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<document id="4630F3B26EF68F8E4F6EA1EB806C4130" ID-DOI="10.5281/zenodo.8222206" ID-ISSN="1536-9307" ID-Zenodo-Dep="8222206" ID-ZooBank="E7C4D357-A2FF-4F8D-B93D-1B9B6CFE3115" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1691436641308" checkinUser="felipe" docAuthor="Teruel, Rolando, Kovařík, František, Lowe, Graeme &amp; Šťáhlavský, František" docDate="2023" docId="03A087CAFFA43216FF5904ED1954DDF8" docLanguage="en" docName="Euscorpius.375.1-22.pdf" docOrigin="Euscorpius 375" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Megacormus seductus Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský 2023, sp. n." docType="treatment" docVersion="3" lastPageNumber="16" masterDocId="FF99FFB2FFAE3207FFA602031C7CD827" masterDocTitle="Two new species of the remarkable scorpion genus Megacormus Karsch, 1881 (Scorpiones: Euscorpiidae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="9" updateTime="1699551157245" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="4E775258A9A7ED25E644ACF27A833198">Two new species of the remarkable scorpion genus Megacormus Karsch, 1881 (Scorpiones: Euscorpiidae)</mods:title>
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<mods:namePart id="8108FA5FBF6B91573C92E7DF5692C4E1">Teruel, Rolando</mods:namePart>
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<mods:title id="E0E0C351B78EBDDF08F6ECE0333EA056">Euscorpius</mods:title>
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<taxonomicName id="4C094D5FFFA4320DFF5904ED1E5ADF2D" authority="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský, 2023" authorityName="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2023" box="[255,550,1773,1802]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="seductus" status="sp. nov.">Megacormus seductus</taxonomicName>
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<paragraph id="8BB636DCFFA4320DFFD505121E86DF6E" blockId="10.[115,769,1773,1900]" pageId="10" pageNumber="9">
(
<figureCitation id="13322A59FFA4320DFFDC05121D78DF0E" box="[122,260,1809,1833]" captionStart-0="Figures 12" captionStart-1="Figures 36" captionStart-2="Figures 716" captionStart-3="Figures 1726" captionStart-4="Figures 2734" captionStart-5="Figures 3542" captionStart-6="Figure 43" captionStartId-0="3.[115,192,2017,2039]" captionStartId-1="6.[113,190,1981,2003]" captionStartId-2="8.[116,193,1960,1982]" captionStartId-3="11.[115,192,1969,1991]" captionStartId-4="12.[116,194,2001,2023]" captionStartId-5="14.[113,190,1957,1979]" captionStartId-6="15.[115,183,2026,2048]" captionTargetBox-0="[113,1474,251,1990]" captionTargetBox-1="[170,1395,231,1967]" captionTargetBox-2="[132,1474,239,1928]" captionTargetBox-3="[153,1471,240,1920]" captionTargetBox-4="[113,1474,217,1991]" captionTargetBox-5="[125,1450,263,1942]" captionTargetBox-6="[113,1474,234,2015]" captionTargetId-0="figure-31@3.[113,1474,251,1990]" captionTargetId-1="figure-53@6.[113,1474,219,1974]" captionTargetId-2="figure-72@8.[113,1474,226,1953]" captionTargetId-3="figure-55@11.[115,1476,221,1958]" captionTargetId-4="figure-62@12.[113,1474,217,1994]" captionTargetId-5="figure-78@14.[113,1474,230,1948]" captionTargetId-6="figure-22@15.[113,1474,231,2017]" captionTargetPageId-0="3" captionTargetPageId-1="6" captionTargetPageId-2="8" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="14" captionTargetPageId-6="15" captionText-0="Figures 12. Megacormus seductus sp. n., male holotype (1) and female paratype with a juvenile (2) in vivo habitus." captionText-1="Figures 36: Megacormus seductus sp. n. Figures 34. Male holotype in dorsal (3) and ventral (4) views. Figures 56. Female paratype in dorsal (5) and ventral (6) views. Scale bars: 10 mm." captionText-2="Figures 716: Megacormus seductus sp. n., male holotype, pedipalp chela dorsal (7), external (8) and ventral (9) views, patella dorsal (10), external (11) and ventral (12) views, femur and trochanter dorsal (13) and ventral (14) views, movable (15) and fixed (16) fingers dentition. Movable finger (15) is under UV light, other figures are under white light." captionText-3="Figures 1726: Megacormus seductus sp. n., female paratype, pedipalp chela dorsal (17), external (18) and ventral (19) views, patella dorsal (20), external (21) and ventral (22) views, femur and trochanter dorsal (23) and ventral (24) views, movable (25) and fixed (26) fingers dentition." captionText-4="Figures 2734: Megacormus seductus sp. n. Figures 2728. Male holotype, carapace and tergites IIV (27) and coxosternal area and sternites (28). Figures 2934. Female paratype, carapace and tergites IIV (29), coxosternal area and sternites (30), and left legs IIV, retrolateral aspect (3134)." captionText-5="Figures 3542. Megacormus seductus sp. n. Figures 35, 3739. Male holotype, telson in lateral (35), metasoma and telson in lateral (37), dorsal (38) and ventral (39) views. Figures 36, 4042. Female paratype, telson in lateral (36), metasoma and telson in lateral (40), dorsal (41) and ventral (42) views. Scale bars: 10 mm (3739, 4042)." captionText-6="Figure 43. Megacormus seductus sp. n., habitat at type locality." figureDoi-0="http://doi.org/10.5281/zenodo.8222208" figureDoi-1="http://doi.org/10.5281/zenodo.8222210" figureDoi-2="http://doi.org/10.5281/zenodo.8222212" figureDoi-3="http://doi.org/10.5281/zenodo.8222214" figureDoi-4="http://doi.org/10.5281/zenodo.8222216" figureDoi-5="http://doi.org/10.5281/zenodo.8222218" figureDoi-6="http://doi.org/10.5281/zenodo.8222222" httpUri-0="https://zenodo.org/record/8222208/files/figure.png" httpUri-1="https://zenodo.org/record/8222210/files/figure.png" httpUri-2="https://zenodo.org/record/8222212/files/figure.png" httpUri-3="https://zenodo.org/record/8222214/files/figure.png" httpUri-4="https://zenodo.org/record/8222216/files/figure.png" httpUri-5="https://zenodo.org/record/8222218/files/figure.png" httpUri-6="https://zenodo.org/record/8222222/files/figure.png" pageId="10" pageNumber="9">Figures 143</figureCitation>
, 5657, 60, 6465, 7273, 7879, 82,
<tableCitation id="C68B0367FFA4320DFD2C05121E85DF0E" box="[650,761,1809,1833]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStartId-0="4.[113,169,1006,1027]" captionStartId-1="20.[111,167,905,926]" captionStartId-2="20.[114,170,1256,1277]" captionText-0="Table 1. Morphometric data for types of Megacormus orizaba sp. n. and Megacormus seductus sp. n. Abbreviations: length (L), width (W), in carapace it corresponds to median width), depth (D)." captionText-1="Table 2. Variation in number of tines on the crown-like structure (cls) of the hemispermatophore of Megacormus species. Data with DNA No. and locality code were obtained from examined hemispermatophores. Other data were obtained from published descriptions or illustrations (1Francke, 1979; 2González-Santillán &amp; Alvarez-Padilla, 2015; 3Sissom, 1994: fig. 8; 4González-Santillán et al., 2017)." captionText-2="Table 3. Pairwise two-sided non-parametric tests of equality of tine counts of crown-like structure (cls) of hemispermatophores between five different Megacormus species. Below diagonal values: z-values from Dunns test; thresholds for significantly different medians were: 1.9600 (regular, shaded), 2.8070 (Bonferroni correction, bold). Above diagonal values: P-values from likelihood ratio tests (P &lt;α = 0.05 shaded; without family-wise correction)." pageId="10" pageNumber="9">Tables 13</tableCitation>
)
<emphasis id="B97DEACEFFA4320DFFDD05321E86DF6E" bold="true" box="[123,762,1841,1865]" pageId="10" pageNumber="9">http://zoobank.org/urn:lsid:zoobank.org:act:4A7487B6-</emphasis>
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<emphasis id="B97DEACEFFA4320DFF5D05571E05DF4B" bold="true" box="[251,633,1876,1900]" pageId="10" pageNumber="9">78AF-4823-A0F5-58580EF8D89B</emphasis>
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<treatmentCitation id="0AA810CDFFA4320DFFD705961EFADF8B" author="SISSOM" box="[113,646,1940,1964]" page="265" pageId="10" pageNumber="9" year="1994">
<taxonomicName id="4C094D5FFFA4320DFFD705961EFADF8B" ID-CoL="72W9F" authority=": Sissom, 1994: 265" authorityName=": Sissom" authorityPageNumber="265" authorityYear="1994" box="[113,646,1940,1964]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="segmentatus">
<emphasis id="B97DEACEFFA4320DFFD705961DF3DF8B" box="[113,399,1941,1964]" italics="true" pageId="10" pageNumber="9">Megacormus segmentatus</emphasis>
:
<bibRefCitation id="EF984B2DFFA4320DFE0305971EFADF8B" author="SISSOM" box="[421,646,1940,1964]" pageId="10" pageNumber="9" pagination="265 - 272" refId="ref14012" refString="SISSOM, W. D. 1994. Systematic studies on the genus Megacormus (Scorpiones, Chactidae, Megacorminae), with descriptions of a new species from Oaxaca, Mexico and of the male of Megacormus segmentatus Pocock. Insecta Mundi, 8: 265 - 272." type="journal article" year="1994">Sissom, 1994: 265</bibRefCitation>
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, 268, 270 (misidentification: specimen and record from Los Tuxtlas).
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TYPEMATERIAL.
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<geoCoordinate id="EE3D501BFFA4320DFC9603C91FBBD9C5" box="[816,967,458,482]" degrees="95" direction="west" minutes="04" orientation="longitude" pageId="10" pageNumber="9" precision="1" seconds="15.7" value="-95.07103">95°04'15.7''W</geoCoordinate>
,
<elevation id="0024D1EFFFA4320DFC7403C81823D9C4" box="[978,1119,459,483]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="9" unit="m" value="200.0">
<quantity id="4CF19B39FFA4320DFC7403C8186BD9C5" box="[978,1047,459,483]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="9" unit="m" value="200.0">200 m</quantity>
a. s. l.
</elevation>
, broadleaf rainforest on volcanic soil, inside rotten logs and stumps,
<date id="FFB7101CFFA4320DFB1603E8195FDA24" box="[1200,1315,491,515]" pageId="10" pageNumber="9" value="2022-10-23">
<collectingDate id="EFF3E9F4FFA4320DFB1603E8195FDA24" box="[1200,1315,491,515]" pageId="10" pageNumber="9" value="2022-10-23">23.X.2022</collectingDate>
</date>
, leg.
