||
<typeStatus id="54B28861FFB0D968F7E36F990D02FE3B" box="[183,286,434,458]" pageId="11" pageNumber="146" type="holotype">holotype</typeStatus>
|
||
); predorsal scale count 0–16*, modally 12; vertebrae 10+14 (1), 11+14 (1), 11+15 (4), 12+14 (1); dorsal pterygiophore pattern 4–221 (in 1); 1 epural; 2 anal pterygiophores anterior to first haemal spine; last haemal spine broadened or forked (with variable amount of sheet of bone between forks).
|
||
</paragraph>
|
||
<paragraph id="8BB636C3FFB0D968F7936C3508B9FABF" blockId="11.[151,1436,151,1358]" pageId="11" pageNumber="146">
|
||
Body moderate, somewhat rounded anteriorly, compressed posteriorly; body depth at anal fin origin 12.3–20.0% of SL. Caudal peduncle length 20.0–27.7% of SL. Caudal peduncle depth 7.7–11.8% of SL. Head rounded to depressed, larger specimens (over
|
||
<quantity id="4CF19B26FFB0D968F5CE6C4D0EF5FD8F" box="[666,745,614,638]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="146" unit="mm" value="30.0">30 mm</quantity>
|
||
SL) with head much wider than deep at preopercular margin, head length 36.9–42.3% of SL; head depth at posterior preopercular margin 35.4–53.5% of HL; width at posterior preopercular margin 49.5–73.7% of HL. Mouth large, terminal and oblique, chin tip anteriormost, jaws forming an angle of about 35º with body axis. Upper jaw length 27.1–43.8% of HL; inner margin of lips smooth; lower lip fused to chin anteriorly, side of lip free; chin flat with no mental frenum. Anterior naris in short tube at edge of or just above upper lip; posterior naris oval to flattened oval, with low rim (or very short tube), occasionally anterior rim slightly produced as low curved flap; nares joined by thin “tube” over nasal rosette. Eyes absent. Snout area broad, flattened, tilted upward slightly, rounded to nearly square anteriorly in dorsal view; snout slightly pointed in lateral view mostly in small specimens, length of snout tip to upper edge of the preopercular margin 65.5–72.9% of HL. Opercle length (upper edge of preopercle to upper margin of opercle) 27.8–39.8% of HL. Preopercular margin bony and broadened into flange posteroventrally, extending from about halfway down posterior margin nearly to rictus; flange present even in small specimens (e.g.
|
||
<quantity id="4CF19B26FFB0D968F5B36DD90F24FBFB" box="[743,824,1010,1034]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="146" unit="mm" value="15.0">15 mm</quantity>
|
||
SL), gently curved outward in some specimens (may not be size-related), flange may be angular or with indentation near corner. Gill opening wide and somewhat variable in anterior extent, reaching forward from just before preopercular rear margin (as in
|
||
<typeStatus id="54B28861FFB0D968F3E66A11090AFBA3" box="[1202,1302,1082,1106]" pageId="11" pageNumber="146" type="holotype">holotype</typeStatus>
|
||
) to halfway between preopercular rear margin and rictus. Tongue large, tip blunt. Teeth in both jaws small, evenly sized, conical and pointed; in two to three rows; teeth of inner rows larger, pointing inward in larger specimens and more upright in smaller. Headpores absent. Sensory papillae conspicuous fleshy bumps, in longitudinal pattern; papillae with thin narrow flap each; thin flaps usually present only on well-preserved specimens (
|
||
<figureCitation id="13322A46FFB0D968F3D26AE108D7FB13" box="[1158,1227,1226,1250]" captionStart="FIGURE 9" captionStartId="10.[151,250,1967,1990]" captionTargetBox="[286,1291,870,1949]" captionTargetPageId="10" captionText="FIGURE 9. Lateral (A) and dorsal (B) views of head of Milyeringa veritas, WAM P. 28262 - 001, 37 mm SL female, showing sensory papillae pattern. Scales indicated." httpUri="https://zenodo.org/record/220766/files/figure.png" pageId="11" pageNumber="146">Fig. 9</figureCitation>
|
||
). Sensory papillae on body on body in variable numbers of irregular vertical (transverse) rows, often interspersed with single papillae and few papillae along hypural crease; papillae on body sometimes with broad to thin narrow flaps (uncertain if absence actual or due to damage); three longitudinal rows of papillae on caudal fin (
|
||
<figureCitation id="13322A46FFB0D968F3106B1D088BFABF" box="[1092,1175,1334,1358]" captionStart="FIGURE 10" captionStartId="11.[151,250,1877,1900]" captionTargetBox="[180,1426,1425,1835]" captionTargetId="figure@11.[151,1436,1412,1857]" captionTargetPageId="11" captionText="FIGURE 10. Composite sketch of Milyeringa veritas showing papillae rows on body and caudal fin; taken from WAM P. 28262 - 001, 40 mm SL male, and WAM P. 133158 - 001, 32.5 mm SL probable female." httpUri="https://zenodo.org/record/220767/files/figure.png" pageId="11" pageNumber="146">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF76664BFFB0D968F7C3697E0815F87B" httpUri="https://zenodo.org/record/220767/files/figure.png" pageId="11" pageNumber="146" targetBox="[180,1426,1425,1835]" targetPageId="11">
|
||
<paragraph id="8BB636C3FFB0D968F7C3697E0815F87B" blockId="11.[151,1436,1877,1930]" pageId="11" pageNumber="146">
|
||
<emphasis id="B97DEAD1FFB0D968F7C3697E0D01F89A" bold="true" box="[151,285,1877,1900]" pageId="11" pageNumber="146">FIGURE 10</emphasis>
|
||
. Composite sketch of
|
||
<taxonomicName id="4C094D40FFB0D968F546697D0ECEF89D" box="[530,722,1878,1900]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB0D968F546697D0ECEF89D" box="[530,722,1878,1900]" italics="true" pageId="11" pageNumber="146">Milyeringa veritas</emphasis>
|
||
</taxonomicName>
|
||
showing papillae rows on body and caudal fin; taken from WAM P.28262-001, 40 mm SL male, and WAM P.133158-001, 32.5 mm SL probable female.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB636C3FFB7D96FF7936EBC0828FF06" blockId="12.[151,1437,151,645]" pageId="12" pageNumber="147">Body covered with small cycloid scales, reaching forward onto head above level of opercle or above rear preopercular margin. Side of head naked. Prepelvic region naked (usually) or with patch of cycloid scales just before pelvic fin bases. Pectoral fin base naked. Belly covered with cycloid scales.</paragraph>
|
||
<paragraph id="8BB636C3FFB7D96FF7936F2F0F85FDE6" blockId="12.[151,1437,151,645]" pageId="12" pageNumber="147">First dorsal fin greatly reduced, roughly triangular, with gap of about three or four scales before the second dorsal fin origin. Posteriormost second dorsal and anal fins short-based, rays taller than first dorsal fin; posterior rays slightly longer than anterior rays but not greatly so, posterior rays falling well short of caudal fin base. Second dorsal fin with all elements segmented. Anal fin usually with spine and segmented rays (three specimens with all elements segmented). Pectoral fin small, slender, pointed, central rays longest, 12.3–28.9% of SL; all rays unbranched. Pelvic fin length 7.7–26.9% of SL; fins small, very slender and pointed, fins extending less than half the distance to anus. Caudal fin oval, with central rays elongate, often greatly so (elongate rays often damaged or broken in available specimens); caudal fin length 18.8–39.2% of SL.</paragraph>
|
||
<paragraph id="8BB636C3FFB7D96FF79D6C0E0812FD75" blockId="12.[151,1437,151,645]" pageId="12" pageNumber="147">
|
||
<emphasis id="B97DEAD1FFB7D96FF79D6C0E0D69FDCD" box="[201,373,548,572]" italics="true" pageId="12" pageNumber="147">Live coloration</emphasis>
|
||
. Photos of living fish are shown in Allen (1982), Merrick and Schmida (1984), Young (1986) and Allen
|
||
<emphasis id="B97DEAD1FFB7D96FF65F6C620D5FFD91" box="[267,323,584,608]" italics="true" pageId="12" pageNumber="147">et al.</emphasis>
|
||
(2002) (
|
||
<figureCitation id="13322A46FFB7D96FF6F76C6C0DE9FDAE" box="[419,501,583,607]" captionStart="FIGURE 11" captionStartId="12.[151,250,1355,1378]" captionTargetBox="[302,1284,680,1333]" captionTargetId="figure@12.[302,1285,679,1334]" captionTargetPageId="12" captionText="FIGURE 11. Captive Milyeringa veritas. Photograph by Douglas Elford, Western Australian Museum." httpUri="https://zenodo.org/record/220768/files/figure.png" pageId="12" pageNumber="147">Fig. 11</figureCitation>
|
||
). Fish white to translucent white posteriorly, with transparent fins, and red gills and pinkish colour from abdominal organs showing through opercle and body wall.
|
||
</paragraph>
|
||
<caption id="DF76664BFFB7D96FF7C36B6008B1FA93" box="[151,1197,1355,1378]" httpUri="https://zenodo.org/record/220768/files/figure.png" pageId="12" pageNumber="147" targetBox="[302,1284,680,1333]" targetPageId="12">
|
||
<paragraph id="8BB636C3FFB7D96FF7C36B6008B1FA93" blockId="12.[151,1197,1355,1378]" box="[151,1197,1355,1378]" pageId="12" pageNumber="147">
|
||
<emphasis id="B97DEAD1FFB7D96FF7C36B600D02FA90" bold="true" box="[151,286,1355,1378]" pageId="12" pageNumber="147">FIGURE 11.</emphasis>
|
||
Captive
|
||
<taxonomicName id="4C094D40FFB7D96FF62D6B670E29FA93" box="[377,565,1356,1378]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="147" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB7D96FF62D6B670E29FA93" box="[377,565,1356,1378]" italics="true" pageId="12" pageNumber="147">Milyeringa veritas</emphasis>
|
||
</taxonomicName>
|
||
. Photograph by Douglas Elford, Western Australian Museum.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB636C3FFB7D96FF79D6BA50981FA3B" blockId="12.[151,1437,1422,2022]" pageId="12" pageNumber="147">
|
||
<emphasis id="B97DEAD1FFB7D96FF79D6BA50DD7FA57" box="[201,459,1422,1446]" italics="true" pageId="12" pageNumber="147">Coloration in alcohol.</emphasis>
|
||
Whitish, with transparent fin membranes. Specimens often with very fine scattered blackish to dark brown melanophores over dorsal surface of cranium and frontals (i.e. would-be interorbital space).
|
||
</paragraph>
|
||
<paragraph id="8BB636C3FFB7D96FF7936BFC0E36F937" blockId="12.[151,1437,1422,2022]" pageId="12" pageNumber="147">
|
||
<emphasis id="B97DEAD1FFB7D96FF7936BFC0DC6FA1E" bold="true" box="[199,474,1494,1519]" pageId="12" pageNumber="147">Results of genotyping.</emphasis>
|
||
The branch of the recovered COI gene tree representing mainland (Cape Range peninsula)
|
||
<taxonomicName id="4C094D40FFB7D96FF6416BD00D8EF9E2" box="[277,402,1531,1555]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="147" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97DEAD1FFB7D96FF6416BD00D8EF9E2" box="[277,402,1531,1555]" italics="true" pageId="12" pageNumber="147">Milyeringa</emphasis>
|
||
</taxonomicName>
|
||
closely resembles Chakrabarty’s (2010) tree in topology and genetic distances within and between two major clades (
|
||
<figureCitation id="13322A46FFB7D96FF68668350E0AF9C7" box="[466,534,1566,1590]" captionStart="FIGURE 3" captionStartId="4.[151,250,1425,1448]" captionTargetBox="[151,1435,373,1404]" captionTargetId="figure@4.[151,1436,373,1405]" captionTargetPageId="4" captionText="FIGURE 3. Milyeringa COI neighbor-joining tree. Collection localities of samples are in parentheses following the sample number. Red indicates the M. brooksi haplotype clade and blue indicates those individuals collected from Kudumurra Well (C- 25)." httpUri="https://zenodo.org/record/220761/files/figure.png" pageId="12" pageNumber="147">Fig. 3</figureCitation>
|
||
) though greater substructure is evident. Most significantly, however, we found both major mitochondrial lineages co-occurring at C-25, Kudamurra Well (samples in blue in the NJ tree,
|
||
<figureCitation id="13322A46FFB7D96FF21E68690992F9AB" box="[1354,1422,1602,1626]" captionStart="FIGURE 3" captionStartId="4.[151,250,1425,1448]" captionTargetBox="[151,1435,373,1404]" captionTargetId="figure@4.[151,1436,373,1405]" captionTargetPageId="4" captionText="FIGURE 3. Milyeringa COI neighbor-joining tree. Collection localities of samples are in parentheses following the sample number. Red indicates the M. brooksi haplotype clade and blue indicates those individuals collected from Kudumurra Well (C- 25)." httpUri="https://zenodo.org/record/220761/files/figure.png" pageId="12" pageNumber="147">Fig. 3</figureCitation>
|
||
). This population shows no concordant allozyme differentiation between individuals of ‘
|
||
<taxonomicName id="4C094D40FFB7D96FF335684C08B1F98F" box="[1121,1197,1639,1662]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="147" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB7D96FF335684C08B1F98F" box="[1121,1197,1639,1662]" italics="true" pageId="12" pageNumber="147">veritas</emphasis>
|
||
</taxonomicName>
|
||
’ haplotype and those of ‘
|
||
<taxonomicName id="4C094D40FFB7D96FF7EA68A00D0DF952" box="[190,273,1675,1699]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="147" phylum="Chordata" rank="species" species="brooksi">
|
||
<emphasis id="B97DEAD1FFB7D96FF7EA68A00D0DF952" box="[190,273,1675,1699]" italics="true" pageId="12" pageNumber="147">brooksi</emphasis>
|
||
</taxonomicName>
|
||
’ haplotype (Adams and Humphreys 1993), a finding consistent with the presence of free gene flow and a single species,
|
||
<taxonomicName id="4C094D40FFB7D96FF60768850E3EF937" box="[339,546,1710,1734]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="147" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB7D96FF60768850E3EF937" box="[339,546,1710,1734]" italics="true" pageId="12" pageNumber="147">Milyeringa veritas</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136548FFB7D96EF79D68F90D28FCC6" lastPageId="13" lastPageNumber="148" pageId="12" pageNumber="147" type="distribution">
|
||
<paragraph id="8BB636C3FFB7D96FF79D68F90FC5F91B" blockId="12.[151,1437,1422,2022]" box="[201,985,1746,1771]" pageId="12" pageNumber="147">
|
||
<emphasis id="B97DEAD1FFB7D96FF79D68F90D7EF91A" bold="true" box="[201,354,1746,1771]" pageId="12" pageNumber="147">Distribution.</emphasis>
|
||
Known from Cape Range peninsula, Western
|
||
<collectingCountry id="F31E7653FFB7D96FF43E68F90FC8F91B" box="[874,980,1746,1770]" name="Australia" pageId="12" pageNumber="147">Australia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB636C3FFB7D96FF79D68DC0DF1F817" blockId="12.[151,1437,1422,2022]" pageId="12" pageNumber="147">
|
||
<emphasis id="B97DEAD1FFB7D96FF79D68DC0D31F8FE" bold="true" box="[201,301,1783,1807]" pageId="12" pageNumber="147">Ecology.</emphasis>
|
||
<taxonomicName id="4C094D40FFB7D96FF66868DD0E0FF8FF" box="[316,531,1782,1806]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="147" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB7D96FF66868DD0E0FF8FF" box="[316,531,1782,1806]" italics="true" pageId="12" pageNumber="147">Milyeringa veritas</emphasis>
|
||
</taxonomicName>
|
||
is listed as vulnerable under the
|
||
<emphasis id="B97DEAD1FFB7D96FF4EE68DC0D86F8C2" italics="true" pageId="12" pageNumber="147">Environment Protection and Biodiversity Conservation Act 1999</emphasis>
|
||
(Commonwealth) and as endangered under Schedule One of the
|
||
<emphasis id="B97DEAD1FFB7D96FF32E69300CD3F8A7" italics="true" pageId="12" pageNumber="147">Wildlife Conservation Act 1950</emphasis>
|
||
(Western
|
||
<collectingCountry id="F31E7653FFB7D96FF61869150DA9F8A7" box="[332,437,1854,1878]" name="Australia" pageId="12" pageNumber="147">Australia</collectingCountry>
|
||
) owing to its small geographic distribution, low populations and vulnerability of its habitat. Two locations in which the species occurs (Bundera Sinkhole and Camerons Cave) are also listed as threatened communities (by the Western Australian Government’s Department of Environment and Conservation). Its distribution lies within the Ningaloo Coast World Heritage Area. The conservation status is discussed by Romero and Vanselow (2000).
|
||
</paragraph>
|
||
<paragraph id="8BB636C3FFB6D96EF7936EBC0EF9FE3E" blockId="13.[151,1437,151,2012]" pageId="13" pageNumber="148">
|
||
The sparse knowledge of the biology of
|
||
<taxonomicName id="4C094D40FFB6D96EF5F86EB30F34FF41" box="[684,808,152,176]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97DEAD1FFB6D96EF5F86EB30F34FF41" box="[684,808,152,176]" italics="true" pageId="13" pageNumber="148">Milyeringa</emphasis>
|
||
</taxonomicName>
|
||
was summarised by Humphreys (2001a).
|
||
<taxonomicName id="4C094D40FFB6D96EF24B6EB30CFFFF25" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB6D96EF24B6EB30CFFFF25" italics="true" pageId="13" pageNumber="148">Milyeringa veritas</emphasis>
|
||
</taxonomicName>
|
||
inhabits the coastal groundwaters of the carbonate karst of the Cape Range peninsula that projects onto the North West Shelf of
|
||
<collectingCountry id="F31E7653FFB6D96EF62B6EF40DF5FF06" box="[383,489,223,247]" name="Australia" pageId="13" pageNumber="148">Australia</collectingCountry>
|
||
. It occurs on the coastal plain and foothills of the peninsula between
|
||
<quantity id="4CF19B26FFB6D96EF3A06EF4095EFF06" box="[1268,1346,223,247]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="148" unit="km" value="0.4">0.4 km</quantity>
|
||
and 4.0 km of the coast. Closer to the coast is an anchialine system, characteristically with marked hydrogeochemical stratification with depth (Seymour
|
||
<emphasis id="B97DEAD1FFB6D96EF5776F020E49FEB1" box="[547,597,296,320]" italics="true" pageId="13" pageNumber="148">et al</emphasis>
|
||
. 2007), but grading to freshwater inland. The fish occupy a wide range of salinity, from freshwater inland to fully marine in the depths of the anchialine system (Humphreys 2001a). Otolith microchemistry indicates that individuals may inhabit waters of widely different salinity through their lives (Humphreys
|
||
<emphasis id="B97DEAD1FFB6D96EF67D6FBE0D46FE5D" box="[297,346,404,428]" italics="true" pageId="13" pageNumber="148">et al</emphasis>
|
||
. 2006). They seem to be opportunistic feeders ingesting prey of epigean origin as well as obligate groundwater species (Humphreys & Feinberg 1995).
|
||
</paragraph>
|
||
<paragraph id="8BB636C3FFB6D96EF79D6FF70D28FCC6" blockId="13.[151,1437,151,2012]" pageId="13" pageNumber="148">
|
||
Anchialine habitats of the
|
||
<typeStatus id="54B28861FFB6D96EF5566FF60E2EFE04" box="[514,562,477,501]" pageId="13" pageNumber="148">type</typeStatus>
|
||
inhabited by
|
||
<taxonomicName id="4C094D40FFB6D96EF58D6FF70F49FE05" box="[729,853,476,500]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97DEAD1FFB6D96EF58D6FF70F49FE05" box="[729,853,476,500]" italics="true" pageId="13" pageNumber="148">Milyeringa</emphasis>
|
||
</taxonomicName>
|
||
are noteworthy for their very diverse stygiobiont crustacean fauna, comprising a characteristic assemblage of higher taxa, the structure of which is highly predictable, frequently extending to the generic composition (Wagner 1994), however far apart in the world they occur (Jaume
|
||
<emphasis id="B97DEAD1FFB6D96EF6606C620D79FD91" box="[308,357,584,608]" italics="true" pageId="13" pageNumber="148">et al</emphasis>
|
||
. 2001); most of their members represent biogeographic and/or phylogenetic relicts (Iliffe 1992). As such, the anchialine fauna of the Cape Range peninsula has an assemblage comparable to anchialine systems of the North Atlantic comprising, inter alia, the higher taxa Remipedia,
|
||
<taxonomicName id="4C094D40FFB6D96EF4846CA40881FD56" box="[976,1181,655,679]" class="Malacostraca" kingdom="Animalia" order="Thermosbaenacea" pageId="13" pageNumber="148" phylum="Arthropoda" rank="order">Thermosbaenacea</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C094D40FFB6D96EF3F96CA40984FD56" authority="(Decapoda)" baseAuthorityName="Decapoda" box="[1197,1432,655,679]" class="Malacostraca" family="Atyidae" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="148" phylum="Arthropoda" rank="family">Atyidae (Decapoda)</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C094D40FFB6D96EF7C36C9F0DB7FD3D" authority="(Amphipoda)" baseAuthorityName="Amphipoda" box="[151,427,692,716]" class="Malacostraca" family="Hadziidae" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="148" phylum="Arthropoda" rank="family">Hadziidae (Amphipoda)</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C094D40FFB6D96EF6EC6C9F0EAEFD3D" authority="(Isopoda)" baseAuthorityName="Isopoda" box="[440,690,692,716]" class="Malacostraca" family="Cirolanidae" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="148" phylum="Arthropoda" rank="family">Cirolanidae (Isopoda)</taxonomicName>
|
||
, Thaumatocypridae (Ostracoda), and the copepod taxa
|
||
<taxonomicName id="4C094D40FFB6D96EF27E6C9F0980FD3D" box="[1322,1436,692,716]" class="Maxillopoda" kingdom="Animalia" order="Calanoida" pageId="13" pageNumber="148" phylum="Arthropoda" rank="order">Calanoida</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C094D40FFB6D96EF7CA6CFC0D5AFD1E" box="[158,326,727,751]" class="Maxillopoda" family="Epacteriscidae" kingdom="Animalia" order="Calanoida" pageId="13" pageNumber="148" phylum="Arthropoda" rank="family">Epacteriscidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C094D40FFB6D96EF6366CFC0E16FD1E" box="[354,522,727,751]" class="Maxillopoda" family="Ridgewayiidae" kingdom="Animalia" order="Calanoida" pageId="13" pageNumber="148" phylum="Arthropoda" rank="family">Ridgewayiidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C094D40FFB6D96EF5356CFC0F24FD1E" box="[609,824,727,751]" class="Maxillopoda" family="Pseudocyclopiidae" kingdom="Animalia" order="Calanoida" pageId="13" pageNumber="148" phylum="Arthropoda" rank="family">Pseudocyclopiidae</taxonomicName>
|
||
),
|
||
<taxonomicName id="4C094D40FFB6D96EF40F6CFC08DFFD1E" box="[859,1219,727,751]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Misophrioida" pageId="13" pageNumber="148" phylum="Arthropoda" rank="order">Misophrioida (Speleophriidae)</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C094D40FFB6D96EF24F6CFC0D4BFCE5" authority="(Halicyclopinae)" baseAuthorityName="Halicyclopinae" class="Maxillopoda" genus="Cyclopoida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="148" phylum="Arthropoda" rank="genus">Cyclopoida (Halicyclopinae)</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C094D40FFB6D96EF6306CD70E2BFCE5" box="[356,567,764,788]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB6D96EF6306CD70E2BFCE5" box="[356,567,764,788]" italics="true" pageId="13" pageNumber="148">Milyeringa veritas</emphasis>
|
||
</taxonomicName>
|
||
is sympatric with the groundwater synbranchid eel
|
||
<taxonomicName id="4C094D40FFB6D96EF3C36CD70D33FCC6" authority="Mees, 1962" authorityName="Mees" authorityYear="1962" class="Actinopterygii" family="Synbranchidae" genus="Ophisternon" kingdom="Animalia" order="Synbranchiformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="candidum">
|
||
<emphasis id="B97DEAD1FFB6D96EF3C36CD70987FCE5" box="[1175,1435,764,788]" italics="true" pageId="13" pageNumber="148">Ophisternon candidum</emphasis>
|
||
(Mees, 1962)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136548FFB6D96EF79D6D6F0F53F82D" pageId="13" pageNumber="148" type="discussion">
|
||
<paragraph id="8BB636C3FFB6D96EF79D6D6F0FB6FBA9" blockId="13.[151,1437,151,2012]" pageId="13" pageNumber="148">
|
||
<emphasis id="B97DEAD1FFB6D96EF79D6D6F0D21FCAD" bold="true" box="[201,317,836,860]" pageId="13" pageNumber="148">Remarks.</emphasis>
|
||
<taxonomicName id="4C094D40FFB6D96EF6176D6F0E0BFCAD" box="[323,535,836,860]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="brooksi">
|
||
<emphasis id="B97DEAD1FFB6D96EF6176D6F0E0BFCAD" box="[323,535,836,860]" italics="true" pageId="13" pageNumber="148">Milyeringa brooksi</emphasis>
|
||
</taxonomicName>
|
||
was described by Chakrabarty (2010), who separated his species from
|
||
<taxonomicName id="4C094D40FFB6D96EF2786D6E0987FCAD" box="[1324,1435,837,860]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB6D96EF2786D6E0987FCAD" box="[1324,1435,837,860]" italics="true" pageId="13" pageNumber="148">M. veritas</emphasis>
|
||
</taxonomicName>
|
||
mainly by its having 10–12 vertical lines of sensory papillae along the body and by “molecular synapomorphies” (which refer to changes in nucleotide position in three genes: ND2, CytB and COI). Chakrabarty also considered that
|
||
<taxonomicName id="4C094D40FFB6D96EF79D6D9A0D5EFC39" box="[201,322,944,968]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="brooksi">
|
||
<emphasis id="B97DEAD1FFB6D96EF79D6D9A0D5EFC39" box="[201,322,944,968]" italics="true" pageId="13" pageNumber="148">M. brooksi</emphasis>
|
||
</taxonomicName>
|
||
was a smaller fish than
|
||
<taxonomicName id="4C094D40FFB6D96EF50D6D9A0ED7FC39" box="[601,715,945,968]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB6D96EF50D6D9A0ED7FC39" box="[601,715,945,968]" italics="true" pageId="13" pageNumber="148">M. veritas</emphasis>
|
||
</taxonomicName>
|
||
(the largest specimen given as
|
||
<quantity id="4CF19B26FFB6D96EF3666D8408B9FC39" box="[1074,1189,943,968]" metricMagnitude="-2" metricUnit="m" metricValue="3.8329999999999997" pageId="13" pageNumber="148" unit="mm" value="38.33">38.33 mm</quantity>
|
||
SL versus
|
||
<quantity id="4CF19B26FFB6D96EF2726D840969FC36" box="[1318,1397,943,967]" metricMagnitude="0" metricUnit="m" metricValue="1.33604" pageId="13" pageNumber="148" unit="in" value="52.6">52.6 in</quantity>
|
||
<taxonomicName id="4C094D40FFB6D96EF22A6D9A0CFFFC1D" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB6D96EF22A6D9A0CFFFC1D" italics="true" pageId="13" pageNumber="148">M. veritas</emphasis>
|
||
</taxonomicName>
|
||
), had a tubular posterior nostril with a skin flap (versus a simple round nostril in
|
||
<taxonomicName id="4C094D40FFB6D96EF3E16DFE0937FC1D" box="[1205,1323,981,1004]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB6D96EF3E16DFE0937FC1D" box="[1205,1323,981,1004]" italics="true" pageId="13" pageNumber="148">M. veritas</emphasis>
|
||
</taxonomicName>
|
||
) and had conspicuous papillae on the dorsal surface of the head (versus a “variable condition” in
|
||
<taxonomicName id="4C094D40FFB6D96EF33C6DD208C4FBE1" box="[1128,1240,1017,1040]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB6D96EF33C6DD208C4FBE1" box="[1128,1240,1017,1040]" italics="true" pageId="13" pageNumber="148">M. veritas</emphasis>
|
||
</taxonomicName>
|
||
). The nostril size and shape in the
|
||
<typeStatus id="54B28861FFB6D96EF60D6A370DA6FBC5" box="[345,442,1052,1076]" pageId="13" pageNumber="148" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C094D40FFB6D96EF6B76A360E41FBC5" box="[483,605,1052,1076]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="brooksi">
|
||
<emphasis id="B97DEAD1FFB6D96EF6B76A360E41FBC5" box="[483,605,1052,1076]" italics="true" pageId="13" pageNumber="148">M. brooksi</emphasis>
|
||
</taxonomicName>
|
||
and the two WAM
|
||
<typeStatus id="54B28861FFB6D96EF4146A360FB7FBC4" box="[832,939,1053,1077]" pageId="13" pageNumber="148" type="paratype">paratypes</typeStatus>
|
||
are basically the same as in
|
||
<emphasis id="B97DEAD1FFB6D96EF3A16A360973FBC5" box="[1269,1391,1053,1076]" italics="true" pageId="13" pageNumber="148">
|
||
<taxonomicName id="4C094D40FFB6D96EF3A16A360977FBC5" box="[1269,1387,1053,1076]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="veritas">M. veritas</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
We cannot distinguish his specimens as a separate species from
|
||
<taxonomicName id="4C094D40FFB6D96EF4666A6A0FBFFBA9" box="[818,931,1089,1112]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB6D96EF4666A6A0FBFFBA9" box="[818,931,1089,1112]" italics="true" pageId="13" pageNumber="148">M. veritas</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB636C3FFB6D96EF79D6A4E097CFAFD" blockId="13.[151,1437,151,2012]" pageId="13" pageNumber="148">
|
||
We observed features in
|
||
<taxonomicName id="4C094D40FFB6D96EF6816A4F0E53FB8D" box="[469,591,1124,1148]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97DEAD1FFB6D96EF6816A4F0E53FB8D" box="[469,591,1124,1148]" italics="true" pageId="13" pageNumber="148">Milyeringa</emphasis>
|
||
</taxonomicName>
|
||
specimens that were missed by Chakrabarty (2010). He indicated that he had
|
||
<specimenCount id="9D0FFD4AFFB6D96EF7C36AA30D30FB51" box="[151,300,1160,1184]" pageId="13" pageNumber="148" type="generic">16 specimens</specimenCount>
|
||
of
|
||
<taxonomicName id="4C094D40FFB6D96EF61B6AA20DA2FB51" box="[335,446,1161,1184]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB6D96EF61B6AA20DA2FB51" box="[335,446,1161,1184]" italics="true" pageId="13" pageNumber="148">M. veritas</emphasis>
|
||
</taxonomicName>
|
||
(sizes given as
|
||
<quantity id="4CF19B26FFB6D96EF53E6AA30F15FB51" box="[618,777,1160,1184]" metricMagnitude="-2" metricUnit="m" metricValue="4.165" metricValueMax="5.26" metricValueMin="3.07" pageId="13" pageNumber="148" unit="mm" value="41.65" valueMax="52.6" valueMin="30.7">30.7–52.6 mm</quantity>
|
||
SL) to compare with his seven specimens of
|
||
<taxonomicName id="4C094D40FFB6D96EF3AD6AA20972FB51" box="[1273,1390,1160,1184]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="brooksi">
|
||
<emphasis id="B97DEAD1FFB6D96EF3AD6AA20972FB51" box="[1273,1390,1160,1184]" italics="true" pageId="13" pageNumber="148">M. brooksi</emphasis>
|
||
</taxonomicName>
|
||
. He apparently did not observe the vertical rows of sensory papillae along the side of the body in
|
||
<taxonomicName id="4C094D40FFB6D96EF3E06A86093AFB35" box="[1204,1318,1197,1220]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB6D96EF3E06A86093AFB35" box="[1204,1318,1197,1220]" italics="true" pageId="13" pageNumber="148">M. veritas</emphasis>
|
||
</taxonomicName>
|
||
which are present in specimens of all sizes and locations. He also describes the head of
|
||
<taxonomicName id="4C094D40FFB6D96EF3576AFA0867FB19" box="[1027,1147,1232,1256]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="brooksi">
|
||
<emphasis id="B97DEAD1FFB6D96EF3576AFA0867FB19" box="[1027,1147,1232,1256]" italics="true" pageId="13" pageNumber="148">M. brooksi</emphasis>
|
||
</taxonomicName>
|
||
as being naked, although scales can be observed on the holotype’s predorsal region, extending forward to above the preopercular margin.
|
||
</paragraph>
|
||
<paragraph id="8BB636C3FFB6D96EF79D6B330D07F9A1" blockId="13.[151,1437,151,2012]" pageId="13" pageNumber="148">
|
||
Chakrabarty’s 2010 DNA sequence data for
|
||
<taxonomicName id="4C094D40FFB6D96EF58B6B330F47FAC1" box="[735,859,1304,1328]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97DEAD1FFB6D96EF58B6B330F47FAC1" box="[735,859,1304,1328]" italics="true" pageId="13" pageNumber="148">Milyeringa</emphasis>
|
||
</taxonomicName>
|
||
were derived from eight specimens from seven populations. The maximum genetic distance found between the two major mitochondrial lineages that represent
|
||
<taxonomicName id="4C094D40FFB6D96EF2296B160CFFFA89" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB6D96EF2296B160CFFFA89" italics="true" pageId="13" pageNumber="148">M. veritas</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C094D40FFB6D96EF6746B4A0D87FA89" box="[288,411,1376,1400]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="brooksi">
|
||
<emphasis id="B97DEAD1FFB6D96EF6746B4A0D87FA89" box="[288,411,1376,1400]" italics="true" pageId="13" pageNumber="148">M. brooksi</emphasis>
|
||
</taxonomicName>
|
||
was 3.2%. The small sample size and low genetic distance (in comparison with genetic distances found among other eleotrid species; e.g. Stevens & Hicks 2009, Thacker 2009) are themselves problematic but, as highlighted in recent discussions of the use of barcoding (e.g. Lohse 2009; Petit & Excoffier 2009), the use of markers from a single uniparentally inherited genome for species delimitation is risky and the inference of a species tree from what is effectively a single gene tree, inadvisable. Multilocus assignment methods, using unlinked markers, have considerably more power in this respect and can also inform on population admixtures.
|
||
</paragraph>
|
||
<paragraph id="8BB636C3FFB6D96EF79368770898F8D6" blockId="13.[151,1437,151,2012]" pageId="13" pageNumber="148">
|
||
Humphreys and Adams (1991) and Adams and Humphreys (1993) subjected seven populations of
|
||
<taxonomicName id="4C094D40FFB6D96EF24B68770CFFF969" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB6D96EF24B68770CFFF969" italics="true" pageId="13" pageNumber="148">Milyeringa veritas</emphasis>
|
||
</taxonomicName>
|
||
(n = 24 & 29, respectively) to comprehensive allozyme analyses at 43 loci and demonstrated that, although allele frequencies and distributions indicated significant population sub-structuring on the peninsula, overall levels of differentiation and the pattern of allele distribution were consistent with the presence of a single biological species on Cape Range peninsula. The mitochondrial data presented here affirm the view established in the 1990s and provide additional evidence for the distinctiveness of the species from Barrow Island.
|
||
</paragraph>
|
||
<paragraph id="8BB636C3FFB6D96EF79369180F53F82D" blockId="13.[151,1437,151,2012]" pageId="13" pageNumber="148">
|
||
<emphasis id="B97DEAD1FFB6D96EF79369180D43F8BD" bold="true" box="[199,351,1843,1868]" pageId="13" pageNumber="148">Conclusions.</emphasis>
|
||
Cape Range peninsula and Barrow Island are home to a distinctive and unique stygofauna including cave-dwelling fishes. The description of a new and highly divergent
|
||
<taxonomicName id="4C094D40FFB6D96EF35369730898F881" box="[1031,1156,1880,1904]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97DEAD1FFB6D96EF35369730898F881" box="[1031,1156,1880,1904]" italics="true" pageId="13" pageNumber="148">Milyeringa</emphasis>
|
||
</taxonomicName>
|
||
from within the range of an already restricted taxon, adds significantly to the regional fauna with matching conservation implications, in the face of increasing and broad-scale mineral exploration. We recognise that further systematic work may be required to understand the evolutionary heritage of
|
||
<taxonomicName id="4C094D40FFB6D96EF52769EF0EECF82D" box="[627,752,1988,2012]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="148" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97DEAD1FFB6D96EF52769EF0EECF82D" box="[627,752,1988,2012]" italics="true" pageId="13" pageNumber="148">Milyeringa</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136548FFB5D96DF7C36EBC0871FE76" pageId="14" pageNumber="149" type="description">
|
||
<paragraph id="8BB636C3FFB5D96DF7C36EBC0D63FF41" blockId="14.[151,383,151,176]" box="[151,383,151,176]" pageId="14" pageNumber="149">
|
||
<emphasis id="B97DEAD1FFB5D96DF7C36EBC0D63FF41" bold="true" box="[151,383,151,176]" pageId="14" pageNumber="149">Acknowledgements</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB636C3FFB5D96DF7C36EF40871FE76" blockId="14.[151,1436,223,392]" pageId="14" pageNumber="149">
|
||
Our thanks to Peter Austin, Australian Minerals Research Centre, CSIRO and the late Peter Blias of SAMA who created microtomography scans and reconstructions of
|
||
<taxonomicName id="4C094D40FFB5D96DF45D6F2E0F61FEED" box="[777,893,261,284]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="149" phylum="Chordata" rank="species" species="justitia">
|
||
<emphasis id="B97DEAD1FFB5D96DF45D6F2E0F61FEED" box="[777,893,261,284]" italics="true" pageId="14" pageNumber="149">M. justitia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C094D40FFB5D96DF4E16F2E0834FEED" box="[949,1064,261,284]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="149" phylum="Chordata" rank="species" species="veritas">
|
||
<emphasis id="B97DEAD1FFB5D96DF4E16F2E0834FEED" box="[949,1064,261,284]" italics="true" pageId="14" pageNumber="149">M. veritas</emphasis>
|
||
</taxonomicName>
|
||
. Our thanks also to Doug Hoese and Tony Gill for their generous sharing of data and valuable discussions on
|
||
<taxonomicName id="4C094D40FFB5D96DF3716F0308BDFEB1" box="[1061,1185,296,320]" class="Actinopterygii" family="Eleotridae" genus="Milyeringa" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="149" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97DEAD1FFB5D96DF3716F0308BDFEB1" box="[1061,1185,296,320]" italics="true" pageId="14" pageNumber="149">Milyeringa</emphasis>
|
||
</taxonomicName>
|
||
osteology. Thanks to Michael Hammer for his discussions with HKL on some points of genetic data. Specimen collection on Barrow Island was facilitated by Western Australian Petroleum Pty Ltd, and Chevron
|
||
<collectingCountry id="F31E7653FFB5D96DF4AB6F440875FE76" box="[1023,1129,367,391]" name="Australia" pageId="14" pageNumber="149">Australia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |