339 lines
37 KiB
XML
339 lines
37 KiB
XML
<document id="2FEFA131DD4FFF8721D118DE10568118" ID="10.11646/zootaxa.4243.1.7SLASH10478" ID-CLB-Dataset="36283" ID-DOI="10.11646/zootaxa.4243.1.7" ID-GBIF-Dataset="1facafb7-179b-4200-ac2f-9d00375a7ce4" ID-ISSN="1175-5326" ID-Zenodo-Dep="398752" ID-ZooBank="96C38B34-BD4C-438B-BC73-FAD6CCBDE212" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1489483589214" checkinUser="plazi" docAuthor="Schädel, Mario & Lechner, Thomas S." docDate="2017" docId="03A087A4FFB8FFB5FF423295FBF811DB" docLanguage="en" docName="zootaxa.4243.1.7SLASH10478.pdf" docOrigin="Zootaxa 4243 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ictinogomphus hassleri Schädel & Lechner, 2017, sp. nov." docType="treatment" docVersion="8" lastPageNumber="159" masterDocId="FF99FFDCFFBCFFB3FFD5316EFFDB1533" masterDocTitle="Two new dragonflies (Odonata: Anisoptera) from the Miocene of Carinthia (Austria)" masterLastPageNumber="164" masterPageNumber="153" pageNumber="157" updateTime="1698437928032" updateUser="ExternalLinkService">
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<mods:titleInfo id="1938C12C3FE8FC0EC17E331FF27B93B4">
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<mods:title id="E8F0113CCB91E9703C12A605E05CE05D">Two new dragonflies (Odonata: Anisoptera) from the Miocene of Carinthia (Austria)</mods:title>
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<mods:namePart id="B4A19642E17CF9AF4073365AA516B5F0">Schädel, Mario</mods:namePart>
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<mods:name id="D48F70EC99B5E9B7EB818F724DA95596" type="personal">
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<mods:namePart id="3462E0A74E3D51F83174458419C6EA48">Lechner, Thomas S.</mods:namePart>
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<mods:typeOfResource id="2858C597DBE5583C11AE6631B527147B">text</mods:typeOfResource>
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<mods:title id="959A509D957690ED3339049E06C7F1AB">Zootaxa</mods:title>
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<mods:part id="CF496734FEDAF633628DA8B45ADE4673">
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<mods:date id="FCC7A1B14078D87E328636042EDC85D5">2017</mods:date>
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<mods:number id="B583A26119190D9F2D2A211E6E4E282C">4243</mods:number>
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<mods:identifier id="B0200BBEDD75C23E90EA70CD7C4EC32C" type="DOI">10.11646/zootaxa.4243.1.7</mods:identifier>
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<treatment id="03A087A4FFB8FFB5FF423295FBF811DB" ID-DOI="http://doi.org/10.5281/zenodo.6033073" ID-GBIF-Taxon="127843942" ID-Zenodo-Dep="6033073" LSID="urn:lsid:plazi:treatment:03A087A4FFB8FFB5FF423295FBF811DB" httpUri="http://treatment.plazi.org/id/03A087A4FFB8FFB5FF423295FBF811DB" lastPageId="6" lastPageNumber="159" pageId="4" pageNumber="157">
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<subSubSection id="C3136539FFB8FFB7FF423295FEE91104" pageId="4" pageNumber="157" type="nomenclature">
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<paragraph id="8BB636B2FFB8FFB7FF423295FDC31125" blockId="4.[151,536,1019,1079]" box="[151,536,1019,1046]" pageId="4" pageNumber="157">
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<heading id="D0FE81DEFFB8FFB7FF423295FDC31125" bold="true" box="[151,536,1019,1046]" fontSize="11" level="1" pageId="4" pageNumber="157" reason="1">
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<emphasis id="B97DEAA0FFB8FFB7FF423295FDC31125" bold="true" box="[151,536,1019,1046]" pageId="4" pageNumber="157">
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<taxonomicName id="4C094D31FFB8FFB7FF423295FE681126" box="[151,435,1019,1045]" class="Insecta" family="Gomphidae" genus="Ictinogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="157" phylum="Arthropoda" rank="species" species="hassleri" status="sp. nov.">
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<emphasis id="B97DEAA0FFB8FFB7FF423295FE681126" bold="true" box="[151,435,1019,1045]" italics="true" pageId="4" pageNumber="157">Ictinogomphus hassleri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24E57DBFFB8FFB7FE6F3292FDC31125" box="[442,536,1020,1046]" pageId="4" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB636B2FFB8FFB7FF423570FEE91104" blockId="4.[151,536,1019,1079]" box="[151,306,1054,1079]" pageId="4" pageNumber="157">
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<figureCitation id="13322A37FFB8FFB7FF423570FEDB1104" box="[151,256,1054,1079]" captionStart="FIGURE 1" captionStartId="5.[151,250,1282,1304]" captionTargetBox="[151,1436,625,1261]" captionTargetId="figure@5.[151,1436,625,1261]" captionTargetPageId="5" captionText="FIGURE 1. Photograph of the hind wing fragment of Ictinogomphus hassleri sp. nov., holotype LMK-Pal- 2016 - 5538." httpUri="https://zenodo.org/record/398753/files/figure.png" pageId="4" pageNumber="157">Figures 1</figureCitation>
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&
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<figureCitation id="13322A37FFB8FFB7FEF13571FEE91104" box="[292,306,1055,1079]" captionStart="FIGURE 2" captionStartId="5.[151,250,1934,1956]" captionTargetBox="[151,1436,1354,1913]" captionTargetId="figure@5.[151,1436,1354,1913]" captionTargetPageId="5" captionText="FIGURE 2. Drawing of the hind wing fragment of Ictinogomphus hassleri sp. nov., holotype LMK-Pal- 2016 - 5538, with terminology of wing venation." httpUri="https://zenodo.org/record/398754/files/figure.png" pageId="4" pageNumber="157">2</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3136539FFB8FFB7FF423508FE0E11F4" pageId="4" pageNumber="157" type="materials_examined">
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<paragraph id="8BB636B2FFB8FFB7FF423508FE0E11F4" blockId="4.[151,1436,1126,2015]" pageId="4" pageNumber="157">