<collectionCode id="ED18AE19FFA4320DFAF803E8190FDA25" box="[1374,1395,491,514]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="9">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B613C81FFA4320DFAD803E818B6DA04" collectionCode="FKCP" collectorName="No." country="Mexico" location="Teruel" pageId="10" pageNumber="9" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<location id="8ED66007FFA4320DFAD803E819BEDA24" LSID="urn:lsid:plazi:treatment:03A087CAFFA43216FF5904ED1954DDF8:8ED66007FFA4320DFAD803E819BEDA24" box="[1406,1474,491,515]" country="Mexico" name="Teruel" pageId="10" pageNumber="9" stateProvince="Veracruz de Ignacio de la Llave">Teruel</location>
et al.,
<specimenCount id="9D0FFD55FFA4320DFCD400081FEFDA04" box="[882,915,523,547]" pageId="10" pageNumber="9" type="male">1♂</specimenCount>
(
<typeStatus id="54B2887EFFA4320DFC070008187FDA04" box="[929,1027,523,547]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
,
<collectorName id="26FC530AFFA4320DFBAB00081849DA05" box="[1037,1077,523,546]" pageId="10" pageNumber="9">No.</collectorName>
2460),
<collectionCode id="ED18AE19FFA4320DFB21000818B6DA04" box="[1159,1226,523,547]" pageId="10" pageNumber="9">FKCP</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B613C81FFA4320DFB7200081F93DA64" collectionCode="FKCP" collectorName="No." country="Mexico" pageId="10" pageNumber="9" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<specimenCount id="9D0FFD55FFA4320DFB7200081885DA04" box="[1236,1273,523,547]" pageId="10" pageNumber="9" type="male">1♂</specimenCount>
<specimenCount id="9D0FFD55FFA4320DFB5F0008196BDA04" box="[1273,1303,523,547]" pageId="10" pageNumber="9" type="female">1♀</specimenCount>
(paratopotypes,
<collectorName id="26FC530AFFA4320DFC9600281F24DA65" box="[816,856,555,578]" pageId="10" pageNumber="9">No.</collectorName>
2461),
<collectionCode id="ED18AE19FFA4320DFC0A00281F93DA64" box="[940,1007,555,579]" pageId="10" pageNumber="9">FKCP</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B613C81FFA4320DFC5C002819F0DA64" box="[1018,1420,555,579]" collectionCode="RTOC" country="Mexico" pageId="10" pageNumber="9" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<specimenCount id="9D0FFD55FFA4320DFC5C00281866DA64" box="[1018,1050,555,579]" pageId="10" pageNumber="9" type="male">1♂</specimenCount>
<specimenCount id="9D0FFD55FFA4320DFBBC002818F9DA64" box="[1050,1157,555,579]" pageId="10" pageNumber="9" type="female">2♀1juv.♀</specimenCount>
(paratopotypes),
<collectionCode id="ED18AE19FFA4320DFAE5002819F0DA64" box="[1347,1420,555,579]" pageId="10" pageNumber="9">RTOC</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B613C81FFA4320DFA3100281F0FDA85" collectingDate="2022-10-22" collectionCode="R" country="Mexico" elevation="400" latitude="18.494223" location="Dos Amates" longLatPrecision="1" longitude="-95.063225" pageId="10" pageNumber="9" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<location id="8ED66007FFA4320DFA3100281FF8DA44" LSID="urn:lsid:plazi:treatment:03A087CAFFA43216FF5904ED1954DDF8:8ED66007FFA4320DFA3100281FF8DA44" country="Mexico" latitude="18.494223" longLatPrecision="1" longitude="-95.063225" name="Dos Amates" pageId="10" pageNumber="9" stateProvince="Veracruz de Ignacio de la Llave">Dos Amates</location>
,
<geoCoordinate id="EE3D501BFFA4320DFC290048185CDA44" box="[911,1056,587,611]" degrees="18" direction="north" minutes="29" orientation="latitude" pageId="10" pageNumber="9" precision="1" seconds="39.2" value="18.494223">18°29'39.2''N</geoCoordinate>
<geoCoordinate id="EE3D501BFFA4320DFB80004918C0DA45" box="[1062,1212,586,610]" degrees="95" direction="west" minutes="03" orientation="longitude" pageId="10" pageNumber="9" precision="1" seconds="47.6" value="-95.063225">95°03'47.6''W</geoCoordinate>
,
<elevation id="0024D1EFFFA4320DFB610048192DDA44" box="[1223,1361,587,611]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="9" unit="m" value="400.0">
<quantity id="4CF19B39FFA4320DFB6100481977DA45" box="[1223,1291,587,611]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="9" unit="m" value="400.0">400 m</quantity>
a. s. l.
</elevation>
, broadleaf rainforest on volcanic soil, inside rotten stump,
<date id="FFB7101CFFA4320DFAED006819C2DAA4" box="[1355,1470,619,643]" pageId="10" pageNumber="9" value="2022-10-22">
<collectingDate id="EFF3E9F4FFA4320DFAED006819C2DAA4" box="[1355,1470,619,643]" pageId="10" pageNumber="9" value="2022-10-22">22.X.2022</collectingDate>
</date>
, leg.
<collectionCode id="ED18AE19FFA4320DFCF800881F0FDA85" box="[862,883,651,674]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="9">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B613C81FFA4320DFCDB00881966DA84" box="[893,1306,651,675]" collectionCode="FKCP" country="Mexico" location="Teruel" pageId="10" pageNumber="9" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<location id="8ED66007FFA4320DFCDB00881FBDDA84" LSID="urn:lsid:plazi:treatment:03A087CAFFA43216FF5904ED1954DDF8:8ED66007FFA4320DFCDB00881FBDDA84" box="[893,961,651,675]" country="Mexico" name="Teruel" pageId="10" pageNumber="9" stateProvince="Veracruz de Ignacio de la Llave">Teruel</location>
et al.,
<specimenCount id="9D0FFD55FFA4320DFBAF00881853DA84" box="[1033,1071,651,675]" pageId="10" pageNumber="9" type="male">3♂</specimenCount>
<specimenCount id="9D0FFD55FFA4320DFB9600881830DA84" box="[1072,1100,651,675]" pageId="10" pageNumber="9" type="female">1♀</specimenCount>
(
<typeStatus id="54B2887EFFA4320DFBFF008818B9DA84" box="[1113,1221,651,675]" pageId="10" pageNumber="9" type="paratype">paratypes</typeStatus>
),
<collectionCode id="ED18AE19FFA4320DFB7100881966DA84" box="[1239,1306,651,675]" pageId="10" pageNumber="9">FKCP</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3136557FFA4320DFC9600C81F91DB44" pageId="10" pageNumber="9" type="etymology">
<paragraph id="8BB636DCFFA4320DFC9600C81F91DB44" blockId="10.[816,1474,715,867]" pageId="10" pageNumber="9">ETYMOLOGY. The selected epithet is a Latin word that literally means “one who lives isolated, apart from others”, and highlights the singular geographical occurrence of this species in Los Tuxtlas Massif, removed from the main distribution area of the genus.</paragraph>
</subSubSection>
<subSubSection id="C3136557FFA4320DFC9601881F99DE65" pageId="10" pageNumber="9" type="diagnosis">
<paragraph id="8BB636DCFFA4320DFC96018818CADD05" blockId="10.[816,1475,907,1315]" pageId="10" pageNumber="9">
DIAGNOSIS. Adult size small for the genus (
<quantity id="4CF19B39FFA4320DFB4B01881922DB85" box="[1261,1374,907,931]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="10" pageNumber="9" unit="mm" value="21.0" valueMax="22.0" valueMin="20.0">2022 mm</quantity>
in males,
<quantity id="4CF19B39FFA4320DFC9601A81FD9DBE5" box="[816,933,939,963]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="2.8" metricValueMin="2.4" pageId="10" pageNumber="9" unit="mm" value="26.0" valueMax="28.0" valueMin="24.0">2428 mm</quantity>
in females). Coloration with base light orange to yellowish brown, densely variegated with dark to blackish brown, paler and less spotted on chelicerae, pedipalp, legs and pectines. Pedipalp patella with 2223 external trichobothria (7
<emphasis id="B97DEACEFFA4320DFCEB06281F1ADC64" box="[845,870,1067,1091]" italics="true" pageId="10" pageNumber="9">eb</emphasis>
, 2
<emphasis id="B97DEACEFFA4320DFC2106281FD7DC64" box="[903,939,1067,1091]" italics="true" pageId="10" pageNumber="9">esb</emphasis>
, 5
<emphasis id="B97DEACEFFA4320DFC6D06281F96DC65" box="[971,1002,1067,1090]" italics="true" pageId="10" pageNumber="9">em</emphasis>
, 45
<emphasis id="B97DEACEFFA4320DFB830628183FDC65" box="[1061,1091,1067,1090]" italics="true" pageId="10" pageNumber="9">est</emphasis>
, 4
<emphasis id="B97DEACEFFA4320DFBC20628180BDC65" box="[1124,1143,1067,1090]" italics="true" pageId="10" pageNumber="9">et</emphasis>
) and 68 ventral trichobothria. Pedipalp fingers with straight proximal dentate margins, without lobe/notch combination in adults of both sexes. Pectines with shaft clearly divided into areas and lamellae by conspicuous sulci; tooth count
<quantity id="4CF19B39FFA4320DFB2706A918BADCE4" box="[1153,1222,1194,1219]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.524" metricValueMin="1.27" pageId="10" pageNumber="9" unit="in" value="5.5" valueMax="6.0" valueMin="5.0">56 in</quantity>
males,
<quantity id="4CF19B39FFA4320DFABD06A9191CDCE4" box="[1307,1376,1194,1219]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="10" pageNumber="9" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">45 in</quantity>
females. Metasomal segment V length/width ratio 2.262.29 in males, 2.292.32 in females. Telson elongated, narrower than metasomal segment V in both sexes.
</paragraph>
<paragraph id="8BB636DCFFA4320DFC9607481F99DE65" blockId="10.[816,1474,1354,1603]" pageId="10" pageNumber="9">
SEXUAL DIMORPHISM. Female differing from male by:
<emphasis id="B97DEACEFFA4320DFADC074919ECDD45" bold="true" box="[1402,1424,1354,1378]" pageId="10" pageNumber="9">1)</emphasis>
size larger (23.927.8 mm);
<emphasis id="B97DEACEFFA4320DFB9107691831DDA5" bold="true" box="[1079,1101,1386,1410]" pageId="10" pageNumber="9">2)</emphasis>
coloration darker and remarkably more densely infuscate (especially on chelicerae);
<emphasis id="B97DEACEFFA4320DFA0A078919BEDD85" bold="true" box="[1452,1474,1418,1442]" pageId="10" pageNumber="9">3)</emphasis>
pedipalps, legs and metasoma slightly longer and slenderer;
<emphasis id="B97DEACEFFA4320DFC9607C91F3BDDC5" bold="true" box="[816,839,1482,1506]" pageId="10" pageNumber="9">4)</emphasis>
mesosoma much wider and more convex-sided;
<emphasis id="B97DEACEFFA4320DFAE907C91919DDC5" bold="true" box="[1359,1381,1482,1506]" pageId="10" pageNumber="9">5)</emphasis>
sternum larger and parallel-sided;
<emphasis id="B97DEACEFFA4320DFBC207E91806DE25" bold="true" box="[1124,1146,1514,1538]" pageId="10" pageNumber="9">6)</emphasis>
genital papillae absent;
<emphasis id="B97DEACEFFA4320DFA0A07E919BEDE25" bold="true" box="[1452,1474,1514,1538]" pageId="10" pageNumber="9">7)</emphasis>
pectines much smaller, with teeth shorter, narrower and fewer in number (45).
</paragraph>
</subSubSection>
<subSubSection id="C3136557FFA43216FC960468188FD958" lastPageId="17" lastPageNumber="16" pageId="10" pageNumber="9" type="description">
<paragraph id="8BB636DCFFA4320DFC960468180EDEA4" blockId="10.[816,1475,1643,2019]" box="[816,1138,1643,1667]" pageId="10" pageNumber="9">
DESCRIPTION (male
<typeStatus id="54B2887EFFA4320DFBA30468181BDEA4" box="[1029,1127,1643,1667]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BB636DCFFA4320AFC9604891EF3DB04" blockId="10.[816,1475,1643,2019]" lastBlockId="13.[113,772,235,2019]" lastPageId="13" lastPageNumber="12" pageId="10" pageNumber="9">
<emphasis id="B97DEACEFFA4320DFC9604891FD7DE85" bold="true" box="[816,939,1674,1698]" pageId="10" pageNumber="9">Coloration</emphasis>
(
<figureCitation id="13322A59FFA4320DFC670488187ADE84" box="[961,1030,1675,1699]" captionStart="Figures 12" captionStartId="3.[115,192,2017,2039]" captionTargetBox="[113,1474,251,1990]" captionTargetId="figure-31@3.[113,1474,251,1990]" captionTargetPageId="3" captionText="Figures 12. Megacormus seductus sp. n., male holotype (1) and female paratype with a juvenile (2) in vivo habitus." figureDoi="http://doi.org/10.5281/zenodo.8222208" httpUri="https://zenodo.org/record/8222208/files/figure.png" pageId="10" pageNumber="9">Fig. 1</figureCitation>
) Base color light orange to yellowish brown, paler on chelicerae, pedipalp, legs and pectines, pedipalp chelae and telson with a reddish hue. Cheliceral manus finely and irregularly reticulate with dark brown, denser distally; fingers deeply infuscate. Pedipalps densely and irregularly marbled with blackish brown, but sparser distally (much denser on femur, sparser on manus), with all carinae deeply infuscate and the base of most trichobothria much paler (yellowish); fingers blackish, with distal third much paler progressively (becoming yellowish at the tip). Carapace symmetrically and densely marbled with dark to blackish brown, anterior and lateral margins mostly pale; eyes and ocular tubercles black. Tergites symmetrically and densely marbled with dark to blackish brown, with a large orange spot along midline that defines a pale dashed line all along IVI, plus a pair of much smaller and fainter posterolateral pale spots on each side that suggest a dark four-stripped pattern for the tergites. Pectines with shaft heavily and irregularly infuscate, but teeth essentially immaculate. Sternites colored and patterned similarly to tergites, but with a solid, narrow black stripe along midline (below ventral median carinae). Legs irregularly marbled with blackish brown, with pale and dark areas distributed in about the same proportion. Metasoma concolorous throughout (i.e., no segments conspicuously darker), densely and irregularly marbled with dark to blackish brown and with carinae deeply infuscate. Telson vesicle densely and irregularly marbled with dark to blackish brown, with four faint pale stripes corresponding to longitudinal furrows; aculeus orange, with distal third blackish.
</paragraph>
<caption id="DF766654FFA5320CFFD505B21CB3D026" ID-DOI="http://doi.org/10.5281/zenodo.8222214" ID-Zenodo-Dep="8222214" httpUri="https://zenodo.org/record/8222214/files/figure.png" pageId="11" pageNumber="10" startId="11.[115,192,1969,1991]" targetBox="[153,1471,240,1920]" targetPageId="11" targetType="figure">
<paragraph id="8BB636DCFFA5320CFFD505B21CB3D026" blockId="11.[115,1473,1969,2049]" pageId="11" pageNumber="10">
<emphasis id="B97DEACEFFA5320CFFD505B21D7EDFE1" bold="true" box="[115,258,1969,1991]" pageId="11" pageNumber="10">Figures 1726</emphasis>
:
<taxonomicName id="4C094D5FFFA5320CFEA905B11D99DFE1" authority="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský, 2023" authorityName="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2023" box="[271,485,1969,1991]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="seductus" status="sp. nov.">
<emphasis id="B97DEACEFFA5320CFEA905B11D99DFE1" box="[271,485,1969,1991]" italics="true" pageId="11" pageNumber="10">Megacormus seductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57B5FFA5320CFE4D05B11E5CDFE0" box="[491,544,1970,1991]" pageId="11" pageNumber="10" rank="species">
<emphasis id="B97DEACEFFA5320CFE4D05B11E7EDFE0" bold="true" box="[491,514,1970,1991]" pageId="11" pageNumber="10">sp</emphasis>
.
<emphasis id="B97DEACEFFA5320CFDA805B11E67DFE0" bold="true" box="[526,539,1970,1991]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
, female paratype, pedipalp chela dorsal (17), external (18) and ventral (19) views, patella dorsal (20), external (21) and ventral (22) views, femur and trochanter dorsal (23) and ventral (24) views, movable (25) and fixed (26) fingers dentition.
</paragraph>
</caption>
<caption id="DF766654FFA2320BFFD205D21D02D007" ID-DOI="http://doi.org/10.5281/zenodo.8222216" ID-Zenodo-Dep="8222216" httpUri="https://zenodo.org/record/8222216/files/figure.png" pageId="12" pageNumber="11" startId="12.[116,194,2001,2023]" targetBox="[113,1474,217,1991]" targetPageId="12" targetType="figure">
<paragraph id="8BB636DCFFA2320BFFD205D21D02D007" blockId="12.[116,1474,2001,2081]" pageId="12" pageNumber="11">
<emphasis id="B97DEACEFFA2320BFFD205D21D74DFC1" bold="true" box="[116,264,2001,2023]" pageId="12" pageNumber="11">Figures 2734</emphasis>
:
<taxonomicName id="4C094D5FFFA2320BFEBE05D11D88DFC0" authority="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský, 2023" authorityName="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2023" box="[280,500,2002,2023]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="seductus" status="sp. nov.">
<emphasis id="B97DEACEFFA2320BFEBE05D11D88DFC0" box="[280,500,2002,2023]" italics="true" pageId="12" pageNumber="11">Megacormus seductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57B5FFA2320BFE5805D11E4BDFC0" box="[510,567,2002,2023]" pageId="12" pageNumber="11" rank="species">
<emphasis id="B97DEACEFFA2320BFE5805D11E69DFC0" bold="true" box="[510,533,2002,2023]" pageId="12" pageNumber="11">sp</emphasis>
.
<emphasis id="B97DEACEFFA2320BFD8205D11E4DDFC0" bold="true" box="[548,561,2002,2023]" pageId="12" pageNumber="11">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B97DEACEFFA2320BFDE605D21EA8DFC1" bold="true" box="[576,724,2001,2023]" pageId="12" pageNumber="11">Figures 2728</emphasis>
. Male holotype, carapace and tergites IIV (27) and coxosternal area and sternites (28).
<emphasis id="B97DEACEFFA2320BFEA005ED1DE5D024" bold="true" box="[262,409,2030,2052]" pageId="12" pageNumber="11">Figures 2934</emphasis>
. Female paratype, carapace and tergites IIV (29), coxosternal area and sternites (30), and left legs IIV, retrolateral aspect (3134).
</paragraph>
</caption>
<paragraph id="8BB636DCFFA3320AFFD701291CC0DBC4" blockId="13.[113,772,235,2019]" pageId="13" pageNumber="12">
<emphasis id="B97DEACEFFA3320AFFD701291C96DB65" bold="true" box="[113,234,810,834]" pageId="13" pageNumber="12">Chelicerae</emphasis>
(
<figureCitation id="13322A59FFA3320AFF5B01281D2DDB64" box="[253,337,811,835]" captionStart="Figures 2734" captionStartId="12.[116,194,2001,2023]" captionTargetBox="[113,1474,217,1991]" captionTargetId="figure-62@12.[113,1474,217,1994]" captionTargetPageId="12" captionText="Figures 2734: Megacormus seductus sp. n. Figures 2728. Male holotype, carapace and tergites IIV (27) and coxosternal area and sternites (28). Figures 2934. Female paratype, carapace and tergites IIV (29), coxosternal area and sternites (30), and left legs IIV, retrolateral aspect (3134)." figureDoi="http://doi.org/10.5281/zenodo.8222216" httpUri="https://zenodo.org/record/8222216/files/figure.png" pageId="13" pageNumber="12">Fig. 27</figureCitation>
). Dentition typical for the genus, teeth short but sharp. Tegument smooth and glossy. Setation very dense ventrally, but essentially lacking dorsally, i.e., reduced to a few macrosetae distally in manus and also along movable finger. Fingers short, robust, evenly curved and subequal in length.
</paragraph>
<paragraph id="8BB636DCFFA3320AFFD701E91C8CDD84" blockId="13.[113,772,235,2019]" pageId="13" pageNumber="12">
<emphasis id="B97DEACEFFA3320AFFD701E91C9CDC25" bold="true" box="[113,224,1002,1026]" pageId="13" pageNumber="12">Pedipalps</emphasis>
(
<figureCitation id="13322A59FFA3320AFF5501E81D10DC24" box="[243,364,1003,1027]" captionStart="Figures 716" captionStartId="8.[116,193,1960,1982]" captionTargetBox="[132,1474,239,1928]" captionTargetId="figure-72@8.[113,1474,226,1953]" captionTargetPageId="8" captionText="Figures 716: Megacormus seductus sp. n., male holotype, pedipalp chela dorsal (7), external (8) and ventral (9) views, patella dorsal (10), external (11) and ventral (12) views, femur and trochanter dorsal (13) and ventral (14) views, movable (15) and fixed (16) fingers dentition. Movable finger (15) is under UV light, other figures are under white light." figureDoi="http://doi.org/10.5281/zenodo.8222212" httpUri="https://zenodo.org/record/8222212/files/figure.png" pageId="13" pageNumber="12">Figs. 716</figureCitation>
). Slightly shorter and slenderer than standard for the genus (3.19 times longer than carapace). Trichobothrial pattern C, neobothriotaxic majorante: femur with all three trichobothria (
<emphasis id="B97DEACEFFA3320AFE0106481DC8DC44" box="[423,436,1099,1123]" italics="true" pageId="13" pageNumber="12">d</emphasis>
,
<emphasis id="B97DEACEFFA3320AFE6306481DADDC45" box="[453,465,1099,1122]" italics="true" pageId="13" pageNumber="12">e</emphasis>
,
<emphasis id="B97DEACEFFA3320AFE4606481D9BDC45" box="[480,487,1099,1122]" italics="true" pageId="13" pageNumber="12">i</emphasis>
) in positions standard for the genus (at basal-most part of segment); patella with 2 dorsal trichobothria (
<emphasis id="B97DEACEFFA3320AFEFE06881D19DC84" box="[344,357,1163,1187]" italics="true" pageId="13" pageNumber="12">d</emphasis>
), one internal trichobothria (
<emphasis id="B97DEACEFFA3320AFD0306881ED0DC85" box="[677,684,1163,1186]" italics="true" pageId="13" pageNumber="12">i</emphasis>
), 21-22 external trichobothria (7/7
<emphasis id="B97DEACEFFA3320AFE3706A81DD6DCE4" box="[401,426,1195,1219]" italics="true" pageId="13" pageNumber="12">eb</emphasis>
, 2/2
<emphasis id="B97DEACEFFA3320AFE4406A81E7ADCE4" box="[482,518,1195,1219]" italics="true" pageId="13" pageNumber="12">esb</emphasis>
, 4/4
<emphasis id="B97DEACEFFA3320AFD9A06A81E27DCE5" box="[572,603,1195,1218]" italics="true" pageId="13" pageNumber="12">em</emphasis>
, 4/5
<emphasis id="B97DEACEFFA3320AFD3406A81ECCDCE5" box="[658,688,1195,1218]" italics="true" pageId="13" pageNumber="12">est</emphasis>
, 4/4
<emphasis id="B97DEACEFFA3320AFD4106A81E86DCE5" box="[743,762,1195,1218]" italics="true" pageId="13" pageNumber="12">et</emphasis>
) and 7/7 ventral trichobothria (
<emphasis id="B97DEACEFFA3320AFE1C06C81DB6DCC5" box="[442,458,1227,1250]" italics="true" pageId="13" pageNumber="12">V</emphasis>
); chela manus with 2 dorsal trichobothria (
<emphasis id="B97DEACEFFA3320AFEB706E81D4EDD24" box="[273,306,1259,1283]" italics="true" pageId="13" pageNumber="12">Db</emphasis>
,
<emphasis id="B97DEACEFFA3320AFEE006E81D1DDD25" box="[326,353,1259,1282]" italics="true" pageId="13" pageNumber="12">Dt</emphasis>
), 10 external trichobothria (3
<emphasis id="B97DEACEFFA3320AFD6206E81E9EDD24" box="[708,738,1259,1283]" italics="true" pageId="13" pageNumber="12">Eb</emphasis>
, 1
<emphasis id="B97DEACEFFA3320AFFD707081CE5DD04" box="[113,153,1291,1315]" italics="true" pageId="13" pageNumber="12">Esb</emphasis>
, 1
<emphasis id="B97DEACEFFA3320AFF1B07081CA3DD05" box="[189,223,1291,1314]" italics="true" pageId="13" pageNumber="12">Est</emphasis>
, 5
<emphasis id="B97DEACEFFA3320AFEA507081D67DD05" box="[259,283,1291,1314]" italics="true" pageId="13" pageNumber="12">Et</emphasis>
, with
<emphasis id="B97DEACEFFA3320AFEC607081DFCDD07" box="[352,384,1291,1314]" italics="true" pageId="13" pageNumber="12">
Et
<subScript id="178D3499FFA3320AFEDE07111DFCDD07" attach="left" box="[376,384,1298,1312]" fontSize="6" pageId="13" pageNumber="12">1</subScript>
</emphasis>
displaced to ventral surface), and 4 ventral trichobothria (with
<emphasis id="B97DEACEFFA3320AFE3F07281DCDDD67" box="[409,433,1323,1346]" italics="true" pageId="13" pageNumber="12">
V
<subScript id="178D3499FFA3320AFE0F07311DCDDD67" attach="left" box="[425,433,1330,1344]" fontSize="6" pageId="13" pageNumber="12">4</subScript>
</emphasis>
displaced to external surface); fixed finger with 4 dorsal trichobothria (
<emphasis id="B97DEACEFFA3320AFD8307481E3CDD44" box="[549,576,1355,1379]" italics="true" pageId="13" pageNumber="12">db</emphasis>
,
<emphasis id="B97DEACEFFA3320AFDE807481E0FDD44" box="[590,627,1355,1379]" italics="true" pageId="13" pageNumber="12">dsb</emphasis>
,
<emphasis id="B97DEACEFFA3320AFD2707481EDCDD44" box="[641,672,1355,1379]" italics="true" pageId="13" pageNumber="12">dst</emphasis>
,
<emphasis id="B97DEACEFFA3320AFD0907481EB8DD44" box="[687,708,1355,1379]" italics="true" pageId="13" pageNumber="12">dt</emphasis>
), two internal trichobothria (
<emphasis id="B97DEACEFFA3320AFEC307681D06DDA4" box="[357,378,1387,1411]" italics="true" pageId="13" pageNumber="12">ib</emphasis>
,
<emphasis id="B97DEACEFFA3320AFE2E07681DEBDDA5" box="[392,407,1387,1410]" italics="true" pageId="13" pageNumber="12">it</emphasis>
) and 4 external trichobothria (
<emphasis id="B97DEACEFFA3320AFD4507681E80DDA4" box="[739,764,1387,1411]" italics="true" pageId="13" pageNumber="12">eb</emphasis>
,
<emphasis id="B97DEACEFFA3320AFFD707881CE9DD84" box="[113,149,1419,1443]" italics="true" pageId="13" pageNumber="12">esb</emphasis>
,
<emphasis id="B97DEACEFFA3320AFF0407881CBCDD85" box="[162,192,1419,1442]" italics="true" pageId="13" pageNumber="12">est</emphasis>
,
<emphasis id="B97DEACEFFA3320AFF6B07881C9CDD85" box="[205,224,1419,1442]" italics="true" pageId="13" pageNumber="12">et</emphasis>
).
</paragraph>
<paragraph id="8BB636DCFFA3320AFFD707A81DD9DFA4" blockId="13.[113,772,235,2019]" pageId="13" pageNumber="12">
<emphasis id="B97DEACEFFA3320AFFD707A81CC4DDE5" box="[113,184,1451,1474]" italics="true" pageId="13" pageNumber="12">Femur</emphasis>
short and robust (length/width ratio = 2.22), almost bare, with all surfaces essentially flat to shallowly convex. Dorsointernal,dorsoexternal, ventrointernal and ventroexternal carinae complete (moderate, coarsely granulose). Tegument densely, roughly and irregularly granulose on dorsal and internal surfaces, coriaceous on external and ventral surfaces.
<emphasis id="B97DEACEFFA3320AFFD704681CC2DEA4" box="[113,190,1643,1667]" italics="true" pageId="13" pageNumber="12">Patella</emphasis>
short and robust (length/width ratio = 2.07), almost bare, with dorsal and ventral surfaces flat, external and internal surfaces markedly convex. Dorsointernal, dorsoexternal, external median, ventrointernal and ventroexternal carinae complete (moderately strong, finely granulose to crenulate). Tegument coriaceous, with abundant fine to medium-sized, rough granules scattered all over (coarser and denser dorsally). Inner dorsal spur large, sharp and curved forwards; inner ventral spur small and sharp.
</paragraph>
<paragraph id="8BB636DCFFA3320AFFD7058819F4DA24" blockId="13.[113,772,235,2019]" lastBlockId="13.[816,1475,235,2019]" pageId="13" pageNumber="12">
<emphasis id="B97DEACEFFA3320AFFD705881CCDDF84" box="[113,177,1931,1955]" italics="true" pageId="13" pageNumber="12">Chela</emphasis>
short and robust, but slenderer than standard for the genus (length/width ratio = 3.37). Manus subtriangular in dorsal view (length/width ratio = 1.25, distally narrower) and oval in lateral view, sparsely setose and prismatic in cross-section; dorsal internal, dorsal marginal, dorsal secondary, digital, external secondary, ventroexternal and ventrointernal carinae complete (moderately strong, finely granulose to crenulate); tegument coriaceous, with a poorly defined reticulate pattern of small, glossy granules scattered all over. Fingers standard for the genus (movable 1.79 times longer than underhand), shallowly curved and sparsely setose, dentition standard for the genus, basal lobe/notch combination entirely absent.
</paragraph>
<paragraph id="8BB636DCFFA3320AFC9600091FCCDB84" blockId="13.[816,1475,235,2019]" pageId="13" pageNumber="12">
<emphasis id="B97DEACEFFA3320AFC9600091FE2DA05" bold="true" box="[816,926,522,546]" pageId="13" pageNumber="12">Carapace</emphasis>
(
<figureCitation id="13322A59FFA3320AFC1600081878DA04" box="[944,1028,523,547]" captionStart="Figures 2734" captionStartId="12.[116,194,2001,2023]" captionTargetBox="[113,1474,217,1991]" captionTargetId="figure-62@12.[113,1474,217,1994]" captionTargetPageId="12" captionText="Figures 2734: Megacormus seductus sp. n. Figures 2728. Male holotype, carapace and tergites IIV (27) and coxosternal area and sternites (28). Figures 2934. Female paratype, carapace and tergites IIV (29), coxosternal area and sternites (30), and left legs IIV, retrolateral aspect (3134)." figureDoi="http://doi.org/10.5281/zenodo.8222216" httpUri="https://zenodo.org/record/8222216/files/figure.png" pageId="13" pageNumber="12">Fig. 27</figureCitation>
). Slightly wider than long (length/width ratio = 0.98), paraboloid in dorsal view. Anterior margin shallowly bilobed, with 23 pairs of white macrosetae plus several minor setae; median notch very shallowly V-shaped. Tegument coriaceous, with abundant small granules scattered all over.All carinae indistinct or absent (even the superciliaries). Furrows: anterior median, central median, posterior median and posterior marginal narrow and moderately deep, fused altogether; posterior laterals narrow and moderately deep; other furrows either absent or undefined. Median ocular tubercle moderately raised, with eyes large and separated by slightly less than one ocular diameter; two pairs of large lateral eyes.
</paragraph>
<paragraph id="8BB636DCFFA3320AFC9601A919EBDC24" blockId="13.[816,1475,235,2019]" pageId="13" pageNumber="12">
<emphasis id="B97DEACEFFA3320AFC9601A91FEFDBE5" bold="true" box="[816,915,938,962]" pageId="13" pageNumber="12">Sternum</emphasis>
(
<figureCitation id="13322A59FFA3320AFC0601A81F91DBE4" box="[928,1005,939,963]" captionStart="Figures 2734" captionStartId="12.[116,194,2001,2023]" captionTargetBox="[113,1474,217,1991]" captionTargetId="figure-62@12.[113,1474,217,1994]" captionTargetPageId="12" captionText="Figures 2734: Megacormus seductus sp. n. Figures 2728. Male holotype, carapace and tergites IIV (27) and coxosternal area and sternites (28). Figures 2934. Female paratype, carapace and tergites IIV (29), coxosternal area and sternites (30), and left legs IIV, retrolateral aspect (3134)." figureDoi="http://doi.org/10.5281/zenodo.8222216" httpUri="https://zenodo.org/record/8222216/files/figure.png" pageId="13" pageNumber="12">Fig. 28</figureCitation>
). Standard for the genus:
<typeStatus id="54B2887EFFA3320AFB5501A8195DDBE4" box="[1267,1313,939,963]" pageId="13" pageNumber="12">type</typeStatus>
2, large, longer than wide and hexagonal in shape. Tegument coriaceous, with some medium-sized, round granules irregularly scattered.
</paragraph>
<paragraph id="8BB636DCFFA3320AFC960609190EDC84" blockId="13.[816,1475,235,2019]" pageId="13" pageNumber="12">
<emphasis id="B97DEACEFFA3320AFC960609187BDC04" bold="true" box="[816,1031,1034,1059]" pageId="13" pageNumber="12">Genital operculum</emphasis>
(
<figureCitation id="13322A59FFA3320AFBB106081815DC04" box="[1047,1129,1035,1059]" captionStart="Figures 2734" captionStartId="12.[116,194,2001,2023]" captionTargetBox="[113,1474,217,1991]" captionTargetId="figure-62@12.[113,1474,217,1994]" captionTargetPageId="12" captionText="Figures 2734: Megacormus seductus sp. n. Figures 2728. Male holotype, carapace and tergites IIV (27) and coxosternal area and sternites (28). Figures 2934. Female paratype, carapace and tergites IIV (29), coxosternal area and sternites (30), and left legs IIV, retrolateral aspect (3134)." figureDoi="http://doi.org/10.5281/zenodo.8222216" httpUri="https://zenodo.org/record/8222216/files/figure.png" pageId="13" pageNumber="12">Fig. 28</figureCitation>
). Medium-sized and prominent, remarkably rhomboidal in shape. Halves neither separated nor fused and almost triangular in shape; tegument smooth and glossy. Genital papillae medium-sized, well-protruding, with tips blunt, round. Pre-pectinal plate apparently absent.
</paragraph>
<paragraph id="8BB636DCFFA3320AFC9606A918C3DD84" blockId="13.[816,1475,235,2019]" pageId="13" pageNumber="12">
<emphasis id="B97DEACEFFA3320AFC9606A91FF1DCE5" bold="true" box="[816,909,1194,1218]" pageId="13" pageNumber="12">Pectines</emphasis>
(
<figureCitation id="13322A59FFA3320AFC3A06A81F90DCE4" box="[924,1004,1195,1219]" captionStart="Figures 2734" captionStartId="12.[116,194,2001,2023]" captionTargetBox="[113,1474,217,1991]" captionTargetId="figure-62@12.[113,1474,217,1994]" captionTargetPageId="12" captionText="Figures 2734: Megacormus seductus sp. n. Figures 2728. Male holotype, carapace and tergites IIV (27) and coxosternal area and sternites (28). Figures 2934. Female paratype, carapace and tergites IIV (29), coxosternal area and sternites (30), and left legs IIV, retrolateral aspect (3134)." figureDoi="http://doi.org/10.5281/zenodo.8222216" httpUri="https://zenodo.org/record/8222216/files/figure.png" pageId="13" pageNumber="12">Fig. 28</figureCitation>
). Size and shape standard for the genus: not reaching coxa-trochanter joint of leg IV, subrectangular and moderately setose, anterior area with 4/4 lamellae, median area with 5/6. Tooth count 6/6, teeth swollen, straight and basally not separated; fulcra entirely absent. Basal plate wider than long; anterior margin widely and deeply V-notched medially, posterior margin essentially straight; tegument smooth, glossy to coriaceous and sparsely granulose.
</paragraph>
<paragraph id="8BB636DCFFA3320AFC9607A81932DF64" blockId="13.[816,1475,235,2019]" pageId="13" pageNumber="12">
<emphasis id="B97DEACEFFA3320AFC9607A81FD4DDE4" bold="true" box="[816,936,1451,1475]" pageId="13" pageNumber="12">Mesosoma</emphasis>
(
<figureCitation id="13322A59FFA3320AFC1207A81849DDE4" box="[948,1077,1451,1475]" captionStart="Figures 2734" captionStartId="12.[116,194,2001,2023]" captionTargetBox="[113,1474,217,1991]" captionTargetId="figure-62@12.[113,1474,217,1994]" captionTargetPageId="12" captionText="Figures 2734: Megacormus seductus sp. n. Figures 2728. Male holotype, carapace and tergites IIV (27) and coxosternal area and sternites (28). Figures 2934. Female paratype, carapace and tergites IIV (29), coxosternal area and sternites (30), and left legs IIV, retrolateral aspect (3134)." figureDoi="http://doi.org/10.5281/zenodo.8222216" httpUri="https://zenodo.org/record/8222216/files/figure.png" pageId="13" pageNumber="12">Figs. 2728</figureCitation>
). Tergites almost bare, anterior margin shallowly convex, posterior margin shallowly produced in the median part; median carina very weak and finely subcrenulate on IVI, submedian and lateral carinae strong and finely serratocrenulate on VII; tegument coriaceous, with abundant small granules scattered all over. Sternites sparsely setose (largely along posterior and lateral margins), posterior margin shallowly concave to widely bilobed; IIIVI acarinate, VII with only ventral median carinae perfectly defined, finely granulose to subcrenulate; tegument coriaceous, with abundant coarse and glossy granules; spiracles small, elongate-oval to short slit-like; V with smooth patch large, paraboloid in shape, translucent to yellowish, glossy and slightly bulky.
</paragraph>
<paragraph id="8BB636DCFFA3320AFC96054819BFDFC5" blockId="13.[816,1475,235,2019]" pageId="13" pageNumber="12">
<emphasis id="B97DEACEFFA3320AFC9605481FD9DF44" bold="true" box="[816,933,1867,1891]" pageId="13" pageNumber="12">Metasoma</emphasis>
(
<figureCitation id="13322A59FFA3320AFC1C0548183DDF44" box="[954,1089,1867,1891]" captionStart="Figures 3542" captionStartId="14.[113,190,1957,1979]" captionTargetBox="[125,1450,263,1942]" captionTargetId="figure-78@14.[113,1474,230,1948]" captionTargetPageId="14" captionText="Figures 3542. Megacormus seductus sp. n. Figures 35, 3739. Male holotype, telson in lateral (35), metasoma and telson in lateral (37), dorsal (38) and ventral (39) views. Figures 36, 4042. Female paratype, telson in lateral (36), metasoma and telson in lateral (40), dorsal (41) and ventral (42) views. Scale bars: 10 mm (3739, 4042)." figureDoi="http://doi.org/10.5281/zenodo.8222218" httpUri="https://zenodo.org/record/8222218/files/figure.png" pageId="13" pageNumber="12">Figs. 3739</figureCitation>
). Size and shape standard for the genus (2.77 times longer than carapace, length/width ratio of segments IV = 0.64, 0.83, 0.89, 1.23 and 2.27), moderately narrower distally and moderately setose; segments IIII wider than long, IVV longer than wide; IIII wider than deep, IVV
</paragraph>
<caption id="DF766654FFA03209FFD705A61ED4DFD3" ID-DOI="http://doi.org/10.5281/zenodo.8222218" ID-Zenodo-Dep="8222218" httpUri="https://zenodo.org/record/8222218/files/figure.png" pageId="14" pageNumber="13" startId="14.[113,190,1957,1979]" targetBox="[125,1450,263,1942]" targetPageId="14" targetType="figure">
<paragraph id="8BB636DCFFA03209FFD705A61ED4DFD3" blockId="14.[113,1471,1957,2037]" pageId="14" pageNumber="13">
<emphasis id="B97DEACEFFA03209FFD705A61D7EDF9D" bold="true" box="[113,258,1957,1979]" pageId="14" pageNumber="13">Figures 3542</emphasis>
.
<taxonomicName id="4C094D5FFFA03209FEA905A51D9BDF9D" authority="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský, 2023" authorityName="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2023" box="[271,487,1957,1979]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="seductus" status="sp. nov.">
<emphasis id="B97DEACEFFA03209FEA905A51D9BDF9D" box="[271,487,1957,1979]" italics="true" pageId="14" pageNumber="13">Megacormus seductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57B5FFA03209FE4805A61E5ADF9C" box="[494,550,1957,1979]" pageId="14" pageNumber="13" rank="species">
<emphasis id="B97DEACEFFA03209FE4805A61E79DF9D" bold="true" box="[494,517,1957,1978]" pageId="14" pageNumber="13">sp</emphasis>
.
<emphasis id="B97DEACEFFA03209FDB505A61E5CDF9D" bold="true" box="[531,544,1957,1978]" pageId="14" pageNumber="13">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B97DEACEFFA03209FD8805A61EE6DF9D" bold="true" box="[558,666,1957,1979]" pageId="14" pageNumber="13">Figures 35</emphasis>
,
<emphasis id="B97DEACEFFA03209FD0E05A61E98DF9D" bold="true" box="[680,740,1957,1979]" pageId="14" pageNumber="13">3739</emphasis>
. Male holotype, telson in lateral (35), metasoma and telson in lateral (37), dorsal (38) and ventral (39) views.
<emphasis id="B97DEACEFFA03209FE6505C11E51DFF0" bold="true" box="[451,557,1986,2008]" pageId="14" pageNumber="13">Figures 36</emphasis>
,
<emphasis id="B97DEACEFFA03209FD9F05C11E09DFF0" bold="true" box="[569,629,1986,2008]" pageId="14" pageNumber="13">4042</emphasis>
. Female paratype, telson in lateral (36), metasoma and telson in lateral (40), dorsal (41) and ventral (42) views. Scale bars: 10 mm (3739, 4042).
</paragraph>
</caption>
<caption id="DF766654FFA13208FFD505E91EA6D027" ID-DOI="http://doi.org/10.5281/zenodo.8222222" ID-Zenodo-Dep="8222222" box="[115,730,2026,2048]" httpUri="https://zenodo.org/record/8222222/files/figure.png" pageId="15" pageNumber="14" startId="15.[115,183,2026,2048]" targetBox="[113,1474,234,2015]" targetPageId="15" targetType="figure">
<paragraph id="8BB636DCFFA13208FFD505E91EA6D027" blockId="15.[115,730,2026,2048]" box="[115,730,2026,2048]" pageId="15" pageNumber="14">
<emphasis id="B97DEACEFFA13208FFD505E91CA8DFD8" bold="true" box="[115,212,2026,2048]" pageId="15" pageNumber="14">Figure 43</emphasis>
.
<taxonomicName id="4C094D5FFFA13208FF4605E81DCBDFD8" authority="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský, 2023" authorityName="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2023" box="[224,439,2026,2048]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="seductus" status="sp. nov.">
<emphasis id="B97DEACEFFA13208FF4605E81DCBDFD8" box="[224,439,2026,2048]" italics="true" pageId="15" pageNumber="14">Megacormus seductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57B5FFA13208FE1A05E81D8DD027" box="[444,497,2027,2048]" pageId="15" pageNumber="14" rank="species">
<emphasis id="B97DEACEFFA13208FE1A05E81DAFD027" bold="true" box="[444,467,2027,2048]" pageId="15" pageNumber="14">sp</emphasis>
.
<emphasis id="B97DEACEFFA13208FE7905E81D90D027" bold="true" box="[479,492,2027,2048]" pageId="15" pageNumber="14">n</emphasis>
.
</taxonomicNameLabel>
, habitat at type locality.
</paragraph>
</caption>
<paragraph id="8BB636DCFFBE3217FFD305F619BED063" blockId="16.[117,1475,2037,2117]" pageId="16" pageNumber="15">
<emphasis id="B97DEACEFFBE3217FFD305F61C83D02D" bold="true" box="[117,255,2037,2059]" pageId="16" pageNumber="15">Figures 4461</emphasis>
.
<taxonomicName id="4C094D5FFFBE3217FEAB05F51DFAD02C" authorityName="Karsch" authorityYear="1881" box="[269,390,2038,2059]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEACEFFBE3217FEAB05F51DFAD02C" box="[269,390,2038,2059]" italics="true" pageId="16" pageNumber="15">Megacormus</emphasis>
</taxonomicName>
, external pedipalp chela of males (44, 46, 48, 50, 52, 54, 56, 58) and females (45, 47, 49, 51, 53, 55, 57, 59), and external patella of male (60) and female (61).
<emphasis id="B97DEACEFFBE3217FDBE0A111EDDD000" bold="true" box="[536,673,2066,2088]" pageId="16" pageNumber="15">Figures 4445</emphasis>
.
<taxonomicName id="4C094D5FFFBE3217FD0A0A101F6FD000" box="[684,787,2066,2088]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="gertschi">
<emphasis id="B97DEACEFFBE3217FD0A0A101F6FD000" box="[684,787,2066,2088]" italics="true" pageId="16" pageNumber="15">M. gertschi</emphasis>
</taxonomicName>
.
<emphasis id="B97DEACEFFBE3217FCB90A111FD4D000" bold="true" box="[799,936,2066,2088]" pageId="16" pageNumber="15">Figures 4647</emphasis>
.
<taxonomicName id="4C094D5FFFBE3217FC150A101859D00F" authorityName=": KovarIk" authorityYear="2019" box="[947,1061,2067,2088]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="granosus">
<emphasis id="B97DEACEFFBE3217FC150A101859D00F" box="[947,1061,2067,2088]" italics="true" pageId="16" pageNumber="15">M. granosus</emphasis>
</taxonomicName>
.
<emphasis id="B97DEACEFFBE3217FB970A1118C5D000" bold="true" box="[1073,1209,2066,2088]" pageId="16" pageNumber="15">Figures 4849</emphasis>
.
<taxonomicName id="4C094D5FFFBE3217FB620A101955D000" authorityName="Sissom" authorityYear="1994" box="[1220,1321,2066,2088]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="grubbsi">
<emphasis id="B97DEACEFFBE3217FB620A101955D000" box="[1220,1321,2066,2088]" italics="true" pageId="16" pageNumber="15">M. grubbsi</emphasis>
</taxonomicName>
.
<emphasis id="B97DEACEFFBE3217FA920A1119C1D000" bold="true" box="[1332,1469,2066,2088]" pageId="16" pageNumber="15">Figures 5051</emphasis>
.
<taxonomicName id="4C094D5FFFBE3217FFD30A331D79D062" authorityName=": Sissom" authorityYear="1994" box="[117,261,2096,2117]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="segmentatus">
<emphasis id="B97DEACEFFBE3217FFD30A331D79D062" box="[117,261,2096,2117]" italics="true" pageId="16" pageNumber="15">M. segmentatus</emphasis>
</taxonomicName>
.
<emphasis id="B97DEACEFFBE3217FEB70A2C1DE6D063" bold="true" box="[273,410,2095,2117]" pageId="16" pageNumber="15">Figures 5253</emphasis>
.
<taxonomicName id="4C094D5FFFBE3217FE030A331D8BD063" box="[421,503,2095,2117]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="xichu">
<emphasis id="B97DEACEFFBE3217FE030A331D8BD063" box="[421,503,2095,2117]" italics="true" pageId="16" pageNumber="15">M. xichu</emphasis>
</taxonomicName>
.
<emphasis id="B97DEACEFFBE3217FDA40A2C1EF7D063" bold="true" box="[514,651,2095,2117]" pageId="16" pageNumber="15">Figures 5455</emphasis>
.
<taxonomicName id="4C094D5FFFBE3217FD310A331F7DD063" authorityName="KovarIk" authorityYear="2019" box="[663,769,2095,2117]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="franckei">
<emphasis id="B97DEACEFFBE3217FD310A331F7DD063" box="[663,769,2095,2117]" italics="true" pageId="16" pageNumber="15">M. franckei</emphasis>
</taxonomicName>
.
<emphasis id="B97DEACEFFBE3217FCAA0A2C1FE9D063" bold="true" box="[780,917,2095,2117]" pageId="16" pageNumber="15">Figures 5657</emphasis>
,
<emphasis id="B97DEACEFFBE3217FC070A2C1FC4D063" bold="true" box="[929,952,2095,2116]" pageId="16" pageNumber="15">60</emphasis>
.
<taxonomicName id="4C094D5FFFBE3217FC620A33184ED063" authority="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský, 2023" authorityName="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2023" box="[964,1074,2095,2117]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="seductus" status="sp. nov.">
<emphasis id="B97DEACEFFBE3217FC620A33184ED063" box="[964,1074,2095,2117]" italics="true" pageId="16" pageNumber="15">M. seductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57B5FFBE3217FB9E0A2C1810D062" box="[1080,1132,2095,2117]" pageId="16" pageNumber="15" rank="species">
<emphasis id="B97DEACEFFBE3217FB9E0A2C1832D063" bold="true" box="[1080,1102,2095,2116]" pageId="16" pageNumber="15">sp</emphasis>
.
<emphasis id="B97DEACEFFBE3217FBFF0A2C181AD063" bold="true" box="[1113,1126,2095,2116]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B97DEACEFFBE3217FBD70A2C1886D063" bold="true" box="[1137,1274,2095,2117]" pageId="16" pageNumber="15">Figures 5859</emphasis>
,
<emphasis id="B97DEACEFFBE3217FAA00A2C1961D063" bold="true" box="[1286,1309,2095,2116]" pageId="16" pageNumber="15">61</emphasis>
.
<taxonomicName id="4C094D5FFFBE3217FA8F0A3319F2D063" authority="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský, 2023" authorityName="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2023" box="[1321,1422,2095,2117]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="orizaba" status="sp. nov.">
<emphasis id="B97DEACEFFBE3217FA8F0A3319F2D063" box="[1321,1422,2095,2117]" italics="true" pageId="16" pageNumber="15">M. orizaba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57B5FFBE3217FA320A2C19BED063" box="[1428,1474,2095,2117]" pageId="16" pageNumber="15" rank="species">
<emphasis id="B97DEACEFFBE3217FA320A2C19D6D063" bold="true" box="[1428,1450,2095,2116]" pageId="16" pageNumber="15">sp</emphasis>
.
<emphasis id="B97DEACEFFBE3217FA130A2C19BED063" bold="true" box="[1461,1474,2095,2116]" pageId="16" pageNumber="15">n</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB636DCFFBF3216FFD702E41EBADA38" blockId="17.[113,772,231,1439]" pageId="17" pageNumber="16">deeper than wide. Segments IIV with seven complete to almost complete carinae, V with five: all weak, irregularly and finely granulose to crenulate, with distalmost denticle conspicuously enlarged; lateral inframedians entirely absent or undefined from surrounding granulation on all segments; ventral submedians entirely absent; ventral median carina perfectly defined all along IV; anal arc weakly granulose; laterodistal lobes of V obsolete, blunt-triangular. Intercarinal tegument coriaceous, with abundant minute and fine granules scattered all over. Dorsal furrow complete, moderately narrow and shallow.</paragraph>
<paragraph id="8BB636DCFFBF3216FFD700241EF1DAD8" blockId="17.[113,772,231,1439]" pageId="17" pageNumber="16">
<emphasis id="B97DEACEFFBF3216FFD700241CC6DA18" bold="true" box="[113,186,551,575]" pageId="17" pageNumber="16">Telson</emphasis>
(
<figureCitation id="13322A59FFBF3216FF6E00241D64DA18" box="[200,280,551,575]" captionStart="Figures 3542" captionStartId="14.[113,190,1957,1979]" captionTargetBox="[125,1450,263,1942]" captionTargetId="figure-78@14.[113,1474,230,1948]" captionTargetPageId="14" captionText="Figures 3542. Megacormus seductus sp. n. Figures 35, 3739. Male holotype, telson in lateral (35), metasoma and telson in lateral (37), dorsal (38) and ventral (39) views. Figures 36, 4042. Female paratype, telson in lateral (36), metasoma and telson in lateral (40), dorsal (41) and ventral (42) views. Scale bars: 10 mm (3739, 4042)." figureDoi="http://doi.org/10.5281/zenodo.8222218" httpUri="https://zenodo.org/record/8222218/files/figure.png" pageId="17" pageNumber="16">Fig. 35</figureCitation>
). Sparsely setose, oval (2.71 times longer than wide, 1.21 times wider than deep); tegument coriaceous, with abundant moderately coarse but weak granules scattered all over and four very shallow longitudinal furrows; distal end of vesicle poorly defined by an obsolete annular ring and vestigial laterodistal shoulders. Aculeus moderately long, sharp and shallowly curved, with tegument glossy.
</paragraph>
<paragraph id="8BB636DCFFBF3216FFD701041DE7DB98" blockId="17.[113,772,231,1439]" pageId="17" pageNumber="16">
<emphasis id="B97DEACEFFBF3216FFD701041CDADB38" bold="true" box="[113,166,775,799]" pageId="17" pageNumber="16">Legs</emphasis>
. Size and slenderness standard for the genus, with all carinae weakly granulose to subcrenulate; tegument smooth coriaceous, with dorsal surface of femur sparsely granulose. Prolateral pedal spurs standard-sized, thick but sharp. Claws standard-sized (at least one-third the length of its respective telotarsus) and well-curved.
</paragraph>
<paragraph id="8BB636DCFFBF3216FFD701C41CB3DDB8" blockId="17.[113,772,231,1439]" pageId="17" pageNumber="16">
<emphasis id="B97DEACEFFBF3216FFD701C41D20DBF8" bold="true" box="[113,348,967,991]" pageId="17" pageNumber="16">Hemispermatophore</emphasis>
. (
<figureCitation id="13322A59FFBF3216FEDF01C41E7FDBF8" box="[377,515,967,991]" captionStart="Figures 6265" captionStartId="18.[109,186,1132,1154]" captionTargetBox="[138,1454,242,1112]" captionTargetId="figure-463@18.[112,1473,221,1121]" captionTargetPageId="18" captionText="Figures 6265. Hemispermatophores of Megacormus spp. Figure 62. M. orizaba sp. n. 19MD (holotype, No. 1621). Figure 63. M. granosus (No. 2462). Figures 6465. M. seductus sp. n. 22MA (holotype and paratopotype, Nos. 2460, 2461).Convex views of left hemispermatophores (62, 63) and right hemispermatophores shown in mirror image (64, 65). Scale bars: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8222224" httpUri="https://zenodo.org/record/8222224/files/figure.png" pageId="17" pageNumber="16">Figs. 6465</figureCitation>
,
<figureCitation id="13322A59FFBF3216FDB301C41E20DBF8" box="[533,604,967,991]" captionStart="Figures 6677" captionStartId="19.[113,187,1941,1963]" captionTargetBox="[118,1474,212,1929]" captionTargetId="figure-160@19.[113,1474,210,1931]" captionTargetPageId="19" captionText="Figures 6677. Hemispermatophore capsules of Megacormus. 66. M. orizaba sp. n. (holotype, No.1621). 6768. M. gertschi (Nos.1619, 1620).69. M. franckei (paratype, No. 1618).7071. M. xichu (Nos.1617, 1626).7273. M. seductus sp. n. (holotype and paratopotype,Nos.24602461).74. M. segmentatus (No.1622).75. M. granosus (No.2462).76. M. grubbsi (No.1623).77. M. grubbsi (No.2463).Convex views ofleftcapsules (66, 6871, 7375 and 77), and right capsules shown in mirror image (67, 72 and 76).Abbreviations: dpd, distal dentate process; dpp, proximal dentate process; cls, crown-like structure; trd, distal transverse ridge; trp, proximal transverse ridge; tmsd, terminal membrane of the sperm duct. Scale bars: 400 μm." figureDoi="http://doi.org/10.5281/zenodo.8222226" httpUri="https://zenodo.org/record/8222226/files/figure.png" pageId="17" pageNumber="16">7273</figureCitation>
). Lamelliform. Overall proportions of distal lamina, capsule, truncal flexure, trunk and pedicel standard for the genus. Distal lamina broad with angulate anterior margin, tapering in distal half; excluding crest, lamina width at basal constriction greater than half maximum lamina width. Capsule with moderately broad proximal transverse ridge; proximal dentate process expanded, either subrectangular with 79 blunt denticles (No. 2460; both left and right capsules), or lanceolate with 710 sharp triangular denticles (No. 2461; both left and right capsules). Distal dentate process broad, laminate, strongly serrated with 1314 (No. 2460) or 18 (No. 2461) triangular denticles. Basal carina with crown-like structure bearing 46 tines. Terminal membrane of sperm duct with numerous fine spicules.
</paragraph>
<paragraph id="8BB636DCFFBF3216FFD707C41DE1DE78" blockId="17.[113,772,1479,2079]" pageId="17" pageNumber="16">
AFFINITIES. The described features distinguish
<taxonomicName id="4C094D5FFFBF3216FDDE07CB1CB1DDD8" authority="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský, 2023" authorityName="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2023" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="seductus" status="sp. nov.">
<emphasis id="B97DEACEFFBF3216FDDE07CB1CB1DDD8" italics="true" pageId="17" pageNumber="16">Megacormus seductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57B5FFBF3216FF7507E41D73DDD9" box="[211,271,1511,1535]" pageId="17" pageNumber="16" rank="species">
<emphasis id="B97DEACEFFBF3216FF7507E41C90DDD8" bold="true" box="[211,236,1511,1535]" pageId="17" pageNumber="16">sp</emphasis>
.
<emphasis id="B97DEACEFFBF3216FF5F07E41D74DDD8" bold="true" box="[249,264,1511,1535]" pageId="17" pageNumber="16">n</emphasis>
.
</taxonomicNameLabel>
from all other species of the genus (see the key below). The validity of
<taxonomicName id="4C094D5FFFBF3216FECC040B1D94DE38" authority="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský, 2023" authorityName="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2023" box="[362,488,1543,1567]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="seductus" status="sp. nov.">
<emphasis id="B97DEACEFFBF3216FECC040B1D94DE38" box="[362,488,1543,1567]" italics="true" pageId="17" pageNumber="16">M. seductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57B5FFBF3216FE4804041E55DE39" box="[494,553,1543,1567]" pageId="17" pageNumber="16" rank="species">
<emphasis id="B97DEACEFFBF3216FE4804041E7BDE38" bold="true" box="[494,519,1543,1567]" pageId="17" pageNumber="16">sp</emphasis>
.
<emphasis id="B97DEACEFFBF3216FDB504041E5EDE38" bold="true" box="[531,546,1543,1567]" pageId="17" pageNumber="16">n</emphasis>
.
</taxonomicNameLabel>
is confirmed also by DNA analysis implemented by Charles University in
<collectingRegion id="49CDF83EFFBF3216FD1F04241F7FDE18" box="[697,771,1575,1599]" country="Czech Republic" name="Praha" pageId="17" pageNumber="16">Prague</collectingRegion>
(manuscript in preparation).
</paragraph>
<paragraph id="8BB636DCFFBF3216FF070464188FD958" blockId="17.[113,772,1479,2079]" lastBlockId="17.[816,1474,231,383]" pageId="17" pageNumber="16">
By the combination of very small adult size and absolute lack of pedipalp lobe/notch combination in adult males, this species resembles only
<taxonomicName id="4C094D5FFFBF3216FECB04AB1D8FDE98" authorityName=": KovarIk" authorityYear="2019" box="[365,499,1704,1727]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="granosus">
<emphasis id="B97DEACEFFBF3216FECB04AB1D8FDE98" box="[365,499,1704,1727]" italics="true" pageId="17" pageNumber="16">M. granosus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C094D5FFFBF3216FD8104AB1EB3DE98" authorityName=": Sissom" authorityYear="1994" box="[551,719,1704,1727]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="segmentatus">
<emphasis id="B97DEACEFFBF3216FD8104AB1EB3DE98" box="[551,719,1704,1727]" italics="true" pageId="17" pageNumber="16">M. segmentatus</emphasis>
</taxonomicName>
, and in fact, it was formerly misidentified as the latter taxon by
<bibRefCitation id="EF984B2DFFBF3216FFD704E41D73DED8" author="SISSOM" box="[113,271,1767,1791]" pageId="17" pageNumber="16" pagination="265 - 272" refId="ref14012" refString="SISSOM, W. D. 1994. Systematic studies on the genus Megacormus (Scorpiones, Chactidae, Megacorminae), with descriptions of a new species from Oaxaca, Mexico and of the male of Megacormus segmentatus Pocock. Insecta Mundi, 8: 265 - 272." type="journal article" year="1994">Sissom (1994)</bibRefCitation>
. However,
<taxonomicName id="4C094D5FFFBF3216FE2304EB1E78DED8" authority="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský, 2023" authorityName="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2023" box="[389,516,1767,1791]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="seductus" status="sp. nov.">
<emphasis id="B97DEACEFFBF3216FE2304EB1E78DED8" box="[389,516,1767,1791]" italics="true" pageId="17" pageNumber="16">M. seductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57B5FFBF3216FDAF04E41E39DED9" box="[521,581,1767,1791]" pageId="17" pageNumber="16" rank="species">
<emphasis id="B97DEACEFFBF3216FDAF04E41E5EDED8" bold="true" box="[521,546,1767,1791]" pageId="17" pageNumber="16">sp</emphasis>
.
<emphasis id="B97DEACEFFBF3216FD8904E41E42DED8" bold="true" box="[559,574,1767,1791]" pageId="17" pageNumber="16">n</emphasis>
.
</taxonomicNameLabel>
occurs in isolated localities (
<figureCitation id="13322A59FFBF3216FF4605041D4CDF38" box="[224,304,1799,1823]" captionStart="Figure 82" captionStartId="22.[113,181,1223,1245]" captionTargetBox="[112,1473,247,1206]" captionTargetId="figure-319@22.[112,1473,247,1206]" captionTargetPageId="22" captionText="Figure 82. Distribution of Megacormus specimens examined for this review (see Comparative material examined in Methods, Material &amp; Abbreviations section)." figureDoi="http://doi.org/10.5281/zenodo.8222230" httpUri="https://zenodo.org/record/8222230/files/figure.png" pageId="17" pageNumber="16">Fig. 82</figureCitation>
). The most morphologically similar species can be regarded as
<taxonomicName id="4C094D5FFFBF3216FE9C052B1DC2DF18" authorityName=": KovarIk" authorityYear="2019" box="[314,446,1832,1855]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="granosus">
<emphasis id="B97DEACEFFBF3216FE9C052B1DC2DF18" box="[314,446,1832,1855]" italics="true" pageId="17" pageNumber="16">M. granosus</emphasis>
</taxonomicName>
, which differs in the following morphological characters that are standard in the species-level taxonomy of the genus:
<emphasis id="B97DEACEFFBF3216FED105641DF8DF58" bold="true" box="[375,388,1895,1919]" pageId="17" pageNumber="16">1</emphasis>
. Size smaller (
<quantity id="4CF19B39FFBF3216FD8005641EE6DF59" box="[550,666,1895,1919]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="17" pageNumber="16" unit="mm" value="16.0" valueMax="18.0" valueMin="14.0">1418 mm</quantity>
in males,
<quantity id="4CF19B39FFBF3216FFD705841C9EDFB9" box="[113,226,1927,1951]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="17" pageNumber="16" unit="mm" value="20.5" valueMax="23.0" valueMin="18.0">1823 mm</quantity>
in females).
<emphasis id="B97DEACEFFBF3216FECC05841D0BDFB8" bold="true" box="[362,375,1927,1951]" pageId="17" pageNumber="16">2</emphasis>
. Entire body and appendages densely covered by very short, rigid, white microsetae in addition to ordinary setae.
<emphasis id="B97DEACEFFBF3216FEB105C41D58DFF8" bold="true" box="[279,292,1991,2015]" pageId="17" pageNumber="16">3</emphasis>
. Pectines with shaft not clearly divided into anterior and median areas and lamellae, all fused altogether into a single triangular sclerite (at most with faint incomplete vestiges of sulci).
<emphasis id="B97DEACEFFBF3216FC5D02E41874D8D8" bold="true" box="[1019,1032,231,255]" pageId="17" pageNumber="16">4</emphasis>
. Pectinal tooth count lower (males 34, females 14).
<emphasis id="B97DEACEFFBF3216FC6103041FA8D938" bold="true" box="[967,980,263,287]" pageId="17" pageNumber="16">5</emphasis>
. Entire body and appendages with intercarinal granulation noticeably finer and more even.
<emphasis id="B97DEACEFFBF3216FA850324194CD918" bold="true" box="[1315,1328,295,319]" pageId="17" pageNumber="16">6</emphasis>
. Adults with mesosoma, metasoma and telson remarkably wider and heavier (more conspicuously so in males).
</paragraph>
</subSubSection>
<subSubSection id="C3136557FFBF3216FC9603A41954DDF8" pageId="17" pageNumber="16" type="distribution">
<paragraph id="8BB636DCFFBF3216FC9603A419FCDAB8" blockId="17.[816,1474,423,671]" pageId="17" pageNumber="16">
DISTRIBUTION. (
<figureCitation id="13322A59FFBF3216FC7C03A41856D998" box="[986,1066,423,447]" captionStart="Figure 82" captionStartId="22.[113,181,1223,1245]" captionTargetBox="[112,1473,247,1206]" captionTargetId="figure-319@22.[112,1473,247,1206]" captionTargetPageId="22" captionText="Figure 82. Distribution of Megacormus specimens examined for this review (see Comparative material examined in Methods, Material &amp; Abbreviations section)." figureDoi="http://doi.org/10.5281/zenodo.8222230" httpUri="https://zenodo.org/record/8222230/files/figure.png" pageId="17" pageNumber="16">Fig. 82</figureCitation>
).
<taxonomicName id="4C094D5FFFBF3216FB9B03AB18C1D998" authority="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský, 2023" authorityName="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2023" box="[1085,1213,423,447]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="seductus" status="sp. nov.">
<emphasis id="B97DEACEFFBF3216FB9B03AB18C1D998" box="[1085,1213,423,447]" italics="true" pageId="17" pageNumber="16">M. seductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57B5FFBF3216FB6203A4197DD999" box="[1220,1281,423,447]" pageId="17" pageNumber="16" rank="species">
<emphasis id="B97DEACEFFBF3216FB6203A418A1D998" bold="true" box="[1220,1245,423,447]" pageId="17" pageNumber="16">sp</emphasis>
.
<emphasis id="B97DEACEFFBF3216FB4D03A41886D998" bold="true" box="[1259,1274,423,447]" pageId="17" pageNumber="16">n</emphasis>
.
</taxonomicNameLabel>
was found in two localities about
<quantity id="4CF19B39FFBF3216FC7903C4185AD9F8" box="[991,1062,455,479]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="16" unit="km" value="10.0">10 km</quantity>
apart from each other, but it may be widespread across Los Tuxtlas area, at least in the relict forest patches. The primary forest has been largely degraded or eliminated all over the area, but several such relicts still persist in the places with steepest slopes and/or privately controlled lands. Los Tuxtlas represents an isolated branch of the Trans-Mexican Neovolcanic Belt, actually its easternmost tip.
</paragraph>
<paragraph id="8BB636DCFFBF3216FC9600C4192FDC98" blockId="17.[816,1475,711,1503]" pageId="17" pageNumber="16">
ECOLOGICAL NOTES. The
<typeStatus id="54B2887EFFBF3216FBE400C41806DAF8" box="[1090,1146,711,735]" pageId="17" pageNumber="16">types</typeStatus>
were all collected inside very old, moist, rotten stumps, in very humid places inside low to mid altitude broadleaf tropical rainforest (
<figureCitation id="13322A59FFBF3216FA840104190ADB38" box="[1314,1398,775,799]" captionStart="Figure 43" captionStartId="15.[115,183,2026,2048]" captionTargetBox="[113,1474,234,2015]" captionTargetId="figure-22@15.[113,1474,231,2017]" captionTargetPageId="15" captionText="Figure 43. Megacormus seductus sp. n., habitat at type locality." figureDoi="http://doi.org/10.5281/zenodo.8222222" httpUri="https://zenodo.org/record/8222222/files/figure.png" pageId="17" pageNumber="16">Fig. 43</figureCitation>
). Each inhabited stump housed a group of 35 individuals of
<taxonomicName id="4C094D5FFFBF3216FA03012B1FF0DB78" authority="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský, 2023" authorityName="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2023" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="seductus" status="sp. nov.">
<emphasis id="B97DEACEFFBF3216FA03012B1FF0DB78" italics="true" pageId="17" pageNumber="16">M. seductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57B5FFBF3216FC3301441FAFDB79" box="[917,979,839,863]" pageId="17" pageNumber="16" rank="species">
<emphasis id="B97DEACEFFBF3216FC3301441FD2DB78" bold="true" box="[917,942,839,863]" pageId="17" pageNumber="16">sp</emphasis>
.
<emphasis id="B97DEACEFFBF3216FC1801441FB1DB78" bold="true" box="[958,973,839,863]" pageId="17" pageNumber="16">n</emphasis>
.
</taxonomicNameLabel>
, mostly adults, which after discovery always played dead and remain motionless to confuse a potential predator (a behavior known as thanatosis or catalepsy). It is noteworthy to mention here that live individuals fluoresced very weakly under UV light, to the point that they were barely distinguishable from the background. A similar observation was reported by
<bibRefCitation id="EF984B2DFFBF3216FC41060418B7DC38" author="SOLEGLAD" box="[999,1227,1031,1055]" pageId="17" pageNumber="16" pagination="251 - 303" refId="ref14062" refString="SOLEGLAD, M. E. 1976. A revision of the scorpion subfamily Megacorminae (Scorpionida, Chactidae). Wassmann Journal of Biology, 34 (2): 251 - 303." type="journal article" year="1976">Soleglad (1976: 286)</bibRefCitation>
for live and preserved specimens of
<taxonomicName id="4C094D5FFFBF3216FC75062B1829DC18" box="[979,1109,1063,1087]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="gertschi">
<emphasis id="B97DEACEFFBF3216FC75062B1F95DC18" box="[979,1001,1064,1087]" italics="true" pageId="17" pageNumber="16">M</emphasis>
.
<emphasis id="B97DEACEFFBF3216FC5906241829DC18" box="[1023,1109,1063,1087]" italics="true" pageId="17" pageNumber="16">gertschi</emphasis>
</taxonomicName>
. Nevertheless, old exuviae and carcass remains did fluoresce as brightly as many other scorpions, and were readily found while searching in the soil; this allowed us to note that this species also inhabits leaf litter and occurs under rocks partially buried in the litter.
</paragraph>
<paragraph id="8BB636DCFFBF3216FCC606C41FC2DCD8" blockId="17.[816,1475,711,1503]" pageId="17" pageNumber="16">Two females were found carrying litters of young, one at each locality.</paragraph>
<paragraph id="8BB636DCFFBF3216FCC607041954DDF8" blockId="17.[816,1475,711,1503]" pageId="17" pageNumber="16">
In both known localities,
<taxonomicName id="4C094D5FFFBF3216FB3A070B195ADD38" authority="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský, 2023" authorityName="Teruel &amp; Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2023" box="[1180,1318,1287,1311]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="seductus" status="sp. nov.">
<emphasis id="B97DEACEFFBF3216FB3A070B195ADD38" box="[1180,1318,1287,1311]" italics="true" pageId="17" pageNumber="16">M. seductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57B5FFBF3216FA9107041902DD39" box="[1335,1406,1287,1311]" pageId="17" pageNumber="16" rank="species">
<emphasis id="B97DEACEFFBF3216FA910704192CDD38" bold="true" box="[1335,1360,1287,1311]" pageId="17" pageNumber="16">sp</emphasis>
.
<emphasis id="B97DEACEFFBF3216FACF07041904DD38" bold="true" box="[1385,1400,1287,1311]" pageId="17" pageNumber="16">n</emphasis>
.
</taxonomicNameLabel>
lives sympatrically with the buthid
<taxonomicName id="4C094D5FFFBF3216FBC907241F0EDD78" authority="(Latreille, 1804)" baseAuthorityName="Latreille" baseAuthorityYear="1804" class="Arachnida" family="Buthidae" genus="Centruroides" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B97DEACEFFBF3216FBC90724192DDD18" box="[1135,1361,1319,1343]" italics="true" pageId="17" pageNumber="16">Centruroides gracilis</emphasis>
(Latreille, 1804)
</taxonomicName>
. At the
<typeStatus id="54B2887EFFBF3216FC7B07441877DD78" box="[989,1035,1351,1375]" pageId="17" pageNumber="16">type</typeStatus>
locality, it also occurs together with
<taxonomicName id="4C094D5FFFBF3216FC9607641889DD58" authority="(Pocock, 1898)" baseAuthorityName="Pocock" baseAuthorityYear="1898" box="[816,1269,1383,1407]" class="Arachnida" family="Buthidae" genus="Centruroides" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="flavopictus">
<emphasis id="B97DEACEFFBF3216FC9607641842DD58" box="[816,1086,1383,1407]" italics="true" pageId="17" pageNumber="16">Centruroides flavopictus</emphasis>
(Pocock, 1898)
</taxonomicName>
and
<taxonomicName id="4C094D5FFFBF3216FA9007641F15DD98" authority="Goodman, Prendini, Francke &amp; Esposito, 2021" authorityName="Goodman, Prendini, Francke &amp; Esposito" authorityYear="2021" class="Arachnida" family="Buthidae" genus="Centruroides" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="catemacoensis">
<emphasis id="B97DEACEFFBF3216FA9007641FB0DDB8" italics="true" pageId="17" pageNumber="16">Centruroides catemacoensis</emphasis>
Goodman, Prendini, Francke &amp; Esposito, 2021
</taxonomicName>
. In Dos Amates, it also lives sympatrically with the diplocentrid
<taxonomicName id="4C094D5FFFBF3216FC1E07C4195FDDF8" authority="Sissom, 1991" authorityName="Sissom" authorityYear="1991" box="[952,1315,1479,1503]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="perezi">
<emphasis id="B97DEACEFFBF3216FC1E07C418F6DDF8" box="[952,1162,1479,1503]" italics="true" pageId="17" pageNumber="16">Diplocentrus perezi</emphasis>
Sissom, 1991
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>