treatments-xml/data/03/A0/87/03A087B3FFADFFEB009D2669FF6B5FF7.xml

<document id="762B4786D642C5036CF56F8D9792626E" ID-CLB-Dataset="24016" ID-DOI="10.1080/00222933.2019.1656784" ID-GBIF-Dataset="9773657e-8372-45b6-b04f-5df047fb3f6b" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670307" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581941904084" checkinUser="valdenar" docAuthor="ı Tan Van Nguyen, Nikolay A. Poyarkov Jr. &amp; Vogel, Gernot" docDate="2019" docId="03A087B3FFADFFEB009D2669FF6B5FF7" docLanguage="en" docName="JNATHIST.53.27–28.1647–1672.pdf.imf" docOrigin="Journal of Natural History 53 (27)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Liopeltis pallidonuchalis ı Tan Van Nguyen &amp; Vogel 2019, sp. nov." docType="treatment" docVersion="10" lastPageNumber="1663" masterDocId="FF99FFCBFFA4FFF9015A276EFF835A3A" masterDocTitle="A new species of the genus Liopeltis Fitzingerı 1843 from Vietnam (Squamata: Colubridae)" masterLastPageNumber="1672" masterPageNumber="1647" pageNumber="1655" updateTime="1698783637265" updateUser="ExternalLinkService">
<mods:mods id="FCA9D265C03A2AFEF84A26A06F07F87B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="84ABFD0581A7E0C4A00725D53EE46FEF">
<mods:title id="9BDEAF3B0BECBFFE680AC582B8E363AD">A new species of the genus Liopeltis Fitzingerı 1843 from Vietnam (Squamata: Colubridae)</mods:title>
</mods:titleInfo>
<mods:name id="404E0E7AA675B4EA8740B78BB9A60BBF" type="personal">
<mods:role id="84A669AB2FE5D996D68683B4CA892D77">
<mods:roleTerm id="BD17330D09D2F209E7736053D34A5C53">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="02C089C7A182BC8A5488F43F6A9FCED6">ı Tan Van Nguyen, Nikolay A. Poyarkov Jr.</mods:namePart>
</mods:name>
<mods:name id="FCBF7D6AAB4D0B5AD1877B464BFE70F2" type="personal">
<mods:role id="E729A5967188278FDFE7FA893820CBAD">
<mods:roleTerm id="C19EE05A0B348EEDD096AE7EED0BD00B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="68470A33E9CCB8D4E995A082F1AF918D">Vogel, Gernot</mods:namePart>
</mods:name>
<mods:typeOfResource id="1B574A9EF0DD30C21B1B5E2DA85E82AB">text</mods:typeOfResource>
<mods:relatedItem id="A26B20AA355FFEE6AD3920E07AA2D422" type="host">
<mods:titleInfo id="CA68B30E95E509C4B43E5038F84E2E17">
<mods:title id="75A307989821D5FFE75B2E5A72BC183B">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="D782AE035B5DCBA7AA29A952F4930C92">
<mods:date id="BDDD1A93FACDF2B8B72CC0E6759C5E80">2019</mods:date>
<mods:detail id="FBBF64F96DF4EAA9DE75ACA1D16BD63B" type="pubDate">
<mods:number id="22686058D076BB1EB8DEF1FC56E61EC8">2019-09-18</mods:number>
</mods:detail>
<mods:detail id="B6A00981FE0DFE3D05B0E6DFBDFEEB1B" type="volume">
<mods:number id="F9BD18B39A80CF29A58CB0E0F0170511">53</mods:number>
</mods:detail>
<mods:detail id="2EEC1C6EF0D0D996CAF1AE0756EE4F4B" type="issue">
<mods:number id="FEA5E3379001E19B05BE57ED55112121">27</mods:number>
</mods:detail>
<mods:extent id="3B57224C2F261D1244811E83221D4A47" unit="page">
<mods:start id="D3AF37B22DA0C240C3796BE2846179C5">1647</mods:start>
<mods:end id="300B9E16F7B441912698FEDAC8A8BA63">1672</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="91FC2969B2E5BA96A8EB0E38E82D34B8">journal article</mods:classification>
<mods:identifier id="4DF26B36D8BF5EA9613332EA04146D63" type="CLB-Dataset">24016</mods:identifier>
<mods:identifier id="E05CAE15D1F79DCF18A2EB7C3CDEC430" type="DOI">10.1080/00222933.2019.1656784</mods:identifier>
<mods:identifier id="F0EBCFC1370E2CC415F5A79D73B30F33" type="GBIF-Dataset">9773657e-8372-45b6-b04f-5df047fb3f6b</mods:identifier>
<mods:identifier id="4FC2E1A716DB2D488C62AEA3B27FCB07" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="D9BAE00186B8041E1322E02CCF387FCC" type="Zenodo-Dep">3670307</mods:identifier>
</mods:mods>
<treatment id="03A087B3FFADFFEB009D2669FF6B5FF7" ID-DOI="http://doi.org/10.5281/zenodo.3671282" ID-GBIF-Taxon="162146578" ID-Zenodo-Dep="3671282" LSID="urn:lsid:plazi:treatment:03A087B3FFADFFEB009D2669FF6B5FF7" httpUri="http://treatment.plazi.org/id/03A087B3FFADFFEB009D2669FF6B5FF7" lastPageId="18" lastPageNumber="1663" pageId="9" pageNumber="1655">
<subSubSection id="C313652EFFADFFF0009D2669FCDE5B1B" box="[455,861,263,289]" pageId="9" pageNumber="1655" type="nomenclature">
<paragraph id="8BB636A5FFADFFF0009D2669FCDE5B1B" blockId="9.[455,861,263,323]" box="[455,861,263,289]" pageId="9" pageNumber="1655">
<heading id="D0FE81C9FFADFFF0009D2669FCDE5B1B" box="[455,861,263,289]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="1655" reason="2">
<taxonomicName id="4C094D26FFADFFF0009D2669FD735B1B" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[455,752,263,289]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="1655" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFADFFF0009D2669FD735B1B" bold="true" box="[455,752,263,289]" italics="true" pageId="9" pageNumber="1655">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFADFFF003A02666FCDE5B1B" box="[762,861,264,289]" pageId="9" pageNumber="1655" rank="species">
<emphasis id="B97DEAB7FFADFFF003A02666FCD55B1B" bold="true" box="[762,854,264,289]" pageId="9" pageNumber="1655">sp. nov</emphasis>
.
</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C313652EFFADFFF000A42644FEB15BA5" pageId="9" pageNumber="1655" type="description">
<paragraph id="8BB636A5FFADFFF000A42644FCA65B79" blockId="9.[455,861,263,323]" box="[510,805,298,323]" pageId="9" pageNumber="1655">
<heading id="D0FE81C9FFADFFF000A42644FCA65B79" box="[510,805,298,323]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="1655" reason="5">
(
<figureCitation id="13322A20FFADFFF0035C2644FD0C5B79" box="[518,655,298,323]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStartId-0="6.[160,225,1017,1040]" captionStartId-1="10.[160,225,1576,1599]" captionStartId-2="12.[160,225,570,593]" captionStartId-3="12.[160,225,1370,1393]" captionStartId-4="13.[160,225,602,625]" captionTargetBox-0="[163,1153,142,985]" captionTargetBox-1="[165,1152,145,1543]" captionTargetBox-2="[163,1152,143,539]" captionTargetBox-3="[163,1153,681,1338]" captionTargetBox-4="[166,1153,143,569]" captionTargetId-0="figure@6.[163,1153,142,985]" captionTargetId-1="figure@10.[163,1153,142,1545]" captionTargetId-2="figure@12.[163,1153,142,539]" captionTargetId-3="figure@12.[163,1153,681,1339]" captionTargetId-4="figure@13.[163,1153,142,570]" captionTargetPageId-0="6" captionTargetPageId-1="10" captionTargetPageId-2="12" captionTargetPageId-3="12" captionTargetPageId-4="13" captionText-0="Figure 1. Known distribution of Liopeltis frenata and related taxa. Star denotes type locality. Question mark denotes and unconfirmed record of Liopeltis cf. frenata from Bhutan. Locality information. Liopeltis pallidonuchalis sp. nov.: 1 – Kon Chu Rang N.R., Gia Lai,Vietnam (type locality of Liopeltis pallidonuchalis sp. nov.; this work); 2 – Ba Na–Nui Chua N.P., Da Nang, Vietnam (as ‘Tourane’, Smith 1943); 3 – Do Quyen, Bach Ma N.P., Thua Thien–Hue, Vietnam (this work); 4 – U Bo, Phong Nha – Ke Bang N.P., Quang Binh, Vietnam (VNUH 9.7.’06–1; Ziegler et al. 2007; Luu et al. 2013); Liopeltis frenata s.str.: 5 – Khasi Hills, Meghalaya,India (restricted type locality of Cyclophis frenatus Günther,1858; Boulenger 1894; Wall 1908); 6 – Medog, Xizang, China (Zhao and Adler 1993; Zhao 2006); 7 – Mouling N.P., Arunachal Pradesh, India (Pawar and Birand 2001); 8 – Naga Hills, Nagaland, India (Ao et al. 2004); 9 – Namdapha N.P., Arunachal Pradesh, India (Pawar and Birand 2001; Krishna et al. 2012); 10 – Putao, Kachin, Myanmar (CAS 225548- 49); 11 – Htingnan, Kachin, Myanmar (Smith 1940); 12 – Myitkyina, Kachin, Myanmar (AMNH R-58541–44); 13 – Huton, Kachin Hills, Kachin, Myanmar (Wall 1925); 14 – Bhamo, Kachin, Myanmar (Smith 1943); 15 – Mansi, Kachin Hills, Kachin, Myanmar (Wall 1925); 16 – Wuliangshan Mt., Yunnan, China (this paper); 17 – Lüchun, Yunnan, China (KIZ79001; Yang and Rao 2008); 18 – Nam Xe, Van Ban, Lao Cai, Vietnam (IEBR 1205; Orlov et al. 2003); 19 – Copia N.R., Thuan Chau, Son La, Vietnam (TBU PAR.109, 116; Pham et al. 2014); 20 – Houaphan, Laos (Teynié et al. 2014); 21 – Xiangkhong, Laos (as ‘Tran-ninh’ in Smith 1943; Deuve 1970); 22 – Bo Kluea,Nan, Thailand (Hauser 2018); Liopeltis cf. stoliczkae (confirmed localities):23 – Samagooting,Nagaland,India (type locality of Ablabes stoliczkae Sclater,1891; ZSI 3955; see Das et al. 1998); 24 – Darjeeling, West Bengal, India (Smith 1943); 25 – Sikkim, India (no exact locality; Smith 1943); 26 – Karin Hills, Kayah, Myanmar (no exact locality; Smith 1943); Liopeltis cf. stoliczkae (unconfirmed localities): 27 – B La, Bao Lam, Lam Dong, Vietnam (IEBR 1293, as Liopeltis cf. tricolor; Orlov et al. 2003); 28 – Pichrada,Modolkiri,Cambodia (FMNH 259193; Stuart et al. 2006); 29 – Lac Xao, Khamkeut, Bolikhamxay, Laos (FMNH 254780; Stuart and Heatwole 2008); Doubtful localities of Liopeltis spp.in Indochina:30 – Lai Chau,Vietnam (mentioned by Hauser 2018 as Liopeltis frenata, but not" captionText-1="Figure 2. Morphology of Liopeltis pallidonuchalis sp. nov. holotype in life (ZMMU R-15682, male). (a) – general lateral view; (b) – head in lateral view; (c) – head in dorsal view; (d) – head in ventral view. Photos by Nikolay A. Poyarkov." captionText-2="Figure 3. Hemipenial morphology of Liopeltis pallidonuchalis sp. nov. holotype (ZMMU R-15682, male). (a) – asulcal side; (b) – sulcal side. Photos by Nikolay A. Poyarkov." captionText-3="Figure 4. Holotype of Liopeltis pallidonuchalis sp. nov. in situ (ZMMU R-15682, male). Photo by Nikolay A. Poyarkov." captionText-4="Figure 5. Paratype of Liopeltis pallidonuchalis sp. nov. (DTU 307, female, roadkill). (a) – head in dorsolateral view; (b) – head in lateral view; (c) – the general view of the roadkill specimen. Photos by The Anh Nguyen." figureDoi-0="http://doi.org/10.5281/zenodo.3670309" figureDoi-1="http://doi.org/10.5281/zenodo.3670311" figureDoi-2="http://doi.org/10.5281/zenodo.3670313" figureDoi-3="http://doi.org/10.5281/zenodo.3670315" figureDoi-4="http://doi.org/10.5281/zenodo.3670317" httpUri-0="https://zenodo.org/record/3670309/files/figure.png" httpUri-1="https://zenodo.org/record/3670311/files/figure.png" httpUri-2="https://zenodo.org/record/3670313/files/figure.png" httpUri-3="https://zenodo.org/record/3670315/files/figure.png" httpUri-4="https://zenodo.org/record/3670317/files/figure.png" pageId="9" pageNumber="1655">
Figures 1 
<emphasis id="B97DEAB7FFADFFF003292644FD025B79" box="[627,641,298,323]" italics="true" pageId="9" pageNumber="1655">–</emphasis>
5
</figureCitation>
; 
<tableCitation id="C68B031EFFADFFF003C42644FC9E5B79" box="[670,797,298,323]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="3.[133,189,463,486]" captionStartId-1="9.[160,216,1145,1168]" captionTargetBox-0="[144,1702,557,1401]" captionTargetBox-1="[171,1145,1210,1717]" captionTargetPageId-0="3" captionTargetPageId-1="9" captionText-0="Table 1. Basic characters for the genera Liopeltis and Gongylosoma compared to the new species. Abbreviations: BR: Brunei; BT: Bhutan; CA: Cambodia; CN: China; ID: Indonesia; IN: India; LA: Laos; ML: Malaysia; MY: Myanmar; NP: Nepal; PL: Philippines; SG: Singapore; SL: Sri Lanka; TL: Thailand; VN: Vietnam; ‘?’: requires verification; n/a: not available." captionText-1="Table 2. Main morphological variation in the type series and referred materials (ref.mat.) of Liopeltis pallidonuchalis sp. nov. See Materials and Methods for other abbreviations." httpUri-0="http://table.plazi.org/id/DF76662DFFA7FFFA01DF26A1FD8F581B" httpUri-1="http://table.plazi.org/id/DF76662DFFADFFF001FA2317FC0B5E97" pageId="9" pageNumber="1655" tableUuid-0="DF76662DFFA7FFFA01DF26A1FD8F581B" tableUuid-1="DF76662DFFADFFF001FA2317FC0B5E97">
Tables 1 
<emphasis id="B97DEAB7FFADFFF0025B2644FC8C5B79" box="[769,783,298,323]" italics="true" pageId="9" pageNumber="1655">–</emphasis>
2
</tableCitation>
)
</heading>
</paragraph>
<paragraph id="8BB636A5FFADFFF001FA26EBFEB15BA5" blockId="9.[160,1155,389,484]" box="[160,306,389,415]" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001FA26EBFEB15BA5" bold="true" box="[160,306,389,415]" italics="true" pageId="9" pageNumber="1655">Chresonymy</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C313652EFFADFFF001FA26C7FD825BDE" pageId="9" pageNumber="1655" type="reference_group">
<paragraph id="8BB636A5FFADFFF001FA26C7FD825BDE" blockId="9.[160,1155,389,484]" pageId="9" pageNumber="1655">
<treatmentCitationGroup id="AB19118BFFADFFF001FA26C7FD825BDE" pageId="9" pageNumber="1655">
<taxonomicName id="4C094D26FFADFFF001FA26C7FED95BFB" baseAuthorityName="Günther" baseAuthorityYear="1858" box="[160,346,425,449]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="1655" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="B97DEAB7FFADFFF001FA26C7FED95BFB" box="[160,346,425,449]" italics="true" pageId="9" pageNumber="1655">Liopeltis frenatus</emphasis>
</taxonomicName>
(partim) 
<emphasis id="B97DEAB7FFADFFF0009F26C7FE515BFB" box="[453,466,425,449]" italics="true" pageId="9" pageNumber="1655">–</emphasis>
<treatmentCitation id="0AA810B4FFADFFF0008126C7FD265BFB" author="Smith MA" box="[475,677,425,449]" page="183" pageId="9" pageNumber="1655" year="1943">
<bibRefCitation id="EF984B54FFADFFF0008126C7FDEE5BFB" author="Smith MA" box="[475,621,425,449]" pageId="9" pageNumber="1655" pagination="583" refId="ref14375" refString="Smith MA. 1943. The Fauna of British Indiai Ceylon and Burmai including the whole of the Indo- Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). London: Taylor and Francis; p. 583." type="book chapter" year="1943">Smith (1943)</bibRefCitation>
: 183
</treatmentCitation>
; 
<treatmentCitation id="0AA810B4FFADFFF003EB26C7FC3B5BFB" author="Ziegler Ti Hendrix Ri Vu NTi Vogt Mi Forster Bi Dang NK" box="[689,952,425,449]" page="10" pageId="9" pageNumber="1655" year="2007">
<bibRefCitation id="EF984B54FFADFFF003EB26C7FC135BFB" author="Ziegler Ti Hendrix Ri Vu NTi Vogt Mi Forster Bi Dang NK" box="[689,912,425,449]" pageId="9" pageNumber="1655" pagination="1 - 40" refId="ref15010" refString="Ziegler Ti Hendrix Ri Vu NTi Vogt Mi Forster Bi Dang NK. 2007. The diversity of a snake community in a karst forest ecosystem in the central Truong Soni Vietnami with an identification key. Zootaxa. 1493: 1 - 40. doi: 10.11646 / zootaxa. 1493.1.1" type="journal article" year="2007">Ziegler et al. (2007)</bibRefCitation>
: 10
</treatmentCitation>
<emphasis id="B97DEAB7FFADFFF002E226C7FC465BFB" box="[952,965,425,449]" italics="true" pageId="9" pageNumber="1655">–</emphasis>
11ı Figure 11 (?); 
<treatmentCitation id="0AA810B4FFADFFF001FA26A2FE7E5BDE" author="Ziegler Ti Hoang QX" box="[160,509,460,484]" page="115" pageId="9" pageNumber="1655" year="2009">
<bibRefCitation id="EF984B54FFADFFF001FA26A2FE465BDE" author="Ziegler Ti Hoang QX" box="[160,453,460,484]" pageId="9" pageNumber="1655" pagination="115" refId="ref15062" refString="Ziegler Ti Hoang QX. 2009. Liopeltis frenatus. Herpetol Rev. 40 (1): 115." type="journal article" year="2009">Ziegler and Hoang (2009)</bibRefCitation>
: 115
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C313652EFFADFFF001FA257AFC255E05" pageId="9" pageNumber="1655" type="materials_examined">
<paragraph id="8BB636A5FFADFFF001FA257AFDCA58AC" blockId="9.[160,1155,532,662]" pageId="9" pageNumber="1655">
<materialsCitation id="3B613CF8FFADFFF001FA257AFDCA58AC" ID-GBIF-Occurrence="2565890781" collectingDate="2016-05-29" collectionCode="ZMMU, R" collectorName="Kon Chu Rang N. &amp; Nikolay A. Poyarkov" country="Vietnam" elevation="1010" location="Provincei" pageId="9" pageNumber="1655" specimenCount="1" stateProvince="Gia Lai" typeStatus="holotype">
<emphasis id="B97DEAB7FFADFFF001FA257AFE915814" bold="true" box="[160,274,532,558]" italics="true" pageId="9" pageNumber="1655">
<typeStatus id="54B28807FFADFFF001FA257AFE8E5814" box="[160,269,532,558]" pageId="9" pageNumber="1655" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED18AE60FFADFFF0007B2578FEEF5814" box="[289,364,534,558]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="1655" type="Museum">ZMMU</collectionCode>
R-15682ı adult male (hemipenis everted) from forest trail in the evergreen montane tropical forest in 
<collectorName id="26FC5373FFADFFF003612556FC8C586A" box="[571,783,568,592]" pageId="9" pageNumber="1655">Kon Chu Rang N.</collectorName>
<collectionCode id="ED18AE60FFADFFF002542556FCA3586A" box="[782,800,568,592]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="1655">R</collectionCode>
. (14.5034° Nı 108.5383° Eı at elevation of 
<elevation id="0024D196FFADFFF000692535FE345849" box="[307,439,603,627]" metricMagnitude="3" metricUnit="m" metricValue="1.01" pageId="9" pageNumber="1655" unit="m" value="1010.0">
<quantity id="4CF19B40FFADFFF000692535FE095849" box="[307,394,603,627]" metricMagnitude="3" metricUnit="m" metricValue="1.01" pageId="9" pageNumber="1655" unit="m" value="1010.0">1010 m</quantity>
asl
</elevation>
)ı 
<collectingRegion id="49CDF847FFADFFF000902535FD9A5849" box="[458,537,603,627]" country="Vietnam" name="Gia Lai" pageId="9" pageNumber="1655">Gia Lai</collectingRegion>
<location id="8ED6607EFFADFFF003792535FD0A5849" LSID="urn:lsid:plazi:treatment:03A087B3FFADFFEB009D2669FF6B5FF7:8ED6607EFFADFFF003792535FD0A5849" box="[547,649,603,627]" country="Vietnam" name="Provincei" pageId="9" pageNumber="1655" stateProvince="Gia Lai">Provinceı</location>
central 
<collectingCountry id="F31E7635FFADFFF003B12535FCCB5849" box="[747,840,603,627]" name="Vietnam" pageId="9" pageNumber="1655">Vietnam</collectingCountry>
(
<figureCitation id="13322A20FFADFFF002012535FC635849" box="[859,992,603,627]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStartId-0="10.[160,225,1576,1599]" captionStartId-1="12.[160,225,570,593]" captionStartId-2="12.[160,225,1370,1393]" captionTargetBox-0="[165,1152,145,1543]" captionTargetBox-1="[163,1152,143,539]" captionTargetBox-2="[163,1153,681,1338]" captionTargetId-0="figure@10.[163,1153,142,1545]" captionTargetId-1="figure@12.[163,1153,142,539]" captionTargetId-2="figure@12.[163,1153,681,1339]" captionTargetPageId-0="10" captionTargetPageId-1="12" captionTargetPageId-2="12" captionText-0="Figure 2. Morphology of Liopeltis pallidonuchalis sp. nov. holotype in life (ZMMU R-15682, male). (a) – general lateral view; (b) – head in lateral view; (c) – head in dorsal view; (d) – head in ventral view. Photos by Nikolay A. Poyarkov." captionText-1="Figure 3. Hemipenial morphology of Liopeltis pallidonuchalis sp. nov. holotype (ZMMU R-15682, male). (a) – asulcal side; (b) – sulcal side. Photos by Nikolay A. Poyarkov." captionText-2="Figure 4. Holotype of Liopeltis pallidonuchalis sp. nov. in situ (ZMMU R-15682, male). Photo by Nikolay A. Poyarkov." figureDoi-0="http://doi.org/10.5281/zenodo.3670311" figureDoi-1="http://doi.org/10.5281/zenodo.3670313" figureDoi-2="http://doi.org/10.5281/zenodo.3670315" httpUri-0="https://zenodo.org/record/3670311/files/figure.png" httpUri-1="https://zenodo.org/record/3670313/files/figure.png" httpUri-2="https://zenodo.org/record/3670315/files/figure.png" pageId="9" pageNumber="1655">
Figures 2 
<emphasis id="B97DEAB7FFADFFF0029F2535FC515849" box="[965,978,603,627]" italics="true" pageId="9" pageNumber="1655">–</emphasis>
4
</figureCitation>
)ı collected by 
<collectorName id="26FC5373FFADFFF001FA2510FE0258AC" box="[160,385,638,662]" pageId="9" pageNumber="1655">Nikolay A. Poyarkov</collectorName>
on 
<date id="FFB71065FFADFFF000EA2510FDC658AC" box="[432,581,638,662]" pageId="9" pageNumber="1655" value="2016-05-29">
<collectingDate id="EFF3E98DFFADFFF000EA2510FDC658AC" box="[432,581,638,662]" pageId="9" pageNumber="1655" value="2016-05-29">29 May 2016</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB636A5FFADFFF001FA25A8FC0259B7" blockId="9.[160,1155,710,909]" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001FA25A8FE9F58DA" bold="true" box="[160,284,710,736]" italics="true" pageId="9" pageNumber="1655">
<typeStatus id="54B28807FFADFFF001FA25A8FE9458DA" box="[160,279,710,736]" pageId="9" pageNumber="1655" type="paratype">Paratypes</typeStatus>
.
</emphasis>
DTU 307 (adult female) roadkill specimen from the road in the evergreen montane tropical forest in Bach Ma N. 
<materialsCitation id="3B613CF8FFADFFF003392584FE9A591F" ID-GBIF-Occurrence="2565890777" elevation="950" location="Ni" municipality="Ei" pageId="9" pageNumber="1655" specimenCount="1" typeStatus="paratype">
P. (16.2035° 
<location id="8ED6607EFFADFFF003AD2584FC8D5938" LSID="urn:lsid:plazi:treatment:03A087B3FFADFFEB009D2669FF6B5FF7:8ED6607EFFADFFF003AD2584FC8D5938" box="[759,782,746,770]" municipality="Ei" name="Ni" pageId="9" pageNumber="1655">Nı</location>
107.8540° 
<collectingMunicipality id="6BD2ACDFFFADFFF002C92584FC255938" box="[915,934,746,770]" pageId="9" pageNumber="1655">Eı</collectingMunicipality>
at an elevation of 
<elevation id="0024D196FFADFFF001FA2463FE90591F" box="[160,275,781,805]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="9" pageNumber="1655" unit="m" value="950.0">
<quantity id="4CF19B40FFADFFF001FA2463FF6B591F" box="[160,232,781,805]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="9" pageNumber="1655" unit="m" value="950.0">950 m</quantity>
asl
</elevation>
)
</materialsCitation>
<materialsCitation id="3B613CF8FFADFFF000402463FF635950" ID-GBIF-Occurrence="2565890780" collectingDate="2018-06-18" collectionCode="ZFMK" collectorName="Thua Thien &amp; Hue Provincei &amp; The Anh Nguyen &amp; Phuong Nhu Hoang &amp; Bach Ma N" country="Vietnam" location="Vietnam" pageId="9" pageNumber="1655" specimenCode="ZFMK 83105" specimenCount="1" typeStatus="paratype">
ı 
<collectorName id="26FC5373FFADFFF0007F2463FE27591F" box="[293,420,781,805]" pageId="9" pageNumber="1655">Thua Thien</collectorName>
<emphasis id="B97DEAB7FFADFFF000FE2463FE32591F" box="[420,433,781,805]" italics="true" pageId="9" pageNumber="1655">–</emphasis>
<collectorName id="26FC5373FFADFFF000E82463FDCD591F" box="[434,590,781,805]" pageId="9" pageNumber="1655">Hue Provinceı</collectorName>
central 
<collectingCountry id="F31E7635FFADFFF003F72463FC89591F" box="[685,778,781,805]" name="Vietnam" pageId="9" pageNumber="1655">Vietnam</collectingCountry>
(
<figureCitation id="13322A20FFADFFF002412463FCFA591F" box="[795,889,781,805]" captionStart="Figure 5" captionStartId="13.[160,225,602,625]" captionTargetBox="[166,1153,143,569]" captionTargetId="figure@13.[163,1153,142,570]" captionTargetPageId="13" captionText="Figure 5. Paratype of Liopeltis pallidonuchalis sp. nov. (DTU 307, female, roadkill). (a) – head in dorsolateral view; (b) – head in lateral view; (c) – the general view of the roadkill specimen. Photos by The Anh Nguyen." figureDoi="http://doi.org/10.5281/zenodo.3670317" httpUri="https://zenodo.org/record/3670317/files/figure.png" pageId="9" pageNumber="1655">Figure 5</figureCitation>
)ı collected by 
<collectorName id="26FC5373FFADFFF005792463FF7B5972" pageId="9" pageNumber="1655">The Anh Nguyen</collectorName>
and 
<collectorName id="26FC5373FFADFFF0006A245EFD935972" box="[304,528,816,840]" pageId="9" pageNumber="1655">Phuong Nhu Hoang</collectorName>
on 
<date id="FFB71065FFADFFF00361245EFD515972" box="[571,722,816,840]" pageId="9" pageNumber="1655" value="2018-06-18">
<collectingDate id="EFF3E98DFFADFFF00361245EFD515972" box="[571,722,816,840]" pageId="9" pageNumber="1655" value="2018-06-18">18 June 2018</collectingDate>
</date>
; 
<specimenCode id="DBAF9EDEFFADFFF00386245EFCE45972" box="[732,871,816,840]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="9" pageNumber="1655" type="Museum">ZFMK 83105</specimenCode>
(adult female) from 
<collectorName id="26FC5373FFADFFF00517245EFF635950" pageId="9" pageNumber="1655">Bach Ma N</collectorName>
</materialsCitation>
.P. (no elevation data available) 
<materialsCitation id="3B613CF8FFADFFF00315243CFCFE59B7" ID-GBIF-Occurrence="2565890783" collectingDate="1998-08" collectorName="Thua Thien &amp; Hue Provincei &amp; Quang Xuan Hoang" country="Vietnam" location="central Vietnami" pageId="9" pageNumber="1655" specimenCount="1" typeStatus="paratype">
ı 
<collectorName id="26FC5373FFADFFF00307243CFD5E5950" box="[605,733,850,874]" pageId="9" pageNumber="1655">Thua Thien</collectorName>
<emphasis id="B97DEAB7FFADFFF00384243CFD685950" box="[734,747,850,874]" italics="true" pageId="9" pageNumber="1655">–</emphasis>
<collectorName id="26FC5373FFADFFF003B1243CFC0A5950" box="[747,905,850,874]" pageId="9" pageNumber="1655">Hue Provinceı</collectorName>
<location id="8ED6607EFFADFFF002CF243CFBCC5950" LSID="urn:lsid:plazi:treatment:03A087B3FFADFFEB009D2669FF6B5FF7:8ED6607EFFADFFF002CF243CFBCC5950" box="[917,1103,850,874]" country="Vietnam" name="central Vietnami" pageId="9" pageNumber="1655">central Vietnamı</location>
collected by 
<collectorName id="26FC5373FFADFFF00048241BFE7759B7" box="[274,500,885,909]" pageId="9" pageNumber="1655">Quang Xuan Hoang</collectorName>
and students before 
<date id="FFB71065FFADFFF003B1241BFCFE59B7" box="[747,893,885,909]" pageId="9" pageNumber="1655" value="1998-08">
<collectingDate id="EFF3E98DFFADFFF003B1241BFCFE59B7" box="[747,893,885,909]" pageId="9" pageNumber="1655" value="1998-08">August 1998</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB636A5FFADFFF001FA24D3FEF859ED" blockId="9.[160,1156,957,1087]" box="[160,379,957,983]" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001FA24D3FEF859ED" bold="true" box="[160,379,957,983]" italics="true" pageId="9" pageNumber="1655">Referred materials</emphasis>
</paragraph>
<paragraph id="8BB636A5FFADFFF001FA248CFC255E05" blockId="9.[160,1156,957,1087]" pageId="9" pageNumber="1655">
<collectionCode id="ED18AE60FFADFFF001FA248CFF6059C0" box="[160,227,994,1018]" pageId="9" pageNumber="1655">VNUH</collectionCode>
9.7. 
<emphasis id="B97DEAB7FFADFFF0004E248FFE9959C3" box="[276,282,993,1017]" italics="true" pageId="9" pageNumber="1655">’</emphasis>
06 
<emphasis id="B97DEAB7FFADFFF0006F248FFEC159C3" box="[309,322,993,1017]" italics="true" pageId="9" pageNumber="1655">–</emphasis>
1 (adult male) roadkill found on the road during the dry season in the U Bo regionı Phong Nha 
<emphasis id="B97DEAB7FFADFFF0009B236AFE4D5E26" box="[449,462,1028,1052]" italics="true" pageId="9" pageNumber="1655">–</emphasis>
Ke Bang N.P. 
<materialsCitation id="3B613CF8FFADFFF00332236AFC215E05" ID-GBIF-Occurrence="2565890773" collectingDate="2006-07-09" collectorName="Astrid Heidrich" country="Vietnam" elevation="600" location="Provincei" pageId="9" pageNumber="1655" specimenCount="1" stateProvince="Quang Binh">
ı 
<collectingRegion id="49CDF847FFADFFF0032D236AFC835E26" box="[631,768,1028,1052]" country="Vietnam" name="Quang Binh" pageId="9" pageNumber="1655">Quang Binh</collectingRegion>
<location id="8ED6607EFFADFFF00256236AFCF15E26" LSID="urn:lsid:plazi:treatment:03A087B3FFADFFEB009D2669FF6B5FF7:8ED6607EFFADFFF00256236AFCF15E26" box="[780,882,1028,1052]" country="Vietnam" name="Provincei" pageId="9" pageNumber="1655" stateProvince="Quang Binh">Provinceı</location>
central 
<collectingCountry id="F31E7635FFADFFF0028C236AFBB05E26" box="[982,1075,1028,1052]" name="Vietnam" pageId="9" pageNumber="1655">Vietnam</collectingCountry>
(at an elevation of 
<elevation id="0024D196FFADFFF000752349FE215E05" box="[303,418,1063,1087]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="1655" unit="m" value="600.0">
<quantity id="4CF19B40FFADFFF000752349FEF45E05" box="[303,375,1063,1087]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="1655" unit="m" value="600.0">600 m</quantity>
asl
</elevation>
)ı collected by 
<collectorName id="26FC5373FFADFFF0031D2349FD6C5E05" box="[583,751,1063,1087]" pageId="9" pageNumber="1655">Astrid Heidrich</collectorName>
on 
<date id="FFB71065FFADFFF002442349FC215E05" box="[798,930,1063,1087]" pageId="9" pageNumber="1655" value="2006-07-09">
<collectingDate id="EFF3E98DFFADFFF002442349FC215E05" box="[798,930,1063,1087]" pageId="9" pageNumber="1655" value="2006-07-09">9 July 2006</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF76662DFFADFFF001FA2317FC0B5E97" ID-Table-UUID="DF76662DFFADFFF001FA2317FC0B5E97" httpUri="http://table.plazi.org/id/DF76662DFFADFFF001FA2317FC0B5E97" pageId="9" pageNumber="1655" targetBox="[171,1145,1210,1717]" targetIsTable="true" targetPageId="9">
<paragraph id="8BB636A5FFADFFF001FA2317FC0B5E97" blockId="9.[160,1156,1144,1197]" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001FA2317FF715EAA" bold="true" box="[160,242,1145,1168]" pageId="9" pageNumber="1655">Table 2.</emphasis>
Main morphological variation in the type series and referred materials (ref.mat.) of 
<taxonomicName id="4C094D26FFADFFF0056C2316FEB35E97" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="1655" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFADFFF0056C2316FEB35E97" italics="true" pageId="9" pageNumber="1655">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFADFFF0006223F8FEF85E97" box="[312,379,1174,1197]" pageId="9" pageNumber="1655" rank="species">
<emphasis id="B97DEAB7FFADFFF0006223F8FEF55E97" bold="true" box="[312,374,1174,1197]" italics="true" pageId="9" pageNumber="1655">sp. nov</emphasis>
.
</taxonomicNameLabel>
See Materials and Methods for other abbreviations.
</paragraph>
</caption>
<paragraph id="8BB636A5FFADFFF001F123D4FBFA5C8F" pageId="9" pageNumber="1655">
<table id="F909C405FFAD000601F123D4FBFA5C8F" box="[171,1145,1210,1717]" gridcols="5" gridrows="21" pageId="9" pageNumber="1655">
<tr id="353934E7FFAD000601F123D4FBFA5EF7" box="[171,1145,1210,1229]" gridrow="0" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F123D4FE0D5EF7" box="[171,398,1210,1229]" gridcol="0" gridrow="0" pageId="9" pageNumber="1655">Number</th>
<th id="76E85D9BFFAD0006008823D4FDDB5EF7" box="[466,600,1210,1229]" gridcol="1" gridrow="0" pageId="9" pageNumber="1655">
ZMMU R 
<emphasis id="B97DEAB7FFADFFF0034023D4FDA65EF7" box="[538,549,1210,1229]" italics="true" pageId="9" pageNumber="1655">–</emphasis>
15682
</th>
<th id="76E85D9BFFAD000603DC23D4FD675EF7" box="[646,740,1210,1229]" gridcol="2" gridrow="0" pageId="9" pageNumber="1655">DTU 307</th>
<th id="76E85D9BFFAD0006027223D4FC135EF7" box="[808,912,1210,1229]" gridcol="3" gridrow="0" pageId="9" pageNumber="1655">ZFMK 83105</th>
<th id="76E85D9BFFAD0006028923D4FBFA5EF7" box="[979,1145,1210,1229]" gridcol="4" gridrow="0" pageId="9" pageNumber="1655">
VNUH 9.7. 
<emphasis id="B97DEAB7FFADFFF0056623D4FBC35EF7" box="[1084,1088,1210,1229]" italics="true" pageId="9" pageNumber="1655">’</emphasis>
06-1
</th>
</tr>
<tr id="353934E7FFAD000601F123B4FBFA5ED7" box="[171,1145,1242,1261]" gridrow="1" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F123B4FE0D5ED7" box="[171,398,1242,1261]" gridcol="0" gridrow="1" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F123B4FF7F5ED7" bold="true" box="[171,252,1242,1261]" italics="true" pageId="9" pageNumber="1655">Type status</emphasis>
</th>
<td id="76E85D9BFFAD0006008823B4FDDB5ED7" box="[466,600,1242,1261]" gridcol="1" gridrow="1" pageId="9" pageNumber="1655">Holotype</td>
<td id="76E85D9BFFAD000603DC23B4FD675ED7" box="[646,740,1242,1261]" gridcol="2" gridrow="1" pageId="9" pageNumber="1655">Paratype</td>
<td id="76E85D9BFFAD0006027223B4FC135ED7" box="[808,912,1242,1261]" gridcol="3" gridrow="1" pageId="9" pageNumber="1655">Paratype</td>
<td id="76E85D9BFFAD0006028923B4FBFA5ED7" box="[979,1145,1242,1261]" gridcol="4" gridrow="1" pageId="9" pageNumber="1655">ref.mat.</td>
</tr>
<tr id="353934E7FFAD000601F1239CFBFA5F3F" box="[171,1145,1266,1285]" gridrow="2" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F1239CFE0D5F3F" box="[171,398,1266,1285]" gridcol="0" gridrow="2" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F1239CFF415F3F" bold="true" box="[171,194,1266,1285]" italics="true" pageId="9" pageNumber="1655">Sex</emphasis>
</th>
<td id="76E85D9BFFAD00060088239CFDDB5F3F" box="[466,600,1266,1285]" gridcol="1" gridrow="2" pageId="9" pageNumber="1655">M</td>
<td id="76E85D9BFFAD000603DC239CFD675F3F" box="[646,740,1266,1285]" gridcol="2" gridrow="2" pageId="9" pageNumber="1655">F</td>
<td id="76E85D9BFFAD00060272239CFC135F3F" box="[808,912,1266,1285]" gridcol="3" gridrow="2" pageId="9" pageNumber="1655">F</td>
<td id="76E85D9BFFAD00060289239CFBFA5F3F" box="[979,1145,1266,1285]" gridcol="4" gridrow="2" pageId="9" pageNumber="1655">M</td>
</tr>
<tr id="353934E7FFAD000601F12264FBFA5F27" box="[171,1145,1290,1309]" gridrow="3" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F12264FE0D5F27" box="[171,398,1290,1309]" gridcol="0" gridrow="3" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F12264FF405F27" bold="true" box="[171,195,1290,1309]" italics="true" pageId="9" pageNumber="1655">SVL</emphasis>
</th>
<td id="76E85D9BFFAD000600882264FDDB5F27" box="[466,600,1290,1309]" gridcol="1" gridrow="3" pageId="9" pageNumber="1655">297</td>
<td id="76E85D9BFFAD000603DC2264FD675F27" box="[646,740,1290,1309]" gridcol="2" gridrow="3" pageId="9" pageNumber="1655">275</td>
<td id="76E85D9BFFAD000602722264FC135F27" box="[808,912,1290,1309]" gridcol="3" gridrow="3" pageId="9" pageNumber="1655">336</td>
<td id="76E85D9BFFAD000602892264FBFA5F27" box="[979,1145,1290,1309]" gridcol="4" gridrow="3" pageId="9" pageNumber="1655">270</td>
</tr>
<tr id="353934E7FFAD000601F1224CFBFA5F0F" box="[171,1145,1314,1333]" gridrow="4" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F1224CFE0D5F0F" box="[171,398,1314,1333]" gridcol="0" gridrow="4" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F1224CFF415F0F" bold="true" box="[171,194,1314,1333]" italics="true" pageId="9" pageNumber="1655">TaL</emphasis>
</th>
<td id="76E85D9BFFAD00060088224CFDDB5F0F" box="[466,600,1314,1333]" gridcol="1" gridrow="4" pageId="9" pageNumber="1655">112</td>
<td id="76E85D9BFFAD000603DC224CFD675F0F" box="[646,740,1314,1333]" gridcol="2" gridrow="4" pageId="9" pageNumber="1655">111</td>
<td id="76E85D9BFFAD00060272224CFC135F0F" box="[808,912,1314,1333]" gridcol="3" gridrow="4" pageId="9" pageNumber="1655">130</td>
<td id="76E85D9BFFAD00060289224CFBFA5F0F" box="[979,1145,1314,1333]" gridcol="4" gridrow="4" pageId="9" pageNumber="1655">116</td>
</tr>
<tr id="353934E7FFAD000601F12254FBFA5F77" box="[171,1145,1338,1357]" gridrow="5" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F12254FE0D5F77" box="[171,398,1338,1357]" gridcol="0" gridrow="5" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F12254FF395F77" bold="true" box="[171,186,1338,1357]" italics="true" pageId="9" pageNumber="1655">TL</emphasis>
</th>
<td id="76E85D9BFFAD000600882254FDDB5F77" box="[466,600,1338,1357]" gridcol="1" gridrow="5" pageId="9" pageNumber="1655">409</td>
<td id="76E85D9BFFAD000603DC2254FD675F77" box="[646,740,1338,1357]" gridcol="2" gridrow="5" pageId="9" pageNumber="1655">386</td>
<td id="76E85D9BFFAD000602722254FC135F77" box="[808,912,1338,1357]" gridcol="3" gridrow="5" pageId="9" pageNumber="1655">466</td>
<td id="76E85D9BFFAD000602892254FBFA5F77" box="[979,1145,1338,1357]" gridcol="4" gridrow="5" pageId="9" pageNumber="1655">386</td>
</tr>
<tr id="353934E7FFAD000601F1223CFBFA5F5F" box="[171,1145,1362,1381]" gridrow="6" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F1223CFE0D5F5F" box="[171,398,1362,1381]" gridcol="0" gridrow="6" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F1223CFF605F5F" bold="true" box="[171,227,1362,1381]" pageId="9" pageNumber="1655">Rel TL</emphasis>
</th>
<td id="76E85D9BFFAD00060088223CFDDB5F5F" box="[466,600,1362,1381]" gridcol="1" gridrow="6" pageId="9" pageNumber="1655">0.274</td>
<td id="76E85D9BFFAD000603DC223CFD675F5F" box="[646,740,1362,1381]" gridcol="2" gridrow="6" pageId="9" pageNumber="1655">2.88</td>
<td id="76E85D9BFFAD00060272223CFC135F5F" box="[808,912,1362,1381]" gridcol="3" gridrow="6" pageId="9" pageNumber="1655">0.279</td>
<td id="76E85D9BFFAD00060289223CFBFA5F5F" box="[979,1145,1362,1381]" gridcol="4" gridrow="6" pageId="9" pageNumber="1655">0.301</td>
</tr>
<tr id="353934E7FFAD000601F12204FBFA5F47" box="[171,1145,1386,1405]" gridrow="7" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F12204FE0D5F47" box="[171,398,1386,1405]" gridcol="0" gridrow="7" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F12204FF4C5F47" bold="true" box="[171,207,1386,1405]" pageId="9" pageNumber="1655">DSR</emphasis>
</th>
<td id="76E85D9BFFAD000600882204FDDB5F47" box="[466,600,1386,1405]" gridcol="1" gridrow="7" pageId="9" pageNumber="1655">15-15-15</td>
<td id="76E85D9BFFAD000603DC2204FD675F47" box="[646,740,1386,1405]" gridcol="2" gridrow="7" pageId="9" pageNumber="1655">15-15-15</td>
<td id="76E85D9BFFAD000602722204FC135F47" box="[808,912,1386,1405]" gridcol="3" gridrow="7" pageId="9" pageNumber="1655">15-15-15</td>
<td id="76E85D9BFFAD000602892204FBFA5F47" box="[979,1145,1386,1405]" gridcol="4" gridrow="7" pageId="9" pageNumber="1655">15-15-15</td>
</tr>
<tr id="353934E7FFAD000601F122ECFBFA5FAF" box="[171,1145,1410,1429]" gridrow="8" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F122ECFE0D5FAF" box="[171,398,1410,1429]" gridcol="0" gridrow="8" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F122ECFF415FAF" bold="true" box="[171,194,1410,1429]" pageId="9" pageNumber="1655">VE</emphasis>
</th>
<td id="76E85D9BFFAD0006008822ECFDDB5FAF" box="[466,600,1410,1429]" gridcol="1" gridrow="8" pageId="9" pageNumber="1655">2 + 133</td>
<td id="76E85D9BFFAD000603DC22ECFD675FAF" box="[646,740,1410,1429]" gridcol="2" gridrow="8" pageId="9" pageNumber="1655">2 + 138</td>
<td id="76E85D9BFFAD0006027222ECFC135FAF" box="[808,912,1410,1429]" gridcol="3" gridrow="8" pageId="9" pageNumber="1655">2 + 138</td>
<td id="76E85D9BFFAD0006028922ECFBFA5FAF" box="[979,1145,1410,1429]" gridcol="4" gridrow="8" pageId="9" pageNumber="1655">4 + 126</td>
</tr>
<tr id="353934E7FFAD000601F122F4FBFA5F97" box="[171,1145,1434,1453]" gridrow="9" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F122F4FE0D5F97" box="[171,398,1434,1453]" gridcol="0" gridrow="9" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F122F4FF425F97" bold="true" box="[171,193,1434,1453]" pageId="9" pageNumber="1655">SC</emphasis>
</th>
<td id="76E85D9BFFAD0006008822F4FDDB5F97" box="[466,600,1434,1453]" gridcol="1" gridrow="9" pageId="9" pageNumber="1655">67</td>
<td id="76E85D9BFFAD000603DC22F4FD675F97" box="[646,740,1434,1453]" gridcol="2" gridrow="9" pageId="9" pageNumber="1655">73</td>
<td id="76E85D9BFFAD0006027222F4FC135F97" box="[808,912,1434,1453]" gridcol="3" gridrow="9" pageId="9" pageNumber="1655">71</td>
<td id="76E85D9BFFAD0006028922F4FBFA5F97" box="[979,1145,1434,1453]" gridcol="4" gridrow="9" pageId="9" pageNumber="1655">69</td>
</tr>
<tr id="353934E7FFAD000601F122DCFBFA5FFF" box="[171,1145,1458,1477]" gridrow="10" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F122DCFE0D5FFF" box="[171,398,1458,1477]" gridcol="0" gridrow="10" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F122DCFF575FFF" bold="true" box="[171,212,1458,1477]" pageId="9" pageNumber="1655">Anal</emphasis>
</th>
<td id="76E85D9BFFAD0006008822DCFDDB5FFF" box="[466,600,1458,1477]" gridcol="1" gridrow="10" pageId="9" pageNumber="1655">2</td>
<td id="76E85D9BFFAD000603DC22DCFD675FFF" box="[646,740,1458,1477]" gridcol="2" gridrow="10" pageId="9" pageNumber="1655">2</td>
<td id="76E85D9BFFAD0006027222DCFC135FFF" box="[808,912,1458,1477]" gridcol="3" gridrow="10" pageId="9" pageNumber="1655">2</td>
<td id="76E85D9BFFAD0006028922DCFBFA5FFF" box="[979,1145,1458,1477]" gridcol="4" gridrow="10" pageId="9" pageNumber="1655">2</td>
</tr>
<tr id="353934E7FFAD000601F122A4FBFA5FE7" box="[171,1145,1482,1501]" gridrow="11" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F122A4FE0D5FE7" box="[171,398,1482,1501]" gridcol="0" gridrow="11" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F122A4FF485FE7" bold="true" box="[171,203,1482,1501]" pageId="9" pageNumber="1655">SLb</emphasis>
</th>
<td id="76E85D9BFFAD0006008822A4FDDB5FE7" box="[466,600,1482,1501]" gridcol="1" gridrow="11" pageId="9" pageNumber="1655">7</td>
<td id="76E85D9BFFAD000603DC22A4FD675FE7" box="[646,740,1482,1501]" gridcol="2" gridrow="11" pageId="9" pageNumber="1655">7</td>
<td id="76E85D9BFFAD0006027222A4FC135FE7" box="[808,912,1482,1501]" gridcol="3" gridrow="11" pageId="9" pageNumber="1655">7</td>
<td id="76E85D9BFFAD0006028922A4FBFA5FE7" box="[979,1145,1482,1501]" gridcol="4" gridrow="11" pageId="9" pageNumber="1655">7</td>
</tr>
<tr id="353934E7FFAD000601F1228CFBFA5FCF" box="[171,1145,1506,1525]" gridrow="12" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F1228CFE0D5FCF" box="[171,398,1506,1525]" gridcol="0" gridrow="12" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F1228CFF495FCF" bold="true" box="[171,202,1506,1525]" pageId="9" pageNumber="1655">SLE</emphasis>
</th>
<td id="76E85D9BFFAD00060088228CFDDB5FCF" box="[466,600,1506,1525]" gridcol="1" gridrow="12" pageId="9" pageNumber="1655">
3 
<emphasis id="B97DEAB7FFADFFF00355228CFD995FCF" box="[527,538,1506,1525]" italics="true" pageId="9" pageNumber="1655">–</emphasis>
4
</td>
<td id="76E85D9BFFAD000603DC228CFD675FCF" box="[646,740,1506,1525]" gridcol="2" gridrow="12" pageId="9" pageNumber="1655">
3 
<emphasis id="B97DEAB7FFADFFF003E0228CFD465FCF" box="[698,709,1506,1525]" italics="true" pageId="9" pageNumber="1655">–</emphasis>
4
</td>
<td id="76E85D9BFFAD00060272228CFC135FCF" box="[808,912,1506,1525]" gridcol="3" gridrow="12" pageId="9" pageNumber="1655">
3 
<emphasis id="B97DEAB7FFADFFF0020C228CFCE25FCF" box="[854,865,1506,1525]" italics="true" pageId="9" pageNumber="1655">–</emphasis>
4
</td>
<td id="76E85D9BFFAD00060289228CFBFA5FCF" box="[979,1145,1506,1525]" gridcol="4" gridrow="12" pageId="9" pageNumber="1655">
3 
<emphasis id="B97DEAB7FFADFFF0057B228CFBAF5FCF" box="[1057,1068,1506,1525]" italics="true" pageId="9" pageNumber="1655">–</emphasis>
4
</td>
</tr>
<tr id="353934E7FFAD000601F12294FBFA5C37" box="[171,1145,1530,1549]" gridrow="13" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F12294FE0D5C37" box="[171,398,1530,1549]" gridcol="0" gridrow="13" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F12294FF445C37" bold="true" box="[171,199,1530,1549]" pageId="9" pageNumber="1655">ILb</emphasis>
</th>
<td id="76E85D9BFFAD000600882294FDDB5C37" box="[466,600,1530,1549]" gridcol="1" gridrow="13" pageId="9" pageNumber="1655">8</td>
<td id="76E85D9BFFAD000603DC2294FD675C37" box="[646,740,1530,1549]" gridcol="2" gridrow="13" pageId="9" pageNumber="1655">8</td>
<td id="76E85D9BFFAD000602722294FC135C37" box="[808,912,1530,1549]" gridcol="3" gridrow="13" pageId="9" pageNumber="1655">8</td>
<td id="76E85D9BFFAD000602892294FBFA5C37" box="[979,1145,1530,1549]" gridcol="4" gridrow="13" pageId="9" pageNumber="1655">7</td>
</tr>
<tr id="353934E7FFAD000601F1217CFBFA5C1F" box="[171,1145,1554,1573]" gridrow="14" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F1217CFE0D5C1F" box="[171,398,1554,1573]" gridcol="0" gridrow="14" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F1217CFF605C1F" bold="true" box="[171,227,1554,1573]" pageId="9" pageNumber="1655">Loreal</emphasis>
</th>
<td id="76E85D9BFFAD00060088217CFDDB5C1F" box="[466,600,1554,1573]" gridcol="1" gridrow="14" pageId="9" pageNumber="1655">0</td>
<td id="76E85D9BFFAD000603DC217CFD675C1F" box="[646,740,1554,1573]" gridcol="2" gridrow="14" pageId="9" pageNumber="1655">1</td>
<td id="76E85D9BFFAD00060272217CFC135C1F" box="[808,912,1554,1573]" gridcol="3" gridrow="14" pageId="9" pageNumber="1655">1</td>
<td id="76E85D9BFFAD00060289217CFBFA5C1F" box="[979,1145,1554,1573]" gridcol="4" gridrow="14" pageId="9" pageNumber="1655">1</td>
</tr>
<tr id="353934E7FFAD000601F12144FBFA5C07" box="[171,1145,1578,1597]" gridrow="15" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F12144FE0D5C07" box="[171,398,1578,1597]" gridcol="0" gridrow="15" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F12144FE0D5C07" bold="true" box="[171,398,1578,1597]" pageId="9" pageNumber="1655">Praefrontal touching SLb</emphasis>
</th>
<td id="76E85D9BFFAD000600882144FDDB5C07" box="[466,600,1578,1597]" gridcol="1" gridrow="15" pageId="9" pageNumber="1655">1</td>
<td id="76E85D9BFFAD000603DC2144FD675C07" box="[646,740,1578,1597]" gridcol="2" gridrow="15" pageId="9" pageNumber="1655">0</td>
<td id="76E85D9BFFAD000602722144FC135C07" box="[808,912,1578,1597]" gridcol="3" gridrow="15" pageId="9" pageNumber="1655">0</td>
<td id="76E85D9BFFAD000602892144FBFA5C07" box="[979,1145,1578,1597]" gridcol="4" gridrow="15" pageId="9" pageNumber="1655">0</td>
</tr>
<tr id="353934E7FFAD000601F1212CFBFA5C6F" box="[171,1145,1602,1621]" gridrow="16" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F1212CFE0D5C6F" box="[171,398,1602,1621]" gridcol="0" gridrow="16" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F1212CFF4F5C6F" bold="true" box="[171,204,1602,1621]" pageId="9" pageNumber="1655">PrO</emphasis>
</th>
<td id="76E85D9BFFAD00060088212CFDDB5C6F" box="[466,600,1602,1621]" gridcol="1" gridrow="16" pageId="9" pageNumber="1655">1</td>
<td id="76E85D9BFFAD000603DC212CFD675C6F" box="[646,740,1602,1621]" gridcol="2" gridrow="16" pageId="9" pageNumber="1655">1</td>
<td id="76E85D9BFFAD00060272212CFC135C6F" box="[808,912,1602,1621]" gridcol="3" gridrow="16" pageId="9" pageNumber="1655">1</td>
<td id="76E85D9BFFAD00060289212CFBFA5C6F" box="[979,1145,1602,1621]" gridcol="4" gridrow="16" pageId="9" pageNumber="1655">1</td>
</tr>
<tr id="353934E7FFAD000601F12134FBFA5C57" box="[171,1145,1626,1645]" gridrow="17" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F12134FE0D5C57" box="[171,398,1626,1645]" gridcol="0" gridrow="17" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F12134FF535C57" bold="true" box="[171,208,1626,1645]" pageId="9" pageNumber="1655">PoO</emphasis>
</th>
<td id="76E85D9BFFAD000600882134FDDB5C57" box="[466,600,1626,1645]" gridcol="1" gridrow="17" pageId="9" pageNumber="1655">2</td>
<td id="76E85D9BFFAD000603DC2134FD675C57" box="[646,740,1626,1645]" gridcol="2" gridrow="17" pageId="9" pageNumber="1655">2</td>
<td id="76E85D9BFFAD000602722134FC135C57" box="[808,912,1626,1645]" gridcol="3" gridrow="17" pageId="9" pageNumber="1655">2</td>
<td id="76E85D9BFFAD000602892134FBFA5C57" box="[979,1145,1626,1645]" gridcol="4" gridrow="17" pageId="9" pageNumber="1655">2</td>
</tr>
<tr id="353934E7FFAD000601F1211CFBFA5CBF" box="[171,1145,1650,1669]" gridrow="18" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F1211CFE0D5CBF" box="[171,398,1650,1669]" gridcol="0" gridrow="18" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F1211CFF3C5CBF" bold="true" box="[171,191,1650,1669]" pageId="9" pageNumber="1655">At</emphasis>
</th>
<td id="76E85D9BFFAD00060088211CFDDB5CBF" box="[466,600,1650,1669]" gridcol="1" gridrow="18" pageId="9" pageNumber="1655">1</td>
<td id="76E85D9BFFAD000603DC211CFD675CBF" box="[646,740,1650,1669]" gridcol="2" gridrow="18" pageId="9" pageNumber="1655">1</td>
<td id="76E85D9BFFAD00060272211CFC135CBF" box="[808,912,1650,1669]" gridcol="3" gridrow="18" pageId="9" pageNumber="1655">1</td>
<td id="76E85D9BFFAD00060289211CFBFA5CBF" box="[979,1145,1650,1669]" gridcol="4" gridrow="18" pageId="9" pageNumber="1655">1</td>
</tr>
<tr id="353934E7FFAD000601F121E4FBFA5CA7" box="[171,1145,1674,1693]" gridrow="19" pageId="9" pageNumber="1655">
<th id="76E85D9BFFAD000601F121E4FE0D5CA7" box="[171,398,1674,1693]" gridcol="0" gridrow="19" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F121E4FF3D5CA7" bold="true" box="[171,190,1674,1693]" pageId="9" pageNumber="1655">Pt</emphasis>
</th>
<td id="76E85D9BFFAD0006008821E4FDDB5CA7" box="[466,600,1674,1693]" gridcol="1" gridrow="19" pageId="9" pageNumber="1655">2</td>
<td id="76E85D9BFFAD000603DC21E4FD675CA7" box="[646,740,1674,1693]" gridcol="2" gridrow="19" pageId="9" pageNumber="1655">2</td>
<td id="76E85D9BFFAD0006027221E4FC135CA7" box="[808,912,1674,1693]" gridcol="3" gridrow="19" pageId="9" pageNumber="1655">2</td>
<td id="76E85D9BFFAD0006028921E4FBFA5CA7" box="[979,1145,1674,1693]" gridcol="4" gridrow="19" pageId="9" pageNumber="1655">2</td>
</tr>
<tr id="353934E7FFAD000601F121CCFBFA5C8F" box="[171,1145,1698,1717]" gridrow="20" pageId="9" pageNumber="1655" rowspan-1="1" rowspan-3="1">
<th id="76E85D9BFFAD000601F121CCFE0D5C8F" box="[171,398,1698,1717]" gridcol="0" gridrow="20" pageId="9" pageNumber="1655">
<emphasis id="B97DEAB7FFADFFF001F121CCFF5B5C8F" bold="true" box="[171,216,1698,1717]" italics="true" pageId="9" pageNumber="1655">Source</emphasis>
</th>
<td id="76E85D9BFFAD000603DC21CCFD675C8F" box="[646,740,1698,1717]" gridcol="2" gridrow="20" pageId="9" pageNumber="1655">This study</td>
<td id="76E85D9BFFAD0006028921CCFBFA5C8F" box="[979,1145,1698,1717]" gridcol="4" gridrow="20" pageId="9" pageNumber="1655">
<bibRefCitation id="EF984B54FFADFFF0028921CCFBFA5C8F" author="Ziegler Ti Hendrix Ri Vu NTi Vogt Mi Forster Bi Dang NK" box="[979,1145,1698,1717]" pageId="9" pageNumber="1655" pagination="1 - 40" refId="ref15010" refString="Ziegler Ti Hendrix Ri Vu NTi Vogt Mi Forster Bi Dang NK. 2007. The diversity of a snake community in a karst forest ecosystem in the central Truong Soni Vietnami with an identification key. Zootaxa. 1493: 1 - 40. doi: 10.11646 / zootaxa. 1493.1.1" type="journal article" year="2007">Ziegler et al. (2007)</bibRefCitation>
</td>
</tr>
</table>
</paragraph>
<caption id="DF76662DFFAEFFF301FA2146FD965C40" ID-DOI="http://doi.org/10.5281/zenodo.3670311" ID-Zenodo-Dep="3670311" httpUri="https://zenodo.org/record/3670311/files/figure.png" pageId="10" pageNumber="1656" startId="10.[160,225,1576,1599]" targetBox="[165,1152,145,1543]" targetPageId="10">
<paragraph id="8BB636A5FFAEFFF301FA2146FD965C40" blockId="10.[160,1156,1575,1658]" pageId="10" pageNumber="1656">
<emphasis id="B97DEAB7FFAEFFF301FA2146FF7E5C05" bold="true" box="[160,253,1576,1599]" pageId="10" pageNumber="1656">Figure 2.</emphasis>
Morphology of 
<taxonomicName id="4C094D26FFAEFFF300FD2149FD0D5C04" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[423,654,1575,1598]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="1656" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFAEFFF300FD2149FD0D5C04" box="[423,654,1575,1598]" italics="true" pageId="10" pageNumber="1656">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFAEFFF303C22146FD6B5C05" box="[664,744,1576,1599]" pageId="10" pageNumber="1656" rank="species">sp. nov.</taxonomicNameLabel>
holotype in life (ZMMU R-15682, male). (a) 
<emphasis id="B97DEAB7FFAEFFF3019B212BFF4D5C66" box="[193,206,1605,1628]" italics="true" pageId="10" pageNumber="1656">–</emphasis>
general lateral view; (b) 
<emphasis id="B97DEAB7FFAEFFF30093212BFE555C66" box="[457,470,1605,1628]" italics="true" pageId="10" pageNumber="1656">–</emphasis>
head in lateral view; (c) 
<emphasis id="B97DEAB7FFAEFFF3038A212BFD5E5C66" box="[720,733,1605,1628]" italics="true" pageId="10" pageNumber="1656">–</emphasis>
head in dorsal view; (d) 
<emphasis id="B97DEAB7FFAEFFF30282212BFC665C66" box="[984,997,1605,1628]" italics="true" pageId="10" pageNumber="1656">–</emphasis>
head in ventral view. Photos by Nikolay A. Poyarkov.
</paragraph>
</caption>
<subSubSection id="C313652EFFAFFFF201FA27E3FCDA5838" pageId="11" pageNumber="1657" type="diagnosis">
<paragraph id="8BB636A5FFAFFFF201FA27E3FCDA5838" blockId="11.[160,1157,141,514]" pageId="11" pageNumber="1657">
<emphasis id="B97DEAB7FFAFFFF201FA27E3FE995A9D" bold="true" box="[160,282,141,167]" italics="true" pageId="11" pageNumber="1657">Diagnosis.</emphasis>
The new species can be separated from its congeners by the following combination of morphological characters: (1) one single (or missing) loreal; (2) one single nasal shield; (3) dorsal scales in 15-15-15 rowsı all smooth; (4) 1 preocuları 2 postoculars and 1 + 2 temporals; (5) 7 supralabialsı of which 3d and 4th in contact with the eye; (6) 8 infralabials; (7) nasal contacting internasal; (8) prefrontal touching or separated from supralabials; (9) ventral scales 126 
<emphasis id="B97DEAB7FFAFFFF203B42652FD785B6E" box="[750,763,316,340]" italics="true" pageId="11" pageNumber="1657">–</emphasis>
138; (10) subcaudal scales 67 
<emphasis id="B97DEAB7FFAFFFF2050F2652FBE15B6E" box="[1109,1122,316,340]" italics="true" pageId="11" pageNumber="1657">–</emphasis>
73ı paired; (11) relative tail length about 
<geoCoordinate id="EE3D5062FFAFFFF2031C2631FD015B4D" box="[582,642,351,375]" degrees="0.274" direction="north" orientation="latitude" pageId="11" pageNumber="1657" precision="55" value="0.274">0.274</geoCoordinate>
<geoCoordinate id="EE3D5062FFAFFFF203D82631FD4C5B4D" box="[642,719,351,375]" degrees="0.301" direction="west" orientation="longitude" pageId="11" pageNumber="1657" precision="55" value="-0.301">
<emphasis id="B97DEAB7FFAFFFF203D82631FD0C5B4D" box="[642,655,351,375]" italics="true" pageId="11" pageNumber="1657">–</emphasis>
0.301
</geoCoordinate>
; (12) an uniform bronze body colouration; (13) a thin postocular stripe extending from eye to end of the neck becoming indistinct posteriorly. The new species can be distinguished from all other congeners by its low subcaudal scales countı and by its characteristic colouration. Detailed comparisons with other species of the genus 
<taxonomicName id="4C094D26FFAFFFF2030F2684FD2E5838" authorityName="Fitzingeri" authorityYear="1843" box="[597,685,490,514]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="1657" phylum="Chordata" rank="genus">
<emphasis id="B97DEAB7FFAFFFF2030F2684FD2E5838" box="[597,685,490,514]" italics="true" pageId="11" pageNumber="1657">Liopeltis</emphasis>
</taxonomicName>
appear below.
</paragraph>
</subSubSection>
<subSubSection id="C313652EFFAFFFF701FA2553FCAC5B86" lastPageId="14" lastPageNumber="1660" pageId="11" pageNumber="1657" type="description">
<paragraph id="8BB636A5FFAFFFF201FA2553FE7E5885" blockId="11.[160,1156,573,1708]" pageId="11" pageNumber="1657">
<emphasis id="B97DEAB7FFAFFFF201FA2553FE41586D" bold="true" box="[160,450,573,599]" italics="true" pageId="11" pageNumber="1657">
Description of 
<typeStatus id="54B28807FFAFFFF200092553FE3E586D" box="[339,445,573,599]" pageId="11" pageNumber="1657" type="holotype">holotype</typeStatus>
.
</emphasis>
Body slender and elongate (
<figureCitation id="13322A20FFAFFFF202402551FCFA586D" box="[794,889,575,599]" captionStart="Figure 2" captionStartId="10.[160,225,1576,1599]" captionTargetBox="[165,1152,145,1543]" captionTargetId="figure@10.[163,1153,142,1545]" captionTargetPageId="10" captionText="Figure 2. Morphology of Liopeltis pallidonuchalis sp. nov. holotype in life (ZMMU R-15682, male). (a) – general lateral view; (b) – head in lateral view; (c) – head in dorsal view; (d) – head in ventral view. Photos by Nikolay A. Poyarkov." figureDoi="http://doi.org/10.5281/zenodo.3670311" httpUri="https://zenodo.org/record/3670311/files/figure.png" pageId="11" pageNumber="1657">Figure 2</figureCitation>
(a))ı somewhat laterally compressed; tail thinı pointed; head smallı moderately distinct from the neck (mostly in dorsal view); snout blunt; nostrils rather largeı in dorsal-lateral positionı round in shape; eye rather largeı pupil circular.
</paragraph>
<paragraph id="8BB636A5FFAFFFF2019A25A4FB0058D8" blockId="11.[160,1156,573,1708]" box="[192,1155,714,738]" pageId="11" pageNumber="1657">
<emphasis id="B97DEAB7FFAFFFF2019A25A4FEA658D8" box="[192,293,714,738]" italics="true" pageId="11" pageNumber="1657">Body size</emphasis>
. HL: 
<quantity id="4CF19B40FFAFFFF2003A25A4FE4558D8" box="[352,454,714,738]" metricMagnitude="-2" metricUnit="m" metricValue="1.98" pageId="11" pageNumber="1657" unit="mm" value="19.8">19.8 mm</quantity>
; SVL: 
<quantity id="4CF19B40FFAFFFF2035325A4FDE958D8" box="[521,618,714,738]" metricMagnitude="-1" metricUnit="m" metricValue="2.9699999999999998" pageId="11" pageNumber="1657" unit="mm" value="297.0">297 mm</quantity>
; TaL: 
<quantity id="4CF19B40FFAFFFF203F125A4FC8F58D8" box="[683,780,714,738]" metricMagnitude="-1" metricUnit="m" metricValue="1.1199999999999999" pageId="11" pageNumber="1657" unit="mm" value="112.0">112 mm</quantity>
; TL: 
<quantity id="4CF19B40FFAFFFF2021A25A4FC2258D8" box="[832,929,714,738]" metricMagnitude="-1" metricUnit="m" metricValue="4.09" pageId="11" pageNumber="1657" unit="mm" value="409.0">409 mm</quantity>
; ratio TaL/TL: 0.274.
</paragraph>
<paragraph id="8BB636A5FFAFFFF2019A2582FBE25970" blockId="11.[160,1156,573,1708]" pageId="11" pageNumber="1657">
<emphasis id="B97DEAB7FFAFFFF2019A2582FEE2593E" box="[192,353,748,772]" italics="true" pageId="11" pageNumber="1657">Body scalation</emphasis>
. Dorsal scale rows 15-15-15ı all smooth; scales on dorsum and flanks somewhat rhomboid in shape; scales of the vertebral row not enlarged; no apical pit detected; 133 ventrals (+2 preventrals); 67 subcaudalsı all paired; anal plate divided.
</paragraph>
<paragraph id="8BB636A5FFAFFFF2019A243AFEB25F80" blockId="11.[160,1156,573,1708]" pageId="11" pageNumber="1657">
<emphasis id="B97DEAB7FFAFFFF2019A243AFEE95956" box="[192,362,852,876]" italics="true" pageId="11" pageNumber="1657">Head scalation</emphasis>
. Rostral heptagonalı wider than highı slightly visible from above (
<figureCitation id="13322A20FFAFFFF201F22419FE8A59B5" box="[168,265,887,911]" captionStart="Figure 2" captionStartId="10.[160,225,1576,1599]" captionTargetBox="[165,1152,145,1543]" captionTargetId="figure@10.[163,1153,142,1545]" captionTargetPageId="10" captionText="Figure 2. Morphology of Liopeltis pallidonuchalis sp. nov. holotype in life (ZMMU R-15682, male). (a) – general lateral view; (b) – head in lateral view; (c) – head in dorsal view; (d) – head in ventral view. Photos by Nikolay A. Poyarkov." figureDoi="http://doi.org/10.5281/zenodo.3670311" httpUri="https://zenodo.org/record/3670311/files/figure.png" pageId="11" pageNumber="1657">Figure 2</figureCitation>
(c)); nasal singleı elongated; nasal surrounded by the first two supralabialsı rostralı internasalı and prefrontal; internasals twoı curvedı wider than longerı in contact with rostral anteriorlyı nasalı and prefrontal; prefrontals twoı prefrontal length comprising less than half of frontal length; prefrontals in contact with internasalsı nasalsı preocularsı and frontal; frontal pentagonalı ca. 1.11 times wider than longı tapering posteriorlyı subequal to the distance from tip of snout to frontal; parietals longer than wideı in contact approximately the length of frontal (
<figureCitation id="13322A20FFAFFFF2025F2329FCE75E65" box="[773,868,1095,1119]" captionStart="Figure 2" captionStartId="10.[160,225,1576,1599]" captionTargetBox="[165,1152,145,1543]" captionTargetId="figure@10.[163,1153,142,1545]" captionTargetPageId="10" captionText="Figure 2. Morphology of Liopeltis pallidonuchalis sp. nov. holotype in life (ZMMU R-15682, male). (a) – general lateral view; (b) – head in lateral view; (c) – head in dorsal view; (d) – head in ventral view. Photos by Nikolay A. Poyarkov." figureDoi="http://doi.org/10.5281/zenodo.3670311" httpUri="https://zenodo.org/record/3670311/files/figure.png" pageId="11" pageNumber="1657">Figure 2</figureCitation>
(c))ı ca. 1.66 times longer than frontal; 1/1 supraocuları distinctly wider than highı in contact with prefrontal; loreal missing; 1/1 preocuları largeı pentagonalı higher than wideı in broad contact with prefrontal; subocular absent; 2/2 small postocularsı upper postocular larger than lower; 7/7 supralabialsı third and fourth entering orbitı sixth largest; 1 + 2/1 + 2 temporalsı anterior one very long and narrow (
<figureCitation id="13322A20FFAFFFF20366239AFD195F36" box="[572,666,1268,1292]" captionStart="Figure 2" captionStartId="10.[160,225,1576,1599]" captionTargetBox="[165,1152,145,1543]" captionTargetId="figure@10.[163,1153,142,1545]" captionTargetPageId="10" captionText="Figure 2. Morphology of Liopeltis pallidonuchalis sp. nov. holotype in life (ZMMU R-15682, male). (a) – general lateral view; (b) – head in lateral view; (c) – head in dorsal view; (d) – head in ventral view. Photos by Nikolay A. Poyarkov." figureDoi="http://doi.org/10.5281/zenodo.3670311" httpUri="https://zenodo.org/record/3670311/files/figure.png" pageId="11" pageNumber="1657">Figure 2</figureCitation>
(b))ı in broad contact with supralabials 5 
<emphasis id="B97DEAB7FFAFFFF20532239AFBF65F36" box="[1128,1141,1268,1292]" italics="true" pageId="11" pageNumber="1657">–</emphasis>
6 and parietalsı upper posterior temporal smaller than lower posteriorı rhomboid; infralabials 8/8 (
<figureCitation id="13322A20FFAFFFF2007A2254FEFE5F68" box="[288,381,1338,1362]" captionStart="Figure 2" captionStartId="10.[160,225,1576,1599]" captionTargetBox="[165,1152,145,1543]" captionTargetId="figure@10.[163,1153,142,1545]" captionTargetPageId="10" captionText="Figure 2. Morphology of Liopeltis pallidonuchalis sp. nov. holotype in life (ZMMU R-15682, male). (a) – general lateral view; (b) – head in lateral view; (c) – head in dorsal view; (d) – head in ventral view. Photos by Nikolay A. Poyarkov." figureDoi="http://doi.org/10.5281/zenodo.3670311" httpUri="https://zenodo.org/record/3670311/files/figure.png" pageId="11" pageNumber="1657">Figure 2</figureCitation>
(d))ı first pair in contact behind small mentalı 1 
<emphasis id="B97DEAB7FFAFFFF202DD2257FC175F6B" box="[903,916,1337,1361]" italics="true" pageId="11" pageNumber="1657">–</emphasis>
4/ 
<quantity id="4CF19B40FFAFFFF202EA2254FC735F68" box="[944,1008,1337,1362]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="11" pageNumber="1657" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">
1 
<emphasis id="B97DEAB7FFAFFFF202E32257FC455F6B" box="[953,966,1337,1361]" italics="true" pageId="11" pageNumber="1657">–</emphasis>
4 in
</quantity>
contact with anterior chin shieldsı fifth infralabial largestı apex directed posteriorly; posterior chin shields longer than anterior onesı separated from each other by a pair of small scales (
<figureCitation id="13322A20FFAFFFF201FD22CCFE855F80" box="[167,262,1442,1466]" captionStart="Figure 2" captionStartId="10.[160,225,1576,1599]" captionTargetBox="[165,1152,145,1543]" captionTargetId="figure@10.[163,1153,142,1545]" captionTargetPageId="10" captionText="Figure 2. Morphology of Liopeltis pallidonuchalis sp. nov. holotype in life (ZMMU R-15682, male). (a) – general lateral view; (b) – head in lateral view; (c) – head in dorsal view; (d) – head in ventral view. Photos by Nikolay A. Poyarkov." figureDoi="http://doi.org/10.5281/zenodo.3670311" httpUri="https://zenodo.org/record/3670311/files/figure.png" pageId="11" pageNumber="1657">Figure 2</figureCitation>
(d)).
</paragraph>
<paragraph id="8BB636A5FFAFFFF2019A22AAFDE15FC5" blockId="11.[160,1156,573,1708]" pageId="11" pageNumber="1657">
<emphasis id="B97DEAB7FFAFFFF2019A22AAFEA65FE6" box="[192,293,1476,1500]" italics="true" pageId="11" pageNumber="1657">Dentition</emphasis>
. Maxillary teeth smallı equalı 20 on left and 22 on the right side (counted directly prior to 
<typeStatus id="54B28807FFAFFFF200002289FE3D5FC5" box="[346,446,1511,1535]" pageId="11" pageNumber="1657" type="holotype">holotype</typeStatus>
preservation).
</paragraph>
<paragraph id="8BB636A5FFAFFFF5019A2164FF595C28" blockId="11.[160,1156,573,1708]" lastBlockId="12.[160,1156,1495,1727]" lastPageId="12" lastPageNumber="1658" pageId="11" pageNumber="1657">
<emphasis id="B97DEAB7FFAFFFF2019A2164FE3E5C18" box="[192,445,1546,1570]" italics="true" pageId="11" pageNumber="1657">Hemipenis morphology</emphasis>
. In everted positionı the organ is singleı not forkedı stoutı and reaches SC 5. Hemipenis is somewhat dumbbell-shapedı its proximal part covered with numerous tooth-shaped proximally oriented hooks (
<figureCitation id="13322A20FFAFFFF202552121FCF15C5D" box="[783,882,1615,1639]" captionStart="Figure 3" captionStartId="12.[160,225,570,593]" captionTargetBox="[163,1152,143,539]" captionTargetId="figure@12.[163,1153,142,539]" captionTargetPageId="12" captionText="Figure 3. Hemipenial morphology of Liopeltis pallidonuchalis sp. nov. holotype (ZMMU R-15682, male). (a) – asulcal side; (b) – sulcal side. Photos by Nikolay A. Poyarkov." figureDoi="http://doi.org/10.5281/zenodo.3670313" httpUri="https://zenodo.org/record/3670313/files/figure.png" pageId="11" pageNumber="1657">Figure 3</figureCitation>
)ı located in 3 
<emphasis id="B97DEAB7FFAFFFF205452121FBAF5C5D" box="[1055,1068,1615,1639]" italics="true" pageId="11" pageNumber="1657">–</emphasis>
5 rows; middle and distal parts of the organ covered with over 30 transverse skinfolds each bearing numerous papillae ending with small horny spinesı which increases in size distally; hemipenis tip with rare spines; sulcus spermaticus distinctı prominent (
<figureCitation id="13322A20FFA8FFF5056622B9FF2C5C28" captionStart="Figure 3" captionStartId="12.[160,225,570,593]" captionTargetBox="[163,1152,143,539]" captionTargetId="figure@12.[163,1153,142,539]" captionTargetPageId="12" captionText="Figure 3. Hemipenial morphology of Liopeltis pallidonuchalis sp. nov. holotype (ZMMU R-15682, male). (a) – asulcal side; (b) – sulcal side. Photos by Nikolay A. Poyarkov." figureDoi="http://doi.org/10.5281/zenodo.3670313" httpUri="https://zenodo.org/record/3670313/files/figure.png" pageId="12" pageNumber="1658">Figure 3</figureCitation>
(b)).
</paragraph>
<caption id="DF76662DFFA8FFF501FA2554FC025855" ID-DOI="http://doi.org/10.5281/zenodo.3670313" ID-Zenodo-Dep="3670313" httpUri="https://zenodo.org/record/3670313/files/figure.png" pageId="12" pageNumber="1658" startId="12.[160,225,570,593]" targetBox="[163,1152,143,539]" targetPageId="12">
<paragraph id="8BB636A5FFA8FFF501FA2554FC025855" blockId="12.[160,1157,570,623]" pageId="12" pageNumber="1658">
<emphasis id="B97DEAB7FFA8FFF501FA2554FF7D586B" bold="true" box="[160,254,570,593]" pageId="12" pageNumber="1658">Figure 3.</emphasis>
Hemipenial morphology of 
<taxonomicName id="4C094D26FFA8FFF5037F2554FC8D586B" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[549,782,570,593]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="1658" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFA8FFF5037F2554FC8D586B" box="[549,782,570,593]" italics="true" pageId="12" pageNumber="1658">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFA8FFF502422554FCED586B" box="[792,878,570,593]" pageId="12" pageNumber="1658" rank="species">
<emphasis id="B97DEAB7FFA8FFF502422554FCEA586B" bold="true" box="[792,873,570,593]" pageId="12" pageNumber="1658">sp. nov</emphasis>
.
</taxonomicNameLabel>
holotype (ZMMU R-15682, male). (a) 
<emphasis id="B97DEAB7FFA8FFF5005C2536FE905855" box="[262,275,600,623]" italics="true" pageId="12" pageNumber="1658">–</emphasis>
asulcal side; (b) 
<emphasis id="B97DEAB7FFA8FFF500E42536FE485855" box="[446,459,600,623]" italics="true" pageId="12" pageNumber="1658">–</emphasis>
sulcal side. Photos by Nikolay A. Poyarkov.
</paragraph>
</caption>
<caption id="DF76662DFFA8FFF501FA2234FEEE5FB5" ID-DOI="http://doi.org/10.5281/zenodo.3670315" ID-Zenodo-Dep="3670315" httpUri="https://zenodo.org/record/3670315/files/figure.png" pageId="12" pageNumber="1658" startId="12.[160,225,1370,1393]" targetBox="[163,1153,681,1338]" targetPageId="12">
<paragraph id="8BB636A5FFA8FFF501FA2234FEEE5FB5" blockId="12.[160,1155,1370,1423]" pageId="12" pageNumber="1658">
<emphasis id="B97DEAB7FFA8FFF501FA2234FF7C5F4B" bold="true" box="[160,255,1370,1393]" pageId="12" pageNumber="1658">Figure 4.</emphasis>
Holotype of 
<taxonomicName id="4C094D26FFA8FFF500D42234FDF45F4B" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[398,631,1370,1393]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="1658" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFA8FFF500D42234FDF45F4B" box="[398,631,1370,1393]" italics="true" pageId="12" pageNumber="1658">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFA8FFF503D82234FD565F4B" box="[642,725,1370,1393]" pageId="12" pageNumber="1658" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B97DEAB7FFA8FFF503852234FC9D5F4B" box="[735,798,1370,1393]" italics="true" pageId="12" pageNumber="1658">in situ</emphasis>
(ZMMU R-15682, male). Photo by Nikolay A. Poyarkov.
</paragraph>
</caption>
<paragraph id="8BB636A5FFA8FFF4019A2173FC685E58" blockId="12.[160,1156,1495,1727]" lastBlockId="13.[160,1156,751,1226]" lastPageId="13" lastPageNumber="1659" pageId="12" pageNumber="1658">
<emphasis id="B97DEAB7FFA8FFF5019A2173FE155C0F" box="[192,406,1565,1589]" italics="true" pageId="12" pageNumber="1658">Colouration (in life)</emphasis>
. In life body and flanks uniform bronze-brown (
<figureCitation id="13322A20FFA8FFF502982173FBA85C0F" box="[962,1067,1565,1589]" captionStart="Figure 2" captionStartId="10.[160,225,1576,1599]" captionTargetBox="[165,1152,145,1543]" captionTargetId="figure@10.[163,1153,142,1545]" captionTargetPageId="10" captionText="Figure 2. Morphology of Liopeltis pallidonuchalis sp. nov. holotype in life (ZMMU R-15682, male). (a) – general lateral view; (b) – head in lateral view; (c) – head in dorsal view; (d) – head in ventral view. Photos by Nikolay A. Poyarkov." figureDoi="http://doi.org/10.5281/zenodo.3670311" httpUri="https://zenodo.org/record/3670311/files/figure.png" pageId="12" pageNumber="1658">Figures 2</figureCitation>
and 
<figureCitation id="13322A20FFA8FFF505322173FBF55C0F" box="[1128,1142,1565,1589]" captionStart="Figure 4" captionStartId="12.[160,225,1370,1393]" captionTargetBox="[163,1153,681,1338]" captionTargetId="figure@12.[163,1153,681,1339]" captionTargetPageId="12" captionText="Figure 4. Holotype of Liopeltis pallidonuchalis sp. nov. in situ (ZMMU R-15682, male). Photo by Nikolay A. Poyarkov." figureDoi="http://doi.org/10.5281/zenodo.3670315" httpUri="https://zenodo.org/record/3670315/files/figure.png" pageId="12" pageNumber="1658">4</figureCitation>
)ı ventrally gradually getting lighter to light-beige background colour of lower flanks; head dorsally somewhat darker bronze-brownı snout beigeı supralabials and ventral surfaces of head immaculate cream-white; pupil blackı iris bicoloured: dorsally goldenı laterally and ventrally copper red-brown (
<figureCitation id="13322A20FFA8FFF5037E21C9FD065C85" box="[548,645,1703,1727]" captionStart="Figure 2" captionStartId="10.[160,225,1576,1599]" captionTargetBox="[165,1152,145,1543]" captionTargetId="figure@10.[163,1153,142,1545]" captionTargetPageId="10" captionText="Figure 2. Morphology of Liopeltis pallidonuchalis sp. nov. holotype in life (ZMMU R-15682, male). (a) – general lateral view; (b) – head in lateral view; (c) – head in dorsal view; (d) – head in ventral view. Photos by Nikolay A. Poyarkov." figureDoi="http://doi.org/10.5281/zenodo.3670311" httpUri="https://zenodo.org/record/3670311/files/figure.png" pageId="12" pageNumber="1658">Figure 2</figureCitation>
(b)); a narrow black postocular stripe of the same width as lower postocular running from the posterior corner of eye along lower postocuları lower margins of primary and lower secondary temporalsı dorsal margins of supralabials 6 
<emphasis id="B97DEAB7FFA9FFF40061245AFECB5976" box="[315,328,820,844]" italics="true" pageId="13" pageNumber="1659">–</emphasis>
7; postocular stripe gently curves dorsally at the level of head basis and becomes less distinct in nuchal area; continues as a pair of short dark grey markings with indistinct borders ca. 5 dorsal scales in length in the neck region (
<figureCitation id="13322A20FFA9FFF402C02414FC7B59A8" box="[922,1016,890,914]" captionStart="Figure 4" captionStartId="12.[160,225,1370,1393]" captionTargetBox="[163,1153,681,1338]" captionTargetId="figure@12.[163,1153,681,1339]" captionTargetPageId="12" captionText="Figure 4. Holotype of Liopeltis pallidonuchalis sp. nov. in situ (ZMMU R-15682, male). Photo by Nikolay A. Poyarkov." figureDoi="http://doi.org/10.5281/zenodo.3670315" httpUri="https://zenodo.org/record/3670315/files/figure.png" pageId="13" pageNumber="1659">Figure 4</figureCitation>
). Rare black spots of irregular shape located in the area of mouth angle (
<figureCitation id="13322A20FFA9FFF4022F24F2FC55598E" box="[885,982,924,948]" captionStart="Figure 2" captionStartId="10.[160,225,1576,1599]" captionTargetBox="[165,1152,145,1543]" captionTargetId="figure@10.[163,1153,142,1545]" captionTargetPageId="10" captionText="Figure 2. Morphology of Liopeltis pallidonuchalis sp. nov. holotype in life (ZMMU R-15682, male). (a) – general lateral view; (b) – head in lateral view; (c) – head in dorsal view; (d) – head in ventral view. Photos by Nikolay A. Poyarkov." figureDoi="http://doi.org/10.5281/zenodo.3670311" httpUri="https://zenodo.org/record/3670311/files/figure.png" pageId="13" pageNumber="1659">Figure 2</figureCitation>
(b))ı posteriorly spots getting elongated and form a narrow-interrupted dark line running along the border between ventrals and dorsal scales for the anterior one-third of body length (
<figureCitation id="13322A20FFA9FFF401FD236AFE935E26" box="[167,272,1028,1052]" captionStart="Figure 2" captionStartId="10.[160,225,1576,1599]" captionTargetBox="[165,1152,145,1543]" captionTargetId="figure@10.[163,1153,142,1545]" captionTargetPageId="10" captionText="Figure 2. Morphology of Liopeltis pallidonuchalis sp. nov. holotype in life (ZMMU R-15682, male). (a) – general lateral view; (b) – head in lateral view; (c) – head in dorsal view; (d) – head in ventral view. Photos by Nikolay A. Poyarkov." figureDoi="http://doi.org/10.5281/zenodo.3670311" httpUri="https://zenodo.org/record/3670311/files/figure.png" pageId="13" pageNumber="1659">Figures 2</figureCitation>
(a) and 4); a shorter and less distinct dark line running from the mouth angle along the border between the second and third rows of dorsal scales for approximately the distance of head length and quickly becoming indistinct (
<figureCitation id="13322A20FFA9FFF4023E2324FC425E58" box="[868,961,1098,1122]" captionStart="Figure 2" captionStartId="10.[160,225,1576,1599]" captionTargetBox="[165,1152,145,1543]" captionTargetId="figure@10.[163,1153,142,1545]" captionTargetPageId="10" captionText="Figure 2. Morphology of Liopeltis pallidonuchalis sp. nov. holotype in life (ZMMU R-15682, male). (a) – general lateral view; (b) – head in lateral view; (c) – head in dorsal view; (d) – head in ventral view. Photos by Nikolay A. Poyarkov." figureDoi="http://doi.org/10.5281/zenodo.3670311" httpUri="https://zenodo.org/record/3670311/files/figure.png" pageId="13" pageNumber="1659">Figure 2</figureCitation>
(a)).
</paragraph>
<caption id="DF76662DFFA9FFF401FA2534FEF35896" ID-DOI="http://doi.org/10.5281/zenodo.3670317" ID-Zenodo-Dep="3670317" httpUri="https://zenodo.org/record/3670317/files/figure.png" pageId="13" pageNumber="1659" startId="13.[160,225,602,625]" targetBox="[166,1153,143,569]" targetPageId="13">
<paragraph id="8BB636A5FFA9FFF401FA2534FEF35896" blockId="13.[160,1155,601,684]" pageId="13" pageNumber="1659">
<emphasis id="B97DEAB7FFA9FFF401FA2534FF7D584B" bold="true" box="[160,254,602,625]" pageId="13" pageNumber="1659">Figure 5.</emphasis>
Paratype of 
<taxonomicName id="4C094D26FFA9FFF400D32537FDF1584A" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[393,626,601,624]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="1659" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFA9FFF400D32537FDF1584A" box="[393,626,601,624]" italics="true" pageId="13" pageNumber="1659">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFA9FFF403262534FD4D584B" box="[636,718,602,625]" pageId="13" pageNumber="1659" rank="species">sp. nov.</taxonomicNameLabel>
(DTU 307, female, roadkill). (a) 
<emphasis id="B97DEAB7FFA9FFF405472534FBA9584B" box="[1053,1066,602,625]" italics="true" pageId="13" pageNumber="1659">–</emphasis>
head in dorsolateral view; (b) 
<emphasis id="B97DEAB7FFA9FFF400232519FE0558B4" box="[377,390,631,654]" italics="true" pageId="13" pageNumber="1659">–</emphasis>
head in lateral view; (c) 
<emphasis id="B97DEAB7FFA9FFF403252519FD0F58B4" box="[639,652,631,654]" italics="true" pageId="13" pageNumber="1659">–</emphasis>
the general view of the roadkill specimen. Photos by The Anh Nguyen.
</paragraph>
</caption>
<paragraph id="8BB636A5FFA9FFF4019A2302FC7B5EF0" blockId="13.[160,1156,751,1226]" pageId="13" pageNumber="1659">
<emphasis id="B97DEAB7FFA9FFF4019A2302FE7C5EBE" box="[192,511,1132,1156]" italics="true" pageId="13" pageNumber="1659">Colouration (in preservative)</emphasis>
. After 3 years in preservative the general colouration pattern did not change; orange tint in the colouration of dorsumı head and eye faded becoming greyish-brown; other features of colouration remain unchanged.
</paragraph>
<paragraph id="8BB636A5FFA9FFF401FA226BFC975C6D" blockId="13.[160,1156,1285,1727]" pageId="13" pageNumber="1659">
<emphasis id="B97DEAB7FFA9FFF401FA226BFE965F25" bold="true" box="[160,277,1285,1311]" italics="true" pageId="13" pageNumber="1659">Variation.</emphasis>
<materialsCitation id="3B613CF8FFA9FFF4007F2269FE865F78" ID-GBIF-Occurrence="2565890779" location="Variation" pageId="13" pageNumber="1659" specimenCount="1">
<location id="8ED6607EFFA9FFF4007F2269FE085F25" LSID="urn:lsid:plazi:treatment:03A087B3FFADFFEB009D2669FF6B5FF7:8ED6607EFFA9FFF4007F2269FE085F25" box="[293,395,1287,1311]" name="Variation" pageId="13" pageNumber="1659">Variation</location>
in measurements and scalation of the type series is presented in 
<tableCitation id="C68B031EFFA9FFF401FA2244FE865F78" box="[160,261,1322,1346]" captionStart="Table 2" captionStartId="9.[160,216,1145,1168]" captionTargetBox="[171,1145,1210,1717]" captionTargetPageId="9" captionText="Table 2. Main morphological variation in the type series and referred materials (ref.mat.) of Liopeltis pallidonuchalis sp. nov. See Materials and Methods for other abbreviations." httpUri="http://table.plazi.org/id/DF76662DFFADFFF001FA2317FC0B5E97" pageId="13" pageNumber="1659" tableUuid="DF76662DFFADFFF001FA2317FC0B5E97">Table 2</tableCitation>
</materialsCitation>
. 
<materialsCitation id="3B613CF8FFA9FFF400792244FE8A5F90" ID-GBIF-Occurrence="2565890771" collectorName="Thua Thien &amp; Hue Province" location="The" pageId="13" pageNumber="1659" specimenCount="2" stateProvince="Gia Lai" typeStatus="holotype">
<location id="8ED6607EFFA9FFF400792244FECE5F78" LSID="urn:lsid:plazi:treatment:03A087B3FFADFFEB009D2669FF6B5FF7:8ED6607EFFA9FFF400792244FECE5F78" box="[291,333,1322,1346]" name="The" pageId="13" pageNumber="1659" stateProvince="Gia Lai">The</location>
<typeStatus id="54B28807FFA9FFF4003C2244FE4F5F78" box="[358,460,1322,1346]" pageId="13" pageNumber="1659" type="holotype">holotype</typeStatus>
from 
<collectingRegion id="49CDF847FFA9FFF4036E2244FC8E5F78" box="[564,781,1322,1346]" country="Vietnam" name="Gia Lai" pageId="13" pageNumber="1659">Gia Lai Province</collectingRegion>
lacks loreal and prefrontal touches second supralabial (
<figureCitation id="13322A20FFA9FFF400AF2223FDDB5F5F" box="[501,600,1357,1381]" captionStart="Figure 2" captionStartId="10.[160,225,1576,1599]" captionTargetBox="[165,1152,145,1543]" captionTargetId="figure@10.[163,1153,142,1545]" captionTargetPageId="10" captionText="Figure 2. Morphology of Liopeltis pallidonuchalis sp. nov. holotype in life (ZMMU R-15682, male). (a) – general lateral view; (b) – head in lateral view; (c) – head in dorsal view; (d) – head in ventral view. Photos by Nikolay A. Poyarkov." figureDoi="http://doi.org/10.5281/zenodo.3670311" httpUri="https://zenodo.org/record/3670311/files/figure.png" pageId="13" pageNumber="1659">Figure 2</figureCitation>
(b))ı while in two 
<typeStatus id="54B28807FFA9FFF402692223FC205F5F" box="[819,931,1357,1381]" pageId="13" pageNumber="1659" type="paratype">paratypes</typeStatus>
from 
<collectorName id="26FC5373FFA9FFF402A82223FBF65F5F" box="[1010,1141,1357,1381]" pageId="13" pageNumber="1659">Thua Thien</collectorName>
<emphasis id="B97DEAB7FFA9FFF4052C2222FB005F5E" box="[1142,1155,1356,1380]" italics="true" pageId="13" pageNumber="1659">–</emphasis>
<collectorName id="26FC5373FFA9FFF401FA2201FEBE5FBD" box="[160,317,1391,1415]" pageId="13" pageNumber="1659">Hue Province</collectorName>
loreal is present and prefrontals are not in contact with supralabials (
<tableCitation id="C68B031EFFA9FFF401F222FCFE835F90" box="[168,256,1426,1450]" captionStart="Table 2" captionStartId="9.[160,216,1145,1168]" captionTargetBox="[171,1145,1210,1717]" captionTargetPageId="9" captionText="Table 2. Main morphological variation in the type series and referred materials (ref.mat.) of Liopeltis pallidonuchalis sp. nov. See Materials and Methods for other abbreviations." httpUri="http://table.plazi.org/id/DF76662DFFADFFF001FA2317FC0B5E97" pageId="13" pageNumber="1659" tableUuid="DF76662DFFADFFF001FA2317FC0B5E97">Table 2</tableCitation>
)
</materialsCitation>
. 
<materialsCitation id="3B613CF8FFA9FFF4004022FCFE0E5FD5" ID-GBIF-Occurrence="2565890772" collectionCode="ZFMK" location="The" pageId="13" pageNumber="1659" specimenCode="ZFMK 83105" specimenCount="3" stateProvince="Gia Lai" typeStatus="paratype">
<location id="8ED6607EFFA9FFF4004022FCFEC75F90" LSID="urn:lsid:plazi:treatment:03A087B3FFADFFEB009D2669FF6B5FF7:8ED6607EFFA9FFF4004022FCFEC75F90" box="[282,324,1426,1450]" name="The" pageId="13" pageNumber="1659" stateProvince="Gia Lai">The</location>
<typeStatus id="54B28807FFA9FFF4000A22FCFE365F90" box="[336,437,1426,1450]" pageId="13" pageNumber="1659" type="paratype">paratype</typeStatus>
DTU 307 has a single small scale separating posterior chin shieldsı while the 
<typeStatus id="54B28807FFA9FFF400DB22DBFE645FF7" box="[385,487,1461,1485]" pageId="13" pageNumber="1659" type="holotype">holotype</typeStatus>
and the 
<typeStatus id="54B28807FFA9FFF4033922DBFD4B5FF7" box="[611,712,1461,1485]" pageId="13" pageNumber="1659" type="paratype">paratype</typeStatus>
<specimenCode id="DBAF9EDEFFA9FFF4038C22DBFCE95FF7" box="[726,874,1461,1485]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="13" pageNumber="1659" type="Museum">ZFMK 83105</specimenCode>
have two small scales anterior to ventrals
</materialsCitation>
. 
<materialsCitation id="3B613CF8FFA9FFF400C522B9FC935C6D" ID-GBIF-Occurrence="2565890778" location="Colouration" pageId="13" pageNumber="1659" specimenCount="2" stateProvince="Gia Lai" typeStatus="paratype">
<location id="8ED6607EFFA9FFF400C522B9FDA65FD5" LSID="urn:lsid:plazi:treatment:03A087B3FFADFFEB009D2669FF6B5FF7:8ED6607EFFA9FFF400C522B9FDA65FD5" box="[415,549,1495,1519]" name="Colouration" pageId="13" pageNumber="1659" stateProvince="Gia Lai">Colouration</location>
of the 
<typeStatus id="54B28807FFA9FFF403D522B9FD775FD5" box="[655,756,1495,1519]" pageId="13" pageNumber="1659" type="paratype">paratype</typeStatus>
DTU 307 is very similar to the colouration oft he 
<typeStatus id="54B28807FFA9FFF400DC2294FE6F5C28" box="[390,492,1530,1554]" pageId="13" pageNumber="1659" type="holotype">holotype</typeStatus>
described above; but a series of 19 small dark-brown paravertebral blotches with indistinct borders is running from the neck region along with the anterior one-third of body length (
<figureCitation id="13322A20FFA9FFF403FD2151FC845C6D" box="[679,775,1599,1623]" captionStart="Figure 5" captionStartId="13.[160,225,602,625]" captionTargetBox="[166,1153,143,569]" captionTargetId="figure@13.[163,1153,142,570]" captionTargetPageId="13" captionText="Figure 5. Paratype of Liopeltis pallidonuchalis sp. nov. (DTU 307, female, roadkill). (a) – head in dorsolateral view; (b) – head in lateral view; (c) – the general view of the roadkill specimen. Photos by The Anh Nguyen." figureDoi="http://doi.org/10.5281/zenodo.3670317" httpUri="https://zenodo.org/record/3670317/files/figure.png" pageId="13" pageNumber="1659">Figure 5</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB636A5FFA9FFF7019A210CFCAC5B86" blockId="13.[160,1156,1285,1727]" lastBlockId="14.[160,1156,142,444]" lastPageId="14" lastPageNumber="1660" pageId="13" pageNumber="1659">
A single road-kill specimenı originally identified as 
<taxonomicName id="4C094D26FFA9FFF40252210CFCF15C40" baseAuthorityName="Guntheri" baseAuthorityYear="1858" box="[776,882,1634,1658]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="1659" phylum="Chordata" rank="species" species="frenata">
<emphasis id="B97DEAB7FFA9FFF40252210CFCF15C40" box="[776,882,1634,1658]" italics="true" pageId="13" pageNumber="1659">L. frenata</emphasis>
</taxonomicName>
ı was reported from the lowland karst forest of Phong Nha 
<emphasis id="B97DEAB7FFA9FFF4036421EAFDC85CA6" box="[574,587,1668,1692]" italics="true" pageId="13" pageNumber="1659">–</emphasis>
Ke Bang N.P. 
<materialsCitation id="3B613CF8FFA9FFF403AD21EBFD755C85" ID-GBIF-Occurrence="2565890782" country="Vietnam" elevation="150" location="Quang Binh Province" pageId="13" pageNumber="1659" specimenCount="1" stateProvince="Quang Binh">
in 
<collectingRegion id="49CDF847FFA9FFF4024D21EBFB8F5CA7" box="[791,1036,1669,1693]" country="Vietnam" name="Quang Binh" pageId="13" pageNumber="1659">Quang Binh Province</collectingRegion>
in central 
<collectingCountry id="F31E7635FFA9FFF401FA21C9FF7E5C85" box="[160,253,1703,1727]" name="Vietnam" pageId="13" pageNumber="1659">Vietnam</collectingCountry>
(
<elevation id="0024D196FFA9FFF4005121C9FE6A5C85" box="[267,489,1703,1727]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="1659" unit="m" value="150.0">
elevation 
<quantity id="4CF19B40FFA9FFF4002621C9FE415C85" box="[380,450,1703,1727]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="1659" unit="m" value="150.0">150 m</quantity>
asl
</elevation>
) by 
<bibRefCitation id="EF984B54FFA9FFF4034121C9FD755C85" author="Ziegler Ti Hendrix Ri Vu NTi Vogt Mi Forster Bi Dang NK" box="[539,758,1703,1727]" pageId="13" pageNumber="1659" pagination="1 - 40" refId="ref15010" refString="Ziegler Ti Hendrix Ri Vu NTi Vogt Mi Forster Bi Dang NK. 2007. The diversity of a snake community in a karst forest ecosystem in the central Truong Soni Vietnami with an identification key. Zootaxa. 1493: 1 - 40. doi: 10.11646 / zootaxa. 1493.1.1" type="journal article" year="2007">Ziegler et al. (2007)</bibRefCitation>
</materialsCitation>
. 
<materialsCitation id="3B613CF8FFA9FFF7025B21C9FB8A5A9C" ID-GBIF-Occurrence="2565890774" country="Vietnam" lastPageId="14" lastPageNumber="1660" location="In" pageId="13" pageNumber="1659" specimenCount="1" stateProvince="Quang Binh">
<location id="8ED6607EFFA9FFF4025B21C9FC955C85" LSID="urn:lsid:plazi:treatment:03A087B3FFADFFEB009D2669FF6B5FF7:8ED6607EFFA9FFF4025B21C9FC955C85" box="[769,790,1703,1727]" country="Vietnam" name="In" pageId="13" pageNumber="1659" stateProvince="Quang Binh">In</location>
generalı morphological data for this specimen agrees well with the diagnosis of 
<taxonomicName id="4C094D26FFAAFFF7038027E0FC615A9C" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[730,994,142,166]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="1660" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFAAFFF7038027E0FC615A9C" box="[730,994,142,166]" italics="true" pageId="14" pageNumber="1660">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFAAFFF702B727E0FB8A5A9C" box="[1005,1033,142,166]" pageId="14" pageNumber="1660" rank="species">sp</taxonomicNameLabel>
</materialsCitation>
. nov. 
<materialsCitation id="3B613CF8FFAAFFF7050827E0FB8C5B0B" ID-GBIF-Occurrence="2565890776" country="Vietnam" latitude="0.274" location="Rel" longLatPrecision="78" longitude="-0.288" pageId="14" pageNumber="1660" specimenCount="3" stateProvince="Quang Binh">
with exception of the following differences (see 
<tableCitation id="C68B031EFFAAFFF703C727DFFD705AF3" box="[669,755,177,201]" captionStart="Table 2" captionStartId="9.[160,216,1145,1168]" captionTargetBox="[171,1145,1210,1717]" captionTargetPageId="9" captionText="Table 2. Main morphological variation in the type series and referred materials (ref.mat.) of Liopeltis pallidonuchalis sp. nov. See Materials and Methods for other abbreviations." httpUri="http://table.plazi.org/id/DF76662DFFADFFF001FA2317FC0B5E97" pageId="14" pageNumber="1660" tableUuid="DF76662DFFADFFF001FA2317FC0B5E97">Table 2</tableCitation>
): comparatively longer tail (
<location id="8ED6607EFFAAFFF7056127DFFBDC5AF3" LSID="urn:lsid:plazi:treatment:03A087B3FFADFFEB009D2669FF6B5FF7:8ED6607EFFAAFFF7056127DFFBDC5AF3" box="[1083,1119,177,201]" country="Vietnam" latitude="0.274" longLatPrecision="78" longitude="-0.288" name="Rel" pageId="14" pageNumber="1660" stateProvince="Quang Binh">Rel</location>
TL 0.301 vs. 
<quantity id="4CF19B40FFAAFFF7005627BAFE335AD6" box="[268,432,211,236]" metricMagnitude="-3" metricUnit="m" metricValue="7.1374" metricValueMax="7.315199999999999" metricValueMin="6.9596" pageId="14" pageNumber="1660" unit="in" value="0.281" valueMax="0.288" valueMin="0.274">
<geoCoordinate id="EE3D5062FFAAFFF7005627BAFECB5AD6" box="[268,328,212,236]" degrees="0.274" direction="north" orientation="latitude" pageId="14" pageNumber="1660" precision="55" value="0.274">0.274</geoCoordinate>
<geoCoordinate id="EE3D5062FFAAFFF7001227BDFE115AD6" box="[328,402,211,236]" degrees="0.288" direction="west" orientation="longitude" pageId="14" pageNumber="1660" precision="55" value="-0.288">
<emphasis id="B97DEAB7FFAAFFF7001227BDFED65AD1" box="[328,341,211,235]" italics="true" pageId="14" pageNumber="1660">–</emphasis>
0.288
</geoCoordinate>
in
</quantity>
the 
<typeStatus id="54B28807FFAAFFF700B227BAFD995AD6" box="[488,538,212,236]" pageId="14" pageNumber="1660">type</typeStatus>
series of the new species)ı four small scales between chin shields anterior to ventrals (vs. 1 
<emphasis id="B97DEAB7FFAAFFF7030F2798FDE15B34" box="[597,610,246,270]" italics="true" pageId="14" pageNumber="1660">–</emphasis>
2 scales in the 
<typeStatus id="54B28807FFAAFFF7024F2798FCC45B34" box="[789,839,246,270]" pageId="14" pageNumber="1660">type</typeStatus>
series of the new species); seven infralabials (vs. eight infralabials in the 
<typeStatus id="54B28807FFAAFFF703F02677FD5F5B0B" box="[682,732,281,305]" pageId="14" pageNumber="1660">type</typeStatus>
series of the new species)
</materialsCitation>
. 
<materialsCitation id="3B613CF8FFAAFFF705412677FD705BA3" ID-GBIF-Occurrence="2565890775" country="Vietnam" location="Taking" pageId="14" pageNumber="1660" specimenCount="1" stateProvince="Quang Binh">
<location id="8ED6607EFFAAFFF705412677FBE55B0B" LSID="urn:lsid:plazi:treatment:03A087B3FFADFFEB009D2669FF6B5FF7:8ED6607EFFAAFFF705412677FBE55B0B" box="[1051,1126,281,305]" country="Vietnam" name="Taking" pageId="14" pageNumber="1660" stateProvince="Quang Binh">Taking</location>
in the account morphological (see above) and ecological (lowland karst forest vs. montane evergreen forest) differences between these two populationsı we tentatively assign the 
<collectingRegion id="49CDF847FFAAFFF701FA26EFFEA65BA3" box="[160,293,385,409]" country="Vietnam" name="Quang Binh" pageId="14" pageNumber="1660">Quang Binh</collectingRegion>
population to 
<taxonomicName id="4C094D26FFAAFFF7009726EFFD535BA3" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[461,720,385,409]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="1660" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFAAFFF7009726EFFD535BA3" box="[461,720,385,409]" italics="true" pageId="14" pageNumber="1660">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFAAFFF7038D26EFFD705BA3" box="[727,755,385,409]" pageId="14" pageNumber="1660" rank="species">sp</taxonomicNameLabel>
</materialsCitation>
. nov. as referred materialsı pending examination of additional specimens and molecular data.
</paragraph>
</subSubSection>
<subSubSection id="C313652EFFAAFFF701FA2689FE4358EB" pageId="14" pageNumber="1660" type="etymology">
<paragraph id="8BB636A5FFAAFFF701FA2689FC635853" blockId="14.[160,1155,487,721]" pageId="14" pageNumber="1660">
<emphasis id="B97DEAB7FFAAFFF701FA2689FEA0583B" bold="true" box="[160,291,487,513]" italics="true" pageId="14" pageNumber="1660">Etymology.</emphasis>
The specific name 
<emphasis id="B97DEAB7FFAAFFF7035E2687FD37583B" box="[516,692,489,513]" italics="true" pageId="14" pageNumber="1660">‘pallidonuchalis’</emphasis>
is a Latinised adjective in the nominative singular (feminine gender)ı derived from Latin 
<emphasis id="B97DEAB7FFAAFFF703922562FCAB581E" box="[712,808,524,548]" italics="true" pageId="14" pageNumber="1660">‘pallidus’</emphasis>
for 
<emphasis id="B97DEAB7FFAAFFF702042562FCE7581E" box="[862,868,524,548]" italics="true" pageId="14" pageNumber="1660">‘</emphasis>
pale 
<emphasis id="B97DEAB7FFAAFFF702C92562FC1A581E" box="[915,921,524,548]" italics="true" pageId="14" pageNumber="1660">’</emphasis>
and Medieval Latin 
<emphasis id="B97DEAB7FFAAFFF701FA2540FF6D587C" box="[160,238,558,582]" italics="true" pageId="14" pageNumber="1660">‘nucha’</emphasis>
ı derived from Arabic 
<emphasis id="B97DEAB7FFAAFFF700B52540FDB2587C" box="[495,561,558,582]" italics="true" pageId="14" pageNumber="1660">‘nuka’</emphasis>
for 
<emphasis id="B97DEAB7FFAAFFF7033F2540FDE8587C" box="[613,619,558,582]" italics="true" pageId="14" pageNumber="1660">‘</emphasis>
nape 
<emphasis id="B97DEAB7FFAAFFF703F82540FD24587C" box="[674,679,558,582]" italics="true" pageId="14" pageNumber="1660">’</emphasis>
ı 
<emphasis id="B97DEAB7FFAAFFF703E22540FD3E587C" box="[696,701,558,582]" italics="true" pageId="14" pageNumber="1660">‘</emphasis>
dorsal surface of neck 
<emphasis id="B97DEAB7FFAAFFF702E62540FC42587C" box="[956,961,558,582]" italics="true" pageId="14" pageNumber="1660">’</emphasis>
ı referring to the pale postocular marking of the new species disappearing in nuchal area.
</paragraph>
<paragraph id="8BB636A5FFAAFFF7019A251AFE4358EB" blockId="14.[160,1155,487,721]" pageId="14" pageNumber="1660">
We suggest the following common names: 
<emphasis id="B97DEAB7FFAAFFF7039B251AFC3358B6" box="[705,944,628,652]" italics="true" pageId="14" pageNumber="1660">Pale-necked Ringneck</emphasis>
(English)ı 
<emphasis id="B97DEAB7FFAAFFF70576251AFEDA5894" italics="true" pageId="14" pageNumber="1660">Rắn đai gáy nhạt màu</emphasis>
(Vietnamese)ı 
<emphasis id="B97DEAB7FFAAFFF7034425F8FC2F5894" box="[542,940,662,686]" italics="true" pageId="14" pageNumber="1660">Blednyi Gladkiy Kamyshovyi Uzh</emphasis>
(Russian)ı 
<emphasis id="B97DEAB7FFAAFFF7051B25F8FECB58EB" italics="true" pageId="14" pageNumber="1660">Blasse Halsbandnatter</emphasis>
(German).
</paragraph>
</subSubSection>
<subSubSection id="C313652EFFAAFFF701FA2593FB005FBD" pageId="14" pageNumber="1660" type="distribution">
<paragraph id="8BB636A5FFAAFFF701FA2593FBDF5E75" blockId="14.[160,1157,765,1103]" pageId="14" pageNumber="1660">
<emphasis id="B97DEAB7FFAAFFF701FA2593FEB0592D" bold="true" box="[160,307,765,791]" italics="true" pageId="14" pageNumber="1660">Distribution.</emphasis>
The distribution of the new species is shown in 
<figureCitation id="13322A20FFAAFFF7023F2591FC40592D" box="[869,963,767,791]" captionStart="Figure 1" captionStartId="6.[160,225,1017,1040]" captionTargetBox="[163,1153,142,985]" captionTargetId="figure@6.[163,1153,142,985]" captionTargetPageId="6" captionText="Figure 1. Known distribution of Liopeltis frenata and related taxa. Star denotes type locality. Question mark denotes and unconfirmed record of Liopeltis cf. frenata from Bhutan. Locality information. Liopeltis pallidonuchalis sp. nov.: 1 – Kon Chu Rang N.R., Gia Lai,Vietnam (type locality of Liopeltis pallidonuchalis sp. nov.; this work); 2 – Ba Na–Nui Chua N.P., Da Nang, Vietnam (as ‘Tourane’, Smith 1943); 3 – Do Quyen, Bach Ma N.P., Thua Thien–Hue, Vietnam (this work); 4 – U Bo, Phong Nha – Ke Bang N.P., Quang Binh, Vietnam (VNUH 9.7.’06–1; Ziegler et al. 2007; Luu et al. 2013); Liopeltis frenata s.str.: 5 – Khasi Hills, Meghalaya,India (restricted type locality of Cyclophis frenatus Günther,1858; Boulenger 1894; Wall 1908); 6 – Medog, Xizang, China (Zhao and Adler 1993; Zhao 2006); 7 – Mouling N.P., Arunachal Pradesh, India (Pawar and Birand 2001); 8 – Naga Hills, Nagaland, India (Ao et al. 2004); 9 – Namdapha N.P., Arunachal Pradesh, India (Pawar and Birand 2001; Krishna et al. 2012); 10 – Putao, Kachin, Myanmar (CAS 225548- 49); 11 – Htingnan, Kachin, Myanmar (Smith 1940); 12 – Myitkyina, Kachin, Myanmar (AMNH R-58541–44); 13 – Huton, Kachin Hills, Kachin, Myanmar (Wall 1925); 14 – Bhamo, Kachin, Myanmar (Smith 1943); 15 – Mansi, Kachin Hills, Kachin, Myanmar (Wall 1925); 16 – Wuliangshan Mt., Yunnan, China (this paper); 17 – Lüchun, Yunnan, China (KIZ79001; Yang and Rao 2008); 18 – Nam Xe, Van Ban, Lao Cai, Vietnam (IEBR 1205; Orlov et al. 2003); 19 – Copia N.R., Thuan Chau, Son La, Vietnam (TBU PAR.109, 116; Pham et al. 2014); 20 – Houaphan, Laos (Teynié et al. 2014); 21 – Xiangkhong, Laos (as ‘Tran-ninh’ in Smith 1943; Deuve 1970); 22 – Bo Kluea,Nan, Thailand (Hauser 2018); Liopeltis cf. stoliczkae (confirmed localities):23 – Samagooting,Nagaland,India (type locality of Ablabes stoliczkae Sclater,1891; ZSI 3955; see Das et al. 1998); 24 – Darjeeling, West Bengal, India (Smith 1943); 25 – Sikkim, India (no exact locality; Smith 1943); 26 – Karin Hills, Kayah, Myanmar (no exact locality; Smith 1943); Liopeltis cf. stoliczkae (unconfirmed localities): 27 – B La, Bao Lam, Lam Dong, Vietnam (IEBR 1293, as Liopeltis cf. tricolor; Orlov et al. 2003); 28 – Pichrada,Modolkiri,Cambodia (FMNH 259193; Stuart et al. 2006); 29 – Lac Xao, Khamkeut, Bolikhamxay, Laos (FMNH 254780; Stuart and Heatwole 2008); Doubtful localities of Liopeltis spp.in Indochina:30 – Lai Chau,Vietnam (mentioned by Hauser 2018 as Liopeltis frenata, but not" figureDoi="http://doi.org/10.5281/zenodo.3670309" httpUri="https://zenodo.org/record/3670309/files/figure.png" pageId="14" pageNumber="1660">Figure 1</figureCitation>
. 
<taxonomicName id="4C094D26FFAAFFF7028A2591FF795903" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="1660" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFAAFFF7028A2591FF795903" italics="true" pageId="14" pageNumber="1660">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFAAFFF7005F244FFEE35903" box="[261,352,801,825]" pageId="14" pageNumber="1660" rank="species">sp. nov.</taxonomicNameLabel>
is presently known from montane evergreen tropical forests of 
<collectingRegion id="49CDF847FFAAFFF7050D244FFF515966" country="Vietnam" name="Kon Tum" pageId="14" pageNumber="1660">Kon Tum</collectingRegion>
<emphasis id="B97DEAB7FFAAFFF70188242AFF5C5966" box="[210,223,836,860]" italics="true" pageId="14" pageNumber="1660">–</emphasis>
<collectingRegion id="49CDF847FFAAFFF70185242AFEB35966" box="[223,304,836,860]" country="Vietnam" name="Gia Lai" pageId="14" pageNumber="1660">Gia Lai</collectingRegion>
Plateau of Truong Son (also known as Annamite) Mountains in Central 
<collectingCountry id="F31E7635FFAAFFF701FA2409FE835945" box="[160,256,871,895]" name="Vietnam" pageId="14" pageNumber="1660">Vietnam</collectingCountry>
. It is relaibly recorded from Kon Chu Rang N.R. in 
<collectingRegion id="49CDF847FFAAFFF702112409FB875945" box="[843,1028,871,895]" country="Vietnam" name="Gia Lai" pageId="14" pageNumber="1660">Gia Lai Province</collectingRegion>
(our data)ı from Bach Ma N.P. in Thua Thien 
<emphasis id="B97DEAB7FFAAFFF7034F24E7FDA1599B" box="[533,546,905,929]" italics="true" pageId="14" pageNumber="1660">–</emphasis>
Hue Province (our data; 
<bibRefCitation id="EF984B54FFAAFFF7026C24E7FBCA599B" author="Ziegler Ti Hoang QX" box="[822,1097,905,929]" pageId="14" pageNumber="1660" pagination="115" refId="ref15062" refString="Ziegler Ti Hoang QX. 2009. Liopeltis frenatus. Herpetol Rev. 40 (1): 115." type="journal article" year="2009">Ziegler and Hoang 2009</bibRefCitation>
) and from Ba Na 
<emphasis id="B97DEAB7FFAAFFF7004724C2FEA959FE" box="[285,298,940,964]" italics="true" pageId="14" pageNumber="1660">–</emphasis>
Nui Chua N.R. in 
<collectingRegion id="49CDF847FFAAFFF700B224C2FDCA59FE" box="[488,585,940,964]" country="Vietnam" name="Da Nang" pageId="14" pageNumber="1660">Da Nang</collectingRegion>
City (historical record from 
<emphasis id="B97DEAB7FFAAFFF702D924C2FC0B59FE" box="[899,904,940,964]" italics="true" pageId="14" pageNumber="1660">‘</emphasis>
Tourane 
<emphasis id="B97DEAB7FFAAFFF702BF24C2FC6859FE" box="[997,1003,940,964]" italics="true" pageId="14" pageNumber="1660">’</emphasis>
ı 
<bibRefCitation id="EF984B54FFAAFFF702AD24C2FBF559FE" author="Smith MA" box="[1015,1142,940,964]" pageId="14" pageNumber="1660" pagination="583" refId="ref14375" refString="Smith MA. 1943. The Fauna of British Indiai Ceylon and Burmai including the whole of the Indo- Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). London: Taylor and Francis; p. 583." type="book chapter" year="1943">Smith 1943</bibRefCitation>
); where it was found at elevations of 
<quantity id="4CF19B40FFAAFFF7030B24A1FD6359DD" box="[593,736,975,999]" metricMagnitude="2" metricUnit="m" metricValue="9.8" metricValueMax="10.1" metricValueMin="9.5" pageId="14" pageNumber="1660" unit="m" value="980.0" valueMax="1010.0" valueMin="950.0">
950 
<emphasis id="B97DEAB7FFAAFFF7032024A1FD0459DD" box="[634,647,975,999]" italics="true" pageId="14" pageNumber="1660">–</emphasis>
1010 m
</quantity>
asl. The new species probably also occurs in karst forests of Phong Nha 
<emphasis id="B97DEAB7FFAAFFF70319249FFDD35E33" box="[579,592,1009,1033]" italics="true" pageId="14" pageNumber="1660">–</emphasis>
Ke Bang N.P.ı 
<collectingRegion id="49CDF847FFAAFFF703A8249FFC605E33" box="[754,995,1009,1033]" country="Vietnam" name="Quang Binh" pageId="14" pageNumber="1660">Quang Binh Province</collectingRegion>
(
<bibRefCitation id="EF984B54FFAAFFF702A8249FFF545E16" author="Ziegler Ti Hendrix Ri Vu NTi Vogt Mi Forster Bi Dang NK" pageId="14" pageNumber="1660" pagination="1 - 40" refId="ref15010" refString="Ziegler Ti Hendrix Ri Vu NTi Vogt Mi Forster Bi Dang NK. 2007. The diversity of a snake community in a karst forest ecosystem in the central Truong Soni Vietnami with an identification key. Zootaxa. 1493: 1 - 40. doi: 10.11646 / zootaxa. 1493.1.1" type="journal article" year="2007">Ziegler et al. 2007</bibRefCitation>
; see Variation) where it occurs at elevations 
<quantity id="4CF19B40FFAAFFF70258237AFCCC5E16" box="[770,847,1044,1068]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="1660" unit="m" value="150.0">150 m</quantity>
aslı however this locality requires further verification. In particuları records from eastern 
<collectingCountry id="F31E7635FFAAFFF7022A2359FC205E75" box="[880,931,1079,1103]" name="Laos" pageId="14" pageNumber="1660">Laos</collectingCountry>
are anticipated.
</paragraph>
<paragraph id="8BB636A5FFAAFFF701FA2314FB005FBD" blockId="14.[160,1155,1146,1415]" pageId="14" pageNumber="1660">
<emphasis id="B97DEAB7FFAAFFF701FA2314FE175EAE" bold="true" box="[160,404,1146,1172]" italics="true" pageId="14" pageNumber="1660">Conservation status.</emphasis>
<taxonomicName id="4C094D26FFAAFFF700FE2312FD295EAE" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[420,682,1148,1172]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="1660" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFAAFFF700FE2312FD295EAE" box="[420,682,1148,1172]" italics="true" pageId="14" pageNumber="1660">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFAAFFF703E92312FC8F5EAE" box="[691,780,1148,1172]" pageId="14" pageNumber="1660" rank="species">sp. nov.</taxonomicNameLabel>
is to date reliably recorded only from two national parks and one nature reserve in central 
<collectingCountry id="F31E7635FFAAFFF7021223F1FC265E8D" box="[840,933,1183,1207]" name="Vietnam" pageId="14" pageNumber="1660">Vietnam</collectingCountry>
and is known only from four specimens. This is a highly secretive snake which appears to be a strict forest specialistı associated with primary undisturbed montane forests. It appears that the new species may be affected by growing anthropogenic pressure and forest destructionı as observed in different areas of central 
<collectingCountry id="F31E7635FFAAFFF703312244FD485F78" box="[619,715,1322,1346]" name="Vietnam" pageId="14" pageNumber="1660">Vietnam</collectingCountry>
. Given the available informationı we suggest 
<taxonomicName id="4C094D26FFAAFFF7005E2222FD8E5F5E" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[260,525,1356,1380]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="1660" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFAAFFF7005E2222FD8E5F5E" box="[260,525,1356,1380]" italics="true" pageId="14" pageNumber="1660">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFAAFFF703402222FDF45F5E" box="[538,631,1356,1380]" pageId="14" pageNumber="1660" rank="species">sp. nov.</taxonomicNameLabel>
be considered as a Vulnerable (VU) species following IUCN 
<emphasis id="B97DEAB7FFAAFFF700132201FECD5FBD" box="[329,334,1391,1415]" italics="true" pageId="14" pageNumber="1660">’</emphasis>
s Red List categories (
<bibRefCitation id="EF984B54FFAAFFF7031A2201FBF55FBD" author="IUCN Standards &amp; Petitions Subcommittee" box="[576,1142,1391,1415]" pageId="14" pageNumber="1660" refId="ref13075" refString="IUCN Standards and Petitions Subcommittee. 2016. Guidelines for using the IUCN red list categories and criteria. Version 12. Prepared by the standards and petitions subcommittee. accessed 2018 Aug 15]. http: // www. iucnredlist. org / documents / RedListGuidelines. pdf." type="journal volume" year="2016">IUCN Standards and Petitions Subcommittee 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C313652EFFAAFFEB01FA22DDFF6B5FF7" lastPageId="18" lastPageNumber="1664" pageId="14" pageNumber="1660" type="biology_ecology">
<paragraph id="8BB636A5FFAAFFF601FA22DDFB955B34" blockId="14.[160,1155,1459,1727]" lastBlockId="15.[160,1157,142,825]" lastPageId="15" lastPageNumber="1661" pageId="14" pageNumber="1660">
<emphasis id="B97DEAB7FFAAFFF701FA22DDFF7D5FF7" bold="true" box="[160,254,1459,1485]" italics="true" pageId="14" pageNumber="1660">Biology.</emphasis>
Our knowledge of the biology of 
<taxonomicName id="4C094D26FFAAFFF703D322DBFC0E5FF7" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[649,909,1461,1485]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="1660" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFAAFFF703D322DBFC0E5FF7" box="[649,909,1461,1485]" italics="true" pageId="14" pageNumber="1660">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFAAFFF702CF22DBFC6F5FF7" box="[917,1004,1461,1485]" pageId="14" pageNumber="1660" rank="species">sp. nov.</taxonomicNameLabel>
is scarce; the species appears to be closely associated with montane evergreen forests covering the 
<collectingRegion id="49CDF847FFAAFFF701FA2294FE855C28" box="[160,262,1530,1554]" country="Vietnam" name="Kon Tum" pageId="14" pageNumber="1660">Kon Tum</collectingRegion>
<emphasis id="B97DEAB7FFAAFFF7005C2294FE905C28" box="[262,275,1530,1554]" italics="true" pageId="14" pageNumber="1660">–</emphasis>
<collectingRegion id="49CDF847FFAAFFF7004E2294FEE35C28" box="[276,352,1530,1554]" country="Vietnam" name="Gia Lai" pageId="14" pageNumber="1660">Gia Lai</collectingRegion>
Plateau of Truong Son Mountains where it was recorded at elevations of 
<quantity id="4CF19B40FFAAFFF701E42173FECB5C0F" box="[190,328,1564,1589]" metricMagnitude="2" metricUnit="m" metricValue="9.8" metricValueMax="10.1" metricValueMin="9.5" pageId="14" pageNumber="1660" unit="m" value="980.0" valueMax="1010.0" valueMin="950.0">
950 
<emphasis id="B97DEAB7FFAAFFF701BD2172FF775C0E" box="[231,244,1564,1588]" italics="true" pageId="14" pageNumber="1660">–</emphasis>
1010 m
</quantity>
asl. It also possibly occurs in lowland karst tropical forests of 
<collectingRegion id="49CDF847FFAAFFF702A42173FB005C0F" box="[1022,1155,1565,1589]" country="Vietnam" name="Quang Binh" pageId="14" pageNumber="1660">Quang Binh</collectingRegion>
karst area (see Variation) where the species was recorded from 
<quantity id="4CF19B40FFAAFFF7022A2151FC345C6D" box="[880,951,1599,1623]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="1660" unit="m" value="150.0">150 m</quantity>
asl. Animals were recorded only in patches of primary undisturbed forest with complete multi-layered canopy and heavy undergrowthı suggesting the new species is a strict forest-dwelling specialist. At the 
<typeStatus id="54B28807FFAAFFF7003E21C9FE155C85" box="[356,406,1703,1727]" pageId="14" pageNumber="1660">type</typeStatus>
locality in Kon Chu Rang N.R. (
<collectingRegion id="49CDF847FFAAFFF703A321C9FC345C85" box="[761,951,1703,1727]" country="Vietnam" name="Gia Lai" pageId="14" pageNumber="1660">Gia Lai Province</collectingRegion>
)ı the forest where the new species was recorded is dominated by large trees of the families 
<taxonomicName id="4C094D26FFABFFF6028227E0FB075A9C" box="[984,1156,142,166]" class="Pinopsida" family="Podocarpaceae" kingdom="Plantae" order="Pinales" pageId="15" pageNumber="1661" phylum="Tracheophyta" rank="family">Podocarpaceae</taxonomicName>
(
<emphasis id="B97DEAB7FFABFFF601FD27DFFD045AF3" box="[167,647,177,201]" italics="true" pageId="15" pageNumber="1661">
<taxonomicName id="4C094D26FFABFFF601FD27DFFEF65AF3" authorityName="Wall. ex Loudon" authorityYear="1838" box="[167,373,177,201]" class="Bivalvia" family="Mytilidae" genus="Dacrydium" kingdom="Animalia" order="Mytiloida" pageId="15" pageNumber="1661" phylum="Mollusca" rank="species" species="elatum">Dacrydium elatum</taxonomicName>
, 
<taxonomicName id="4C094D26FFABFFF600DE27DFFD045AF3" authorityName="de Laub" authorityYear="1969" baseAuthorityName="Blume" box="[388,647,177,201]" class="Pinopsida" family="Podocarpaceae" genus="Dacrycarpus" kingdom="Plantae" order="Pinales" pageId="15" pageNumber="1661" phylum="Tracheophyta" rank="species" species="imbricatus">Dacrycarpus imbricatus</taxonomicName>
</emphasis>
)ı Magnoliaceaeı 
<taxonomicName id="4C094D26FFABFFF6021327DFFC535AF3" box="[841,976,177,201]" class="Magnoliopsida" family="Burseraceae" kingdom="Plantae" order="Sapindales" pageId="15" pageNumber="1661" phylum="Tracheophyta" rank="family">Burseraceae</taxonomicName>
(
<taxonomicName id="4C094D26FFABFFF602B827DFFBCE5AF3" box="[994,1101,177,201]" class="Gastropoda" family="Strombidae" genus="Canarium" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="1661" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAB7FFABFFF602B827DFFBCE5AF3" box="[994,1101,177,201]" italics="true" pageId="15" pageNumber="1661">Canarium</emphasis>
</taxonomicName>
sp.)ı 
<taxonomicName id="4C094D26FFABFFF601FA27BAFE975AD6" box="[160,276,212,236]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="15" pageNumber="1661" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
(
<taxonomicName id="4C094D26FFABFFF6007127BAFE115AD6" box="[299,402,212,236]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" order="Myrtales" pageId="15" pageNumber="1661" phylum="Tracheophyta" rank="genus">
<emphasis id="B97DEAB7FFABFFF6007127BAFE115AD6" box="[299,402,212,236]" italics="true" pageId="15" pageNumber="1661">Syzygium</emphasis>
</taxonomicName>
sp.)ı 
<taxonomicName id="4C094D26FFABFFF6008627BAFD1E5AD6" box="[476,669,212,236]" class="Magnoliopsida" family="Hamamelidaceae" kingdom="Plantae" order="Saxifragales" pageId="15" pageNumber="1661" phylum="Tracheophyta" rank="family">Hamamelidaceae</taxonomicName>
(
<emphasis id="B97DEAB7FFABFFF603EE27BAFCB45AD6" box="[692,823,212,236]" italics="true" pageId="15" pageNumber="1661">Simingtonia</emphasis>
sp.)ı 
<taxonomicName id="4C094D26FFABFFF602DB27BAFC715AD6" box="[897,1010,212,236]" class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" order="Laurales" pageId="15" pageNumber="1661" phylum="Tracheophyta" rank="family">Lauraceae</taxonomicName>
(
<taxonomicName id="4C094D26FFABFFF6055327BAFBC45AD6" box="[1033,1095,212,236]" class="Magnoliopsida" family="Lauraceae" genus="Litsea" kingdom="Plantae" order="Laurales" pageId="15" pageNumber="1661" phylum="Tracheophyta" rank="genus">
<emphasis id="B97DEAB7FFABFFF6055327BAFBC45AD6" box="[1033,1095,212,236]" italics="true" pageId="15" pageNumber="1661">Litsea</emphasis>
</taxonomicName>
sp.)ı Rhodoliaceae (
<emphasis id="B97DEAB7FFABFFF6001D2798FE245B34" box="[327,423,246,270]" italics="true" pageId="15" pageNumber="1661">Rhodolia</emphasis>
sp.)ı Fagaceaeı 
<taxonomicName id="4C094D26FFABFFF603062798FD735B34" box="[604,752,246,270]" class="Magnoliopsida" family="Sterculiaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="15" pageNumber="1661" phylum="Tracheophyta" rank="family">Sterculiaceae</taxonomicName>
(
<taxonomicName id="4C094D26FFABFFF6025A2798FCEF5B34" box="[768,876,246,270]" class="Magnoliopsida" family="Malvaceae" genus="Scaphium" kingdom="Plantae" order="Malvales" pageId="15" pageNumber="1661" phylum="Tracheophyta" rank="genus">
<emphasis id="B97DEAB7FFABFFF6025A2798FCEF5B34" box="[768,876,246,270]" italics="true" pageId="15" pageNumber="1661">Scaphium</emphasis>
</taxonomicName>
sp.) (
<figureCitation id="13322A20FFABFFF602F62798FB8A5B34" box="[940,1033,246,270]" captionStart="Figure 6" captionStartId="15.[160,225,1675,1698]" captionTargetBox="[162,1152,901,1644]" captionTargetId="figure@15.[162,1153,901,1644]" captionTargetPageId="15" captionText="Figure 6. Natural habitat of Liopeltis pallidonuchalis sp. nov. in Kon Chu Rang N.R., Gia Lai Province, Vietnam. Photo by Alina V. Alexandrova." figureDoi="http://doi.org/10.5281/zenodo.3670319" httpUri="https://zenodo.org/record/3670319/files/figure.png" pageId="15" pageNumber="1661">Figure 6</figureCitation>
).
</paragraph>
<paragraph id="8BB636A5FFABFFF6019A2677FDD7581E" blockId="15.[160,1157,142,825]" pageId="15" pageNumber="1661">
As with other species of the genusı 
<taxonomicName id="4C094D26FFABFFF603002677FCDC5B0B" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[602,863,281,305]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="1661" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFABFFF603002677FCDC5B0B" box="[602,863,281,305]" italics="true" pageId="15" pageNumber="1661">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFABFFF602322677FC435B0B" box="[872,960,281,305]" pageId="15" pageNumber="1661" rank="species">sp. nov.</taxonomicNameLabel>
was found to be a very secretive speciesı with terrestrial and nocturnal activity; the only specimen that we received alive was recorded at dusk around 19:30 h at an air temperature of 20 
<emphasis id="B97DEAB7FFABFFF605182630FBCC5B4C" box="[1090,1103,350,374]" italics="true" pageId="15" pageNumber="1661">–</emphasis>
22°C and 100% humidity; three other known specimens are roadkills. The 
<typeStatus id="54B28807FFABFFF602F626EFFB985BA3" box="[940,1051,385,409]" pageId="15" pageNumber="1661" type="holotype">holotypes</typeStatus>
stomach contained remains of a spiderı which we tentatively identified as a member of the family 
<taxonomicName id="4C094D26FFABFFF601FA26A8FECA5BE4" authority="Gen." authorityName="Gen." box="[160,329,454,478]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="1661" phylum="Arthropoda" rank="family">Lycosidae Gen.</taxonomicName>
sp. The adult female ZFMK 83105 described by 
<bibRefCitation id="EF984B54FFABFFF6023F26A8FB075BE4" author="Ziegler Ti Hoang QX" box="[869,1156,454,478]" pageId="15" pageNumber="1661" pagination="115" refId="ref15062" refString="Ziegler Ti Hoang QX. 2009. Liopeltis frenatus. Herpetol Rev. 40 (1): 115." type="journal article" year="2009">Ziegler and Hoang (2009)</bibRefCitation>
contained four eggs of 
<quantity id="4CF19B40FFABFFF600FF2687FD9F583B" box="[421,540,489,513]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="15" pageNumber="1661" unit="mm" value="26.5" valueMax="28.0" valueMin="25.0">
25 
<emphasis id="B97DEAB7FFABFFF6009A2687FE4E583B" box="[448,461,489,513]" italics="true" pageId="15" pageNumber="1661">–</emphasis>
28 mm
</quantity>
in length and 
<quantity id="4CF19B40FFABFFF6039A2687FC98583B" box="[704,795,489,513]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="15" pageNumber="1661" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">
7 
<emphasis id="B97DEAB7FFABFFF603972687FD59583B" box="[717,730,489,513]" italics="true" pageId="15" pageNumber="1661">–</emphasis>
9 mm
</quantity>
in width. Nothing else is known about the biology of the new species.
</paragraph>
<paragraph id="8BB636A5FFABFFE9019A2540FBEF5B34" blockId="15.[160,1157,142,825]" lastBlockId="16.[160,1155,142,270]" lastPageId="16" lastPageNumber="1662" pageId="15" pageNumber="1661">
Other species of snakes recorded syntopically with the new species at the 
<typeStatus id="54B28807FFABFFF6050B2540FB00587C" box="[1105,1155,558,582]" pageId="15" pageNumber="1661">type</typeStatus>
locality in Kon Chu Rang N.R. (
<collectingRegion id="49CDF847FFABFFF60379253FFD6A5853" box="[547,745,593,617]" country="Vietnam" name="Gia Lai" pageId="15" pageNumber="1661">Gia Lai Province</collectingRegion>
) included 
<taxonomicName id="4C094D26FFABFFF6022A253FFF6858B1" authority="(Blyth)" baseAuthorityName="Blyth" baseAuthorityYear="1856" class="Reptilia" family="Colubridae" genus="Rhabdophis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="1661" phylum="Chordata" rank="species" species="nigrocinctus">
<emphasis id="B97DEAB7FFABFFF6022A253FFB005853" box="[880,1155,593,617]" italics="true" pageId="15" pageNumber="1661">Rhabdophis nigrocinctus</emphasis>
(Blyth)
</taxonomicName>
ı 
<taxonomicName id="4C094D26FFABFFF601AF251DFDF058B1" authority="(Boulenger)" baseAuthorityName="Boulenger" baseAuthorityYear="1899" box="[245,627,627,651]" class="Reptilia" family="Colubridae" genus="Sinonatrix" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="1661" phylum="Chordata" rank="species" species="percarinata">
<emphasis id="B97DEAB7FFABFFF601AF251DFE6458B1" box="[245,487,627,651]" italics="true" pageId="15" pageNumber="1661">Sinonatrix percarinata</emphasis>
(Boulenger)
</taxonomicName>
ı 
<taxonomicName id="4C094D26FFABFFF60327251DFC4358B1" authority="(Gunther)" baseAuthorityName="Gunther" box="[637,960,627,651]" class="Reptilia" family="Viperidae" genus="Ovophis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="1661" phylum="Chordata" rank="species" species="monticola">
<emphasis id="B97DEAB7FFABFFF60327251DFCC958B1" box="[637,842,627,651]" italics="true" pageId="15" pageNumber="1661">Ovophis monticola</emphasis>
(Günther)
</taxonomicName>
ı 
<taxonomicName id="4C094D26FFABFFF60290251DFEA65894" authority="(Boulenger)" baseAuthorityName="Boulenger" baseAuthorityYear="1905" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="1661" phylum="Chordata" rank="species" species="hamptoni">
<emphasis id="B97DEAB7FFABFFF60290251DFB0058B1" box="[970,1155,627,651]" italics="true" pageId="15" pageNumber="1661">Pareas hamptoni</emphasis>
(Boulenger)
</taxonomicName>
ı 
<taxonomicName id="4C094D26FFABFFF6006925F8FBCD5894" authority="(Davidi Baini Nguyeni Orlovi Vogeli Vu &amp; Ziegler)" box="[307,1102,662,686]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="15" pageNumber="1661" phylum="Chordata" rank="species" species="leucomystax">
<emphasis id="B97DEAB7FFABFFF6006925F8FD8E5894" box="[307,525,662,686]" italics="true" pageId="15" pageNumber="1661">Hebius leucomystax</emphasis>
(Davidı Bainı Nguyenı Orlovı Vogelı Vu &amp; Ziegler)
</taxonomicName>
and 
<taxonomicName id="4C094D26FFABFFF601FA25D7FCC458EB" authority="Davidi Vidal &amp; Pauwels. In Bach Ma N. P." authorityName="Davidi Vidal &amp; Pauwels. In Bach Ma N. P." box="[160,839,697,721]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="1661" phylum="Chordata" rank="species" species="vogeli">
<emphasis id="B97DEAB7FFABFFF601FA25D7FEF058EB" box="[160,371,697,721]" italics="true" pageId="15" pageNumber="1661">Trimeresurus vogeli</emphasis>
Davidı Vidal &amp; Pauwels. In Bach Ma N.P.
</taxonomicName>
(Thua Thien 
<emphasis id="B97DEAB7FFABFFF6028D25D7FC6758EB" box="[983,996,697,721]" italics="true" pageId="15" pageNumber="1661">–</emphasis>
<collectingRegion id="49CDF847FFABFFF602BE25D7FBFE58EB" box="[996,1149,697,721]" inferredBySuffix="true" pageId="15" pageNumber="1661">Hue Province</collectingRegion>
) the new species was recorded in sympatry with 
<emphasis id="B97DEAB7FFABFFF603B425B5FC4A58C9" box="[750,969,731,755]" italics="true" pageId="15" pageNumber="1661">Calamaria concolor</emphasis>
Orlovı Nguyenı Nguyenı Ananjeva &amp; Hoı 
<taxonomicName id="4C094D26FFABFFF600E52590FC21592C" authority="Dumerili Bibron &amp; Dumerili" authorityName="Dumerili Bibron &amp; Dumerili" box="[447,930,766,790]" class="Gastropoda" family="Strombidae" genus="Canarium" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="1661" phylum="Mollusca" rank="species" species="pavimentata">
<emphasis id="B97DEAB7FFABFFF600E52590FDE4592C" box="[447,615,766,790]" italics="true" pageId="15" pageNumber="1661">C. pavimentata</emphasis>
Dumérilı Bibron &amp; Dumérilı
</taxonomicName>
<taxonomicName id="4C094D26FFABFFF602F12590FEF15903" authority="(Bourret)" baseAuthorityName="Bourret" baseAuthorityYear="1935" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="1661" phylum="Chordata" rank="species" species="ngansonensis">
<emphasis id="B97DEAB7FFABFFF602F12590FF795903" italics="true" pageId="15" pageNumber="1661">Dendrelaphis ngansonensis</emphasis>
(Bourret)
</taxonomicName>
ı 
<taxonomicName id="4C094D26FFABFFF600DD244FFD605903" authority="(Anderson)" baseAuthorityName="Anderson" baseAuthorityYear="1879" box="[391,739,801,825]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="1661" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B97DEAB7FFABFFF600DD244FFDD05903" box="[391,595,801,825]" italics="true" pageId="15" pageNumber="1661">Lycodon fasciatus</emphasis>
(Anderson)
</taxonomicName>
ı 
<taxonomicName id="4C094D26FFABFFF603A2244FFBB65903" authority="(Blyth)" baseAuthorityName="Blyth" baseAuthorityYear="1854" box="[760,1077,801,825]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="1661" phylum="Chordata" rank="species" species="catenatus">
<emphasis id="B97DEAB7FFABFFF603A2244FFC5B5903" box="[760,984,801,825]" italics="true" pageId="15" pageNumber="1661">Oligodon catenatus</emphasis>
(Blyth)
</taxonomicName>
ı 
<taxonomicName id="4C094D26FFABFFE90510244FFEF65A9C" authority="(Gunther)" baseAuthorityName="Gunther" baseAuthorityYear="1858" class="Reptilia" family="Colubridae" genus="Ptyas" kingdom="Animalia" lastPageId="16" lastPageNumber="1662" order="Squamata" pageId="15" pageNumber="1661" phylum="Chordata" rank="species" species="carinata">
<emphasis id="B97DEAB7FFABFFE90510244FFF795A9C" italics="true" lastPageId="16" lastPageNumber="1662" pageId="15" pageNumber="1661">Ptyas carinata</emphasis>
(Günther)
</taxonomicName>
ı 
<taxonomicName id="4C094D26FFB4FFE900DE27E0FDE05A9C" authority="(Schlegel)" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[388,611,142,166]" class="Reptilia" family="Colubridae" genus="Ptyas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="korros">
<emphasis id="B97DEAB7FFB4FFE900DE27E0FE655A9C" box="[388,486,142,166]" italics="true" pageId="16" pageNumber="1662">P. korros</emphasis>
(Schlegel)
</taxonomicName>
ı 
<taxonomicName id="4C094D26FFB4FFE9032B27E0FC105A9C" authority="(Blyth)" baseAuthorityName="Blyth" baseAuthorityYear="1854" box="[625,915,142,166]" class="Reptilia" family="Colubridae" genus="Ptyas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="nigromarginata">
<emphasis id="B97DEAB7FFB4FFE9032B27E0FCBF5A9C" box="[625,828,142,166]" italics="true" pageId="16" pageNumber="1662">P. nigromarginata</emphasis>
(Blyth)
</taxonomicName>
ı 
<emphasis id="B97DEAB7FFB4FFE902F827E0FB005A9C" box="[930,1155,142,166]" italics="true" pageId="16" pageNumber="1662">
<taxonomicName id="4C094D26FFB4FFE902F827E0FBFC5A9C" box="[930,1151,142,166]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="leucomystax">Hebius leucomystax</taxonomicName>
,
</emphasis>
<taxonomicName id="4C094D26FFB4FFE901FA27DFFD8A5AF3" authority="(Blyth)" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[160,521,177,201]" class="Reptilia" family="Colubridae" genus="Pseudoxenodon" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="macrops">
<emphasis id="B97DEAB7FFB4FFE901FA27DFFE305AF3" box="[160,435,177,201]" italics="true" pageId="16" pageNumber="1662">Pseudoxenodon macrops</emphasis>
(Blyth)
</taxonomicName>
ı 
<taxonomicName id="4C094D26FFB4FFE9034C27DFFCD55AF3" authority="(Gray)" baseAuthorityName="Gray" baseAuthorityYear="1853" box="[534,854,177,201]" class="Reptilia" family="Colubridae" genus="Sibynophis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="collaris">
<emphasis id="B97DEAB7FFB4FFE9034C27DFFD085AF3" box="[534,651,177,201]" italics="true" pageId="16" pageNumber="1662">Sibynophis</emphasis>
cf. 
<emphasis id="B97DEAB7FFB4FFE903E327DFFC865AF3" box="[697,773,177,201]" italics="true" pageId="16" pageNumber="1662">collaris</emphasis>
(Gray)
</taxonomicName>
ı 
<taxonomicName id="4C094D26FFB4FFE9023927DFFD365AD6" authority="Kuchi Kiziriani Nguyeni Lawsoni Donnelly &amp; Mebsi" class="Reptilia" family="Elapidae" genus="Bungarus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="slowinskii">
<emphasis id="B97DEAB7FFB4FFE9023927DFFBB85AF3" box="[867,1083,177,201]" italics="true" pageId="16" pageNumber="1662">Bungarus slowinskii</emphasis>
Kuchı Kizirianı Nguyenı Lawsonı Donnelly &amp; Mebsı
</taxonomicName>
<taxonomicName id="4C094D26FFB4FFE9039F27BAFBAA5AD6" authority="(Cantor)" baseAuthorityName="Cantor" baseAuthorityYear="1836" box="[709,1065,212,236]" class="Reptilia" family="Elapidae" genus="Ophiophagus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="hannah">
<emphasis id="B97DEAB7FFB4FFE9039F27BAFC3F5AD6" box="[709,956,212,236]" italics="true" pageId="16" pageNumber="1662">Ophiophagus hannah</emphasis>
(Cantor)
</taxonomicName>
ı 
<emphasis id="B97DEAB7FFB4FFE9056127BAFE6F5B34" italics="true" pageId="16" pageNumber="1662">
<taxonomicName id="4C094D26FFB4FFE9056127BAFE8F5B34" baseAuthorityName="Boulenger" baseAuthorityYear="1905" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="hamptoni">Pareas hamptoni</taxonomicName>
, 
<taxonomicName id="4C094D26FFB4FFE900432798FE6A5B34" box="[281,489,246,270]" class="Reptilia" family="Viperidae" genus="Ovophis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="monticola">Ovophis monticola</taxonomicName>
,
</emphasis>
<taxonomicName id="4C094D26FFB4FFE900AF2798FCD75B34" authority="(Smith)" baseAuthorityName="Smith" baseAuthorityYear="1930" box="[501,852,246,270]" class="Arachnida" family="Lycosidae" genus="Protobothrops" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="1662" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="B97DEAB7FFB4FFE900AF2798FD745B34" box="[501,759,246,270]" italics="true" pageId="16" pageNumber="1662">Protobothrops cornutus</emphasis>
(Smith)
</taxonomicName>
ı and 
<taxonomicName id="4C094D26FFB4FFE902C92798FBE65B34" box="[915,1125,246,270]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="vogeli">
<emphasis id="B97DEAB7FFB4FFE902C92798FBE65B34" box="[915,1125,246,270]" italics="true" pageId="16" pageNumber="1662">Trimeresurus vogeli</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF76662DFFABFFF601FA21E5FDBB5C85" ID-DOI="http://doi.org/10.5281/zenodo.3670319" ID-Zenodo-Dep="3670319" httpUri="https://zenodo.org/record/3670319/files/figure.png" pageId="15" pageNumber="1661" startId="15.[160,225,1675,1698]" targetBox="[162,1152,901,1644]" targetPageId="15">
<paragraph id="8BB636A5FFABFFF601FA21E5FDBB5C85" blockId="15.[160,1157,1674,1727]" pageId="15" pageNumber="1661">
<emphasis id="B97DEAB7FFABFFF601FA21E5FF785C98" bold="true" box="[160,251,1675,1698]" pageId="15" pageNumber="1661">Figure 6.</emphasis>
Natural habitat of 
<taxonomicName id="4C094D26FFABFFF600E121E4FD235C9B" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[443,672,1674,1697]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="1661" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFABFFF600E121E4FD235C9B" box="[443,672,1674,1697]" italics="true" pageId="15" pageNumber="1661">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFABFFF603FD21E5FD755C98" box="[679,758,1675,1698]" pageId="15" pageNumber="1661" rank="species">sp. nov.</taxonomicNameLabel>
in Kon Chu Rang N.R., Gia Lai Province, Vietnam. Photo by Alina V. Alexandrova.
</paragraph>
</caption>
<paragraph id="8BB636A5FFB4FFE901FA2655FC905895" blockId="16.[160,1156,315,1727]" pageId="16" pageNumber="1662">
<emphasis id="B97DEAB7FFB4FFE901FA2655FEC35B6F" bold="true" box="[160,320,315,341]" italics="true" pageId="16" pageNumber="1662">Comparisons.</emphasis>
Comparative morphological data for the new species and currently recognised members of the genera 
<taxonomicName id="4C094D26FFB4FFE903652631FD145B4D" authorityName="Fitzingeri" authorityYear="1843" box="[575,663,351,375]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="genus">
<emphasis id="B97DEAB7FFB4FFE903652631FD145B4D" box="[575,663,351,375]" italics="true" pageId="16" pageNumber="1662">Liopeltis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C094D26FFB4FFE9038C2631FCEF5B4D" authorityName="Fitzingeri" authorityYear="1843" box="[726,876,351,375]" class="Reptilia" family="Colubridae" genus="Gongylosoma" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="genus">
<emphasis id="B97DEAB7FFB4FFE9038C2631FCEF5B4D" box="[726,876,351,375]" italics="true" pageId="16" pageNumber="1662">Gongylosoma</emphasis>
</taxonomicName>
is presented in 
<tableCitation id="C68B031EFFB4FFE905732631FBFD5B4D" box="[1065,1150,351,375]" captionStart="Table 1" captionStartId="3.[133,189,463,486]" captionTargetBox="[144,1702,557,1401]" captionTargetPageId="3" captionText="Table 1. Basic characters for the genera Liopeltis and Gongylosoma compared to the new species. Abbreviations: BR: Brunei; BT: Bhutan; CA: Cambodia; CN: China; ID: Indonesia; IN: India; LA: Laos; ML: Malaysia; MY: Myanmar; NP: Nepal; PL: Philippines; SG: Singapore; SL: Sri Lanka; TL: Thailand; VN: Vietnam; ‘?’: requires verification; n/a: not available." httpUri="http://table.plazi.org/id/DF76662DFFA7FFFA01DF26A1FD8F581B" pageId="16" pageNumber="1662" tableUuid="DF76662DFFA7FFFA01DF26A1FD8F581B">Table 1</tableCitation>
. We decided to include the new species into the genus 
<taxonomicName id="4C094D26FFB4FFE9025326ECFCE25BA0" authorityName="Fitzingeri" authorityYear="1843" box="[777,865,386,410]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="genus">
<emphasis id="B97DEAB7FFB4FFE9025326ECFCE25BA0" box="[777,865,386,410]" italics="true" pageId="16" pageNumber="1662">Liopeltis</emphasis>
</taxonomicName>
rather than 
<taxonomicName id="4C094D26FFB4FFE902B726ECFB005BA0" authorityName="Fitzingeri" authorityYear="1843" box="[1005,1155,386,410]" class="Reptilia" family="Colubridae" genus="Gongylosoma" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="genus">
<emphasis id="B97DEAB7FFB4FFE902B726ECFB005BA0" box="[1005,1155,386,410]" italics="true" pageId="16" pageNumber="1662">Gongylosoma</emphasis>
</taxonomicName>
for the following reasons: the number of dorsal scalesı and the prefrontal touching the upper labialsı both characters are unknown in the genus 
<taxonomicName id="4C094D26FFB4FFE9021D26A9FC5E5BE5" authorityName="Fitzingeri" authorityYear="1843" box="[839,989,455,479]" class="Reptilia" family="Colubridae" genus="Gongylosoma" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="genus">
<emphasis id="B97DEAB7FFB4FFE9021D26A9FC5E5BE5" box="[839,989,455,479]" italics="true" pageId="16" pageNumber="1662">Gongylosoma</emphasis>
</taxonomicName>
. According to 
<bibRefCitation id="EF984B54FFB4FFE901FA2684FEC75838" author="Leviton AE" box="[160,324,490,514]" pageId="16" pageNumber="1662" pagination="367 - 381" refId="ref13384" refString="Leviton AE. 1964. Contributions to a review of Philippine snakesi II. The snakes of the genera Liopeltis and Sibynophis. Philipp J Sci. 92: 367 - 381." type="journal article" year="1964">Leviton (1964)</bibRefCitation>
the head shape of the new species resembles the genus 
<taxonomicName id="4C094D26FFB4FFE902B22684FBFD5838" authorityName="Fitzingeri" authorityYear="1843" box="[1000,1150,490,514]" class="Reptilia" family="Colubridae" genus="Gongylosoma" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="genus">
<emphasis id="B97DEAB7FFB4FFE902B22684FBFD5838" box="[1000,1150,490,514]" italics="true" pageId="16" pageNumber="1662">Gongylosoma</emphasis>
</taxonomicName>
. Howeverı Leviton did only compare the two species 
<taxonomicName id="4C094D26FFB4FFE9024E2563FBCE581F" authority="(Boiei 1827)" baseAuthorityName="Boiei" baseAuthorityYear="1827" box="[788,1101,525,549]" class="Reptilia" family="Colubridae" genus="Gongylosoma" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="baliodeirum">
<emphasis id="B97DEAB7FFB4FFE9024E2563FC3A581F" box="[788,953,525,549]" italics="true" pageId="16" pageNumber="1662">G. baliodeirum</emphasis>
(
<bibRefCitation id="EF984B54FFB4FFE902912563FBC6581F" box="[971,1093,525,549]" pageId="16" pageNumber="1662" refId="ref12251" type="journal article">Boieı 1827</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C094D26FFB4FFE901FA2541FE8A587D" baseAuthorityName="Schlegeli" baseAuthorityYear="1837" box="[160,265,559,583]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="tricolor">
<emphasis id="B97DEAB7FFB4FFE901FA2541FE8A587D" box="[160,265,559,583]" italics="true" pageId="16" pageNumber="1662">L. tricolor</emphasis>
</taxonomicName>
. Several other 
<taxonomicName id="4C094D26FFB4FFE900E72541FD96587D" authorityName="Fitzingeri" authorityYear="1843" box="[445,533,559,583]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="genus">
<emphasis id="B97DEAB7FFB4FFE900E72541FD96587D" box="[445,533,559,583]" italics="true" pageId="16" pageNumber="1662">Liopeltis</emphasis>
</taxonomicName>
species do rather have the 
<emphasis id="B97DEAB7FFB4FFE9023E2541FCEA587D" box="[868,873,559,583]" italics="true" pageId="16" pageNumber="1662">‘</emphasis>
shortı deep and convex 
<emphasis id="B97DEAB7FFB4FFE905242541FB07587D" box="[1150,1156,559,583]" italics="true" pageId="16" pageNumber="1662">’</emphasis>
head profile than the 
<emphasis id="B97DEAB7FFB4FFE900CE253CFE1A5850" box="[404,409,594,618]" italics="true" pageId="16" pageNumber="1662">‘</emphasis>
long shallowed and flattened 
<emphasis id="B97DEAB7FFB4FFE903B8253CFD6B5850" box="[738,744,594,618]" italics="true" pageId="16" pageNumber="1662">’</emphasis>
shape which Leviton defined for the genus 
<taxonomicName id="4C094D26FFB4FFE901B1251BFEC058B7" authorityName="Fitzingeri" authorityYear="1843" box="[235,323,629,653]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="genus">
<emphasis id="B97DEAB7FFB4FFE901B1251BFEC058B7" box="[235,323,629,653]" italics="true" pageId="16" pageNumber="1662">Liopeltis</emphasis>
</taxonomicName>
. Presentlyı based entirely on morphological reasons we are not convinced about the necessity of separation of these two genera.
</paragraph>
<paragraph id="8BB636A5FFB4FFE9019A25D4FE595998" blockId="16.[160,1156,315,1727]" pageId="16" pageNumber="1662">
Morphologically 
<taxonomicName id="4C094D26FFB4FFE900D625D4FD1D58E8" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[396,670,698,722]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFB4FFE900D625D4FD1D58E8" box="[396,670,698,722]" italics="true" pageId="16" pageNumber="1662">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFB4FFE903EE25D4FC9958E8" box="[692,794,698,722]" pageId="16" pageNumber="1662" rank="species">sp. nov.</taxonomicNameLabel>
is most closely resembling 
<taxonomicName id="4C094D26FFB4FFE901FA25B2FEAB58CE" baseAuthorityName="Guntheri" baseAuthorityYear="1858" box="[160,296,732,756]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="calamaria">
<emphasis id="B97DEAB7FFB4FFE901FA25B2FEAB58CE" box="[160,296,732,756]" italics="true" pageId="16" pageNumber="1662">L. calamaria</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C094D26FFB4FFE9003D25B2FE5358CE" baseAuthorityName="Guntheri" baseAuthorityYear="1858" box="[359,464,732,756]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="frenata">
<emphasis id="B97DEAB7FFB4FFE9003D25B2FE5358CE" box="[359,464,732,756]" italics="true" pageId="16" pageNumber="1662">L. frenata</emphasis>
</taxonomicName>
. The main characters which separate it from 
<taxonomicName id="4C094D26FFB4FFE902BA25B2FBEA58CE" baseAuthorityName="Guntheri" baseAuthorityYear="1858" box="[992,1129,732,756]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="calamaria">
<emphasis id="B97DEAB7FFB4FFE902BA25B2FBEA58CE" box="[992,1129,732,756]" italics="true" pageId="16" pageNumber="1662">L. calamaria</emphasis>
</taxonomicName>
is the colouration; the fact that the nasals are fused with the internasals in 
<taxonomicName id="4C094D26FFB4FFE902962591FBD1592D" baseAuthorityName="Guntheri" baseAuthorityYear="1858" box="[972,1106,767,791]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="calamaria">
<emphasis id="B97DEAB7FFB4FFE902962591FBD1592D" box="[972,1106,767,791]" italics="true" pageId="16" pageNumber="1662">L. calamaria</emphasis>
</taxonomicName>
and the prefrontal is widely separated from the prefrontal by the postnasal. 
<materialsCitation id="3B613CF8FFB4FFE9029E244CFF2D5945" ID-GBIF-Occurrence="2565890770" collectorName="Beside" county="Indiani" location="Nepalese" municipality="Lankan" pageId="16" pageNumber="1662" specimenCount="1" stateProvince="Surrey">
<collectorName id="26FC5373FFB4FFE9029E244CFB8E5900" box="[964,1037,802,826]" pageId="16" pageNumber="1662">Beside</collectorName>
that there is a huge distribution gap between the 
<collectingCounty id="62D74E29FFB4FFE90327242AFD4A5966" box="[637,713,836,860]" pageId="16" pageNumber="1662">Indianı</collectingCounty>
<collectingRegion id="49CDF847FFB4FFE9038E242AFD735966" box="[724,752,836,860]" country="United Kingdom" name="Surrey" pageId="16" pageNumber="1662">Sri</collectingRegion>
<collectingMunicipality id="6BD2ACDFFFB4FFE903A6242AFCCF5966" box="[764,844,836,860]" pageId="16" pageNumber="1662">Lankan</collectingMunicipality>
and 
<location id="8ED6607EFFB4FFE902D4242AFC765966" LSID="urn:lsid:plazi:treatment:03A087B3FFADFFEB009D2669FF6B5FF7:8ED6607EFFB4FFE902D4242AFC765966" box="[910,1013,836,860]" county="Indiani" municipality="Lankan" name="Nepalese" pageId="16" pageNumber="1662" stateProvince="Surrey">Nepalese</location>
localities of 
<taxonomicName id="4C094D26FFB4FFE901FA2409FF2D5945" baseAuthorityName="Guntheri" baseAuthorityYear="1858" box="[160,174,871,895]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="calamaria">
<emphasis id="B97DEAB7FFB4FFE901FA2409FF2D5945" box="[160,174,871,895]" italics="true" pageId="16" pageNumber="1662">L</emphasis>
</taxonomicName>
</materialsCitation>
<emphasis id="B97DEAB7FFB4FFE901F42409FEAF5945" box="[174,300,871,895]" italics="true" pageId="16" pageNumber="1662">. calamaria</emphasis>
and the 
<typeStatus id="54B28807FFB4FFE900FE2409FE555945" box="[420,470,871,895]" pageId="16" pageNumber="1662">type</typeStatus>
locality of 
<taxonomicName id="4C094D26FFB4FFE9033E2409FCED5945" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[612,878,871,895]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFB4FFE9033E2409FCED5945" box="[612,878,871,895]" italics="true" pageId="16" pageNumber="1662">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFB4FFE902212409FC5F5945" box="[891,988,871,895]" pageId="16" pageNumber="1662" rank="species">sp. nov.</taxonomicNameLabel>
ı which makes a conspecifity very unlikely.
</paragraph>
<paragraph id="8BB636A5FFB4FFE9019A24C2FCD95C6D" blockId="16.[160,1156,315,1727]" pageId="16" pageNumber="1662">
The main differences of the new species from 
<taxonomicName id="4C094D26FFB4FFE903B324C2FCD659FE" baseAuthorityName="Guntheri" baseAuthorityYear="1858" box="[745,853,940,964]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="frenata">
<emphasis id="B97DEAB7FFB4FFE903B324C2FCD659FE" box="[745,853,940,964]" italics="true" pageId="16" pageNumber="1662">L. frenata</emphasis>
</taxonomicName>
are the lower number of ventralsı subcaudals and the colouration as well. For photos of the type specimen of 
<taxonomicName id="4C094D26FFB4FFE901FA249CFD8D5E30" authority="Guntheri 1858" authorityName="Guntheri" authorityYear="1858" box="[160,526,1010,1034]" class="Squamata" family="Colubridae" genus="Cyclophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="B97DEAB7FFB4FFE901FA249CFEE45E30" box="[160,359,1010,1034]" italics="true" pageId="16" pageNumber="1662">Cyclophis frenatus</emphasis>
Güntherı 1858
</taxonomicName>
(BMNH 1946.1.1.72) see 
<figureCitation id="13322A20FFB4FFE9027F249CFC025E30" box="[805,897,1010,1034]" captionStart="Figure 7" captionStartId="17.[160,225,1045,1068]" captionTargetBox="[164,1150,143,1012]" captionTargetId="figure@17.[163,1153,142,1013]" captionTargetPageId="17" captionText="Figure 7. Morphology of Cyclophis frenatus Günther, 1858 holotype in preservative (BMNH 1946.1.1.72, male; from ‘Mountains near Affghanistan’ [= Afghanistan], corrected to ‘Khasi Hills’ by Boulenger 1894). A – general dorsal view; B – general ventral view; C – head in dorsal view; D – head in ventral view; E – head in lateral view. Photos by Gernot Vogel." figureDoi="http://doi.org/10.5281/zenodo.3670321" httpUri="https://zenodo.org/record/3670321/files/figure.png" pageId="16" pageNumber="1662">Figure 7</figureCitation>
. 
<taxonomicName id="4C094D26FFB4FFE902D7249CFF355E16" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFB4FFE902D7249CFF355E16" italics="true" pageId="16" pageNumber="1662">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFB4FFE901E7237AFE955E16" box="[189,278,1044,1068]" pageId="16" pageNumber="1662" rank="species">sp. nov.</taxonomicNameLabel>
ı differs from 
<taxonomicName id="4C094D26FFB4FFE900FD237AFD8D5E16" baseAuthorityName="Guntheri" baseAuthorityYear="1858" box="[423,526,1044,1068]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="frenata">
<emphasis id="B97DEAB7FFB4FFE900FD237AFD8D5E16" box="[423,526,1044,1068]" italics="true" pageId="16" pageNumber="1662">L. frenata</emphasis>
</taxonomicName>
by a significantly lower number of ventrals (126 
<emphasis id="B97DEAB7FFB4FFE90573237AFBB55E16" box="[1065,1078,1044,1068]" italics="true" pageId="16" pageNumber="1662">–</emphasis>
138 vs. 140 
<emphasis id="B97DEAB7FFB4FFE901932359FF555E75" box="[201,214,1079,1103]" italics="true" pageId="16" pageNumber="1662">–</emphasis>
172). The difference in the number of subcaudals seems to be slimı but in 
<taxonomicName id="4C094D26FFB4FFE905462359FB075E75" baseAuthorityName="Guntheri" baseAuthorityYear="1858" box="[1052,1156,1079,1103]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="frenata">
<emphasis id="B97DEAB7FFB4FFE905462359FB075E75" box="[1052,1156,1079,1103]" italics="true" pageId="16" pageNumber="1662">L. frenata</emphasis>
</taxonomicName>
has a large sexual dimorphism in the number of subcaudals. So the lower part of the range of the subcaudals in 
<taxonomicName id="4C094D26FFB4FFE900862312FDC55EAE" baseAuthorityName="Guntheri" baseAuthorityYear="1858" box="[476,582,1148,1172]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="frenata">
<emphasis id="B97DEAB7FFB4FFE900862312FDC55EAE" box="[476,582,1148,1172]" italics="true" pageId="16" pageNumber="1662">L. frenata</emphasis>
</taxonomicName>
corresponds to female specimens and the lowest number is given by 
<bibRefCitation id="EF984B54FFB4FFE900D223F1FDC85E8D" author="Boulenger GA" box="[392,587,1183,1207]" pageId="16" pageNumber="1662" refId="ref12293" refString="Boulenger GA. 1890. The Fauna of British Indiai including ceylon and Burma. Reptile and Batrachia. England: Taylor Francis Lond.; p. 541." type="journal volume" year="1890">Boulenger (1890)</bibRefCitation>
as 
<quantity id="4CF19B40FFB4FFE9032E23F1FD555E8D" box="[628,726,1183,1207]" metricMagnitude="0" metricUnit="m" metricValue="2.3241" metricValueMax="2.4384" metricValueMin="2.2098" pageId="16" pageNumber="1662" unit="in" value="91.5" valueMax="96.0" valueMin="87.0">
87 
<emphasis id="B97DEAB7FFB4FFE903D523F1FD1F5E8D" box="[655,668,1183,1207]" italics="true" pageId="16" pageNumber="1662">–</emphasis>
96 in
</quantity>
both sexesı by 
<bibRefCitation id="EF984B54FFB4FFE902D723F1FB9D5E8D" author="Smith MA" box="[909,1054,1183,1207]" pageId="16" pageNumber="1662" pagination="583" refId="ref14375" refString="Smith MA. 1943. The Fauna of British Indiai Ceylon and Burmai including the whole of the Indo- Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). London: Taylor and Francis; p. 583." type="book chapter" year="1943">Smith (1943)</bibRefCitation>
and 
<bibRefCitation id="EF984B54FFB4FFE9050023F1FF655EE0" author="Das I" pageId="16" pageNumber="1662" refId="ref12497" refString="Das I. 2010. A field guide to the reptiles of south-east Asia. London: New Holland Publishers (UK) i Ltd.; p. 376." type="journal volume" year="2010">Das (2010)</bibRefCitation>
as 
<quantity id="4CF19B40FFB4FFE9005623ACFEF85EE0" box="[268,379,1218,1242]" metricMagnitude="0" metricUnit="m" metricValue="2.2225" metricValueMax="2.667" metricValueMin="1.778" pageId="16" pageNumber="1662" unit="in" value="87.5" valueMax="105.0" valueMin="70.0">
70 
<emphasis id="B97DEAB7FFB4FFE9007223ACFEB65EE0" box="[296,309,1218,1242]" italics="true" pageId="16" pageNumber="1662">–</emphasis>
105 in
</quantity>
both sexesı by 
<bibRefCitation id="EF984B54FFB4FFE9037623ACFD415EE0" author="Deuve J" box="[556,706,1218,1242]" pageId="16" pageNumber="1662" pagination="1 - 252" refId="ref12625" refString="Deuve J. 1970. Serpents du Laos. Paris: Memoirs O. R. S. T. O. M.; Vol. 39. p. 1 - 252. [in French]." type="journal article" year="1970">Deuve (1970)</bibRefCitation>
as 
<quantity id="4CF19B40FFB4FFE903B223ACFCD45EE0" box="[744,855,1218,1242]" metricMagnitude="0" metricUnit="m" metricValue="2.413" metricValueMax="2.6162" metricValueMin="2.2098" pageId="16" pageNumber="1662" unit="in" value="95.0" valueMax="103.0" valueMin="87.0">
87 
<emphasis id="B97DEAB7FFB4FFE9025E23ACFC925EE0" box="[772,785,1218,1242]" italics="true" pageId="16" pageNumber="1662">–</emphasis>
103 in
</quantity>
both sexesı by 
<bibRefCitation id="EF984B54FFB4FFE9055223ACFF655EC7" author="Orlov NLi Ryabov SAi Nguyen SVi Nguyen TQ" pageId="16" pageNumber="1662" pagination="217 - 240" refId="ref13955" refString="Orlov NLi Ryabov SAi Nguyen SVi Nguyen TQ. 2003. New records and data on the poorly known snakes of Vietnam. Russ J Herpetol. 10: 217 - 240." type="journal article" year="2003">Orlov et al. (2003)</bibRefCitation>
as 
<quantity id="4CF19B40FFB4FFE9004B238BFEC85EC7" box="[273,331,1253,1277]" metricMagnitude="0" metricUnit="m" metricValue="2.1336" pageId="16" pageNumber="1662" unit="in" value="84.0">84 in</quantity>
the maleı by 
<bibRefCitation id="EF984B54FFB4FFE900AA238BFD615EC7" author="Yang Di Rao D" box="[496,738,1253,1277]" pageId="16" pageNumber="1662" refId="ref14912" refString="Yang Di Rao D. 2008. Amphibia and Reptilia of Yunnan Kunming. Yunnan Publishing Group Corporationi Yunnan Science; p. 411. in Chinese." type="book" year="2008">Yang and Rao (2008)</bibRefCitation>
as 
<quantity id="4CF19B40FFB4FFE90256238BFCC45EC7" box="[780,839,1253,1277]" metricMagnitude="0" metricUnit="m" metricValue="2.3622" pageId="16" pageNumber="1662" unit="in" value="93.0">93 in</quantity>
the femaleı by 
<bibRefCitation id="EF984B54FFB4FFE9055A238BFF655F25" author="Pham AVi Nguyen SLHi Nguyen TQ" pageId="16" pageNumber="1662" pagination="771 - 777" refId="ref14112" refString="Pham AVi Nguyen SLHi Nguyen TQ. 2014. New records of snakes (Squamata: Serpentes) from Son La Province Vietnam. Herpetol Notes. 7: 771 - 777." type="journal article" year="2014">Pham et al. (2014)</bibRefCitation>
as 
<quantity id="4CF19B40FFB4FFE9004E2269FEDE5F25" box="[276,349,1287,1311]" metricMagnitude="0" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="1662" unit="in" value="100.0">100 in</quantity>
the maleı 
<quantity id="4CF19B40FFB4FFE900BB2269FD9E5F25" box="[481,541,1287,1311]" metricMagnitude="0" metricUnit="m" metricValue="2.3876" pageId="16" pageNumber="1662" unit="in" value="94.0">94 in</quantity>
the female. We counted 
<quantity id="4CF19B40FFB4FFE902172269FC0A5F25" box="[845,905,1287,1311]" metricMagnitude="0" metricUnit="m" metricValue="2.5146" pageId="16" pageNumber="1662" unit="in" value="99.0">99 in</quantity>
a single male of the 
<typeStatus id="54B28807FFB4FFE901FA2244FE875F78" box="[160,260,1322,1346]" pageId="16" pageNumber="1662" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C094D26FFB4FFE900762244FE165F78" baseAuthorityName="Guntheri" baseAuthorityYear="1858" box="[300,405,1322,1346]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="frenata">
<emphasis id="B97DEAB7FFB4FFE900762244FE165F78" box="[300,405,1322,1346]" italics="true" pageId="16" pageNumber="1662">L. frenata</emphasis>
</taxonomicName>
ı which is much higher than the 
<quantity id="4CF19B40FFB4FFE902562244FCED5F78" box="[780,878,1322,1346]" metricMagnitude="0" metricUnit="m" metricValue="1.7271999999999998" metricValueMax="1.7526" metricValueMin="1.7018" pageId="16" pageNumber="1662" unit="in" value="68.0" valueMax="69.0" valueMin="67.0">
67 
<emphasis id="B97DEAB7FFB4FFE9027D2244FCB75F78" box="[807,820,1322,1346]" italics="true" pageId="16" pageNumber="1662">–</emphasis>
69 in
</quantity>
the males and 
<quantity id="4CF19B40FFB4FFE9057B2244FB005F78" box="[1057,1155,1322,1346]" metricMagnitude="0" metricUnit="m" metricValue="1.8288" metricValueMax="1.8542" metricValueMin="1.8034" pageId="16" pageNumber="1662" unit="in" value="72.0" valueMax="73.0" valueMin="71.0">
71 
<emphasis id="B97DEAB7FFB4FFE905672244FBC95F78" box="[1085,1098,1322,1346]" italics="true" pageId="16" pageNumber="1662">–</emphasis>
73 in
</quantity>
the females of 
<taxonomicName id="4C094D26FFB4FFE900082223FDDA5F5F" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[338,601,1357,1381]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFB4FFE900082223FDDA5F5F" box="[338,601,1357,1381]" italics="true" pageId="16" pageNumber="1662">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFB4FFE9033E2223FD3D5F5F" box="[612,702,1357,1381]" pageId="16" pageNumber="1662" rank="species">sp. nov.</taxonomicNameLabel>
In 
<taxonomicName id="4C094D26FFB4FFE903B22223FC6C5F5F" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[744,1007,1357,1381]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFB4FFE903B22223FC6C5F5F" box="[744,1007,1357,1381]" italics="true" pageId="16" pageNumber="1662">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFB4FFE902A02223FBD75F5F" box="[1018,1108,1357,1381]" pageId="16" pageNumber="1662" rank="species">sp. nov.</taxonomicNameLabel>
the postocular stripe is thin whereas it is very thick in 
<taxonomicName id="4C094D26FFB4FFE903942201FCB65FBD" baseAuthorityName="Guntheri" baseAuthorityYear="1858" box="[718,821,1391,1415]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="frenata">
<emphasis id="B97DEAB7FFB4FFE903942201FCB65FBD" box="[718,821,1391,1415]" italics="true" pageId="16" pageNumber="1662">L. frenata</emphasis>
</taxonomicName>
(see 
<figureCitation id="13322A20FFB4FFE902352201FC735FBD" box="[879,1008,1391,1415]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStartId-0="17.[160,225,1045,1068]" captionStartId-1="18.[160,225,830,853]" captionTargetBox-0="[164,1150,143,1012]" captionTargetBox-1="[164,1151,143,797]" captionTargetId-0="figure@17.[163,1153,142,1013]" captionTargetId-1="figure@18.[163,1153,142,798]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="Figure 7. Morphology of Cyclophis frenatus Günther, 1858 holotype in preservative (BMNH 1946.1.1.72, male; from ‘Mountains near Affghanistan’ [= Afghanistan], corrected to ‘Khasi Hills’ by Boulenger 1894). A – general dorsal view; B – general ventral view; C – head in dorsal view; D – head in ventral view; E – head in lateral view. Photos by Gernot Vogel." captionText-1="Figure 8. Liopeltis frenata (Günther, 1858) from Wuliangshan Mt., Yunnan, China, in life. (a) – general lateral view; (b) – head in lateral view. Photos by Chung-Wei You." figureDoi-0="http://doi.org/10.5281/zenodo.3670321" figureDoi-1="http://doi.org/10.5281/zenodo.3670323" httpUri-0="https://zenodo.org/record/3670321/files/figure.png" httpUri-1="https://zenodo.org/record/3670323/files/figure.png" pageId="16" pageNumber="1662">
Figures 7 
<emphasis id="B97DEAB7FFB4FFE9028F2201FC615FBD" box="[981,994,1391,1415]" italics="true" pageId="16" pageNumber="1662">–</emphasis>
8
</figureCitation>
). Besides the postocular stripeı in 
<taxonomicName id="4C094D26FFB4FFE900D722FCFE745F90" baseAuthorityName="Guntheri" baseAuthorityYear="1858" box="[397,503,1426,1450]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="frenata">
<emphasis id="B97DEAB7FFB4FFE900D722FCFE745F90" box="[397,503,1426,1450]" italics="true" pageId="16" pageNumber="1662">L. frenata</emphasis>
</taxonomicName>
3 
<emphasis id="B97DEAB7FFB4FFE9035422FCFD985F90" box="[526,539,1426,1450]" italics="true" pageId="16" pageNumber="1662">–</emphasis>
4 narrower lateral dark stripes are presentı which run from the neck region posteriorly along the anterior half of the body (
<figureCitation id="13322A20FFB4FFE9029822DBFBA85FF7" box="[962,1067,1461,1485]" captionStart="Figure 7" captionStartId="17.[160,225,1045,1068]" captionTargetBox="[164,1150,143,1012]" captionTargetId="figure@17.[163,1153,142,1013]" captionTargetPageId="17" captionText="Figure 7. Morphology of Cyclophis frenatus Günther, 1858 holotype in preservative (BMNH 1946.1.1.72, male; from ‘Mountains near Affghanistan’ [= Afghanistan], corrected to ‘Khasi Hills’ by Boulenger 1894). A – general dorsal view; B – general ventral view; C – head in dorsal view; D – head in ventral view; E – head in lateral view. Photos by Gernot Vogel." figureDoi="http://doi.org/10.5281/zenodo.3670321" httpUri="https://zenodo.org/record/3670321/files/figure.png" pageId="16" pageNumber="1662">Figures 7</figureCitation>
and 
<figureCitation id="13322A20FFB4FFE9053222DBFBF55FF7" box="[1128,1142,1461,1485]" captionStart="Figure 8" captionStartId="18.[160,225,830,853]" captionTargetBox="[164,1151,143,797]" captionTargetId="figure@18.[163,1153,142,798]" captionTargetPageId="18" captionText="Figure 8. Liopeltis frenata (Günther, 1858) from Wuliangshan Mt., Yunnan, China, in life. (a) – general lateral view; (b) – head in lateral view. Photos by Chung-Wei You." figureDoi="http://doi.org/10.5281/zenodo.3670323" httpUri="https://zenodo.org/record/3670323/files/figure.png" pageId="16" pageNumber="1662">8</figureCitation>
); such dark markings are absent in the new species. Finallyı molecular data suggest that there is a deep divergence between cyt 
<emphasis id="B97DEAB7FFB4FFE903242294FD0F5C28" box="[638,652,1530,1554]" italics="true" pageId="16" pageNumber="1662">b</emphasis>
gene sequences of 
<taxonomicName id="4C094D26FFB4FFE902272294FC6B5C28" baseAuthorityName="Guntheri" baseAuthorityYear="1858" box="[893,1000,1530,1554]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" sensu="stricto" species="frenata">
<emphasis id="B97DEAB7FFB4FFE902272294FC6B5C28" box="[893,1000,1530,1554]" italics="true" pageId="16" pageNumber="1662">L. frenata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFB4FFE902A82294FB005C28" box="[1010,1155,1530,1554]" pageId="16" pageNumber="1662" sensu="stricto">sensu stricto</taxonomicNameLabel>
from northern 
<collectingCountry id="F31E7635FFB4FFE900132173FE315C0F" box="[329,434,1565,1589]" name="Myanmar" pageId="16" pageNumber="1662">Myanmar</collectingCountry>
and 
<taxonomicName id="4C094D26FFB4FFE900B42173FD705C0F" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[494,755,1565,1589]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFB4FFE900B42173FD705C0F" box="[494,755,1565,1589]" italics="true" pageId="16" pageNumber="1662">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFB4FFE903A12173FCD05C0F" box="[763,851,1565,1589]" pageId="16" pageNumber="1662" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="B97DEAB7FFB4FFE902392173FCF25C0F" box="[867,881,1565,1589]" italics="true" pageId="16" pageNumber="1662">p</emphasis>
= 15.3 
<emphasis id="B97DEAB7FFB4FFE9029A2172FC4E5C0E" box="[960,973,1564,1588]" italics="true" pageId="16" pageNumber="1662">–</emphasis>
15.6%)ı which is exceeding the species-level of divergence in colubrid snakes.
</paragraph>
<paragraph id="8BB636A5FFB4FFE8019A210CFD705C4C" blockId="16.[160,1156,315,1727]" lastBlockId="17.[160,1157,1214,1724]" lastPageId="17" lastPageNumber="1663" pageId="16" pageNumber="1662">
<taxonomicName id="4C094D26FFB4FFE9019A210CFE4A5C40" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[192,457,1634,1658]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFB4FFE9019A210CFE4A5C40" box="[192,457,1634,1658]" italics="true" pageId="16" pageNumber="1662">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFB4FFE9008F210CFDB15C40" box="[469,562,1634,1658]" pageId="16" pageNumber="1662" rank="species">sp. nov.</taxonomicNameLabel>
differs from 
<taxonomicName id="4C094D26FFB4FFE9038B210CFCDF5C40" baseAuthorityName="Boettgeri" baseAuthorityYear="1897" box="[721,860,1634,1658]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="1662" phylum="Chordata" rank="species" species="philippina">
<emphasis id="B97DEAB7FFB4FFE9038B210CFCDF5C40" box="[721,860,1634,1658]" italics="true" pageId="16" pageNumber="1662">L. philippina</emphasis>
</taxonomicName>
by having regularly one loreal (vs. missing); 7 supralabialsı 3rd and 4th in contact with the eye (vs. 8 supralabialsı fourth and fifth in contact with the eye); slight lower number of ventral scales (126 
<emphasis id="B97DEAB7FFB4FFE9051721C9FBD95C85" box="[1101,1114,1703,1727]" italics="true" pageId="16" pageNumber="1662">–</emphasis>
138 vs. 139 
<emphasis id="B97DEAB7FFB5FFE801B423D0FF785EEC" box="[238,251,1214,1238]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
150)ı lower number of subcaudals (67 
<emphasis id="B97DEAB7FFB5FFE803FF23D0FD315EEC" box="[677,690,1214,1238]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
73 vs. 110 
<emphasis id="B97DEAB7FFB5FFE8027E23D0FCB25EEC" box="[804,817,1214,1238]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
119)ı dorsal scales in 15-15-15 rows (vs. 15-15-13 rows)ı postocular stripe is thin (vs. four longitudinal brown stripes begin on neck and continue along body). 
<taxonomicName id="4C094D26FFB5FFE803D6226AFC115F26" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[652,914,1284,1308]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="1663" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFB5FFE803D6226AFC115F26" box="[652,914,1284,1308]" italics="true" pageId="17" pageNumber="1663">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFB5FFE802C6226AFC755F26" box="[924,1014,1284,1308]" pageId="17" pageNumber="1663" rank="species">sp. nov.</taxonomicNameLabel>
differs from 
<taxonomicName id="4C094D26FFB5FFE801FA2248FF715F04" baseAuthorityName="Guntheri" baseAuthorityYear="1860" box="[160,242,1318,1342]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="1663" phylum="Chordata" rank="species" species="rappi">
<emphasis id="B97DEAB7FFB5FFE801FA2248FF715F04" box="[160,242,1318,1342]" italics="true" pageId="17" pageNumber="1663">L. rappi</emphasis>
</taxonomicName>
by having one nasal (vs. two nasals)ı temporals 1 + 2 (vs. 1 + 1)ı higher number of supralabials (7 vs. 6)ı lower number of ventral scales (126 
<emphasis id="B97DEAB7FFB5FFE8023A2227FCEE5F5B" box="[864,877,1353,1377]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
138 vs. 176 
<emphasis id="B97DEAB7FFB5FFE802AB2227FC7D5F5B" box="[1009,1022,1353,1377]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
195)ı a thin postocular stripe (vs. none); 
<taxonomicName id="4C094D26FFB5FFE800B92202FD6B5FBE" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[483,744,1388,1412]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="1663" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFB5FFE800B92202FD6B5FBE" box="[483,744,1388,1412]" italics="true" pageId="17" pageNumber="1663">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFB5FFE803AA2202FCCB5FBE" box="[752,840,1388,1412]" pageId="17" pageNumber="1663" rank="species">sp. nov.</taxonomicNameLabel>
differs from 
<taxonomicName id="4C094D26FFB5FFE802862202FBDC5FBE" baseAuthorityName="Sclateri" baseAuthorityYear="1891" box="[988,1119,1388,1412]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="1663" phylum="Chordata" rank="species" species="stoliczkae">
<emphasis id="B97DEAB7FFB5FFE802862202FBDC5FBE" box="[988,1119,1388,1412]" italics="true" pageId="17" pageNumber="1663">L. stoliczkae</emphasis>
</taxonomicName>
by having 7 supralabialsı 3rd and 4th in contact with the eye (vs. 8 supralabialsı fourth and fifth in contact with the eye); lower number of ventral scales (126 
<emphasis id="B97DEAB7FFB5FFE802F622DFFC3A5FF3" box="[940,953,1457,1481]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
138 vs. 148 
<emphasis id="B97DEAB7FFB5FFE8051A22DFFBCE5FF3" box="[1088,1101,1457,1481]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
154); lower number of subcaudals (67 
<emphasis id="B97DEAB7FFB5FFE8035422BAFD985FD6" box="[526,539,1492,1516]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
73 vs. 116 
<emphasis id="B97DEAB7FFB5FFE803D722BAFD195FD6" box="[653,666,1492,1516]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
134); dorsal scales in 15-15-15 rows (vs. 15- 15-13 rows); 
<taxonomicName id="4C094D26FFB5FFE8006F2298FDB85C34" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[309,571,1526,1550]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="1663" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFB5FFE8006F2298FDB85C34" box="[309,571,1526,1550]" italics="true" pageId="17" pageNumber="1663">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFB5FFE8031E2298FD1D5C34" box="[580,670,1526,1550]" pageId="17" pageNumber="1663" rank="species">sp. nov.</taxonomicNameLabel>
differs from 
<taxonomicName id="4C094D26FFB5FFE8026E2298FC1F5C34" baseAuthorityName="Schlegeli" baseAuthorityYear="1837" box="[820,924,1526,1550]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="1663" phylum="Chordata" rank="species" species="tricolor">
<emphasis id="B97DEAB7FFB5FFE8026E2298FC1F5C34" box="[820,924,1526,1550]" italics="true" pageId="17" pageNumber="1663">L. tricolor</emphasis>
</taxonomicName>
by having regularly one loreal (vs. missing); slight lower number of ventral scales (126 
<emphasis id="B97DEAB7FFB5FFE802F72177FC395C0B" box="[941,954,1561,1585]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
138 vs. 140 
<emphasis id="B97DEAB7FFB5FFE8051A2177FBCE5C0B" box="[1088,1101,1561,1585]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
187)ı lower number of subcaudals (67 
<emphasis id="B97DEAB7FFB5FFE803542152FD985C6E" box="[526,539,1596,1620]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
73 vs. 103 
<emphasis id="B97DEAB7FFB5FFE803D72152FD195C6E" box="[653,666,1596,1620]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
137)ı dorsal scales in 15-15-15 rows (vs. 15- 15-13 rows)ı postocular stripe is thin (vs. very thick).
</paragraph>
<caption id="DF76662DFFB5FFE801FA237BFCF35EBE" ID-DOI="http://doi.org/10.5281/zenodo.3670321" ID-Zenodo-Dep="3670321" httpUri="https://zenodo.org/record/3670321/files/figure.png" pageId="17" pageNumber="1663" startId="17.[160,225,1045,1068]" targetBox="[164,1150,143,1012]" targetPageId="17">
<paragraph id="8BB636A5FFB5FFE801FA237BFCF35EBE" blockId="17.[160,1155,1044,1156]" pageId="17" pageNumber="1663">
<emphasis id="B97DEAB7FFB5FFE801FA237BFE805E16" bold="true" box="[160,259,1045,1068]" pageId="17" pageNumber="1663">Figure 7.</emphasis>
Morphology of 
<taxonomicName id="4C094D26FFB5FFE800E5237AFC9F5E16" authority="Gunther, 1858" authorityName="Guntheri" authorityYear="1858" box="[447,796,1044,1068]" class="Squamata" family="Colubridae" genus="Cyclophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="17" pageNumber="1663" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="B97DEAB7FFB5FFE800E5237AFDF45E11" box="[447,631,1044,1067]" italics="true" pageId="17" pageNumber="1663">Cyclophis frenatus</emphasis>
<bibRefCitation id="EF984B54FFB5FFE803DD237BFC9F5E16" author="Gunther A" box="[647,796,1045,1068]" pageId="17" pageNumber="1663" refId="ref12855" refString="Gunther A. 1858. Catalogue of Colubrine snakes of the British Museum. London; p. 281." type="journal volume" year="1858">Günther, 1858</bibRefCitation>
</taxonomicName>
holotype in preservative (BMNH 1946.1.1.72, male; from 
<emphasis id="B97DEAB7FFB5FFE800C9235CFE1B5E73" box="[403,408,1074,1097]" italics="true" pageId="17" pageNumber="1663">‘</emphasis>
Mountains near Affghanistan 
<emphasis id="B97DEAB7FFB5FFE803E3235CFD3D5E73" box="[697,702,1074,1097]" italics="true" pageId="17" pageNumber="1663">’</emphasis>
[= Afghanistan], corrected to 
<emphasis id="B97DEAB7FFB5FFE802A9235CFC7B5E73" box="[1011,1016,1074,1097]" italics="true" pageId="17" pageNumber="1663">‘</emphasis>
Khasi Hills 
<emphasis id="B97DEAB7FFB5FFE80507235CFBE15E73" box="[1117,1122,1074,1097]" italics="true" pageId="17" pageNumber="1663">’</emphasis>
by 
<bibRefCitation id="EF984B54FFB5FFE801FA233EFEBD5E5D" author="Boulenger GA" box="[160,318,1104,1127]" pageId="17" pageNumber="1663" pagination="382" refId="ref12323" refString="Boulenger GA. 1894. Catalogue of the snakes in the British Museum (Natural History). London: Br Mus; Vol. 2. p. 382." type="journal article" year="1894">Boulenger 1894</bibRefCitation>
). A 
<emphasis id="B97DEAB7FFB5FFE800312321FEFB5E5C" box="[363,376,1103,1126]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
general dorsal view; B 
<emphasis id="B97DEAB7FFB5FFE803302321FDF45E5C" box="[618,631,1103,1126]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
general ventral view; C 
<emphasis id="B97DEAB7FFB5FFE802282321FCFC5E5C" box="[882,895,1103,1126]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
head in dorsal view; D 
<emphasis id="B97DEAB7FFB5FFE8052C2321FB005E5C" box="[1142,1155,1103,1126]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
head in ventral view; E 
<emphasis id="B97DEAB7FFB5FFE800CA2303FE1E5EBE" box="[400,413,1133,1156]" italics="true" pageId="17" pageNumber="1663">–</emphasis>
head in lateral view. Photos by Gernot Vogel.
</paragraph>
</caption>
<paragraph id="8BB636A5FFB5FFEB019A21EFFF6B5FF7" blockId="17.[160,1157,1214,1724]" lastBlockId="18.[160,1157,940,1485]" lastPageId="18" lastPageNumber="1664" pageId="17" pageNumber="1663">
<taxonomicName id="4C094D26FFB5FFE8019A21EFFE455CA3" authority="ı Tan Van Nguyen &amp; Vogel, 2019" authorityName="ı Tan Van Nguyen &amp; Vogel" authorityYear="2019" box="[192,454,1665,1689]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="1663" phylum="Chordata" rank="species" species="pallidonuchalis" status="sp. nov.">
<emphasis id="B97DEAB7FFB5FFE8019A21EFFE455CA3" box="[192,454,1665,1689]" italics="true" pageId="17" pageNumber="1663">Liopeltis pallidonuchalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CCFFB5FFE8009521EFFDAB5CA3" box="[463,552,1665,1689]" pageId="17" pageNumber="1663" rank="species">sp. nov.</taxonomicNameLabel>
differs from members of the genus 
<taxonomicName id="4C094D26FFB5FFE8029721EFFBE05CA3" authorityName="Fitzingeri" authorityYear="1843" box="[973,1123,1665,1689]" class="Reptilia" family="Colubridae" genus="Gongylosoma" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="1663" phylum="Chordata" rank="genus">
<emphasis id="B97DEAB7FFB5FFE8029721EFFBE05CA3" box="[973,1123,1665,1689]" italics="true" pageId="17" pageNumber="1663">Gongylosoma</emphasis>
</taxonomicName>
as follows: 
<taxonomicName id="4C094D26FFB5FFE801A421CAFE1D5C86" baseAuthorityName="Boiei" baseAuthorityYear="1827" box="[254,414,1700,1724]" class="Reptilia" family="Colubridae" genus="Gongylosoma" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="1663" phylum="Chordata" rank="species" species="baliodeirum">
<emphasis id="B97DEAB7FFB5FFE801A421CAFE1D5C86" box="[254,414,1700,1724]" italics="true" pageId="17" pageNumber="1663">G. baliodeirum</emphasis>
</taxonomicName>
by having one nasal (vs. two)ı dorsal scales in 15-15-15 rows (vs. 13-13-13 rows)ı 7 supralabialsı third and fourth in contact with the eye (vs. 6 
<emphasis id="B97DEAB7FFB6FFEB054224C2FBA659FE" box="[1048,1061,940,964]" italics="true" pageId="18" pageNumber="1664">–</emphasis>
7 supralabialsı fourth and fifth in contact with the eye)ı a thin postocularstripe (vs. none); 
<taxonomicName id="4C094D26FFB6FFEB01FA249CFE755E30" authority="(Petersi 1871)" baseAuthorityName="Petersi" baseAuthorityYear="1871" box="[160,502,1010,1034]" class="Reptilia" family="Colubridae" genus="Gongylosoma" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="1664" phylum="Chordata" rank="species" species="longicaudum">
<emphasis id="B97DEAB7FFB6FFEB01FA249CFED35E30" box="[160,336,1010,1034]" italics="true" pageId="18" pageNumber="1664">G. longicaudum</emphasis>
(
<bibRefCitation id="EF984B54FFB6FFEB003A249CFE6D5E30" box="[352,494,1010,1034]" pageId="18" pageNumber="1664" refId="ref14059" type="journal article">Petersı 1871</bibRefCitation>
)
</taxonomicName>
by having dorsal scales in 15-15-15 rows (vs. 13-13-13)ı lower number of subcaudals (67 
<emphasis id="B97DEAB7FFB6FFEB0346237AFDAA5E16" box="[540,553,1044,1068]" italics="true" pageId="18" pageNumber="1664">–</emphasis>
73 vs. 91 
<emphasis id="B97DEAB7FFB6FFEB03CE237AFD225E16" box="[660,673,1044,1068]" italics="true" pageId="18" pageNumber="1664">–</emphasis>
120)ı postocular streak thin and dark (vs. light)ı no stripes at least anteriorly (vs. stripes present); 
<taxonomicName id="4C094D26FFB6FFEB02692359FEA85E48" authority="Grismeri Das &amp; Leongi 2003" authorityName="Grismeri Das &amp; Leongi" authorityYear="2003" class="Reptilia" family="Colubridae" genus="Gongylosoma" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="1664" phylum="Chordata" rank="species" species="mukutense">
<emphasis id="B97DEAB7FFB6FFEB02692359FC495E75" box="[819,970,1079,1103]" italics="true" pageId="18" pageNumber="1664">G. mukutense</emphasis>
Grismerı Das &amp; Leongı 2003
</taxonomicName>
by having one nasal (vs. two nasals)ı dorsal scales in 15-15-15 rows (vs. 13- 13-13 rows)ı temporals 1 + 2 (vs. 1 + 1)ı lower number of subcaudals (67 
<emphasis id="B97DEAB7FFB6FFEB055A2312FB8E5EAE" box="[1024,1037,1148,1172]" italics="true" pageId="18" pageNumber="1664">–</emphasis>
73 vs. 99)ı postocular streak thin and dark (vs. light)ı no stripes at least anteriorly (vs. stripes present); 
<taxonomicName id="4C094D26FFB6FFEB005423ACFD035EE0" authority="(Stoliczkai 1870)" baseAuthorityName="Stoliczkai" baseAuthorityYear="1870" box="[270,640,1218,1242]" class="Reptilia" family="Colubridae" genus="Gongylosoma" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="1664" phylum="Chordata" rank="species" species="nicobariense">
<emphasis id="B97DEAB7FFB6FFEB005423ACFE3A5EE0" box="[270,441,1218,1242]" italics="true" pageId="18" pageNumber="1664">G. nicobariense</emphasis>
(
<bibRefCitation id="EF984B54FFB6FFEB009623ACFDFA5EE0" box="[460,633,1218,1242]" pageId="18" pageNumber="1664" refId="ref14445" type="journal article">Stoliczkaı 1870</bibRefCitation>
)
</taxonomicName>
one nasal (vs. two nasals)ı dorsal scales in 15-15-15 rows (vs. 17 
<emphasis id="B97DEAB7FFB6FFEB00D5238AFE1F5EC6" box="[399,412,1252,1276]" italics="true" pageId="18" pageNumber="1664">–</emphasis>
17-17 rows)ı regularly one loreal (vs. missing)ı temporals 1 + 2 (vs. 2 + 2)ı lower number of ventrals (126 
<emphasis id="B97DEAB7FFB6FFEB03132269FDD55F25" box="[585,598,1287,1311]" italics="true" pageId="18" pageNumber="1664">–</emphasis>
138 vs. 192)ı lower number of subcaudals (67 
<emphasis id="B97DEAB7FFB6FFEB05012269FBEB5F25" box="[1115,1128,1287,1311]" italics="true" pageId="18" pageNumber="1664">–</emphasis>
73 vs. 84)ı postocular streak thin and dark (vs. light); 
<taxonomicName id="4C094D26FFB6FFEB03942244FB8E5F78" authority="(Theobaldi 1868)" baseAuthorityName="Theobaldi" baseAuthorityYear="1868" box="[718,1037,1322,1346]" class="Reptilia" family="Colubridae" genus="Gongylosoma" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="1664" phylum="Chordata" rank="species" species="scriptum">
<emphasis id="B97DEAB7FFB6FFEB03942244FCCA5F78" box="[718,841,1322,1346]" italics="true" pageId="18" pageNumber="1664">G. scriptum</emphasis>
(
<bibRefCitation id="EF984B54FFB6FFEB020C2244FB865F78" box="[854,1029,1322,1346]" pageId="18" pageNumber="1664" refId="ref14712" type="journal article">Theobaldı 1868</bibRefCitation>
)
</taxonomicName>
by having one nasal (vs. two)ı 7 supralabials third and fourth in contact with the eye (vs. 8 supralabialsı 3rd and 5th in contact with the eye)ı lower number of subcaudals (67 
<emphasis id="B97DEAB7FFB6FFEB05012201FBEB5FBD" box="[1115,1128,1391,1415]" italics="true" pageId="18" pageNumber="1664">–</emphasis>
73 vs. 87 
<emphasis id="B97DEAB7FFB6FFEB01BB22FCFF6D5F90" box="[225,238,1426,1450]" italics="true" pageId="18" pageNumber="1664">–</emphasis>
98)ı dorsal scales in 15-15-15 rows (vs. 13-13-13 rows)ı postocular streak thin (vs. none).
</paragraph>
</subSubSection>
</treatment>
</document>