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<emphasis id="B97DEAA0FFB8FFB7FF423508FED0114C" bold="true" box="[151,267,1126,1151]" pageId="4" pageNumber="157">Holotype.</emphasis>
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<materialsCitation id="3B613CEFFFB8FFB7FEC73508FE0E11F4" ID-GBIF-Occurrence="1439650064" country="Austria" location="State Museum" municipality="Klagenfurt" pageId="4" pageNumber="157" specimenCount="1" stateProvince="Karnten" typeStatus="holotype">
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The
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<typeStatus id="54B28810FFB8FFB7FE903508FE7C114C" box="[325,423,1126,1151]" pageId="4" pageNumber="157" type="holotype">holotype</typeStatus>
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is stored in the private collection of Dr. Andreas Hassler. The deposition of the specimen is projected in the
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<location id="8ED66069FFB8FFB7FEA535E5FDC81191" LSID="urn:lsid:plazi:treatment:03A087A4FFB8FFB5FF423295FBF811DB:8ED66069FFB8FFB7FEA535E5FDC81191" box="[368,531,1163,1187]" country="Austria" municipality="Klagenfurt" name="State Museum" pageId="4" pageNumber="157" stateProvince="Karnten">State Museum</location>
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of
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<collectingRegion id="49CDF850FFB8FFB7FDE835E5FD411191" box="[573,666,1163,1186]" country="Austria" name="Karnten" pageId="4" pageNumber="157">Kärnten</collectingRegion>
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,
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<collectingMunicipality id="6BD2ACC8FFB8FFB7FD7D35E4FCFF1190" box="[680,804,1162,1187]" pageId="4" pageNumber="157">Klagenfurt</collectingMunicipality>
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,
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<collectingCountry id="F31E7622FFB8FFB7FCE735E4FC5E1190" box="[818,901,1162,1187]" name="Austria" pageId="4" pageNumber="157">Austria</collectingCountry>
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under collection number LMK-Pal-2016-5538 (Old working-title: SIH-14).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3136539FFB8FFB7FF1235BCFA561344" pageId="4" pageNumber="157" type="diagnosis">
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<paragraph id="8BB636B2FFB8FFB7FF1235BCFB8C1049" blockId="4.[151,1436,1126,2015]" pageId="4" pageNumber="157">
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<emphasis id="B97DEAA0FFB8FFB7FF1235BCFE9A11D8" bold="true" box="[199,321,1234,1259]" pageId="4" pageNumber="157">Diagnosis.</emphasis>
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This new fossil species is very similar to modern representatives of the genera
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<taxonomicName id="4C094D31FFB8FFB7FB1035BDFAB011D8" box="[1221,1387,1235,1259]" class="Insecta" family="Gomphidae" genus="Ictinogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="157" phylum="Arthropoda" rank="genus">
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<emphasis id="B97DEAA0FFB8FFB7FB1035BDFAB011D8" box="[1221,1387,1235,1259]" italics="true" pageId="4" pageNumber="157">Ictinogomphus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C094D31FFB8FFB7FF423598FE21103C" authority="Fraser, 1939" authorityName="Fraser" authorityYear="1939" box="[151,506,1270,1295]" class="Insecta" family="Gomphidae" genus="Sinictinogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="157" phylum="Arthropoda" rank="genus">
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<emphasis id="B97DEAA0FFB8FFB7FF423598FEBB103D" box="[151,352,1270,1294]" italics="true" pageId="4" pageNumber="157">Sinictinogomphus</emphasis>
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<bibRefCitation id="EF984B43FFB8FFB7FEBC3599FE21103C" author="Fraser" box="[361,506,1271,1295]" pageId="4" pageNumber="157" refString="Fraser, F. C. (1939) A note on the generic characters of Ictinogomphus Cowley (Odonata). Proceedings of the Royal Entomological Society. Series B, Taxonomy, 8 (2), 21 - 24." type="journal article" year="1939">Fraser, 1939</bibRefCitation>
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</taxonomicName>
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. It differs from other
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<taxonomicName id="4C094D31FFB8FFB7FD2F3598FC5B103C" box="[762,896,1270,1295]" class="Insecta" family="Gomphidae" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="157" phylum="Arthropoda" rank="subFamily" subFamily="Lindeniinae">Lindeniinae</taxonomicName>
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genera in the shape of the triangulum and the position of the Tspl-origin. The unique compilation of character states may not separate the fossil from all members of
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<taxonomicName id="4C094D31FFB8FFB7FF603450FE801065" box="[181,347,1342,1366]" class="Insecta" family="Gomphidae" genus="Ictinogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="157" phylum="Arthropoda" rank="genus">
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<emphasis id="B97DEAA0FFB8FFB7FF603450FE801065" box="[181,347,1342,1366]" italics="true" pageId="4" pageNumber="157">Ictinogomphus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C094D31FFB8FFB7FE463450FD871065" box="[403,604,1342,1366]" class="Insecta" family="Gomphidae" genus="Sinictinogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="157" phylum="Arthropoda" rank="genus">
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<emphasis id="B97DEAA0FFB8FFB7FE463450FD871065" box="[403,604,1342,1366]" italics="true" pageId="4" pageNumber="157">Sinictinogomphus</emphasis>
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</taxonomicName>
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, but the remoteness in time and geographical range strongly suggests that the described specimen is not conspecific in the sense of a biological species concept.
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</paragraph>
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<paragraph id="8BB636B2FFB8FFB7FF1234E8FCC91338" blockId="4.[151,1436,1126,2015]" pageId="4" pageNumber="157">
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The
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<taxonomicName id="4C094D31FFB8FFB7FF2934E8FE5910AC" box="[252,386,1414,1439]" class="Insecta" family="Gomphidae" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="157" phylum="Arthropoda" rank="subFamily" subFamily="Lindeniinae">Lindeniinae</taxonomicName>
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fossil described by
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<bibRefCitation id="EF984B43FFB8FFB7FDBF34E9FC8110AC" author="Huang" box="[618,858,1414,1439]" pageId="4" pageNumber="157" refString="Huang, D. Y. & Nel, A. (2009) First fossil record of a Lindeniidae from the Miocene Shanwang Formation of China (Odonata, Anisoptera). Bulletin de la Societe entomologique de France, 114 (4), 441 - 443." type="journal article" year="2009">Huang & Nel (2009)</bibRefCitation>
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from the Miocene of
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<collectingCountry id="F31E7622FFB8FFB7FB8C34E8FB4710AC" box="[1113,1180,1414,1439]" name="China" pageId="4" pageNumber="157">China</collectingCountry>
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has a distinctly larger hind wing than our specimen and 3 to 4 rows of cells between the origin of the secondary branch of IR2 and RP´´instead of only two rows as in our specimen (which is probably due to the larger size). With a length of
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<quantity id="4CF19B57FFB8FFB7FACB34A1FAB410D5" box="[1310,1391,1487,1511]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="157" unit="mm" value="40.0">40 mm</quantity>
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, the hind wing is distinctly larger than the Chinese specimen.
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</paragraph>
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<paragraph id="8BB636B2FFB8FFB7FF123779FA561344" blockId="4.[151,1436,1126,2015]" pageId="4" pageNumber="157">
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Our specimen is very similar to the specimen described by
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<bibRefCitation id="EF984B43FFB8FFB7FCBF3778FBAF131C" author="Nel" box="[874,1140,1558,1583]" pageId="4" pageNumber="157" refString="Nel, A. & Paicheler, J. C. (1994 a) Les Gomphidae fossiles. Un inventaire critique (Odonata: Gomphidae). Annales de la Societe entomologique de France, 30 (1), 55 - 77." type="journal article" year="1994">Nel & Paicheler (1994)</bibRefCitation>
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from the latest Oligocene sediments in Bes-Konak,
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<collectingCountry id="F31E7622FFB8FFB7FE6F3754FDD71360" box="[442,524,1594,1619]" name="Turkey" pageId="4" pageNumber="157">Turkey</collectingCountry>
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. The Bes-Konak specimen differs only in the manifestation of intercalated veins and the oblique pterostigmal brace vein, based on the area of the fossil wings that is preserved in both specimens.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3136539FFB8FFB7FF1237ECFF0D1234" pageId="4" pageNumber="157" type="reference_group">
|
||
<paragraph id="8BB636B2FFB8FFB7FF1237ECFF0D1234" blockId="4.[151,1436,1126,2015]" pageId="4" pageNumber="157">
|
||
The
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||
<taxonomicName id="4C094D31FFB8FFB7FF2B37EDFE7F13A8" ID-CoL="53FW" box="[254,420,1667,1691]" class="Insecta" family="Gomphidae" genus="Ictinogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="157" phylum="Arthropoda" rank="genus">
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<emphasis id="B97DEAA0FFB8FFB7FF2B37EDFE7F13A8" box="[254,420,1667,1691]" italics="true" pageId="4" pageNumber="157">Ictinogomphus</emphasis>
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</taxonomicName>
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specimen from Seifhennersdorf (Early Oligocene,
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<collectingCountry id="F31E7622FFB8FFB7FC2B37EDFBB113A8" box="[1022,1130,1667,1691]" name="Germany" pageId="4" pageNumber="157">Germany</collectingCountry>
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||
;
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||
<bibRefCitation id="EF984B43FFB8FFB7FBAF37ECFA4C13A8" author="Prokop" box="[1146,1431,1666,1691]" pageId="4" pageNumber="157" refString="Prokop, J. & Fikacek, M. (2007) An annotated list of early Oligocene insect fauna from Seifhennersdorf (Saxony, Germany). Acta Musei Nationalis Pragae, Series B Historia Naturalis, 63 (2 - 4), 209 - 217." type="journal article" year="2007">Prokop & Fikacek 2007</bibRefCitation>
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||
;
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||
<bibRefCitation id="EF984B43FFB8FFB7FF4237C8FE57138C" author="Prokop" box="[151,396,1702,1727]" pageId="4" pageNumber="157" refString="Prokop, J., Pecharova, M. & Nel, A. (2016) New Cenozoic dragonflies from the Most Basin and Stredohori Complex volcanic area (Czech Republic, Germany). Journal of Natural History, 50 (37 - 38), 2311 - 2326." type="journal article" year="2016">
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||
Prokop
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<emphasis id="B97DEAA0FFB8FFB7FF2E37C9FEE4138D" box="[251,319,1702,1726]" italics="true" pageId="4" pageNumber="157">et al.</emphasis>
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||
2016
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||
</bibRefCitation>
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||
) cannot be clearly differentiated from our herein-described specimen, because the Seifhennersdorf specimen is represented by a forewing in contrast to our specimen that is represented by a hind wing.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3136539FFB8FFB5FF12367CFDA5172B" lastPageId="6" lastPageNumber="159" pageId="4" pageNumber="157" type="description">
|
||
<paragraph id="8BB636B2FFB8FFB7FF12367CFEB31241" blockId="4.[151,1436,1126,2015]" pageId="4" pageNumber="157">
|
||
<emphasis id="B97DEAA0FFB8FFB7FF12367CFE821218" bold="true" box="[199,345,1810,1835]" pageId="4" pageNumber="157">Description.</emphasis>
|
||
An almost complete wing lacking only anal region and basalmost part of ACuf as well as area basal to discoidal triangle. Total length
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||
<quantity id="4CF19B57FFB8FFB7FD9A3658FD68127D" box="[591,691,1846,1871]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" pageId="4" pageNumber="157" unit="mm" value="32.5">32.5 mm</quantity>
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||
and total width
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||
<quantity id="4CF19B57FFB8FFB7FCB23659FC16127D" box="[871,973,1847,1871]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="4" pageNumber="157" unit="mm" value="11.1">11.1 mm</quantity>
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||
. Preservation better towards wingtip than in more basal part.
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||
</paragraph>
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||
<paragraph id="8BB636B2FFB8FFB7FF123610FEC712EC" blockId="4.[151,1436,1126,2015]" pageId="4" pageNumber="157">
|
||
Anterior wing margin from base to nodus (CA & CP & ScA´) with short spines; nodus
|
||
<quantity id="4CF19B57FFB8FFB7FB633611FAC712A5" box="[1206,1308,1919,1943]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="4" pageNumber="157" unit="mm" value="14.1">14.1 mm</quantity>
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||
from wing base and
|
||
<quantity id="4CF19B57FFB8FFB7FF2B36CDFEB91289" box="[254,354,1955,1979]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="4" pageNumber="157" unit="mm" value="18.4">18.4 mm</quantity>
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||
from tip (at 43% of the wing length); secondary antenodal crossveins at least in distal part of ans not aligned.
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFB9FFB6FF1231F9FB20142E" blockId="5.[151,1436,151,573]" pageId="5" pageNumber="158">
|
||
Pterostigma with parallel anterior and posterior margin;
|
||
<quantity id="4CF19B57FFB9FFB6FCEE31F9FC4A159C" box="[827,913,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="5" pageNumber="158" unit="mm" value="5.7">5.7 mm</quantity>
|
||
in proximodistal dimension and maximum
|
||
<quantity id="4CF19B57FFB9FFB6FAAC31F6FF1815E7" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="158" unit="mm" value="0.9">0.9 mm</quantity>
|
||
in anteroposterior dimension; posterior margin of pterostigma broader than anterior margin; proximal margin orthogonal and distal margin oblique with an angle of 134° to posterior margin. Pterostigmal brace vein broad and pointing posterior at a right angle to RA +; Preradial and Interradial field both with one row of cells.
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFB9FFB6FF123049FD58149F" blockId="5.[151,1436,151,573]" pageId="5" pageNumber="158">
|
||
Distal part of antesubnodal space not free; at least one crossvein between RA and RP´
|
||
<quantity id="4CF19B57FFB9FFB6FB603046FAD6140C" box="[1205,1293,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="158" unit="mm" value="2.4">2.4 mm</quantity>
|
||
proximal of subnodus; distance between RP´–RP´´furca and subnodus
|
||
<quantity id="4CF19B57FFB9FFB6FC853022FC751457" box="[848,942,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="5" pageNumber="158" unit="mm" value="4.8">4.8 mm</quantity>
|
||
. Interradial sector with distinct secondary branch
|
||
<quantity id="4CF19B57FFB9FFB6FF39301EFE9E14B4" box="[236,325,368,392]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="5" pageNumber="158" unit="mm" value="7.8">7.8 mm</quantity>
|
||
distal from RP´–RP´´furca; two intercalated veins between IR2 + and its secondary branch and between same secondary branch and RP´´-.
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFB9FFB6FF1230D6FE82170E" blockId="5.[151,1436,151,573]" pageId="5" pageNumber="158">RP1 and RP2 parallel and with one row of cells in between just near to level of proximal margin of pterostigma; intercalated vein between RP1 and RP2 from mid-pterostigmal level to posterior wing margin dividing area between outer veins more or less centrally; numerous small cells anterior and posterior of this intercalated vein.</paragraph>
|
||
<caption id="DF76663AFFB9FFB6FF42346CFA97102B" box="[151,1356,1282,1304]" httpUri="https://zenodo.org/record/398753/files/figure.png" pageId="5" pageNumber="158" targetBox="[151,1436,625,1261]" targetPageId="5">
|
||
<paragraph id="8BB636B2FFB9FFB6FF42346CFA97102B" blockId="5.[151,1356,1282,1304]" box="[151,1356,1282,1304]" pageId="5" pageNumber="158">
|
||
<emphasis id="B97DEAA0FFB9FFB6FF42346CFEC81024" bold="true" box="[151,275,1282,1304]" pageId="5" pageNumber="158">FIGURE 1.</emphasis>
|
||
Photograph of the hind wing fragment of
|
||
<taxonomicName id="4C094D31FFB9FFB6FD6B346CFC71102B" box="[702,938,1282,1304]" class="Insecta" family="Gomphidae" genus="Ictinogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="hassleri" status="sp. nov.">
|
||
<emphasis id="B97DEAA0FFB9FFB6FD6B346CFC71102B" box="[702,938,1282,1304]" italics="true" pageId="5" pageNumber="158">Ictinogomphus hassleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97DEAA0FFB9FFB6FC64346CFBDA1024" bold="true" box="[945,1025,1282,1303]" pageId="5" pageNumber="158">
|
||
<taxonomicNameLabel id="A24E57DBFFB9FFB6FC64346CFBDA1024" box="[945,1025,1282,1303]" pageId="5" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype LMK-Pal-2016-5538.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF76663AFFB9FFB6FF4236E0FE1712F0" httpUri="https://zenodo.org/record/398754/files/figure.png" pageId="5" pageNumber="158" targetBox="[151,1436,1354,1913]" targetPageId="5">
|
||
<paragraph id="8BB636B2FFB9FFB6FF4236E0FE1712F0" blockId="5.[151,1435,1934,1987]" pageId="5" pageNumber="158">
|
||
<emphasis id="B97DEAA0FFB9FFB6FF4236E0FECD1297" bold="true" box="[151,278,1934,1956]" pageId="5" pageNumber="158">FIGURE 2.</emphasis>
|
||
Drawing of the hind wing fragment of
|
||
<taxonomicName id="4C094D31FFB9FFB6FD1136E0FC6E1297" box="[708,949,1934,1956]" class="Insecta" family="Gomphidae" genus="Ictinogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="hassleri" status="sp. nov.">
|
||
<emphasis id="B97DEAA0FFB9FFB6FD1136E0FC6E1297" box="[708,949,1934,1956]" italics="true" pageId="5" pageNumber="158">Ictinogomphus hassleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97DEAA0FFB9FFB6FC6A36E1FBC81297" bold="true" box="[959,1043,1935,1956]" pageId="5" pageNumber="158">
|
||
<taxonomicNameLabel id="A24E57DBFFB9FFB6FC6A36E1FBC81297" box="[959,1043,1935,1956]" pageId="5" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype LMK-Pal-2016-5538, with terminology of wing venation.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB636B2FFBAFFB5FF1231F9FCE415C4" blockId="6.[151,1437,151,1257]" pageId="6" pageNumber="159">
|
||
Distance between arculus and RP furca (origin of RP´& RP´´)
|
||
<quantity id="4CF19B57FFBAFFB5FC4631F9FC36159C" box="[915,1005,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="6" pageNumber="159" unit="mm" value="6.3">6.3 mm</quantity>
|
||
; premedial field with one row of cells and with more or less parallel boundaries except for the very posterodistal part; posterodistal part of premedial field with two rows of cells and broader than anteroproximal part.
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFBAFFB5FF12306AFCBA142F" blockId="6.[151,1437,151,1257]" box="[199,865,260,285]" pageId="6" pageNumber="159">At least five antefurcal crossveins between RP - and MA +.</paragraph>
|
||
<paragraph id="8BB636B2FFBAFFB5FF123046FBA21473" blockId="6.[151,1437,151,1257]" box="[199,1145,295,320]" pageId="6" pageNumber="159">
|
||
Htc
|
||
<quantity id="4CF19B57FFBAFFB5FF233046FE97140C" box="[246,332,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="6" pageNumber="159" unit="mm" value="3.9">3.9 mm</quantity>
|
||
long and
|
||
<quantity id="4CF19B57FFBAFFB5FE6E3046FDCA140C" box="[443,529,296,320]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="159" unit="mm" value="0.8">0.8 mm</quantity>
|
||
wide; anterior margin of htc (MA +) distinctly curved.
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFBAFFB5FF123022FD1F14E3" blockId="6.[151,1437,151,1257]" pageId="6" pageNumber="159">
|
||
Discoidal triangle longitudinally stretched and divided by at least two secondary crossveins; basal side
|
||
<quantity id="4CF19B57FFBAFFB5FA923022FA461457" box="[1351,1437,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="159" unit="mm" value="1.5">1.5 mm</quantity>
|
||
long; costal side
|
||
<quantity id="4CF19B57FFBAFFB5FE8C301EFE6814B4" box="[345,435,368,392]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="159" unit="mm" value="3.2">3.2 mm</quantity>
|
||
; posterodistal side
|
||
<quantity id="4CF19B57FFBAFFB5FD5B3001FD3314B4" box="[654,744,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="159" unit="mm" value="3.6">3.6 mm</quantity>
|
||
; posterodistal side (ddv) almost straight with a slight kink at origin of the distinct Tspl. Two rows of cells in basal part of postdiscoidal space. Strong secondary branch of MA +
|
||
<quantity id="4CF19B57FFBAFFB5FF4230D9FF3614FC" box="[151,237,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="159" unit="mm" value="4.5">4.5 mm</quantity>
|
||
distal of distal angle of discoidal triangle.
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFBAFFB5FF1230B2FDA5172B" blockId="6.[151,1437,151,1257]" pageId="6" pageNumber="159">
|
||
Gaff (CuA from posterior angle of discoidal triangle to CuA’/CuA’’-branching)
|
||
<quantity id="4CF19B57FFBAFFB5FB9830B2FB7814C7" box="[1101,1187,476,501]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="159" unit="mm" value="0.6">0.6 mm</quantity>
|
||
long (not prolonged); CuA´+ with 2 distinct secondary branches.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136539FFBAFFB5FF12334AFEAF17B6" pageId="6" pageNumber="159" type="materials_examined">
|
||
<paragraph id="8BB636B2FFBAFFB5FF12334AFEAF17B6" blockId="6.[151,1437,151,1257]" pageId="6" pageNumber="159">
|
||
<materialsCitation id="3B613CEFFFBAFFB5FF12334AFEAF17B6" ID-GBIF-Occurrence="1439650055" country="Austria" county="Fine" latitude="46.798145" location="St. Andra" longLatPrecision="1" longitude="14.806603" municipality="Wolfsberg" pageId="6" pageNumber="159" specimenCount="1" stateProvince="Carinthia" typeStatus="holotype">
|
||
<emphasis id="B97DEAA0FFBAFFB5FF12334AFDC3170E" bold="true" box="[199,536,548,573]" pageId="6" pageNumber="159">
|
||
<typeStatus id="54B28810FFBAFFB5FF12334AFED9170E" box="[199,258,548,573]" pageId="6" pageNumber="159">Type</typeStatus>
|
||
locality and horizon.
|
||
</emphasis>
|
||
Claypit Schaßbach (
|
||
<geoCoordinate id="EE3D5075FFBAFFB5FCCE334AFC1D170E" box="[795,966,548,573]" direction="north" orientation="latitude" pageId="6" pageNumber="159" precision="1" value="46.798145">46°47'53.32"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3D5075FFBAFFB5FC0F334AFB59170F" box="[986,1154,548,572]" direction="east" orientation="longitude" pageId="6" pageNumber="159" precision="1" value="14.806603">14°48'23.77"E</geoCoordinate>
|
||
),
|
||
<location id="8ED66069FFBAFFB5FB4B334AFACE170E" LSID="urn:lsid:plazi:treatment:03A087A4FFB8FFB5FF423295FBF811DB:8ED66069FFBAFFB5FB4B334AFACE170E" box="[1182,1301,548,573]" country="Austria" county="Fine" latitude="46.798145" longLatPrecision="1" longitude="14.806603" municipality="Wolfsberg" name="St. Andra" pageId="6" pageNumber="159" stateProvince="Carinthia">St. Andrä</location>
|
||
,
|
||
<collectingMunicipality id="6BD2ACC8FFBAFFB5FAFD334AFA47170E" box="[1320,1436,548,573]" pageId="6" pageNumber="159">Wolfsberg</collectingMunicipality>
|
||
(district),
|
||
<collectingRegion id="49CDF850FFBAFFB5FED03329FEAB1753" box="[261,368,583,608]" country="Austria" name="Karnten" pageId="6" pageNumber="159">Carinthia</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31E7622FFBAFFB5FEAE3329FE0A1753" box="[379,465,583,608]" name="Austria" pageId="6" pageNumber="159">Austria</collectingCountry>
|
||
.
|
||
<collectingCounty id="62D74E3EFFBAFFB5FE093329FDD51753" box="[476,526,583,608]" pageId="6" pageNumber="159">Fine</collectingCounty>
|
||
laminated mudstone of the lacustrine lower part of the Mühldorf Formation. Age late Early Miocene.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136539FFBAFFB5FF1233E1FB981626" pageId="6" pageNumber="159" type="etymology">
|
||
<paragraph id="8BB636B2FFBAFFB5FF1233E1FD0117FE" blockId="6.[151,1437,151,1257]" pageId="6" pageNumber="159">
|
||
<emphasis id="B97DEAA0FFBAFFB5FF1233E1FE95179B" bold="true" box="[199,334,655,680]" pageId="6" pageNumber="159">Etymology.</emphasis>
|
||
The species name
|
||
<taxonomicName id="4C094D31FFBAFFB5FDF733FEFDA0179B" box="[546,635,656,680]" class="Insecta" family="Gomphidae" genus="Ictinogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="159" phylum="Arthropoda" rank="species" species="hassleri">
|
||
<emphasis id="B97DEAA0FFBAFFB5FDF733FEFDA0179B" box="[546,635,656,680]" italics="true" pageId="6" pageNumber="159">hassleri</emphasis>
|
||
</taxonomicName>
|
||
refers to the finder of the specimen and local specialist for the field site Dr. Andreas Hassler, St. Andrä, Wolfsberg,
|
||
<collectingCountry id="F31E7622FFBAFFB5FD5533DAFD0D17FE" box="[640,726,692,717]" name="Austria" pageId="6" pageNumber="159">Austria</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFBAFFB5FF1233B9FB981626" blockId="6.[151,1437,151,1257]" pageId="6" pageNumber="159">
|
||
<emphasis id="B97DEAA0FFBAFFB5FF1233B9FE1017C3" bold="true" box="[199,459,727,752]" pageId="6" pageNumber="159">Wingspan estimation.</emphasis>
|
||
The wingspan for the whole animal is estimated as about
|
||
<quantity id="4CF19B57FFBAFFB5FBB133B6FB6D17DC" box="[1124,1206,728,752]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="159" unit="mm" value="70.0">70 mm</quantity>
|
||
, taking into account that the forewing is a bit longer than the hind wing in recent
|
||
<taxonomicName id="4C094D31FFBAFFB5FCE83392FC381627" box="[829,995,764,788]" class="Insecta" family="Gomphidae" genus="Ictinogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="159" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97DEAA0FFBAFFB5FCE83392FC381627" box="[829,995,764,788]" italics="true" pageId="6" pageNumber="159">Ictinogomphus</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136539FFBAFFB5FF123271FD3C16B3" pageId="6" pageNumber="159" type="description">
|
||
<paragraph id="8BB636B2FFBAFFB5FF123271FB0E160B" blockId="6.[151,1437,151,1257]" box="[199,1237,799,824]" pageId="6" pageNumber="159">
|
||
<emphasis id="B97DEAA0FFBAFFB5FF123271FE72160B" bold="true" box="[199,425,799,824]" pageId="6" pageNumber="159">Sex determination.</emphasis>
|
||
The sex determination is not possible lacking the basal part of the wing.
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFBAFFB5FF12322AFD3C16B3" blockId="6.[151,1437,151,1257]" pageId="6" pageNumber="159">
|
||
<emphasis id="B97DEAA0FFBAFFB5FF12322AFE00166E" bold="true" box="[199,475,836,861]" pageId="6" pageNumber="159">Phylogenetic affinities.</emphasis>
|
||
<taxonomicName id="4C094D31FFBAFFB5FE32322AFD21166E" authority="Selys, 1854" authorityName="Selys" authorityYear="1854" box="[487,762,836,861]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Anisoptera" kingdom="Animalia" order="Malvales" pageId="6" pageNumber="159" phylum="Tracheophyta" rank="genus">Anisoptera Selys, 1854</taxonomicName>
|
||
: strong expansion of the cubito-anal field, presence of a discoidal triangle and a hypertriangle (
|
||
<bibRefCitation id="EF984B43FFBAFFB5FD933209FD0016B3" author="Bechly" box="[582,731,871,896]" pageId="6" pageNumber="159" refString="Bechly, G. (2007) Chapter 11.5 Odonata: damselflies and dragonflies. In: Martill, D. M., Bechly, G. & Loveridge, R. F. (Eds.), The Crato fossil beds of Brazil: window into an ancient world. Cambridge University Press, Cambridge, pp. 184 - 222." type="book chapter" year="2007">Bechly 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136539FFBAFFB5FF1232E2FC5F1107" pageId="6" pageNumber="159" type="reference_group">
|
||
<paragraph id="8BB636B2FFBAFFB5FF1232E2FBFF16FB" blockId="6.[151,1437,151,1257]" pageId="6" pageNumber="159">
|
||
<taxonomicName id="4C094D31FFBAFFB5FF1232E2FE201697" ID-CoL="623L5" authority="Rambur, 1842" authorityName="Rambur" authorityYear="1842" box="[199,507,908,933]" class="Insecta" family="Gomphidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="159" phylum="Arthropoda" rank="family">Gomphidae Rambur, 1842</taxonomicName>
|
||
: posterodistal side of the triangulum with a slight angle; presence of a trigonal supplement; distinctly curved anterior side of hypertriangle (
|
||
<bibRefCitation id="EF984B43FFBAFFB5FCE832C1FC0A16FB" author="Bechly" box="[829,977,943,968]" pageId="6" pageNumber="159" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata) unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der * Odonata. Petalura, special Volume 2, 1 - 402." type="journal article" year="1996">Bechly 1996</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF984B43FFBAFFB5FC0932DEFBC216FB" author="Bechly" box="[988,1049,944,968]" pageId="6" pageNumber="159" refString="Bechly, G. (2003) Phylogenetic systematics of Odonata. In: Schorr, M. & Lindeboom, M. (Eds.), Dragonfly Research, Folume 1. Zerf-Tubingen (CD-ROM)." type="book chapter" year="2003">2003</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFBAFFB5FF1232BAFC5F1107" blockId="6.[151,1437,151,1257]" pageId="6" pageNumber="159">
|
||
<taxonomicName id="4C094D31FFBAFFB5FF1232BAFE9616DE" box="[199,333,980,1005]" class="Insecta" family="Gomphidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="159" phylum="Arthropoda" rank="subFamily" subFamily="Lindeniinae">Lindeniinae</taxonomicName>
|
||
Jacobson & Bianchi, 1905: IR2 and MA both with a distinct secondary branch; hind wing discoidal triangle elongated in distal direction; distal side of triangulum sigmoidal shaped with a slight kink at origin of Tspl; triangulum with more than two cells (
|
||
<bibRefCitation id="EF984B43FFBAFFB5FD363572FCA31107" author="Bechly" box="[739,888,1052,1077]" pageId="6" pageNumber="159" refString="Bechly, G. (2003) Phylogenetic systematics of Odonata. In: Schorr, M. & Lindeboom, M. (Eds.), Dragonfly Research, Folume 1. Zerf-Tubingen (CD-ROM)." type="book chapter" year="2003">Bechly 2003</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136539FFBAFFB5FF123551FBF811DB" pageId="6" pageNumber="159" type="description">
|
||
<paragraph id="8BB636B2FFBAFFB5FF123551FCD21193" blockId="6.[151,1437,151,1257]" pageId="6" pageNumber="159">
|
||
According to the slight differences in wing morphology and the large distance between the two fossil sites it seems not very likely that
|
||
<taxonomicName id="4C094D31FFBAFFB5FE1F350AFD09114F" box="[458,722,1124,1148]" class="Insecta" family="Gomphidae" genus="Ictinogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="159" phylum="Arthropoda" rank="species" species="hassleri">
|
||
<emphasis id="B97DEAA0FFBAFFB5FE1F350AFD09114F" box="[458,722,1124,1148]" italics="true" pageId="6" pageNumber="159">Ictinogomphus hassleri</emphasis>
|
||
</taxonomicName>
|
||
is truly conspecific but is closely related to the undetermined
|
||
<taxonomicName id="4C094D31FFBAFFB5FF4235E9FEC61193" box="[151,285,1159,1184]" class="Insecta" family="Gomphidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="159" phylum="Arthropoda" rank="subFamily" subFamily="Lindeniinae">Lindeniinae</taxonomicName>
|
||
fossil from
|
||
<collectingCountry id="F31E7622FFBAFFB5FE7035E9FE2E1193" box="[421,501,1159,1184]" name="Turkey" pageId="6" pageNumber="159">Turkey</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF984B43FFBAFFB5FDD635E9FD271193" author="Nel" box="[515,764,1159,1184]" pageId="6" pageNumber="159" refString="Nel, A. & Paicheler, J. C. (1994 a) Les Gomphidae fossiles. Un inventaire critique (Odonata: Gomphidae). Annales de la Societe entomologique de France, 30 (1), 55 - 77." type="journal article" year="1994">Nel & Paicheler 1994</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFBAFFB5FF1235C2FBF811DB" blockId="6.[151,1437,151,1257]" pageId="6" pageNumber="159">
|
||
Although our specimen resembles modern species of both
|
||
<taxonomicName id="4C094D31FFBAFFB5FCB035C2FBD011F7" box="[869,1035,1196,1220]" class="Insecta" family="Gomphidae" genus="Ictinogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="159" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97DEAA0FFBAFFB5FCB035C2FBD011F7" box="[869,1035,1196,1220]" italics="true" pageId="6" pageNumber="159">Ictinogomphus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C094D31FFBAFFB5FB9335C2FAD411F7" box="[1094,1295,1196,1220]" class="Insecta" family="Gomphidae" genus="Sinictinogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="159" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97DEAA0FFBAFFB5FB9335C2FAD411F7" box="[1094,1295,1196,1220]" italics="true" pageId="6" pageNumber="159">Sinictinogomphus</emphasis>
|
||
</taxonomicName>
|
||
, we suggest
|
||
<taxonomicName id="4C094D31FFBAFFB5FF4235BEFEE611DB" box="[151,317,1232,1256]" class="Insecta" family="Gomphidae" genus="Ictinogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="159" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97DEAA0FFBAFFB5FF4235BEFEE611DB" box="[151,317,1232,1256]" italics="true" pageId="6" pageNumber="159">Ictinogomphus</emphasis>
|
||
</taxonomicName>
|
||
as the most suitable genus name due to the older age of the name.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |